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<document id="F4F171F14EBAFEB8508B0B58E998F7DA" ID-DOI="10.11646/zootaxa.5323.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8209415" ID-ZooBank="C022FD72-9C54-4A89-A880-3D99F52C8499" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691006370328" checkinUser="plazi" docAuthor="Tan, Ming Kai, Salvador, Jewel Anne G., Sabang, Aira Mae M., Bahoy, Daphne Cayle M., Nuñeza, Olga Macas &amp; Robillard, Tony" docDate="2023" docId="03B587EEE76767670CFEC08A281FFE28" docLanguage="en" docName="zootaxa.5323.3.1.pdf" docOrigin="Zootaxa 5323 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5323.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mistshenkoana higaonon Tan &amp; Salvador &amp; Sabang &amp; Bahoy &amp; Nuñeza &amp; Robillard 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="333" masterDocId="FF8CFF96E77A67470C69C3582A4BFFA1" masterDocTitle="Taxonomy and bioacoustics of little-known Grylloidea crickets (Orthoptera, Ensifera) from Mindanao, Philippines" masterLastPageNumber="348" masterPageNumber="301" pageNumber="330" updateTime="1691007180022" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B42C983759332722848DA8E228B4BF9E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="153EF2A7970DF40C031A29DDF4E3FC9D">
<mods:title id="7A49B70B79598DF205F22D29B2383B1B">Taxonomy and bioacoustics of little-known Grylloidea crickets (Orthoptera, Ensifera) from Mindanao, Philippines</mods:title>
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<mods:namePart id="D772E7ED6D79CAE9B86FFE20DB417298">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="05C2072251BC40BC4D65E17F11D01053">Block 207 A, Woodleigh Link, Singapore 361207, Republic of Singapore</mods:affiliation>
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<mods:namePart id="CD0168BDB10078218439B9758D7DCEA7">Salvador, Jewel Anne G.</mods:namePart>
<mods:affiliation id="3312B0D80D565D157D1B75D43ADE53A5">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
</mods:name>
<mods:name id="6D70DE2EE99803469AC94DDEA7F2A42C" type="personal">
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</mods:role>
<mods:namePart id="A3DD42F34C630424C2A58FFE72CF747D">Sabang, Aira Mae M.</mods:namePart>
<mods:affiliation id="0949E3CADE4E7C935BA1C8F5D7C782D3">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
</mods:name>
<mods:name id="CCF7444CA323F0A9F9ACEA57FE9475AA" type="personal">
<mods:role id="AD528740B9A4ECE7F622A9EF2315FF5B">
<mods:roleTerm id="DEB2BCE7CA1ED5E1149CD07667D8923D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="64792020891E71D77D53A34165C14A55">Bahoy, Daphne Cayle M.</mods:namePart>
<mods:affiliation id="263F74B3F054E01373AC89629FD48D3F">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
</mods:name>
<mods:name id="860E681262C7F60E94A85FC507C78782" type="personal">
<mods:role id="140CFC0FBAAE2AB073FF39CEDD68A2C4">
<mods:roleTerm id="7B09B13CF08A3041A2379E62B8054C8C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E8CB5D0FA45C1DDD8106AB8E7654E98B">Nuñeza, Olga Macas</mods:namePart>
<mods:affiliation id="041255F653129178CEAFED0D269A9D8E">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="8852C73F6D14555D6F8C901ED56D27D0">Robillard, Tony</mods:namePart>
<mods:affiliation id="2F3E32C7E3E048B746C88B8AC37063DF">Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
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<mods:titleInfo id="AE85A823DA7B55AC00AFB31FCEE2A12B">
<mods:title id="AFACC1BBDA97B60A2A72EAE53C0D7F7A">Zootaxa</mods:title>
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<mods:part id="B21CBF73FE62395AFF083CEAFFB7EE5C">
<mods:date id="4812A093139A2090360D1645F3A655BB">2023</mods:date>
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<mods:number id="CFDFF52D807F7CCAD96115CF54EE3051">2023-08-02</mods:number>
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<mods:number id="B9C13FB2ACC425E6041471B4BFD59A43">5323</mods:number>
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<mods:number id="4A2C98B5A022931A5AD69EF8812650B7">3</mods:number>
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<mods:url id="EE8CB551C07E6367FBB8E367C4FCFBD9">http://dx.doi.org/10.11646/zootaxa.5323.3.1</mods:url>
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<mods:classification id="CEA470FA6C15A96D3F4159D248390057">journal article</mods:classification>
<mods:identifier id="9234C16723ECE78338F8467A1CA3AC30" type="DOI">10.11646/zootaxa.5323.3.1</mods:identifier>
<mods:identifier id="D1EB8DBC87C01ACD843DDA11142C660B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7A7890776BB759821D18FC9048E53B7B" type="Zenodo-Dep">8209415</mods:identifier>
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<treatment id="03B587EEE76767670CFEC08A281FFE28" LSID="urn:lsid:plazi:treatment:03B587EEE76767670CFEC08A281FFE28" httpUri="http://treatment.plazi.org/id/03B587EEE76767670CFEC08A281FFE28" lastPageId="32" lastPageNumber="333" pageId="29" pageNumber="330">
<subSubSection id="C3066573E767675A0CFEC08A294FFC4C" box="[151,772,978,1005]" pageId="29" pageNumber="330" type="nomenclature">
<paragraph id="8BA336F8E767675A0CFEC08A294FFC4C" blockId="29.[151,772,978,1041]" box="[151,772,978,1005]" pageId="29" pageNumber="330">
<heading id="D0EB8194E767675A0CFEC08A294FFC4C" bold="true" box="[151,772,978,1005]" fontSize="11" level="1" pageId="29" pageNumber="330" reason="1">
<emphasis id="B968EAEAE767675A0CFEC08A294FFC4C" bold="true" box="[151,772,978,1005]" pageId="29" pageNumber="330">
<taxonomicName id="4C1C4D7BE767675A0CFEC08A2848FC4C" authority="Tan" authorityName="Tan &amp; Salvador &amp; Sabang &amp; Bahoy &amp; Nuñeza &amp; Robillard" authorityYear="2023" box="[151,515,978,1005]" class="Insecta" family="Gryllidae" genus="Mistshenkoana" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="higaonon">
<emphasis id="B968EAEAE767675A0CFEC08A2B87FC4C" bold="true" box="[151,460,978,1005]" italics="true" pageId="29" pageNumber="330">Mistshenkoana higaonon</emphasis>
Tan
</taxonomicName>
&amp;
<collectingCountry id="F30B7668E767675A0E43C08A28D1FC4D" box="[554,666,978,1004]" name="El Salvador" pageId="29" pageNumber="330">Salvador</collectingCountry>
,
<taxonomicNameLabel id="A25B5791E767675A0ECEC08B294FFC4C" box="[679,772,979,1005]" pageId="29" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3066573E767675A0CFEC0AF2B96FB00" pageId="29" pageNumber="330" type="description">
<paragraph id="8BA336F8E767675A0CFEC0AF2B6CFBB0" blockId="29.[151,772,978,1041]" box="[151,295,1015,1041]" pageId="29" pageNumber="330">
(
<figureCitation id="13272A7DE767675A0CF6C0AF2ABDFBB0" box="[159,246,1015,1041]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="29" pageNumber="330">Figs 26</figureCitation>
,
<figureCitation id="13272A7DE767675A0D6BC0AF2B54FBB0" box="[258,287,1015,1041]" captionStart="FIGURE 27" captionStartId="31.[151,250,1545,1569]" captionTargetBox="[151,1436,787,1513]" captionTargetId="figure-281@31.[151,1436,777,1522]" captionTargetPageId="31" captionText="FIGURE 27. Mistshenkoana higaonon sp. nov. ♁ genitalia in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8209485" httpUri="https://zenodo.org/record/8209485/files/figure.png" pageId="29" pageNumber="330">27</figureCitation>
)
</paragraph>
<paragraph id="8BA336F8E767675A0CFEC7662B96FB00" blockId="29.[151,1437,1086,2013]" pageId="29" pageNumber="330">
<emphasis id="B968EAEAE767675A0CFEC7662BD7FBF9" bold="true" box="[151,412,1086,1112]" pageId="29" pageNumber="330">Specimens examined.</emphasis>
<materialsCitation id="3B743CA5E767675A0DC0C7672B96FB00" collectingDate="2023-02-16" collectionCode="PNM" collectorName="J. A. G. Salvador" country="Philippines" county="Mindanao" latitude="8.70115" location="Barangay Lunotan" longLatPrecision="1" longitude="125.01004" municipality="Gingoog City" pageId="29" pageNumber="330" specimenCount="1" stateProvince="Misamis Oriental" typeStatus="holotype">
<emphasis id="B968EAEAE767675A0DC0C7672854FBF8" bold="true" box="[425,543,1087,1113]" pageId="29" pageNumber="330">
<typeStatus id="54A7885AE767675A0DC0C7672851FBF8" box="[425,538,1087,1113]" pageId="29" pageNumber="330">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F30B7668E767675A0E45C767289AFBF8" box="[556,721,1087,1113]" name="Philippines" pageId="29" pageNumber="330">PHILIPPINES</collectingCountry>
• ♁;
<collectingCounty id="62C24E74E767675A0F77C76729D9FBF8" box="[798,914,1087,1113]" pageId="29" pageNumber="330">Mindanao</collectingCounty>
,
<collectingRegion id="49D8F81AE767675A0FCAC7672E24FBF8" box="[931,1135,1087,1113]" country="Philippines" name="Misamis Oriental" pageId="29" pageNumber="330">Misamis Oriental</collectingRegion>
,
<collectingMunicipality id="6BC7AC82E767675A08E9C7672F6BFBF8" box="[1152,1312,1087,1113]" pageId="29" pageNumber="330">Gingoog City</collectingMunicipality>
,
<location id="8EC36023E767675A0958C7672ABCFBDC" LSID="urn:lsid:plazi:treatment:03B587EEE76767670CFEC08A281FFE28:8EC36023E767675A0958C7672ABCFBDC" country="Philippines" county="Mindanao" latitude="8.70115" longLatPrecision="1" longitude="125.01004" municipality="Gingoog City" name="Barangay Lunotan" pageId="29" pageNumber="330" stateProvince="Misamis Oriental">Barangay Lunotan</location>
, near
<location id="8EC36023E767675A0D54C73B2B95FBDC" LSID="urn:lsid:plazi:treatment:03B587EEE76767670CFEC08A281FFE28:8EC36023E767675A0D54C73B2B95FBDC" box="[317,478,1123,1149]" country="Philippines" county="Mindanao" latitude="8.70115" longLatPrecision="1" longitude="125.01004" municipality="Gingoog City" name="Mt Sumagaya" pageId="29" pageNumber="330" stateProvince="Misamis Oriental">Mt Sumagaya</location>
;
<geoCoordinate id="EE28503FE767675A0D82C73A2816FBDC" box="[491,605,1122,1149]" degrees="8.70115" direction="north" orientation="latitude" pageId="29" pageNumber="330" precision="1" value="8.70115">N8.70115</geoCoordinate>
,
<geoCoordinate id="EE28503FE767675A0E00C73A28BFFBDC" box="[617,756,1122,1149]" degrees="125.01004" direction="east" orientation="longitude" pageId="29" pageNumber="330" precision="1" value="125.01004">E125.01004</geoCoordinate>
, 1206.0±7.0 m.a.s.l.;
<date id="FFA21038E767675A0F83C73B2EFCFBDC" box="[1002,1207,1123,1149]" pageId="29" pageNumber="330" value="2023-02-16">
<collectingDate id="EFE6E9D0E767675A0F83C73B2EFCFBDC" box="[1002,1207,1123,1149]" pageId="29" pageNumber="330" value="2023-02-16">16 February 2023</collectingDate>
</date>
, 20h09; coll.
<collectorName id="26E9532EE767675A093DC73B2AB5FB00" pageId="29" pageNumber="330">J.A.G. Salvador</collectorName>
; MIN.23.67 (
<collectionCode id="ED0DAE3DE767675A0DFCC7DF2B9EFB00" box="[405,469,1159,1185]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="29" pageNumber="330">PNM</collectionCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3066573E767675A0CAEC7F228AEFA44" pageId="29" pageNumber="330" type="diagnosis">
<paragraph id="8BA336F8E767675A0CAEC7F22BC6FA90" blockId="29.[151,1437,1086,2013]" pageId="29" pageNumber="330">
<emphasis id="B968EAEAE767675A0CAEC7F22B09FB65" bold="true" box="[199,322,1194,1220]" pageId="29" pageNumber="330">Diagnosis.</emphasis>
The new species differs from the aforementioned species and
<collectingCountry id="F30B7668E767675A084AC7F32EEAFB64" box="[1059,1185,1195,1221]" name="Philippines" pageId="29" pageNumber="330">Philippines</collectingCountry>
congeners by similar characters mentioned in
<taxonomicName id="4C1C4D7BE767675A0DC3C79728FBFB48" authority="Tan &amp; Salvador &amp; Sabang &amp; Bahoy &amp; Nuñeza &amp; Robillard, 2023" authorityName="Tan &amp; Salvador &amp; Sabang &amp; Bahoy &amp; Nuñeza &amp; Robillard" authorityYear="2023" box="[426,688,1231,1257]" class="Insecta" family="Gryllidae" genus="Mistshenkoana" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="lunotan" status="sp. nov.">
<emphasis id="B968EAEAE767675A0DC3C79728FBFB48" box="[426,688,1231,1257]" italics="true" pageId="29" pageNumber="330">Mistshenkoana lunotan</emphasis>
</taxonomicName>
<emphasis id="B968EAEAE767675A0EDEC797295AFB48" bold="true" box="[695,785,1231,1257]" pageId="29" pageNumber="330">
<taxonomicNameLabel id="A25B5791E767675A0EDEC797295AFB48" box="[695,785,1231,1257]" pageId="29" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but it also differs by the head and pronotum having more dark-brown spots; the FW dorsal field with three large distinct white oblique and elongated spots and the FIIIs with mottled dark patterns.
</paragraph>
<paragraph id="8BA336F8E767675A0CAEC66328AEFA44" blockId="29.[151,1437,1086,2013]" pageId="29" pageNumber="330">
From
<taxonomicName id="4C1C4D7BE767675A0D62C663285AFAF4" authority="Tan &amp; Salvador &amp; Sabang &amp; Bahoy &amp; Nuñeza &amp; Robillard, 2023" authorityName="Tan &amp; Salvador &amp; Sabang &amp; Bahoy &amp; Nuñeza &amp; Robillard" authorityYear="2023" box="[267,529,1339,1365]" class="Insecta" family="Gryllidae" genus="Mistshenkoana" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="lunotan" status="sp. nov.">
<emphasis id="B968EAEAE767675A0D62C663285AFAF4" box="[267,529,1339,1365]" italics="true" pageId="29" pageNumber="330">Mistshenkoana lunotan</emphasis>
</taxonomicName>
<emphasis id="B968EAEAE767675A0E71C663283AFAF4" bold="true" box="[536,625,1339,1365]" pageId="29" pageNumber="330">
<taxonomicNameLabel id="A25B5791E767675A0E71C663283AFAF4" box="[536,625,1339,1365]" pageId="29" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the male genitalia of this new species also differ by the pseudepiphallic lophi, in profile view, having dorsal margin somewhat straight throughout but ventral margin at middle bent, and basal half tapering more than apical half (instead of simply triangular); and by the pseudepiphallic parameres, in ventral view, with posterior half having a small spine-like process and in profile view, ventral lobe broader than dorsal lobe (instead of about as broad as each other).
</paragraph>
</subSubSection>
<subSubSection id="C3066573E767675A0CAEC6B72ED0F98C" pageId="29" pageNumber="330" type="etymology">
<paragraph id="8BA336F8E767675A0CAEC6B72ED0F98C" blockId="29.[151,1437,1086,2013]" pageId="29" pageNumber="330">
<emphasis id="B968EAEAE767675A0CAEC6B72B05F9A8" bold="true" box="[199,334,1519,1545]" pageId="29" pageNumber="330">Etymology.</emphasis>
The species is named after the Higaonon tribe of
<collectingRegion id="49D8F81AE767675A0FE1C6B72982F9A8" box="[904,969,1519,1545]" country="Philippines" name="Iligan" pageId="29" pageNumber="330">Iligan</collectingRegion>
which the species coexists with; noun in apposition. Higaonon is also derived from the native words higa (living), goan (mountains).
</paragraph>
</subSubSection>
<subSubSection id="C3066573E76767670CAEC56E2823FF59" lastPageId="32" lastPageNumber="333" pageId="29" pageNumber="330" type="description">
<paragraph id="8BA336F8E76767580CAEC56E2B1CFE70" blockId="29.[151,1437,1086,2013]" lastBlockId="31.[151,1437,151,753]" lastPageId="31" lastPageNumber="332" pageId="29" pageNumber="330">
<emphasis id="B968EAEAE767675A0CAEC56E2B12F9F1" bold="true" box="[199,345,1590,1616]" pageId="29" pageNumber="330">Description.</emphasis>
Habitus typical of the genus, medium sized, cream-coloured to yellow brown with dark-brown mottled patterns (
<figureCitation id="13272A7DE767675A0D09C5032846F9D4" box="[352,525,1627,1653]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="29" pageNumber="330">Figs 26A, 26B</figureCitation>
). Head with dorsum dark brown mottled, pubescent. Rostrum finely pubescent, dark coloured, about 0.9 times as wide as scapes (
<figureCitation id="13272A7DE767675A0EAEC5272964F938" box="[711,815,1663,1689]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="29" pageNumber="330">Fig. 26A</figureCitation>
). Scapes yellow brown with brown patterns. Eyes view distinctly taller than long (
<figureCitation id="13272A7DE767675A0DAAC5FB2860F91C" box="[451,555,1699,1725]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="29" pageNumber="330">Fig. 26C</figureCitation>
). Median ocellus oval; lateral ocelli located near eyes. Maxillary palpi cream-coloured; with apical segment longest, dorso-posteriorly dark-coloured, flattened and somewhat rectangular, with obliquely-truncated apex; with subapical segment shorter and faintly expanding apically; third segment longer than subapical segment, cylindrical and slightly expanding apically (
<figureCitation id="13272A7DE767675A0F18C4572997F888" box="[881,988,1807,1833]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="29" pageNumber="330">Fig. 26D</figureCitation>
). Face in anterior view as tall as wide; mostly cream-coloured except between scapes (
<figureCitation id="13272A7DE767675A0ED2C46B2968F8EC" box="[699,803,1843,1869]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="29" pageNumber="330">Fig. 26E</figureCitation>
). Gena cream-coloured, posterior of eye with two dark brown bands (
<figureCitation id="13272A7DE767675A0D50C40F2BEAF8D0" box="[313,417,1879,1905]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="29" pageNumber="330">Fig. 26C</figureCitation>
). Pronotal disk brown with dense dark-brown mottled patterns; 1.4 times as wide as long, widening posteriorly (posterior margin 1.6 times as wide as anterior margin); densely and finely pubescent, with a row of setae along posterior and anterior margins; anterior margin of disk straight; posterior margin of disk broadly convex in middle (
<figureCitation id="13272A7DE767675A0D0EC49B2B84F87C" box="[359,463,1987,2013]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="29" pageNumber="330">Fig. 26A</figureCitation>
). Pronotal lateral lobe 1.2 times as long as high, cream-coloured with some dark-brown mottled patterns (
<figureCitation id="13272A7DE76567580D35C3CF2B8FFF10" box="[348,452,151,177]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="31" pageNumber="332">Fig. 26C</figureCitation>
). Metanotal gland absent. TI swollen; with inner tympanum open and having very large elongate, oblong tympanal membrane; with outer tympanum obscure (nearly distinguishable). TIII with 5 inner and 4 outer long subapical spurs, 3 larger inner and 3 small outer apical spurs; before most proximal subapical spurs and between subapical spurs with small spines. Legs generally cream-coloured with irregular and sparse dark spots; Fs with more dark spots in the apical half; Ts generally with more dark spots. FIII with a dark brown mottled patterns on inner and outer parts; in dorsal view with three oblique bands, apical band thickest, followed by basal and middle bands. TIII in dorsal view with dark patches and spots. TaIII basal segment cream-coloured with basal and apical parts dark coloured. Thoracic and abdominal segments cream-coloured; abdominal tergite with some dark colouration.
</paragraph>
<caption id="DF636670E76467590CFEC4CF2BEDF859" ID-DOI="http://doi.org/10.5281/zenodo.8209483" ID-Zenodo-Dep="8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="30" pageNumber="331" startId="30.[151,250,1943,1967]" targetBox="[177,1410,196,1914]" targetPageId="30" targetType="figure">
<paragraph id="8BA336F8E76467590CFEC4CF2BEDF859" blockId="30.[151,1437,1943,2040]" pageId="30" pageNumber="331">
<emphasis id="B968EAEAE76467590CFEC4CF2B69F80E" bold="true" box="[151,290,1943,1967]" pageId="30" pageNumber="331">FIGURE 26.</emphasis>
<taxonomicName id="4C1C4D7BE76467590D42C4CF2866F80E" authority="Tan &amp; Salvador &amp; Sabang &amp; Bahoy &amp; Nuñeza &amp; Robillard, 2023" authorityName="Tan &amp; Salvador &amp; Sabang &amp; Bahoy &amp; Nuñeza &amp; Robillard" authorityYear="2023" box="[299,557,1943,1967]" class="Insecta" family="Gryllidae" genus="Mistshenkoana" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="higaonon" status="sp. nov.">
<emphasis id="B968EAEAE76467590D42C4CF2866F80E" box="[299,557,1943,1967]" italics="true" pageId="30" pageNumber="331">Mistshenkoana higaonon</emphasis>
</taxonomicName>
<emphasis id="B968EAEAE76467590E5CC4C028C3F811" bold="true" box="[565,648,1944,1968]" pageId="30" pageNumber="331">
<taxonomicNameLabel id="A25B5791E76467590E5CC4C028C3F811" box="[565,648,1944,1968]" pageId="30" pageNumber="331" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E).
</paragraph>
</caption>
<paragraph id="8BA336F8E76567580CAEC2832907FD50" blockId="31.[151,1437,151,753]" pageId="31" pageNumber="332">
<emphasis id="B968EAEAE76567580CAEC2832B40FE54" bold="true" box="[199,267,475,501]" pageId="31" pageNumber="332">Male.</emphasis>
FW (
<figureCitation id="13272A7DE76567580D23C2832BA7FE54" box="[330,492,475,501]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="31" pageNumber="332">Figs 26F, 26G</figureCitation>
) covering abdomen and surpassing apex of FIII. Colouration: cells generally yellow brown, veins brown; cross-veins at basal half typically cream-coloured, at apical half with faint black patch around cross-veins. Along M with three elongated white/ cream-coloured spots: at the basal third (smallest), in the middle (largest) and near apical third. Lateral field with intercalary triangle transparent, otherwise yellow brown with brown veins and cream-coloured cross-veins (
<figureCitation id="13272A7DE76567580E31C133288FFD24" box="[600,708,619,645]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="31" pageNumber="332">Fig. 26G</figureCitation>
). Venation: without stridulatory organs, with longitudinal veins parallel; lateral field with R and M narrowly spaced apart, mostly parallel and with cross-veins cream-coloured; with R and Sc spaced apart, parallel and with cross-veins (especially in the basal half); branches of Sc generally parallel. Hind wings clearly surpassing FWs (
<figureCitation id="13272A7DE76567580EFFC18F290BFD50" box="[662,832,727,753]" captionStart="FIGURE 26" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[177,1410,196,1914]" captionTargetId="figure-17@30.[177,1410,181,1919]" captionTargetPageId="30" captionText="FIGURE 26. Mistshenkoana higaonon sp. nov.: dorsal (A) and lateral (B) habitus; head and pronotum in lateral view (C); maxillary palps in lateral view (D); face in anterior view (E); FWs in dorsal (F) and lateral (G) views. Scale bars: 5 mm (A, B), 2 mm (F, G), 1 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.8209483" httpUri="https://zenodo.org/record/8209483/files/figure.png" pageId="31" pageNumber="332">Figs 26A, 26B</figureCitation>
).
</paragraph>
<caption id="DF636670E76567580CFEC5512FC1F980" ID-DOI="http://doi.org/10.5281/zenodo.8209485" ID-Zenodo-Dep="8209485" box="[151,1418,1545,1572]" httpUri="https://zenodo.org/record/8209485/files/figure.png" pageId="31" pageNumber="332" startId="31.[151,250,1545,1569]" targetBox="[151,1436,787,1513]" targetPageId="31" targetType="figure">
<paragraph id="8BA336F8E76567580CFEC5512FC1F980" blockId="31.[151,1418,1545,1572]" box="[151,1418,1545,1572]" pageId="31" pageNumber="332">
<emphasis id="B968EAEAE76567580CFEC5512B6BF980" bold="true" box="[151,288,1545,1569]" pageId="31" pageNumber="332">FIGURE 27.</emphasis>
<taxonomicName id="4C1C4D7BE76567580D4FC552286DF983" authority="Tan &amp; Salvador &amp; Sabang &amp; Bahoy &amp; Nuñeza &amp; Robillard, 2023" authorityName="Tan &amp; Salvador &amp; Sabang &amp; Bahoy &amp; Nuñeza &amp; Robillard" authorityYear="2023" box="[294,550,1546,1570]" class="Insecta" family="Gryllidae" genus="Mistshenkoana" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="332" phylum="Arthropoda" rank="species" species="higaonon" status="sp. nov.">
<emphasis id="B968EAEAE76567580D4FC552286DF983" box="[294,550,1546,1570]" italics="true" pageId="31" pageNumber="332">Mistshenkoana higaonon</emphasis>
</taxonomicName>
<emphasis id="B968EAEAE76567580E45C5522837F983" bold="true" box="[556,636,1546,1570]" pageId="31" pageNumber="332">
<taxonomicNameLabel id="A25B5791E76567580E45C5522837F983" box="[556,636,1546,1570]" pageId="31" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♁ genitalia in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BA336F8E76567580CAEC508291BF873" blockId="31.[151,1437,1616,2002]" pageId="31" pageNumber="332">
Subgenital plate typical of genus, elongated and tapering apically; cream-coloured. Male genitalia as shown in
<figureCitation id="13272A7DE76567580CFEC52C2AA2F92F" box="[151,233,1652,1678]" captionStart="FIGURE 27" captionStartId="31.[151,250,1545,1569]" captionTargetBox="[151,1436,787,1513]" captionTargetId="figure-281@31.[151,1436,777,1522]" captionTargetPageId="31" captionText="FIGURE 27. Mistshenkoana higaonon sp. nov. ♁ genitalia in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8209485" httpUri="https://zenodo.org/record/8209485/files/figure.png" pageId="31" pageNumber="332">Fig. 27</figureCitation>
: pseudepiphallus [epiphallus] not so elongated, tapering slightly posteriorly; at posterior end divided into two parts and roundly and very narrowly excised; posterior end produced into pseudepiphallic lophi [apical epiphallic lobe]. Pseudepiphallic lophi [apical epiphallic lobe], in dorsal and ventral views, long and slender and pointing posterior-externally with apex obtuse; inner margins with a few setae. In profile view, dorsal margin somewhat straight throughout, ventral margin at middle bent; basal half tapering more than apical half; apical half slender triangular; apex subacute; ventral margin with a few setae. Pseudepiphallic paramere [ectoparamere], in ventral view, elongated and slender; with anterior half (also the ventral lobe) rhombic, at middle folded, with posterior half (also the dorsal lobe) with rounded apex and a small spine-like process at the inner-apical part. In profile view, ventral lobe (vl) broader than dorsal lobe (dl), between the lobes rounded emarginated. Ectophallic fold [rachis] small, slender and straight, taper posteriorly with apex obtuse.
</paragraph>
<paragraph id="8BA336F8E75A67670CAEC3CE2BE9FF11" blockId="32.[151,1437,150,393]" box="[199,418,150,176]" pageId="32" pageNumber="333">
<emphasis id="B968EAEAE75A67670CAEC3CE2B6EFF11" bold="true" box="[199,293,150,176]" pageId="32" pageNumber="333">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BA336F8E75A67670CAEC3E22823FF59" blockId="32.[151,1437,150,393]" pageId="32" pageNumber="333">
<emphasis id="B968EAEAE75A67670CAEC3E22BAFFF75" bold="true" box="[199,484,186,212]" pageId="32" pageNumber="333">Measurements (in mm).</emphasis>
BL = 11.3; BWL = 18.9; HL = 0.8; PronL = 1.6; PronW = 2.2; FWL = 12.3; FWW = 2.3; HWT = 4.5; FIIIL = 8.1; TIIIL = 8.4.
</paragraph>
</subSubSection>
<subSubSection id="C3066573E75A67670CAEC25A2BE2FEBD" box="[199,425,258,284]" pageId="32" pageNumber="333" type="biology_ecology">
<paragraph id="8BA336F8E75A67670CAEC25A2BE2FEBD" blockId="32.[151,1437,150,393]" box="[199,425,258,284]" pageId="32" pageNumber="333">
<emphasis id="B968EAEAE75A67670CAEC25A2B67FEBD" bold="true" box="[199,300,258,284]" pageId="32" pageNumber="333">Ecology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3066573E75A67670CAEC27E2F7BFEE0" box="[199,1328,294,321]" pageId="32" pageNumber="333" type="materials_examined">
<paragraph id="8BA336F8E75A67670CAEC27E2F7BFEE0" blockId="32.[151,1437,150,393]" box="[199,1328,294,321]" pageId="32" pageNumber="333">
<materialsCitation id="3B743CA5E75A67670CAEC27E2F7BFEE0" box="[199,1328,294,321]" country="Philippines" county="Mindanao Island" location="Barangay Lunotan" municipality="Gingoog City" pageId="32" pageNumber="333" specimenCount="1" stateProvince="Misamis Oriental" typeStatus="holotype">
<emphasis id="B968EAEAE75A67670CAEC27E2B2DFEE1" bold="true" box="[199,358,294,320]" pageId="32" pageNumber="333">
<typeStatus id="54A7885AE75A67670CAEC27E2B49FEE1" box="[199,258,294,320]" pageId="32" pageNumber="333">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F30B7668E75A67670D05C27E285FFEE1" box="[364,532,294,320]" name="Philippines" pageId="32" pageNumber="333">PHILIPPINES</collectingCountry>
,
<collectingCounty id="62C24E74E75A67670E76C27E2894FEE1" box="[543,735,294,320]" pageId="32" pageNumber="333">Mindanao Island</collectingCounty>
,
<collectingRegion id="49D8F81AE75A67670E83C27E29FAFEE1" box="[746,945,294,320]" country="Philippines" name="Misamis Oriental" pageId="32" pageNumber="333">Misamis Oriental</collectingRegion>
,
<collectingMunicipality id="6BC7AC82E75A67670FD2C27E2E1EFEE1" box="[955,1109,294,320]" pageId="32" pageNumber="333">Gingoog City</collectingMunicipality>
,
<location id="8EC36023E75A67670808C27F2F7BFEE0" LSID="urn:lsid:plazi:treatment:03B587EEE76767670CFEC08A281FFE28:8EC36023E75A67670808C27F2F7BFEE0" box="[1121,1328,295,321]" country="Philippines" county="Mindanao Island" municipality="Gingoog City" name="Barangay Lunotan" pageId="32" pageNumber="333" stateProvince="Misamis Oriental">Barangay Lunotan</location>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3066573E75A67670CAEC212281FFE28" pageId="32" pageNumber="333" type="distribution">
<paragraph id="8BA336F8E75A67670CAEC2122E39FEC5" blockId="32.[151,1437,150,393]" box="[199,1138,330,356]" pageId="32" pageNumber="333">
<emphasis id="B968EAEAE75A67670CAEC2122B2AFEC5" bold="true" box="[199,353,330,356]" pageId="32" pageNumber="333">Distribution.</emphasis>
<collectingCountry id="F30B7668E75A67670D0EC2122847FEC5" box="[359,524,330,356]" name="Philippines" pageId="32" pageNumber="333">PHILIPPINES</collectingCountry>
(Mindanao Island: Gingoog City in
<collectingRegion id="49D8F81AE75A67670FCCC2122E20FEC5" box="[933,1131,330,356]" country="Philippines" name="Misamis Oriental" pageId="32" pageNumber="333">Misamis Oriental</collectingRegion>
)
</paragraph>
<paragraph id="8BA336F8E75A67670CAEC236281FFE28" blockId="32.[151,1437,150,393]" box="[199,596,366,393]" pageId="32" pageNumber="333">
<emphasis id="B968EAEAE75A67670CAEC2362B28FE29" bold="true" box="[199,355,366,392]" pageId="32" pageNumber="333">Calling song.</emphasis>
This species is mute.
</paragraph>
</subSubSection>
</treatment>
</document>
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