162 lines
17 KiB
XML
162 lines
17 KiB
XML
<document id="AC6774A533D029744C7BE8A79B1FE15A" ID-CLB-Dataset="36820" ID-DOI="10.11646/zootaxa.3829.1.1" ID-GBIF-Dataset="a4b69269-5a4c-4b5b-9f89-6b314d3ba284" ID-ISSN="1175-5326" ID-Zenodo-Dep="286556" ID-ZooBank="75210CDC-AC6A-4624-A6F1-1BC969BC7CAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460452709399" checkinUser="plazi" docAuthor="Pyron, Robert Alexander & Wallach, Van" docDate="2014" docId="03B587DDC102B175CFD7CC1DFB54FDBA" docLanguage="en" docName="zt03829p081.pdf" docOrigin="Zootaxa 3829 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Acutotyphlops Wallach 1995" docType="treatment" docVersion="7" lastPageNumber="58" masterDocId="FF8CFFA5C13AB14CCF40CA0EFFB9FFA2" masterDocTitle="Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence" masterLastPageNumber="81" masterPageNumber="1" pageNumber="57" updateTime="1698338434491" updateUser="ExternalLinkService">
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<mods:titleInfo id="C0561A204AA67CE0001F51B35E2A8B77">
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<mods:title id="2E391C6E9F4F10A5DDBCEB35C7B73855">Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence</mods:title>
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<mods:name id="72CDD61B3A9C1E603A51EAA62DED03B6" type="personal">
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<mods:namePart id="F6BB7B2EA1BD5A96684DD8982E7C7439">Pyron, Robert Alexander</mods:namePart>
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<mods:name id="E283931E04E6A618E104D071255F89D9" type="personal">
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<mods:namePart id="0DA1D8185C8845B8E35DBDF3B0D0DB38">Wallach, Van</mods:namePart>
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<mods:title id="A43008F354A6A8D5703D60733E8314C4">Zootaxa</mods:title>
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<mods:part id="4029CD2F76B5B0CF96EDBAB210DDF438">
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<mods:date id="E751E4D5EAF06D6CF6072CD64965AC07">2014</mods:date>
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<mods:detail id="100ADBFC1BFB9F38B90DEF82A9194CFE" type="volume">
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<mods:number id="CD16CED31387309D3797BE149EAE93F4">3829</mods:number>
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<mods:identifier id="764076BD17A6B2BEF95F99CE00ECC6DE" type="DOI">10.11646/zootaxa.3829.1.1</mods:identifier>
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<treatment id="03B587DDC102B175CFD7CC1DFB54FDBA" ID-DOI="http://doi.org/10.5281/zenodo.6128004" ID-GBIF-Taxon="127668698" ID-Zenodo-Dep="6128004" LSID="urn:lsid:plazi:treatment:03B587DDC102B175CFD7CC1DFB54FDBA" httpUri="http://treatment.plazi.org/id/03B587DDC102B175CFD7CC1DFB54FDBA" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
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<subSubSection id="C3066540C102B174CFD7CC1DFE42F98F" box="[151,507,1555,1581]" pageId="56" pageNumber="57" type="nomenclature">
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<paragraph id="8BA336CBC102B174CFD7CC1DFE42F98F" blockId="56.[151,507,1555,1581]" box="[151,507,1555,1581]" pageId="56" pageNumber="57">
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<heading id="D0EB81A7C102B174CFD7CC1DFE42F98F" bold="true" box="[151,507,1555,1581]" fontSize="11" level="1" pageId="56" pageNumber="57" reason="1">
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<taxonomicName id="4C1C4D48C102B174CFD7CC1DFE42F98F" authority="Wallach, 1995" authorityName="Wallach" authorityYear="1995" box="[151,507,1555,1581]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="56" pageNumber="57" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C102B174CFD7CC1DFE42F98F" bold="true" box="[151,507,1555,1581]" pageId="56" pageNumber="57">
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<emphasis id="B968EAD9C102B174CFD7CC1DFEFDF98F" bold="true" box="[151,324,1555,1581]" italics="true" pageId="56" pageNumber="57">Acutotyphlops</emphasis>
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<bibRefCitation id="EF8D4B3AC102B174CE0BCC1DFE42F98F" author="Wallach" box="[331,507,1555,1581]" pageId="56" pageNumber="57" refString="Wallach, V. (1995) A new genus for the Ramphotyphlops subocularis species group (Serpentes: Typhlopidae), with description of a new species. Asiatic Herpetological Research, 6, 132 - 150." type="journal article" year="1995">Wallach, 1995</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C102B174CFD7CC57FE9AF91B" pageId="56" pageNumber="57" type="materials_examined">
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<paragraph id="8BA336CBC102B174CFD7CC57FCBDF9D0" blockId="56.[151,1436,1625,2010]" box="[151,772,1625,1650]" pageId="56" pageNumber="57">
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<emphasis id="B968EAD9C102B174CFD7CC57FE8BF9D0" bold="true" box="[151,306,1625,1650]" pageId="56" pageNumber="57">
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<typeStatus id="54A78869C102B174CFD7CC57FF6BF9D0" box="[151,210,1625,1650]" pageId="56" pageNumber="57">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C1C4D48C102B174CE79CC57FCBDF9D0" authority="Wallach, 1995" authorityName="Wallach" authorityYear="1995" box="[313,772,1625,1650]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="kunuaensis">
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<emphasis id="B968EAD9C102B174CE79CC57FDE4F9D0" box="[313,605,1625,1650]" italics="true" pageId="56" pageNumber="57">Acutotyphlops kunuaensis</emphasis>
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<bibRefCitation id="EF8D4B3AC102B174CD23CC57FCBDF9D0" author="Wallach" box="[611,772,1625,1650]" pageId="56" pageNumber="57" refString="Wallach, V. (1995) A new genus for the Ramphotyphlops subocularis species group (Serpentes: Typhlopidae), with description of a new species. Asiatic Herpetological Research, 6, 132 - 150." type="journal article" year="1995">Wallach, 1995</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BA336CBC102B174CF87CC73FE9AF91B" blockId="56.[151,1436,1625,2010]" pageId="56" pageNumber="57">
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<emphasis id="B968EAD9C102B174CF87CC73FE37F934" bold="true" box="[199,398,1661,1686]" pageId="56" pageNumber="57">Species content.</emphasis>
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<emphasis id="B968EAD9C102B174CEDDCC73FA94F934" box="[413,1325,1661,1686]" italics="true" pageId="56" pageNumber="57">
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<taxonomicName id="4C1C4D48C102B174CEDDCC73FD7DF934" box="[413,708,1661,1686]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="banaorum">Acutotyphlops banaorum</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C102B174CD98CC70FC22F934" box="[728,923,1661,1686]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="infralabialis">Ac. infralabialis</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C102B174CCEECC73FBD1F934" box="[942,1128,1661,1686]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="kunuaensis">
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<emphasis id="B968EAD9C102B174CCEECC73FBD1F934" bold="true" box="[942,1128,1661,1686]" italics="true" pageId="56" pageNumber="57">Ac. kunuaensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C102B174CBC0CC70FA91F934" box="[1152,1320,1661,1686]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="solomonis">Ac. solomonis</taxonomicName>
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,
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</emphasis>
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and
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<emphasis id="B968EAD9C102B174CA36CC73FE9AF91B" italics="true" pageId="56" pageNumber="57">
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<taxonomicName id="4C1C4D48C102B174CA36CC73FEA4F918" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="subocularis">
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<emphasis id="B968EAD9C102B174CA36CC73FEA4F918" bold="true" italics="true" pageId="56" pageNumber="57">Ac. subocularis</emphasis>
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</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C102B175CF87CCCBFE6FFE2A" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="diagnosis">
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<paragraph id="8BA336CBC102B175CF87CCCBFDC7FEBF" blockId="56.[151,1436,1625,2010]" lastBlockId="57.[151,1437,151,536]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
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<emphasis id="B968EAD9C102B174CF87CCCBFEF8F97C" bold="true" box="[199,321,1733,1758]" pageId="56" pageNumber="57">Diagnosis.</emphasis>
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<taxonomicName id="4C1C4D48C102B174CE08CCCBFE51F97C" box="[328,488,1733,1758]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="56" pageNumber="57" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C102B174CE08CCCBFE51F97C" box="[328,488,1733,1758]" italics="true" pageId="56" pageNumber="57">Acutotyphlops</emphasis>
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</taxonomicName>
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can be distinguished from all other typhlopoids by its inverted V-shape lower jaw or frontorostral and paired prefrontals shields. Small- to large-sized (total length
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<quantity id="4CE49B2EC102B174CB54CCE4FB17F8A0" box="[1044,1198,1770,1794]" metricMagnitude="-1" metricUnit="m" metricValue="2.94" metricValueMax="4.87" metricValueMin="1.01" pageId="56" pageNumber="57" unit="mm" value="294.0" valueMax="487.0" valueMin="101.0">101–487 mm</quantity>
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), stout- to moderatebodied (length/width ratio 18–58) snakes with 26–36 scale rows (with reduction), 334–542 total middorsals, short to long tail (1.0–7.7% total length) with 12–30 subcaudals (length/width ratio 1.0–3.3), and apical spine small to thorn-like. Dorsal and lateral head profiles usually pointed (rarely tapered), very narrow rostral (0.10–0.26 head width), inferior nasal suture in contact with first or second supralabial, preocular or subocular in contact with second and third supralabials, eye small, without visible pupil, or reduced to faint eyespot, T-III
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<collectingCountry id="F30B765BC102B174CA79CD90FAD8F814" box="[1337,1377,1950,1974]" name="Cyprus" pageId="56" pageNumber="57">SIP</collectingCountry>
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, and postoculars 3–5. Lateral tongue papillae absent; left lung absent, tracheal, cardiac and right lungs multicameral (with 21–80 + 2–12 + 5–17 chambers, respectively); testes segmented; hemipenis protrusible (retracted organ with 3–9 coils), retrocloacal sacs present; and rectal caecum absent. Coloration usually bicolored with dark brown dorsum and pale yellow or gold venter (with sharp demarcation between them), one species golden orange with numerous, irregular black spots on dorsum.
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</paragraph>
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<paragraph id="8BA336CBC103B175CF87CB29FE6FFE2A" blockId="57.[151,1437,151,536]" pageId="57" pageNumber="58">
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<emphasis id="B968EAD9C103B175CF87CB29FE51FEE2" bold="true" box="[199,488,295,320]" pageId="57" pageNumber="58">Phylogenetic definition.</emphasis>
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Includes the MRCA of
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<taxonomicName id="4C1C4D48C103B175CC58CB29FBFAFEE2" box="[792,1091,295,320]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="kunuaensis">
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<emphasis id="B968EAD9C103B175CC58CB29FBFAFEE2" box="[792,1091,295,320]" italics="true" pageId="57" pageNumber="58">Acutotyphlops kunuaensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C1C4D48C103B175CBC6CB26FA85FEE2" box="[1158,1340,295,320]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="subocularis">
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<emphasis id="B968EAD9C103B175CBC6CB26FA85FEE2" box="[1158,1340,295,320]" italics="true" pageId="57" pageNumber="58">Ac. subocularis</emphasis>
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</taxonomicName>
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and all descendants thereof, and all species more closely related to
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<taxonomicName id="4C1C4D48C103B175CC78CB42FC58FEC7" box="[824,993,332,357]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="kunuaensis">
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<emphasis id="B968EAD9C103B175CC78CB42FC58FEC7" box="[824,993,332,357]" italics="true" pageId="57" pageNumber="58">Ac. kunuaensis</emphasis>
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</taxonomicName>
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than to the
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<typeStatus id="54A78869C103B175CB26CB43FB2FFEC7" box="[1126,1174,333,357]" pageId="57" pageNumber="58">type</typeStatus>
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species of the 15 other typhlopid genera listed here.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C103B175CF87CB9AFC57FE0F" box="[199,1006,404,429]" pageId="57" pageNumber="58" type="etymology">
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<paragraph id="8BA336CBC103B175CF87CB9AFC57FE0F" blockId="57.[151,1437,151,536]" box="[199,1006,404,429]" pageId="57" pageNumber="58">
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<emphasis id="B968EAD9C103B175CF87CB9AFEF7FE0F" bold="true" box="[199,334,404,429]" pageId="57" pageNumber="58">Etymology.</emphasis>
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From the Latin for pointed
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<emphasis id="B968EAD9C103B175CDC6CB9AFD58FE09" box="[646,737,404,427]" italics="true" pageId="57" pageNumber="58">(acutos)</emphasis>
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, referring to the rostral.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C103B175CF87CBB9FBBFFE57" pageId="57" pageNumber="58" type="distribution">
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<paragraph id="8BA336CBC103B175CF87CBB9FBBFFE57" blockId="57.[151,1437,151,536]" pageId="57" pageNumber="58">
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<emphasis id="B968EAD9C103B175CF87CBB9FED9FE72" bold="true" box="[199,352,439,464]" pageId="57" pageNumber="58">Distribution.</emphasis>
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Primarily eastern
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<collectingCountry id="F30B765BC103B175CD71CBB6FCB7FE72" box="[561,782,439,464]" name="Papua New Guinea" pageId="57" pageNumber="58">Papua New Guinea</collectingCountry>
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, the
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<collectingCountry id="F30B765BC103B175CC04CBB9FBB9FE72" box="[836,1024,439,464]" name="Solomon Islands" pageId="57" pageNumber="58">Solomon Islands</collectingCountry>
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and Louisiade Archipelago, but one species (
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<taxonomicName id="4C1C4D48C103B175CFB8CBD2FDABFE57" box="[248,530,476,501]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="banaorum">
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<emphasis id="B968EAD9C103B175CFB8CBD2FDABFE57" box="[248,530,476,501]" italics="true" pageId="57" pageNumber="58">Acutotyphlops banaorum</emphasis>
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</taxonomicName>
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) occurs in the
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<collectingCountry id="F30B765BC103B175CDF9CBD2FC83FE57" box="[697,826,476,501]" name="Philippines" pageId="57" pageNumber="58">Philippines</collectingCountry>
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, ~
|
||
<quantity id="4CE49B2EC103B175CC1BCBD2FC07FE57" box="[859,958,476,501]" metricMagnitude="6" metricUnit="m" metricValue="4.0" pageId="57" pageNumber="58" unit="km" value="4000.0">4000 km</quantity>
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away.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C103B175CF87CBF1FB54FDBA" box="[199,1261,511,536]" pageId="57" pageNumber="58" type="discussion">
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<paragraph id="8BA336CBC103B175CF87CBF1FB54FDBA" blockId="57.[151,1437,151,536]" box="[199,1261,511,536]" pageId="57" pageNumber="58">
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<emphasis id="B968EAD9C103B175CF87CBF1FE82FDBA" bold="true" box="[199,315,511,536]" pageId="57" pageNumber="58">Remarks.</emphasis>
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This group apparently contains additional undescribed species (
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<bibRefCitation id="EF8D4B3AC103B175CB49CBF1FB58FDBA" author="Hedges" box="[1033,1249,511,536]" pageId="57" pageNumber="58" refString="Hedges, S. B., Marion, A. B., Lipp, K. M., Marin, J. & Vidal, N. (2014) A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology, 49, 1 - 61." type="journal article" year="2014">
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Hedges
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<emphasis id="B968EAD9C103B175CB25C80FFB27FDBA" box="[1125,1182,511,536]" italics="true" pageId="57" pageNumber="58">et al.</emphasis>
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2014
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</bibRefCitation>
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||
).
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</paragraph>
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</subSubSection>
|
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</treatment>
|
||
</document> |