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<document id="EB3B784DADEED8DF1D06F185F72C55B5" ID-ISSN="1175-5326" ID-ZooBank="9913C60F-5F0D-42ED-A5E6-109EB5156B4F" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629832609958" checkinUser="felipe" docAuthor="Behounek, G., Han, H. L. &amp; V. S. Kononenko" docDate="2012" docId="0F360A170A12FFEAA8810E75E6CEFC8A" docLanguage="en" docName="zt03587p088.pdf" docOrigin="Zootaxa 3587" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Flavala secunda Behounek, Han &amp; Kononenko 2012, sp. n." docType="treatment" docVersion="3" lastPageNumber="82" masterDocId="F30F726F0A11FFEEA8160C2FE415FFA7" masterDocTitle="Two new genera and three new species of the Pantheinae from East Asia and China (Lepidoptera, Noctuidae). Contribution to revision of Pantheinae VIII" masterLastPageNumber="88" masterPageNumber="78" pageNumber="81" updateTime="1699118637777" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="623710D4BCD90FCF52E07B5747D6BCDC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E14EA4C0400D85A4F955206081D6F629">
<mods:title id="DAE6F6539D2AD061E1DF0B686EED6EC1">Two new genera and three new species of the Pantheinae from East Asia and China (Lepidoptera, Noctuidae). Contribution to revision of Pantheinae VIII</mods:title>
</mods:titleInfo>
<mods:name id="B91C5A72091FC89B9634932E0BC432B7" type="personal">
<mods:role id="8D135B6F56BE9B48823A3DEE223D7CF2">
<mods:roleTerm id="E6C27C20A8C5DA1C961AD8FD9091C3AB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B15BC780AB64C58DB32792E8C8FDDBA7">Behounek, G.</mods:namePart>
<mods:affiliation id="A7875F57A2AA9BC62178B0B412EF20AC">D- 85567 Grafing / Munich, Sudetenstrasse 6, Germany. E-mail: Gottfried. Behounek @ t-online. de</mods:affiliation>
</mods:name>
<mods:name id="520E4AAE2C6610275F6729307BF32DDE" type="personal">
<mods:role id="8AA15B150D5D2F1CD7562B5933D994F3">
<mods:roleTerm id="2BED1DF9C2EADA5E6B74575F44CB2F78">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EF267587DB6FB9F152583BC33554198E">Han, H. L.</mods:namePart>
<mods:affiliation id="769EA2ED491DF67472AA7A8E278721DC">School of Forestry, Northeast Forestry University, Harbin, CH- 150014 China. E-mail: hanhuilin @ yahoo. com. cn</mods:affiliation>
</mods:name>
<mods:name id="2C3C41EAFBF8518E4741D4B175CC7EA2" type="personal">
<mods:role id="E9CAF3F9113B3992585D3BFDA301E41C">
<mods:roleTerm id="58743F99CF5D45A481E7F645B0F8ACB2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="93F9766977691FF991E8CEA0957C9F9B">V. S. Kononenko</mods:namePart>
<mods:affiliation id="E55D4B32AD7743FC45974BDB936106EC">Laboratory of Entomology, Institute of Biology and Soil Science Far Eastern Branch of Russian Academy of Sciences, RF- 690022 Vladivostok, Russia. E-mail: kononenko @ ibss. dvo. ru, vlad _ kononenko @ mail. ru</mods:affiliation>
</mods:name>
<mods:typeOfResource id="58201F01DB5F1E744EAEE58AB2553241">text</mods:typeOfResource>
<mods:relatedItem id="844CF2EE7A090DEFD5F23B4F6D723778" type="host">
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<mods:title id="0952BF0F848CB8A12D3E86A01321EECC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="486EC9550641BF2C1B530A02C38A0ED3">
<mods:date id="416FBF79304CB0FE2527E0FB352B5F02">2012</mods:date>
<mods:detail id="4C574485172D308E5B8398C09E1FA7E6" type="pubDate">
<mods:number id="F0F2EF7021DBFC7D96F61826C0E27C35">2012-12-17</mods:number>
</mods:detail>
<mods:detail id="73C4D480D409C814B140393FA263C966" type="volume">
<mods:number id="6F0F4077EEDA6FA73840B35013954AE0">3587</mods:number>
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<mods:start id="93AB5D558D10D36338FF284A0E80C541">78</mods:start>
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<mods:classification id="EC59763CFDDD3200E82D081E586320C5">journal article</mods:classification>
<mods:identifier id="0C9ABBE82C08A1A20CA0633625E7D9C3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6D8BA714F2FAC50EE3C5C40B59977C6A" type="ZooBank">9913C60F-5F0D-42ED-A5E6-109EB5156B4F</mods:identifier>
</mods:mods>
<treatment id="0F360A170A12FFEAA8810E75E6CEFC8A" ID-DOI="http://doi.org/10.5281/zenodo.5259260" ID-Zenodo-Dep="5259260" LSID="urn:lsid:plazi:treatment:0F360A170A12FFEAA8810E75E6CEFC8A" httpUri="http://treatment.plazi.org/id/0F360A170A12FFEAA8810E75E6CEFC8A" lastPageId="4" lastPageNumber="82" pageId="3" pageNumber="81">
<subSubSection id="CF85E88A0A12FFEDA8810E75E725FDD3" box="[151,816,602,628]" pageId="3" pageNumber="81" type="nomenclature">
<paragraph id="8720BB010A12FFEDA8810E75E725FDD3" blockId="3.[151,816,602,661]" box="[151,816,602,628]" pageId="3" pageNumber="81">
<heading id="DC680C6D0A12FFEDA8810E75E725FDD3" bold="true" box="[151,816,602,628]" fontSize="11" level="1" pageId="3" pageNumber="81" reason="1">
<emphasis id="B5EB67130A12FFEDA8810E75E725FDD3" bold="true" box="[151,816,602,628]" pageId="3" pageNumber="81">
<taxonomicName id="409FC0820A12FFEDA8810E75E6F5FDD3" authority="Behounek, Han &amp; Kononenko" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[151,736,602,628]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="secunda" status="sp. nov.">
<emphasis id="B5EB67130A12FFEDA8810E75E54BFDD3" bold="true" box="[151,350,602,628]" italics="true" pageId="3" pageNumber="81">Flavala secunda</emphasis>
Behounek, Han &amp; Kononenko
</taxonomicName>
,
<taxonomicNameLabel id="AED8DA680A12FFEDAAF80E75E725FDD3" box="[750,816,602,628]" pageId="3" pageNumber="81" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CF85E88A0A12FFEDA8880E52E556FD32" box="[158,323,637,661]" pageId="3" pageNumber="81" type="description">
<paragraph id="8720BB010A12FFEDA8880E52E556FD32" blockId="3.[151,816,602,661]" box="[158,323,637,661]" pageId="3" pageNumber="81">
(
<figureCitation id="1FA4A7840A12FFEDA8B00E52E505FD32" box="[166,272,637,661]" captionStart="FIGURES 1–8" captionStartId="6.[151,263,1841,1864]" captionTargetBox="[151,1436,193,1821]" captionTargetId="figure-84@6.[151,1436,193,1821]" captionTargetPageId="6" captionText="FIGURES 1–8. Imagines. 1. Anacronicta caliginea, female, Russia, Primorye; 2. A. nitida, male, Japan; 3. Flavala flavala, male, North India, genit. prep. 7421; 4. F. flavala, male, underside, Nepal; 5. F. crypta, male, holotype, China, West Sichuan, genit. prep. 8031; 6. F. crypta, male, underside, paratype, North Vietnam; 7. F. secunda, male, holotype, North Vietnam, genit. prep. 7419; 8. Ditto, underside." pageId="3" pageNumber="81">Figs. 7, 8</figureCitation>
,
<figureCitation id="1FA4A7840A12FFEDA90B0E52E52FFD32" box="[285,314,637,661]" captionStart="FIGURES 14–16" captionStartId="8.[151,263,1841,1864]" captionTargetBox="[151,1418,193,1813]" captionTargetId="figure-55@8.[151,1436,193,1821]" captionTargetPageId="8" captionText="FIGURES 14–16. Flavala spp., male genitalia, details of structure (A—uncus, B—cornuti). 14. Flavala flavala, Myanmar, genit. prep. 7420GB; 15. F. crypta, holotype, West China, genit. prep. 8031GB; 16. F. secunda, holotype, North Vietnam, genit. prep. 7419GB." pageId="3" pageNumber="81">16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CF85E88A0A12FFEDA8810EEDE5D0FC20" pageId="3" pageNumber="81" type="materials_examined">
<paragraph id="8720BB010A12FFEDA8810EEDE5D0FC20" blockId="3.[151,1437,706,2014]" pageId="3" pageNumber="81">
<emphasis id="B5EB67130A12FFEDA8810EEDE595FD7C" bold="true" box="[151,384,706,731]" inLineHeading="true" pageId="3" pageNumber="81" reason="1">Material examined.</emphasis>
<materialsCitation id="37F7B15C0A12FFEDA9900EEDE113FD5A" collectingDate="2008-08" collectionCode="ZSM" collectorName="Behounek &amp; Munchen &amp; Lep" country="Vietnam" county="North" elevation="2400" location="Mt. Fan Si Pan" municipality="Sa Pa" pageId="3" pageNumber="81" specimenCount="1" stateProvince="Lao Cai" typeStatus="holotype">
<typeStatus id="582405A30A12FFEDA9900EEDE5E6FD7D" box="[390,499,706,730]" pageId="3" pageNumber="81" type="holotype">Holotype</typeStatus>
: male,
<collectingCounty id="6E41C38D0A12FFEDAA540EEDE696FD7D" box="[578,643,706,730]" pageId="3" pageNumber="81">North</collectingCounty>
<collectingCountry id="FF88FB910A12FFEDAA9C0EEDE6F9FD7D" box="[650,748,706,730]" name="Vietnam" pageId="3" pageNumber="81">Vietnam</collectingCountry>
, Prov.
<collectingRegion id="455B75E30A12FFEDAB2E0EECE784FD7D" box="[824,913,706,730]" country="Vietnam" name="Lao Cai" pageId="3" pageNumber="81">Lao Cai</collectingRegion>
/
<collectingMunicipality id="6744217B0A12FFEDAB8C0EEDE7CEFD7D" box="[922,987,706,730]" pageId="3" pageNumber="81">Sa Pa</collectingMunicipality>
,
<location id="8240EDDA0A12FFEDABF00EECE085FD7D" LSID="urn:lsid:plazi:treatment:0F360A170A12FFEAA8810E75E6CEFC8A:8240EDDA0A12FFEDABF00EECE085FD7D" box="[998,1168,706,730]" country="Vietnam" county="North" municipality="Sa Pa" name="Mt. Fan Si Pan" pageId="3" pageNumber="81" stateProvince="Lao Cai">Mt. Fan Si Pan</location>
,
<quantity id="406716E40A12FFEDAC8D0EEDE0E7FD7D" box="[1179,1266,706,730]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="81" unit="m" value="2400.0">
<elevation id="0CB25C320A12FFEDAC8D0EEDE0E7FD7D" box="[1179,1266,706,730]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="81" unit="m" value="2400.0">2400 m</elevation>
</quantity>
,
<date id="F3219DC10A12FFEDACE80EEDE177FD7D" box="[1278,1378,706,730]" pageId="3" pageNumber="81" value="2008-08">
<collectingDate id="E36564290A12FFEDACE80EEDE177FD7D" box="[1278,1378,706,730]" pageId="3" pageNumber="81" value="2008-08">viii.2008</collectingDate>
</date>
(leg. local coll., ex coll Becher), coll.
<collectorName id="2A6ADED70A12FFEDAA1B0ECAE694FD5A" box="[525,641,741,765]" pageId="3" pageNumber="81">Behounek</collectorName>
,
<collectionCode id="E18E23C40A12FFEDAA9B0ECAE6D3FD5A" box="[653,710,741,765]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="81" type="Museum">ZSM</collectionCode>
<collectorName id="2A6ADED70A12FFEDAAD90ECAE729FD5A" box="[719,828,741,765]" pageId="3" pageNumber="81">München</collectorName>
; genit. prep. 7419, BC
<collectionCode id="E18E23C40A12FFEDAC510ECAE095FD5A" box="[1095,1152,741,765]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="81" type="Museum">ZSM</collectionCode>
<collectorName id="2A6ADED70A12FFEDAC9E0EC9E0A6FD5A" box="[1160,1203,742,765]" pageId="3" pageNumber="81">Lep</collectorName>
48680
</materialsCitation>
.
<materialsCitation id="37F7B15C0A12FFEDAD050EC9E73FFCB8" collectingDate="2008-08" collectionCode="ZSM" collectorName="Becher" country="Vietnam" county="North" elevation="2400" location="Mt. Fan Si Pan" municipality="Sa Pa" pageId="3" pageNumber="81" specimenCount="2" specimenCount-male="2" stateProvince="Lao Cai" typeStatus="paratype">
<typeStatus id="582405A30A12FFEDAD050EC9E196FD59" box="[1299,1411,742,766]" pageId="3" pageNumber="81" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="919970880A12FFEDAD980ECAE4CEFCB8" pageId="3" pageNumber="81" type="male">2 males</specimenCount>
, same locality and collector,
<date id="F3219DC10A12FFEDAA320F28E686FCB8" box="[548,659,775,799]" pageId="3" pageNumber="81" value="2008-08">
<collectingDate id="E36564290A12FFEDAA320F28E686FCB8" box="[548,659,775,799]" pageId="3" pageNumber="81" value="2008-08">viii. 2008</collectingDate>
</date>
, coll.
<collectorName id="2A6ADED70A12FFEDAAC10F28E73FFCB8" box="[727,810,775,799]" pageId="3" pageNumber="81">Becher</collectorName>
</materialsCitation>
;
<materialsCitation id="37F7B15C0A12FFEDAB200F28E5E0FCC2" collectingDate="1995-04" collectorName="Sinyaev &amp; Simonov &amp; A. Schintlmeister &amp; Museum Witt" country="Vietnam" county="Mt" elevation="1700" location="Cha Pa" municipality="West" pageId="3" pageNumber="81" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="919970880A12FFEDAB200F28E790FCB8" box="[822,901,775,799]" pageId="3" pageNumber="81" type="male">1 male</specimenCount>
, N
<collectingCountry id="FF88FB910A12FFEDABBB0F28E01AFCB8" box="[941,1039,775,799]" name="Vietnam" pageId="3" pageNumber="81">Vietnam</collectingCountry>
,
<collectingCounty id="6E41C38D0A12FFEDAC0C0F27E02FFCB8" box="[1050,1082,776,799]" pageId="3" pageNumber="81">Mt</collectingCounty>
Fan-si-pan,
<collectingMunicipality id="6744217B0A12FFEDACDF0F27E115FCB8" box="[1225,1280,776,799]" pageId="3" pageNumber="81">West</collectingMunicipality>
side,
<location id="8240EDDA0A12FFEDAD520F28E18DFCB8" LSID="urn:lsid:plazi:treatment:0F360A170A12FFEAA8810E75E6CEFC8A:8240EDDA0A12FFEDAD520F28E18DFCB8" box="[1348,1432,775,799]" country="Vietnam" county="Mt" municipality="West" name="Cha Pa" pageId="3" pageNumber="81">Cha Pa</location>
, 22˚20’N 103˚40’E,
<quantity id="406716E40A12FFEDA96B0F05E60AFCE5" box="[381,543,810,835]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="3" pageNumber="81" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">
<elevation id="0CB25C320A12FFEDA96B0F05E60AFCE5" box="[381,543,810,835]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="3" pageNumber="81" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">
1600
<emphasis id="B5EB67130A12FFEDA9A30F04E5D6FCE4" bold="true" box="[437,451,811,835]" pageId="3" pageNumber="81">–</emphasis>
1800 m
</elevation>
</quantity>
,
<date id="F3219DC10A12FFEDAA390F05E690FCE5" box="[559,645,810,834]" pageId="3" pageNumber="81" value="1995-04">
<collectingDate id="E36564290A12FFEDAA390F05E690FCE5" box="[559,645,810,834]" pageId="3" pageNumber="81" value="1995-04">iv.1995</collectingDate>
</date>
, leg.
<collectorName id="2A6ADED70A12FFEDAADF0F05E736FCE5" box="[713,803,810,834]" pageId="3" pageNumber="81">Sinyaev</collectorName>
&amp;
<collectorName id="2A6ADED70A12FFEDAB460F05E7A3FCE5" box="[848,950,810,834]" pageId="3" pageNumber="81">Simonov</collectorName>
, ex coll.
<collectorName id="2A6ADED70A12FFEDAC3E0F05E0F3FCE5" box="[1064,1254,810,834]" pageId="3" pageNumber="81">A.Schintlmeister</collectorName>
,
<collectorName id="2A6ADED70A12FFEDACE00F04E181FCE5" box="[1270,1428,810,834]" pageId="3" pageNumber="81">Museum Witt</collectorName>
/ genit. prep. 7331GB (coll. GB)
</materialsCitation>
;
<materialsCitation id="37F7B15C0A12FFEDA9E90F62E5D4FC20" collectingDate="2010-03" collectionCode="T, AB" collectorName="Coll. Becher" country="Vietnam" county="North" elevation="2300" location="Mt. Fan Si Pan" municipality="Sa Pa" pageId="3" pageNumber="81" specimenCount="1" specimenCount-male="1" stateProvince="Lao Cai" typeStatus="paratype">
<specimenCount id="919970880A12FFEDA9E90F62E658FCC2" box="[511,589,845,869]" pageId="3" pageNumber="81" type="male">1 male</specimenCount>
,
<collectingCounty id="6E41C38D0A12FFEDAA410F62E68DFCC2" box="[599,664,845,869]" pageId="3" pageNumber="81">North</collectingCounty>
<collectingCountry id="FF88FB910A12FFEDAA890F62E714FCC2" box="[671,769,845,869]" name="Vietnam" pageId="3" pageNumber="81">Vietnam</collectingCountry>
, Prov.
<collectingRegion id="455B75E30A12FFEDAB5A0F61E7B0FCC2" box="[844,933,845,869]" country="Vietnam" name="Lao Cai" pageId="3" pageNumber="81">Lao Cai</collectingRegion>
/
<collectingMunicipality id="6744217B0A12FFEDABBB0F62E7F8FCC2" box="[941,1005,845,869]" pageId="3" pageNumber="81">Sa Pa</collectingMunicipality>
,
<location id="8240EDDA0A12FFEDABEE0F61E0B4FCC2" LSID="urn:lsid:plazi:treatment:0F360A170A12FFEAA8810E75E6CEFC8A:8240EDDA0A12FFEDABEE0F61E0B4FCC2" box="[1016,1185,845,869]" country="Vietnam" county="North" municipality="Sa Pa" name="Mt. Fan Si Pan" pageId="3" pageNumber="81" stateProvince="Lao Cai">Mt. Fan Si Pan</location>
,
<quantity id="406716E40A12FFEDACBD0F62E114FCC2" box="[1195,1281,845,869]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="81" unit="m" value="2300.0">
<elevation id="0CB25C320A12FFEDACBD0F62E114FCC2" box="[1195,1281,845,869]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="81" unit="m" value="2300.0">2300 m</elevation>
</quantity>
,
<date id="F3219DC10A12FFEDAD1B0F62E176FCC2" box="[1293,1379,845,869]" pageId="3" pageNumber="81" value="2010-03">
<collectingDate id="E36564290A12FFEDAD1B0F62E176FCC2" box="[1293,1379,845,869]" pageId="3" pageNumber="81" value="2010-03">iii.2010</collectingDate>
</date>
(leg.
<collectionCode id="E18E23C40A12FFEDA8810F5FE4BCFC20" box="[151,169,880,903]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="81">T</collectionCode>
. Ihle)
<collectorName id="2A6ADED70A12FFEDA8E60F40E568FC20" box="[240,381,879,903]" pageId="3" pageNumber="81">Coll. Becher</collectorName>
(
<collectionCode id="E18E23C40A12FFEDA99B0F40E5ACFC20" box="[397,441,879,903]" pageId="3" pageNumber="81">AB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CF85E88A0A12FFEDA8D10FBDE58BFA28" pageId="3" pageNumber="81" type="diagnosis">
<paragraph id="8720BB010A12FFEDA8D10FBDE58BFA28" blockId="3.[151,1437,706,2014]" pageId="3" pageNumber="81">
<emphasis id="B5EB67130A12FFEDA8D10FBDE554FC0C" bold="true" box="[199,321,914,939]" inLineHeading="true" pageId="3" pageNumber="81" reason="1">Diagnosis.</emphasis>
The new species is the second hidden species in the
<taxonomicName id="409FC0820A12FFEDABF00FBCE04DFC0C" authorityName="flavala (Moore" authorityYear="1867" baseAuthorityName="flavala (Moore" baseAuthorityYear="1867" box="[998,1112,915,939]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="flavala">
<emphasis id="B5EB67130A12FFEDABF00FBCE04DFC0C" box="[998,1112,915,939]" italics="true" pageId="3" pageNumber="81">F. flavala</emphasis>
</taxonomicName>
group. Externally it looks intermediate between
<taxonomicName id="409FC0820A12FFEDA9980F99E5EDFC6A" authorityName="flavala (Moore" authorityYear="1867" baseAuthorityName="flavala (Moore" baseAuthorityYear="1867" box="[398,504,949,973]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="flavala">
<emphasis id="B5EB67130A12FFEDA9980F99E5EDFC6A" box="[398,504,949,973]" italics="true" pageId="3" pageNumber="81">F. flavala</emphasis>
</taxonomicName>
and
<taxonomicName id="409FC0820A12FFEDAA390F99E687FC6A" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[559,658,950,973]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDAA390F99E687FC6A" box="[559,658,950,973]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
, but more similar to the latter. Ground colour of forewing darker and somewhat more yellowish than in
<taxonomicName id="409FC0820A12FFEDAA0C0FF6E66BFC57" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[538,638,985,1008]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDAA0C0FF6E66BFC57" box="[538,638,985,1008]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
(ash-grey in
<taxonomicName id="409FC0820A12FFEDAB020FF6E76DFC57" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[788,888,985,1008]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDAB020FF6E76DFC57" box="[788,888,985,1008]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
, dark yellowish-grey in
<taxonomicName id="409FC0820A12FFEDAC9A0FF6E0E0FC57" authorityName="flavala (Moore" authorityYear="1867" baseAuthorityName="flavala (Moore" baseAuthorityYear="1867" box="[1164,1269,984,1008]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="flavala">
<emphasis id="B5EB67130A12FFEDAC9A0FF6E0E0FC57" box="[1164,1269,984,1008]" italics="true" pageId="3" pageNumber="81">F. flavala</emphasis>
</taxonomicName>
); wing pattern more coarse. Antemedial line incurved inwardly (in
<taxonomicName id="409FC0820A12FFEDAAF50FD4E752FBB5" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[739,839,1019,1042]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDAAF50FD4E752FBB5" box="[739,839,1019,1042]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
straight); postmedial line more diffused, less distinct than in
<taxonomicName id="409FC0820A12FFEDA8FB0831E545FB92" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[237,336,1054,1077]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDA8FB0831E545FB92" box="[237,336,1054,1077]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
, subterminal line more distinct than in
<taxonomicName id="409FC0820A12FFEDAB190831E766FB92" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[783,883,1054,1077]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDAB190831E766FB92" box="[783,883,1054,1077]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
, but less than in
<taxonomicName id="409FC0820A12FFEDAC200831E0B4FB92" authorityName="flavala (Moore" authorityYear="1867" baseAuthorityName="flavala (Moore" baseAuthorityYear="1867" box="[1078,1185,1053,1077]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="flavala">
<emphasis id="B5EB67130A12FFEDAC200831E0B4FB92" box="[1078,1185,1053,1077]" italics="true" pageId="3" pageNumber="81">F. flavala</emphasis>
</taxonomicName>
. Abdomen yellowishgrey (ach-grey in
<taxonomicName id="409FC0820A12FFEDA94B086EE5AAFBFF" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[349,447,1089,1112]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDA94B086EE5AAFBFF" box="[349,447,1089,1112]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
). Hindwing ground colour more yellowish than in
<taxonomicName id="409FC0820A12FFEDABE0086EE04CFBFF" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[1014,1113,1089,1112]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDABE0086EE04CFBFF" box="[1014,1113,1089,1112]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
, but less yellowish than in
<taxonomicName id="409FC0820A12FFEDAD91086EE4F0FBDC" authorityName="flavala (Moore" authorityYear="1867" baseAuthorityName="flavala (Moore" baseAuthorityYear="1867" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="flavala">
<emphasis id="B5EB67130A12FFEDAD91086EE4F0FBDC" italics="true" pageId="3" pageNumber="81">F. flavala</emphasis>
</taxonomicName>
; less suffused with grey compared with both species; terminal band lies somewhat remote from the wing margin separated by a pale greyish suffusion (in
<taxonomicName id="409FC0820A12FFEDAADF08A9E73AFB3A" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[713,815,1158,1181]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDAADF08A9E73AFB3A" box="[713,815,1158,1181]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
it is almost adjacent to the wing margin, in
<taxonomicName id="409FC0820A12FFEDAD2608A9E189FB3A" authorityName="flavala (Moore" authorityYear="1867" baseAuthorityName="flavala (Moore" baseAuthorityYear="1867" box="[1328,1436,1157,1181]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="flavala">
<emphasis id="B5EB67130A12FFEDAD2608A9E189FB3A" box="[1328,1436,1157,1181]" italics="true" pageId="3" pageNumber="81">F. flavala</emphasis>
</taxonomicName>
clearly separated from wing margin by a distinct yellow field. Underside of wings more yellowish than in
<taxonomicName id="409FC0820A12FFEDAD240886E181FB67" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[1330,1428,1193,1216]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDAD240886E181FB67" box="[1330,1428,1193,1216]" italics="true" pageId="3" pageNumber="81">F.crypta</emphasis>
</taxonomicName>
, with less developed dark elements of pattern: forewing with narrower terminal band and less expressed discal spot in comparison to
<taxonomicName id="409FC0820A12FFEDA94D08C1E5D0FAA2" authorityName="flavala (Moore" authorityYear="1867" baseAuthorityName="flavala (Moore" baseAuthorityYear="1867" box="[347,453,1261,1285]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="flavala">
<emphasis id="B5EB67130A12FFEDA94D08C1E5D0FAA2" box="[347,453,1261,1285]" italics="true" pageId="3" pageNumber="81">F. flavala</emphasis>
</taxonomicName>
and
<taxonomicName id="409FC0820A12FFEDA9EA08C1E64AFAA2" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[508,607,1262,1285]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDA9EA08C1E64AFAA2" box="[508,607,1262,1285]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
; hindwing with weak diffused terminal band and small discal spot.,Male genitalia almost identical with
<taxonomicName id="409FC0820A12FFEDA9E7093EE64FFA8F" authorityName="flavala (Moore" authorityYear="1867" baseAuthorityName="flavala (Moore" baseAuthorityYear="1867" box="[497,602,1296,1320]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="flavala">
<emphasis id="B5EB67130A12FFEDA9E7093EE64FFA8F" box="[497,602,1296,1320]" italics="true" pageId="3" pageNumber="81">F. flavala</emphasis>
</taxonomicName>
, and
<taxonomicName id="409FC0820A12FFEDAA8E093EE6EFFA8F" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[664,762,1297,1320]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDAA8E093EE6EFFA8F" box="[664,762,1297,1320]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
, however, in
<taxonomicName id="409FC0820A12FFEDAB85093EE7E3FA8F" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[915,1014,1297,1320]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDAB85093EE7E3FA8F" box="[915,1014,1297,1320]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
, the cucullus is longer and narrow, in
<taxonomicName id="409FC0820A12FFEDA881091CE518FAEC" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[151,269,1331,1355]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="secunda">
<emphasis id="B5EB67130A12FFEDA881091CE518FAEC" box="[151,269,1331,1355]" italics="true" pageId="3" pageNumber="81">F. secunda</emphasis>
</taxonomicName>
it is shorter and broader, but not as broad as in
<taxonomicName id="409FC0820A12FFEDAB09091CE79DFAEC" authorityName="flavala (Moore" authorityYear="1867" baseAuthorityName="flavala (Moore" baseAuthorityYear="1867" box="[799,904,1331,1355]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="flavala">
<emphasis id="B5EB67130A12FFEDAB09091CE79DFAEC" box="[799,904,1331,1355]" italics="true" pageId="3" pageNumber="81">F. flavala</emphasis>
</taxonomicName>
. Distal end of valva, after saccular extension in
<taxonomicName id="409FC0820A12FFEDA8810979E4EEFACA" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[151,251,1366,1389]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B5EB67130A12FFEDA8810979E4EEFACA" box="[151,251,1366,1389]" italics="true" pageId="3" pageNumber="81">F. crypta</emphasis>
</taxonomicName>
narrow, have more or less tapered, pointed ending. In
<taxonomicName id="409FC0820A12FFEDAB770979E7CDFACA" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[865,984,1365,1389]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="secunda">
<emphasis id="B5EB67130A12FFEDAB770979E7CDFACA" box="[865,984,1365,1389]" italics="true" pageId="3" pageNumber="81">F. secunda</emphasis>
</taxonomicName>
the end of valva is more blunt, but in
<taxonomicName id="409FC0820A12FFEDAD910979E4F0FA37" authorityName="flavala (Moore" authorityYear="1867" baseAuthorityName="flavala (Moore" baseAuthorityYear="1867" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="flavala">
<emphasis id="B5EB67130A12FFEDAD910979E4F0FA37" italics="true" pageId="3" pageNumber="81">F. flavala</emphasis>
</taxonomicName>
it is more blunt.
</paragraph>
</subSubSection>
<subSubSection id="CF85E88A0A12FFEDA8D109B5E67BF830" pageId="3" pageNumber="81" type="description">
<paragraph id="8720BB010A12FFEDA8D109B5E67BF830" blockId="3.[151,1437,706,2014]" pageId="3" pageNumber="81">
<emphasis id="B5EB67130A12FFEDA8D109B5E54DFA14" bold="true" box="[199,344,1434,1459]" inLineHeading="true" pageId="3" pageNumber="81" reason="1">Description.</emphasis>
Adult (
<figureCitation id="1FA4A7840A12FFEDA9A309B5E632FA15" box="[437,551,1434,1458]" captionStart="FIGURES 1–8" captionStartId="6.[151,263,1841,1864]" captionTargetBox="[151,1436,193,1821]" captionTargetId="figure-84@6.[151,1436,193,1821]" captionTargetPageId="6" captionText="FIGURES 1–8. Imagines. 1. Anacronicta caliginea, female, Russia, Primorye; 2. A. nitida, male, Japan; 3. Flavala flavala, male, North India, genit. prep. 7421; 4. F. flavala, male, underside, Nepal; 5. F. crypta, male, holotype, China, West Sichuan, genit. prep. 8031; 6. F. crypta, male, underside, paratype, North Vietnam; 7. F. secunda, male, holotype, North Vietnam, genit. prep. 7419; 8. Ditto, underside." pageId="3" pageNumber="81">Figs. 7, 8</figureCitation>
). Wingspan
<quantity id="406716E40A12FFEDAAAD09B5E72FFA15" box="[699,826,1434,1459]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.1" metricValueMin="3.8" pageId="3" pageNumber="81" unit="mm" value="39.5" valueMax="41.0" valueMin="38.0">
38
<emphasis id="B5EB67130A12FFEDAAC109B4E6F0FA14" bold="true" box="[727,741,1435,1459]" pageId="3" pageNumber="81">–</emphasis>
41 mm
</quantity>
. Head and thorax covered with short whitish scales; vestiture of thorax formed by hair-like scales; patagia and tegulae whitish; ground colour of forewing yellowishbrown-grey, wing pattern formed by blackish lines; basal field, costal area and space behind reniform with whitish suffusion; subbasal field brownish-grey, with whitish suffusion; antemedial line black, distinct, slightly incurved outwardly; medial field yellowish brown-grey, with indistinct diffused medial shadow, darker in costal area; orbicular very small, as whitish spot; reniform distinct, formed by two whitish spots and black spot between, with dark surrounding line; postmedial line blackish, thin, dentate, diffused, costal area behind reniform and around postmedial line with whitish suffusion; subterminal field brownish-grey; subterminal line dark, diffused, surrounded with thin outer whitish line; terminal field brownish grey; terminal line as row of dark diffused streaks; cilia greyish-brown. Hindwing pale yellowish-grey, with weak greyish suffusion along inner margin, indistinct discal spot and diffused terminal band along outer margin, separated from margin by yellowish-grey field; cilia pale, yellowish-grey, dark opposite veins. Underside pale yellowish, with brownish-grey terminal band and blackish suffusion around distal part of discal cell on forewing and wide greyish-brown terminal band distantly along outer margin and discal mark on hindwing. Male genitalia (
<figureCitation id="1FA4A7840A12FFEDAB9B0B72E7F4F8D2" box="[909,993,1885,1909]" captionStart="FIGURES 14–16" captionStartId="8.[151,263,1841,1864]" captionTargetBox="[151,1418,193,1813]" captionTargetId="figure-55@8.[151,1436,193,1821]" captionTargetPageId="8" captionText="FIGURES 14–16. Flavala spp., male genitalia, details of structure (A—uncus, B—cornuti). 14. Flavala flavala, Myanmar, genit. prep. 7420GB; 15. F. crypta, holotype, West China, genit. prep. 8031GB; 16. F. secunda, holotype, North Vietnam, genit. prep. 7419GB." pageId="3" pageNumber="81">Fig. 16</figureCitation>
) as described for the genus and in the paragraph “Diagnosis”. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="CF85E88A0A12FFEDA8D10B8DE7D1F81D" box="[199,964,1954,1979]" pageId="3" pageNumber="81" type="distribution">
<paragraph id="8720BB010A12FFEDA8D10B8DE7D1F81D" blockId="3.[151,1437,706,2014]" box="[199,964,1954,1979]" pageId="3" pageNumber="81">
<emphasis id="B5EB67130A12FFEDA8D10B8DE575F81D" box="[199,352,1954,1979]" inLineHeading="true" pageId="3" pageNumber="81" reason="1">
<emphasis id="B5EB67130A12FFEDA8D10B8DE54CF81C" bold="true" box="[199,345,1954,1979]" pageId="3" pageNumber="81">Distribution</emphasis>
.
</emphasis>
(Map 22). North
<collectingCountry id="FF88FB910A12FFEDAA300B8DE69DF81D" box="[550,648,1954,1978]" name="Vietnam" pageId="3" pageNumber="81">Vietnam</collectingCountry>
, probably Southwest
<collectingCountry id="FF88FB910A12FFEDAB6F0B8DE7AAF81D" box="[889,959,1954,1978]" name="China" pageId="3" pageNumber="81">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CF85E88A0A12FFEAA8D10BEAE6CEFC8A" lastPageId="4" lastPageNumber="82" pageId="3" pageNumber="81" type="etymology">
<paragraph id="8720BB010A12FFEDA8D10BEAE117F87A" blockId="3.[151,1437,706,2014]" box="[199,1282,1989,2014]" pageId="3" pageNumber="81">
<emphasis id="B5EB67130A12FFEDA8D10BEAE558F87A" bold="true" box="[199,333,1989,2013]" inLineHeading="true" pageId="3" pageNumber="81" reason="1">Etymology.</emphasis>
The species name
<taxonomicName id="409FC0820A12FFEDAA340BEAE66BF87A" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[546,638,1989,2013]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="secunda">
<emphasis id="B5EB67130A12FFEDAA340BEAE66BF87A" box="[546,638,1989,2013]" italics="true" pageId="3" pageNumber="81">secunda</emphasis>
</taxonomicName>
means the second “hidden species” in the genus
<taxonomicName id="409FC0820A12FFEDACB20BEAE0EEF87A" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[1188,1275,1989,2013]" class="Insecta" family="Noctuidae" genus="Flavala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B5EB67130A12FFEDACB20BEAE0EEF87A" box="[1188,1275,1989,2013]" italics="true" pageId="3" pageNumber="81">Flavala</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8720BB010A15FFEAA8810F37E6CEFC8A" blockId="4.[151,731,792,814]" box="[151,731,792,814]" pageId="4" pageNumber="82">
<emphasis id="B5EB67130A15FFEAA8810F37E4F4FC8A" bold="true" box="[151,225,792,813]" pageId="4" pageNumber="82">TREE.</emphasis>
Diagram of DNA barcoding tree for
<taxonomicName id="409FC0820A15FFEAAA4A0F37E6CEFC8A" box="[604,731,792,814]" class="Insecta" family="Noctuidae" genus="Flavala" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="82" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B5EB67130A15FFEAAA4A0F37E6BEFC89" box="[604,683,792,814]" italics="true" pageId="4" pageNumber="82">Flavala</emphasis>
spp.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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