240 lines
39 KiB
XML
240 lines
39 KiB
XML
<document id="EFBA8EB56697429F0F41F459652F046B" ID-CLB-Dataset="8111" ID-DOI="10.37520/aemnp.2020.26" ID-GBIF-Dataset="1a61f2ea-5825-49cd-afeb-bb9d9f9be24a" ID-ISSN="1804-6487" ID-Zenodo-Dep="4489332" ID-ZooBank="7CF8BFF6-034F-475E-9A3A-1FA8AF862824" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612221451257" checkinUser="felipe" docAuthor="Zettel, Herbert & Laciny, Alice" docDate="2020" docId="03FC501AFFB96854FC1A4AF8FB86F980" docLanguage="en" docName="ActaEntMusNatPrag.60.2.419-425.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Rhagovelia kalalaoensis Zettel & Laciny 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="423" masterDocId="FFC52862FFBA6850FFA24F03FF83FFD5" masterDocTitle="Rhagovelia species from Nosy Boraha east of Madagascar (Hemiptera: Heteroptera: Veliidae)" masterLastPageNumber="425" masterPageNumber="419" pageNumber="422" updateTime="1698926184911" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="A52C36C90A2C1DF698EDE17F3C416FA7">Rhagovelia species from Nosy Boraha east of Madagascar (Hemiptera: Heteroptera: Veliidae)</mods:title>
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<mods:name id="D386748939588C475EDD98EF9D48B3DD" type="personal">
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<mods:namePart id="8F1651F4BB5F8672D1E89DE407671648">Zettel, Herbert</mods:namePart>
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<mods:namePart id="B02DD64B86C5F9311B6D82C8B19BD2E4">Laciny, Alice</mods:namePart>
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<mods:typeOfResource id="B23CD218B9D2A4CFE1E6744FEBBA4349">text</mods:typeOfResource>
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<mods:title id="F430FD48CB1F919DC9DA91299B45EE0F">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="9F6D249F5714039BA02C4219024655A9">
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<mods:date id="DE4278CAF91BE6C115697F98438DA54F">2020</mods:date>
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<mods:number id="0FF9AE83FB328AB6DBF100AA912D3726">2020-07-06</mods:number>
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<mods:classification id="A30ECD32891DB23C052A8842CBFA669F">journal article</mods:classification>
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<mods:identifier id="3AAD4DE6F1EEFCB75A955A3EEA7FA374" type="DOI">10.37520/aemnp.2020.26</mods:identifier>
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<treatment id="03FC501AFFB96854FC1A4AF8FB86F980" ID-DOI="http://doi.org/10.5281/zenodo.4550763" ID-GBIF-Taxon="177426402" ID-Zenodo-Dep="4550763" LSID="urn:lsid:plazi:treatment:03FC501AFFB96854FC1A4AF8FB86F980" httpUri="http://treatment.plazi.org/id/03FC501AFFB96854FC1A4AF8FB86F980" lastPageId="4" lastPageNumber="423" pageId="3" pageNumber="422">
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<subSubSection id="C34FB287FFB96853FC1A4AF8FAA3F9C6" box="[952,1312,1531,1555]" pageId="3" pageNumber="422" type="nomenclature">
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<paragraph id="8BEAE10CFFB96853FC1A4AF8FAA3F9C6" blockId="3.[952,1312,1531,1555]" box="[952,1312,1531,1555]" pageId="3" pageNumber="422">
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<heading id="D0A25660FFB96853FC1A4AF8FAA3F9C6" box="[952,1312,1531,1555]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="422" reason="3">
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<emphasis id="B9213D1EFFB96853FC1A4AF8FAA3F9C6" bold="true" box="[952,1312,1531,1555]" pageId="3" pageNumber="422">
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<taxonomicName id="4C559A8FFFB96853FC1A4AF8FB40F9C6" authority="Zettel & Laciny, 2020" authorityName="Zettel & Laciny" authorityYear="2020" box="[952,1219,1531,1555]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="kalalaoensis" status="sp. nov.">
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<emphasis id="B9213D1EFFB96853FC1A4AF8FB40F9C6" bold="true" box="[952,1219,1531,1555]" italics="true" pageId="3" pageNumber="422">Rhagovelia kalalaoensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2128065FFB96853FB684AFFFAA3F9C6" box="[1226,1312,1532,1555]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34FB287FFB96853FC574919FB60F9F8" box="[1013,1251,1562,1581]" pageId="3" pageNumber="422" type="description">
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<paragraph id="8BEAE10CFFB96853FC574919FB60F9F8" blockId="3.[828,1437,1562,1676]" box="[1013,1251,1562,1581]" pageId="3" pageNumber="422">
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(
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<figureCitation id="136EFD89FFB96853FC594919FBB1F9F8" box="[1019,1074,1562,1581]" captionStart="Figs 1–4. 1–2" captionStartId="2.[151,188,1715,1734]" captionTargetBox="[151,1436,189,1697]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 1–4. 1–2 – Rhagovelia madli sp. nov.: 1 – habitus of holotype, apterous male; 2 – paratype, apterous female; 3 – R. kalalaoensis sp. nov., holotype, apterous male; 4 – R. reuteri Hoberlandt, 1951, small apterous male with reduced metatibial tooth." figureDoi="http://doi.org/10.5281/zenodo.4489336" httpUri="https://zenodo.org/record/4489336/files/figure.png" pageId="3" pageNumber="422">Figs 3</figureCitation>
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,
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<figureCitation id="136EFD89FFB96853FB9E4919FB1EF9F8" box="[1084,1181,1562,1581]" captionStart="Figs 5–12" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 5–12. Characteristics of males: 5–6 – right foreleg: 5 – Rhagovelia madli sp. nov., 6 – R. kalalaoensis sp. nov.; 7–8 – right hind leg: 7 – R. madli sp. nov. 8 – R. kalalaoensis sp. nov.; 9–10 – pygophore, dorsal view: 9 – R. madli sp. nov. 10 – R. kalalaoensis sp. nov.; 11–12 – left paramere, lateral view: 11 – R. madli sp. nov. 12 – R. kalalaoensis sp. nov. Pilosity in Figs 5–6 partly, in Figs 7–10 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="3" pageNumber="422">6, 8, 10, 12</figureCitation>
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,
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<figureCitation id="136EFD89FFB96853FB044919FB5FF9F8" box="[1190,1244,1562,1581]" captionStart="Figs 13–18" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 13–18.Characteristics of females: 13–16 – apex of abdomen, lateral view: 13 – Rhagovelia madli sp. nov., 14 – R. kalalaoensis sp. nov., 15 – R. ambra Polhemus & Andersen, 2015, 16 – R. aquacola Polhemus & Andersen, 2015; 17–18 – tergites 7 and 8, dorsal view: 17 – R. kalalaoensis sp. nov., 18 – R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="3" pageNumber="422">14, 17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34FB287FFB96853FC9E4947FBA5F959" pageId="3" pageNumber="422" type="materials_examined">
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<paragraph id="8BEAE10CFFB96853FC9E4947FBA5F959" blockId="3.[828,1437,1562,1676]" pageId="3" pageNumber="422">
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<emphasis id="B9213D1EFFB96853FC9E4947FC43F982" bold="true" box="[828,960,1604,1623]" pageId="3" pageNumber="422">
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<typeStatus id="54EE5FAEFFB96853FC9E4947FCEAF982" box="[828,873,1604,1623]" pageId="3" pageNumber="422">Type</typeStatus>
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material.
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</emphasis>
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HƟΓƟΤΥΡΕ (apterous ♁) and ΡΑRΑΤΥΡΕඌ (1 apterous ♁, 1 apterous
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<specimenCount id="9D532A85FFB96853FC294958FC18F9A7" box="[907,923,1627,1650]" pageId="3" pageNumber="422" type="female">♀</specimenCount>
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):
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<collectingCountry id="F342A19CFFB96853FC0C495DFB95F9A4" box="[942,1046,1630,1649]" name="Madagascar" pageId="3" pageNumber="422">Madagascar</collectingCountry>
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, Nosy Boraha: Forêt de Kalalao,
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<date id="FFEBC7CCFFB96853FA9F495DFA1BF9A4" box="[1341,1432,1630,1649]" pageId="3" pageNumber="422" value="1992-10-27">27.X.1992</date>
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, leg. M. Madl (all
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<collectionCode id="ED4479C9FFB96853FC70497AFB9FF959" box="[978,1052,1657,1676]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="422" type="Museum">NHMW</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34FB287FFB96854FC9E49A7FB4BF9C0" lastPageId="4" lastPageNumber="423" pageId="3" pageNumber="422" type="description">
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<paragraph id="8BEAE10CFFB96853FC9E49A7FAA5F80E" blockId="3.[828,1437,1699,2075]" pageId="3" pageNumber="422">
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<emphasis id="B9213D1EFFB96853FC9E49A7FB05F96E" bold="true" box="[828,1158,1699,1724]" pageId="3" pageNumber="422">
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Description.
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<emphasis id="B9213D1EFFB96853FC6C49A0FBFCF969" bold="true" box="[974,1151,1699,1724]" italics="true" pageId="3" pageNumber="422">Male (apterous)</emphasis>
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.
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</emphasis>
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<emphasis id="B9213D1EFFB96853FB2F49A7FAABF96E" box="[1165,1320,1700,1723]" italics="true" pageId="3" pageNumber="422">Measurements</emphasis>
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(
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<typeStatus id="54EE5FAEFFB96853FA9549A7FA1AF96E" box="[1335,1433,1700,1723]" pageId="3" pageNumber="422" type="holotype">holotype</typeStatus>
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,
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<typeStatus id="54EE5FAEFFB96853FC9E49C7FC1EF90E" box="[828,925,1732,1755]" pageId="3" pageNumber="422" type="paratype">paratype</typeStatus>
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): Body length 2.91, 2.55; maximum body width (at metapleura) 1.06, 1.03. Head length 0.45, 0.42, width 0.80, 0.78; minimum eye distance 0.13, 0.12. Pronotum length 0.83, 0.81, width 1.00, 1.00. Lengths of antennomeres (
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<typeStatus id="54EE5FAEFFB96853FC254847FC6AF88E" box="[903,1001,1860,1883]" pageId="3" pageNumber="422" type="holotype">holotype</typeStatus>
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), I 0.77, II 0.45, III 0.47, IV 0.51. Lengths of leg segments (
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<typeStatus id="54EE5FAEFFB96853FC534867FBD6F8AE" box="[1009,1109,1892,1915]" pageId="3" pageNumber="422" type="holotype">holotype</typeStatus>
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): profemur 0.94, protibia 1.00, protarsus 0.02 + 0.03 + 0.24, mesofemur 1.59, mesotibia 1.47, mesotarsus 0.10 + 0.50 + 0.72, metafemur 1.34, metatibia 1.44, metatarsus 0.05 + 0.08 + 0.29.
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</paragraph>
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<paragraph id="8BEAE10CFFB96854FCF848E7FEE3FE00" blockId="3.[828,1437,1699,2075]" lastBlockId="4.[151,761,190,2069]" lastPageId="4" lastPageNumber="423" pageId="3" pageNumber="422">
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<emphasis id="B9213D1EFFB96853FCF848E7FC25F82E" box="[858,934,2020,2043]" italics="true" pageId="3" pageNumber="422">Colour</emphasis>
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(
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<figureCitation id="136EFD89FFB96853FC1648E7FC76F82E" box="[948,1013,2020,2043]" captionStart="Figs 1–4. 1–2" captionStartId="2.[151,188,1715,1734]" captionTargetBox="[151,1436,189,1697]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 1–4. 1–2 – Rhagovelia madli sp. nov.: 1 – habitus of holotype, apterous male; 2 – paratype, apterous female; 3 – R. kalalaoensis sp. nov., holotype, apterous male; 4 – R. reuteri Hoberlandt, 1951, small apterous male with reduced metatibial tooth." figureDoi="http://doi.org/10.5281/zenodo.4489336" httpUri="https://zenodo.org/record/4489336/files/figure.png" pageId="3" pageNumber="422">Fig. 3</figureCitation>
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). Trunk chiefly black. Pronotum anteriorly with transverse yellow stripe medially interrupted by narrow blackish brown line. Tergite 6 with small yellowish brown spot, tergite 7 with large yellow spot. Laterotergites 3–6 and sternites 3–6 with narrow yellow lateral margins. Propleuron and acetabula yellow. Venter bicoloured; mesosternum until sternite 6 black, other parts yellow (segment 8 slightly darker). Antenna black, basal third of antennomere 1 yellow. Legs largely black; all coxae, pro- and metatrochanter, basal third of profemur, and base of metafemur yellow.
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</paragraph>
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<paragraph id="8BEAE10CFFBE6854FF174EDDFEBAFD20" blockId="4.[151,761,190,2069]" pageId="4" pageNumber="423">
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<emphasis id="B9213D1EFFBE6854FF174EDDFE93FE20" box="[181,272,478,501]" italics="true" pageId="4" pageNumber="423">Pilosity.</emphasis>
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Body dorsally with dispersed, short, grey, appressed pubescence and inconspicuous, dispersed, grey, subdecumbent short setae; only head, sides of pronotum, and posterior margin of segment 8 with some long black setae. Tubercle of protrochanter with tuft of black setae. Profemur on posterior face with basal row of three slightly thickened setae; posterior row of flexor side with short stout setae. Other black setae on legs normal, slender, of varying length.
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</paragraph>
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<paragraph id="8BEAE10CFFBE6854FF174DFDFD7AFA20" blockId="4.[151,761,190,2069]" pageId="4" pageNumber="423">
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<emphasis id="B9213D1EFFBE6854FF174DFDFEAAFCC0" box="[181,297,766,789]" italics="true" pageId="4" pageNumber="423">Structures.</emphasis>
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Jugum not flattened, narrow, with few black spicula laterally and posteriorly. Proepisternum with few black spicula anteriorly. Pronotum more than twice as long as head, covering mesonotum. Protrochanter (
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<figureCitation id="136EFD89FFBE6854FD6E4C5DFF26FC40" captionStart="Figs 5–12" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 5–12. Characteristics of males: 5–6 – right foreleg: 5 – Rhagovelia madli sp. nov., 6 – R. kalalaoensis sp. nov.; 7–8 – right hind leg: 7 – R. madli sp. nov. 8 – R. kalalaoensis sp. nov.; 9–10 – pygophore, dorsal view: 9 – R. madli sp. nov. 10 – R. kalalaoensis sp. nov.; 11–12 – left paramere, lateral view: 11 – R. madli sp. nov. 12 – R. kalalaoensis sp. nov. Pilosity in Figs 5–6 partly, in Figs 7–10 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="4" pageNumber="423">Fig. 6</figureCitation>
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) with subapical tubercle. Profemur (
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<figureCitation id="136EFD89FFBE6854FD854C7DFDE5FC40" box="[551,614,894,917]" captionStart="Figs 5–12" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 5–12. Characteristics of males: 5–6 – right foreleg: 5 – Rhagovelia madli sp. nov., 6 – R. kalalaoensis sp. nov.; 7–8 – right hind leg: 7 – R. madli sp. nov. 8 – R. kalalaoensis sp. nov.; 9–10 – pygophore, dorsal view: 9 – R. madli sp. nov. 10 – R. kalalaoensis sp. nov.; 11–12 – left paramere, lateral view: 11 – R. madli sp. nov. 12 – R. kalalaoensis sp. nov. Pilosity in Figs 5–6 partly, in Figs 7–10 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="4" pageNumber="423">Fig. 6</figureCitation>
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) with distinct concavity on extensor side. Metafemur (
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<figureCitation id="136EFD89FFBE6854FD954C9DFDF0FC60" box="[567,627,926,949]" captionStart="Figs 5–12" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 5–12. Characteristics of males: 5–6 – right foreleg: 5 – Rhagovelia madli sp. nov., 6 – R. kalalaoensis sp. nov.; 7–8 – right hind leg: 7 – R. madli sp. nov. 8 – R. kalalaoensis sp. nov.; 9–10 – pygophore, dorsal view: 9 – R. madli sp. nov. 10 – R. kalalaoensis sp. nov.; 11–12 – left paramere, lateral view: 11 – R. madli sp. nov. 12 – R. kalalaoensis sp. nov. Pilosity in Figs 5–6 partly, in Figs 7–10 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="4" pageNumber="423">Fig. 8</figureCitation>
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) moderately incrassate, ca. 3.3–3.4 times as long as wide, on flexor side with rich dentition: basal row consisting of 13–14 densely set short spines only weakly increasing in length distally, their distance smaller than their length; posterodistal row consisting of 11–13 teeth strongly decreasing in size distally, the first one considerably longer than the second; anterodistal row consisting of 7–9 minute spines. Metatibia (
|
||
<figureCitation id="136EFD89FFBE6854FEAE4B9DFECFFB60" box="[268,332,1182,1205]" captionStart="Figs 5–12" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 5–12. Characteristics of males: 5–6 – right foreleg: 5 – Rhagovelia madli sp. nov., 6 – R. kalalaoensis sp. nov.; 7–8 – right hind leg: 7 – R. madli sp. nov. 8 – R. kalalaoensis sp. nov.; 9–10 – pygophore, dorsal view: 9 – R. madli sp. nov. 10 – R. kalalaoensis sp. nov.; 11–12 – left paramere, lateral view: 11 – R. madli sp. nov. 12 – R. kalalaoensis sp. nov. Pilosity in Figs 5–6 partly, in Figs 7–10 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="4" pageNumber="423">Fig. 8</figureCitation>
|
||
) straight, on flexor side with fine toothlike structures along entire length, with very short apical spine. Pregenital segments of abdomen relatively slender. Tergites relatively narrow; tergites 1–5 matt, 6 with small shiny spot at middle, 7 almost entirely shiny (except margins). In
|
||
<typeStatus id="54EE5FAEFFBE6854FF5A4A3DFEDBFA80" box="[248,344,1342,1365]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
, fifth tergite 2.1 times as wide as long; seventh tergite 1.8 times as long as sixth and 1.0 times as long as wide. Sternites almost unmodified; sternite 7 between paired, very shallow lateral impressions very weakly convex, hind margin broadly concave. Segment 8 small, subcylindrical, with indistinct ventromedial carina.
|
||
</paragraph>
|
||
<paragraph id="8BEAE10CFFBE6854FF174AFDFD7AF960" blockId="4.[151,761,190,2069]" pageId="4" pageNumber="423">
|
||
Genital segments small and weakly modified. Pygophore slender subovate. Proctiger (
|
||
<figureCitation id="136EFD89FFBE6854FDB9491DFDEEF9E0" box="[539,621,1566,1589]" captionStart="Figs 5–12" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 5–12. Characteristics of males: 5–6 – right foreleg: 5 – Rhagovelia madli sp. nov., 6 – R. kalalaoensis sp. nov.; 7–8 – right hind leg: 7 – R. madli sp. nov. 8 – R. kalalaoensis sp. nov.; 9–10 – pygophore, dorsal view: 9 – R. madli sp. nov. 10 – R. kalalaoensis sp. nov.; 11–12 – left paramere, lateral view: 11 – R. madli sp. nov. 12 – R. kalalaoensis sp. nov. Pilosity in Figs 5–6 partly, in Figs 7–10 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="4" pageNumber="423">Fig. 10</figureCitation>
|
||
) moderately slender, basolateral lobes moderately protruding and posteriorly angled. Paramere (
|
||
<figureCitation id="136EFD89FFBE6854FE38495DFE67F9A0" box="[410,484,1630,1653]" captionStart="Figs 5–12" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 5–12. Characteristics of males: 5–6 – right foreleg: 5 – Rhagovelia madli sp. nov., 6 – R. kalalaoensis sp. nov.; 7–8 – right hind leg: 7 – R. madli sp. nov. 8 – R. kalalaoensis sp. nov.; 9–10 – pygophore, dorsal view: 9 – R. madli sp. nov. 10 – R. kalalaoensis sp. nov.; 11–12 – left paramere, lateral view: 11 – R. madli sp. nov. 12 – R. kalalaoensis sp. nov. Pilosity in Figs 5–6 partly, in Figs 7–10 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="4" pageNumber="423">Fig. 12</figureCitation>
|
||
) small, proximal part very short and setose, middle part curved, bearing short setae ventrally, distal part slightly widened and broadly rounded.
|
||
</paragraph>
|
||
<paragraph id="8BEAE10CFFBE6854FF1749BEFD24F880" blockId="4.[151,761,190,2069]" pageId="4" pageNumber="423">
|
||
<emphasis id="B9213D1EFFBE6854FF1749BEFE16F900" bold="true" box="[181,405,1725,1749]" pageId="4" pageNumber="423">
|
||
<emphasis id="B9213D1EFFBE6854FF1749BEFE0EF900" bold="true" box="[181,397,1725,1749]" italics="true" pageId="4" pageNumber="423">Female (apterous)</emphasis>
|
||
.
|
||
</emphasis>
|
||
The single teneral female has shrivelled during drying so that exact measurements and a description of the exact position of connexiva in a natural condition cannot be provided. However, the most important structural details allow a diagnosis of this morph:
|
||
</paragraph>
|
||
<paragraph id="8BEAE10CFFBE6854FF17485DFEC1F860" blockId="4.[151,761,190,2069]" pageId="4" pageNumber="423">
|
||
<emphasis id="B9213D1EFFBE6854FF17485DFE82F8A0" box="[181,257,1886,1909]" italics="true" pageId="4" pageNumber="423">Colour</emphasis>
|
||
similar as in male, but sternite 7 almost entirely yellow (except margin) and yellow base of metafemur slightly smaller.
|
||
</paragraph>
|
||
<paragraph id="8BEAE10CFFBE6854FF1748BDFC2DFEA0" blockId="4.[151,761,190,2069]" lastBlockId="4.[828,1437,190,1621]" pageId="4" pageNumber="423">
|
||
<emphasis id="B9213D1EFFBE6854FF1748BDFE85F800" box="[181,262,1982,2005]" italics="true" pageId="4" pageNumber="423">Pilosity</emphasis>
|
||
similar to that of male, but without hair tuft on protrochanter, with slender setae on profemur, and pubescence on sternites strongly reduced. Tergites with sparse pubescence; tergite 7 (
|
||
<figureCitation id="136EFD89FFBE6854FBDA4FBDFB45FF00" box="[1144,1222,190,213]" captionStart="Figs 13–18" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 13–18.Characteristics of females: 13–16 – apex of abdomen, lateral view: 13 – Rhagovelia madli sp. nov., 14 – R. kalalaoensis sp. nov., 15 – R. ambra Polhemus & Andersen, 2015, 16 – R. aquacola Polhemus & Andersen, 2015; 17–18 – tergites 7 and 8, dorsal view: 17 – R. kalalaoensis sp. nov., 18 – R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="4" pageNumber="423">Fig. 17</figureCitation>
|
||
) with long, posteriorly directed tuft of black setae, the longest setae reaching apical third of tergite 8. Apical tuft of setae on connexiva moderately developed. Tergite 8, proctiger, and gonocoxa 1 with short pilosity, only the latter with a few moderately long setae.
|
||
</paragraph>
|
||
<paragraph id="8BEAE10CFFBE6854FCF84E7DFBAFFCE0" blockId="4.[828,1437,190,1621]" pageId="4" pageNumber="423">
|
||
<emphasis id="B9213D1EFFBE6854FCF84E7DFBF7FE40" box="[858,1140,382,405]" italics="true" pageId="4" pageNumber="423">Structural characteristics.</emphasis>
|
||
Head and thorax similar to those of male. Protrochanter without tubercle. Metafemur more slender than in male, on flexor side with distal row consisting of 6 teeth. Metatibia without teeth on flexor side. Abdomen strongly modified. Connexiva without any swellings. The narrow posterior tergites suggest that the connexival margins are strongly convergent posteriorly in dorsal aspect. Tergites 1–3 convex, 4–8 flat; tergite 7 (
|
||
<figureCitation id="136EFD89FFBE6854FCE64D7DFC13FD40" box="[836,912,638,661]" captionStart="Figs 13–18" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 13–18.Characteristics of females: 13–16 – apex of abdomen, lateral view: 13 – Rhagovelia madli sp. nov., 14 – R. kalalaoensis sp. nov., 15 – R. ambra Polhemus & Andersen, 2015, 16 – R. aquacola Polhemus & Andersen, 2015; 17–18 – tergites 7 and 8, dorsal view: 17 – R. kalalaoensis sp. nov., 18 – R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="4" pageNumber="423">Fig. 17</figureCitation>
|
||
) narrow, almost parallel-sided (weakly narrowed posteriorly), about 1.5 times as long as wide at base; tergite 8 (
|
||
<figureCitation id="136EFD89FFBE6854FCF54DBDFC27FD00" box="[855,932,702,725]" captionStart="Figs 13–18" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 13–18.Characteristics of females: 13–16 – apex of abdomen, lateral view: 13 – Rhagovelia madli sp. nov., 14 – R. kalalaoensis sp. nov., 15 – R. ambra Polhemus & Andersen, 2015, 16 – R. aquacola Polhemus & Andersen, 2015; 17–18 – tergites 7 and 8, dorsal view: 17 – R. kalalaoensis sp. nov., 18 – R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="4" pageNumber="423">Fig. 17</figureCitation>
|
||
) roundish, about as wide as long and about one fourth wider than base of tergite 7. Posterior corner of sternite 7 (
|
||
<figureCitation id="136EFD89FFBE6854FC0B4DFDFC77FCC0" box="[937,1012,766,789]" captionStart="Figs 13–18" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 13–18.Characteristics of females: 13–16 – apex of abdomen, lateral view: 13 – Rhagovelia madli sp. nov., 14 – R. kalalaoensis sp. nov., 15 – R. ambra Polhemus & Andersen, 2015, 16 – R. aquacola Polhemus & Andersen, 2015; 17–18 – tergites 7 and 8, dorsal view: 17 – R. kalalaoensis sp. nov., 18 – R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="4" pageNumber="423">Fig. 14</figureCitation>
|
||
) only slightly acute (almost rectangular) and weakly protruded.
|
||
</paragraph>
|
||
<paragraph id="8BEAE10CFFBE6854FCF84C3DFB4FFC80" blockId="4.[828,1437,190,1621]" box="[858,1228,829,853]" pageId="4" pageNumber="423">
|
||
<emphasis id="B9213D1EFFBE6854FCF84C3DFBD7FC80" bold="true" box="[858,1108,829,853]" italics="true" pageId="4" pageNumber="423">Macropterous morphs.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="8BEAE10CFFBE6854FC9E4C5DFB4BF9C0" blockId="4.[828,1437,190,1621]" pageId="4" pageNumber="423">
|
||
<emphasis id="B9213D1EFFBE6854FC9E4C5DFBA1FCA0" bold="true" box="[828,1058,862,885]" pageId="4" pageNumber="423">Comparative notes.</emphasis>
|
||
<taxonomicName id="4C559A8FFFBE6854FB8E4C5DFAB9FCA0" authority="Zettel & Laciny, 2020" authorityName="Zettel & Laciny" authorityYear="2020" box="[1068,1338,862,885]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="kalalaoensis" status="sp. nov.">
|
||
<emphasis id="B9213D1EFFBE6854FB8E4C5DFAB9FCA0" box="[1068,1338,862,885]" italics="true" pageId="4" pageNumber="423">Rhagovelia kalalaoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2128065FFBE6854FAE74C5DFA1EFCA0" box="[1349,1437,862,885]" pageId="4" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is a member of the
|
||
<taxonomicName id="4C559A8FFFBE6854FBB64C7DFBF2FC40" authorityName="Hoberlandt" authorityYear="1941" box="[1044,1137,894,917]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="tesari">
|
||
<emphasis id="B9213D1EFFBE6854FBB64C7DFBF2FC40" box="[1044,1137,894,917]" italics="true" pageId="4" pageNumber="423">R. tesari</emphasis>
|
||
</taxonomicName>
|
||
group. It is very similar to
|
||
<taxonomicName id="4C559A8FFFBE6854FC9E4C9DFA89FC60" authority="Polhemus & Andersen, 2015" authorityName="Polhemus & Andersen" authorityYear="2015" box="[828,1290,926,949]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="aquacola">
|
||
<emphasis id="B9213D1EFFBE6854FC9E4C9DFC42FC60" box="[828,961,926,949]" italics="true" pageId="4" pageNumber="423">R. aquacola</emphasis>
|
||
<bibRefCitation id="EFC49CFDFFBE6854FC684C9DFA89FC60" author="POLHEMUS D. A. & ANDERSEN N. M." box="[970,1290,926,949]" pageId="4" pageNumber="423" pagination="115 - 220" refId="ref6614" refString="POLHEMUS D. A. & ANDERSEN N. M. 2015: Rhagovelia of Madagascar and nearby Indian Ocean Islands (Hemiptera: Veliidae), Part 2: Revision of the tesari species group. Insect Systematics and Evolution 46: 115 - 220." type="journal article" year="2015">Polhemus & Andersen, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
, but the new species is much smaller and its legs are darker. For direct comparison we used apterous specimens of
|
||
<taxonomicName id="4C559A8FFFBE6854FAB84CDDFA1FFC20" authorityName="Polhemus & Andersen" authorityYear="2015" box="[1306,1436,990,1013]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="aquacola">
|
||
<emphasis id="B9213D1EFFBE6854FAB84CDDFA1FFC20" box="[1306,1436,990,1013]" italics="true" pageId="4" pageNumber="423">R. aquacola</emphasis>
|
||
</taxonomicName>
|
||
originating from the
|
||
<typeStatus id="54EE5FAEFFBE6854FBB34CFDFBBDFBC0" box="[1041,1086,1022,1045]" pageId="4" pageNumber="423">type</typeStatus>
|
||
area deposited in the Natural History Museum
|
||
<collectingRegion id="49912FEEFFBE6854FC684B1DFB94FBE0" box="[970,1047,1054,1077]" country="Austria" name="Wien" pageId="4" pageNumber="423">Vienna</collectingRegion>
|
||
. The two species share the armature of the legs, especially the characteristic hairy tubercle on the protrochanter of males (
|
||
<figureCitation id="136EFD89FFBE6854FB9C4B5DFB03FBA0" box="[1086,1152,1118,1141]" captionStart="Figs 5–12" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 5–12. Characteristics of males: 5–6 – right foreleg: 5 – Rhagovelia madli sp. nov., 6 – R. kalalaoensis sp. nov.; 7–8 – right hind leg: 7 – R. madli sp. nov. 8 – R. kalalaoensis sp. nov.; 9–10 – pygophore, dorsal view: 9 – R. madli sp. nov. 10 – R. kalalaoensis sp. nov.; 11–12 – left paramere, lateral view: 11 – R. madli sp. nov. 12 – R. kalalaoensis sp. nov. Pilosity in Figs 5–6 partly, in Figs 7–10 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="4" pageNumber="423">Fig. 6</figureCitation>
|
||
), and the long, posteriorly directed pilosity at the hind margin of tergite 7 of females (
|
||
<figureCitation id="136EFD89FFBE6854FCE14B9DFC39FB60" box="[835,954,1182,1205]" captionStart="Figs 13–18" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 13–18.Characteristics of females: 13–16 – apex of abdomen, lateral view: 13 – Rhagovelia madli sp. nov., 14 – R. kalalaoensis sp. nov., 15 – R. ambra Polhemus & Andersen, 2015, 16 – R. aquacola Polhemus & Andersen, 2015; 17–18 – tergites 7 and 8, dorsal view: 17 – R. kalalaoensis sp. nov., 18 – R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="4" pageNumber="423">Figs 17, 18</figureCitation>
|
||
). The paramere shapes are identical. The main structural difference is found in the apterous female: in
|
||
<taxonomicName id="4C559A8FFFBE6854FA274BBDFC42FB20" authority="Zettel & Laciny, 2020" authorityName="Zettel & Laciny" authorityYear="2020" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="kalalaoensis" status="sp. nov.">
|
||
<emphasis id="B9213D1EFFBE6854FA274BBDFA16FB00" box="[1413,1429,1214,1237]" italics="true" pageId="4" pageNumber="423">R</emphasis>
|
||
.
|
||
<emphasis id="B9213D1EFFBE6854FC9E4BDDFC42FB20" box="[828,961,1246,1269]" italics="true" pageId="4" pageNumber="423">kalalaoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2128065FFBE6854FC684BDDFBA3FB20" box="[970,1056,1246,1269]" pageId="4" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
the connexiva are slender all over their length (with strong swelling on segments 4 and
|
||
<quantity id="4CAD4CE9FFBE6854FAD14BFDFA1FFAC0" box="[1395,1436,1278,1301]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="423" unit="in" value="5.0">5 in</quantity>
|
||
<taxonomicName id="4C559A8FFFBE6854FC9E4A1DFC38FAE0" authorityName="Polhemus & Andersen" authorityYear="2015" box="[828,955,1310,1333]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="aquacola">
|
||
<emphasis id="B9213D1EFFBE6854FC9E4A1DFC38FAE0" box="[828,955,1310,1333]" italics="true" pageId="4" pageNumber="423">R. aquacola</emphasis>
|
||
</taxonomicName>
|
||
), tergite 7 is narrower than tergite 8 (subequal in
|
||
<taxonomicName id="4C559A8FFFBE6854FCF74A3DFC56FA80" authorityName="Polhemus & Andersen" authorityYear="2015" box="[853,981,1342,1365]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="aquacola">
|
||
<emphasis id="B9213D1EFFBE6854FCF74A3DFC56FA80" box="[853,981,1342,1365]" italics="true" pageId="4" pageNumber="423">R. aquacola</emphasis>
|
||
</taxonomicName>
|
||
; comp.
|
||
<figureCitation id="136EFD89FFBE6854FB8A4A3DFB47FA80" box="[1064,1220,1342,1365]" captionStart="Figs 13–18" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 13–18.Characteristics of females: 13–16 – apex of abdomen, lateral view: 13 – Rhagovelia madli sp. nov., 14 – R. kalalaoensis sp. nov., 15 – R. ambra Polhemus & Andersen, 2015, 16 – R. aquacola Polhemus & Andersen, 2015; 17–18 – tergites 7 and 8, dorsal view: 17 – R. kalalaoensis sp. nov., 18 – R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="4" pageNumber="423">Figs 17 and 18</figureCitation>
|
||
), and the connexival corner of sternite 7 is less protruded in lateral aspect (comp.
|
||
<figureCitation id="136EFD89FFBE6854FC9E4A7DFC5AFA40" box="[828,985,1406,1429]" captionStart="Figs 13–18" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 13–18.Characteristics of females: 13–16 – apex of abdomen, lateral view: 13 – Rhagovelia madli sp. nov., 14 – R. kalalaoensis sp. nov., 15 – R. ambra Polhemus & Andersen, 2015, 16 – R. aquacola Polhemus & Andersen, 2015; 17–18 – tergites 7 and 8, dorsal view: 17 – R. kalalaoensis sp. nov., 18 – R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="4" pageNumber="423">Figs 14 and 16</figureCitation>
|
||
). Using the key by PƟΓΗΕΜUඌ & AΝĐΕRඌΕΝ (2015),
|
||
<taxonomicName id="4C559A8FFFBE6854FC334A9DFBB6FA60" authority="Zettel & Laciny, 2020" authorityName="Zettel & Laciny" authorityYear="2020" box="[913,1077,1438,1461]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="kalalaoensis" status="sp. nov.">
|
||
<emphasis id="B9213D1EFFBE6854FC334A9DFC22FA60" box="[913,929,1438,1461]" italics="true" pageId="4" pageNumber="423">R</emphasis>
|
||
.
|
||
<emphasis id="B9213D1EFFBE6854FC124A9DFBB6FA60" box="[944,1077,1438,1461]" italics="true" pageId="4" pageNumber="423">kalalaoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2128065FFBE6854FB9F4A9DFB12FA60" box="[1085,1169,1438,1461]" pageId="4" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
keys out with
|
||
<taxonomicName id="4C559A8FFFBE6854FA904A9DFBFBFA00" authority="Polhemus & Andersen, 2015" authorityName="Polhemus & Andersen" authorityYear="2015" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="brincki">
|
||
<emphasis id="B9213D1EFFBE6854FA904A9DFA1EFA60" box="[1330,1437,1438,1461]" italics="true" pageId="4" pageNumber="423">R. brincki</emphasis>
|
||
<bibRefCitation id="EFC49CFDFFBE6854FC9E4ABDFBFBFA00" author="POLHEMUS D. A. & ANDERSEN N. M." box="[828,1144,1470,1493]" pageId="4" pageNumber="423" pagination="115 - 220" refId="ref6614" refString="POLHEMUS D. A. & ANDERSEN N. M. 2015: Rhagovelia of Madagascar and nearby Indian Ocean Islands (Hemiptera: Veliidae), Part 2: Revision of the tesari species group. Insect Systematics and Evolution 46: 115 - 220." type="journal article" year="2015">Polhemus & Andersen, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
because of its small body size, but strongly differs from it by the paramere shape and other aforementioned characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34FB287FFBE6854FC9E491DFB86F980" pageId="4" pageNumber="423" type="etymology">
|
||
<paragraph id="8BEAE10CFFBE6854FC9E491DFB86F980" blockId="4.[828,1437,190,1621]" pageId="4" pageNumber="423">
|
||
<emphasis id="B9213D1EFFBE6854FC9E491DFC3FF9E0" bold="true" box="[828,956,1566,1589]" pageId="4" pageNumber="423">Etymology.</emphasis>
|
||
Named after the
|
||
<typeStatus id="54EE5FAEFFBE6854FB23491DFB2CF9E0" box="[1153,1199,1566,1589]" pageId="4" pageNumber="423">type</typeStatus>
|
||
locality, the Forêt de Kalalao; adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |