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<document ID-DOI="10.11646/zootaxa.4766.1.1" ID-GBIF-Dataset="ed4d1bf6-284a-4898-952a-58943d5a3026" ID-ISSN="1175-5334" ID-Zenodo-Dep="3763907" ID-ZooBank="urn:lsid:zoobank.org:pub:B1B5DA41-5035-4783-8D47-28857B6305AE" approvalRequired="56" approvalRequired_for_textStreams="56" checkinTime="1587670592452" checkinUser="carolina" docAuthor="Ketwetsuriya, Chatchalerm, Karapunar, Baran, Charoentitirat, Thasinee &amp; Nützel, Al- Exander" docDate="2020" docId="03B587AB4F15155DFF517A5BFD1BFCDA" docLanguage="en" docName="zootaxa.4766.1.1.pdf.imf" docOrigin="Zootaxa 4766 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@1ef1a562" docStyleName="zootaxa.v2_0.2013.journal_article" docTitle="Platyzona gradata Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel 2020, sp. nov." docType="treatment" docVersion="5" lastPageNumber="36" masterDocId="FF8CFFD34F37157EFFC6787EFFCBFF85" masterDocTitle="Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane" masterLastPageNumber="47" masterPageNumber="1" pageNumber="35" updateTime="1643439188523" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ketwetsuriya, Chatchalerm</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Karapunar, Baran</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Charoentitirat, Thasinee</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nützel, Al- Exander</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-04-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4766</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:end>47</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.4766.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">ed4d1bf6-284a-4898-952a-58943d5a3026</mods:identifier>
<mods:identifier type="ISSN">1175-5334</mods:identifier>
<mods:identifier type="Zenodo-Dep">3763907</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:B1B5DA41-5035-4783-8D47-28857B6305AE</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3803829" ID-GBIF-Taxon="163737176" ID-Zenodo-Dep="3803829" LSID="urn:lsid:plazi:treatment:03B587AB4F15155DFF517A5BFD1BFCDA" httpUri="http://treatment.plazi.org/id/03B587AB4F15155DFF517A5BFD1BFCDA" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<subSubSection box="[151,471,549,576]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph blockId="34.[151,471,549,611]" box="[151,471,549,576]" pageId="34" pageNumber="35">
<heading bold="true" box="[151,471,549,576]" fontSize="10" level="2" pageId="34" pageNumber="35" reason="2">
<emphasis bold="true" box="[151,471,549,576]" pageId="34" pageNumber="35">
<taxonomicName authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[151,370,549,576]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="gradata" status="sp. nov.">
<emphasis bold="true" box="[151,370,549,576]" italics="true" pageId="34" pageNumber="35">Platyzona gradata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[378,471,550,576]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,249,585,611]" pageId="34" pageNumber="35" type="description">
<paragraph blockId="34.[151,471,549,611]" box="[151,249,585,611]" pageId="34" pageNumber="35">
(
<figureCitation box="[160,241,585,611]" captionStart="FIGURE 23" captionStartId="34.[151,250,1804,1829]" captionTargetBox="[166,1421,1245,1779]" captionTargetId="figure@34.[166,1421,1245,1779]" captionTargetPageId="34" captionText="FIGURE 23. Platyzona gradata sp. nov., AF, ESKU-19-LP 11, holotype. Scale bars represent: 2 mm (AB, EF); 1 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.3763953" httpUri="https://zenodo.org/record/3763953/files/figure.png" pageId="34" pageNumber="35">Fig. 23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,842,657,684]" pageId="34" pageNumber="35" type="etymology">
<paragraph blockId="34.[151,1437,657,1224]" box="[151,842,657,684]" pageId="34" pageNumber="35">
<emphasis bold="true" box="[151,286,658,684]" pageId="34" pageNumber="35">Etymology.</emphasis>
From Latin gradata, because of the gradate spire.
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="materials_examined">
<paragraph blockId="34.[151,1437,657,1224]" box="[199,745,693,720]" pageId="34" pageNumber="35">
<materialsCitation ID-GBIF-Occurrence="2608527129" box="[199,745,693,720]" collectionCode="LP" pageId="34" pageNumber="35" specimenCode="ESKU-19-LP 11" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[199,315,694,720]" pageId="34" pageNumber="35">
<typeStatus box="[199,310,694,720]" pageId="34" pageNumber="35">Holotype</typeStatus>
.
</emphasis>
Only
<specimenCount box="[385,542,693,720]" pageId="34" pageNumber="35">one specimen</specimenCount>
:
<specimenCode box="[553,741,693,719]" pageId="34" pageNumber="35">ESKU-19-LP 11</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph blockId="34.[151,1437,657,1224]" box="[199,1154,728,756]" pageId="34" pageNumber="35">
Dimensions (mm): ESKU-19-
<collectionCode box="[534,568,729,755]" country="Russia" name="Laboratory of Palaeontology" pageId="34" pageNumber="35">LP</collectionCode>
11: height = 11.6; width = 8.3; apical angle =
<emphasis box="[1087,1099,730,755]" italics="true" pageId="34" pageNumber="35">c</emphasis>
. 72
<superScript attach="none" box="[1142,1147,728,742]" fontSize="6" pageId="34" pageNumber="35">º</superScript>
.
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="distribution">
<paragraph blockId="34.[151,1437,657,1224]" pageId="34" pageNumber="35">
<emphasis bold="true" box="[199,673,765,792]" pageId="34" pageNumber="35">Type locality and stratigraphical range.</emphasis>
Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about
<quantity box="[311,383,801,827]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="34" pageNumber="35" unit="km" value="13.0">13 km</quantity>
east of
<collectingRegion box="[474,674,801,827]" country="Thailand" name="Lop Buri" pageId="34" pageNumber="35">Lopburi Province</collectingRegion>
, Central
<collectingCountry box="[777,877,801,827]" name="Thailand" pageId="34" pageNumber="35">Thailand</collectingCountry>
(
<figureCitation box="[893,960,801,827]" captionStart="FIGURE" captionStartId="1.[151,168,1975,2001]" captionTargetBox="[243,1345,1164,1941]" captionTargetId="figure@1.[236,1350,1160,1948]" captionTargetPageId="1" captionText="FIGURE 1. Geological map of the study location at Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about 13 km east of Lopburi Province, Central Thailand." figureDoi="http://doi.org/10.5281/zenodo.3763909" httpUri="https://zenodo.org/record/3763909/files/figure.png" pageId="34" pageNumber="35">Fig. 1</figureCitation>
), Khao Khad Formation,
<collectingRegion box="[1255,1350,801,827]" country="Thailand" name="Sara Buri" pageId="34" pageNumber="35">Saraburi</collectingRegion>
Group, Middle Permian, Roadian.
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="description">
<paragraph blockId="34.[151,1437,657,1224]" pageId="34" pageNumber="35">
<emphasis bold="true" box="[199,345,873,900]" pageId="34" pageNumber="35">Description.</emphasis>
Moderately high-spired, turbiniform shell consisting of 6 whorls (apex missing) with a step-like whorl profile; early teleoconch whorls (first three whorls) rounded, convex, with an ornament of faint spiral threads (maybe due to preservation); upper whorl surface of later whorls developing wide, steeply inclined ramp; ramp slightly convex to straight ornamented by
<emphasis box="[618,630,982,1007]" italics="true" pageId="34" pageNumber="35">c</emphasis>
. 8 to 10 equally spaced spiral threads; selenizone wide and flat, approximately
<quantity box="[235,321,1017,1044]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="34" pageNumber="35" unit="mm" value="0.8">0.8 mm</quantity>
wide, covering almost half of whorl height of body-whorl, situated below ramp; adapical border of ramp at about mid-whorl, forming also angulated border of ramp; outer whorl surface of selenizone almost straight, parallel to shell axis, bordered by two spiral threads followed by strong spiral cords; selenizone with curved transverse lunulae; adapical spiral rib forming periphery; abapical spiral somewhat above lower suture; whorls embracing somewhat below abapical spiral rib; suture distinctly impressed; base flatly convex with numerous evenly spaced spiral threads, shallowly phaneromphalous; aperture not well-preserved, seemingly with straight columellar lip.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3763953" ID-Zenodo-Dep="3763953" httpUri="https://zenodo.org/record/3763953/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,250,1804,1829]" targetBox="[166,1421,1245,1779]" targetPageId="34">
<paragraph blockId="34.[151,1436,1804,1865]" pageId="34" pageNumber="35">
<emphasis bold="true" box="[151,291,1804,1829]" pageId="34" pageNumber="35">FIGURE 23.</emphasis>
<taxonomicName authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[300,490,1805,1829]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="gradata" status="sp. nov.">
<emphasis box="[300,490,1805,1829]" italics="true" pageId="34" pageNumber="35">Platyzona gradata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[499,583,1805,1828]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
, AF, ESKU-19-LP 11, holotype. Scale bars represent: 2 mm (AB, EF); 1 mm (CD).
</paragraph>
</caption>
<subSubSection lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="discussion">
<paragraph blockId="34.[151,1436,1911,2046]" pageId="34" pageNumber="35">
<emphasis bold="true" box="[199,315,1911,1937]" pageId="34" pageNumber="35">Remarks.</emphasis>
<taxonomicName box="[326,438,1911,1937]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="34" pageNumber="35" phylum="Mollusca" rank="genus">
<emphasis box="[326,438,1911,1937]" italics="true" pageId="34" pageNumber="35">Platyzona</emphasis>
</taxonomicName>
is reported from
<collectingCountry box="[648,748,1911,1937]" name="Thailand" pageId="34" pageNumber="35">Thailand</collectingCountry>
for the first time. Several species of this genus have been reported from other Permian assemblages from Asia e.g., from
<collectingCountry box="[875,990,1947,1973]" name="Cambodia" pageId="34" pageNumber="35">Cambodia</collectingCountry>
(
<bibRefCitation author="Mansuy" box="[1007,1178,1947,1973]" firstAuthor="Mansuy" pageId="34" pageNumber="35" pagination="1 - 133" refId="ref26022" refString="Mansuy, H. (1913 a) Faunes des calcaires a Productus de l'Indochine. Memoires du Service Geologique de l'Indochine, 2 (4), 1 - 133." type="journal article" year="1913" yearSuffix="a">
Mansuy 1913
<emphasis box="[1164,1178,1948,1973]" italics="true" pageId="34" pageNumber="35">a</emphasis>
</bibRefCitation>
;
<bibRefCitation author="Delpey" box="[1195,1343,1947,1973]" firstAuthor="Delpey" pageId="34" pageNumber="35" pagination="345 - 369" refId="ref23861" refString="Delpey, G. (1941) Les Gasteropodes permiens du Cambodge. Journal de Conchyologie, 84, 255 - 278, 345 - 369." type="journal article" year="1941">Delpey 1941</bibRefCitation>
),
<collectingRegion box="[1365,1432,1947,1973]" country="Malaysia" name="Perak" pageId="34" pageNumber="35">Perak</collectingRegion>
,
<collectingCountry box="[151,254,1983,2009]" name="Malaysia" pageId="34" pageNumber="35">Malaysia</collectingCountry>
(
<bibRefCitation author="Batten" box="[269,411,1983,2009]" firstAuthor="Batten" pageId="34" pageNumber="35" pagination="1 - 44" refId="ref23042" refString="Batten, R. L. (1972) Permian gastropods and chitons from Perak, Malaysia. Part 1. Chitons, bellerophontids, euomphalids and pleurotomarians. Bulletin of the American Museum of Natural History, 147, 1 - 44." type="journal article" year="1972">Batten 1972</bibRefCitation>
), South
<collectingCountry box="[504,571,1983,2009]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
(
<bibRefCitation author="Pan &amp; Erwin" box="[586,801,1983,2009]" firstAuthor="Pan" pageId="34" pageNumber="35" pagination="1 - 49" refId="ref27138" refString="Pan, H. - Z. &amp; Erwin, D. H. (2002) Gastropods from the Permian of Guangxi and Yunnan provinces, South China. Journal of Paleontology, 78 (Supplement 56), 1 - 49. https: // doi. org / 10.1666 / 0022 - 3360 (2002) 76 [1: GFTPOG] 2.0. CO; 2" type="journal article" year="2002">Pan &amp; Erwin 2002</bibRefCitation>
) and from the Akasaka Limestone,
<collectingCountry box="[1202,1266,1984,2010]" name="Japan" pageId="34" pageNumber="35">Japan</collectingCountry>
(Nützel &amp;
<bibRefCitation author="Nakazawa" firstAuthor="Nakazawa" pageId="34" pageNumber="35" pagination="103 - 169" refId="ref27024" refString="Nu ¨ tzel, A. &amp; Nakazawa, K. (2012) Permian (Capitanian) gastropods from the Akasaka Limestone (Gifu Prefecture, Japan). Journal of Systematic Palaeontology, 10, 103 - 169. https: // doi. org / 10.1080 / 14772019.2010.549659" type="journal article" year="2012">Nakazawa 2012</bibRefCitation>
). It has also been reported from the
<collectingCountry box="[705,759,2019,2045]" name="United States of America" pageId="34" pageNumber="35">USA</collectingCountry>
(
<bibRefCitation author="Batten" box="[774,916,2019,2045]" firstAuthor="Batten" pageId="34" pageNumber="35" pagination="1 - 64" refId="ref23162" refString="Batten, R. L. (1989) Permian Gastropoda of the Southwestern United States. 7. Pleurotomariacea: Eotomariidae, Lophospiriidae, Gosseletinidae. American Museum Novitates, 2958, 1 - 64." type="journal article" year="1989">Batten 1989</bibRefCitation>
).
</paragraph>
<paragraph blockId="35.[151,1437,153,863]" pageId="35" pageNumber="36">
The single specimen at hand resembles the
<typeStatus box="[705,801,153,179]" pageId="35" pageNumber="36">paratype</typeStatus>
of
<taxonomicName authority="(Pan &amp; Yu, 1993)" baseAuthorityName="Pan &amp; Yu" baseAuthorityYear="1993" box="[844,1328,153,179]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="dongluoensis">
<emphasis box="[844,1114,153,179]" italics="true" pageId="35" pageNumber="36">Platyzona dongluoensis</emphasis>
(
<bibRefCitation author="Pan &amp; Yu" box="[1131,1320,153,179]" firstAuthor="Pan" pageId="35" pageNumber="36" refId="ref27245" refString="Pan, Y. - T. &amp; Yu, W. Z. (1993) Permian Gastropoda of China. Ocean Press, Beijing, 68 pp." type="book" year="1993">Pan &amp; Yu, 1993</bibRefCitation>
)
</taxonomicName>
from the Upper Permian Changxing Formation of south
<collectingCountry box="[686,753,189,215]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
(
<bibRefCitation author="Pan &amp; Yu" box="[769,947,189,215]" firstAuthor="Pan" pageId="35" pageNumber="36" refId="ref27245" refString="Pan, Y. - T. &amp; Yu, W. Z. (1993) Permian Gastropoda of China. Ocean Press, Beijing, 68 pp." type="book" year="1993">Pan &amp; Yu 1993</bibRefCitation>
, pl. 8, fig. 3ac). However, the
<typeStatus box="[1307,1405,189,215]" pageId="35" pageNumber="36">holotype</typeStatus>
of this species differs considerably from the
<typeStatus box="[619,715,225,251]" pageId="35" pageNumber="36">paratype</typeStatus>
and from the present specimen in having a narrow selenizone at mid-whorl and in other characters (
<bibRefCitation author="Pan &amp; Yu" box="[544,720,261,287]" firstAuthor="Pan" pageId="35" pageNumber="36" refId="ref27245" refString="Pan, Y. - T. &amp; Yu, W. Z. (1993) Permian Gastropoda of China. Ocean Press, Beijing, 68 pp." type="book" year="1993">Pan &amp; Yu 1993</bibRefCitation>
, pl. 8, fig. 2ab). The
<typeStatus box="[966,1062,261,287]" pageId="35" pageNumber="36">paratype</typeStatus>
of
<taxonomicName baseAuthorityName="Pan &amp; Yu" baseAuthorityYear="1993" box="[1100,1275,261,287]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="dongluoensis">
<emphasis box="[1100,1275,261,287]" italics="true" pageId="35" pageNumber="36">P. dongluoensis</emphasis>
</taxonomicName>
represents the genus
<taxonomicName box="[223,335,297,323]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">
<emphasis box="[223,335,297,323]" italics="true" pageId="35" pageNumber="36">Platyzona</emphasis>
</taxonomicName>
but the
<typeStatus box="[425,523,297,323]" pageId="35" pageNumber="36">holotype</typeStatus>
and hence the species itself does not, the species reported by
<bibRefCitation author="Pan &amp; Erwin" box="[1210,1436,297,323]" firstAuthor="Pan" pageId="35" pageNumber="36" pagination="1 - 49" refId="ref27138" refString="Pan, H. - Z. &amp; Erwin, D. H. (2002) Gastropods from the Permian of Guangxi and Yunnan provinces, South China. Journal of Paleontology, 78 (Supplement 56), 1 - 49. https: // doi. org / 10.1666 / 0022 - 3360 (2002) 76 [1: GFTPOG] 2.0. CO; 2" type="journal article" year="2002">Pan &amp; Erwin (2002)</bibRefCitation>
as of
<taxonomicName baseAuthorityName="Pan &amp; Yu" baseAuthorityYear="1993" box="[213,392,333,359]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="dongluoensis">
<emphasis box="[213,230,334,359]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis box="[244,392,333,359]" italics="true" pageId="35" pageNumber="36">dongluoensis</emphasis>
</taxonomicName>
from the Late Permian of
<collectingCountry box="[692,759,333,359]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
resembles the misidentified
<typeStatus box="[1082,1189,333,359]" pageId="35" pageNumber="36">paratypes</typeStatus>
of this species and
<taxonomicName authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="gradata" status="sp. nov.">
<emphasis box="[1412,1429,334,359]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis box="[151,240,369,395]" italics="true" pageId="35" pageNumber="36">gradata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[247,334,369,395]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
and could be conspecific.
</paragraph>
<paragraph blockId="35.[151,1437,153,863]" pageId="35" pageNumber="36">
<taxonomicName authority="Batten, 1972" authorityName="Batten" authorityYear="1972" box="[199,548,405,431]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="nodohumerosa">
<emphasis box="[199,216,406,431]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis box="[231,396,405,431]" italics="true" pageId="35" pageNumber="36">nodohumerosa</emphasis>
<bibRefCitation author="Batten" box="[404,548,405,431]" firstAuthor="Batten" pageId="35" pageNumber="36" pagination="1 - 44" refId="ref23042" refString="Batten, R. L. (1972) Permian gastropods and chitons from Perak, Malaysia. Part 1. Chitons, bellerophontids, euomphalids and pleurotomarians. Bulletin of the American Museum of Natural History, 147, 1 - 44." type="journal article" year="1972">Batten, 1972</bibRefCitation>
</taxonomicName>
from the Permian of
<collectingCountry box="[792,895,405,431]" name="Malaysia" pageId="35" pageNumber="36">Malaysia</collectingCountry>
resembles
<taxonomicName authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[1022,1140,405,431]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="gradata" status="sp. nov.">
<emphasis box="[1022,1140,405,431]" italics="true" pageId="35" pageNumber="36">P. gradata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1147,1235,405,431]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
but has a nodular ornament and distinct spiral cords or lirae on the selenizone.
<taxonomicName authority="(Reed, 1927)" baseAuthorityName="Reed" baseAuthorityYear="1927" box="[851,1165,441,467]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="eulkaiensis">
<emphasis box="[851,868,442,467]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis box="[884,1008,441,467]" italics="true" pageId="35" pageNumber="36">eulkaiensis</emphasis>
(
<bibRefCitation author="Reed" box="[1025,1156,441,467]" firstAuthor="Reed" pageId="35" pageNumber="36" pagination="129 - 158" refId="ref27357" refString="Reed Cowper, F. R. (1927) Upper Carboniferous fossils from Argentina. Carnegie Institute of Washington, 381, 129 - 158." type="journal article" year="1927">Reed, 1927</bibRefCitation>
)
</taxonomicName>
as illustrated by
<bibRefCitation author="Batten" firstAuthor="Batten" pageId="35" pageNumber="36" pagination="1 - 44" refId="ref23042" refString="Batten, R. L. (1972) Permian gastropods and chitons from Perak, Malaysia. Part 1. Chitons, bellerophontids, euomphalids and pleurotomarians. Bulletin of the American Museum of Natural History, 147, 1 - 44." type="journal article" year="1972">Batten (1972)</bibRefCitation>
from the Permian of
<collectingCountry box="[467,570,477,503]" name="Malaysia" pageId="35" pageNumber="36">Malaysia</collectingCountry>
and by Nützel &amp;
<bibRefCitation author="Nakazawa" box="[770,969,477,503]" firstAuthor="Nakazawa" pageId="35" pageNumber="36" pagination="103 - 169" refId="ref27024" refString="Nu ¨ tzel, A. &amp; Nakazawa, K. (2012) Permian (Capitanian) gastropods from the Akasaka Limestone (Gifu Prefecture, Japan). Journal of Systematic Palaeontology, 10, 103 - 169. https: // doi. org / 10.1080 / 14772019.2010.549659" type="journal article" year="2012">Nakazawa (2012)</bibRefCitation>
form the Permian of
<collectingCountry box="[1209,1273,477,503]" name="Japan" pageId="35" pageNumber="36">Japan</collectingCountry>
is also similar but has a narrower selenizone that is lower on the whorls (just above the suture) and a weaker spiral ornament.
</paragraph>
<paragraph blockId="35.[151,1437,153,863]" pageId="35" pageNumber="36">
Three Late Permian species from South
<collectingCountry box="[646,713,549,575]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
have been assigned to
<taxonomicName box="[968,1080,549,575]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">
<emphasis box="[968,1080,549,575]" italics="true" pageId="35" pageNumber="36">Platyzona</emphasis>
</taxonomicName>
by
<bibRefCitation author="Pan &amp; Erwin" box="[1121,1351,549,575]" firstAuthor="Pan" pageId="35" pageNumber="36" pagination="1 - 49" refId="ref27138" refString="Pan, H. - Z. &amp; Erwin, D. H. (2002) Gastropods from the Permian of Guangxi and Yunnan provinces, South China. Journal of Paleontology, 78 (Supplement 56), 1 - 49. https: // doi. org / 10.1666 / 0022 - 3360 (2002) 76 [1: GFTPOG] 2.0. CO; 2" type="journal article" year="2002">Pan &amp; Erwin (2002)</bibRefCitation>
:
<taxonomicName authority="Pan &amp; Erwin, 2002" authorityName="Pan &amp; Erwin" authorityYear="2002" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="pulchella">
<emphasis box="[1361,1378,550,575]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis italics="true" pageId="35" pageNumber="36">pulchella</emphasis>
<bibRefCitation author="Pan &amp; Erwin" box="[226,439,585,611]" firstAuthor="Pan" pageId="35" pageNumber="36" pagination="1 - 49" refId="ref27138" refString="Pan, H. - Z. &amp; Erwin, D. H. (2002) Gastropods from the Permian of Guangxi and Yunnan provinces, South China. Journal of Paleontology, 78 (Supplement 56), 1 - 49. https: // doi. org / 10.1666 / 0022 - 3360 (2002) 76 [1: GFTPOG] 2.0. CO; 2" type="journal article" year="2002">Pan &amp; Erwin, 2002</bibRefCitation>
</taxonomicName>
lacks spiral cords and has more pronounced growth-lines.
<taxonomicName authority="Pan &amp; Erwin, 2002" authorityName="Pan &amp; Erwin" authorityYear="2002" box="[1090,1411,585,611]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="nitella">
<emphasis box="[1090,1107,586,611]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis box="[1120,1192,585,611]" italics="true" pageId="35" pageNumber="36">nitella</emphasis>
<bibRefCitation author="Pan &amp; Erwin" box="[1198,1411,585,611]" firstAuthor="Pan" pageId="35" pageNumber="36" pagination="1 - 49" refId="ref27138" refString="Pan, H. - Z. &amp; Erwin, D. H. (2002) Gastropods from the Permian of Guangxi and Yunnan provinces, South China. Journal of Paleontology, 78 (Supplement 56), 1 - 49. https: // doi. org / 10.1666 / 0022 - 3360 (2002) 76 [1: GFTPOG] 2.0. CO; 2" type="journal article" year="2002">Pan &amp; Erwin, 2002</bibRefCitation>
</taxonomicName>
is broader and has more pronounced lunulae.
<taxonomicName authority="Pan &amp; Erwin, 2002" authorityName="Pan &amp; Erwin" authorityYear="2002" box="[637,997,621,647]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="luculenta">
<emphasis box="[637,654,622,647]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis box="[668,772,621,647]" italics="true" pageId="35" pageNumber="36">luculenta</emphasis>
<bibRefCitation author="Pan &amp; Erwin" box="[780,997,621,647]" firstAuthor="Pan" pageId="35" pageNumber="36" pagination="1 - 49" refId="ref27138" refString="Pan, H. - Z. &amp; Erwin, D. H. (2002) Gastropods from the Permian of Guangxi and Yunnan provinces, South China. Journal of Paleontology, 78 (Supplement 56), 1 - 49. https: // doi. org / 10.1666 / 0022 - 3360 (2002) 76 [1: GFTPOG] 2.0. CO; 2" type="journal article" year="2002">Pan &amp; Erwin, 2002</bibRefCitation>
</taxonomicName>
is based on an early juvenile specimen (protoconch and 1.5 teleoconch whorls). In contrast to
<taxonomicName authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[769,887,657,683]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="gradate" status="sp. nov.">
<emphasis box="[769,786,658,683]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis box="[800,887,657,683]" italics="true" pageId="35" pageNumber="36">gradate</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[895,984,657,683]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
, it has strong spiral cords on the earliest teleoconch and the selenizone is higher on the whorls.
</paragraph>
<paragraph blockId="35.[151,1437,153,863]" pageId="35" pageNumber="36">
Four Permian species from SW
<collectingCountry box="[569,623,729,755]" name="United States of America" pageId="35" pageNumber="36">USA</collectingCountry>
have been assigned to
<taxonomicName box="[893,1005,729,755]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">
<emphasis box="[893,1005,729,755]" italics="true" pageId="35" pageNumber="36">Platyzona</emphasis>
</taxonomicName>
by
<bibRefCitation author="Batten" box="[1052,1213,729,755]" firstAuthor="Batten" pageId="35" pageNumber="36" pagination="1 - 64" refId="ref23162" refString="Batten, R. L. (1989) Permian Gastropoda of the Southwestern United States. 7. Pleurotomariacea: Eotomariidae, Lophospiriidae, Gosseletinidae. American Museum Novitates, 2958, 1 - 64." type="journal article" year="1989">Batten (1989)</bibRefCitation>
:
<taxonomicName authority="Batten, 1989" authorityName="Batten" authorityYear="1989" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="rotunda">
<emphasis box="[1226,1243,730,755]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis box="[1259,1347,729,755]" italics="true" pageId="35" pageNumber="36">rotunda</emphasis>
<bibRefCitation author="Batten" firstAuthor="Batten" pageId="35" pageNumber="36" pagination="1 - 64" refId="ref23162" refString="Batten, R. L. (1989) Permian Gastropoda of the Southwestern United States. 7. Pleurotomariacea: Eotomariidae, Lophospiriidae, Gosseletinidae. American Museum Novitates, 2958, 1 - 64." type="journal article" year="1989">Batten, 1989</bibRefCitation>
</taxonomicName>
is broader and has entirely rounded whorls.
<taxonomicName authority="Batten, 1989" authorityName="Batten" authorityYear="1989" box="[706,1003,765,791]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="cancellata">
<emphasis box="[706,723,766,791]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis box="[737,854,765,791]" italics="true" pageId="35" pageNumber="36">cancellata</emphasis>
<bibRefCitation author="Batten" box="[860,1003,765,791]" firstAuthor="Batten" pageId="35" pageNumber="36" pagination="1 - 64" refId="ref23162" refString="Batten, R. L. (1989) Permian Gastropoda of the Southwestern United States. 7. Pleurotomariacea: Eotomariidae, Lophospiriidae, Gosseletinidae. American Museum Novitates, 2958, 1 - 64." type="journal article" year="1989">Batten, 1989</bibRefCitation>
</taxonomicName>
has a much narrower selenizone and a cancellate ornament.
<taxonomicName authority="Batten, 1989" authorityName="Batten" authorityYear="1989" box="[388,651,801,827]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species">
<emphasis box="[388,405,802,827]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis box="[418,502,801,827]" italics="true" pageId="35" pageNumber="36">pagoda</emphasis>
<bibRefCitation author="Batten" box="[508,651,801,827]" firstAuthor="Batten" pageId="35" pageNumber="36" pagination="1 - 64" refId="ref23162" refString="Batten, R. L. (1989) Permian Gastropoda of the Southwestern United States. 7. Pleurotomariacea: Eotomariidae, Lophospiriidae, Gosseletinidae. American Museum Novitates, 2958, 1 - 64." type="journal article" year="1989">Batten, 1989</bibRefCitation>
</taxonomicName>
is more slender and has a concave selenizone and has entirely rounded whorls.
<taxonomicName authority="Batten" authorityName="Batten" authorityYear="1989" box="[241,477,837,863]" class="Gastropoda" family="Goniasmatidae" genus="Platyzona" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="anguispira">
<emphasis box="[241,258,838,863]" italics="true" pageId="35" pageNumber="36">P</emphasis>
.
<emphasis box="[273,394,838,863]" italics="true" pageId="35" pageNumber="36">anguispira</emphasis>
Batten
</taxonomicName>
, has uncoiled whorls.
</paragraph>
</subSubSection>
</treatment>
</document>