treatments-xml/data/03/B5/87/03B587EEFFC4FFF646A15AB6CB180CCF.xml

<document id="246A649421FDC93F908D13314A739A67" ID-CLB-Dataset="46484" ID-DOI="10.5281/zenodo.278111" ID-GBIF-Dataset="bb46bb18-3c39-4396-9651-1edcb19e37d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="278111" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460381463121" checkinUser="plazi" docAuthor="Kranz-Baltensperger, Yvonne" docDate="2011" docId="03B587EEFFC4FFF646A15AB6CB180CCF" docLanguage="en" docName="zt02939p049.pdf" docOrigin="Zootaxa 2939" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ischnothyreus fobor Kranz-Baltensperger, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="46" masterDocId="FF8CFF96FFDEFFDB46365A21C9370F1B" masterDocTitle="The oonopid spider genus Ischnothyreus in Borneo (Oonopidae, Araneae)" masterLastPageNumber="49" masterPageNumber="1" pageNumber="27" updateTime="1698296904597" updateUser="ExternalLinkService">
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<mods:title id="7B3A8A6820161221FD37E7F0CC63CE4D">The oonopid spider genus Ischnothyreus in Borneo (Oonopidae, Araneae)</mods:title>
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<mods:date id="D9EDB1BD43A054DB96BA7B70C1CBD93F">2011</mods:date>
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<treatment id="03B587EEFFC4FFF646A15AB6CB180CCF" ID-DOI="http://doi.org/10.5281/zenodo.6189536" ID-GBIF-Taxon="119414312" ID-Zenodo-Dep="6189536" LSID="urn:lsid:plazi:treatment:03B587EEFFC4FFF646A15AB6CB180CCF" httpUri="http://treatment.plazi.org/id/03B587EEFFC4FFF646A15AB6CB180CCF" lastPageId="45" lastPageNumber="46" pageId="26" pageNumber="27">
<subSubSection id="C3066573FFC4FFC146A15AB6C8670FC9" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BA336F8FFC4FFC146A15AB6C8C70FAA" blockId="26.[151,496,151,210]" box="[151,496,151,177]" pageId="26" pageNumber="27">
<heading id="D0EB8194FFC4FFC146A15AB6C8C70FAA" bold="true" box="[151,496,151,177]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B968EAEAFFC4FFC146A15AB6C8C70FAA" bold="true" box="[151,496,151,177]" pageId="26" pageNumber="27">
<taxonomicName id="4C1C4D7BFFC4FFC146A15AB6C8BB0FAA" ID-CoL="3Q65J" box="[151,396,151,177]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fobor" status="sp. nov.">
<emphasis id="B968EAEAFFC4FFC146A15AB6C8BB0FAA" bold="true" box="[151,396,151,177]" italics="true" pageId="26" pageNumber="27">Ischnothyreus fobor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFC4FFC147A55AB6C8C70FAA" box="[403,496,151,177]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA336F8FFC4FFC146A15A9BC8670FC9" blockId="26.[151,496,151,210]" box="[151,336,186,210]" pageId="26" pageNumber="27">
(
<figureCitation id="13272A7DFFC4FFC146A95A9BC8170FC9" box="[159,288,186,210]" captionStart-0="FIGURE 20" captionStart-1="FIGURE 21" captionStartId-0="26.[151,250,1873,1896]" captionStartId-1="27.[151,250,1812,1835]" captionTargetBox-0="[181,1387,270,1849]" captionTargetBox-1="[151,1421,188,1790]" captionTargetId-0="figure@26.[181,1405,266,1850]" captionTargetId-1="figure@27.[151,1436,153,1791]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionText-0="FIGURE 20. Ischnothyreus fobor sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, F, left palp, prolateral view; G, left chelicera, anterior view. vp, ventral projection; stp, strong-toothlike projection; vpr, ventral protuberance; l, lobe; e, embolus; mo, membranous outgrowth. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G = 0.1 mm." captionText-1="FIGURE 21. Ischnothyreus fobor sp. nov. female. A, habitus, dorsal view; B, habitus, lateral view; C, carapace, dorsal view; D, carapace, ventral view; E, carapace, frontal view; F, genital area, ventral view, G, genital area, dorsal view. a, apodeme; wt, winding tube; gls, goblet-like structure. Scale bars A, B = 1 mm; C, D, E = 0.5 mm; F, G = 0.1 mm." httpUri-0="https://zenodo.org/record/278131/files/figure.png" httpUri-1="https://zenodo.org/record/278132/files/figure.png" pageId="26" pageNumber="27">Figs 20–21</figureCitation>
, 
<figureCitation id="13272A7DFFC4FFC1471D5A9BC87F0FC9" box="[299,328,186,210]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="26" pageNumber="27">35</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF636670FFC4FFC146A15D70C8B508D0" httpUri="https://zenodo.org/record/278131/files/figure.png" pageId="26" pageNumber="27" targetBox="[181,1387,270,1849]" targetPageId="26">
<paragraph id="8BA336F8FFC4FFC146A15D70C8B508D0" blockId="26.[151,1436,1873,1995]" pageId="26" pageNumber="27">
<emphasis id="B968EAEAFFC4FFC146A15D70C817087D" bold="true" box="[151,288,1873,1896]" pageId="26" pageNumber="27">FIGURE 20.</emphasis>
<taxonomicName id="4C1C4D7BFFC4FFC147135D70C8C7087C" box="[293,496,1873,1895]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fobor" status="sp. nov.">
<emphasis id="B968EAEAFFC4FFC147135D70C8C7087C" box="[293,496,1873,1895]" italics="true" pageId="26" pageNumber="27">Ischnothyreus fobor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFC4FFC147C05D73CB72087C" box="[502,581,1874,1895]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, F, left palp, prolateral view; G, left chelicera, anterior view. vp, ventral projection; stp, strong-toothlike projection; vpr, ventral protuberance; l, lobe; e, embolus; mo, membranous outgrowth. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G = 0.1 mm.
</paragraph>
</caption>
<caption id="DF636670FFC5FFC046A15D35CDBE0876" httpUri="https://zenodo.org/record/278132/files/figure.png" pageId="27" pageNumber="28" targetBox="[151,1421,188,1790]" targetPageId="27">
<paragraph id="8BA336F8FFC5FFC046A15D35CDBE0876" blockId="27.[151,1436,1812,1901]" pageId="27" pageNumber="28">
<emphasis id="B968EAEAFFC5FFC046A15D35C8170832" bold="true" box="[151,288,1812,1835]" pageId="27" pageNumber="28">FIGURE 21.</emphasis>
<taxonomicName id="4C1C4D7BFFC5FFC047115D35C8C40831" box="[295,499,1812,1834]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="fobor" status="sp. nov.">
<emphasis id="B968EAEAFFC5FFC047115D35C8C40831" box="[295,499,1812,1834]" italics="true" pageId="27" pageNumber="28">Ischnothyreus fobor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFC5FFC047CC5D34CB7D0831" box="[506,586,1813,1834]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
female. A, habitus, dorsal view; B, habitus, lateral view; C, carapace, dorsal view; D, carapace, ventral view; E, carapace, frontal view; F, genital area, ventral view, G, genital area, dorsal view. a, apodeme; wt, winding tube; gls, goblet-like structure. Scale bars A, B = 1 mm; C, D, E = 0.5 mm; F, G = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C3066573FFC2FFC746A15AB8CC6D0D44" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BA336F8FFC2FFC746A15AB8C8C90FEC" blockId="28.[151,1437,151,1616]" pageId="28" pageNumber="29">
<emphasis id="B968EAEAFFC2FFC746A15AB8C8840FAB" box="[151,435,152,176]" italics="true" pageId="28" pageNumber="29">
<typeStatus id="54A7885AFFC2FFC746A15AB8C9FC0FAB" box="[151,203,153,176]" pageId="28" pageNumber="29">Type</typeStatus>
material: 
<typeStatus id="54A7885AFFC2FFC7477A5AB9C8840FAB" box="[332,435,152,176]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
: male (PBI_OON 00016082), 
<collectingCountry id="F30B7668FFC2FFC745125AB6CA8D0FB4" box="[804,954,151,175]" name="Malaysia" pageId="28" pageNumber="29">MALAYSIA</collectingCountry>
: Sabah: 
<taxonomicName id="4C1C4D7BFFC2FFC742145AB9CD430FB4" box="[1058,1140,152,175]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="48" phylum="Arthropoda" rank="species" species="danum">Danum</taxonomicName>
Valley Field Centre, 
<date id="FFA21038FFC2FFC7434F5AB6C83B0FCF" pageId="28" pageNumber="29" value="1991-05-06" valueMax="1991-05-16" valueMin="1991-05-06">6.– 16.V.1991</date>
, leg. C.L. &amp; P.R. Deeleman, 
<collectionCode id="ED0DAE3DFFC2FFC744625A9CCB9B0FCF" LSID="urn:lsid:biocol.org:col:34992" box="[596,684,189,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
. 
<typeStatus id="54A7885AFFC2FFC744815A9CCA1E0FCF" box="[695,809,189,212]" pageId="28" pageNumber="29" type="paratype">
<emphasis id="B968EAEAFFC2FFC744815A9CCA1E0FCF" box="[695,809,189,212]" italics="true" pageId="28" pageNumber="29">Paratypes</emphasis>
</typeStatus>
: 
<specimenCount id="9D1AFD71FFC2FFC7450E5A9DCAB90FCF" box="[824,910,188,212]" pageId="28" pageNumber="29" type="male">2 males</specimenCount>
&amp; 
<specimenCount id="9D1AFD71FFC2FFC745825A9DCD170FCF" box="[948,1056,188,212]" pageId="28" pageNumber="29" type="female">2 females</specimenCount>
(PBI_OON 00032229), collected together with 
<typeStatus id="54A7885AFFC2FFC747045AFEC8A00FEC" box="[306,407,223,247]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED0DAE3DFFC2FFC747945AC1C8CD0FEC" LSID="urn:lsid:biocol.org:col:34992" box="[418,506,224,247]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
.
</paragraph>
<paragraph id="8BA336F8FFC2FFC746F15B25CC6D0D44" blockId="28.[151,1437,151,1616]" pageId="28" pageNumber="29">
<emphasis id="B968EAEAFFC2FFC746F15B25CB200E07" box="[199,535,260,284]" italics="true" pageId="28" pageNumber="29">Additional material examined</emphasis>
: 
<specimenCount id="9D1AFD71FFC2FFC744115B25CB490E07" box="[551,638,260,284]" pageId="28" pageNumber="29" type="male">2 males</specimenCount>
(PBI_OON 00031469), 
<collectingCountry id="F30B7668FFC2FFC745A05B25CD1B0E07" box="[918,1068,260,284]" name="Malaysia" pageId="28" pageNumber="29">MALAYSIA</collectingCountry>
: Sabah: Kinabalu National Park, 
<quantity id="4CE49B1DFFC2FFC746A15B06C9D00E24" box="[151,231,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="28" pageNumber="29" unit="m" value="1550.0">1550m</quantity>
, 
<date id="FFA21038FFC2FFC746C25B06C8870E24" box="[244,432,295,319]" pageId="28" pageNumber="29" value="1980-07-21" valueMax="1980-07-27" valueMin="1980-07-21">21.–27.VII.1980</date>
, leg. C.L. &amp; P.R. Deeleman (
<collectionCode id="ED0DAE3DFFC2FFC745365B06CA6C0E24" LSID="urn:lsid:biocol.org:col:34992" box="[768,859,295,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
). 
<specimenCount id="9D1AFD71FFC2FFC745595B06CA8B0E24" box="[879,956,295,319]" pageId="28" pageNumber="29" type="male">1 male</specimenCount>
(PBI_OON 00032231), Bru-88/12. BRU- NEI DARUSSALAM: Brunei-Muara District: près du pont sur le ruisseau Sungai Lubang 
<taxonomicName id="4C1C4D7BFFC2FFC742AA5B6CCDD70E7F" box="[1180,1248,333,356]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="48" phylum="Arthropoda" rank="species" species="barus">Barus</taxonomicName>
, route venant de Tutong, 
<quantity id="4CE49B1DFFC2FFC746C35B4EC8030E9C" box="[245,308,367,391]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="28" pageNumber="29" unit="km" value="33.0">33km</quantity>
de Bandar Seri Begawan, prélèvement de sol au pied de deux grands arbres proches des habitations, env. 
<quantity id="4CE49B1DFFC2FFC746FA5BB5C9CA0EB7" box="[204,253,404,428]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="m" value="20.0">20m</quantity>
(translated: close to bridge over Sungai Lubang 
<taxonomicName id="4C1C4D7BFFC2FFC745155BB4CA530EB7" box="[803,868,405,428]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="48" phylum="Arthropoda" rank="species" species="barus">Barus</taxonomicName>
river, route from Tutong, 
<quantity id="4CE49B1DFFC2FFC742BC5BB5CDE60EB7" box="[1162,1233,404,428]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="28" pageNumber="29" unit="km" value="33.0">33 km</quantity>
from Bandar Seri Begawan, soil extraction at base of two large trees close to habitations, ca. 
<quantity id="4CE49B1DFFC2FFC745D75B96CD200ED4" box="[993,1047,439,463]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="m" value="20.0">20m</quantity>
); 
<date id="FFA21038FFC2FFC7421D5B96CD990ED4" box="[1067,1198,439,463]" pageId="28" pageNumber="29" value="1988-11-16">16.XI.1988</date>
; leg. B. Hauser (Berlese à Seri Begawan) (
<collectionCode id="ED0DAE3DFFC2FFC747AB5BFDC8CE0EEF" box="[413,505,476,500]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="28" pageNumber="29">MHNG</collectionCode>
), 
<specimenCount id="9D1AFD71FFC2FFC744385BFDCB450EEF" box="[526,626,476,500]" pageId="28" pageNumber="29" type="female">1 female</specimenCount>
(PBI_OON 00015355), Bru-88/33. 
<collectingCountry id="F30B7668FFC2FFC742225BFCCC7C0EEF" box="[1044,1355,476,500]" name="Brunei" pageId="28" pageNumber="29">BRUNEI DARUSSALAM</collectingCountry>
: Belait District: Labi Hills Forest Reserve, Teraja, 
<quantity id="4CE49B1DFFC2FFC744B25BDECBF40D0C" box="[644,707,511,535]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="28" pageNumber="29" unit="km" value="42.0">42km</quantity>
S de Sungai Liang (
<quantity id="4CE49B1DFFC2FFC745995BDECAC70D0C" box="[943,1008,511,535]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="29" unit="km" value="12.0">12km</quantity>
au Sud de Labi), environs de Rumah Panjang, forêt primaire (translated: Labi Hills Forest Reserve, Teraja, 
<quantity id="4CE49B1DFFC2FFC7459F5805CADF0D27" box="[937,1000,548,572]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="28" pageNumber="29" unit="km" value="42.0">42km</quantity>
south of Sungai Liang (
<quantity id="4CE49B1DFFC2FFC742CE5805CC0E0D27" box="[1272,1337,548,572]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="29" unit="km" value="12.0">12km</quantity>
south of Labi), proximity of Rumah Panjang), mixed dipterocarp forest, 
<quantity id="4CE49B1DFFC2FFC745565866CAA30D44" box="[864,916,583,607]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="29" unit="m" value="40.0">40m</quantity>
; 
<date id="FFA21038FFC2FFC745965866CD150D44" box="[928,1058,583,607]" pageId="28" pageNumber="29" value="1988-11-22">22.XI.1988</date>
, leg. C. Lienhard (
<collectionCode id="ED0DAE3DFFC2FFC742C45866CC790D44" box="[1266,1358,583,607]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="28" pageNumber="29">MHNG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC2FFC746F1584DCDEA0D9F" box="[199,1245,620,644]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8BA336F8FFC2FFC746F1584DCDEA0D9F" blockId="28.[151,1437,151,1616]" box="[199,1245,620,644]" pageId="28" pageNumber="29">
<emphasis id="B968EAEAFFC2FFC746F1584DC8080D9F" box="[199,319,620,644]" italics="true" pageId="28" pageNumber="29">Etymology</emphasis>
: The specific name, a noun in apposition, is an abbreviation of &quot;found in Borneo&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC2FFC746F158B0C9CB0DF4" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BA336F8FFC2FFC746F158B0C9CB0DF4" blockId="28.[151,1437,151,1616]" pageId="28" pageNumber="29">
<emphasis id="B968EAEAFFC2FFC746F158B0C80E0DB3" box="[199,313,657,680]" italics="true" pageId="28" pageNumber="29">Diagnosis</emphasis>
: The male of 
<taxonomicName id="4C1C4D7BFFC2FFC747E358B0CB1E0DB3" box="[469,553,656,680]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fobor">
<emphasis id="B968EAEAFFC2FFC747E358B0CB1E0DB3" box="[469,553,656,680]" italics="true" pageId="28" pageNumber="29">I. fobor</emphasis>
</taxonomicName>
can be recognized by the palpal bulb with its extremely long and thin embolus (
<figureCitation id="13272A7DFFC2FFC746A95895C9C60DD7" box="[159,241,692,716]" captionStart="FIGURE 20" captionStartId="26.[151,250,1873,1896]" captionTargetBox="[181,1387,270,1849]" captionTargetId="figure@26.[181,1405,266,1850]" captionTargetPageId="26" captionText="FIGURE 20. Ischnothyreus fobor sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, F, left palp, prolateral view; G, left chelicera, anterior view. vp, ventral projection; stp, strong-toothlike projection; vpr, ventral protuberance; l, lobe; e, embolus; mo, membranous outgrowth. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278131/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
E–F). The female genital area is characterized by a small strongly winding tube running posteriorly (
<figureCitation id="13272A7DFFC2FFC743595895C9840DF4" captionStart="FIGURE 21" captionStartId="27.[151,250,1812,1835]" captionTargetBox="[151,1421,188,1790]" captionTargetId="figure@27.[151,1436,153,1791]" captionTargetPageId="27" captionText="FIGURE 21. Ischnothyreus fobor sp. nov. female. A, habitus, dorsal view; B, habitus, lateral view; C, carapace, dorsal view; D, carapace, ventral view; E, carapace, frontal view; F, genital area, ventral view, G, genital area, dorsal view. a, apodeme; wt, winding tube; gls, goblet-like structure. Scale bars A, B = 1 mm; C, D, E = 0.5 mm; F, G = 0.1 mm." httpUri="https://zenodo.org/record/278132/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
F–G).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC2FFC746F158DCC9CB0937" pageId="28" pageNumber="29" type="description">
<paragraph id="8BA336F8FFC2FFC746F158DCC8CF0BDF" blockId="28.[151,1437,151,1616]" pageId="28" pageNumber="29">
<emphasis id="B968EAEAFFC2FFC746F158DCC87C0C0F" box="[199,331,765,788]" italics="true" pageId="28" pageNumber="29">Description</emphasis>
: Male (
<typeStatus id="54A7885AFFC2FFC7479058DDCB390C0F" box="[422,526,764,788]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFC2FFC7448558DDCA2B0C0F" box="[691,796,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="28" pageNumber="29" unit="mm" value="1.66">1.66 mm</quantity>
. 
<emphasis id="B968EAEAFFC2FFC7451E58DCCAA10C0F" box="[808,918,765,788]" italics="true" pageId="28" pageNumber="29">Carapace</emphasis>
: domed in lateral view, fovea present, lateral margin slightly rebordered. 
<emphasis id="B968EAEAFFC2FFC747F85901CB1F0C23" box="[462,552,800,824]" italics="true" pageId="28" pageNumber="29">Clypeus</emphasis>
: curved downwards in frontal view. 
<emphasis id="B968EAEAFFC2FFC745F45900CAC00C23" box="[962,1015,801,824]" italics="true" pageId="28" pageNumber="29">Eyes</emphasis>
: ALE touching. 
<emphasis id="B968EAEAFFC2FFC742855901CC390C2C" box="[1203,1294,800,823]" italics="true" pageId="28" pageNumber="29">Sternum</emphasis>
: setae abundant. 
<emphasis id="B968EAEAFFC2FFC746E05965C86E0C47" box="[214,345,836,860]" italics="true" pageId="28" pageNumber="29">Mouthparts</emphasis>
: chelicerae slightly divergent, without prominent process at base of fangs, fang groove with few small and one larger denticles (
<figureCitation id="13272A7DFFC2FFC744325946CB600C64" box="[516,599,871,895]" captionStart="FIGURE 20" captionStartId="26.[151,250,1873,1896]" captionTargetBox="[181,1387,270,1849]" captionTargetId="figure@26.[181,1405,266,1850]" captionTargetPageId="26" captionText="FIGURE 20. Ischnothyreus fobor sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, F, left palp, prolateral view; G, left chelicera, anterior view. vp, ventral projection; stp, strong-toothlike projection; vpr, ventral protuberance; l, lobe; e, embolus; mo, membranous outgrowth. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278131/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
G). Anterior margin of labium indented at middle. Anteromedian tip of endites with one strong, tooth-like projection (stp) (
<figureCitation id="13272A7DFFC2FFC744D659ADCA060CBF" box="[736,817,908,932]" captionStart="FIGURE 20" captionStartId="26.[151,250,1873,1896]" captionTargetBox="[181,1387,270,1849]" captionTargetId="figure@26.[181,1405,266,1850]" captionTargetPageId="26" captionText="FIGURE 20. Ischnothyreus fobor sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, F, left palp, prolateral view; G, left chelicera, anterior view. vp, ventral projection; stp, strong-toothlike projection; vpr, ventral protuberance; l, lobe; e, embolus; mo, membranous outgrowth. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278131/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
D). 
<emphasis id="B968EAEAFFC2FFC7455059ADCAEE0CBF" box="[870,985,908,932]" italics="true" pageId="28" pageNumber="29">Abdomen:</emphasis>
scutum extending far dorsal of pedicel. Dorsal scutum covering 1/2 to 3/4 of abdomen, fused to epigastric scutum. Epigastric and postepigastric scutum short, almost rectangular, covering about 2/3 of abdominal length. Spinneret scutum present, incomplete ring, with fringe of needle-like setae. 
<emphasis id="B968EAEAFFC2FFC747F159D8C8CB0B0B" box="[455,508,1017,1040]" italics="true" pageId="28" pageNumber="29">Legs</emphasis>
: spine formula: femora: I p0-1-1; II p0-0-1; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFC2FFC7476B5E3DC8F00B2F" box="[349,455,1052,1076]" italics="true" pageId="28" pageNumber="29">Genitalia</emphasis>
: Palp strongly sclerotized, proximal segments dark brown, trochanter with large ventral projection (vp) (
<figureCitation id="13272A7DFFC2FFC747665E61C8950B43" box="[336,418,1088,1112]" captionStart="FIGURE 20" captionStartId="26.[151,250,1873,1896]" captionTargetBox="[181,1387,270,1849]" captionTargetId="figure@26.[181,1405,266,1850]" captionTargetPageId="26" captionText="FIGURE 20. Ischnothyreus fobor sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, F, left palp, prolateral view; G, left chelicera, anterior view. vp, ventral projection; stp, strong-toothlike projection; vpr, ventral protuberance; l, lobe; e, embolus; mo, membranous outgrowth. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278131/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
D–F), femur normal size, patella shorter than femur, not enlarged, cymbium dark brown, fused with bulb, bulb dark brown, more than two times as long as cymbium, stout, tapering apically, with ventral protuberance (vpr) and lobe (l), embolus (e) very long and filamentous, originating between several membranous outgrowths (mo) (
<figureCitation id="13272A7DFFC2FFC747545E8DC8850BDF" box="[354,434,1196,1220]" captionStart="FIGURE 20" captionStartId="26.[151,250,1873,1896]" captionTargetBox="[181,1387,270,1849]" captionTargetId="figure@26.[181,1405,266,1850]" captionTargetPageId="26" captionText="FIGURE 20. Ischnothyreus fobor sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, F, left palp, prolateral view; G, left chelicera, anterior view. vp, ventral projection; stp, strong-toothlike projection; vpr, ventral protuberance; l, lobe; e, embolus; mo, membranous outgrowth. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278131/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
E–F).
</paragraph>
<paragraph id="8BA336F8FFC2FFC746F15EF1C9CB0937" blockId="28.[151,1437,151,1616]" pageId="28" pageNumber="29">
Female (
<typeStatus id="54A7885AFFC2FFC7471E5EF1C8B80BF3" box="[296,399,1232,1256]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFC2FFC744065EF1CBA00BF3" box="[560,663,1232,1256]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="28" pageNumber="29" unit="mm" value="1.68">1.68 mm</quantity>
. As in male except as noted. 
<emphasis id="B968EAEAFFC2FFC745D65EF1CD790BFC" box="[992,1102,1232,1255]" italics="true" pageId="28" pageNumber="29">Carapace</emphasis>
: without any pattern, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view. 
<emphasis id="B968EAEAFFC2FFC745925ED5CAC90A17" box="[932,1022,1268,1292]" italics="true" pageId="28" pageNumber="29">Clypeus</emphasis>
: straight in frontal view. 
<emphasis id="B968EAEAFFC2FFC743165ED4CC620A17" box="[1312,1365,1269,1292]" italics="true" pageId="28" pageNumber="29">Eyes</emphasis>
: ALE separated by less than their radius. 
<emphasis id="B968EAEAFFC2FFC744155F39CB980A2B" box="[547,687,1304,1328]" italics="true" pageId="28" pageNumber="29">Mouthparts:</emphasis>
labium not fused to sternum, anterior margin not indented at middle. Endites unmodified. Femur of female palp with three ventral spines, tibia with three trichobothria. 
<emphasis id="B968EAEAFFC2FFC7431C5F1DCCA30A4F" box="[1322,1428,1340,1364]" italics="true" pageId="28" pageNumber="29">Abdomen</emphasis>
: dorsal scutum covering about 1/2 of abdomen, not fused to epigastric scutum. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length, with short posteriorly directed lateral apodemes (a) (
<figureCitation id="13272A7DFFC2FFC743125FA5CC420A87" box="[1316,1397,1412,1436]" captionStart="FIGURE 21" captionStartId="27.[151,250,1812,1835]" captionTargetBox="[151,1421,188,1790]" captionTargetId="figure@27.[151,1436,153,1791]" captionTargetPageId="27" captionText="FIGURE 21. Ischnothyreus fobor sp. nov. female. A, habitus, dorsal view; B, habitus, lateral view; C, carapace, dorsal view; D, carapace, ventral view; E, carapace, frontal view; F, genital area, ventral view, G, genital area, dorsal view. a, apodeme; wt, winding tube; gls, goblet-like structure. Scale bars A, B = 1 mm; C, D, E = 0.5 mm; F, G = 0.1 mm." httpUri="https://zenodo.org/record/278132/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
F). Dense patch of setae anterior to spinnerets present. 
<emphasis id="B968EAEAFFC2FFC744E25F88CA3E0ADB" box="[724,777,1449,1472]" italics="true" pageId="28" pageNumber="29">Legs</emphasis>
: spine formula: femora: I p0-1-1; r0-1-1; II p0-0-1; r0-0-1; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFC2FFC7452C5FEDCAB40AFF" box="[794,899,1484,1508]" italics="true" pageId="28" pageNumber="29">Genitalia</emphasis>
: simple, in the middle of the anterior margin of the postepigastric scutum starts a winding tube (wt), running posteriorly, ending in globlet-like structure (gls) (
<figureCitation id="13272A7DFFC2FFC743585FD1C9840937" captionStart="FIGURE 21" captionStartId="27.[151,250,1812,1835]" captionTargetBox="[151,1421,188,1790]" captionTargetId="figure@27.[151,1436,153,1791]" captionTargetPageId="27" captionText="FIGURE 21. Ischnothyreus fobor sp. nov. female. A, habitus, dorsal view; B, habitus, lateral view; C, carapace, dorsal view; D, carapace, ventral view; E, carapace, frontal view; F, genital area, ventral view, G, genital area, dorsal view. a, apodeme; wt, winding tube; gls, goblet-like structure. Scale bars A, B = 1 mm; C, D, E = 0.5 mm; F, G = 0.1 mm." httpUri="https://zenodo.org/record/278132/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
F–G).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC2FFC746F15C19C86709C5" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8BA336F8FFC2FFC746F15C19CBA2094B" blockId="28.[151,1437,151,1616]" box="[199,661,1592,1616]" pageId="28" pageNumber="29">
<emphasis id="B968EAEAFFC2FFC746F15C19C867094B" box="[199,336,1592,1616]" italics="true" pageId="28" pageNumber="29">Distribution</emphasis>
: Sabah and 
<collectingCountry id="F30B7668FFC2FFC747EF5C19CB12094B" box="[473,549,1592,1616]" name="Brunei" pageId="28" pageNumber="29">Brunei</collectingCountry>
(
<figureCitation id="13272A7DFFC2FFC744035C19CBBF094B" box="[565,648,1592,1616]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="28" pageNumber="29">Fig. 35</figureCitation>
).
</paragraph>
<paragraph id="8BA336F8FFC2FFC746A15C85C86709C5" blockId="28.[151,497,1700,1758]" pageId="28" pageNumber="29">
<taxonomicName id="4C1C4D7BFFC2FFC746A15C85C8A709A5" box="[151,400,1700,1726]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hooki" status="sp. nov.">
<emphasis id="B968EAEAFFC2FFC746A15C85C8A709A5" bold="true" box="[151,400,1700,1726]" italics="true" pageId="28" pageNumber="29">Ischnothyreus hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFC2FFC747A05C85C8C609A5" box="[406,497,1700,1726]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13272A7DFFC2FFC746A95CE7C81709C5" box="[159,288,1734,1758]" captionStart-0="FIGURE 22" captionStart-1="FIGURE 23" captionStartId-0="29.[151,250,1883,1906]" captionStartId-1="30.[151,250,1874,1897]" captionTargetBox-0="[151,1412,233,1858]" captionTargetBox-1="[151,1436,225,1833]" captionTargetId-0="figure@29.[151,1436,195,1858]" captionTargetId-1="figure@30.[151,1436,193,1833]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="FIGURE 22. Ischnothyreus hooki sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, carapace, dorsal view; F, carapace, ventral view; G, I, left palp, retrolateral view; H, J, left palp, prolateral view. stp, strong, tooth-like projection; e, embolus; l, lobe; vpr, ventral protuberance; elo, ear-like outgrowth. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." captionText-1="FIGURE 23. Ischnothyreus hooki sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, abdomen, ventral view; E, G, genital area, ventral view; F, H, genital area, dorsal view. a, apodeme; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G, H = 0.1 mm." httpUri-0="https://zenodo.org/record/278133/files/figure.png" httpUri-1="https://zenodo.org/record/278134/files/figure.png" pageId="28" pageNumber="29">Figs 22–23</figureCitation>
, 
<figureCitation id="13272A7DFFC2FFC7471D5CE7C87F09C5" box="[299,328,1734,1758]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="28" pageNumber="29">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC2FFC446A15D31C8620E24" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BA336F8FFC2FFC746A15D31CA2C08AD" blockId="28.[151,1436,1806,1974]" pageId="28" pageNumber="29">
<emphasis id="B968EAEAFFC2FFC746A15D31C89B083D" box="[151,428,1806,1831]" italics="true" pageId="28" pageNumber="29">
<typeStatus id="54A7885AFFC2FFC746A15D31C9FC083C" box="[151,203,1808,1831]" pageId="28" pageNumber="29">Type</typeStatus>
material: 
<typeStatus id="54A7885AFFC2FFC747735D2FC89B083D" box="[325,428,1806,1830]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
: male (PBI_OON 00031692), 
<collectingCountry id="F30B7668FFC2FFC745385D2FCA9C083D" box="[782,939,1806,1830]" name="Indonesia" pageId="28" pageNumber="29">INDONESIA</collectingCountry>
: East 
<taxonomicName id="4C1C4D7BFFC2FFC745C65D2FCD40083D" box="[1008,1143,1806,1830]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
: Tenggarong, light forest, leaflitter, 
<date id="FFA21038FFC2FFC747345D13C8BE0851" box="[258,393,1842,1866]" pageId="28" pageNumber="29" value="1980-08-04">4.VIII.1980</date>
, coll. Deeleman, 
<collectionCode id="ED0DAE3DFFC2FFC7447D5D12CB940851" LSID="urn:lsid:biocol.org:col:34992" box="[587,675,1843,1866]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
. 
<typeStatus id="54A7885AFFC2FFC7449B5D15CA280850" box="[685,799,1844,1867]" pageId="28" pageNumber="29" type="paratype">
<emphasis id="B968EAEAFFC2FFC7449B5D15CA280850" box="[685,799,1844,1867]" italics="true" pageId="28" pageNumber="29">Paratypes</emphasis>
</typeStatus>
: 
<specimenCount id="9D1AFD71FFC2FFC7451B5D13CA4E0851" box="[813,889,1842,1866]" pageId="28" pageNumber="29" type="male">1 male</specimenCount>
(PBI_OON 00031363), 
<collectingCountry id="F30B7668FFC2FFC742BA5D13CC1E0851" box="[1164,1321,1842,1866]" name="Indonesia" pageId="28" pageNumber="29">INDONESIA</collectingCountry>
: East 
<taxonomicName id="4C1C4D7BFFC2FFC7435C5D13C9C10875" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
: Sepaku, 
<quantity id="4CE49B1DFFC2FFC747545D77C8960875" box="[354,417,1878,1902]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="29" unit="km" value="40.0">40km</quantity>
NNW from Balikpapan, degraded primary forest, 
<date id="FFA21038FFC2FFC745E95D77CD510875" box="[991,1126,1878,1902]" pageId="28" pageNumber="29" value="1980-08-02">2.VIII.1980</date>
, coll. Deeleman, 
<collectionCode id="ED0DAE3DFFC2FFC7431D5D76CCB40875" LSID="urn:lsid:biocol.org:col:34992" box="[1323,1411,1879,1902]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
. 
<specimenCount id="9D1AFD71FFC2FFC743B85D77C9D90889" pageId="28" pageNumber="29" type="female">2 females</specimenCount>
(PBI_OON 00031769), 
<collectingCountry id="F30B7668FFC2FFC744325D5BCB960889" box="[516,673,1914,1938]" name="Indonesia" pageId="28" pageNumber="29">INDONESIA</collectingCountry>
: East 
<taxonomicName id="4C1C4D7BFFC2FFC744D45D5BCA5E0889" box="[738,873,1914,1938]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
: Sepaku, 
<quantity id="4CE49B1DFFC2FFC745E35D5BCD230889" box="[981,1044,1914,1938]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="29" unit="km" value="40.0">40km</quantity>
NNW of Balikpapan, primary forest, dry period, 21./ 
<date id="FFA21038FFC2FFC747595DBFC8CB08AD" box="[367,508,1950,1974]" pageId="28" pageNumber="29" value="1982-07-22">22.VII.1982</date>
, coll. Deeleman, 
<collectionCode id="ED0DAE3DFFC2FFC744895DBECA2008AD" LSID="urn:lsid:biocol.org:col:34992" box="[703,791,1951,1974]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
.
</paragraph>
<caption id="DF636670FFC3FFC646A15D7ACBF808D5" httpUri="https://zenodo.org/record/278133/files/figure.png" pageId="29" pageNumber="30" targetBox="[151,1412,233,1858]" targetPageId="29">
<paragraph id="8BA336F8FFC3FFC646A15D7ACBF808D5" blockId="29.[151,1436,1883,1998]" pageId="29" pageNumber="30">
<emphasis id="B968EAEAFFC3FFC646A15D7AC816086A" bold="true" box="[151,289,1883,1906]" pageId="29" pageNumber="30">FIGURE 22.</emphasis>
<taxonomicName id="4C1C4D7BFFC3FFC647115D7DC8C30869" box="[295,500,1884,1906]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="hooki" status="sp. nov.">
<emphasis id="B968EAEAFFC3FFC647115D7DC8C30869" box="[295,500,1884,1906]" italics="true" pageId="29" pageNumber="30">Ischnothyreus hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFC3FFC647CD5D7CCB7C0869" box="[507,587,1885,1906]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, carapace, dorsal view; F, carapace, ventral view; G, I, left palp, retrolateral view; H, J, left palp, prolateral view. stp, strong, tooth-like projection; e, embolus; l, lobe; vpr, ventral protuberance; elo, ear-like outgrowth. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm.
</paragraph>
</caption>
<caption id="DF636670FFC0FFC546A15D73CBE508BD" httpUri="https://zenodo.org/record/278134/files/figure.png" pageId="30" pageNumber="31" targetBox="[151,1436,225,1833]" targetPageId="30">
<paragraph id="8BA336F8FFC0FFC546A15D73CBE508BD" blockId="30.[151,1436,1874,1958]" pageId="30" pageNumber="31">
<emphasis id="B968EAEAFFC0FFC546A15D73C8280873" bold="true" box="[151,287,1874,1897]" pageId="30" pageNumber="31">FIGURE 23.</emphasis>
<taxonomicName id="4C1C4D7BFFC0FFC547135D72C8C60872" box="[293,497,1875,1897]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="hooki" status="sp. nov.">
<emphasis id="B968EAEAFFC0FFC547135D72C8C60872" box="[293,497,1875,1897]" italics="true" pageId="30" pageNumber="31">Ischnothyreus hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFC0FFC547C15D75CB710872" box="[503,582,1876,1897]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, abdomen, ventral view; E, G, genital area, ventral view; F, H, genital area, dorsal view. a, apodeme; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G, H = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA336F8FFC1FFC446F15AB9C8620E24" blockId="31.[151,1437,151,1257]" pageId="31" pageNumber="32">
<emphasis id="B968EAEAFFC1FFC446F15AB9CB170FAB" box="[199,544,152,176]" italics="true" pageId="31" pageNumber="32">Additional material examined</emphasis>
: 
<specimenCount id="9D1AFD71FFC1FFC444035AB6CBAC0FB4" box="[565,667,151,175]" pageId="31" pageNumber="32" type="female">1 female</specimenCount>
(PBI_OON 00031777), 
<collectingCountry id="F30B7668FFC1FFC445F75AB6CD690FB4" box="[961,1118,151,175]" name="Indonesia" pageId="31" pageNumber="32">INDONESIA</collectingCountry>
: East 
<taxonomicName id="4C1C4D7BFFC1FFC4429D5AB6CC050FB4" box="[1195,1330,151,175]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
: Sepaku, 
<quantity id="4CE49B1DFFC1FFC446A15A9DC9E10FCF" box="[151,214,188,212]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="32" unit="km" value="40.0">40km</quantity>
NNW of Balikpapan, degraded rainforest, litter, 
<date id="FFA21038FFC1FFC445755A9DCADA0FCF" box="[835,1005,188,212]" pageId="31" pageNumber="32" value="1980-08-02" valueMax="1980-08-05" valueMin="1980-08-02">2.–5.VIII.1980</date>
, coll. Deeleman (
<collectionCode id="ED0DAE3DFFC1FFC442F95A9DCC1D0FCF" LSID="urn:lsid:biocol.org:col:34992" box="[1231,1322,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="31" pageNumber="32">RMNH</collectionCode>
). 
<specimenCount id="9D1AFD71FFC1FFC443705A9DCCAC0FCF" box="[1350,1435,188,212]" pageId="31" pageNumber="32" type="male">1 male</specimenCount>
(PBI_OON 00031628), 
<collectingCountry id="F30B7668FFC1FFC4479E5AFECB720FEC" box="[424,581,223,247]" name="Indonesia" pageId="31" pageNumber="32">INDONESIA</collectingCountry>
: East 
<taxonomicName id="4C1C4D7BFFC1FFC444BF5AFECA270FEC" box="[649,784,223,247]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
: Sungai Wain Protection Forest, ca. 
<quantity id="4CE49B1DFFC1FFC442985AFECDC00FEC" box="[1198,1271,223,247]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="km" value="15.0">15 km</quantity>
N of Balikpapan (
<geoCoordinate id="EE28503FFFC1FFC446E55B25C87C0E07" box="[211,331,260,284]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="15" value="-1.1433333">1°08’36”S</geoCoordinate>
, 
<geoCoordinate id="EE28503FFFC1FFC4476D5B25C8C50E07" box="[347,498,260,284]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="15" value="116.849724">116°50’59”E</geoCoordinate>
), 
<quantity id="4CE49B1DFFC1FFC4443C5B25CB090E07" box="[522,574,260,284]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="31" pageNumber="32" unit="m" value="80.0">80m</quantity>
, secondary forest adjacent to primary forest, 5./ 
<date id="FFA21038FFC1FFC4425A5B25CDEE0E07" box="[1132,1241,260,284]" pageId="31" pageNumber="32" value="2008-10-07">7.X.2008</date>
, leg. P. Schwendinger (
<collectionCode id="ED0DAE3DFFC1FFC446DB5B06C87E0E24" box="[237,329,295,319]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="31" pageNumber="32">MHNG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC1FFC446F15B6DCC210E9C" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8BA336F8FFC1FFC446F15B6DCC210E9C" blockId="31.[151,1437,151,1257]" pageId="31" pageNumber="32">
<emphasis id="B968EAEAFFC1FFC446F15B6DC8080E7F" box="[199,319,332,356]" italics="true" pageId="31" pageNumber="32">Etymology</emphasis>
: The specific name is derived from Captain Hook from the novel Peter Pan, written by James M. Barrie, and refers to the similarity of the distal part of the male palp with the left arm of Captain Hook.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC1FFC446F15BB4CC5B0ED4" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BA336F8FFC1FFC446F15BB4CC5B0ED4" blockId="31.[151,1437,151,1257]" pageId="31" pageNumber="32">
<emphasis id="B968EAEAFFC1FFC446F15BB4C80E0EB7" box="[199,313,405,428]" italics="true" pageId="31" pageNumber="32">Diagnosis</emphasis>
: The male of 
<taxonomicName id="4C1C4D7BFFC1FFC447EC5BB4CB070EB7" box="[474,560,404,428]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="hooki">
<emphasis id="B968EAEAFFC1FFC447EC5BB4CB070EB7" box="[474,560,404,428]" italics="true" pageId="31" pageNumber="32">I. hooki</emphasis>
</taxonomicName>
can be recognized by the hook-like embolus (
<figureCitation id="13272A7DFFC1FFC442095BB5CDA70EB7" box="[1087,1168,404,428]" captionStart="FIGURE 22" captionStartId="29.[151,250,1883,1906]" captionTargetBox="[151,1412,233,1858]" captionTargetId="figure@29.[151,1436,195,1858]" captionTargetPageId="29" captionText="FIGURE 22. Ischnothyreus hooki sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, carapace, dorsal view; F, carapace, ventral view; G, I, left palp, retrolateral view; H, J, left palp, prolateral view. stp, strong, tooth-like projection; e, embolus; l, lobe; vpr, ventral protuberance; elo, ear-like outgrowth. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278133/files/figure.png" pageId="31" pageNumber="32">Fig. 22</figureCitation>
G–J). With its strongly winding tube, the female genital area of 
<taxonomicName id="4C1C4D7BFFC1FFC4446D5B98CB860ECB" box="[603,689,440,464]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="hooki">
<emphasis id="B968EAEAFFC1FFC4446D5B98CB860ECB" box="[603,689,440,464]" italics="true" pageId="31" pageNumber="32">I. hooki</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C1C4D7BFFC1FFC445755B98CAA10ECB" box="[835,918,440,464]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="fobor">
<emphasis id="B968EAEAFFC1FFC445755B98CAA10ECB" box="[835,918,440,464]" italics="true" pageId="31" pageNumber="32">I. fobor</emphasis>
</taxonomicName>
, but the winding tube in 
<taxonomicName id="4C1C4D7BFFC1FFC442995B98CC330ECB" box="[1199,1284,440,464]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="hooki">
<emphasis id="B968EAEAFFC1FFC442995B98CD8F0ECB" box="[1199,1208,441,464]" italics="true" pageId="31" pageNumber="32">I</emphasis>
. 
<emphasis id="B968EAEAFFC1FFC442F05B99CC330ECB" box="[1222,1284,440,464]" italics="true" pageId="31" pageNumber="32">hooki</emphasis>
</taxonomicName>
is larger.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC1FFC446F15BFCCB020B67" pageId="31" pageNumber="32" type="description">
<paragraph id="8BA336F8FFC1FFC446F15BFCCDCC0C2C" blockId="31.[151,1437,151,1257]" pageId="31" pageNumber="32">
<emphasis id="B968EAEAFFC1FFC446F15BFCC87C0EEF" box="[199,331,477,500]" italics="true" pageId="31" pageNumber="32">Description</emphasis>
: Male (
<typeStatus id="54A7885AFFC1FFC447925BFDCB3B0EEF" box="[420,524,476,500]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFC1FFC444865BFDCA2F0EEF" box="[688,792,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="31" pageNumber="32" unit="mm" value="1.33">1.33 mm</quantity>
. 
<emphasis id="B968EAEAFFC1FFC445155BFCCAA60EEF" box="[803,913,477,500]" italics="true" pageId="31" pageNumber="32">Carapace</emphasis>
: broadly oval in dorsal view. 
<emphasis id="B968EAEAFFC1FFC442D35BFDCC080EEF" box="[1253,1343,476,500]" italics="true" pageId="31" pageNumber="32">Clypeus</emphasis>
: curved downwards in frontal view, low, ALE separated from edge of carapace by less than their radius. 
<emphasis id="B968EAEAFFC1FFC442F35820CDCD0D03" box="[1221,1274,513,536]" italics="true" pageId="31" pageNumber="32">Eyes</emphasis>
: PLE circular, posterior eye row straight from above, procurved from front. 
<emphasis id="B968EAEAFFC1FFC4455F5804CAF30D27" box="[873,964,549,572]" italics="true" pageId="31" pageNumber="32">Sternum</emphasis>
: setae abundant. 
<emphasis id="B968EAEAFFC1FFC442A25805CC280D27" box="[1172,1311,548,572]" italics="true" pageId="31" pageNumber="32">Mouthparts:</emphasis>
chelicerae without prominent process at base of fangs. Labium triangular. Anteromedian tip of endites with one strong, toothlike projection (stp) (
<figureCitation id="13272A7DFFC1FFC447B5584DC8E30D9F" box="[387,468,620,644]" captionStart="FIGURE 22" captionStartId="29.[151,250,1883,1906]" captionTargetBox="[151,1412,233,1858]" captionTargetId="figure@29.[151,1436,195,1858]" captionTargetPageId="29" captionText="FIGURE 22. Ischnothyreus hooki sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, carapace, dorsal view; F, carapace, ventral view; G, I, left palp, retrolateral view; H, J, left palp, prolateral view. stp, strong, tooth-like projection; e, embolus; l, lobe; vpr, ventral protuberance; elo, ear-like outgrowth. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278133/files/figure.png" pageId="31" pageNumber="32">Fig. 22</figureCitation>
B, C, F). 
<emphasis id="B968EAEAFFC1FFC44472584DCB990D9F" box="[580,686,620,644]" italics="true" pageId="31" pageNumber="32">Abdomen</emphasis>
: scutum extending far dorsal of pedicel. 
<emphasis id="B968EAEAFFC1FFC4424F584CCD990D9F" box="[1145,1198,621,644]" italics="true" pageId="31" pageNumber="32">Legs</emphasis>
: spine formula: femora: I p0-1-1; r0-0-1; II v0-0-2; tibiae: I, II 
<date id="FFA21038FFC1FFC4444058AECB9B0DBC" box="[630,684,655,679]" pageId="31" pageNumber="32">v4-2</date>
-2; metatarsi: I, II 
<date id="FFA21038FFC1FFC4454258AECA9D0DBC" box="[884,938,655,679]" pageId="31" pageNumber="32">v2-2</date>
-0. 
<emphasis id="B968EAEAFFC1FFC445FB58B1CD000DB3" box="[973,1079,656,680]" italics="true" pageId="31" pageNumber="32">Genitalia</emphasis>
: Sperm pore situated in front of anterior spiracles. Palps strongly sclerotized, proximal segments brown, trochanter with ventral projection, patella about as long as femur, not enlarged, cymbium brown, fused with bulb, bulb brown, more than two times as long as cymbium, stout, tapering apically, with ventral protuberance (vpr) (
<figureCitation id="13272A7DFFC1FFC445A758DDCAD50C0F" box="[913,994,764,788]" captionStart="FIGURE 22" captionStartId="29.[151,250,1883,1906]" captionTargetBox="[151,1412,233,1858]" captionTargetId="figure@29.[151,1436,195,1858]" captionTargetPageId="29" captionText="FIGURE 22. Ischnothyreus hooki sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, carapace, dorsal view; F, carapace, ventral view; G, I, left palp, retrolateral view; H, J, left palp, prolateral view. stp, strong, tooth-like projection; e, embolus; l, lobe; vpr, ventral protuberance; elo, ear-like outgrowth. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278133/files/figure.png" pageId="31" pageNumber="32">Fig. 22</figureCitation>
H, J) and lobe (l) (
<figureCitation id="13272A7DFFC1FFC4428B58DDCC250C0F" box="[1213,1298,764,788]" captionStart="FIGURE 22" captionStartId="29.[151,250,1883,1906]" captionTargetBox="[151,1412,233,1858]" captionTargetId="figure@29.[151,1436,195,1858]" captionTargetPageId="29" captionText="FIGURE 22. Ischnothyreus hooki sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, carapace, dorsal view; F, carapace, ventral view; G, I, left palp, retrolateral view; H, J, left palp, prolateral view. stp, strong, tooth-like projection; e, embolus; l, lobe; vpr, ventral protuberance; elo, ear-like outgrowth. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278133/files/figure.png" pageId="31" pageNumber="32">Fig. 22</figureCitation>
. G, I), distal part with ear-like outgrowth (elo) prolaterally (
<figureCitation id="13272A7DFFC1FFC44492593ECBC30C2C" box="[676,756,799,823]" captionStart="FIGURE 22" captionStartId="29.[151,250,1883,1906]" captionTargetBox="[151,1412,233,1858]" captionTargetId="figure@29.[151,1436,195,1858]" captionTargetPageId="29" captionText="FIGURE 22. Ischnothyreus hooki sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, carapace, dorsal view; F, carapace, ventral view; G, I, left palp, retrolateral view; H, J, left palp, prolateral view. stp, strong, tooth-like projection; e, embolus; l, lobe; vpr, ventral protuberance; elo, ear-like outgrowth. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278133/files/figure.png" pageId="31" pageNumber="32">Fig. 22</figureCitation>
H, J), embolus (e) short hook (
<figureCitation id="13272A7DFFC1FFC44264593ECD940C2C" box="[1106,1187,799,823]" captionStart="FIGURE 22" captionStartId="29.[151,250,1883,1906]" captionTargetBox="[151,1412,233,1858]" captionTargetId="figure@29.[151,1436,195,1858]" captionTargetPageId="29" captionText="FIGURE 22. Ischnothyreus hooki sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, carapace, dorsal view; F, carapace, ventral view; G, I, left palp, retrolateral view; H, J, left palp, prolateral view. stp, strong, tooth-like projection; e, embolus; l, lobe; vpr, ventral protuberance; elo, ear-like outgrowth. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278133/files/figure.png" pageId="31" pageNumber="32">Fig. 22</figureCitation>
G, I, J).
</paragraph>
<paragraph id="8BA336F8FFC1FFC446F15965CB020B67" blockId="31.[151,1437,151,1257]" pageId="31" pageNumber="32">
Female (
<typeStatus id="54A7885AFFC1FFC4471C5965C8A60C47" box="[298,401,836,860]" pageId="31" pageNumber="32" type="paratype">paratype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFC1FFC444015965CBA80C47" box="[567,671,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="31" pageNumber="32" unit="mm" value="1.8">1.80 mm</quantity>
. As in male except as noted. 
<emphasis id="B968EAEAFFC1FFC445C45964CD570C47" box="[1010,1120,837,860]" italics="true" pageId="31" pageNumber="32">Carapace</emphasis>
: without any pattern. 
<emphasis id="B968EAEAFFC1FFC443695964CCA30C47" box="[1375,1428,837,860]" italics="true" pageId="31" pageNumber="32">Eyes</emphasis>
: posterior eye row procurved from above. 
<emphasis id="B968EAEAFFC1FFC4445D5949CBC10C9B" box="[619,758,872,896]" italics="true" pageId="31" pageNumber="32">Mouthparts:</emphasis>
endites unmodified. Femur of female palp with four ventral spines, tibia with three trichobothria. 
<emphasis id="B968EAEAFFC1FFC4440159ADCB960CBF" box="[567,673,908,932]" italics="true" pageId="31" pageNumber="32">Abdomen</emphasis>
: scutum not extending far dorsal of pedicel. Dorsal scutum covering less than 1/2 of abdomen, less than 1/4 abdomen width, not fused to epigastric scutum. Postepigastric scutum widely hexagonal, only around epigastric furrow, with short posteriorly directed lateral apodemes (a) (
<figureCitation id="13272A7DFFC1FFC4431759F5CC450CF7" box="[1313,1394,980,1004]" captionStart="FIGURE 23" captionStartId="30.[151,250,1874,1897]" captionTargetBox="[151,1436,225,1833]" captionTargetId="figure@30.[151,1436,193,1833]" captionTargetPageId="30" captionText="FIGURE 23. Ischnothyreus hooki sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, abdomen, ventral view; E, G, genital area, ventral view; F, H, genital area, dorsal view. a, apodeme; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G, H = 0.1 mm." httpUri="https://zenodo.org/record/278134/files/figure.png" pageId="31" pageNumber="32">Fig. 23</figureCitation>
E). Dense patch of setae anterior to spinnerets present. 
<emphasis id="B968EAEAFFC1FFC444EE59D8CA3A0B0B" box="[728,781,1017,1040]" italics="true" pageId="31" pageNumber="32">Legs</emphasis>
: spine formula: femora: I p0-2-1; r0-0-1; II p0-0-2; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFC1FFC444F95E3DCA0F0B2F" box="[719,824,1052,1076]" italics="true" pageId="31" pageNumber="32">Genitalia</emphasis>
: the posterior margin of the epigastric scutum is lined with numerous needle-like setae. In the middle of the postepigastric scutum is a strongly winding tube (wt) (
<figureCitation id="13272A7DFFC1FFC4437A5E61CCAB0B43" box="[1356,1436,1088,1112]" captionStart="FIGURE 23" captionStartId="30.[151,250,1874,1897]" captionTargetBox="[151,1436,225,1833]" captionTargetId="figure@30.[151,1436,193,1833]" captionTargetPageId="30" captionText="FIGURE 23. Ischnothyreus hooki sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, abdomen, ventral view; E, G, genital area, ventral view; F, H, genital area, dorsal view. a, apodeme; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G, H = 0.1 mm." httpUri="https://zenodo.org/record/278134/files/figure.png" pageId="31" pageNumber="32">Fig. 23</figureCitation>
E–H) visible, which runs posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC1FFC446F15EA9CBBF0BBB" box="[199,648,1160,1184]" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8BA336F8FFC1FFC446F15EA9CBBF0BBB" blockId="31.[151,1437,151,1257]" box="[199,648,1160,1184]" pageId="31" pageNumber="32">
<emphasis id="B968EAEAFFC1FFC446F15EA9C8670BBB" box="[199,336,1160,1184]" italics="true" pageId="31" pageNumber="32">Distribution</emphasis>
: East 
<taxonomicName id="4C1C4D7BFFC1FFC447A05EA9CB2E0BBB" box="[406,537,1160,1184]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
(
<figureCitation id="13272A7DFFC1FFC4441E5EA9CB4C0BBB" box="[552,635,1160,1184]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="31" pageNumber="32">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC1FFC446F15E8DC8670A6A" pageId="31" pageNumber="32" type="discussion">
<paragraph id="8BA336F8FFC1FFC446F15E8DCA760BF3" blockId="31.[151,1437,151,1257]" pageId="31" pageNumber="32">
<emphasis id="B968EAEAFFC1FFC446F15E8DC81E0BDF" box="[199,297,1196,1220]" italics="true" pageId="31" pageNumber="32">Remarks</emphasis>
: Male and female of this species were not collected together. However, since the somatic morphology is very similar they are tentatively considered as conspecific.
</paragraph>
<paragraph id="8BA336F8FFC1FFC446A15F17C8670A6A" blockId="31.[151,474,1334,1393]" pageId="31" pageNumber="32">
<taxonomicName id="4C1C4D7BFFC1FFC446A15F17C84E0A4B" box="[151,377,1334,1360]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="jojo" status="sp. nov.">
<emphasis id="B968EAEAFFC1FFC446A15F17C84E0A4B" bold="true" box="[151,377,1334,1360]" italics="true" pageId="31" pageNumber="32">Ischnothyreus jojo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFC1FFC447B65F17C8ED0A4B" box="[384,474,1334,1360]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13272A7DFFC1FFC446A95F78C8170A6A" box="[159,288,1369,1393]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStartId-0="32.[151,250,1855,1878]" captionStartId-1="33.[151,250,1860,1883]" captionTargetBox-0="[151,1418,205,1828]" captionTargetBox-1="[151,1434,204,1830]" captionTargetId-0="figure@32.[151,1436,193,1828]" captionTargetId-1="figure@33.[151,1436,193,1831]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionText-0="FIGURE 24. Ischnothyreus jojo sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, anterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, toothlike projection; pbp, plate-like basal process; sse, saw-shaped extension; vp, ventral projection; vpr, ventral protuberance; l, lobe; spa, small pointed apophysis. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm." captionText-1="FIGURE 25. Ischnothyreus jojo sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, abdomen, ventral view; E, F, genital area, ventral view; G, genital area, dorsal view. cse, curved, sclerotized extensions; a, apodeme; uss, u-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm." httpUri-0="https://zenodo.org/record/278135/files/figure.png" httpUri-1="https://zenodo.org/record/278136/files/figure.png" pageId="31" pageNumber="32">Figs 24–25</figureCitation>
, 
<figureCitation id="13272A7DFFC1FFC4471D5F78C87F0A6A" box="[299,328,1369,1393]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="31" pageNumber="32">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC1FFC446A15FBCC832097C" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BA336F8FFC1FFC446A15FBCCD0D0AE7" blockId="31.[151,1437,1436,2036]" pageId="31" pageNumber="32">
<emphasis id="B968EAEAFFC1FFC446A15FBCC8820AAF" box="[151,437,1436,1460]" italics="true" pageId="31" pageNumber="32">
<typeStatus id="54A7885AFFC1FFC446A15FBCC9FC0AAF" box="[151,203,1437,1460]" pageId="31" pageNumber="32">Type</typeStatus>
material: 
<typeStatus id="54A7885AFFC1FFC447795FBDC8820AAF" box="[335,437,1436,1460]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
</emphasis>
: male (PBI_OON 00032184), 
<collectingCountry id="F30B7668FFC1FFC4451C5FBDCAF00AAF" box="[810,967,1436,1460]" name="Indonesia" pageId="31" pageNumber="32">INDONESIA</collectingCountry>
: Central 
<taxonomicName id="4C1C4D7BFFC1FFC4420E5FBDCD880AAF" box="[1080,1215,1436,1460]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
: Kaharian, 
<geoCoordinate id="EE28503FFFC1FFC4437D5FBDCCA00AAF" box="[1355,1431,1436,1460]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="925" value="-2.0333333">2°02'S</geoCoordinate>
, 
<geoCoordinate id="EE28503FFFC1FFC446A15F9EC8370ACC" box="[151,256,1471,1495]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="925" value="113.666664">113°40'E</geoCoordinate>
, 
<date id="FFA21038FFC1FFC447395F9EC8840ACC" box="[271,435,1471,1495]" pageId="31" pageNumber="32" value="1985-09-02" valueMax="1985-09-16" valueMin="1985-09-02">2.–16.IX.1985</date>
, swampy primary forest, leaflitter, leg. Suh. Djojosudharmo, coll. Deeleman, 
<collectionCode id="ED0DAE3DFFC1FFC443775FE1CCAE0ACC" LSID="urn:lsid:biocol.org:col:34992" box="[1345,1433,1472,1495]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="31" pageNumber="32">RMNH</collectionCode>
. 
<typeStatus id="54A7885AFFC1FFC446A15FC4C83E0AE7" box="[151,265,1509,1532]" pageId="31" pageNumber="32" type="paratype">
<emphasis id="B968EAEAFFC1FFC446A15FC4C83E0AE7" box="[151,265,1509,1532]" italics="true" pageId="31" pageNumber="32">Paratypes</emphasis>
</typeStatus>
: 
<specimenCount id="9D1AFD71FFC1FFC447215FC5C8B40AE7" box="[279,387,1508,1532]" pageId="31" pageNumber="32" type="female">3 females</specimenCount>
(PBI_OON32181), collected together with 
<typeStatus id="54A7885AFFC1FFC445595FC5CAE30AE7" box="[879,980,1508,1532]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED0DAE3DFFC1FFC445E95FC4CD000AE7" LSID="urn:lsid:biocol.org:col:34992" box="[991,1079,1509,1532]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="31" pageNumber="32">RMNH</collectionCode>
.
</paragraph>
<paragraph id="8BA336F8FFC1FFC446F15C29C832097C" blockId="31.[151,1437,1436,2036]" pageId="31" pageNumber="32">
<emphasis id="B968EAEAFFC1FFC446F15C29CB23093B" box="[199,532,1544,1568]" italics="true" pageId="31" pageNumber="32">Additional material examined</emphasis>
: 
<specimenCount id="9D1AFD71FFC1FFC444145C26CBB90904" box="[546,654,1543,1567]" pageId="31" pageNumber="32" type="female">2 females</specimenCount>
(PBI_OON 00031468), 
<collectingCountry id="F30B7668FFC1FFC445975C26CD090904" box="[929,1086,1543,1567]" name="Indonesia" pageId="31" pageNumber="32">INDONESIA</collectingCountry>
: Central 
<taxonomicName id="4C1C4D7BFFC1FFC442975C26CC1F0904" box="[1185,1320,1543,1567]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
: Tumbang Tahai, 
<geoCoordinate id="EE28503FFFC1FFC446DA5C0DC80F095F" box="[236,312,1580,1604]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="925" value="-2.0333333">2°02'S</geoCoordinate>
, 
<geoCoordinate id="EE28503FFFC1FFC4477B5C0DC882095F" box="[333,437,1580,1604]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="925" value="113.583336">113°35'E</geoCoordinate>
, primary marshy forest, 
<date id="FFA21038FFC1FFC444DB5C0DCAA5095F" box="[749,914,1580,1604]" pageId="31" pageNumber="32" value="1985-09-02" valueMax="1985-09-13" valueMin="1985-09-02">2.–13.IX.1985</date>
, coll. Deeleman, leg. Suh. Djojosudharmo (
<collectionCode id="ED0DAE3DFFC1FFC446A95C6EC9CD097C" LSID="urn:lsid:biocol.org:col:34992" box="[159,250,1615,1639]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="31" pageNumber="32">RMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC1FFC446F15C55CAF60997" box="[199,961,1652,1676]" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8BA336F8FFC1FFC446F15C55CAF60997" blockId="31.[151,1437,1436,2036]" box="[199,961,1652,1676]" pageId="31" pageNumber="32">
<emphasis id="B968EAEAFFC1FFC446F15C55C8080997" box="[199,319,1652,1676]" italics="true" pageId="31" pageNumber="32">Etymology</emphasis>
: The specific name is an arbitrary combination of letters.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC1FFC446F15CB8CD9209EC" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BA336F8FFC1FFC446F15CB8CD9209EC" blockId="31.[151,1437,1436,2036]" pageId="31" pageNumber="32">
<emphasis id="B968EAEAFFC1FFC446F15CB8C80E09AB" box="[199,313,1689,1712]" italics="true" pageId="31" pageNumber="32">Diagnosis</emphasis>
: The male of 
<taxonomicName id="4C1C4D7BFFC1FFC447EE5CB8CB2C09AB" box="[472,539,1689,1712]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="jojo">
<emphasis id="B968EAEAFFC1FFC447EE5CB8CB2C09AB" box="[472,539,1689,1712]" italics="true" pageId="31" pageNumber="32">I. jojo</emphasis>
</taxonomicName>
can be recognized by the plate-like basal process and the saw-like extension at the base of the cheliceral fang (
<figureCitation id="13272A7DFFC1FFC4443E5C9DCB6C09CF" box="[520,603,1724,1748]" captionStart="FIGURE 24" captionStartId="32.[151,250,1855,1878]" captionTargetBox="[151,1418,205,1828]" captionTargetId="figure@32.[151,1436,193,1828]" captionTargetPageId="32" captionText="FIGURE 24. Ischnothyreus jojo sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, anterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, toothlike projection; pbp, plate-like basal process; sse, saw-shaped extension; vp, ventral projection; vpr, ventral protuberance; l, lobe; spa, small pointed apophysis. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278135/files/figure.png" pageId="31" pageNumber="32">Fig. 24</figureCitation>
E). The female genital area is characterized by the posteriorly running, strongly winding tube, anteriorly covered by a u-shaped, sclerotized structure (
<figureCitation id="13272A7DFFC1FFC4423F5CFECD6E09EC" box="[1033,1113,1759,1783]" captionStart="FIGURE 25" captionStartId="33.[151,250,1860,1883]" captionTargetBox="[151,1434,204,1830]" captionTargetId="figure@33.[151,1436,193,1831]" captionTargetPageId="33" captionText="FIGURE 25. Ischnothyreus jojo sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, abdomen, ventral view; E, F, genital area, ventral view; G, genital area, dorsal view. cse, curved, sclerotized extensions; a, apodeme; uss, u-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278136/files/figure.png" pageId="31" pageNumber="32">Fig. 25</figureCitation>
E–G).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFC1FFF946F15D24CBB30DC1" lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32" type="description">
<paragraph id="8BA336F8FFC1FFF946F15D24CD3B0E0C" blockId="31.[151,1437,1436,2036]" lastBlockId="34.[151,1437,151,765]" lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32">
<emphasis id="B968EAEAFFC1FFC446F15D24C87C0807" box="[199,331,1797,1820]" italics="true" pageId="31" pageNumber="32">Description</emphasis>
: Male (
<typeStatus id="54A7885AFFC1FFC447935D25CB3A0807" box="[421,525,1796,1820]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFC1FFC444875D25CA2D0807" box="[689,794,1796,1820]" metricMagnitude="-3" metricUnit="m" metricValue="1.76" pageId="31" pageNumber="32" unit="mm" value="1.76">1.76 mm</quantity>
. 
<emphasis id="B968EAEAFFC1FFC445105D25CAB70807" box="[806,896,1796,1820]" italics="true" pageId="31" pageNumber="32">Clypeus</emphasis>
: low, ALE separated from edge of carapace by less than their radius. 
<emphasis id="B968EAEAFFC1FFC447BB5D08C8F5085B" box="[397,450,1833,1856]" italics="true" pageId="31" pageNumber="32">Eyes</emphasis>
: PME largest. 
<emphasis id="B968EAEAFFC1FFC4445D5D09CBD9085B" box="[619,750,1832,1856]" italics="true" pageId="31" pageNumber="32">Mouthparts</emphasis>
: chelicerae slightly divergent, process at base of fangs platelike (pbp), with inwards pointing hook distally, and with a saw-shaped extension (sse), fang groove with few small and one larger denticles (
<figureCitation id="13272A7DFFC1FFC447875D51CB360893" box="[433,513,1904,1928]" captionStart="FIGURE 24" captionStartId="32.[151,250,1855,1878]" captionTargetBox="[151,1418,205,1828]" captionTargetId="figure@32.[151,1436,193,1828]" captionTargetPageId="32" captionText="FIGURE 24. Ischnothyreus jojo sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, anterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, toothlike projection; pbp, plate-like basal process; sse, saw-shaped extension; vp, ventral projection; vpr, ventral protuberance; l, lobe; spa, small pointed apophysis. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278135/files/figure.png" pageId="31" pageNumber="32">Fig. 24</figureCitation>
D–E). Anterior margin of labium indented at middle. Anteromedian tip of endites with one strong, tooth-like projection (stp) (
<figureCitation id="13272A7DFFC1FFC444AB5DB5CBC508B7" box="[669,754,1940,1964]" captionStart="FIGURE 24" captionStartId="32.[151,250,1855,1878]" captionTargetBox="[151,1418,205,1828]" captionTargetId="figure@32.[151,1436,193,1828]" captionTargetPageId="32" captionText="FIGURE 24. Ischnothyreus jojo sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, anterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, toothlike projection; pbp, plate-like basal process; sse, saw-shaped extension; vp, ventral projection; vpr, ventral protuberance; l, lobe; spa, small pointed apophysis. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278135/files/figure.png" pageId="31" pageNumber="32">Fig. 24</figureCitation>
B). 
<emphasis id="B968EAEAFFC1FFC4451A5DB5CAA108B7" box="[812,918,1940,1964]" italics="true" pageId="31" pageNumber="32">Abdomen</emphasis>
: epigastric and postepigastric scutum fused. Postepigastric scutum long, almost rectangular, covering about 3/4 of abdominal length. 
<emphasis id="B968EAEAFFC1FFC4429A5D98CDD408CB" box="[1196,1251,1977,2000]" italics="true" pageId="31" pageNumber="32">Legs</emphasis>
: spine formula: femora: I p0-1-1; r0-0-2; II p0-0-1; r0-0-1; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFC1FFC442C45DFDCC6B08EF" box="[1266,1372,2012,2036]" italics="true" pageId="31" pageNumber="32">Genitalia</emphasis>
: Palp strongly sclerotized, proximal segments brown, trochanter with small ventral projection (vp) (
<figureCitation id="13272A7DFFFCFFF942E45AB6CC120FB4" box="[1234,1317,151,175]" captionStart="FIGURE 24" captionStartId="32.[151,250,1855,1878]" captionTargetBox="[151,1418,205,1828]" captionTargetId="figure@32.[151,1436,193,1828]" captionTargetPageId="32" captionText="FIGURE 24. Ischnothyreus jojo sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, anterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, toothlike projection; pbp, plate-like basal process; sse, saw-shaped extension; vp, ventral projection; vpr, ventral protuberance; l, lobe; spa, small pointed apophysis. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278135/files/figure.png" pageId="34" pageNumber="35">Fig. 24</figureCitation>
F), femur normal size, patella about as long as femur, not enlarged, cymbium brown, fused with bulb, bulb brown, 1 to 1.5 times as long as cymbium, stout, tapering apically, with ventral protuberance (vpr) and lobe (l), apex of bulb roundish with small pointed apophyses (spa), embolus not discernible (
<figureCitation id="13272A7DFFFCFFF945445ADECAF40E0C" box="[882,963,255,279]" captionStart="FIGURE 24" captionStartId="32.[151,250,1855,1878]" captionTargetBox="[151,1418,205,1828]" captionTargetId="figure@32.[151,1436,193,1828]" captionTargetPageId="32" captionText="FIGURE 24. Ischnothyreus jojo sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, anterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, toothlike projection; pbp, plate-like basal process; sse, saw-shaped extension; vp, ventral projection; vpr, ventral protuberance; l, lobe; spa, small pointed apophysis. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278135/files/figure.png" pageId="34" pageNumber="35">Fig. 24</figureCitation>
F–G).
</paragraph>
<caption id="DF636670FFFEFFFB46A15D1ECD6B08A9" httpUri="https://zenodo.org/record/278135/files/figure.png" pageId="32" pageNumber="33" targetBox="[151,1418,205,1828]" targetPageId="32">
<paragraph id="8BA336F8FFFEFFFB46A15D1ECD6B08A9" blockId="32.[151,1436,1855,1970]" pageId="32" pageNumber="33">
<emphasis id="B968EAEAFFFEFFFB46A15D1EC828084E" bold="true" box="[151,287,1855,1878]" pageId="32" pageNumber="33">FIGURE 24.</emphasis>
<taxonomicName id="4C1C4D7BFFFEFFFB47125D61C8E9084D" box="[292,478,1856,1878]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="jojo" status="sp. nov.">
<emphasis id="B968EAEAFFFEFFFB47125D61C8E9084D" box="[292,478,1856,1878]" italics="true" pageId="32" pageNumber="33">Ischnothyreus jojo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFFEFFFB47D25D60CB05084D" box="[484,562,1857,1878]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, anterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, toothlike projection; pbp, plate-like basal process; sse, saw-shaped extension; vp, ventral projection; vpr, ventral protuberance; l, lobe; spa, small pointed apophysis. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm.
</paragraph>
</caption>
<caption id="DF636670FFFFFFFA46A15D65CDD4088C" httpUri="https://zenodo.org/record/278136/files/figure.png" pageId="33" pageNumber="34" targetBox="[151,1434,204,1830]" targetPageId="33">
<paragraph id="8BA336F8FFFFFFFA46A15D65CDD4088C" blockId="33.[151,1436,1860,1943]" pageId="33" pageNumber="34">
<emphasis id="B968EAEAFFFFFFFA46A15D65C8170842" bold="true" box="[151,288,1860,1883]" pageId="33" pageNumber="34">FIGURE 25.</emphasis>
<taxonomicName id="4C1C4D7BFFFFFFFA47115D65C8D40841" box="[295,483,1860,1882]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="jojo" status="sp. nov.">
<emphasis id="B968EAEAFFFFFFFA47115D65C8D40841" box="[295,483,1860,1882]" italics="true" pageId="33" pageNumber="34">Ischnothyreus jojo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFFFFFFA47DC5D64CB0E0841" box="[490,569,1861,1882]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, abdomen, ventral view; E, F, genital area, ventral view; G, genital area, dorsal view. cse, curved, sclerotized extensions; a, apodeme; uss, u-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA336F8FFFCFFF946F15B03CBB30DC1" blockId="34.[151,1437,151,765]" pageId="34" pageNumber="35">
Female (
<typeStatus id="54A7885AFFFCFFF9471E5B03C8B80E21" box="[296,399,290,314]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFFCFFF944065B03CBA00E21" box="[560,663,290,314]" metricMagnitude="-3" metricUnit="m" metricValue="2.13" pageId="34" pageNumber="35" unit="mm" value="2.13">2.13 mm</quantity>
. As in male except as noted. 
<emphasis id="B968EAEAFFFCFFF945D65B02CD790E21" box="[992,1102,291,314]" italics="true" pageId="34" pageNumber="35">Carapace</emphasis>
: without any pattern, broadly oval in dorsal view, pars cephalica slightly elevated in lateral view. 
<emphasis id="B968EAEAFFFCFFF945955B64CACA0E46" box="[931,1021,325,349]" italics="true" pageId="34" pageNumber="35">Clypeus</emphasis>
: curved downwards in frontal view. 
<emphasis id="B968EAEAFFFCFFF946A15B48C9FB0E9B" box="[151,204,361,384]" italics="true" pageId="34" pageNumber="35">Eyes</emphasis>
: posterior eye row straight from above. 
<emphasis id="B968EAEAFFFCFFF944935B49CA370E64" box="[677,768,360,383]" italics="true" pageId="34" pageNumber="35">Sternum</emphasis>
: setae abundant. 
<emphasis id="B968EAEAFFFCFFF945F85B49CD6D0E9B" box="[974,1114,360,384]" italics="true" pageId="34" pageNumber="35">Mouthparts:</emphasis>
chelicerae straight; without modification Labium triangular. Endites unmodified. Femur of female palp with four ventral spines, tibia with three trichobothria. 
<emphasis id="B968EAEAFFFCFFF947405B8CC8D70EDE" box="[374,480,429,453]" italics="true" pageId="34" pageNumber="35">Abdomen</emphasis>
: dorsal scutum covering less than 1/2 of abdomen, less than 1/4 abdomen width, not fused to epigastric scutum. Epigastric scutum slightly protruding. Postepigastric scutum widely hexagonal, only around epigastric furrow, not fused to epigastric scutum, with short posteriorly directed lateral apodemes (a) (
<figureCitation id="13272A7DFFFCFFF943585BD3C9840D36" captionStart="FIGURE 25" captionStartId="33.[151,250,1860,1883]" captionTargetBox="[151,1434,204,1830]" captionTargetId="figure@33.[151,1436,193,1831]" captionTargetPageId="33" captionText="FIGURE 25. Ischnothyreus jojo sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, abdomen, ventral view; E, F, genital area, ventral view; G, genital area, dorsal view. cse, curved, sclerotized extensions; a, apodeme; uss, u-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278136/files/figure.png" pageId="34" pageNumber="35">Fig. 25</figureCitation>
E–G). Dense patch of setae anterior to spinnerets present. 
<emphasis id="B968EAEAFFFCFFF945765837CA420D36" box="[832,885,534,557]" italics="true" pageId="34" pageNumber="35">Legs</emphasis>
: spine formula: femora: I p0-1-2; r0-0-2; II p0-0- 1; r0-0-1; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFFCFFF945B05819CAC70D4B" box="[902,1008,568,592]" italics="true" pageId="34" pageNumber="35">Genitalia</emphasis>
: the posterior margin of the epigastric scutum is lined with numerous, long, needle-like setae. The epigastric furrow is narrow. The margin of the postepigastric scutum is slightly thickened. In its middle starts a dark thin, winding tube (wt) running posteriorly. The tube is covered ventrally by a sclerotized u-shaped structure (uss). On both sides of the tube are small, curved sclerotized extensions (cse) (
<figureCitation id="13272A7DFFFCFFF947A158E3C8DF0DC1" box="[407,488,706,730]" captionStart="FIGURE 25" captionStartId="33.[151,250,1860,1883]" captionTargetBox="[151,1434,204,1830]" captionTargetId="figure@33.[151,1436,193,1831]" captionTargetPageId="33" captionText="FIGURE 25. Ischnothyreus jojo sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, abdomen, ventral view; E, F, genital area, ventral view; G, genital area, dorsal view. cse, curved, sclerotized extensions; a, apodeme; uss, u-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G = 0.1 mm." httpUri="https://zenodo.org/record/278136/files/figure.png" pageId="34" pageNumber="35">Fig. 25</figureCitation>
E–G) visible.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFFCFFF946F158C4C8670C9D" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BA336F8FFFCFFF946F158C4CB9C0DE6" blockId="34.[151,1437,151,765]" box="[199,683,741,765]" pageId="34" pageNumber="35">
<emphasis id="B968EAEAFFFCFFF946F158C4C8670DE6" box="[199,336,741,765]" italics="true" pageId="34" pageNumber="35">Distribution</emphasis>
: Central 
<taxonomicName id="4C1C4D7BFFFCFFF9478E58C4CB0C0DE6" box="[440,571,741,765]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
(
<figureCitation id="13272A7DFFFCFFF9447C58C4CBAA0DE6" box="[586,669,741,765]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="34" pageNumber="35">Fig. 35</figureCitation>
).
</paragraph>
<paragraph id="8BA336F8FFFCFFF946A1596DC8670C9D" blockId="34.[151,567,844,902]" pageId="34" pageNumber="35">
<taxonomicName id="4C1C4D7BFFFCFFF946A1596DC8E20C7D" box="[151,469,844,870]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="kalimantan" status="sp. nov.">
<emphasis id="B968EAEAFFFCFFF946A1596DC8E20C7D" bold="true" box="[151,469,844,870]" italics="true" pageId="34" pageNumber="35">Ischnothyreus kalimantan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFFCFFF947EA596DCB000C7D" box="[476,567,844,870]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13272A7DFFFCFFF946A9594FC8170C9D" box="[159,288,878,902]" captionStart-0="FIGURE 26" captionStart-1="FIGURE 27" captionStartId-0="35.[151,250,1857,1880]" captionStartId-1="36.[151,250,1893,1916]" captionTargetBox-0="[151,1414,183,1816]" captionTargetBox-1="[151,1419,193,1855]" captionTargetId-0="figure@35.[151,1436,167,1816]" captionTargetId-1="figure@36.[151,1436,193,1856]" captionTargetPageId-0="35" captionTargetPageId-1="36" captionText-0="FIGURE 26. Ischnothyreus kalimantan sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, tooth-like projection; ssp, slightly sclerotized process; m, membrane; l, lobe; spa, small pointed apophysis; vp, ventral projection; vpr, ventral protuberance. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F = 0.1 mm; G = 0.05 mm." captionText-1="FIGURE 27. Ischnothyreus kalimantan sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, abdomen, ventral view; F, G, genital area, ventral view; H, genital area, dorsal view. a, apodeme; wt, winding tube; ts, translucent sluice. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; F, G, H = 0.1 mm." httpUri-0="https://zenodo.org/record/278137/files/figure.png" httpUri-1="https://zenodo.org/record/278138/files/figure.png" pageId="34" pageNumber="35">Figs 26–27</figureCitation>
, 
<figureCitation id="13272A7DFFFCFFF9471D594FC87F0C9D" box="[299,328,878,902]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="34" pageNumber="35">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFFCFFF946A15993CDB50B17" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BA336F8FFFCFFF946A15993CDB50B17" blockId="34.[151,1437,945,2003]" pageId="34" pageNumber="35">
<emphasis id="B968EAEAFFFCFFF946A15993C8820CD2" box="[151,437,945,969]" italics="true" pageId="34" pageNumber="35">
<typeStatus id="54A7885AFFFCFFF946A15993C9FC0CD2" box="[151,203,946,969]" pageId="34" pageNumber="35">Type</typeStatus>
material: 
<typeStatus id="54A7885AFFFCFFF947795990C8820CD2" box="[335,437,945,969]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
</emphasis>
: male (PBI_OON 00032180), 
<collectingCountry id="F30B7668FFFCFFF9451C5990CAF00CD2" box="[810,967,945,969]" name="Indonesia" pageId="34" pageNumber="35">INDONESIA</collectingCountry>
: Central 
<taxonomicName id="4C1C4D7BFFFCFFF9420E5990CD880CD2" box="[1080,1215,945,969]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
: Kaharian, 
<geoCoordinate id="EE28503FFFFCFFF9437D5990CCA00CD2" box="[1355,1431,945,969]" direction="south" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="-2.0333333">2°02'S</geoCoordinate>
, 
<geoCoordinate id="EE28503FFFFCFFF946A159F3C8370CF1" box="[151,256,978,1002]" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="925" value="113.666664">113°40'E</geoCoordinate>
, 
<date id="FFA21038FFFCFFF9473959F3C8840CF1" box="[271,435,978,1002]" pageId="34" pageNumber="35" value="1985-09-02" valueMax="1985-09-16" valueMin="1985-09-02">2.–16.IX.1985</date>
, swampy primary forest, leaflitter, leg. Suh. Djojosudharmo, coll. Deeleman, 
<collectionCode id="ED0DAE3DFFFCFFF9437759F2CCAE0CF1" LSID="urn:lsid:biocol.org:col:34992" box="[1345,1433,979,1002]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="34" pageNumber="35">RMNH</collectionCode>
. 
<typeStatus id="54A7885AFFFCFFF946A159D4C83E0B17" box="[151,265,1013,1036]" pageId="34" pageNumber="35" type="paratype">
<emphasis id="B968EAEAFFFCFFF946A159D4C83E0B17" box="[151,265,1013,1036]" italics="true" pageId="34" pageNumber="35">Paratypes</emphasis>
</typeStatus>
: 
<specimenCount id="9D1AFD71FFFCFFF9472159D5C85A0B17" box="[279,365,1012,1036]" pageId="34" pageNumber="35" type="male">2 males</specimenCount>
&amp; 
<specimenCount id="9D1AFD71FFFCFFF947A759D5C8CA0B17" box="[401,509,1012,1036]" pageId="34" pageNumber="35" type="female">5 females</specimenCount>
(PBI_OON 00032183), collected with 
<typeStatus id="54A7885AFFFCFFF9458159D5CD2B0B17" box="[951,1052,1012,1036]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED0DAE3DFFFCFFF9421159D4CD480B17" LSID="urn:lsid:biocol.org:col:34992" box="[1063,1151,1013,1036]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="34" pageNumber="35">RMNH</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFFCFFF946F15E34CB3E0B55" pageId="34" pageNumber="35" type="etymology">
<paragraph id="8BA336F8FFFCFFF946F15E34CB3E0B55" blockId="34.[151,1437,945,2003]" pageId="34" pageNumber="35">
<emphasis id="B968EAEAFFFCFFF946F15E34C8080B36" box="[199,319,1045,1069]" italics="true" pageId="34" pageNumber="35">Etymology</emphasis>
: The specific name is a noun in apposition and refers to the Indonesian name of Borneo, the island where this species was collected.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFFCFFF946F15E78CDBD0BAE" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="8BA336F8FFFCFFF946F15E78CDBD0BAE" blockId="34.[151,1437,945,2003]" pageId="34" pageNumber="35">
<emphasis id="B968EAEAFFFCFFF946F15E78C80E0B6B" box="[199,313,1113,1136]" italics="true" pageId="34" pageNumber="35">Diagnosis</emphasis>
: The male of 
<taxonomicName id="4C1C4D7BFFFCFFF947E55E78CB5F0B6B" box="[467,616,1112,1136]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="kalimantan">
<emphasis id="B968EAEAFFFCFFF947E55E78CB5F0B6B" box="[467,616,1112,1136]" italics="true" pageId="34" pageNumber="35">I. kalimantan</emphasis>
</taxonomicName>
can be recognized by the distal part of the palpal bulb, which is blunt and entirely surrounded by a translucent membrane (
<figureCitation id="13272A7DFFFCFFF944885E5BCA380B89" box="[702,783,1146,1170]" captionStart="FIGURE 26" captionStartId="35.[151,250,1857,1880]" captionTargetBox="[151,1414,183,1816]" captionTargetId="figure@35.[151,1436,167,1816]" captionTargetPageId="35" captionText="FIGURE 26. Ischnothyreus kalimantan sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, tooth-like projection; ssp, slightly sclerotized process; m, membrane; l, lobe; spa, small pointed apophysis; vp, ventral projection; vpr, ventral protuberance. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F = 0.1 mm; G = 0.05 mm." httpUri="https://zenodo.org/record/278137/files/figure.png" pageId="34" pageNumber="35">Fig. 26</figureCitation>
F–G). The female genital area is protruding, the strongly winding tube ends in a rather large sluice, which is folded underneath itself (
<figureCitation id="13272A7DFFFCFFF945C65EBCCD760BAE" box="[1008,1089,1181,1205]" captionStart="FIGURE 27" captionStartId="36.[151,250,1893,1916]" captionTargetBox="[151,1419,193,1855]" captionTargetId="figure@36.[151,1436,193,1856]" captionTargetPageId="36" captionText="FIGURE 27. Ischnothyreus kalimantan sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, abdomen, ventral view; F, G, genital area, ventral view; H, genital area, dorsal view. a, apodeme; wt, winding tube; ts, translucent sluice. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; F, G, H = 0.1 mm." httpUri="https://zenodo.org/record/278138/files/figure.png" pageId="34" pageNumber="35">Fig. 27</figureCitation>
F–H).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFFCFFF946F15EE0CC2408AB" pageId="34" pageNumber="35" type="description">
<paragraph id="8BA336F8FFFCFFF946F15EE0CC150929" blockId="34.[151,1437,945,2003]" pageId="34" pageNumber="35">
<emphasis id="B968EAEAFFFCFFF946F15EE0C87C0BC3" box="[199,331,1217,1240]" italics="true" pageId="34" pageNumber="35">Description</emphasis>
: Male (
<typeStatus id="54A7885AFFFCFFF947975E9ECB3D0BCC" box="[417,522,1215,1239]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFFCFFF944915E9ECA3A0BCC" box="[679,781,1215,1239]" metricMagnitude="-3" metricUnit="m" metricValue="1.79" pageId="34" pageNumber="35" unit="mm" value="1.79">1.79 mm</quantity>
. 
<emphasis id="B968EAEAFFFCFFF945215EE0CA7B0BC3" box="[791,844,1217,1240]" italics="true" pageId="34" pageNumber="35">Eyes</emphasis>
: all subequal. 
<emphasis id="B968EAEAFFFCFFF945D85EE1CD7D0BCC" box="[1006,1098,1216,1239]" italics="true" pageId="34" pageNumber="35">Sternum</emphasis>
: as long as wide. 
<emphasis id="B968EAEAFFFCFFF943265EE1CCAB0BC3" box="[1296,1436,1216,1240]" italics="true" pageId="34" pageNumber="35">Mouthparts:</emphasis>
chelicerae straight, fang groove with few small and one larger denticles. Anteriorly directed, slightly sclerotized process (ssp) at base of fangs (
<figureCitation id="13272A7DFFFCFFF947D85F24CB090A06" box="[494,574,1285,1309]" captionStart="FIGURE 26" captionStartId="35.[151,250,1857,1880]" captionTargetBox="[151,1414,183,1816]" captionTargetId="figure@35.[151,1436,167,1816]" captionTargetPageId="35" captionText="FIGURE 26. Ischnothyreus kalimantan sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, tooth-like projection; ssp, slightly sclerotized process; m, membrane; l, lobe; spa, small pointed apophysis; vp, ventral projection; vpr, ventral protuberance. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F = 0.1 mm; G = 0.05 mm." httpUri="https://zenodo.org/record/278137/files/figure.png" pageId="34" pageNumber="35">Fig. 26</figureCitation>
E). Labium triangular, anterior margin indented at middle. Anteromedian tip of endites with one strong, tooth-like projection (stp) (
<figureCitation id="13272A7DFFFCFFF944C45F06CA750A24" box="[754,834,1319,1343]" captionStart="FIGURE 26" captionStartId="35.[151,250,1857,1880]" captionTargetBox="[151,1414,183,1816]" captionTargetId="figure@35.[151,1436,167,1816]" captionTargetPageId="35" captionText="FIGURE 26. Ischnothyreus kalimantan sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, tooth-like projection; ssp, slightly sclerotized process; m, membrane; l, lobe; spa, small pointed apophysis; vp, ventral projection; vpr, ventral protuberance. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F = 0.1 mm; G = 0.05 mm." httpUri="https://zenodo.org/record/278137/files/figure.png" pageId="34" pageNumber="35">Fig. 26</figureCitation>
B). 
<emphasis id="B968EAEAFFFCFFF945445F09CAEC0A5B" box="[882,987,1320,1344]" italics="true" pageId="34" pageNumber="35">Abdomen</emphasis>
: scutum extending far dorsal of pedicel. Dorsal scutum orange-brown, covering 1/2 to 3/4 of abdomen width. Epigastric and postepigastric scutum fused, postepigastric scutum short, almost rectangular, covering about 2/3 of abdominal length. 
<emphasis id="B968EAEAFFFCFFF9424C5F4FCD980A9E" box="[1146,1199,1390,1413]" italics="true" pageId="34" pageNumber="35">Legs</emphasis>
: spine formula: femora: I p0-1-1; r0-1-2; II p0-0-1; r0-1-2; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0; 
<emphasis id="B968EAEAFFFCFFF942C65FB1CC6D0AB3" box="[1264,1370,1424,1448]" italics="true" pageId="34" pageNumber="35">Genitalia</emphasis>
: Palp strongly sclerotized, proximal segments brown, trochanter with ventral projection (vp) (
<figureCitation id="13272A7DFFFCFFF942485F93CDE70AD1" box="[1150,1232,1458,1482]" captionStart="FIGURE 26" captionStartId="35.[151,250,1857,1880]" captionTargetBox="[151,1414,183,1816]" captionTargetId="figure@35.[151,1436,167,1816]" captionTargetPageId="35" captionText="FIGURE 26. Ischnothyreus kalimantan sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, tooth-like projection; ssp, slightly sclerotized process; m, membrane; l, lobe; spa, small pointed apophysis; vp, ventral projection; vpr, ventral protuberance. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F = 0.1 mm; G = 0.05 mm." httpUri="https://zenodo.org/record/278137/files/figure.png" pageId="34" pageNumber="35">Fig. 26</figureCitation>
F), femur normal size, patella longer than femur, not enlarged, cymbium brown, fused with bulb, bulb brown, 1 to 1.5 times as long as cymbium, stout, tapering apically, with ventral protuberance (vpr) and lobe (l), distal part of bulb blunt, with several small pointed apophyses (spa), entirely surrounded by translucent membrane (m) (
<figureCitation id="13272A7DFFFCFFF942BF5C3BCDEE0929" box="[1161,1241,1562,1586]" captionStart="FIGURE 26" captionStartId="35.[151,250,1857,1880]" captionTargetBox="[151,1414,183,1816]" captionTargetId="figure@35.[151,1436,167,1816]" captionTargetPageId="35" captionText="FIGURE 26. Ischnothyreus kalimantan sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, tooth-like projection; ssp, slightly sclerotized process; m, membrane; l, lobe; spa, small pointed apophysis; vp, ventral projection; vpr, ventral protuberance. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F = 0.1 mm; G = 0.05 mm." httpUri="https://zenodo.org/record/278137/files/figure.png" pageId="34" pageNumber="35">Fig. 26</figureCitation>
F–G).
</paragraph>
<paragraph id="8BA336F8FFFCFFF946F15C1CCC2408AB" blockId="34.[151,1437,945,2003]" pageId="34" pageNumber="35">
Female (
<typeStatus id="54A7885AFFFCFFF9471F5C1CC8A7094E" box="[297,400,1597,1621]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFFCFFF944055C1CCBAB094E" box="[563,668,1597,1621]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" pageId="34" pageNumber="35" unit="mm" value="1.88">1.88 mm</quantity>
. As in male except as noted. 
<emphasis id="B968EAEAFFFCFFF945DD5C1FCD6E094E" box="[1003,1113,1598,1621]" italics="true" pageId="34" pageNumber="35">Carapace</emphasis>
: orange-brown, without any pattern, broadly oval in dorsal view, pars cephalica slightly elevated in lateral view, fovea present. Endites unmodified. Femur of female palp with five ventral spines, tibia with three trichobothria. 
<emphasis id="B968EAEAFFFCFFF942955CA2CC3A0980" box="[1187,1293,1667,1691]" italics="true" pageId="34" pageNumber="35">Abdomen</emphasis>
: scutum not extending far dorsal of pedicel. Dorsal scutum covering about 1/2 of abdomen. Epigastric scutum strongly protruding. Postepigastric scutum widely hexagonal, only around epigastric furrow, not fused to epigastric scutum, with short, posteriorly directed lateral apodemes (a) (
<figureCitation id="13272A7DFFFCFFF944815CCBCA3E0819" box="[695,777,1770,1794]" captionStart="FIGURE 27" captionStartId="36.[151,250,1893,1916]" captionTargetBox="[151,1419,193,1855]" captionTargetId="figure@36.[151,1436,193,1856]" captionTargetPageId="36" captionText="FIGURE 27. Ischnothyreus kalimantan sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, abdomen, ventral view; F, G, genital area, ventral view; H, genital area, dorsal view. a, apodeme; wt, winding tube; ts, translucent sluice. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; F, G, H = 0.1 mm." httpUri="https://zenodo.org/record/278138/files/figure.png" pageId="34" pageNumber="35">Fig. 27</figureCitation>
F–H). Dense patch of setae anterior to spinnerets present. 
<emphasis id="B968EAEAFFFCFFF946A15D2FC9FB083E" box="[151,204,1806,1829]" italics="true" pageId="34" pageNumber="35">Legs</emphasis>
: spine formula: femora: I p0-1-1; r0-1-2; II p0-0-1; r0-1-2; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFFCFFF946DC5D11C8630853" box="[234,340,1840,1864]" italics="true" pageId="34" pageNumber="35">Genitalia</emphasis>
: the posterior margin of the epigastric scutum is lined with numerous, long, needle-like setae. The epigastric furrow is wide. The margin of the postepigastric scutum is slightly thickened. In its middle starts a dark, thin, strongly winding tube (wt) running posteriorly, prolonged with a translucent sluice (ts), distally folded underneath itself (
<figureCitation id="13272A7DFFFCFFF947145DB9C84508AB" box="[290,370,1944,1968]" captionStart="FIGURE 27" captionStartId="36.[151,250,1893,1916]" captionTargetBox="[151,1419,193,1855]" captionTargetId="figure@36.[151,1436,193,1856]" captionTargetPageId="36" captionText="FIGURE 27. Ischnothyreus kalimantan sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, abdomen, ventral view; F, G, genital area, ventral view; H, genital area, dorsal view. a, apodeme; wt, winding tube; ts, translucent sluice. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; F, G, H = 0.1 mm." httpUri="https://zenodo.org/record/278138/files/figure.png" pageId="34" pageNumber="35">Fig. 27</figureCitation>
F–H). The sluice is ventrally lined with numerous, needle-like setae on both sides.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFFCFFFE46F15D9AC8670FC9" lastPageId="37" lastPageNumber="38" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BA336F8FFFCFFF946F15D9ACB9C08C9" blockId="34.[151,1437,945,2003]" box="[199,683,1978,2003]" pageId="34" pageNumber="35">
<emphasis id="B968EAEAFFFCFFF946F15D9AC86708C8" box="[199,336,1979,2003]" italics="true" pageId="34" pageNumber="35">Distribution</emphasis>
: Central 
<taxonomicName id="4C1C4D7BFFFCFFF9478E5D9BCB0C08C9" box="[440,571,1978,2002]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
(
<figureCitation id="13272A7DFFFCFFF9447C5D9BCBAA08C9" box="[586,669,1978,2002]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="34" pageNumber="35">Fig. 35</figureCitation>
).
</paragraph>
<caption id="DF636670FFFDFFF846A15D60CDD408AE" httpUri="https://zenodo.org/record/278137/files/figure.png" pageId="35" pageNumber="36" targetBox="[151,1414,183,1816]" targetPageId="35">
<paragraph id="8BA336F8FFFDFFF846A15D60CDD408AE" blockId="35.[151,1436,1857,1973]" pageId="35" pageNumber="36">
<emphasis id="B968EAEAFFFDFFF846A15D60C828084C" bold="true" box="[151,287,1857,1880]" pageId="35" pageNumber="36">FIGURE 26.</emphasis>
<taxonomicName id="4C1C4D7BFFFDFFF847125D63CB1D0843" box="[292,554,1858,1880]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kalimantan" status="sp. nov.">
<emphasis id="B968EAEAFFFDFFF847125D63CB1D0843" box="[292,554,1858,1880]" italics="true" pageId="35" pageNumber="36">Ischnothyreus kalimantan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFFDFFF844195D62CB490843" box="[559,638,1859,1880]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, retrolateral view; G, left bulb, prolateral view. stp, strong, tooth-like projection; ssp, slightly sclerotized process; m, membrane; l, lobe; spa, small pointed apophysis; vp, ventral projection; vpr, ventral protuberance. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F = 0.1 mm; G = 0.05mm.
</paragraph>
</caption>
<caption id="DF636670FFFAFFFF46A15D44CDD108DB" httpUri="https://zenodo.org/record/278138/files/figure.png" pageId="36" pageNumber="37" targetBox="[151,1419,193,1855]" targetPageId="36">
<paragraph id="8BA336F8FFFAFFFF46A15D44CDD108DB" blockId="36.[151,1437,1893,1984]" pageId="36" pageNumber="37">
<emphasis id="B968EAEAFFFAFFFF46A15D44C8150861" bold="true" box="[151,290,1893,1916]" pageId="36" pageNumber="37">FIGURE 27.</emphasis>
<taxonomicName id="4C1C4D7BFFFAFFFF471C5D44CB060860" box="[298,561,1893,1915]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kalimantan" status="sp. nov.">
<emphasis id="B968EAEAFFFAFFFF471C5D44CB060860" box="[298,561,1893,1915]" italics="true" pageId="36" pageNumber="37">Ischnothyreus kalimantan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFFAFFFF440F5D47CBBD0860" box="[569,650,1894,1915]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, abdomen, ventral view; F, G, genital area, ventral view; H, genital area, dorsal view. a, apodeme; wt, winding tube; ts, translucent sluice. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; F, G, H = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA336F8FFFBFFFE46A15AB6C8670FC9" blockId="37.[151,519,151,210]" pageId="37" pageNumber="38">
<taxonomicName id="4C1C4D7BFFFBFFFE46A15AB6C8910FAA" box="[151,422,151,177]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="matang" status="sp. nov.">
<emphasis id="B968EAEAFFFBFFFE46A15AB6C8910FAA" bold="true" box="[151,422,151,177]" italics="true" pageId="37" pageNumber="38">Ischnothyreus matang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFFBFFFE479B5AB9CB300FA9" box="[429,519,152,178]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13272A7DFFFBFFFE46A95A9BC8170FC9" box="[159,288,186,210]" captionStart-0="FIGURE 28" captionStart-1="FIGURE 29" captionStartId-0="37.[151,250,1790,1813]" captionStartId-1="38.[151,250,1417,1440]" captionTargetBox-0="[151,1393,533,1746]" captionTargetBox-1="[153,1430,151,1375]" captionTargetId-0="figure@37.[151,1436,499,1768]" captionTargetId-1="figure@38.[153,1438,129,1388]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionText-0="FIGURE 28. Ischnothyreus matang sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, left palp, retrolateral view; F, left palp, prolateral view. stp, strong, tooth-like projection; m, membrane; vp, ventral projection; vpr, ventral protuberance; l, lobe; sa, spine-like apophysis. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F = 0.1 mm." captionText-1="FIGURE 29. Ischnothyreus matang sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, D, genital area, ventral view; E, genital area, dorsal view. wt, winding tube; gls, goblet-like structure. Scale bars A, B = 1 mm; C, D, E = 0.1 mm." httpUri-0="https://zenodo.org/record/278139/files/figure.png" httpUri-1="https://zenodo.org/record/278140/files/figure.png" pageId="37" pageNumber="38">Figs 28–29</figureCitation>
, 
<figureCitation id="13272A7DFFFBFFFE471D5A9BC87F0FC9" box="[299,328,186,210]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="37" pageNumber="38">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFFBFFFE46A15B25C9C60E84" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="8BA336F8FFFBFFFE46A15B25C9C60E84" blockId="37.[151,1436,258,450]" pageId="37" pageNumber="38">
<emphasis id="B968EAEAFFFBFFFE46A15B25C8900E00" box="[151,423,259,283]" italics="true" pageId="37" pageNumber="38">
<typeStatus id="54A7885AFFFBFFFE46A15B25C9FC0E00" box="[151,203,260,283]" pageId="37" pageNumber="38">Type</typeStatus>
material: 
<typeStatus id="54A7885AFFFBFFFE47765B22C8900E00" box="[320,423,259,283]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
</emphasis>
: male (PBI_OON 00031470), 
<collectingCountry id="F30B7668FFFBFFFE44C85B23CAA30E01" box="[766,916,258,282]" name="Malaysia" pageId="37" pageNumber="38">MALAYSIA</collectingCountry>
: 
<collectingCountry id="F30B7668FFFBFFFE45A85B23CD340E01" box="[926,1027,258,282]" name="Malaysia" pageId="37" pageNumber="38">Sarawak</collectingCountry>
: 
<taxonomicName id="4C1C4D7BFFFBFFFE423A5B22CD530E00" box="[1036,1124,259,283]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="46" phylum="Arthropoda" rank="species" species="matang">Matang</taxonomicName>
, 
<quantity id="4CE49B1DFFFBFFFE42595B23CD850E01" box="[1135,1202,258,282]" metricMagnitude="2" metricUnit="m" metricValue="1.22" pageId="37" pageNumber="38" unit="m" value="122.0">122m</quantity>
, leaflitter, 
<date id="FFA21038FFFBFFFE43115B23CCA00E01" box="[1319,1431,258,282]" pageId="37" pageNumber="38" value="1985-04-04">4.IV.1985</date>
, coll. Deeleman, leg. C.L. &amp; P.R. Deeleman, 
<collectionCode id="ED0DAE3DFFFBFFFE44805B04CA390E27" LSID="urn:lsid:biocol.org:col:34992" box="[694,782,293,316]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="37" pageNumber="38">RMNH</collectionCode>
. 
<typeStatus id="54A7885AFFFBFFFE45295B04CAA60E27" box="[799,913,293,316]" pageId="37" pageNumber="38" type="paratype">
<emphasis id="B968EAEAFFFBFFFE45295B04CAA60E27" box="[799,913,293,316]" italics="true" pageId="37" pageNumber="38">Paratypes</emphasis>
</typeStatus>
: 
<specimenCount id="9D1AFD71FFFBFFFE45905B05CD390E27" box="[934,1038,292,316]" pageId="37" pageNumber="38" type="female">1 female</specimenCount>
(PBI_OON 00032185), collected together with 
<typeStatus id="54A7885AFFFBFFFE47005B64C8AC0E46" box="[310,411,325,349]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED0DAE3DFFFBFFFE47915B67C8C80E46" LSID="urn:lsid:biocol.org:col:34992" box="[423,511,326,349]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="37" pageNumber="38">RMNH</collectionCode>
. 
<specimenCount id="9D1AFD71FFFBFFFE443C5B64CB5A0E46" box="[522,621,325,349]" pageId="37" pageNumber="38" type="female">1 female</specimenCount>
(PBI_OON 00016081), 
<collectingCountry id="F30B7668FFFBFFFE45B15B64CD2A0E46" box="[903,1053,325,349]" name="Malaysia" pageId="37" pageNumber="38">MALAYSIA</collectingCountry>
: 
<collectingCountry id="F30B7668FFFBFFFE421E5B64CDBA0E46" box="[1064,1165,325,349]" name="Malaysia" pageId="37" pageNumber="38">Sarawak</collectingCountry>
: Santubong, 
<quantity id="4CE49B1DFFFBFFFE43165B64CC680E46" box="[1312,1375,325,349]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="38" unit="km" value="20.0">20km</quantity>
N of Kuching, Camp Permai, 
<geoCoordinate id="EE28503FFFFBFFFE47835B47CB310E65" box="[437,518,358,382]" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="1.7666667">1°46'N</geoCoordinate>
, 
<geoCoordinate id="EE28503FFFFBFFFE44235B47CB490E65" box="[533,638,358,382]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="110.316666">110°19'E</geoCoordinate>
, 
<quantity id="4CE49B1DFFFBFFFE44BB5B47CBF60E65" box="[653,705,358,382]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="38" unit="m" value="10.0">10m</quantity>
, 
<date id="FFA21038FFFBFFFE44F95B47CAB00E65" box="[719,903,358,382]" pageId="37" pageNumber="38" value="2003-08-05" valueMax="2003-08-10" valueMin="2003-08-05">5.–10.VIII.2003</date>
, leg. A. Schulz (winkler-extraction), 
<collectingCountry id="F30B7668FFFBFFFE43005B47CC6C0E65" box="[1334,1371,358,382]" name="American Samoa" pageId="37" pageNumber="38">AS</collectingCountry>
/03-2, 
<collectionCode id="ED0DAE3DFFFBFFFE46A15BA6C9DA0E84" box="[151,237,391,415]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="37" pageNumber="38">MHNG</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFFBFFFE46F15B88CD560EDA" box="[199,1121,425,450]" pageId="37" pageNumber="38" type="etymology">
<paragraph id="8BA336F8FFFBFFFE46F15B88CD560EDA" blockId="37.[151,1436,258,450]" box="[199,1121,425,450]" pageId="37" pageNumber="38">
<emphasis id="B968EAEAFFFBFFFE46F15B88C8080EDA" box="[199,319,425,449]" italics="true" pageId="37" pageNumber="38">Etymology</emphasis>
: The specific name is a noun in apposition taken from the 
<typeStatus id="54A7885AFFFBFFFE45E65B8BCD370ED9" box="[976,1024,426,450]" pageId="37" pageNumber="38">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="DF636670FFFBFFFE46A15CDFC852086B" httpUri="https://zenodo.org/record/278139/files/figure.png" pageId="37" pageNumber="38" targetBox="[151,1393,533,1746]" targetPageId="37">
<paragraph id="8BA336F8FFFBFFFE46A15CDFC852086B" blockId="37.[151,1436,1790,1904]" pageId="37" pageNumber="38">
<emphasis id="B968EAEAFFFBFFFE46A15CDFC8170808" bold="true" box="[151,288,1790,1813]" pageId="37" pageNumber="38">FIGURE 28.</emphasis>
<taxonomicName id="4C1C4D7BFFFBFFFE47105CDFCB31080F" box="[294,518,1790,1812]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="matang" status="sp. nov.">
<emphasis id="B968EAEAFFFBFFFE47105CDFCB31080F" box="[294,518,1790,1812]" italics="true" pageId="37" pageNumber="38">Ischnothyreus matang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFFBFFFE443B5CDECB6B080F" box="[525,604,1791,1812]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, left palp, retrolateral view; F, left palp, prolateral view. stp, strong, tooth-like projection; m, membrane; vp, ventral projection; vpr, ventral protuberance; l, lobe; sa, spine-like apophysis. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F = 0.1 mm.
</paragraph>
</caption>
<caption id="DF636670FFF8FFFD46A15FA8CCA50AA4" httpUri="https://zenodo.org/record/278140/files/figure.png" pageId="38" pageNumber="39" targetBox="[153,1430,151,1375]" targetPageId="38">
<paragraph id="8BA336F8FFF8FFFD46A15FA8CCA50AA4" blockId="38.[151,1436,1417,1471]" pageId="38" pageNumber="39">
<emphasis id="B968EAEAFFF8FFFD46A15FA8C8170A84" bold="true" box="[151,288,1417,1440]" pageId="38" pageNumber="39">FIGURE 29.</emphasis>
<taxonomicName id="4C1C4D7BFFF8FFFD47105FABCB300ABB" box="[294,519,1418,1440]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="matang" status="sp. nov.">
<emphasis id="B968EAEAFFF8FFFD47105FABCB300ABB" box="[294,519,1418,1440]" italics="true" pageId="38" pageNumber="39">Ischnothyreus matang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFF8FFFD443B5FAACB6B0ABB" box="[525,604,1419,1440]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
female. A, habitus, dorsal view; B, habitus, ventral view; C, D, genital area, ventral view; E, genital area, dorsal view. wt, winding tube; gls, goblet-like structure. Scale bars A, B = 1 mm; C, D, E = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C3066573FFF8FFFD46F15FCCC85A0950" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="8BA336F8FFF8FFFD46F15FCCC85A0950" blockId="38.[151,1436,1515,2007]" pageId="38" pageNumber="39">
<emphasis id="B968EAEAFFF8FFFD46F15FCCC80E091F" box="[199,313,1517,1540]" italics="true" pageId="38" pageNumber="39">Diagnosis</emphasis>
: The male of 
<taxonomicName id="4C1C4D7BFFF8FFFD47E05FCCCB76091F" box="[470,577,1517,1540]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="matang">
<emphasis id="B968EAEAFFF8FFFD47E05FCCCB76091F" box="[470,577,1517,1540]" italics="true" pageId="38" pageNumber="39">I. matang</emphasis>
</taxonomicName>
can be recognized by the large ventral lobe and the collar-like membrane on the palpal bulb (
<figureCitation id="13272A7DFFF8FFFD477A5C2EC8AB093C" box="[332,412,1551,1575]" captionStart="FIGURE 28" captionStartId="37.[151,250,1790,1813]" captionTargetBox="[151,1393,533,1746]" captionTargetId="figure@37.[151,1436,499,1768]" captionTargetPageId="37" captionText="FIGURE 28. Ischnothyreus matang sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, left palp, retrolateral view; F, left palp, prolateral view. stp, strong, tooth-like projection; m, membrane; vp, ventral projection; vpr, ventral protuberance; l, lobe; sa, spine-like apophysis. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F = 0.1 mm." httpUri="https://zenodo.org/record/278139/files/figure.png" pageId="38" pageNumber="39">Fig. 28</figureCitation>
E–F). The female genital area shows a strongly winding tube, ending in a goblet-like structure (
<figureCitation id="13272A7DFFF8FFFD46E45C12C8150950" box="[210,290,1587,1611]" captionStart="FIGURE 29" captionStartId="38.[151,250,1417,1440]" captionTargetBox="[153,1430,151,1375]" captionTargetId="figure@38.[153,1438,129,1388]" captionTargetPageId="38" captionText="FIGURE 29. Ischnothyreus matang sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, D, genital area, ventral view; E, genital area, dorsal view. wt, winding tube; gls, goblet-like structure. Scale bars A, B = 1 mm; C, D, E = 0.1 mm." httpUri="https://zenodo.org/record/278140/files/figure.png" pageId="38" pageNumber="39">Fig. 29</figureCitation>
D–E).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF8FFFC46F15C79CC7A0EEF" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="description">
<paragraph id="8BA336F8FFF8FFFD46F15C79CACA08CC" blockId="38.[151,1436,1515,2007]" pageId="38" pageNumber="39">
<emphasis id="B968EAEAFFF8FFFD46F15C79C87C0974" box="[199,331,1624,1647]" italics="true" pageId="38" pageNumber="39">Description</emphasis>
: Male (
<typeStatus id="54A7885AFFF8FFFD47975C76CB3E0974" box="[417,521,1623,1647]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFF8FFFD44915C76CA390974" box="[679,782,1623,1647]" metricMagnitude="-3" metricUnit="m" metricValue="1.59" pageId="38" pageNumber="39" unit="mm" value="1.59">1.59 mm</quantity>
. Fovea present. 
<emphasis id="B968EAEAFFF8FFFD45F75C76CD2C0974" box="[961,1051,1623,1647]" italics="true" pageId="38" pageNumber="39">Clypeus</emphasis>
: low, ALE separated from edge of carapace by less than their radius. 
<emphasis id="B968EAEAFFF8FFFD441F5C5DCBB30988" box="[553,644,1660,1683]" italics="true" pageId="38" pageNumber="39">Sternum</emphasis>
: wider than long. 
<emphasis id="B968EAEAFFF8FFFD456B5C5DCADE098F" box="[861,1001,1660,1684]" italics="true" pageId="38" pageNumber="39">Mouthparts:</emphasis>
chelicerae slightly divergent, without prominent process at base of fangs, fang groove with few denticles. Anteromedian tip of endites with one strong, tooth-like projection (stp) (
<figureCitation id="13272A7DFFF8FFFD47FF5CE2CB2C09C0" box="[457,539,1731,1755]" captionStart="FIGURE 28" captionStartId="37.[151,250,1790,1813]" captionTargetBox="[151,1393,533,1746]" captionTargetId="figure@37.[151,1436,499,1768]" captionTargetPageId="37" captionText="FIGURE 28. Ischnothyreus matang sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, left palp, retrolateral view; F, left palp, prolateral view. stp, strong, tooth-like projection; m, membrane; vp, ventral projection; vpr, ventral protuberance; l, lobe; sa, spine-like apophysis. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F = 0.1 mm." httpUri="https://zenodo.org/record/278139/files/figure.png" pageId="38" pageNumber="39">Fig. 28</figureCitation>
D). 
<emphasis id="B968EAEAFFF8FFFD44665CE5CB8D09C7" box="[592,698,1732,1756]" italics="true" pageId="38" pageNumber="39">Abdomen</emphasis>
: scutum extending far dorsal of pedicel. Dorsal scutum covering less than 1/2 of abdomen, more than 1/2 to most of abdomen width, fused to epigastric scutum. Epigastric scutum and postepigastric scutum fused. 
<emphasis id="B968EAEAFFF8FFFD443A5D2CCB76083F" box="[524,577,1805,1828]" italics="true" pageId="38" pageNumber="39">Legs</emphasis>
: spine formula: I p0-1-2; r0-0-1; tibiae: I p2-1-1; r2-1-1; metatarsi: I p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFF8FFFD46DA5D0EC861085C" box="[236,342,1839,1863]" italics="true" pageId="38" pageNumber="39">Genitalia</emphasis>
: Sperm pore situated in front of anterior spiracles. Palp strongly sclerotized, proximal segments brown, trochanter with large ventral projection (vp) (
<figureCitation id="13272A7DFFF8FFFD44DC5D72CA0C0870" box="[746,827,1875,1899]" captionStart="FIGURE 28" captionStartId="37.[151,250,1790,1813]" captionTargetBox="[151,1393,533,1746]" captionTargetId="figure@37.[151,1436,499,1768]" captionTargetPageId="37" captionText="FIGURE 28. Ischnothyreus matang sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, left palp, retrolateral view; F, left palp, prolateral view. stp, strong, tooth-like projection; m, membrane; vp, ventral projection; vpr, ventral protuberance; l, lobe; sa, spine-like apophysis. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F = 0.1 mm." httpUri="https://zenodo.org/record/278139/files/figure.png" pageId="38" pageNumber="39">Fig. 28</figureCitation>
E), patella about as long as femur, not enlarged, cymbium brown, fused with bulb, bulb brown, more than two times as long as cymbium, stout, tapering apically, with double ventral protuberance (vpr) and lobe (l), distal part with small, spine-like apophysis (sa) dorsally. Collar-like membrane (m) on retrolateral side, embolus (e) slightly curved (
<figureCitation id="13272A7DFFF8FFFD45505D9ECA8108CC" box="[870,950,1983,2007]" captionStart="FIGURE 28" captionStartId="37.[151,250,1790,1813]" captionTargetBox="[151,1393,533,1746]" captionTargetId="figure@37.[151,1436,499,1768]" captionTargetPageId="37" captionText="FIGURE 28. Ischnothyreus matang sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, dorsal view; D, carapace, ventral view; E, left palp, retrolateral view; F, left palp, prolateral view. stp, strong, tooth-like projection; m, membrane; vp, ventral projection; vpr, ventral protuberance; l, lobe; sa, spine-like apophysis. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F = 0.1 mm." httpUri="https://zenodo.org/record/278139/files/figure.png" pageId="38" pageNumber="39">Fig. 28</figureCitation>
E–F).
</paragraph>
<paragraph id="8BA336F8FFF9FFFC46F15AB6CC7A0EEF" blockId="39.[151,1436,151,644]" pageId="39" pageNumber="40">
Female (
<typeStatus id="54A7885AFFF9FFFC471E5AB6C8B80FB4" box="[296,399,151,175]" pageId="39" pageNumber="40" type="paratype">paratype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFF9FFFC44065AB6CBA00FB4" box="[560,663,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="39" pageNumber="40" unit="mm" value="1.9">1.90 mm</quantity>
. As in male except as noted. 
<emphasis id="B968EAEAFFF9FFFC45D65AB9CD790FB4" box="[992,1102,152,175]" italics="true" pageId="39" pageNumber="40">Carapace</emphasis>
: without any pattern, broadly oval in dorsal view. Fovea absent. 
<emphasis id="B968EAEAFFF9FFFC44175A9CCBB00FCF" box="[545,647,189,212]" italics="true" pageId="39" pageNumber="40">Sternum:</emphasis>
longer than wide, setae abundant. Labium triangular, anterior margin indented at middle. Endites unmodified. 
<emphasis id="B968EAEAFFF9FFFC44565AC1CBFD0FE3" box="[608,714,224,248]" italics="true" pageId="39" pageNumber="40">Abdomen</emphasis>
: Dorsal scutum less than 1/4 abdomen width. Epigastric scutum strongly protruding. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length, not fused to epigastric scutum, with short posteriorly directed lateral apodemes (a) (
<figureCitation id="13272A7DFFF9FFFC422D5B06CD5C0E24" box="[1051,1131,295,319]" captionStart="FIGURE 29" captionStartId="38.[151,250,1417,1440]" captionTargetBox="[153,1430,151,1375]" captionTargetId="figure@38.[153,1438,129,1388]" captionTargetPageId="38" captionText="FIGURE 29. Ischnothyreus matang sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, D, genital area, ventral view; E, genital area, dorsal view. wt, winding tube; gls, goblet-like structure. Scale bars A, B = 1 mm; C, D, E = 0.1 mm." httpUri="https://zenodo.org/record/278140/files/figure.png" pageId="39" pageNumber="40">Fig. 29</figureCitation>
D). Spinneret scutum with a fringe of needle-like setae. Dense patch of setae anterior to spinnerets present. 
<emphasis id="B968EAEAFFF9FFFC422D5B6CCD670E7F" box="[1051,1104,333,356]" italics="true" pageId="39" pageNumber="40">Legs</emphasis>
: spine formula: femora: I p0- 1-2; r0-0-1; II p0-0-1; r0-0-1; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFF9FFFC426B5B51CDF10E93" box="[1117,1222,368,392]" italics="true" pageId="39" pageNumber="40">Genitalia</emphasis>
: the posterior margin of the epigastric scutum is lined with numerous, long, needle-like setae. The epigastric furrow is rather wide. The margin of the postepigastric scutum is thickened. In its middle starts a thin, strongly winding tube (wt), ending straight in elongated, narrow, goblet-like structure (gls), which is protruding from the scutum (
<figureCitation id="13272A7DFFF9FFFC428E5BFDCC350EEF" box="[1208,1282,476,500]" captionStart="FIGURE 29" captionStartId="38.[151,250,1417,1440]" captionTargetBox="[153,1430,151,1375]" captionTargetId="figure@38.[153,1438,129,1388]" captionTargetPageId="38" captionText="FIGURE 29. Ischnothyreus matang sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, D, genital area, ventral view; E, genital area, dorsal view. wt, winding tube; gls, goblet-like structure. Scale bars A, B = 1 mm; C, D, E = 0.1 mm." httpUri="https://zenodo.org/record/278140/files/figure.png" pageId="39" pageNumber="40">Fig 29</figureCitation>
D–E).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF9FFFC46F15821CB180D0C" box="[199,559,511,536]" pageId="39" pageNumber="40" type="distribution">
<paragraph id="8BA336F8FFF9FFFC46F15821CB180D0C" blockId="39.[151,1436,151,644]" box="[199,559,511,536]" pageId="39" pageNumber="40">
<emphasis id="B968EAEAFFF9FFFC46F15821C8670D03" box="[199,336,512,536]" italics="true" pageId="39" pageNumber="40">Distribution</emphasis>
: 
<collectingCountry id="F30B7668FFF9FFFC47685BDEC8890D0C" box="[350,446,511,535]" name="Malaysia" pageId="39" pageNumber="40">Sarawak</collectingCountry>
(
<figureCitation id="13272A7DFFF9FFFC47F85BDECB160D0C" box="[462,545,511,535]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="39" pageNumber="40">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF9FFFC46F15805C8670C09" pageId="39" pageNumber="40" type="discussion">
<paragraph id="8BA336F8FFF9FFFC46F15805CA700D9F" blockId="39.[151,1436,151,644]" pageId="39" pageNumber="40">
<emphasis id="B968EAEAFFF9FFFC46F15805C8290D27" box="[199,286,548,572]" italics="true" pageId="39" pageNumber="40">Remark</emphasis>
: Since only one male specimen (the 
<typeStatus id="54A7885AFFF9FFFC44F95805CA040D27" box="[719,819,548,572]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
) and two female specimens are currently available, a more accurate examination (preparation of the chelicerae, for example) was not possible; further examinations should be done as soon as more specimens become available.
</paragraph>
<paragraph id="8BA336F8FFF9FFFC46A158F6C8670C09" blockId="39.[151,523,727,786]" pageId="39" pageNumber="40">
<taxonomicName id="4C1C4D7BFFF9FFFC46A158F6C89D0DEA" box="[151,426,727,753]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mulumi" status="sp. nov.">
<emphasis id="B968EAEAFFF9FFFC46A158F6C89D0DEA" bold="true" box="[151,426,727,753]" italics="true" pageId="39" pageNumber="40">Ischnothyreus mulumi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFF9FFFC478658F9CB3C0DE9" box="[432,523,728,754]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13272A7DFFF9FFFC46A958DBC8170C09" box="[159,288,762,786]" captionStart-0="FIGURE 30" captionStart-1="FIGURE 31" captionStartId-0="40.[151,250,1847,1870]" captionStartId-1="41.[151,250,1852,1875]" captionTargetBox-0="[151,1427,212,1823]" captionTargetBox-1="[151,1424,203,1824]" captionTargetId-0="figure@40.[151,1436,195,1823]" captionTargetId-1="figure@41.[151,1436,193,1826]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURE 30. Ischnothyreus mulumi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, lateral view; G, left chelicera, posterior view; H, left palp, retrolateral view; I, left bulb, prolateral view; J, left bulb, retrolateral view. stp, strong, tooth-like projection; vp, ventral projection; b, brush; bp, basal process; vpr, ventral protuberance; l, lobe; clm, collar-like membrane; bsa, beak-like apophysis; ri, ribbed. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; G = 0.05 mm; F, H = 0.1 mm; I, J = 0.01 mm." captionText-1="FIGURE 31. Ischnothyreus mulumi sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, abdomen, ventral view; G, H, genital area, ventral view; I, J, genital area, dorsal view. a, apodeme; wt, winding tube; hc, half circle. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri-0="https://zenodo.org/record/278141/files/figure.png" httpUri-1="https://zenodo.org/record/278142/files/figure.png" pageId="39" pageNumber="40">Figs 30–31</figureCitation>
, 
<figureCitation id="13272A7DFFF9FFFC471D58DBC87F0C09" box="[299,328,762,786]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="39" pageNumber="40">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF9FFFC46A15965CDE30A49" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="8BA336F8FFF9FFFC46A15965CAC00CB9" blockId="39.[151,1437,834,2010]" pageId="39" pageNumber="40">
<emphasis id="B968EAEAFFF9FFFC46A15965C8860C40" box="[151,433,835,859]" italics="true" pageId="39" pageNumber="40">
<typeStatus id="54A7885AFFF9FFFC46A15965C9FC0C40" box="[151,203,836,859]" pageId="39" pageNumber="40">Type</typeStatus>
material: 
<typeStatus id="54A7885AFFF9FFFC477C5962C8860C40" box="[330,433,835,859]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
</emphasis>
: male (PBI_OON 00012470), 
<collectingCountry id="F30B7668FFF9FFFC452B5963CA740C41" box="[797,835,834,858]" name="American Samoa" pageId="39" pageNumber="40">AS</collectingCountry>
/03-7. 
<collectingCountry id="F30B7668FFF9FFFC45B85963CD130C41" box="[910,1060,834,858]" name="Malaysia" pageId="39" pageNumber="40">MALAYSIA</collectingCountry>
: 
<collectingCountry id="F30B7668FFF9FFFC42055963CDAF0C41" box="[1075,1176,834,858]" name="Malaysia" pageId="39" pageNumber="40">Sarawak</collectingCountry>
: Mulu, National Park, 
<quantity id="4CE49B1DFFF9FFFC46A15947C9D30C65" box="[151,228,870,894]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="40" unit="km" value="100.0">100km</quantity>
SEE of Miri, 
<geoCoordinate id="EE28503FFFF9FFFC47A05947C8DF0C65" box="[406,488,870,894]" direction="north" orientation="latitude" pageId="39" pageNumber="40" precision="925" value="4.0">4°00'N</geoCoordinate>
, 
<geoCoordinate id="EE28503FFFF9FFFC47CE5947CB560C65" box="[504,609,870,894]" direction="east" orientation="longitude" pageId="39" pageNumber="40" precision="925" value="114.816666">114°49'E</geoCoordinate>
, 
<quantity id="4CE49B1DFFF9FFFC44475947CB830C65" box="[625,692,870,894]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="40" unit="m" value="200.0">200m</quantity>
, 
<date id="FFA21038FFF9FFFC44F25947CABD0C65" box="[708,906,870,894]" pageId="39" pageNumber="40" value="2003-08-19" valueMax="2003-08-24" valueMin="2003-08-19">19.–24.VIII.2003</date>
, leg. A. Schulz (winkler-extraction), 
<collectionCode id="ED0DAE3DFFF9FFFC43755947CCAE0C65" box="[1347,1433,870,894]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="39" pageNumber="40">MHNG</collectionCode>
. 
<typeStatus id="54A7885AFFF9FFFC46A159ADC83E0CB8" box="[151,265,908,931]" pageId="39" pageNumber="40" type="paratype">
<emphasis id="B968EAEAFFF9FFFC46A159ADC83E0CB8" box="[151,265,908,931]" italics="true" pageId="39" pageNumber="40">Paratypes</emphasis>
</typeStatus>
: 
<specimenCount id="9D1AFD71FFF9FFFC472159ABC8B40CB9" box="[279,387,906,930]" pageId="39" pageNumber="40" type="female">4 females</specimenCount>
(PBI_OON 00032196), same data as 
<typeStatus id="54A7885AFFF9FFFC451B59ABCAA50CB9" box="[813,914,906,930]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED0DAE3DFFF9FFFC45AA59ABCAC50CB9" box="[924,1010,906,930]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="39" pageNumber="40">MHNG</collectionCode>
.
</paragraph>
<paragraph id="8BA336F8FFF9FFFC46F1598FCDE30A49" blockId="39.[151,1437,834,2010]" pageId="39" pageNumber="40">
<emphasis id="B968EAEAFFF9FFFC46F1598FCB220CDD" box="[199,533,942,966]" italics="true" pageId="39" pageNumber="40">Additional material examined</emphasis>
: 
<specimenCount id="9D1AFD71FFF9FFFC4415598FCB590CDD" box="[547,622,942,966]" pageId="39" pageNumber="40" type="male">1 male</specimenCount>
&amp; 
<specimenCount id="9D1AFD71FFF9FFFC44A4598FCBC50CDD" box="[658,754,942,966]" pageId="39" pageNumber="40" type="female">1 female</specimenCount>
(PBI_OON 00036304), 
<collectingCountry id="F30B7668FFF9FFFC4231598ECC0C0CDD" box="[1031,1339,942,966]" name="Brunei" pageId="39" pageNumber="40">BRUNEI DARUSSALAM</collectingCountry>
: Temburong District: Ashton Trail near Kuala Belalong Field Studies Centre, 
<quantity id="4CE49B1DFFF9FFFC423059F3CD710CF1" box="[1030,1094,978,1002]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="39" pageNumber="40" unit="km" value="21.0">21km</quantity>
SSW Bangar, 
<geoCoordinate id="EE28503FFFF9FFFC433159F3CCAF0CF1" box="[1287,1432,978,1002]" direction="north" orientation="latitude" pageId="39" pageNumber="40" precision="1" value="4.5418835">04°32.513'N</geoCoordinate>
, 
<geoCoordinate id="EE28503FFFF9FFFC46A159D7C8060B15" box="[151,305,1014,1038]" direction="east" orientation="longitude" pageId="39" pageNumber="40" precision="1" value="115.155">115°09.300'E</geoCoordinate>
, 
<quantity id="4CE49B1DFFF9FFFC470B59D7C8800B15" box="[317,439,1014,1038]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="39" pageNumber="40" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">100–150m</quantity>
, primary mixed dipterocarp forest, sifting leaf litter, 
<date id="FFA21038FFF9FFFC423A59D7CD400B15" box="[1036,1143,1014,1038]" pageId="39" pageNumber="40" value="2009-10-01">1.X.2009</date>
, C. Griswold &amp; N. Chousou Polydouri UT004, CASENT 9036044 (
<collectionCode id="ED0DAE3DFFF9FFFC44A15E3BCBE40B29" LSID="urn:lsid:biocol.org:col:34700" box="[663,723,1050,1074]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="39" pageNumber="40">CAS</collectionCode>
). 
<specimenCount id="9D1AFD71FFF9FFFC44DD5E3BCA6B0B29" box="[747,860,1050,1074]" pageId="39" pageNumber="40" type="female">2 females</specimenCount>
(PBI_OON 00036326), same data as PBI_OON 0 0 0 36304 except miniwinkler extraction of concentrated leaf litter, 
<date id="FFA21038FFF9FFFC45ED5E1FCD730B4D" box="[987,1092,1086,1110]" pageId="39" pageNumber="40" value="2009-10-03">3.X.2009</date>
(
<collectionCode id="ED0DAE3DFFF9FFFC426B5E1FCDAE0B4D" LSID="urn:lsid:biocol.org:col:34700" box="[1117,1177,1086,1110]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="39" pageNumber="40">CAS</collectionCode>
). 
<specimenCount id="9D1AFD71FFF9FFFC42805E1FCC3B0B4D" box="[1206,1292,1086,1110]" pageId="39" pageNumber="40" type="male">1 male</specimenCount>
(PBI_OON 00036335), same data as PBI_OON 0 0 0 36304 except 
<date id="FFA21038FFF9FFFC45375E43CAB20B61" box="[769,901,1122,1146]" pageId="39" pageNumber="40" value="2009-10-04" valueMax="2009-10-09" valueMin="2009-10-04">4–9.X.2009</date>
(
<collectionCode id="ED0DAE3DFFF9FFFC45A05E43CAE50B61" LSID="urn:lsid:biocol.org:col:34700" box="[918,978,1122,1146]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="39" pageNumber="40">CAS</collectionCode>
). 
<specimenCount id="9D1AFD71FFF9FFFC45D05E43CD7E0B61" box="[998,1097,1122,1146]" pageId="39" pageNumber="40" type="female">1 female</specimenCount>
(PBI_OON 00036333), same data as PBI_OON 0 0 0 36304 (
<collectionCode id="ED0DAE3DFFF9FFFC47D85EA7CB1D0B85" LSID="urn:lsid:biocol.org:col:34700" box="[494,554,1158,1182]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="39" pageNumber="40">CAS</collectionCode>
). 
<specimenCount id="9D1AFD71FFF9FFFC44085EA7CBBD0B85" box="[574,650,1158,1182]" pageId="39" pageNumber="40" type="male">1 male</specimenCount>
(PBI_OON 00036325), same data as PBI_OON 0 0 0 36326 (
<collectionCode id="ED0DAE3DFFF9FFFC430B5EA7CC4E0B85" LSID="urn:lsid:biocol.org:col:34700" box="[1341,1401,1158,1182]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="39" pageNumber="40">CAS</collectionCode>
). 
<specimenCount id="9D1AFD71FFF9FFFC43B85EA7C9D40BD9" pageId="39" pageNumber="40" type="female">1 female</specimenCount>
(PBI_OON 00035260), same data as PBI_OON 0 0 0 36326 (
<collectionCode id="ED0DAE3DFFF9FFFC45BE5E8BCAF30BD9" LSID="urn:lsid:biocol.org:col:34700" box="[904,964,1194,1218]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="39" pageNumber="40">CAS</collectionCode>
). 
<specimenCount id="9D1AFD71FFF9FFFC45E15E8BCD0F0BD9" box="[983,1080,1194,1218]" pageId="39" pageNumber="40" type="female">1 female</specimenCount>
(PBI_OON 00031652), INDO- 
<collectionCode id="ED0DAE3DFFF9FFFC46A15EEFC9D90BFD" box="[151,238,1230,1254]" pageId="39" pageNumber="40">NESIA</collectionCode>
: East 
<taxonomicName id="4C1C4D7BFFF9FFFC470E5EEFC8880BFD" box="[312,447,1230,1254]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
: Berau Distr., Hutan Wisata Sei Tangap, ca. 
<quantity id="4CE49B1DFFF9FFFC45E75EEFCD390BFD" box="[977,1038,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="39" pageNumber="40" unit="km" value="8.0">8 km</quantity>
W of Tanjungredeb (
<geoCoordinate id="EE28503FFFF9FFFC432F5EEFCCA00BFD" box="[1305,1431,1230,1254]" direction="north" orientation="latitude" pageId="39" pageNumber="40" precision="15" value="2.1344445">2°08’04”N</geoCoordinate>
, 
<geoCoordinate id="EE28503FFFF9FFFC46A15ED3C81A0A11" box="[151,301,1266,1290]" direction="east" orientation="longitude" pageId="39" pageNumber="40" precision="15" value="117.410835">117°24’39”E</geoCoordinate>
), 
<quantity id="4CE49B1DFFF9FFFC47755ED3C8430A11" box="[323,372,1266,1290]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="39" pageNumber="40" unit="m" value="30.0">30m</quantity>
(primary forest), 
<date id="FFA21038FFF9FFFC44775ED3CB9C0A11" box="[577,683,1266,1290]" pageId="39" pageNumber="40" value="2008-10-02">2.X.2008</date>
, leg. P. Schwendinger (
<collectionCode id="ED0DAE3DFFF9FFFC45815ED3CD240A11" box="[951,1043,1266,1290]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="39" pageNumber="40">MHNG</collectionCode>
). 
<specimenCount id="9D1AFD71FFF9FFFC421E5ED3CDBD0A11" box="[1064,1162,1266,1290]" pageId="39" pageNumber="40" type="female">1 female</specimenCount>
(PBI_OON 00031644), 
<collectingCountry id="F30B7668FFF9FFFC46A15F37C8030A35" box="[151,308,1302,1326]" name="Indonesia" pageId="39" pageNumber="40">INDONESIA</collectingCountry>
: East 
<taxonomicName id="4C1C4D7BFFF9FFFC47BE5F37CB380A35" box="[392,527,1302,1326]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
: Berau Distr., Hutan Mayang Mangurai, ca. 
<quantity id="4CE49B1DFFF9FFFC42765F37CDA70A35" box="[1088,1168,1302,1326]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="39" pageNumber="40" unit="km" value="15.0">15 km</quantity>
SW of Tanjungredeb (
<geoCoordinate id="EE28503FFFF9FFFC46965F1BC8290A49" box="[160,286,1338,1362]" direction="north" orientation="latitude" pageId="39" pageNumber="40" precision="15" value="2.103611">2°06’13”N</geoCoordinate>
, 
<geoCoordinate id="EE28503FFFF9FFFC471F5F1BC8880A49" box="[297,447,1338,1362]" direction="east" orientation="longitude" pageId="39" pageNumber="40" precision="15" value="117.40139">117°24’05”E</geoCoordinate>
), 
<quantity id="4CE49B1DFFF9FFFC47E25F1BCB320A49" box="[468,517,1338,1362]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="40" unit="m" value="20.0">20m</quantity>
(secondary forest), 
<date id="FFA21038FFF9FFFC44D35F1BCA500A49" box="[741,871,1338,1362]" pageId="39" pageNumber="40" value="2008-09-30">30.IX.2008</date>
, leg. P. Schwendinger (
<collectionCode id="ED0DAE3DFFF9FFFC425A5F1BCDFF0A49" box="[1132,1224,1338,1362]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="39" pageNumber="40">MHNG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF9FFFC46F15F7FCD560A6D" box="[199,1121,1374,1399]" pageId="39" pageNumber="40" type="etymology">
<paragraph id="8BA336F8FFF9FFFC46F15F7FCD560A6D" blockId="39.[151,1437,834,2010]" box="[199,1121,1374,1399]" pageId="39" pageNumber="40">
<emphasis id="B968EAEAFFF9FFFC46F15F7FC8080A6D" box="[199,319,1374,1398]" italics="true" pageId="39" pageNumber="40">Etymology</emphasis>
: The specific name is a noun in apposition taken from the 
<typeStatus id="54A7885AFFF9FFFC45E65F7ECD370A6C" box="[976,1024,1375,1399]" pageId="39" pageNumber="40">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF9FFFC46F15FA5CC140AF9" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="8BA336F8FFF9FFFC46F15FA5CC140AF9" blockId="39.[151,1437,834,2010]" pageId="39" pageNumber="40">
<emphasis id="B968EAEAFFF9FFFC46F15FA5C80E0A80" box="[199,313,1412,1435]" italics="true" pageId="39" pageNumber="40">Diagnosis</emphasis>
: The male of 
<taxonomicName id="4C1C4D7BFFF9FFFC47EF5FA5CB720A80" box="[473,581,1411,1435]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mulumi">
<emphasis id="B968EAEAFFF9FFFC47EF5FA5CB720A80" box="[473,581,1411,1435]" italics="true" pageId="39" pageNumber="40">I. mulumi</emphasis>
</taxonomicName>
can easily be recognized by the unusual, crown-shaped terminal part of the bulb, with a beak-like apophysis on the ventral side (
<figureCitation id="13272A7DFFF9FFFC44D85F87CA080AA5" box="[750,831,1446,1470]" captionStart="FIGURE 30" captionStartId="40.[151,250,1847,1870]" captionTargetBox="[151,1427,212,1823]" captionTargetId="figure@40.[151,1436,195,1823]" captionTargetPageId="40" captionText="FIGURE 30. Ischnothyreus mulumi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, lateral view; G, left chelicera, posterior view; H, left palp, retrolateral view; I, left bulb, prolateral view; J, left bulb, retrolateral view. stp, strong, tooth-like projection; vp, ventral projection; b, brush; bp, basal process; vpr, ventral protuberance; l, lobe; clm, collar-like membrane; bsa, beak-like apophysis; ri, ribbed. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; G = 0.05 mm; F, H = 0.1 mm; I, J = 0.01 mm." httpUri="https://zenodo.org/record/278141/files/figure.png" pageId="39" pageNumber="40">Fig. 30</figureCitation>
H–J). The female genital area is characterized by the strongly winding tube, ending posteriorly in a circle, surrounded by numerous short setae (
<figureCitation id="13272A7DFFF9FFFC42B85FEBCDE90AF9" box="[1166,1246,1482,1506]" captionStart="FIGURE 31" captionStartId="41.[151,250,1852,1875]" captionTargetBox="[151,1424,203,1824]" captionTargetId="figure@41.[151,1436,193,1826]" captionTargetPageId="41" captionText="FIGURE 31. Ischnothyreus mulumi sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, abdomen, ventral view; G, H, genital area, ventral view; I, J, genital area, dorsal view. a, apodeme; wt, winding tube; hc, half circle. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278142/files/figure.png" pageId="39" pageNumber="40">Fig. 31</figureCitation>
G–J).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF9FFF146F15FD1CAB20ED4" lastPageId="42" lastPageNumber="43" pageId="39" pageNumber="40" type="description">
<paragraph id="8BA336F8FFF9FFFC46F15FD1CCA408C1" blockId="39.[151,1437,834,2010]" pageId="39" pageNumber="40">
<emphasis id="B968EAEAFFF9FFFC46F15FD1C87C091C" box="[199,331,1520,1543]" italics="true" pageId="39" pageNumber="40">Description</emphasis>
: Male (
<typeStatus id="54A7885AFFF9FFFC47915FCFCB38091D" box="[423,527,1518,1542]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFF9FFFC44805FCFCA17091D" box="[694,800,1518,1542]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="39" pageNumber="40" unit="mm" value="1.53">1.53 mm</quantity>
. 
<emphasis id="B968EAEAFFF9FFFC451B5FCECAAC091D" box="[813,923,1519,1542]" italics="true" pageId="39" pageNumber="40">Carapace</emphasis>
: orange-brown, broadly oval in dorsal view, domed in lateral view, surface of elevated portion of pars cephalica strongly reticulate, sides strongly reticulate. 
<emphasis id="B968EAEAFFF9FFFC46A15C16C9C40955" box="[151,243,1591,1614]" italics="true" pageId="39" pageNumber="40">Sternum</emphasis>
: as long as wide, setae abundant. 
<emphasis id="B968EAEAFFF9FFFC444C5C17CBCA0955" box="[634,765,1590,1614]" italics="true" pageId="39" pageNumber="40">Mouthparts</emphasis>
: chelicerae with anteriorly directed process (bp) at base of fangs and brush (b) on retromargin (
<figureCitation id="13272A7DFFF9FFFC441F5C7BCB4F0969" box="[553,632,1626,1650]" captionStart="FIGURE 30" captionStartId="40.[151,250,1847,1870]" captionTargetBox="[151,1427,212,1823]" captionTargetId="figure@40.[151,1436,195,1823]" captionTargetPageId="40" captionText="FIGURE 30. Ischnothyreus mulumi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, lateral view; G, left chelicera, posterior view; H, left palp, retrolateral view; I, left bulb, prolateral view; J, left bulb, retrolateral view. stp, strong, tooth-like projection; vp, ventral projection; b, brush; bp, basal process; vpr, ventral protuberance; l, lobe; clm, collar-like membrane; bsa, beak-like apophysis; ri, ribbed. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; G = 0.05 mm; F, H = 0.1 mm; I, J = 0.01 mm." httpUri="https://zenodo.org/record/278141/files/figure.png" pageId="39" pageNumber="40">Fig. 30</figureCitation>
G), fang groove with few denticles. Anterior margin of labium indented at middle. Anteromedian tip of endites with one strong, tooth-like projection (stp) (
<figureCitation id="13272A7DFFF9FFFC42035C5FCDB3098D" box="[1077,1156,1662,1686]" captionStart="FIGURE 30" captionStartId="40.[151,250,1847,1870]" captionTargetBox="[151,1427,212,1823]" captionTargetId="figure@40.[151,1436,195,1823]" captionTargetPageId="40" captionText="FIGURE 30. Ischnothyreus mulumi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, lateral view; G, left chelicera, posterior view; H, left palp, retrolateral view; I, left bulb, prolateral view; J, left bulb, retrolateral view. stp, strong, tooth-like projection; vp, ventral projection; b, brush; bp, basal process; vpr, ventral protuberance; l, lobe; clm, collar-like membrane; bsa, beak-like apophysis; ri, ribbed. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; G = 0.05 mm; F, H = 0.1 mm; I, J = 0.01 mm." httpUri="https://zenodo.org/record/278141/files/figure.png" pageId="39" pageNumber="40">Fig. 30</figureCitation>
E). 
<emphasis id="B968EAEAFFF9FFFC42855C5FCC2A098D" box="[1203,1309,1662,1686]" italics="true" pageId="39" pageNumber="40">Abdomen</emphasis>
: book lung covers large, elliptical. Pedicel tube medium, scutum extending far dorsal of pedicel. Dorsal scutum strongly sclerotized, orange, anteriorly and posteriorly darkened, covering full length of abdomen, middle surface and sides striated. Epigastric and postepigastric scutum strongly sclerotized, fused, postepigastric scutum long, almost rectangular, covering about 3/4 of abdominal length. Spinneret scutum with fringe of long setae. 
<emphasis id="B968EAEAFFF9FFFC42DC5D31CC28083C" box="[1258,1311,1808,1831]" italics="true" pageId="39" pageNumber="40">Legs</emphasis>
: spine formula: femora: I p0-1-1; r0-1-0; II p0-0-1; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFF9FFFC42C75D12CC6C0850" box="[1265,1371,1843,1867]" italics="true" pageId="39" pageNumber="40">Genitalia</emphasis>
: Palp strongly sclerotized, proximal segments brown, trochanter with ventral projection (vp) (
<figureCitation id="13272A7DFFF9FFFC42585D77CD890875" box="[1134,1214,1878,1902]" captionStart="FIGURE 30" captionStartId="40.[151,250,1847,1870]" captionTargetBox="[151,1427,212,1823]" captionTargetId="figure@40.[151,1436,195,1823]" captionTargetPageId="40" captionText="FIGURE 30. Ischnothyreus mulumi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, lateral view; G, left chelicera, posterior view; H, left palp, retrolateral view; I, left bulb, prolateral view; J, left bulb, retrolateral view. stp, strong, tooth-like projection; vp, ventral projection; b, brush; bp, basal process; vpr, ventral protuberance; l, lobe; clm, collar-like membrane; bsa, beak-like apophysis; ri, ribbed. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; G = 0.05 mm; F, H = 0.1 mm; I, J = 0.01 mm." httpUri="https://zenodo.org/record/278141/files/figure.png" pageId="39" pageNumber="40">Fig. 30</figureCitation>
E), patella about as long as femur, not enlarged, cymbium brown, fused with bulb, bulb brown, more than twice as long as cymbium, stout, tapering apically, with ventral protuberance (vpr) and lobe (l) (
<figureCitation id="13272A7DFFF9FFFC45955DBFCAC308AD" box="[931,1012,1950,1974]" captionStart="FIGURE 30" captionStartId="40.[151,250,1847,1870]" captionTargetBox="[151,1427,212,1823]" captionTargetId="figure@40.[151,1436,195,1823]" captionTargetPageId="40" captionText="FIGURE 30. Ischnothyreus mulumi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, lateral view; G, left chelicera, posterior view; H, left palp, retrolateral view; I, left bulb, prolateral view; J, left bulb, retrolateral view. stp, strong, tooth-like projection; vp, ventral projection; b, brush; bp, basal process; vpr, ventral protuberance; l, lobe; clm, collar-like membrane; bsa, beak-like apophysis; ri, ribbed. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; G = 0.05 mm; F, H = 0.1 mm; I, J = 0.01 mm." httpUri="https://zenodo.org/record/278141/files/figure.png" pageId="39" pageNumber="40">Fig. 30</figureCitation>
H), end of bulb enlarged, ribbed (ri), crown-shaped. Membranous, beak-shaped apophysis (bsa) and collar-like membrane (clm) ventrally (
<figureCitation id="13272A7DFFF9FFFC433F5DE3CC6E08C1" box="[1289,1369,1986,2010]" captionStart="FIGURE 30" captionStartId="40.[151,250,1847,1870]" captionTargetBox="[151,1427,212,1823]" captionTargetId="figure@40.[151,1436,195,1823]" captionTargetPageId="40" captionText="FIGURE 30. Ischnothyreus mulumi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, lateral view; G, left chelicera, posterior view; H, left palp, retrolateral view; I, left bulb, prolateral view; J, left bulb, retrolateral view. stp, strong, tooth-like projection; vp, ventral projection; b, brush; bp, basal process; vpr, ventral protuberance; l, lobe; clm, collar-like membrane; bsa, beak-like apophysis; ri, ribbed. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; G = 0.05 mm; F, H = 0.1 mm; I, J = 0.01 mm." httpUri="https://zenodo.org/record/278141/files/figure.png" pageId="39" pageNumber="40">
Fig. 
<date id="FFA21038FFF9FFFC430B5DE3CC6E08C1" box="[1341,1369,1986,2010]" pageId="39" pageNumber="40" value="1930-01">30</date>
</figureCitation>
I–J).
</paragraph>
<caption id="DF636670FFF6FFF346A15D16CD1E08D3" httpUri="https://zenodo.org/record/278141/files/figure.png" pageId="40" pageNumber="41" targetBox="[151,1427,212,1823]" targetPageId="40">
<paragraph id="8BA336F8FFF6FFF346A15D16CD1E08D3" blockId="40.[151,1437,1847,1992]" pageId="40" pageNumber="41">
<emphasis id="B968EAEAFFF6FFF346A15D16C8280857" bold="true" box="[151,287,1847,1870]" pageId="40" pageNumber="41">FIGURE 30.</emphasis>
<taxonomicName id="4C1C4D7BFFF6FFF347135D16CB320856" box="[293,517,1847,1869]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mulumi" status="sp. nov.">
<emphasis id="B968EAEAFFF6FFF347135D16CB320856" box="[293,517,1847,1869]" italics="true" pageId="40" pageNumber="41">Ischnothyreus mulumi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFF6FFF3443D5D19CB6D0856" box="[523,602,1848,1869]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, lateral view; G, left chelicera, posterior view; H, left palp, retrolateral view; I, left bulb, prolateral view; J, left bulb, retrolateral view. stp, strong, tooth-like projection; vp, ventral projection; b, brush; bp, basal process; vpr, ventral protuberance; l, lobe; clm, collar-like membrane; bsa, beak-like apophysis; ri, ribbed. Scale bars A, B, C = 1 mm; D, E = 0.5 mm; G = 0.05 mm; F, H = 0.1 mm; I, J = 0.01 mm.
</paragraph>
</caption>
<caption id="DF636670FFF7FFF246A15D1DCCBA088C" httpUri="https://zenodo.org/record/278142/files/figure.png" pageId="41" pageNumber="42" targetBox="[151,1424,203,1824]" targetPageId="41">
<paragraph id="8BA336F8FFF7FFF246A15D1DCCBA088C" blockId="41.[151,1436,1852,1943]" pageId="41" pageNumber="42">
<emphasis id="B968EAEAFFF7FFF246A15D1DC817084A" bold="true" box="[151,288,1852,1875]" pageId="41" pageNumber="42">FIGURE 31.</emphasis>
<taxonomicName id="4C1C4D7BFFF7FFF247115D1DCB300849" box="[295,519,1852,1874]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mulumi" status="sp. nov.">
<emphasis id="B968EAEAFFF7FFF247115D1DCB300849" box="[295,519,1852,1874]" italics="true" pageId="41" pageNumber="42">Ischnothyreus mulumi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFF7FFF2443B5D1CCB6A0849" box="[525,605,1853,1874]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, abdomen, ventral view; G, H, genital area, ventral view; I, J, genital area, dorsal view. a, apodeme; wt, winding tube; hc, half circle. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA336F8FFF7FFF146F15DE6CAB20ED4" blockId="41.[199,1436,1991,2016]" lastBlockId="42.[151,1437,151,536]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
Female (
<typeStatus id="54A7885AFFF7FFF2471D5DE6C8A508C4" box="[299,402,1991,2015]" pageId="41" pageNumber="42" type="paratype">paratype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFF7FFF2440C5DE6CB9408C4" box="[570,675,1991,2015]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="41" pageNumber="42" unit="mm" value="1.66">1.66 mm</quantity>
. As in male except as noted. 
<emphasis id="B968EAEAFFF7FFF245CB5DE9CD5C08C4" box="[1021,1131,1992,2015]" italics="true" pageId="41" pageNumber="42">Carapace</emphasis>
: without any pattern, pars cephalica strongly elevated in lateral view. 
<emphasis id="B968EAEAFFF4FFF144A25AB9CBC70FB4" box="[660,752,152,175]" italics="true" pageId="42" pageNumber="43">Sternum</emphasis>
: longer than wide. 
<emphasis id="B968EAEAFFF4FFF145EF5AB9CD6B0FAB" box="[985,1116,152,176]" italics="true" pageId="42" pageNumber="43">Mouthparts</emphasis>
: without basal process and brush. Labium anterior margin not indented at middle. Endites unmodified. Femur of female palp with three ventral spines. 
<emphasis id="B968EAEAFFF4FFF146DD5AC1C8620FE3" box="[235,341,224,248]" italics="true" pageId="42" pageNumber="43">Abdomen</emphasis>
: epigastric scutum slightly protruding (arrow) (
<figureCitation id="13272A7DFFF4FFF145585AFECA880FEC" box="[878,959,223,247]" captionStart="FIGURE 31" captionStartId="41.[151,250,1852,1875]" captionTargetBox="[151,1424,203,1824]" captionTargetId="figure@41.[151,1436,193,1826]" captionTargetPageId="41" captionText="FIGURE 31. Ischnothyreus mulumi sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, abdomen, ventral view; G, H, genital area, ventral view; I, J, genital area, dorsal view. a, apodeme; wt, winding tube; hc, half circle. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278142/files/figure.png" pageId="42" pageNumber="43">Fig. 31</figureCitation>
C). Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length, not fused to epigastric scutum, with short, curved, posteriorly directed lateral apodemes (a) (
<figureCitation id="13272A7DFFF4FFF147D85B06CB080E24" box="[494,575,295,319]" captionStart="FIGURE 31" captionStartId="41.[151,250,1852,1875]" captionTargetBox="[151,1424,203,1824]" captionTargetId="figure@41.[151,1436,193,1826]" captionTargetPageId="41" captionText="FIGURE 31. Ischnothyreus mulumi sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, abdomen, ventral view; G, H, genital area, ventral view; I, J, genital area, dorsal view. a, apodeme; wt, winding tube; hc, half circle. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278142/files/figure.png" pageId="42" pageNumber="43">Fig. 31</figureCitation>
F). Dense patch of setae anterior to spinnerets present. 
<emphasis id="B968EAEAFFF4FFF142825B08CDDE0E5B" box="[1204,1257,297,320]" italics="true" pageId="42" pageNumber="43">Legs</emphasis>
: spine formula: femora: I p0-1-1; r0-1-0; II p0-0-1; v0-0-1; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFF4FFF143375B6DCC5C0E7F" box="[1281,1387,332,356]" italics="true" pageId="42" pageNumber="43">Genitalia</emphasis>
: the posterior margin of the epigastric scutum is lined with numerous needle-like setae. The margin of the postepigastric scutum is thickened. At its middle starts a dark, thin, strongly winding tube running backwards (wt), ending in a dark, half-circle (hc), surrounded by numerous setae (
<figureCitation id="13272A7DFFF4FFF144C65B96CA770ED4" box="[752,832,439,463]" captionStart="FIGURE 31" captionStartId="41.[151,250,1852,1875]" captionTargetBox="[151,1424,203,1824]" captionTargetId="figure@41.[151,1436,193,1826]" captionTargetPageId="41" captionText="FIGURE 31. Ischnothyreus mulumi sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, abdomen, ventral view; G, H, genital area, ventral view; I, J, genital area, dorsal view. a, apodeme; wt, winding tube; hc, half circle. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278142/files/figure.png" pageId="42" pageNumber="43">Fig. 31</figureCitation>
G–J).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF4FFF146F15BFDCA4E0EEF" box="[199,889,476,500]" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8BA336F8FFF4FFF146F15BFDCA4E0EEF" blockId="42.[151,1437,151,536]" box="[199,889,476,500]" pageId="42" pageNumber="43">
<emphasis id="B968EAEAFFF4FFF146F15BFDC8670EEF" box="[199,336,476,500]" italics="true" pageId="42" pageNumber="43">Distribution</emphasis>
: 
<collectingCountry id="F30B7668FFF4FFF147685BFDC8F60EEF" box="[350,449,476,500]" name="Malaysia" pageId="42" pageNumber="43">Sarawak</collectingCountry>
, 
<collectingCountry id="F30B7668FFF4FFF147FB5BFDCB2E0EEF" box="[461,537,476,500]" name="Brunei" pageId="42" pageNumber="43">Brunei</collectingCountry>
and East 
<taxonomicName id="4C1C4D7BFFF4FFF144B15BFDCA3D0EEF" box="[647,778,476,500]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="46" phylum="Arthropoda" rank="species" species="kalimantan">Kalimantan</taxonomicName>
(
<figureCitation id="13272A7DFFF4FFF1452F5BFDCA5B0EEF" box="[793,876,476,500]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="42" pageNumber="43">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF4FFF146F15821C8670DBD" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8BA336F8FFF4FFF146F15821CC0B0D0C" blockId="42.[151,1437,151,536]" box="[199,1340,511,536]" pageId="42" pageNumber="43">
<emphasis id="B968EAEAFFF4FFF146F15821C8290D03" box="[199,286,512,536]" italics="true" pageId="42" pageNumber="43">Remark</emphasis>
: The material collected in 
<collectingCountry id="F30B7668FFF4FFF1447D5BDECBAD0D0C" box="[587,666,511,535]" name="Brunei" pageId="42" pageNumber="43">Brunei</collectingCountry>
, 2009, is kept in 97% ethanol. The specimens are very dark.
</paragraph>
<paragraph id="8BA336F8FFF4FFF146A1584DC8670DBD" blockId="42.[151,468,620,678]" pageId="42" pageNumber="43">
<taxonomicName id="4C1C4D7BFFF4FFF146A1584DC8450D9D" box="[151,370,620,646]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="rex" status="sp. nov.">
<emphasis id="B968EAEAFFF4FFF146A1584DC8450D9D" bold="true" box="[151,370,620,646]" italics="true" pageId="42" pageNumber="43">Ischnothyreus rex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFF4FFF1474F584DC8E30D9D" box="[377,468,620,646]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13272A7DFFF4FFF146A958AFC8170DBD" box="[159,288,654,678]" captionStart-0="FIGURE 32" captionStart-1="FIGURE 33" captionStartId-0="43.[151,250,1825,1848]" captionStartId-1="44.[151,250,1877,1900]" captionTargetBox-0="[151,1429,219,1800]" captionTargetBox-1="[151,1429,251,1855]" captionTargetId-0="figure@43.[151,1436,195,1800]" captionTargetId-1="figure@44.[151,1436,193,1856]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURE 32. Ischnothyreus rex sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, frontal view; G, H, left chelicera, anterior view; I, J, left palp, prolateral view. blc, book-lung covers; stp, strong, tooth-like projection; vp, ventral projection; csp, crown-shaped process; vpr, ventral protuberance; l, lobe; mo, membranous outgrowth; e, embolus. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." captionText-1="FIGURE 33. Ischnothyreus rex sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, G, I, genital area, ventral view; F, left palp, prolateral view; H, genital area, dorsal view. sls, small, lateral sclerite; a, apodeme; yss, y-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G, H, I = 0.1 mm." httpUri-0="https://zenodo.org/record/278143/files/figure.png" httpUri-1="https://zenodo.org/record/278144/files/figure.png" pageId="42" pageNumber="43">Figs 32–33</figureCitation>
, 
<figureCitation id="13272A7DFFF4FFF1471D58AFC87F0DBD" box="[299,328,654,678]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="42" pageNumber="43">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF4FFF146A158F9C8AC0C2D" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8BA336F8FFF4FFF146A158F9C8AC0C2D" blockId="42.[151,1437,726,1975]" pageId="42" pageNumber="43">
<emphasis id="B968EAEAFFF4FFF146A158F9C89B0DF5" box="[151,428,726,751]" italics="true" pageId="42" pageNumber="43">
<typeStatus id="54A7885AFFF4FFF146A158F9C9FE0DF4" box="[151,201,728,751]" pageId="42" pageNumber="43">Type</typeStatus>
material: 
<typeStatus id="54A7885AFFF4FFF1477358F7C89B0DF5" box="[325,428,726,750]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
</emphasis>
: male (PBI_OON 00016064), 
<collectingCountry id="F30B7668FFF4FFF1453B58F7CA940DF5" box="[781,931,726,750]" name="Malaysia" pageId="42" pageNumber="43">MALAYSIA</collectingCountry>
: Sabah: Kinabalu National Park, Power Station, 
<quantity id="4CE49B1DFFF4FFF146E558DBC8140C09" box="[211,291,762,786]" metricMagnitude="3" metricUnit="m" metricValue="1.96" pageId="42" pageNumber="43" unit="m" value="1960.0">1960m</quantity>
, 
<date id="FFA21038FFF4FFF1470458DBC8AD0C09" box="[306,410,762,786]" pageId="42" pageNumber="43" value="1991-05-04">4.V.1991</date>
, leg. C.L. &amp; P.R. Deeleman, 
<collectionCode id="ED0DAE3DFFF4FFF144C658DACA7F0C09" LSID="urn:lsid:biocol.org:col:34992" box="[752,840,763,786]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="42" pageNumber="43">RMNH</collectionCode>
. 
<typeStatus id="54A7885AFFF4FFF1456258DDCA8C0C08" box="[852,955,764,787]" pageId="42" pageNumber="43" type="paratype">
<emphasis id="B968EAEAFFF4FFF1456258DDCA8C0C08" box="[852,955,764,787]" italics="true" pageId="42" pageNumber="43">Paratype</emphasis>
</typeStatus>
: female (PBI_OON 00016065), collected with 
<typeStatus id="54A7885AFFF4FFF146E6593FC8020C2D" box="[208,309,798,822]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED0DAE3DFFF4FFF14776593EC8AF0C2D" LSID="urn:lsid:biocol.org:col:34992" box="[320,408,799,822]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="42" pageNumber="43">RMNH</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF4FFF146F15962CB9E0C65" pageId="42" pageNumber="43" type="etymology">
<paragraph id="8BA336F8FFF4FFF146F15962CB9E0C65" blockId="42.[151,1437,726,1975]" pageId="42" pageNumber="43">
<emphasis id="B968EAEAFFF4FFF146F15962C8080C40" box="[199,319,835,859]" italics="true" pageId="42" pageNumber="43">Etymology</emphasis>
: The specific epithet is a noun in apposition meaning “king” in Latin. It refers to the crown-shaped process on the base of the male cheliceral fang.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF4FFF146F159ADCA000B29" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8BA336F8FFF4FFF146F159ADCA000B29" blockId="42.[151,1437,726,1975]" pageId="42" pageNumber="43">
<emphasis id="B968EAEAFFF4FFF146F159ADC80E0CB8" box="[199,313,908,931]" italics="true" pageId="42" pageNumber="43">Diagnosis</emphasis>
: Very large species. The male of 
<taxonomicName id="4C1C4D7BFFF4FFF1448659ADCBDC0CB8" box="[688,747,908,931]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="B968EAEAFFF4FFF1448659ADCBDC0CB8" box="[688,747,908,931]" italics="true" pageId="42" pageNumber="43">I. rex</emphasis>
</taxonomicName>
resembles the male of 
<taxonomicName id="4C1C4D7BFFF4FFF145D959ADCD740CB8" box="[1007,1091,907,931]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="fobor">
<emphasis id="B968EAEAFFF4FFF145D959ADCD740CB8" box="[1007,1091,907,931]" italics="true" pageId="42" pageNumber="43">I. fobor</emphasis>
</taxonomicName>
in having a very long and thin embolus (
<figureCitation id="13272A7DFFF4FFF14733598FC8620CDD" box="[261,341,942,966]" captionStart="FIGURE 32" captionStartId="43.[151,250,1825,1848]" captionTargetBox="[151,1429,219,1800]" captionTargetId="figure@43.[151,1436,195,1800]" captionTargetPageId="43" captionText="FIGURE 32. Ischnothyreus rex sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, frontal view; G, H, left chelicera, anterior view; I, J, left palp, prolateral view. blc, book-lung covers; stp, strong, tooth-like projection; vp, ventral projection; csp, crown-shaped process; vpr, ventral protuberance; l, lobe; mo, membranous outgrowth; e, embolus. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278143/files/figure.png" pageId="42" pageNumber="43">
Fig. 
<date id="FFA21038FFF4FFF1470F598FC8620CDD" box="[313,341,942,966]" pageId="42" pageNumber="43" value="1932-01">32</date>
</figureCitation>
I–J), but can easily be distinguished from the latter by the crown-shaped basal process on the cheliceral fang (
<figureCitation id="13272A7DFFF4FFF1471E59F3C84D0CF1" box="[296,378,978,1002]" captionStart="FIGURE 32" captionStartId="43.[151,250,1825,1848]" captionTargetBox="[151,1429,219,1800]" captionTargetId="figure@43.[151,1436,195,1800]" captionTargetPageId="43" captionText="FIGURE 32. Ischnothyreus rex sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, frontal view; G, H, left chelicera, anterior view; I, J, left palp, prolateral view. blc, book-lung covers; stp, strong, tooth-like projection; vp, ventral projection; csp, crown-shaped process; vpr, ventral protuberance; l, lobe; mo, membranous outgrowth; e, embolus. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278143/files/figure.png" pageId="42" pageNumber="43">Fig. 32</figureCitation>
F–H). The female of 
<taxonomicName id="4C1C4D7BFFF4FFF1444359F5CB870CF0" box="[629,688,980,1003]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="B968EAEAFFF4FFF1444359F5CB870CF0" box="[629,688,980,1003]" italics="true" pageId="42" pageNumber="43">I. rex</emphasis>
</taxonomicName>
resembles the female of 
<taxonomicName id="4C1C4D7BFFF4FFF145E759F5CD120CF0" box="[977,1061,979,1003]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="flippi">
<emphasis id="B968EAEAFFF4FFF145E759F5CD120CF0" box="[977,1061,979,1003]" italics="true" pageId="42" pageNumber="43">I. flippi</emphasis>
</taxonomicName>
in having a sclerotized y-shaped structure in the genital area, but can be distinguished from the latter by the presence of a dark circle, which seems to be a prolongation of the y-shaped structure (
<figureCitation id="13272A7DFFF4FFF144955E3BCBC40B29" box="[675,755,1050,1074]" captionStart="FIGURE 33" captionStartId="44.[151,250,1877,1900]" captionTargetBox="[151,1429,251,1855]" captionTargetId="figure@44.[151,1436,193,1856]" captionTargetPageId="44" captionText="FIGURE 33. Ischnothyreus rex sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, G, I, genital area, ventral view; F, left palp, prolateral view; H, genital area, dorsal view. sls, small, lateral sclerite; a, apodeme; yss, y-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G, H, I = 0.1 mm." httpUri="https://zenodo.org/record/278144/files/figure.png" pageId="42" pageNumber="43">Fig. 33</figureCitation>
G–I).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF4FFF146F15E1ECDEC0889" pageId="42" pageNumber="43" type="description">
<paragraph id="8BA336F8FFF4FFF146F15E1ECAF8091D" blockId="42.[151,1437,726,1975]" pageId="42" pageNumber="43">
<emphasis id="B968EAEAFFF4FFF146F15E1EC87C0B4D" box="[199,331,1087,1110]" italics="true" pageId="42" pageNumber="43">Description</emphasis>
: Male (
<typeStatus id="54A7885AFFF4FFF1479D5E1FCB230B4D" box="[427,532,1086,1110]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFF4FFF144F75E1FCA1B0B4D" box="[705,812,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="2.94" pageId="42" pageNumber="43" unit="mm" value="2.94">2.94 mm</quantity>
. 
<emphasis id="B968EAEAFFF4FFF1450D5E1ECA9E0B4D" box="[827,937,1087,1110]" italics="true" pageId="42" pageNumber="43">Carapace</emphasis>
: orange-brown, slightly elevated in lateral view, surface and sides of elevated portion of pars cephalica finely reticulate, fovea present, lateral margin rebordered. 
<emphasis id="B968EAEAFFF4FFF146D05EA6C8750B85" box="[230,322,1159,1182]" italics="true" pageId="42" pageNumber="43">Sternum</emphasis>
: longer than wide, pale orange, uniform, not fused to carapace, surface smooth, setae abundant. 
<emphasis id="B968EAEAFFF4FFF146A15E8AC82D0BD8" box="[151,282,1195,1219]" italics="true" pageId="42" pageNumber="43">Mouthparts</emphasis>
: chelicerae slightly divergent, with a prominent, crown-shaped process (csp) at the base of the fangs (
<figureCitation id="13272A7DFFF4FFF146A95EEFC9C70BFD" box="[159,240,1230,1254]" captionStart="FIGURE 32" captionStartId="43.[151,250,1825,1848]" captionTargetBox="[151,1429,219,1800]" captionTargetId="figure@43.[151,1436,195,1800]" captionTargetPageId="43" captionText="FIGURE 32. Ischnothyreus rex sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, frontal view; G, H, left chelicera, anterior view; I, J, left palp, prolateral view. blc, book-lung covers; stp, strong, tooth-like projection; vp, ventral projection; csp, crown-shaped process; vpr, ventral protuberance; l, lobe; mo, membranous outgrowth; e, embolus. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278143/files/figure.png" pageId="42" pageNumber="43">Fig. 32</figureCitation>
F–H). Anteromedian tip of endites with one strong, tooth-like projection (stp) (
<figureCitation id="13272A7DFFF4FFF1425D5EEFCD8B0BFD" box="[1131,1212,1230,1254]" captionStart="FIGURE 32" captionStartId="43.[151,250,1825,1848]" captionTargetBox="[151,1429,219,1800]" captionTargetId="figure@43.[151,1436,195,1800]" captionTargetPageId="43" captionText="FIGURE 32. Ischnothyreus rex sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, frontal view; G, H, left chelicera, anterior view; I, J, left palp, prolateral view. blc, book-lung covers; stp, strong, tooth-like projection; vp, ventral projection; csp, crown-shaped process; vpr, ventral protuberance; l, lobe; mo, membranous outgrowth; e, embolus. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278143/files/figure.png" pageId="42" pageNumber="43">Fig. 32</figureCitation>
E). 
<emphasis id="B968EAEAFFF4FFF142DD5EEFCC620BFD" box="[1259,1365,1230,1254]" italics="true" pageId="42" pageNumber="43">Abdomen</emphasis>
: book lung covers large, elliptical (blc) (
<figureCitation id="13272A7DFFF4FFF144265ED3CB570A11" box="[528,608,1266,1290]" captionStart="FIGURE 32" captionStartId="43.[151,250,1825,1848]" captionTargetBox="[151,1429,219,1800]" captionTargetId="figure@43.[151,1436,195,1800]" captionTargetPageId="43" captionText="FIGURE 32. Ischnothyreus rex sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, frontal view; G, H, left chelicera, anterior view; I, J, left palp, prolateral view. blc, book-lung covers; stp, strong, tooth-like projection; vp, ventral projection; csp, crown-shaped process; vpr, ventral protuberance; l, lobe; mo, membranous outgrowth; e, embolus. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278143/files/figure.png" pageId="42" pageNumber="43">Fig. 32</figureCitation>
C). Scutum extending far dorsal of pedicel. Dorsal scutum yellow-brown, fused to epigastric scutum. Spinneret scutum present, incomplete ring, with fringe of long setae. Colulus present. 
<emphasis id="B968EAEAFFF4FFF146A15F1DC9FB0A48" box="[151,204,1340,1363]" italics="true" pageId="42" pageNumber="43">Legs</emphasis>
: spine formula: femora: I p0-1-1; II p0-1-0; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. Each tibia with three trichobothria, each metatarsus with one trichobothrium. 
<emphasis id="B968EAEAFFF4FFF1458A5F7FCD110A6D" box="[956,1062,1374,1398]" italics="true" pageId="42" pageNumber="43">Genitalia</emphasis>
: Palp strongly sclerotized, proximal segments almost black, trochanter with ventral projection (vp) (
<figureCitation id="13272A7DFFF4FFF145A05FA3CAD00A81" box="[918,999,1410,1434]" captionStart="FIGURE 32" captionStartId="43.[151,250,1825,1848]" captionTargetBox="[151,1429,219,1800]" captionTargetId="figure@43.[151,1436,195,1800]" captionTargetPageId="43" captionText="FIGURE 32. Ischnothyreus rex sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, frontal view; G, H, left chelicera, anterior view; I, J, left palp, prolateral view. blc, book-lung covers; stp, strong, tooth-like projection; vp, ventral projection; csp, crown-shaped process; vpr, ventral protuberance; l, lobe; mo, membranous outgrowth; e, embolus. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278143/files/figure.png" pageId="42" pageNumber="43">Fig. 32</figureCitation>
E), patella about as long as femur, not enlarged, cymbium almost black, fused with bulb, bulb almost black, 1 to 1.5 times as long as cymbium, stout, tapering apically, with ventral protuberance (vpr) and lobe (l), embolus (e) extremely long and thin, filamentous, base of embolus with several membranous outgrowths (mo) (
<figureCitation id="13272A7DFFF4FFF145735FCFCAA2091D" box="[837,917,1518,1542]" captionStart="FIGURE 32" captionStartId="43.[151,250,1825,1848]" captionTargetBox="[151,1429,219,1800]" captionTargetId="figure@43.[151,1436,195,1800]" captionTargetPageId="43" captionText="FIGURE 32. Ischnothyreus rex sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, frontal view; G, H, left chelicera, anterior view; I, J, left palp, prolateral view. blc, book-lung covers; stp, strong, tooth-like projection; vp, ventral projection; csp, crown-shaped process; vpr, ventral protuberance; l, lobe; mo, membranous outgrowth; e, embolus. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm." httpUri="https://zenodo.org/record/278143/files/figure.png" pageId="42" pageNumber="43">
Fig. 
<date id="FFA21038FFF4FFF1454F5FCFCAA2091D" box="[889,917,1518,1542]" pageId="42" pageNumber="43" value="1932-01">32</date>
</figureCitation>
I–J).
</paragraph>
<paragraph id="8BA336F8FFF4FFF146F15C33CDEC0889" blockId="42.[151,1437,726,1975]" pageId="42" pageNumber="43">
Female (
<typeStatus id="54A7885AFFF4FFF1471E5C33C8B80931" box="[296,399,1554,1578]" pageId="42" pageNumber="43" type="paratype">paratype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFF4FFF144065C33CBA00931" box="[560,663,1554,1578]" metricMagnitude="-3" metricUnit="m" metricValue="3.62" pageId="42" pageNumber="43" unit="mm" value="3.62">3.62 mm</quantity>
. As in male except as noted. 
<emphasis id="B968EAEAFFF4FFF145D65C32CD790931" box="[992,1102,1555,1578]" italics="true" pageId="42" pageNumber="43">Carapace</emphasis>
: without any pattern, broadly oval in dorsal view. 
<emphasis id="B968EAEAFFF4FFF1474F5C17CB330955" box="[377,516,1590,1614]" italics="true" pageId="42" pageNumber="43">Mouthparts:</emphasis>
chelicerae without prominent basal process. Anteromedian tip of endites unmodified. Femur of female palp with six ventral spines, tibia with three trichobothria, end of tarsus finger-like (
<figureCitation id="13272A7DFFF4FFF143045C7BCCB60969" box="[1330,1409,1626,1650]" captionStart="FIGURE 33" captionStartId="44.[151,250,1877,1900]" captionTargetBox="[151,1429,251,1855]" captionTargetId="figure@44.[151,1436,193,1856]" captionTargetPageId="44" captionText="FIGURE 33. Ischnothyreus rex sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, G, I, genital area, ventral view; F, left palp, prolateral view; H, genital area, dorsal view. sls, small, lateral sclerite; a, apodeme; yss, y-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G, H, I = 0.1 mm." httpUri="https://zenodo.org/record/278144/files/figure.png" pageId="42" pageNumber="43">Fig. 33</figureCitation>
F, arrow). 
<emphasis id="B968EAEAFFF4FFF146C65C5FC86D098D" box="[240,346,1662,1686]" italics="true" pageId="42" pageNumber="43">Abdomen</emphasis>
: pedicel tube medium. Dorsal scutum covering less than 1/2 of abdomen, not fused to epigastric scutum. Epigastric scutum slightly protruding, with small lateral sclerites (sls) (
<figureCitation id="13272A7DFFF4FFF142175C83CD4409A1" box="[1057,1139,1698,1722]" captionStart="FIGURE 33" captionStartId="44.[151,250,1877,1900]" captionTargetBox="[151,1429,251,1855]" captionTargetId="figure@44.[151,1436,193,1856]" captionTargetPageId="44" captionText="FIGURE 33. Ischnothyreus rex sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, G, I, genital area, ventral view; F, left palp, prolateral view; H, genital area, dorsal view. sls, small, lateral sclerite; a, apodeme; yss, y-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G, H, I = 0.1 mm." httpUri="https://zenodo.org/record/278144/files/figure.png" pageId="42" pageNumber="43">Fig. 33</figureCitation>
E). Postepigastric scutum widely hexagonal, only around epigastric furrow, not fused to epigastric scutum, with short, posteriorly directed lateral apodemes (a) (
<figureCitation id="13272A7DFFF4FFF147BD5CCBC8EB0819" box="[395,476,1770,1794]" captionStart="FIGURE 33" captionStartId="44.[151,250,1877,1900]" captionTargetBox="[151,1429,251,1855]" captionTargetId="figure@44.[151,1436,193,1856]" captionTargetPageId="44" captionText="FIGURE 33. Ischnothyreus rex sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, G, I, genital area, ventral view; F, left palp, prolateral view; H, genital area, dorsal view. sls, small, lateral sclerite; a, apodeme; yss, y-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G, H, I = 0.1 mm." httpUri="https://zenodo.org/record/278144/files/figure.png" pageId="42" pageNumber="43">Fig. 33</figureCitation>
E, G–I). 
<emphasis id="B968EAEAFFF4FFF144715CCDCB4B0818" box="[583,636,1772,1795]" italics="true" pageId="42" pageNumber="43">Legs</emphasis>
: spine formula: femora: I p0-1-1; r0-0-2; II p0-0-1; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFF4FFF144775D2FCB9C083D" box="[577,683,1806,1830]" italics="true" pageId="42" pageNumber="43">Genitalia</emphasis>
: the posterior margin of the epigastric scutum is lined with numerous needle-like setae. The margin of the postepigastric scutum is thickened. At its middle is a y-shaped, sclerotized structure visible (yss), whose posterior part leads to a tube. After overlapping itself, the tube runs as a circle, surrounds the y-shaped structure and finally leads to the strongly winding tube (wt) (
<figureCitation id="13272A7DFFF4FFF1421F5D5BCD4E0889" box="[1065,1145,1914,1938]" captionStart="FIGURE 33" captionStartId="44.[151,250,1877,1900]" captionTargetBox="[151,1429,251,1855]" captionTargetId="figure@44.[151,1436,193,1856]" captionTargetPageId="44" captionText="FIGURE 33. Ischnothyreus rex sp. nov. female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, G, I, genital area, ventral view; F, left palp, prolateral view; H, genital area, dorsal view. sls, small, lateral sclerite; a, apodeme; yss, y-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G, H, I = 0.1 mm." httpUri="https://zenodo.org/record/278144/files/figure.png" pageId="42" pageNumber="43">Fig. 33</figureCitation>
E, G–I).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF4FFF646F15DBEC8110FC9" lastPageId="45" lastPageNumber="46" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8BA336F8FFF4FFF146F15DBECB2408AD" blockId="42.[151,1437,726,1975]" box="[199,531,1950,1975]" pageId="42" pageNumber="43">
<emphasis id="B968EAEAFFF4FFF146F15DBEC86708AC" box="[199,336,1951,1975]" italics="true" pageId="42" pageNumber="43">Distribution</emphasis>
: Sabah (
<figureCitation id="13272A7DFFF4FFF147845DBFCB3108AD" box="[434,518,1950,1974]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="42" pageNumber="43">Fig. 35</figureCitation>
).
</paragraph>
<caption id="DF636670FFF5FFF046A15D00C83708A8" httpUri="https://zenodo.org/record/278143/files/figure.png" pageId="43" pageNumber="44" targetBox="[151,1429,219,1800]" targetPageId="43">
<paragraph id="8BA336F8FFF5FFF046A15D00C83708A8" blockId="43.[151,1436,1825,1971]" pageId="43" pageNumber="44">
<emphasis id="B968EAEAFFF5FFF046A15D00C828082C" bold="true" box="[151,287,1825,1848]" pageId="43" pageNumber="44">FIGURE 32.</emphasis>
<taxonomicName id="4C1C4D7BFFF5FFF047135D03C8EF0823" box="[293,472,1826,1848]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="rex" status="sp. nov.">
<emphasis id="B968EAEAFFF5FFF047135D03C8EF0823" box="[293,472,1826,1848]" italics="true" pageId="43" pageNumber="44">Ischnothyreus rex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFF5FFF047E85D02CB1A0823" box="[478,557,1827,1848]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
male. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, dorsal view; E, carapace, ventral view; F, carapace, frontal view; G, H, left chelicera, anterior view; I, J, left palp, prolateral view. blc, book-lung covers; stp, strong, tooth-like projection; vp, ventral projection; csp, crown-shaped process; vpr, ventral protuberance; l, lobe; mo, membranous outgrowth; e, embolus. Scale bars A, B, C = 1 mm; D, E, F = 0.5 mm; G, H, I, J = 0.1 mm.
</paragraph>
</caption>
<caption id="DF636670FFF2FFF746A15D74C9DB08CA" httpUri="https://zenodo.org/record/278144/files/figure.png" pageId="44" pageNumber="45" targetBox="[151,1429,251,1855]" targetPageId="44">
<paragraph id="8BA336F8FFF2FFF746A15D74C9DB08CA" blockId="44.[151,1436,1877,2001]" pageId="44" pageNumber="45">
<emphasis id="B968EAEAFFF2FFF746A15D74C8160871" bold="true" box="[151,289,1877,1900]" pageId="44" pageNumber="45">FIGURE 33.</emphasis>
<taxonomicName id="4C1C4D7BFFF2FFF7471E5D74C8EB0870" box="[296,476,1877,1899]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="rex" status="sp. nov.">
<emphasis id="B968EAEAFFF2FFF7471E5D74C8EB0870" box="[296,476,1877,1899]" italics="true" pageId="44" pageNumber="45">Ischnothyreus rex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFF2FFF747D25D77CB040870" box="[484,563,1878,1899]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
female. A, habitus, dorsal view; B, habitus, ventral view; C, habitus lateral view; D, carapace, frontal view; E, G, I, genital area, ventral view; F, left palp, prolateral view; H, genital area, dorsal view. sls, small, lateral sclerite; a, apodeme; yss, y-shaped structure; wt, winding tube. Scale bars A, B, C = 1 mm; D = 0.5 mm; E, F, G, H, I = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA336F8FFF3FFF646A15AB6C8110FC9" blockId="45.[151,502,151,210]" pageId="45" pageNumber="46">
<taxonomicName id="4C1C4D7BFFF3FFF646A15AB6C8A30FAA" box="[151,404,151,177]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="serapi" status="sp. nov.">
<emphasis id="B968EAEAFFF3FFF646A15AB6C8A30FAA" bold="true" box="[151,404,151,177]" italics="true" pageId="45" pageNumber="46">Ischnothyreus serapi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5791FFF3FFF647AD5AB9C8C10FA9" box="[411,502,152,178]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13272A7DFFF3FFF646A95A9BC9C20FC9" box="[159,245,186,210]" captionStart="FIGURE 34" captionStartId="46.[151,250,1885,1908]" captionTargetBox="[151,1416,246,1855]" captionTargetId="figure@46.[151,1436,193,1856]" captionTargetPageId="46" captionText="FIGURE 34. Ischnothyreus serapi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, ventral view; G, left palp, prolateral view; H, left palp, retrolateral view. stp, strong, tooth-like projection; sbp, small, basal process; b, brush; csb, crown-shaped basis; ri, ribbed; vpr, ventral protuberance; l, lobe; pa, pointed apophysis. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G, H = 0.05 mm." httpUri="https://zenodo.org/record/278145/files/figure.png" pageId="45" pageNumber="46">Figs 34</figureCitation>
, 
<figureCitation id="13272A7DFFF3FFF647375A9BC8290FC9" box="[257,286,186,210]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="45" pageNumber="46">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF3FFF646A15ADBCD0A0E66" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="8BA336F8FFF3FFF646A15ADBCD0A0E66" blockId="45.[151,1436,249,981]" pageId="45" pageNumber="46">
<emphasis id="B968EAEAFFF3FFF646A15ADBC8990E0A" box="[151,430,249,273]" italics="true" pageId="45" pageNumber="46">
<typeStatus id="54A7885AFFF3FFF646A15ADBC9FE0E0A" box="[151,201,250,273]" pageId="45" pageNumber="46">Type</typeStatus>
material: 
<typeStatus id="54A7885AFFF3FFF647715AD8C8990E0A" box="[327,430,249,273]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
</emphasis>
: male (PBI_OON 00032200), 
<collectingCountry id="F30B7668FFF3FFF645225AD8CA9D0E0A" box="[788,938,249,273]" name="Malaysia" pageId="45" pageNumber="46">MALAYSIA</collectingCountry>
: 
<collectingCountry id="F30B7668FFF3FFF645815AD8CD2B0E0A" box="[951,1052,249,273]" name="Malaysia" pageId="45" pageNumber="46">Sarawak</collectingCountry>
: route Kuching– 
<taxonomicName id="4C1C4D7BFFF3FFF642EB5AD8CC040E0A" box="[1245,1331,249,273]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="matang">Matang</taxonomicName>
, Gunung 
<taxonomicName id="4C1C4D7BFFF3FFF646A15B3CC9D50E2E" box="[151,226,285,309]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="48" phylum="Arthropoda" rank="species" species="serapi">Serapi</taxonomicName>
, prélèvement du sol dans la forêt le long de la route vers la station TV (translated: soil extraction in forest along street in direction of television station), 
<quantity id="4CE49B1DFFF3FFF6449F5B60CBDB0E42" box="[681,748,321,345]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="45" pageNumber="46" unit="m" value="670.0">670m</quantity>
, 
<date id="FFA21038FFF3FFF644CF5B60CA410E42" box="[761,886,321,345]" pageId="45" pageNumber="46" value="1987-12-09">9.XII.1987</date>
, leg. B. Hauser (Berlese à Kuching), Sar-87/64, 
<collectionCode id="ED0DAE3DFFF3FFF646A15B44C9DA0E66" box="[151,237,357,381]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="45" pageNumber="46">MHNG</collectionCode>
. 
<typeStatus id="54A7885AFFF3FFF646C15B47C86A0E66" box="[247,349,358,381]" pageId="45" pageNumber="46" type="paratype">
<emphasis id="B968EAEAFFF3FFF646C15B47C86A0E66" box="[247,349,358,381]" italics="true" pageId="45" pageNumber="46">Paratype</emphasis>
</typeStatus>
: male (PBI_OON 00016218), same data as for 
<typeStatus id="54A7885AFFF3FFF645425B44CAEE0E66" box="[884,985,357,381]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED0DAE3DFFF3FFF645D25B44CD0D0E66" box="[996,1082,357,381]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="45" pageNumber="46">MHNG</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF3FFF646F15BA8CD560EBB" box="[199,1121,392,417]" pageId="45" pageNumber="46" type="etymology">
<paragraph id="8BA336F8FFF3FFF646F15BA8CD560EBB" blockId="45.[151,1436,249,981]" box="[199,1121,392,417]" pageId="45" pageNumber="46">
<emphasis id="B968EAEAFFF3FFF646F15BA8C8080EBA" box="[199,319,393,417]" italics="true" pageId="45" pageNumber="46">Etymology</emphasis>
: The specific name is a noun in apposition taken from the 
<typeStatus id="54A7885AFFF3FFF645E65BA8CD370EBA" box="[976,1024,393,417]" pageId="45" pageNumber="46">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF3FFF646F15B8AC9ED0EF8" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="8BA336F8FFF3FFF646F15B8AC9ED0EF8" blockId="45.[151,1436,249,981]" pageId="45" pageNumber="46">
<emphasis id="B968EAEAFFF3FFF646F15B8AC80E0ED9" box="[199,313,427,450]" italics="true" pageId="45" pageNumber="46">Diagnosis</emphasis>
: The male of 
<taxonomicName id="4C1C4D7BFFF3FFF647EC5B8ACB0F0ED9" box="[474,568,427,450]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="serapi">
<emphasis id="B968EAEAFFF3FFF647EC5B8ACB0F0ED9" box="[474,568,427,450]" italics="true" pageId="45" pageNumber="46">I. serapi</emphasis>
</taxonomicName>
can be recognized by the anvil-shaped distal part of the palpal bulb (
<figureCitation id="13272A7DFFF3FFF6437C5B8BCCAB0ED9" box="[1354,1436,426,450]" captionStart="FIGURE 34" captionStartId="46.[151,250,1885,1908]" captionTargetBox="[151,1416,246,1855]" captionTargetId="figure@46.[151,1436,193,1856]" captionTargetPageId="46" captionText="FIGURE 34. Ischnothyreus serapi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, ventral view; G, left palp, prolateral view; H, left palp, retrolateral view. stp, strong, tooth-like projection; sbp, small, basal process; b, brush; csb, crown-shaped basis; ri, ribbed; vpr, ventral protuberance; l, lobe; pa, pointed apophysis. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G, H = 0.05 mm." httpUri="https://zenodo.org/record/278145/files/figure.png" pageId="45" pageNumber="46">Fig. 34</figureCitation>
F–H).
</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF3FFF646F15BCFC8A90CA8" pageId="45" pageNumber="46" type="description">
<paragraph id="8BA336F8FFF3FFF646F15BCFC9FF0C94" blockId="45.[151,1436,249,981]" pageId="45" pageNumber="46">
<emphasis id="B968EAEAFFF3FFF646F15BCFC87C0D1E" box="[199,331,494,517]" italics="true" pageId="45" pageNumber="46">Description</emphasis>
: Male (
<typeStatus id="54A7885AFFF3FFF647935BCDCB3A0D1F" box="[421,525,492,516]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
). Total length 
<quantity id="4CE49B1DFFF3FFF644845BCDCA2C0D1F" box="[690,795,492,516]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="45" pageNumber="46" unit="mm" value="1.15">1.15 mm</quantity>
. 
<emphasis id="B968EAEAFFF3FFF645115BCCCAA20D1F" box="[807,917,493,516]" italics="true" pageId="45" pageNumber="46">Carapace</emphasis>
: whole carapace strongly reticulate. 
<emphasis id="B968EAEAFFF3FFF6430D5BCFCC470D1E" box="[1339,1392,494,517]" italics="true" pageId="45" pageNumber="46">Eyes</emphasis>
: all subequal, PLE circular. 
<emphasis id="B968EAEAFFF3FFF647935831CB1F0D33" box="[421,552,528,552]" italics="true" pageId="45" pageNumber="46">Mouthparts</emphasis>
: chelicerae, endites and labium yellow. Chelicerae with small process (sbp) at base of the fangs. Brush (b) on retromargin near the basal process (
<figureCitation id="13272A7DFFF3FFF645495815CAF80D57" box="[895,975,564,588]" captionStart="FIGURE 34" captionStartId="46.[151,250,1885,1908]" captionTargetBox="[151,1416,246,1855]" captionTargetId="figure@46.[151,1436,193,1856]" captionTargetPageId="46" captionText="FIGURE 34. Ischnothyreus serapi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, ventral view; G, left palp, prolateral view; H, left palp, retrolateral view. stp, strong, tooth-like projection; sbp, small, basal process; b, brush; csb, crown-shaped basis; ri, ribbed; vpr, ventral protuberance; l, lobe; pa, pointed apophysis. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G, H = 0.05 mm." httpUri="https://zenodo.org/record/278145/files/figure.png" pageId="45" pageNumber="46">Fig. 34</figureCitation>
E). Anteromedian tip of endites with one strong, tooth-like projection (stp) (
<figureCitation id="13272A7DFFF3FFF6442C5879CB5D0D6B" box="[538,618,600,624]" captionStart="FIGURE 34" captionStartId="46.[151,250,1885,1908]" captionTargetBox="[151,1416,246,1855]" captionTargetId="figure@46.[151,1436,193,1856]" captionTargetPageId="46" captionText="FIGURE 34. Ischnothyreus serapi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, ventral view; G, left palp, prolateral view; H, left palp, retrolateral view. stp, strong, tooth-like projection; sbp, small, basal process; b, brush; csb, crown-shaped basis; ri, ribbed; vpr, ventral protuberance; l, lobe; pa, pointed apophysis. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G, H = 0.05 mm." httpUri="https://zenodo.org/record/278145/files/figure.png" pageId="45" pageNumber="46">Fig. 34</figureCitation>
B). 
<emphasis id="B968EAEAFFF3FFF644AD5879CA320D6B" box="[667,773,600,624]" italics="true" pageId="45" pageNumber="46">Abdomen</emphasis>
: dorsal scutum strongly sclerotized, covering full length of abdomen, more than 1/2 to most of abdomen width, middle surface striated, sides striated. Epigastric and postepigastric scutum strongly sclerotized, fused. 
<emphasis id="B968EAEAFFF3FFF644415880CB9B0DA3" box="[631,684,673,696]" italics="true" pageId="45" pageNumber="46">Legs</emphasis>
: spine formula: femora: I p0-1-1; r0-0-1; II p0-0-1; tibiae: I, II p2- 1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0. 
<emphasis id="B968EAEAFFF3FFF644BB58E5CBC10DC7" box="[653,758,708,732]" italics="true" pageId="45" pageNumber="46">Genitalia</emphasis>
: Palp strongly sclerotized, proximal segments brown, trochanter normal size, with ventral projection (vp), femur normal size, patella about as long as femur, not enlarged, cymbium brown, fused with bulb, bulb brown, more than two times as long as cymbium, stout, tapering apically, with ventral protuberance (vpr) and lobe (l), distal part of bulb laterally enlarged, anvil-shaped, ribbed (ri), with crown-shaped basis (csb) and small, pointed apophyses distally (pa) (
<figureCitation id="13272A7DFFF3FFF645995972CD360C70" box="[943,1025,851,875]" captionStart="FIGURE 34" captionStartId="46.[151,250,1885,1908]" captionTargetBox="[151,1416,246,1855]" captionTargetId="figure@46.[151,1436,193,1856]" captionTargetPageId="46" captionText="FIGURE 34. Ischnothyreus serapi sp. nov. male. A, habitus, dorsal view; B, habitus, ventral view; C, carapace, lateral view; D, carapace, frontal view; E, left chelicera, posterior view; F, left palp, ventral view; G, left palp, prolateral view; H, left palp, retrolateral view. stp, strong, tooth-like projection; sbp, small, basal process; b, brush; csb, crown-shaped basis; ri, ribbed; vpr, ventral protuberance; l, lobe; pa, pointed apophysis. Scale bars A, B = 1 mm; C, D = 0.5 mm; E, F, G, H = 0.05 mm." httpUri="https://zenodo.org/record/278145/files/figure.png" pageId="45" pageNumber="46">Fig. 34</figureCitation>
F–H). Embolus not clearly discernible.
</paragraph>
<paragraph id="8BA336F8FFF3FFF646F159BAC8A90CA8" blockId="45.[151,1436,249,981]" box="[199,414,923,947]" pageId="45" pageNumber="46">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3066573FFF3FFF646F1599CCB180CCF" box="[199,559,956,981]" pageId="45" pageNumber="46" type="distribution">
<paragraph id="8BA336F8FFF3FFF646F1599CCB180CCF" blockId="45.[151,1436,249,981]" box="[199,559,956,981]" pageId="45" pageNumber="46">
<emphasis id="B968EAEAFFF3FFF646F1599CC8670CCE" box="[199,336,957,981]" italics="true" pageId="45" pageNumber="46">Distribution</emphasis>
: 
<collectingCountry id="F30B7668FFF3FFF64768599DC8890CCF" box="[350,446,956,980]" name="Malaysia" pageId="45" pageNumber="46">Sarawak</collectingCountry>
(
<figureCitation id="13272A7DFFF3FFF647F8599DCB160CCF" box="[462,545,956,980]" captionStart="FIGURE 35" captionStartId="47.[151,260,1425,1450]" captionTargetBox="[228,1363,156,1406]" captionTargetId="figure@47.[205,1392,121,1430]" captionTargetPageId="47" captionText="FIGURE 35. Records of Ischnothyreus species treated in the present study." httpUri="https://zenodo.org/record/278146/files/figure.png" pageId="45" pageNumber="46">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>