583 lines
72 KiB
XML
583 lines
72 KiB
XML
<document id="27636616BF4E9407B0DD21080070275F" ID-CLB-Dataset="32547" ID-DOI="10.11646/zootaxa.4299.4.4" ID-GBIF-Dataset="9cdbf4b8-5a10-4e26-bbcc-c8dcdba8a722" ID-ISSN="1175-5326" ID-Zenodo-Dep="839420" ID-ZooBank="3437A45D-7D3E-4B21-BBCB-BD0B5EDAD44E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502006210608" checkinUser="plazi" docAuthor="Britz, Ralf, Adamson, Eleanor, Raghavan, Rajeev, Ali, Anvar & Dahanukar, Neelesh" docDate="2017" docId="03C27A4F8861FF96FF04CCDAFC83FEA9" docLanguage="en" docName="zootaxa.4299.4.4.pdf.imf" docOrigin="Zootaxa 4299 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Channa pseudomarulius Günther 1861" docType="treatment" docVersion="8" lastPageNumber="535" masterDocId="FFFB02378862FF91FF93CC07FFDDFFB6" masterDocTitle="Channa pseudomarulius, a valid species of snakehead from the Western Ghats region of peninsular India (Teleostei: Channidae), with comments on Ophicephalus grandinosus, O. theophrasti and O. leucopunctatus" masterLastPageNumber="545" masterPageNumber="529" pageNumber="532" updateTime="1698466249722" updateUser="ExternalLinkService">
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<mods:title id="ECD930E68D73B5468411E4C9773CBC92">Channa pseudomarulius, a valid species of snakehead from the Western Ghats region of peninsular India (Teleostei: Channidae), with comments on Ophicephalus grandinosus, O. theophrasti and O. leucopunctatus</mods:title>
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<mods:name id="15D20A6346914952B5083D4AD5E3CFF6" type="personal">
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<mods:roleTerm id="17E84682C8A96FE29FDF5A5C954CEE8A">Author</mods:roleTerm>
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<mods:namePart id="04708EEE412DA748C9076F54A237C8B8">Britz, Ralf</mods:namePart>
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<mods:name id="BBFA7172E0ADB6C578512E3852B6DECD" type="personal">
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<mods:roleTerm id="F8334454FCA6FAF97C00C9DCDABEDB25">Author</mods:roleTerm>
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<mods:namePart id="E2E00E09FCE19C448E8B924F5B803AA6">Adamson, Eleanor</mods:namePart>
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<mods:roleTerm id="B2C453543C8938EC9699103834E7A07F">Author</mods:roleTerm>
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<mods:namePart id="809DF52DDAA44006EE495AB5239E359A">Raghavan, Rajeev</mods:namePart>
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<mods:namePart id="0A4FD0F1BEFFC08C125C9C061E6AC96B">Ali, Anvar</mods:namePart>
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<mods:name id="A95186BCEA373EE334408DF9BF236832" type="personal">
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<mods:roleTerm id="89A5266842DC608702E1F518223D00D7">Author</mods:roleTerm>
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<mods:namePart id="B23C0D21B4B02B9C6BE2586628E84FC8">Dahanukar, Neelesh</mods:namePart>
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<mods:typeOfResource id="F67B46C416E19AA0D1781E9DBFB7A29E">text</mods:typeOfResource>
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<mods:title id="22C11B3E516EA1619A8E85D38834EE73">Zootaxa</mods:title>
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<mods:date id="4DDB09A8AD9FFEEDAE32F349CA11FAB4">2017</mods:date>
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<mods:number id="11C1A60BA3ACDC048539B046B15BBE78">4299</mods:number>
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<mods:identifier id="D8068B5A4F910B1B781CF217837D8597" type="DOI">10.11646/zootaxa.4299.4.4</mods:identifier>
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<treatment id="03C27A4F8861FF96FF04CCDAFC83FEA9" ID-DOI="http://doi.org/10.5281/zenodo.6045575" ID-GBIF-Taxon="132630606" ID-Zenodo-Dep="6045575" LSID="urn:lsid:plazi:treatment:03C27A4F8861FF96FF04CCDAFC83FEA9" httpUri="http://treatment.plazi.org/id/03C27A4F8861FF96FF04CCDAFC83FEA9" lastPageId="7" lastPageNumber="535" pageId="3" pageNumber="532">
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<subSubSection id="C37198D28861FF92FF04CCDAFEF3FEAE" pageId="3" pageNumber="532" type="nomenclature">
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<paragraph id="8BD4CB598861FF92FF04CCDAFE1FFF41" blockId="3.[151,450,221,280]" box="[151,450,221,247]" pageId="3" pageNumber="532">
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<heading id="D09C7C358861FF92FF04CCDAFE1FFF41" bold="true" box="[151,450,221,247]" fontSize="11" level="1" pageId="3" pageNumber="532" reason="1">
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<taxonomicName id="4C6BB0DA8861FF92FF04CCDAFE1FFF41" ID-CoL="49WB5" authorityName="Günther" authorityYear="1861" box="[151,450,221,247]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="532" phylum="Chordata" rank="species" species="pseudomarulius">
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<emphasis id="B91F174B8861FF92FF04CCDAFE1FFF41" bold="true" box="[151,450,221,247]" italics="true" pageId="3" pageNumber="532">Channa pseudomarulius</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BD4CB598861FF92FF04CCF8FEF3FEAE" blockId="3.[151,450,221,280]" box="[151,302,255,280]" pageId="3" pageNumber="532">
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(
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<figureCitation id="1350D7DC8861FF92FF0DCCF8FED6FEAE" box="[158,267,255,280]" captionStart="FIGURE 2" captionStartId="4.[151,250,1605,1627]" captionTargetBox="[151,1436,293,1583]" captionTargetId="figure@4.[151,1436,265,1583]" captionTargetPageId="4" captionText="FIGURE 2. Channa pseudomarulius, holotype (BMNH 1854.6. 13.123, 200 mm SL), lateral view (A); juvenile (KUFOS 2017. KUT. 15, 56 mm SL), lateral view (B); juvenile (ZRC 41657, 100.5 mm SL), lateral view (C); adult (ZRC 22869, 363 mm SL), lateral view." httpUri="https://zenodo.org/record/839424/files/figure.png" pageId="3" pageNumber="532">Figures 2</figureCitation>
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,
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||
<figureCitation id="1350D7DC8861FF92FE84CD07FEF8FEAE" box="[279,293,256,280]" captionStart="FIGURE 3" captionStartId="6.[151,250,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 3. Channa pseudomarulius, colouration in life. Juvenile (A), KUFOS 2017. KUT. 15, ca. 55 mm SL, Kerala, Kuttiyadi river; juvenile (B) ca. 80 mm SL, Kerala, Irrity, not preserved; subadult (C), 164 mm SL, Kerala, Pamba River, not preserved." httpUri="https://zenodo.org/record/839426/files/figure.png" pageId="3" pageNumber="532">3</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C37198D28861FF92FF04CD40FAE6FE1E" pageId="3" pageNumber="532" type="diagnosis">
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<paragraph id="8BD4CB598861FF92FF04CD40FAE6FE1E" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="532">
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<emphasis id="B91F174B8861FF92FF04CD40FECCFED6" bold="true" box="[151,273,327,352]" pageId="3" pageNumber="532">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C6BB0DA8861FF92FE8ACD4FFDF3FED6" authorityName="Günther" authorityYear="1861" box="[281,558,328,352]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="532" phylum="Chordata" rank="species" species="pseudomarulius">
|
||
<emphasis id="B91F174B8861FF92FE8ACD4FFDF3FED6" box="[281,558,328,352]" italics="true" pageId="3" pageNumber="532">Channa pseudomarulius</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C6BB0DA8861FF92FCF7CD4FFC36FED6" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[868,1003,328,352]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="532" phylum="Chordata" rank="species" species="marulius">
|
||
<emphasis id="B91F174B8861FF92FCF7CD4FFC36FED6" box="[868,1003,328,352]" italics="true" pageId="3" pageNumber="532">C. marulius</emphasis>
|
||
</taxonomicName>
|
||
by fewer lateral line scales (59–61 vs 63–65), fewer dorsal- (47–50 vs 52–56) and anal-fin (29–33 vs 33–37) rays and fewer vertebrae (55 vs 59–63) (
|
||
<tableCitation id="C6E9FEE28861FF92FF0DCD88FF28FE1E" box="[158,245,399,424]" captionStart="TABLE 2" captionStartId="7.[151,239,1406,1429]" captionText="TABLE 2. Meristic information for Channa pseudomarulius, C. marulius, the lectotype of Ophiocephalus grandinosus (MNHN A 1959) and the holotype of O. theophrasti (MNHN A 668)." pageId="3" pageNumber="532">Table 2</tableCitation>
|
||
,
|
||
<figureCitation id="1350D7DC8861FF92FE92CD88FE99FE1E" box="[257,324,399,424]" captionStart="FIGURE 7" captionStartId="11.[151,250,875,897]" captionTargetBox="[169,1393,212,835]" captionTargetId="figure@11.[151,1436,193,854]" captionTargetPageId="11" captionText="FIGURE 7. Bar graphs of selected meristic characters illustrating differences between Channa pseudomarulius and C. marulius (data includes lectotype of O. grandinosus and holotype of O. theophrasti as C. marulius)." httpUri="https://zenodo.org/record/839434/files/figure.png" pageId="3" pageNumber="532">Fig. 7</figureCitation>
|
||
), and by a shorter body/longer head especially in specimens over
|
||
<quantity id="4C9366BC8861FF92FBB4CD97FB59FE11" box="[1063,1156,400,424]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="532" unit="mm" value="200.0">200 mm</quantity>
|
||
SL (see below).
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C37198D28861FF96FF54CDB3FC83FEA9" lastPageId="7" lastPageNumber="536" pageId="3" pageNumber="532" type="description">
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<paragraph id="8BD4CB598861FF92FF54CDB3FCBCFCEE" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="532">
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<emphasis id="B91F174B8861FF92FF54CDB3FE84FE7B" bold="true" box="[199,345,436,461]" pageId="3" pageNumber="532">Description.</emphasis>
|
||
For general appearance, see
|
||
<figureCitation id="1350D7DC8861FF92FD20CDB3FCFDFE7A" box="[691,800,436,461]" captionStart="FIGURE 2" captionStartId="4.[151,250,1605,1627]" captionTargetBox="[151,1436,293,1583]" captionTargetId="figure@4.[151,1436,265,1583]" captionTargetPageId="4" captionText="FIGURE 2. Channa pseudomarulius, holotype (BMNH 1854.6. 13.123, 200 mm SL), lateral view (A); juvenile (KUFOS 2017. KUT. 15, 56 mm SL), lateral view (B); juvenile (ZRC 41657, 100.5 mm SL), lateral view (C); adult (ZRC 22869, 363 mm SL), lateral view." httpUri="https://zenodo.org/record/839424/files/figure.png" pageId="3" pageNumber="532">Figures 2</figureCitation>
|
||
and
|
||
<figureCitation id="1350D7DC8861FF92FCF3CDB3FCB2FE7A" box="[864,879,436,460]" captionStart="FIGURE 3" captionStartId="6.[151,250,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 3. Channa pseudomarulius, colouration in life. Juvenile (A), KUFOS 2017. KUT. 15, ca. 55 mm SL, Kerala, Kuttiyadi river; juvenile (B) ca. 80 mm SL, Kerala, Irrity, not preserved; subadult (C), 164 mm SL, Kerala, Pamba River, not preserved." httpUri="https://zenodo.org/record/839426/files/figure.png" pageId="3" pageNumber="532">3</figureCitation>
|
||
, for morphometric information Table 1 and for meristic data
|
||
<tableCitation id="C6E9FEE28861FF92FEBECDD0FE5FFE46" box="[301,386,471,496]" captionStart="TABLE 2" captionStartId="7.[151,239,1406,1429]" captionText="TABLE 2. Meristic information for Channa pseudomarulius, C. marulius, the lectotype of Ophiocephalus grandinosus (MNHN A 1959) and the holotype of O. theophrasti (MNHN A 668)." pageId="3" pageNumber="532">Table 2</tableCitation>
|
||
. Dorsal profile rising in straight line from tip of snout to anterior base of dorsal fin, continuing in horizontal line to middle of dorsal fin, sloping ventrally from there to posterior dorsal-fin base. Ventral profile falling in straight line to vertical through base of pectoral fin, continuing in horizontal line parallel to dorsal profile up to posterior third of anal fin, rising from there to caudal-fin base. Dorsal fin with long base, 2/3 of SL, originating at vertical through pelvic-fin base and extending to vertical through posterior base of anal fin. Anal fin long, 2/5 of SL, originating behind vertical through distal tip of adpressed pectoral fin and ending at vertical through posterior dorsal-fin base. Head length about 1/3 of SL (Fig. 2,3). Eye located in anterior 1/3 of head. Lips prominent; mouth large, with posterior angle of gape behind vertical through posterior rim of orbit. Upper jaw with several rows of villiform teeth along its length, few canines near symphysis. Lower jaw with several rows of small recurved teeth at front, continuing posteriorly as single row, with several prominent canines interspersed. Vomer and palatine with several rows of small, recurved, conical teeth.
|
||
</paragraph>
|
||
<paragraph id="8BD4CB598861FF92FF54CF63FBDCFC73" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="532">
|
||
Lateral-line scales
|
||
<quantity id="4C9366BC8861FF92FE35CF63FDD3FCCB" box="[422,526,868,893]" metricMagnitude="0" metricUnit="m" metricValue="1.5112999999999999" metricValueMax="1.5493999999999999" metricValueMin="1.4731999999999998" pageId="3" pageNumber="532" unit="in" value="59.5" valueMax="61.0" valueMin="58.0">58–61 in</quantity>
|
||
total; 15–18 scales pre-drop, 2–4 forming drop, and 39–42 scales postdrop. Dorsal-fin rays 47–50. Anal-fin rays 29–33. Pectoral fin with 16–18 rays almost reaching vertical through vent when adpressed. Pelvic-fin rays 6. Caudal fin rounded, with 14 principal rays.
|
||
</paragraph>
|
||
<paragraph id="8BD4CB598861FF92FF54CFC8FE30FA0B" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="532">
|
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<emphasis id="B91F174B8861FF92FF54CFC8FE82FC5E" bold="true" box="[199,351,975,1000]" pageId="3" pageNumber="532">Colouration.</emphasis>
|
||
In preservative (
|
||
<figureCitation id="1350D7DC8861FF92FD8FCFC8FDBDFC5E" box="[540,608,975,1000]" captionStart="FIGURE 2" captionStartId="4.[151,250,1605,1627]" captionTargetBox="[151,1436,293,1583]" captionTargetId="figure@4.[151,1436,265,1583]" captionTargetPageId="4" captionText="FIGURE 2. Channa pseudomarulius, holotype (BMNH 1854.6. 13.123, 200 mm SL), lateral view (A); juvenile (KUFOS 2017. KUT. 15, 56 mm SL), lateral view (B); juvenile (ZRC 41657, 100.5 mm SL), lateral view (C); adult (ZRC 22869, 363 mm SL), lateral view." httpUri="https://zenodo.org/record/839424/files/figure.png" pageId="3" pageNumber="532">Fig. 2</figureCitation>
|
||
), small juveniles of
|
||
<quantity id="4C9366BC8861FF92FCD0CFC8FC61FC51" box="[835,956,975,1000]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="532" unit="mm" value="55.0" valueMax="60.0" valueMin="50.0">50–60 mm</quantity>
|
||
SL with dorsum dark brown from snout to end of dorsal-fin base, followed ventrally by conspicuous light-brown longitudinal band extending from eye to base of caudal fin, limited ventrally by slightly wider, dark-brown to black mid-lateral band; ventral area of head and body greyish white. Fins cream, with ventral half of caudal fin darker at base; caudal-fin ocellus absent. Juveniles of about
|
||
<quantity id="4C9366BC8861FF92FEFEC867FE16FBCE" box="[365,459,1120,1144]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="532" unit="mm" value="100.0">100 mm</quantity>
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SL with dark brown colour on top and sides of head ventrally to gular area (
|
||
<figureCitation id="1350D7DC8861FF92FAA5C858FAA6FBCE" box="[1334,1403,1119,1144]" captionStart="FIGURE 2" captionStartId="4.[151,250,1605,1627]" captionTargetBox="[151,1436,293,1583]" captionTargetId="figure@4.[151,1436,265,1583]" captionTargetPageId="4" captionText="FIGURE 2. Channa pseudomarulius, holotype (BMNH 1854.6. 13.123, 200 mm SL), lateral view (A); juvenile (KUFOS 2017. KUT. 15, 56 mm SL), lateral view (B); juvenile (ZRC 41657, 100.5 mm SL), lateral view (C); adult (ZRC 22869, 363 mm SL), lateral view." httpUri="https://zenodo.org/record/839424/files/figure.png" pageId="3" pageNumber="532">Fig. 2</figureCitation>
|
||
B). Ventral side of head and throat whitish cream. Body dorsally dark brown, set off from midlateral series of dark brown blotches by narrow lighter line (a remnant of the small juvenile light stripe) running just below anterior part of lateral line before lateral-line drop and above it after drop. Series of up to seven large dark brown blotches midlaterally (a remnant of the dark brown longitudinal stripe of smaller juveniles) alternating with lighter narrow bars forming as dorsal extensions from lighter ventral side of abdominal and caudal areas of body. Pectoral and pelvic fins cream. Dorsal-fin membrane brownish with pattern of white, sometimes connected, lines. Anal fin whitish cream with few irregular grey spots in posterior part of fin. Caudal fin with dark brown blotch at base, a dark blotch with conspicuous white rim forming ocellus on dorsal part of fin and four cream to white vertical lines separating darker stripes from each other.
|
||
</paragraph>
|
||
<paragraph id="8BD4CB598861FF92FF54C9CEFC90F9C6" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="532">
|
||
In adults of around
|
||
<quantity id="4C9366BC8861FF92FE3FC9CFFE2BFA56" box="[428,502,1480,1504]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="532" unit="cm" value="30.0">30 cm</quantity>
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||
, dorsal and lateral area of head and body dark brown reaching ventrally to about level of lateral line (
|
||
<figureCitation id="1350D7DC8861FF92FE15C9EBFE10F9B2" box="[390,461,1516,1541]" captionStart="FIGURE 2" captionStartId="4.[151,250,1605,1627]" captionTargetBox="[151,1436,293,1583]" captionTargetId="figure@4.[151,1436,265,1583]" captionTargetPageId="4" captionText="FIGURE 2. Channa pseudomarulius, holotype (BMNH 1854.6. 13.123, 200 mm SL), lateral view (A); juvenile (KUFOS 2017. KUT. 15, 56 mm SL), lateral view (B); juvenile (ZRC 41657, 100.5 mm SL), lateral view (C); adult (ZRC 22869, 363 mm SL), lateral view." httpUri="https://zenodo.org/record/839424/files/figure.png" pageId="3" pageNumber="532">Fig. 2</figureCitation>
|
||
C). Lower third of head yellowish beige, body ventral to lateral line light yellowish brown, surrounding series of four or five dark brown to black blotches. Blotches extending dorsally to or just beyond lateral line. Pectoral, dorsal, caudal, and anal fins brown, but pectoral fin with dark brown vertical mark at base. Pelvic fin yellowish beige. Caudal-fin ocellus indistinct.
|
||
</paragraph>
|
||
<paragraph id="8BD4CB598861FF92FF54CA7AFE42F8B6" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="532">
|
||
In life (
|
||
<figureCitation id="1350D7DC8861FF92FEB3CA7BFEBBF922" box="[288,358,1660,1685]" captionStart="FIGURE 3" captionStartId="6.[151,250,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 3. Channa pseudomarulius, colouration in life. Juvenile (A), KUFOS 2017. KUT. 15, ca. 55 mm SL, Kerala, Kuttiyadi river; juvenile (B) ca. 80 mm SL, Kerala, Irrity, not preserved; subadult (C), 164 mm SL, Kerala, Pamba River, not preserved." httpUri="https://zenodo.org/record/839426/files/figure.png" pageId="3" pageNumber="532">Fig. 3</figureCitation>
|
||
), Small juveniles of
|
||
<quantity id="4C9366BC8861FF92FDCFCA7BFD05F922" box="[604,728,1660,1685]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="532" unit="mm" value="55.0" valueMax="60.0" valueMin="50.0">50–60 mm</quantity>
|
||
SL with bright yellowish orange longitudinal band from eye (including iris) to caudal-fin base. Cheek and side of abdomen with golden iridescence. Middle and lower half of body with dark brown longitudinal band originating on opercle and extending onto base of caudal fin. Otherwise fin membranes hyaline.
|
||
</paragraph>
|
||
<paragraph id="8BD4CB598861FF95FF54CB0AFE0DFF63" blockId="3.[151,1437,327,2009]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="533" pageId="3" pageNumber="532">
|
||
Larger juveniles of ca.
|
||
<quantity id="4C9366BC8861FF92FE40CB0BFDF9F892" box="[467,548,1804,1828]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="532" unit="mm" value="80.0">80 mm</quantity>
|
||
SL light brown on top and sides with darker marbling dorsally from head to caudal peduncle (
|
||
<figureCitation id="1350D7DC8861FF92FECDCB28FE7FF8FE" box="[350,418,1839,1864]" captionStart="FIGURE 3" captionStartId="6.[151,250,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 3. Channa pseudomarulius, colouration in life. Juvenile (A), KUFOS 2017. KUT. 15, ca. 55 mm SL, Kerala, Kuttiyadi river; juvenile (B) ca. 80 mm SL, Kerala, Irrity, not preserved; subadult (C), 164 mm SL, Kerala, Pamba River, not preserved." httpUri="https://zenodo.org/record/839426/files/figure.png" pageId="3" pageNumber="532">Fig. 3</figureCitation>
|
||
A). Light stripe running from eye to end of caudal peduncle separating darker dorsal side from series of light brown mid-lateral blotches. Blotches less conspicuous anteriorly in series, separated from each other by orange golden blotches. Ventral side of head and body whitish cream. Pectoral and pelvic fins hyaline. Dorsal- fin membrane with irregular pattern of spots often rimmed by lighter yellowish lines, otherwise hyaline. Anal fin hyaline. Caudal with dark brown base and very conspicuous ocellus on dorsal part of fin formed by large black spherical spot rimmed by narrow yellow line. Additional dorsoventrally running oblique yellowish lines on otherwise brown caudal fin.
|
||
</paragraph>
|
||
<caption id="DF149BD18866FF95FF04CA42FE9BF92E" httpUri="https://zenodo.org/record/839424/files/figure.png" pageId="4" pageNumber="533" targetBox="[151,1436,293,1583]" targetPageId="4">
|
||
<paragraph id="8BD4CB598866FF95FF04CA42FE9BF92E" blockId="4.[151,1435,1605,1688]" pageId="4" pageNumber="533">
|
||
<emphasis id="B91F174B8866FF95FF04CA42FECAF9EC" bold="true" box="[151,279,1605,1627]" pageId="4" pageNumber="533">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C6BB0DA8866FF95FEB2CA42FDC3F9ED" authorityName="Günther" authorityYear="1861" box="[289,542,1605,1627]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="533" phylum="Chordata" rank="species" species="pseudomarulius">
|
||
<emphasis id="B91F174B8866FF95FEB2CA42FDC3F9ED" box="[289,542,1605,1627]" italics="true" pageId="4" pageNumber="533">Channa pseudomarulius</emphasis>
|
||
</taxonomicName>
|
||
, holotype (BMNH 1854.6.13.123, 200 mm SL), lateral view (A); juvenile (KUFOS 2017.KUT.15, 56 mm SL), lateral view (B); juvenile (ZRC 41657, 100.5 mm SL), lateral view (C); adult (ZRC 22869, 363 mm SL), lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD4CB598866FF95FF54CAB8FBEFF806" blockId="4.[151,1437,1727,1968]" pageId="4" pageNumber="533">
|
||
Subadults (
|
||
<quantity id="4C9366BC8866FF95FED7CAB8FE7FF961" box="[324,418,1727,1752]" metricMagnitude="-1" metricUnit="m" metricValue="1.64" pageId="4" pageNumber="533" unit="mm" value="164.0">164 mm</quantity>
|
||
SL) similar to juveniles in colouration of dorsal side of head and body, but lateral series of dark brown mid-lateral blotches more conspicuous and separated by bright orange blotches (
|
||
<figureCitation id="1350D7DC8866FF95FB23CAE4FB2EF94A" box="[1200,1267,1763,1788]" captionStart="FIGURE 3" captionStartId="6.[151,250,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 3. Channa pseudomarulius, colouration in life. Juvenile (A), KUFOS 2017. KUT. 15, ca. 55 mm SL, Kerala, Kuttiyadi river; juvenile (B) ca. 80 mm SL, Kerala, Irrity, not preserved; subadult (C), 164 mm SL, Kerala, Pamba River, not preserved." httpUri="https://zenodo.org/record/839426/files/figure.png" pageId="4" pageNumber="533">Fig. 3</figureCitation>
|
||
B). Mid-lateral dark blotches also with numerous white spots, more numerous in lower half of blotch, but also present above blotches in posterior dorsal half of caudal area. Pectoral and pelvic fins cream hyaline. Dorsal fin with series of dark spots at base and numerous white spots arranged in roughly three irregular longitudinal lines. Anal fin cream with series of white spots arranged in lines. Caudal fin brownish with several irregular vertical lines of white markings. Ocellus less conspicuous than in juveniles and appearing slightly faded.
|
||
</paragraph>
|
||
<paragraph id="8BD4CB598867FF94FFEECDC9FE44FE49" blockId="5.[125,1838,461,511]" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECDC9FF34FE52" bold="true" box="[125,233,462,484]" pageId="5" pageNumber="534">TABLE].</emphasis>
|
||
Morphometric information for
|
||
<taxonomicName id="4C6BB0DA8867FF94FDB2CDCAFCCEFE55" authorityName="Günther" authorityYear="1861" box="[545,787,461,483]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="534" phylum="Chordata" rank="species" species="pseudomarulius">
|
||
<emphasis id="B91F174B8867FF94FDB2CDCAFCCEFE55" box="[545,787,461,483]" italics="true" pageId="5" pageNumber="534">Channa pseudomarulius</emphasis>
|
||
</taxonomicName>
|
||
(n=11),
|
||
<taxonomicName id="4C6BB0DA8867FF94FCFACDC9FC03FE55" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[873,990,461,483]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="534" phylum="Chordata" rank="species" species="marulius">
|
||
<emphasis id="B91F174B8867FF94FCFACDC9FC03FE55" box="[873,990,461,483]" italics="true" pageId="5" pageNumber="534">C. marulius</emphasis>
|
||
</taxonomicName>
|
||
, the lectotype of
|
||
<taxonomicName id="4C6BB0DA8867FF94FB1ACDCAFA42FE55" box="[1161,1439,461,483]" family="Channidae" genus="Ophiocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="534" phylum="Chordata" rank="species" species="grandinosus">
|
||
<emphasis id="B91F174B8867FF94FB1ACDCAFA42FE55" box="[1161,1439,461,483]" italics="true" pageId="5" pageNumber="534">Ophiocephalus grandinosus</emphasis>
|
||
</taxonomicName>
|
||
(MNHN A1959) anđ the holotype of
|
||
<taxonomicName id="4C6BB0DA8867FF94F886CDC9FF33FE49" authorityName="Valenciennes in Jaquemont" authorityYear="1838" family="Channidae" genus="Ophicephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="534" phylum="Chordata" rank="species" species="theophrasti">
|
||
<emphasis id="B91F174B8867FF94F886CDC9FF33FE49" italics="true" pageId="5" pageNumber="534">O. theophrasti</emphasis>
|
||
</taxonomicName>
|
||
(MNHN A 668).
|
||
</paragraph>
|
||
<paragraph id="8BD4CB598867FF94FFEECE38F97BF902" pageId="5" pageNumber="534">
|
||
<table id="F96B39F98867006EFFEECE00F8F3F902" box="[125,1838,519,1716]" gridcols="7" gridrows="24" pageId="5" pageNumber="534">
|
||
<tr id="355BC91B8867006EFFEECE00F8F3FDE3" box="[125,1838,519,597]" gridrow="0" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECE00FE0AFDE3" box="[125,471,519,597]" gridcol="0" gridrow="0" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECE38FEACFDE3" bold="true" box="[125,369,575,597]" pageId="5" pageNumber="534">Standard length in mm</emphasis>
|
||
</th>
|
||
<th id="768AA0678867006EFDD7CE00FCDEFDE3" box="[580,771,519,597]" gridcol="1" gridrow="0" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FDD7CE00FD48FD8F" bold="true" pageId="5" pageNumber="534">
|
||
<taxonomicName id="4C6BB0DA8867FF94FDD7CE00FCDEFDAA" authorityName="Günther" authorityYear="1861" box="[580,771,519,540]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="534" phylum="Chordata" rank="species" species="pseudomarulius">
|
||
<emphasis id="B91F174B8867FF94FDD7CE00FCDEFDAA" bold="true" box="[580,771,519,540]" italics="true" pageId="5" pageNumber="534">C. pseudomarulius</emphasis>
|
||
</taxonomicName>
|
||
(n = ]])
|
||
</emphasis>
|
||
97.6!363.0
|
||
</th>
|
||
<th id="768AA0678867006EFCF3CE00FC0BFDE3" box="[864,982,519,597]" gridcol="2" gridrow="0" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FCF3CE24FC0BFD8F" bold="true" box="[864,982,547,569]" pageId="5" pageNumber="534">Mean ± SD</emphasis>
|
||
</th>
|
||
<th id="768AA0678867006EFBB9CE00FB7EFDE3" box="[1066,1187,519,597]" gridcol="3" gridrow="0" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FBB9CE00FBA1FD8F" bold="true" pageId="5" pageNumber="534">
|
||
<taxonomicName id="4C6BB0DA8867FF94FBB9CE00FB7EFDAA" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[1066,1187,519,540]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="534" phylum="Chordata" rank="species" species="marulius">
|
||
<emphasis id="B91F174B8867FF94FBB9CE00FB7EFDAA" bold="true" box="[1066,1187,519,540]" italics="true" pageId="5" pageNumber="534">C. marulius</emphasis>
|
||
</taxonomicName>
|
||
(n = 35)
|
||
</emphasis>
|
||
40.6!564.0
|
||
</th>
|
||
<th id="768AA0678867006EFB67CE00FAB4FDE3" box="[1268,1385,519,597]" gridcol="4" gridrow="0" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FB67CE24FAB4FD8F" bold="true" box="[1268,1385,547,569]" pageId="5" pageNumber="534">Mean ± SD</emphasis>
|
||
</th>
|
||
<th id="768AA0678867006EFA2DCE00F992FDE3" box="[1470,1615,519,597]" gridcol="5" gridrow="0" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FA2DCE00F9C5FD8F" bold="true" pageId="5" pageNumber="534">
|
||
<taxonomicName id="4C6BB0DA8867FF94FA2DCE00F992FDAA" authorityName="Valenciennes in Jaquemont" authorityYear="1838" box="[1470,1615,519,540]" family="Channidae" genus="Ophicephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="534" phylum="Chordata" rank="species" species="theophrasti">
|
||
<emphasis id="B91F174B8867FF94FA2DCE00F992FDAA" bold="true" box="[1470,1615,519,540]" italics="true" pageId="5" pageNumber="534">O. theophrasti</emphasis>
|
||
</taxonomicName>
|
||
holotype
|
||
</emphasis>
|
||
330.0
|
||
</th>
|
||
<th id="768AA0678867006EF914CE00F8F3FDE3" box="[1671,1838,519,597]" gridcol="6" gridrow="0" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94F914CE00F93BFD8F" bold="true" pageId="5" pageNumber="534">
|
||
<taxonomicName id="4C6BB0DA8867FF94F914CE00F8F9FDAA" box="[1671,1828,519,540]" family="Channidae" genus="Ophiocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="534" phylum="Chordata" rank="species" species="grandinosus">
|
||
<emphasis id="B91F174B8867FF94F914CE00F8F9FDAA" bold="true" box="[1671,1828,519,540]" italics="true" pageId="5" pageNumber="534">O. grandinosus</emphasis>
|
||
</taxonomicName>
|
||
lectotype
|
||
</emphasis>
|
||
564.0
|
||
</th>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECE7BF8F3FD06" box="[125,1838,636,688]" gridrow="1" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECE7BFE0AFD06" box="[125,471,636,688]" gridcol="0" gridrow="1" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FF6ECE7BFF20FD06" bold="true" pageId="5" pageNumber="534">In percent of SL Head length</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CE7BFCDEFD06" box="[580,771,636,688]" gridcol="1" gridrow="1" pageId="5" pageNumber="534">29.2!32.9</td>
|
||
<td id="768AA0678867006EFCF3CE7BFC0BFD06" box="[864,982,636,688]" gridcol="2" gridrow="1" pageId="5" pageNumber="534">31.1 ± 1.3</td>
|
||
<td id="768AA0678867006EFBB9CE7BFB7EFD06" box="[1066,1187,636,688]" gridcol="3" gridrow="1" pageId="5" pageNumber="534">23.8!35.7</td>
|
||
<td id="768AA0678867006EFB67CE7BFAB4FD06" box="[1268,1385,636,688]" gridcol="4" gridrow="1" pageId="5" pageNumber="534">30.2 ± 2.5</td>
|
||
<td id="768AA0678867006EFA2DCE7BF992FD06" box="[1470,1615,636,688]" gridcol="5" gridrow="1" pageId="5" pageNumber="534">27.5</td>
|
||
<td id="768AA0678867006EF914CE7BF8F3FD06" box="[1671,1838,636,688]" gridcol="6" gridrow="1" pageId="5" pageNumber="534">23.8</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECEBFF8F3FD78" box="[125,1838,696,718]" gridrow="2" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECEBFFE0AFD78" box="[125,471,696,718]" gridcol="0" gridrow="2" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECEBFFF2AFD78" bold="true" box="[125,247,696,718]" pageId="5" pageNumber="534">Head depth</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CEBFFCDEFD78" box="[580,771,696,718]" gridcol="1" gridrow="2" pageId="5" pageNumber="534">10.5!15.8</td>
|
||
<td id="768AA0678867006EFCF3CEBFFC0BFD78" box="[864,982,696,718]" gridcol="2" gridrow="2" pageId="5" pageNumber="534">13.0 ± 1.8</td>
|
||
<td id="768AA0678867006EFBB9CEBFFB7EFD78" box="[1066,1187,696,718]" gridcol="3" gridrow="2" pageId="5" pageNumber="534">10.1!15.2</td>
|
||
<td id="768AA0678867006EFB67CEBFFAB4FD78" box="[1268,1385,696,718]" gridcol="4" gridrow="2" pageId="5" pageNumber="534">12.6 ± 1.4</td>
|
||
<td id="768AA0678867006EFA2DCEBFF992FD78" box="[1470,1615,696,718]" gridcol="5" gridrow="2" pageId="5" pageNumber="534">13.1</td>
|
||
<td id="768AA0678867006EF914CEBFF8F3FD78" box="[1671,1838,696,718]" gridcol="6" gridrow="2" pageId="5" pageNumber="534">11.6</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECED1F8F3FC9F" box="[125,1838,726,809]" gridrow="3" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECED1FE0AFC9F" box="[125,471,726,809]" gridcol="0" gridrow="3" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECED0FF28FC9F" bold="true" pageId="5" pageNumber="534">Head width Body depth Body width</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CED1FCDEFC9F" box="[580,771,726,809]" gridcol="1" gridrow="3" pageId="5" pageNumber="534">12.5!17.8 14.5!18.6 6.5!18.1</td>
|
||
<td id="768AA0678867006EFCF3CED1FC0BFC9F" box="[864,982,726,809]" gridcol="2" gridrow="3" pageId="5" pageNumber="534">15.8 ± 1.8 16.6 ± 1.5 13.4 ± 3.6</td>
|
||
<td id="768AA0678867006EFBB9CED1FB7EFC9F" box="[1066,1187,726,809]" gridcol="3" gridrow="3" pageId="5" pageNumber="534">13.3!18.2 12.1!18.5 7.2!12.5</td>
|
||
<td id="768AA0678867006EFB67CED1FAB4FC9F" box="[1268,1385,726,809]" gridcol="4" gridrow="3" pageId="5" pageNumber="534">15.6 ± 0.8 15.5 ± 1.6 10.4 ± 1.6</td>
|
||
<td id="768AA0678867006EFA2DCED1F992FC9F" box="[1470,1615,726,809]" gridcol="5" gridrow="3" pageId="5" pageNumber="534">15.7 14.8 11.6</td>
|
||
<td id="768AA0678867006EF914CED1F8F3FC9F" box="[1671,1838,726,809]" gridcol="6" gridrow="3" pageId="5" pageNumber="534">15.4 14.1 10.6</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECF36F8F3FCF1" box="[125,1838,817,839]" gridrow="4" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECF36FE0AFCF1" box="[125,471,817,839]" gridcol="0" gridrow="4" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECF36FEF6FCF1" bold="true" box="[125,299,817,839]" pageId="5" pageNumber="534">Predorsal length</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CF36FCDEFCF1" box="[580,771,817,839]" gridcol="1" gridrow="4" pageId="5" pageNumber="534">30.9!35.5</td>
|
||
<td id="768AA0678867006EFCF3CF36FC0BFCF1" box="[864,982,817,839]" gridcol="2" gridrow="4" pageId="5" pageNumber="534">32.9 ± 1.6</td>
|
||
<td id="768AA0678867006EFBB9CF36FB7EFCF1" box="[1066,1187,817,839]" gridcol="3" gridrow="4" pageId="5" pageNumber="534">26.2!39.7</td>
|
||
<td id="768AA0678867006EFB67CF36FAB4FCF1" box="[1268,1385,817,839]" gridcol="4" gridrow="4" pageId="5" pageNumber="534">33.0 ± 2.8</td>
|
||
<td id="768AA0678867006EFA2DCF36F992FCF1" box="[1470,1615,817,839]" gridcol="5" gridrow="4" pageId="5" pageNumber="534">31.2</td>
|
||
<td id="768AA0678867006EF914CF36F8F3FCF1" box="[1671,1838,817,839]" gridcol="6" gridrow="4" pageId="5" pageNumber="534">26.2</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECF48F8F3FCD3" box="[125,1838,847,869]" gridrow="5" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECF48FE0AFCD3" box="[125,471,847,869]" gridcol="0" gridrow="5" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECF48FECAFCD3" bold="true" box="[125,279,847,869]" pageId="5" pageNumber="534">Preanal length</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CF48FCDEFCD3" box="[580,771,847,869]" gridcol="1" gridrow="5" pageId="5" pageNumber="534">48.8!56.6</td>
|
||
<td id="768AA0678867006EFCF3CF48FC0BFCD3" box="[864,982,847,869]" gridcol="2" gridrow="5" pageId="5" pageNumber="534">52.5 ± 2.5</td>
|
||
<td id="768AA0678867006EFBB9CF48FB7EFCD3" box="[1066,1187,847,869]" gridcol="3" gridrow="5" pageId="5" pageNumber="534">47.6!62.1</td>
|
||
<td id="768AA0678867006EFB67CF48FAB4FCD3" box="[1268,1385,847,869]" gridcol="4" gridrow="5" pageId="5" pageNumber="534">53.1 ± 2.9</td>
|
||
<td id="768AA0678867006EFA2DCF48F992FCD3" box="[1470,1615,847,869]" gridcol="5" gridrow="5" pageId="5" pageNumber="534">52.4</td>
|
||
<td id="768AA0678867006EF914CF48F8F3FCD3" box="[1671,1838,847,869]" gridcol="6" gridrow="5" pageId="5" pageNumber="534">51.1</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECF6AF8F3FC14" box="[125,1838,877,930]" gridrow="6" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECF6AFE0AFC14" box="[125,471,877,930]" gridcol="0" gridrow="6" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECF69FEFAFC14" bold="true" pageId="5" pageNumber="534">Prepectoral length Prepelvic length</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CF6AFCDEFC14" box="[580,771,877,930]" gridcol="1" gridrow="6" pageId="5" pageNumber="534">29.9!33.6 30.7!35.2</td>
|
||
<td id="768AA0678867006EFCF3CF6AFC0BFC14" box="[864,982,877,930]" gridcol="2" gridrow="6" pageId="5" pageNumber="534">31.2 ± 1.2 33.3 ± 1.4</td>
|
||
<td id="768AA0678867006EFBB9CF6AFB7EFC14" box="[1066,1187,877,930]" gridcol="3" gridrow="6" pageId="5" pageNumber="534">25.7!35.8 29.0!38.1</td>
|
||
<td id="768AA0678867006EFB67CF6AFAB4FC14" box="[1268,1385,877,930]" gridcol="4" gridrow="6" pageId="5" pageNumber="534">30.7 ± 2.7 33.3 ± 2.7</td>
|
||
<td id="768AA0678867006EFA2DCF6AF992FC14" box="[1470,1615,877,930]" gridcol="5" gridrow="6" pageId="5" pageNumber="534">28.1 31.2</td>
|
||
<td id="768AA0678867006EF914CF6AF8F3FC14" box="[1671,1838,877,930]" gridcol="6" gridrow="6" pageId="5" pageNumber="534">25.7 29.3</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECFADF8F3FC76" box="[125,1838,938,960]" gridrow="7" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECFADFE0AFC76" box="[125,471,938,960]" gridcol="0" gridrow="7" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECFADFEB9FC76" bold="true" box="[125,356,938,960]" pageId="5" pageNumber="534">Dorsal-fin base length</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CFADFCDEFC76" box="[580,771,938,960]" gridcol="1" gridrow="7" pageId="5" pageNumber="534">60.9!71.0</td>
|
||
<td id="768AA0678867006EFCF3CFADFC0BFC76" box="[864,982,938,960]" gridcol="2" gridrow="7" pageId="5" pageNumber="534">65.5 ± 3.2</td>
|
||
<td id="768AA0678867006EFBB9CFADFB7EFC76" box="[1066,1187,938,960]" gridcol="3" gridrow="7" pageId="5" pageNumber="534">60.5!72.5</td>
|
||
<td id="768AA0678867006EFB67CFADFAB4FC76" box="[1268,1385,938,960]" gridcol="4" gridrow="7" pageId="5" pageNumber="534">64.4 ± 3.2</td>
|
||
<td id="768AA0678867006EFA2DCFADF992FC76" box="[1470,1615,938,960]" gridcol="5" gridrow="7" pageId="5" pageNumber="534">65.2</td>
|
||
<td id="768AA0678867006EF914CFADF8F3FC76" box="[1671,1838,938,960]" gridcol="6" gridrow="7" pageId="5" pageNumber="534">70.6</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECFCFF8F3FC68" box="[125,1838,968,990]" gridrow="8" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECFCFFE0AFC68" box="[125,471,968,990]" gridcol="0" gridrow="8" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECFCFFE8CFC68" bold="true" box="[125,337,968,990]" pageId="5" pageNumber="534">Anal-fin base length</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CFCFFCDEFC68" box="[580,771,968,990]" gridcol="1" gridrow="8" pageId="5" pageNumber="534">38.9!45.3</td>
|
||
<td id="768AA0678867006EFCF3CFCFFC0BFC68" box="[864,982,968,990]" gridcol="2" gridrow="8" pageId="5" pageNumber="534">42.1 ± 2.2</td>
|
||
<td id="768AA0678867006EFBB9CFCFFB7EFC68" box="[1066,1187,968,990]" gridcol="3" gridrow="8" pageId="5" pageNumber="534">33.7!47.5</td>
|
||
<td id="768AA0678867006EFB67CFCFFAB4FC68" box="[1268,1385,968,990]" gridcol="4" gridrow="8" pageId="5" pageNumber="534">41.5 ± 3.1</td>
|
||
<td id="768AA0678867006EFA2DCFCFF992FC68" box="[1470,1615,968,990]" gridcol="5" gridrow="8" pageId="5" pageNumber="534">45.5</td>
|
||
<td id="768AA0678867006EF914CFCFF8F3FC68" box="[1671,1838,968,990]" gridcol="6" gridrow="8" pageId="5" pageNumber="534">47.5</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECFE1F8F3FBAD" box="[125,1838,998,1051]" gridrow="9" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECFE1FE0AFBAD" box="[125,471,998,1051]" gridcol="0" gridrow="9" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECFE0FEF4FBAD" bold="true" pageId="5" pageNumber="534">Pectoral-fin length Pelvic-fin length</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CFE1FCDEFBAD" box="[580,771,998,1051]" gridcol="1" gridrow="9" pageId="5" pageNumber="534">15.1!17.7 10.2!12.1</td>
|
||
<td id="768AA0678867006EFCF3CFE1FC0BFBAD" box="[864,982,998,1051]" gridcol="2" gridrow="9" pageId="5" pageNumber="534">16.5 ± 1.1 11.2 ± 0.5</td>
|
||
<td id="768AA0678867006EFBB9CFE1FB7EFBAD" box="[1066,1187,998,1051]" gridcol="3" gridrow="9" pageId="5" pageNumber="534">13.9!17.6 9.2!11.6</td>
|
||
<td id="768AA0678867006EFB67CFE1FAB4FBAD" box="[1268,1385,998,1051]" gridcol="4" gridrow="9" pageId="5" pageNumber="534">15.8 ± 1.0 10.4 ± 0.7</td>
|
||
<td id="768AA0678867006EFA2DCFE1F992FBAD" box="[1470,1615,998,1051]" gridcol="5" gridrow="9" pageId="5" pageNumber="534">16.3 9.8</td>
|
||
<td id="768AA0678867006EF914CFE1F8F3FBAD" box="[1671,1838,998,1051]" gridcol="6" gridrow="9" pageId="5" pageNumber="534">15.7 9.2</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEEC824F8F3FB8F" box="[125,1838,1059,1081]" gridrow="10" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEEC824FE0AFB8F" box="[125,471,1059,1081]" gridcol="0" gridrow="10" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEEC824FEA5FB8F" bold="true" box="[125,376,1059,1081]" pageId="5" pageNumber="534">Caudal-peduncle length</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7C824FCDEFB8F" box="[580,771,1059,1081]" gridcol="1" gridrow="10" pageId="5" pageNumber="534">5.6!8.5</td>
|
||
<td id="768AA0678867006EFCF3C824FC0BFB8F" box="[864,982,1059,1081]" gridcol="2" gridrow="10" pageId="5" pageNumber="534">6.9 ± 0.9</td>
|
||
<td id="768AA0678867006EFBB9C824FB7EFB8F" box="[1066,1187,1059,1081]" gridcol="3" gridrow="10" pageId="5" pageNumber="534">6.1!9.2</td>
|
||
<td id="768AA0678867006EFB67C824FAB4FB8F" box="[1268,1385,1059,1081]" gridcol="4" gridrow="10" pageId="5" pageNumber="534">7.5 ± 0.7</td>
|
||
<td id="768AA0678867006EFA2DC824F992FB8F" box="[1470,1615,1059,1081]" gridcol="5" gridrow="10" pageId="5" pageNumber="534">7.3</td>
|
||
<td id="768AA0678867006EF914C824F8F3FB8F" box="[1671,1838,1059,1081]" gridcol="6" gridrow="10" pageId="5" pageNumber="534">6.7</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEEC846F8F3FBE1" box="[125,1838,1089,1111]" gridrow="11" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEEC846FE0AFBE1" box="[125,471,1089,1111]" gridcol="0" gridrow="11" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEEC846FEAFFBE1" bold="true" box="[125,370,1089,1111]" pageId="5" pageNumber="534">Caudal-peduncle depth</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7C846FCDEFBE1" box="[580,771,1089,1111]" gridcol="1" gridrow="11" pageId="5" pageNumber="534">7.2!10.7</td>
|
||
<td id="768AA0678867006EFCF3C846FC0BFBE1" box="[864,982,1089,1111]" gridcol="2" gridrow="11" pageId="5" pageNumber="534">8.1 ± 0.9</td>
|
||
<td id="768AA0678867006EFBB9C846FB7EFBE1" box="[1066,1187,1089,1111]" gridcol="3" gridrow="11" pageId="5" pageNumber="534">6.5!8.3</td>
|
||
<td id="768AA0678867006EFB67C846FAB4FBE1" box="[1268,1385,1089,1111]" gridcol="4" gridrow="11" pageId="5" pageNumber="534">7.2 ± 0.4</td>
|
||
<td id="768AA0678867006EFA2DC846F992FBE1" box="[1470,1615,1089,1111]" gridcol="5" gridrow="11" pageId="5" pageNumber="534">7.1</td>
|
||
<td id="768AA0678867006EF914C846F8F3FBE1" box="[1671,1838,1089,1111]" gridcol="6" gridrow="11" pageId="5" pageNumber="534">7.9</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEEC858F8F3FB22" box="[125,1838,1119,1172]" gridrow="12" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEEC858FE0AFB22" box="[125,471,1119,1172]" gridcol="0" gridrow="12" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEEC867FED5FB22" bold="true" pageId="5" pageNumber="534">Snout length Eye diameter</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7C858FCDEFB22" box="[580,771,1119,1172]" gridcol="1" gridrow="12" pageId="5" pageNumber="534">0.5!0.6 3.2!6.1</td>
|
||
<td id="768AA0678867006EFCF3C858FC0BFB22" box="[864,982,1119,1172]" gridcol="2" gridrow="12" pageId="5" pageNumber="534">0.6 ± 0.0 4.3 ± 1.0</td>
|
||
<td id="768AA0678867006EFBB9C858FB7EFB22" box="[1066,1187,1119,1172]" gridcol="3" gridrow="12" pageId="5" pageNumber="534">4.1!6.9 2.5!5.2</td>
|
||
<td id="768AA0678867006EFB67C858FAB4FB22" box="[1268,1385,1119,1172]" gridcol="4" gridrow="12" pageId="5" pageNumber="534">6.1 ± 0.6 4.1 ± 0.7</td>
|
||
<td id="768AA0678867006EFA2DC858F992FB22" box="[1470,1615,1119,1172]" gridcol="5" gridrow="12" pageId="5" pageNumber="534">5.8 3.2</td>
|
||
<td id="768AA0678867006EF914C858F8F3FB22" box="[1671,1838,1119,1172]" gridcol="6" gridrow="12" pageId="5" pageNumber="534">4.3 2.5</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEEC89BF8F3FB04" box="[125,1838,1180,1202]" gridrow="13" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEEC89BFE0AFB04" box="[125,471,1180,1202]" gridcol="0" gridrow="13" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEEC89BFEF6FB04" bold="true" box="[125,299,1180,1202]" pageId="5" pageNumber="534">Preorbital depth</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7C89BFCDEFB04" box="[580,771,1180,1202]" gridcol="1" gridrow="13" pageId="5" pageNumber="534">5.3!6.6</td>
|
||
<td id="768AA0678867006EFCF3C89BFC0BFB04" box="[864,982,1180,1202]" gridcol="2" gridrow="13" pageId="5" pageNumber="534">5.7 ± 0.4</td>
|
||
<td id="768AA0678867006EFBB9C89BFB7EFB04" box="[1066,1187,1180,1202]" gridcol="3" gridrow="13" pageId="5" pageNumber="534">5.1!6.5</td>
|
||
<td id="768AA0678867006EFB67C89BFAB4FB04" box="[1268,1385,1180,1202]" gridcol="4" gridrow="13" pageId="5" pageNumber="534">5.8 ± 0.4</td>
|
||
<td id="768AA0678867006EFA2DC89BF992FB04" box="[1470,1615,1180,1202]" gridcol="5" gridrow="13" pageId="5" pageNumber="534">5.3</td>
|
||
<td id="768AA0678867006EF914C89BF8F3FB04" box="[1671,1838,1180,1202]" gridcol="6" gridrow="13" pageId="5" pageNumber="534">5.7</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEEC8BDF8F3FB66" box="[125,1838,1210,1232]" gridrow="14" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEEC8BDFE0AFB66" box="[125,471,1210,1232]" gridcol="0" gridrow="14" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEEC8BDFEEEFB66" bold="true" box="[125,307,1210,1232]" pageId="5" pageNumber="534">Postorbital depth</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7C8BDFCDEFB66" box="[580,771,1210,1232]" gridcol="1" gridrow="14" pageId="5" pageNumber="534">7.5!10.2</td>
|
||
<td id="768AA0678867006EFCF3C8BDFC0BFB66" box="[864,982,1210,1232]" gridcol="2" gridrow="14" pageId="5" pageNumber="534">8.5 ± 0.9</td>
|
||
<td id="768AA0678867006EFBB9C8BDFB7EFB66" box="[1066,1187,1210,1232]" gridcol="3" gridrow="14" pageId="5" pageNumber="534">6.7!9.5</td>
|
||
<td id="768AA0678867006EFB67C8BDFAB4FB66" box="[1268,1385,1210,1232]" gridcol="4" gridrow="14" pageId="5" pageNumber="534">8.3 ± 0.8</td>
|
||
<td id="768AA0678867006EFA2DC8BDF992FB66" box="[1470,1615,1210,1232]" gridcol="5" gridrow="14" pageId="5" pageNumber="534">8.3</td>
|
||
<td id="768AA0678867006EF914C8BDF8F3FB66" box="[1671,1838,1210,1232]" gridcol="6" gridrow="14" pageId="5" pageNumber="534">7.1</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEEC8DFF8F3FABB" box="[125,1838,1240,1293]" gridrow="15" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEEC8DFFE0AFABB" box="[125,471,1240,1293]" gridcol="0" gridrow="15" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEEC8DEFEE8FABB" bold="true" pageId="5" pageNumber="534">Interorbital width Upper jaw length</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7C8DFFCDEFABB" box="[580,771,1240,1293]" gridcol="1" gridrow="15" pageId="5" pageNumber="534">6.0!8.1 10.5!13.5</td>
|
||
<td id="768AA0678867006EFCF3C8DFFC0BFABB" box="[864,982,1240,1293]" gridcol="2" gridrow="15" pageId="5" pageNumber="534">6.9 ± 0.7 11.9 ± 0.9</td>
|
||
<td id="768AA0678867006EFBB9C8DFFB7EFABB" box="[1066,1187,1240,1293]" gridcol="3" gridrow="15" pageId="5" pageNumber="534">6.5!8.1 8.9!13.1</td>
|
||
<td id="768AA0678867006EFB67C8DFFAB4FABB" box="[1268,1385,1240,1293]" gridcol="4" gridrow="15" pageId="5" pageNumber="534">7.3 ± 0.4 11.7 ± 1.0</td>
|
||
<td id="768AA0678867006EFA2DC8DFF992FABB" box="[1470,1615,1240,1293]" gridcol="5" gridrow="15" pageId="5" pageNumber="534">7.2 10.9</td>
|
||
<td id="768AA0678867006EF914C8DFF8F3FABB" box="[1671,1838,1240,1293]" gridcol="6" gridrow="15" pageId="5" pageNumber="534">7.1 8.9</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEEC934F8F3FAFF" box="[125,1838,1331,1353]" gridrow="16" pageId="5" pageNumber="534" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
||
<th id="768AA0678867006EFFEEC934FE0AFAFF" box="[125,471,1331,1353]" gridcol="0" gridrow="16" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FF5CC934FE0AFAFF" bold="true" box="[207,471,1331,1353]" pageId="5" pageNumber="534">In percent of head length</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEEC956F8F3FA30" box="[125,1838,1361,1414]" gridrow="17" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEEC956FE0AFA30" box="[125,471,1361,1414]" gridcol="0" gridrow="17" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEEC955FED5FA30" bold="true" pageId="5" pageNumber="534">Snout length Eye diameter</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7C956FCDEFA30" box="[580,771,1361,1414]" gridcol="1" gridrow="17" pageId="5" pageNumber="534">16.9!21.2 10.9!18.8</td>
|
||
<td id="768AA0678867006EFCF3C956FC0BFA30" box="[864,982,1361,1414]" gridcol="2" gridrow="17" pageId="5" pageNumber="534">19.2 ± 1.6 13.7 ± 2.8</td>
|
||
<td id="768AA0678867006EFBB9C956FB7EFA30" box="[1066,1187,1361,1414]" gridcol="3" gridrow="17" pageId="5" pageNumber="534">14.0!23.6 10.1!15.4</td>
|
||
<td id="768AA0678867006EFB67C956FAB4FA30" box="[1268,1385,1361,1414]" gridcol="4" gridrow="17" pageId="5" pageNumber="534">20.2 ± 1.6 13.4 ± 1.4</td>
|
||
<td id="768AA0678867006EFA2DC956F992FA30" box="[1470,1615,1361,1414]" gridcol="5" gridrow="17" pageId="5" pageNumber="534">21.1 11.7</td>
|
||
<td id="768AA0678867006EF914C956F8F3FA30" box="[1671,1838,1361,1414]" gridcol="6" gridrow="17" pageId="5" pageNumber="534">18.3 10.4</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEEC989F8F3FA12" box="[125,1838,1422,1444]" gridrow="18" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEEC989FE0AFA12" box="[125,471,1422,1444]" gridcol="0" gridrow="18" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEEC989FEF6FA12" bold="true" box="[125,299,1422,1444]" pageId="5" pageNumber="534">Preorbital depth</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7C989FCDEFA12" box="[580,771,1422,1444]" gridcol="1" gridrow="18" pageId="5" pageNumber="534">16.3!20.7</td>
|
||
<td id="768AA0678867006EFCF3C989FC0BFA12" box="[864,982,1422,1444]" gridcol="2" gridrow="18" pageId="5" pageNumber="534">18.3 ± 1.4</td>
|
||
<td id="768AA0678867006EFBB9C989FB7EFA12" box="[1066,1187,1422,1444]" gridcol="3" gridrow="18" pageId="5" pageNumber="534">17.1!23.9</td>
|
||
<td id="768AA0678867006EFB67C989FAB4FA12" box="[1268,1385,1422,1444]" gridcol="4" gridrow="18" pageId="5" pageNumber="534">19.3 ± 1.3</td>
|
||
<td id="768AA0678867006EFA2DC989F992FA12" box="[1470,1615,1422,1444]" gridcol="5" gridrow="18" pageId="5" pageNumber="534">19.3</td>
|
||
<td id="768AA0678867006EF914C989F8F3FA12" box="[1671,1838,1422,1444]" gridcol="6" gridrow="18" pageId="5" pageNumber="534">23.9</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEEC9ABF8F3FA74" box="[125,1838,1452,1474]" gridrow="19" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEEC9ABFE0AFA74" box="[125,471,1452,1474]" gridcol="0" gridrow="19" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEEC9ABFEEEFA74" bold="true" box="[125,307,1452,1474]" pageId="5" pageNumber="534">Postorbital depth</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7C9ABFCDEFA74" box="[580,771,1452,1474]" gridcol="1" gridrow="19" pageId="5" pageNumber="534">23.2!31.3</td>
|
||
<td id="768AA0678867006EFCF3C9ABFC0BFA74" box="[864,982,1452,1474]" gridcol="2" gridrow="19" pageId="5" pageNumber="534">27.4 ± 2.6</td>
|
||
<td id="768AA0678867006EFBB9C9ABFB7EFA74" box="[1066,1187,1452,1474]" gridcol="3" gridrow="19" pageId="5" pageNumber="534">23.2!32.1</td>
|
||
<td id="768AA0678867006EFB67C9ABFAB4FA74" box="[1268,1385,1452,1474]" gridcol="4" gridrow="19" pageId="5" pageNumber="534">27.6 ± 1.9</td>
|
||
<td id="768AA0678867006EFA2DC9ABF992FA74" box="[1470,1615,1452,1474]" gridcol="5" gridrow="19" pageId="5" pageNumber="534">30.3</td>
|
||
<td id="768AA0678867006EF914C9ABF8F3FA74" box="[1671,1838,1452,1474]" gridcol="6" gridrow="19" pageId="5" pageNumber="534">29.9</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEEC9CDF8F3FA49" box="[125,1838,1482,1535]" gridrow="20" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEEC9CDFE0AFA49" box="[125,471,1482,1535]" gridcol="0" gridrow="20" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEEC9CCFEE6FA49" bold="true" pageId="5" pageNumber="534">Head depth Interorbital width</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7C9CDFCDEFA49" box="[580,771,1482,1535]" gridcol="1" gridrow="20" pageId="5" pageNumber="534">32.8!52.5 18.2!25.3</td>
|
||
<td id="768AA0678867006EFCF3C9CDFC0BFA49" box="[864,982,1482,1535]" gridcol="2" gridrow="20" pageId="5" pageNumber="534">41.9 ± 6.4 22.1 ± 2.0</td>
|
||
<td id="768AA0678867006EFBB9C9CDFB7EFA49" box="[1066,1187,1482,1535]" gridcol="3" gridrow="20" pageId="5" pageNumber="534">35.8!49.0 21.9!30.0</td>
|
||
<td id="768AA0678867006EFB67C9CDFAB4FA49" box="[1268,1385,1482,1535]" gridcol="4" gridrow="20" pageId="5" pageNumber="534">41.8 ± 4.3 24.3 ± 1.7</td>
|
||
<td id="768AA0678867006EFA2DC9CDF992FA49" box="[1470,1615,1482,1535]" gridcol="5" gridrow="20" pageId="5" pageNumber="534">47.5 26.0</td>
|
||
<td id="768AA0678867006EF914C9CDF8F3FA49" box="[1671,1838,1482,1535]" gridcol="6" gridrow="20" pageId="5" pageNumber="534">48.8 30.0</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECA00F8F3F9AB" box="[125,1838,1543,1565]" gridrow="21" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECA00FE0AF9AB" box="[125,471,1543,1565]" gridcol="0" gridrow="21" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECA00FEE8F9AB" bold="true" box="[125,309,1543,1565]" pageId="5" pageNumber="534">Upper jaw length</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CA00FCDEF9AB" box="[580,771,1543,1565]" gridcol="1" gridrow="21" pageId="5" pageNumber="534">34.9!41.3</td>
|
||
<td id="768AA0678867006EFCF3CA00FC0BF9AB" box="[864,982,1543,1565]" gridcol="2" gridrow="21" pageId="5" pageNumber="534">38.3 ± 2.1</td>
|
||
<td id="768AA0678867006EFBB9CA00FB7EF9AB" box="[1066,1187,1543,1565]" gridcol="3" gridrow="21" pageId="5" pageNumber="534">36.2!41.1</td>
|
||
<td id="768AA0678867006EFB67CA00FAB4F9AB" box="[1268,1385,1543,1565]" gridcol="4" gridrow="21" pageId="5" pageNumber="534">38.8 ± 1.1</td>
|
||
<td id="768AA0678867006EFA2DCA00F992F9AB" box="[1470,1615,1543,1565]" gridcol="5" gridrow="21" pageId="5" pageNumber="534">39.5</td>
|
||
<td id="768AA0678867006EF914CA00F8F3F9AB" box="[1671,1838,1543,1565]" gridcol="6" gridrow="21" pageId="5" pageNumber="534">37.4</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECA43F8F3F920" box="[125,1838,1604,1686]" gridrow="22" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECA43FE0AF920" box="[125,471,1604,1686]" gridcol="0" gridrow="22" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FEA2CA43FF18F920" bold="true" pageId="5" pageNumber="534">Ratios Depth/width of body SL/HL</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CA43FCDEF920" box="[580,771,1604,1686]" gridcol="1" gridrow="22" pageId="5" pageNumber="534">1.0!2.3 3.0!3.4</td>
|
||
<td id="768AA0678867006EFCF3CA43FC0BF920" box="[864,982,1604,1686]" gridcol="2" gridrow="22" pageId="5" pageNumber="534">1.3 ± 0.4 3.2 ± 0.1</td>
|
||
<td id="768AA0678867006EFBB9CA43FB7EF920" box="[1066,1187,1604,1686]" gridcol="3" gridrow="22" pageId="5" pageNumber="534">1.1!2.0 2.8!4.2</td>
|
||
<td id="768AA0678867006EFB67CA43FAB4F920" box="[1268,1385,1604,1686]" gridcol="4" gridrow="22" pageId="5" pageNumber="534">1.5 ± 0.2 3.3 ± 0.3</td>
|
||
<td id="768AA0678867006EFA2DCA43F992F920" box="[1470,1615,1604,1686]" gridcol="5" gridrow="22" pageId="5" pageNumber="534">1.3 3.6</td>
|
||
<td id="768AA0678867006EF914CA43F8F3F920" box="[1671,1838,1604,1686]" gridcol="6" gridrow="22" pageId="5" pageNumber="534">1.3 4.2</td>
|
||
</tr>
|
||
<tr id="355BC91B8867006EFFEECA99F8F3F902" box="[125,1838,1694,1716]" gridrow="23" pageId="5" pageNumber="534">
|
||
<th id="768AA0678867006EFFEECA99FE0AF902" box="[125,471,1694,1716]" gridcol="0" gridrow="23" pageId="5" pageNumber="534">
|
||
<emphasis id="B91F174B8867FF94FFEECA99FEC5F902" bold="true" box="[125,280,1694,1716]" pageId="5" pageNumber="534">SL/body depth</emphasis>
|
||
</th>
|
||
<td id="768AA0678867006EFDD7CA99FCDEF902" box="[580,771,1694,1716]" gridcol="1" gridrow="23" pageId="5" pageNumber="534">5.4!6.9</td>
|
||
<td id="768AA0678867006EFCF3CA99FC0BF902" box="[864,982,1694,1716]" gridcol="2" gridrow="23" pageId="5" pageNumber="534">6.1 ± 0.6</td>
|
||
<td id="768AA0678867006EFBB9CA99FB7EF902" box="[1066,1187,1694,1716]" gridcol="3" gridrow="23" pageId="5" pageNumber="534">5.4!8.3</td>
|
||
<td id="768AA0678867006EFB67CA99FAB4F902" box="[1268,1385,1694,1716]" gridcol="4" gridrow="23" pageId="5" pageNumber="534">6.5 ± 0.7</td>
|
||
<td id="768AA0678867006EFA2DCA99F992F902" box="[1470,1615,1694,1716]" gridcol="5" gridrow="23" pageId="5" pageNumber="534">6.8</td>
|
||
<td id="768AA0678867006EF914CA99F8F3F902" box="[1671,1838,1694,1716]" gridcol="6" gridrow="23" pageId="5" pageNumber="534">7.1</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF149BD18864FF97FF04C9DBFA4FF9A6" httpUri="https://zenodo.org/record/839426/files/figure.png" pageId="6" pageNumber="535" targetBox="[151,1436,193,1478]" targetPageId="6">
|
||
<paragraph id="8BD4CB598864FF97FF04C9DBFA4FF9A6" blockId="6.[151,1436,1500,1552]" pageId="6" pageNumber="535">
|
||
<emphasis id="B91F174B8864FF97FF04C9DBFECFFA47" bold="true" box="[151,274,1500,1522]" pageId="6" pageNumber="535">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C6BB0DA8864FF97FE8BC9DBFDD2FA44" authorityName="Günther" authorityYear="1861" box="[280,527,1500,1522]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="pseudomarulius">
|
||
<emphasis id="B91F174B8864FF97FE8BC9DBFDD2FA44" box="[280,527,1500,1522]" italics="true" pageId="6" pageNumber="535">Channa pseudomarulius</emphasis>
|
||
</taxonomicName>
|
||
, colouration in life. Juvenile (A), KUFOS 2017.KUT.15, ca. 55 mm SL, Kerala, Kuttiyadi river; juvenile (B) ca. 80 mm SL, Kerala, Irrity, not preserved; subadult (C), 164 mm SL, Kerala, Pamba River, not preserved.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD4CB598864FF97FF54CA3BFA9CF89B" blockId="6.[151,1436,1596,2018]" pageId="6" pageNumber="535">
|
||
<emphasis id="B91F174B8864FF97FF54CA3BFDC3F9E3" bold="true" box="[199,542,1596,1621]" pageId="6" pageNumber="535">Comparative morphometry.</emphasis>
|
||
In general appearance,
|
||
<taxonomicName id="4C6BB0DA8864FF97FCD6CA3AFB82F9E3" authorityName="Günther" authorityYear="1861" box="[837,1119,1597,1621]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="pseudomarulius">
|
||
<emphasis id="B91F174B8864FF97FCD6CA3AFB82F9E3" box="[837,1119,1597,1621]" italics="true" pageId="6" pageNumber="535">Channa pseudomarulius</emphasis>
|
||
</taxonomicName>
|
||
is shorter bodied than
|
||
<emphasis id="B91F174B8864FF97FA11CA3AFEDFF9CF" italics="true" pageId="6" pageNumber="535">
|
||
<taxonomicName id="4C6BB0DA8864FF97FA11CA3AFF23F9CF" baseAuthorityName="Hamilton" baseAuthorityYear="1822" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="marulius">C. marulius</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
This is reflected by comparatively lower number of vertebrae and lateral-line scales (
|
||
<figureCitation id="1350D7DC8864FF97FB70CA66FAFBF9CF" box="[1251,1318,1633,1658]" captionStart="FIGURE 7" captionStartId="11.[151,250,875,897]" captionTargetBox="[169,1393,212,835]" captionTargetId="figure@11.[151,1436,193,854]" captionTargetPageId="11" captionText="FIGURE 7. Bar graphs of selected meristic characters illustrating differences between Channa pseudomarulius and C. marulius (data includes lectotype of O. grandinosus and holotype of O. theophrasti as C. marulius)." httpUri="https://zenodo.org/record/839434/files/figure.png" pageId="6" pageNumber="535">Fig 7</figureCitation>
|
||
.).
|
||
<taxonomicName id="4C6BB0DA8864FF97FAD0CA66FE97F92B" authorityName="Günther" authorityYear="1861" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="pseudomarulius">
|
||
<emphasis id="B91F174B8864FF97FAD0CA66FE97F92B" italics="true" pageId="6" pageNumber="535">Channa pseudomarulius</emphasis>
|
||
</taxonomicName>
|
||
also has fewer dorsal- and anal-fin rays, although the
|
||
<typeStatus id="54D075FB8864FF97FC35CA82FBD5F928" box="[934,1032,1669,1694]" pageId="6" pageNumber="535" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6BB0DA8864FF97FBBFCA82FB22F92B" authorityName="Günther" authorityYear="1861" box="[1068,1279,1669,1693]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="pseudomarulius">
|
||
<emphasis id="B91F174B8864FF97FBBFCA82FB22F92B" box="[1068,1279,1669,1693]" italics="true" pageId="6" pageNumber="535">C. pseudomarulius</emphasis>
|
||
</taxonomicName>
|
||
specimen has 33 anal rays, overlapping with the lowest counts for
|
||
<taxonomicName id="4C6BB0DA8864FF97FD6CCAAEFC5AF977" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[767,903,1705,1729]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="marulius">
|
||
<emphasis id="B91F174B8864FF97FD6CCAAEFC5AF977" box="[767,903,1705,1729]" italics="true" pageId="6" pageNumber="535">C. marulius</emphasis>
|
||
</taxonomicName>
|
||
specimens (
|
||
<figureCitation id="1350D7DC8864FF97FB84CAAEFB86F977" box="[1047,1115,1705,1730]" captionStart="FIGURE 7" captionStartId="11.[151,250,875,897]" captionTargetBox="[169,1393,212,835]" captionTargetId="figure@11.[151,1436,193,854]" captionTargetPageId="11" captionText="FIGURE 7. Bar graphs of selected meristic characters illustrating differences between Channa pseudomarulius and C. marulius (data includes lectotype of O. grandinosus and holotype of O. theophrasti as C. marulius)." httpUri="https://zenodo.org/record/839434/files/figure.png" pageId="6" pageNumber="535">Fig 7</figureCitation>
|
||
.). This overall shorter body length means that its head appears comparatively “larger”, as can be seen in
|
||
<figureCitation id="1350D7DC8864FF97FC7CCACAFBEFF953" box="[1007,1074,1741,1766]" captionStart="FIGURE 8" captionStartId="11.[151,250,1365,1387]" captionTargetBox="[167,1425,998,1329]" captionTargetId="figure@11.[151,1436,978,1344]" captionTargetPageId="11" captionText="FIGURE 8. Scatter plot of head length versus standard length, loess lines show trend for each species. The appearance of larger head sizes in Channa pseudomarulius specimens is related to the shorter body of this species in comparison to C. marulius, as reflected by lower vertebral, lateral line scale, and dorsal- and anal-fin ray counts (see Figure 7). This difference in head size is especially apparent when comparing larger specimens (> 200 mm)." httpUri="https://zenodo.org/record/839436/files/figure.png" pageId="6" pageNumber="535">Fig. 8</figureCitation>
|
||
, this difference expressed as the ratio of HL/SL is significant among larger specimens (>
|
||
<quantity id="4C9366BC8864FF97FC81CAF6FCB6F8BF" box="[786,875,1777,1801]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="535" unit="mm" value="200.0">200mm</quantity>
|
||
) (t = -4.482, df = 18.102, p = <0.0003), although not significant when available data from smaller fishes are included (t = -1.6489, df = 35.122, p = 0.1081).
|
||
</paragraph>
|
||
<paragraph id="8BD4CB598864FF96FF54CB3EFC83FEA9" blockId="6.[151,1436,1596,2018]" lastBlockId="7.[151,1436,151,287]" lastPageId="7" lastPageNumber="536" pageId="6" pageNumber="535">
|
||
<materialsCitation id="3B03C1048864FF96FF54CB3EFC83FEA9" ID-GBIF-Occurrence="1585880164" collectorName="Fu's F S" lastPageId="7" lastPageNumber="536" location="north east India" municipality="south west India" pageId="6" pageNumber="535" specimenCount="1" stateProvince="Western">
|
||
<emphasis id="B91F174B8864FF97FF54CB3EFE46F8E4" bold="true" box="[199,411,1849,1874]" pageId="6" pageNumber="535">Molecular results</emphasis>
|
||
. CO1 sequence data revealed two genetic groups; 1)
|
||
<taxonomicName id="4C6BB0DA8864FF97FB9BCB3EFAC3F8E7" authorityName="Günther" authorityYear="1861" box="[1032,1310,1849,1873]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="pseudomarulius">
|
||
<emphasis id="B91F174B8864FF97FB9BCB3EFAC3F8E7" box="[1032,1310,1849,1873]" italics="true" pageId="6" pageNumber="535">Channa pseudomarulius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B91F174B8864FF97FAA2CB3DFAE2F8E7" box="[1329,1343,1850,1873]" italics="true" pageId="6" pageNumber="535">n</emphasis>
|
||
= 11, 4 haplotypes
|
||
<emphasis id="B91F174B8864FF97FE89CB59FEF5F8C3" box="[282,296,1886,1909]" italics="true" pageId="6" pageNumber="535">p</emphasis>
|
||
1–
|
||
<emphasis id="B91F174B8864FF97FED7CB59FE8FF8C3" box="[324,338,1886,1909]" italics="true" pageId="6" pageNumber="535">p</emphasis>
|
||
4) collected exclusively in
|
||
<collectingMunicipality id="6BB051238864FF97FD1FCB5AFC91F8C0" box="[652,844,1885,1910]" pageId="6" pageNumber="535">south west India</collectingMunicipality>
|
||
, and; 2)
|
||
<taxonomicName id="4C6BB0DA8864FF97FC26CB5AFBE0F8C3" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[949,1085,1885,1909]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="marulius">
|
||
<emphasis id="B91F174B8864FF97FC26CB5AFBE0F8C3" box="[949,1085,1885,1909]" italics="true" pageId="6" pageNumber="535">C. marulius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B91F174B8864FF97FBC3CB59FB83F8C3" box="[1104,1118,1886,1909]" italics="true" pageId="6" pageNumber="535">n</emphasis>
|
||
=15, 8 haplotypes
|
||
<emphasis id="B91F174B8864FF97FAD3CB59FA89F8C3" box="[1344,1364,1886,1909]" italics="true" pageId="6" pageNumber="535">m</emphasis>
|
||
1–
|
||
<emphasis id="B91F174B8864FF97FAE3CB59FA59F8C3" box="[1392,1412,1886,1909]" italics="true" pageId="6" pageNumber="535">m</emphasis>
|
||
8) collected throughout central and
|
||
<location id="8EB49D828864FF97FD82CB86FD19F82C" LSID="urn:lsid:plazi:treatment:03C27A4F8861FF96FF04CCDAFC83FEA9:8EB49D828864FF97FD82CB86FD19F82C" box="[529,708,1921,1946]" municipality="south west India" name="north east India" pageId="6" pageNumber="535" stateProvince="Western">north east India</location>
|
||
and
|
||
<collectingRegion id="49AF05BB8864FF97FD6CCB85FC87F82F" box="[767,858,1922,1945]" country="Sri Lanka" name="Western" pageId="6" pageNumber="535">Western</collectingRegion>
|
||
Myanmar (see
|
||
<figureCitation id="1350D7DC8864FF97FB82CB86FB8AF82F" box="[1041,1111,1921,1946]" captionStart="FIGURE 4" captionStartId="7.[151,250,1161,1183]" captionTargetBox="[203,1371,342,1126]" captionTargetId="figure@7.[203,1371,342,1126]" captionTargetPageId="7" captionText="FIGURE 4. Median-joining networks show the hypothesised evolutionary relationships between CO 1 haplotypes (615 bps) for Channa pseudomarulius (left) and C. marulius (right). In each network, circles represent individual haplotypes, lines between haplotypes represent single base pair mutations, and small filled circles represent hypothesised intermediate haplotypes not sampled in this study. There was a minimum of 49 mutational differences between haplotypes from each species. Colour of each haplotype corresponds to geographical locations illustrated on map of Indian subcontinent, unfilled circles represent haplotypes that were observed in samples of unverified origin (all C. pseudomarulius were from Kerala / South India, the unverified C. marulius was reportedly from an unknown location in Andhra Pradesh)." httpUri="https://zenodo.org/record/839428/files/figure.png" pageId="6" pageNumber="535">Fig. 4</figureCitation>
|
||
). Within each group genetic divergence was low (maximum distances of 0.003 and 0.008 for
|
||
<taxonomicName id="4C6BB0DA8864FF97FCFDCBA2FB9FF80B" authorityName="Günther" authorityYear="1861" box="[878,1090,1957,1981]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="pseudomarulius">
|
||
<emphasis id="B91F174B8864FF97FCFDCBA2FB9FF80B" box="[878,1090,1957,1981]" italics="true" pageId="6" pageNumber="535">C. pseudomarulius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6BB0DA8864FF97FBEACBA2FB23F80B" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[1145,1278,1957,1981]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="535" phylum="Chordata" rank="species" species="marulius">
|
||
<emphasis id="B91F174B8864FF97FBEACBA2FB23F80B" box="[1145,1278,1957,1981]" italics="true" pageId="6" pageNumber="535">C. marulius</emphasis>
|
||
</taxonomicName>
|
||
respectively), while divergence between groups was an order of magnitude higher (net divergence 0.082, mean divergence 0.084), equating to a minimum of 8.1% uncorrected divergence (50 mutations). Nucleotide diversity was generally low (π = 1.09, 2.34 for
|
||
<taxonomicName id="4C6BB0DA8865FF96FE27CCBBFD51FF62" authorityName="Günther" authorityYear="1861" box="[436,652,188,212]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="536" phylum="Chordata" rank="species" species="pseudomarulius">
|
||
<emphasis id="B91F174B8865FF96FE27CCBBFD51FF62" box="[436,652,188,212]" italics="true" pageId="7" pageNumber="536">C. pseudomarulius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6BB0DA8865FF96FD59CCBBFC8EFF62" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[714,851,188,212]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="536" phylum="Chordata" rank="species" species="marulius">
|
||
<emphasis id="B91F174B8865FF96FD59CCBBFC8EFF62" box="[714,851,188,212]" italics="true" pageId="7" pageNumber="536">C. marulius</emphasis>
|
||
</taxonomicName>
|
||
respectively) and
|
||
<collectorName id="269EAE8F8865FF96FBBECCBBFBA2FF6F" box="[1069,1151,188,217]" pageId="7" pageNumber="536">
|
||
Fu’s F
|
||
<emphasis id="B91F174B8865FF96FBE4CCCCFBA2FF6F" box="[1143,1151,203,217]" italics="true" pageId="7" pageNumber="536">S</emphasis>
|
||
</collectorName>
|
||
returned non-significant results when each species was analysed as a single ‘population’ (p values>0.05) providing no evidence for genetic signatures of demographic changes in recent evolutionary time.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |