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<mods:title id="20F3E73BF21FCA711BEF2AE9D26067D2">Review of the Oriental genus Indocloeon Müller-Liebenau 1982 (Ephemeroptera: Baetidae) with descriptions of two new species</mods:title>
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<mods:namePart id="A71C525C3137A94D17559D8384E3DB9B">Kluge, Nikita J.</mods:namePart>
<mods:affiliation id="10185ECF00A9E678EA331DE14F06DCDF">Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia Website: http: // www. insecta. bio. spbu. ru. n. kluge @ spbu. ru; https: // orcid. org / 0000 - 0001 - 9741 - 7790</mods:affiliation>
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<mods:namePart id="023CAB3A466E02C8D40B6C55F45B0C74">Suttinun, Chanaporn</mods:namePart>
<mods:affiliation id="EC9369CC5F89E1C659BE161217BDFF2A">Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand. chana. suttinun @ gmail. com; https: // orcid. org / 0000 - 0002 - 0380 - 1554</mods:affiliation>
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<heading id="D0EB81D8F203FF91ECABF8E70F26F8F1" bold="true" box="[151,841,1838,1865]" fontSize="11" level="1" pageId="4" pageNumber="455" reason="1">
<taxonomicName id="4C1C4D37F203FF91ECABF8E70F26F8F1" ID-CoL="3PN48" authority="Muller-Liebenau 1982" authorityName="Muller-Liebenau" authorityYear="1982" box="[151,841,1838,1865]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="primum" subGenus="Indocloeon">
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<emphasis id="B968EAA6F203FF91ECABF8E70D4FF8F0" bold="true" box="[151,288,1838,1865]" italics="true" pageId="4" pageNumber="455">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F203FF91ED0DF8E70DD5F8F0" bold="true" box="[305,442,1838,1865]" italics="true" pageId="4" pageNumber="455">Indocloeon</emphasis>
)
<emphasis id="B968EAA6F203FF91EDF7F8E70E44F8F1" bold="true" box="[459,555,1838,1864]" italics="true" pageId="4" pageNumber="455">primum</emphasis>
<bibRefCitation id="EF8D4B45F203FF91EE0FF8E70F26F8F1" author="Muller-Liebenau, I." box="[563,841,1838,1864]" pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
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(
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,
<figureCitation id="13272A31F203FF91ED2CF8980D37F8D5" box="[272,344,1873,1900]" captionStart="FIGURES 1524" captionStartId="9.[151,264,1843,1868]" captionTargetBox="[151,1430,181,1817]" captionTargetId="figure@9.[151,1436,181,1817]" captionTargetPageId="9" captionText="FIGURES 1524. Larva of Indocloeon (Indocloeon) primum. 1520, posterior margins of abdominal terga I, II, IV, VII, IX and X (black triangle shows median line on tergum IX); 21, abdominal tergum VI; 22, abdominal sternum III; 23, cercus; 24, scales on abdominal sternum, enlarged." figureDoi="http://doi.org/10.5281/zenodo.3839337" httpUri="https://zenodo.org/record/3839337/files/figure.png" pageId="4" pageNumber="455">1524</figureCitation>
,
<figureCitation id="13272A31F203FF91ED58F8980DC2F8D5" box="[356,429,1873,1900]" captionStart="FIGURES 3539" captionStartId="11.[151,264,1880,1905]" captionTargetBox="[151,1436,489,1856]" captionTargetId="figure@11.[151,1436,489,1856]" captionTargetPageId="11" captionText="FIGURES 3539. Imagines of Indocloeon (Indocloeon) primum. 35, antennae of female imago; 36, abdominal terga and sterna of male imago (genitals removed); 37, male imago; 38, abdominal terga and sterna of female imago; 39, female imago." figureDoi="http://doi.org/10.5281/zenodo.3839341" httpUri="https://zenodo.org/record/3839341/files/figure.png" pageId="4" pageNumber="455">3539</figureCitation>
,
<figureCitation id="13272A31F203FF91ED84F89B0E6EF8D5" box="[440,513,1874,1900]" captionStart="FIGURES 4649" captionStartId="14.[151,264,1772,1797]" captionTargetBox="[151,1431,460,1743]" captionTargetId="figure@14.[151,1436,453,1747]" captionTargetPageId="14" captionText="FIGURES 4649. Male genitals of Indocloeon s. str. 46, imaginal genitals of Indocloeon (Indocloeon) secundum sp. n. (holotype) (1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 47, their subimaginal exuviae; 48, gonostylus of Indocloeon (Indocloeon) primum; 49, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.3839347" httpUri="https://zenodo.org/record/3839347/files/figure.png" pageId="4" pageNumber="455">4849</figureCitation>
,
<figureCitation id="13272A31F203FF91EE30F8980E3CF8D5" box="[524,595,1873,1900]" captionStart="FIGURES 5052" captionStartId="15.[151,264,1805,1830]" captionTargetBox="[151,1436,345,1780]" captionTargetId="figure@15.[151,1436,345,1780]" captionTargetPageId="15" captionText="FIGURES 5052. Egg of Indocloeon (Indocloeon) primum. 50, general view; 5152, enlarged areas marked by frames on Fig. 50." figureDoi="http://doi.org/10.5281/zenodo.3839349" httpUri="https://zenodo.org/record/3839349/files/figure.png" pageId="4" pageNumber="455">5052</figureCitation>
)
</paragraph>
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<treatmentCitation id="0ABD10A5F203FF91ECABF85F0E13F817" author="Muller-Liebenau, I." box="[151,636,1942,1967]" page="127" pageId="4" pageNumber="455" year="1982">
<taxonomicName id="4C1C4D37F203FF91ECABF85F0E13F817" ID-CoL="3PN48" authority="Muller-Liebenau 1982: 127" authorityName="Muller-Liebenau" authorityPageNumber="127" authorityYear="1982" box="[151,636,1942,1967]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="primum">
<emphasis id="B968EAA6F203FF91ECABF85F0D31F817" box="[151,350,1942,1966]" italics="true" pageId="4" pageNumber="455">Indocloeon primum</emphasis>
<bibRefCitation id="EF8D4B45F203FF91ED59F85F0E13F817" author="Muller-Liebenau, I." box="[357,636,1942,1967]" pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982: 127</bibRefCitation>
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(larva);
<treatmentCitation id="0ABD10A5F203FF91EEEFF85F0F11F816" author="Kluge, N. J." box="[723,894,1942,1967]" page="367" pageId="4" pageNumber="455" year="2012">
<bibRefCitation id="EF8D4B45F203FF91EEEFF85F0F11F816" author="Kluge, N. J." box="[723,894,1942,1967]" pageId="4" pageNumber="455" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012: 367</bibRefCitation>
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(larva, subimago,
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and
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imagines, egg).
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<emphasis id="B968EAA6F203FF91ECABF8150D16F84E" bold="true" box="[151,377,2012,2039]" pageId="4" pageNumber="455">Material examined</emphasis>
(
<collectionCode id="ED0DAE71F203FF91EDB4F8140DD3F84E" box="[392,444,2013,2039]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="455" type="Museum">ZIN</collectionCode>
).
<materialsCitation id="3B743CE9F203FF91EDF3F81409F7F84E" ID-GBIF-Occurrence="2622593302" box="[463,1432,2013,2039]" country="Sri Lanka" location="Badulla District" pageId="4" pageNumber="455" specimenCount="1" stateProvince="Uva">
<collectingCountry id="F30B7624F203FF91EDF3F8140E07F84E" box="[463,616,2013,2039]" name="Sri Lanka" pageId="4" pageNumber="455">SRI LANKA</collectingCountry>
:
<collectingRegion id="49D8F856F203FF91EE4EF8140F61F84E" box="[626,782,2013,2039]" country="Sri Lanka" name="Uva" pageId="4" pageNumber="455">Uva Province</collectingRegion>
,
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, Haputale, small stream in tea plantation
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,
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<paragraph id="8BA336B4F202FF90ECABFF500EA8FEDE" blockId="5.[151,1437,152,1295]" pageId="5" pageNumber="456">
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running from
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,
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,
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,
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</date>
, coll.
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&amp;
<collectorName id="26E95362F202FF90E822FF5008FAFF0A" box="[1054,1173,153,179]" pageId="5" pageNumber="456">L. Sheyko</collectorName>
: 19 L-S-I
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</materialsCitation>
, 12 L-S-I
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, 3 L-S
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, 3 L-S
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,
<specimenCount id="9D1AFD3DF202FF90ED5CFF740DB5FF6E" box="[352,474,189,215]" pageId="5" pageNumber="456" type="larva">128 larvae</specimenCount>
;
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<collectingRegion id="49D8F856F202FF90EDDBFF740EC8FF6E" box="[487,679,189,215]" country="Zambia" name="Central" pageId="5" pageNumber="456">Central Province</collectingRegion>
,
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,
<location id="8EC3606FF202FF90EF51FF740FA7FF6E" LSID="urn:lsid:plazi:treatment:03B587A2F203FF90ECABF8E70DB0FAB6:8EC3606FF202FF90EF51FF740FA7FF6E" box="[877,968,189,215]" municipality="Matale District" name="Sigiriya" pageId="5" pageNumber="456" stateProvince="Central">Sigiriya</location>
,
<date id="FFA21074F202FF90EFE9FF740801FF6E" box="[981,1134,189,215]" pageId="5" pageNumber="456" value="2011-01-28" valueMax="2011-01-30" valueMin="2011-01-28">
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: 1 S-I
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;
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<collectingRegion id="49D8F856F202FF90E8EBFF7409F7FF6E" box="[1239,1432,189,215]" country="Zambia" name="Central" pageId="5" pageNumber="456">Central Province</collectingRegion>
,
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,
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, very small stream close to guesthouse «Green House» near beginning of steps to
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(
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),
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<geoCoordinate id="EE285073F202FF90EE05FECC0EAEFEA6" box="[569,705,261,287]" degrees="80" direction="east" minutes="31" orientation="longitude" pageId="5" pageNumber="456" precision="15" seconds="13" value="80.52028">80°3113”E</geoCoordinate>
,
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<collectingDate id="EFE6E99CF202FF90EEF2FECC0F0AFEA6" box="[718,869,261,287]" pageId="5" pageNumber="456" value="2011-01-24" valueMax="2011-01-26" valueMin="2011-01-24">2426.I.2011</collectingDate>
</date>
, coll.
<collectorName id="26E95362F202FF90EF96FECC087EFEA6" box="[938,1041,261,287]" pageId="5" pageNumber="456">N. Kluge</collectorName>
&amp;
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: 3 L-S-I
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, 1 S-I
<specimenCount id="9D1AFD3DF202FF90E951FECC09EEFEA6" box="[1389,1409,261,287]" pageId="5" pageNumber="456" type="male"></specimenCount>
, 1 S-I
<specimenCount id="9D1AFD3DF202FF90ECFDFEE00CBEFEFA" box="[193,209,297,323]" pageId="5" pageNumber="456" type="female"></specimenCount>
,
<specimenCount id="9D1AFD3DF202FF90ECE2FEE00D53FEFA" box="[222,316,297,323]" pageId="5" pageNumber="456" type="larva">6 larvae</specimenCount>
;
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<collectingMunicipality id="6BC7ACCEF202FF90ED75FEE00DD3FEFA" box="[329,444,297,323]" pageId="5" pageNumber="456">Delhausie</collectingMunicipality>
,
<location id="8EC3606FF202FF90EDF4FEE00EC6FEFA" LSID="urn:lsid:plazi:treatment:03B587A2F203FF90ECABF8E70DB0FAB6:8EC3606FF202FF90EDF4FEE00EC6FEFA" box="[456,681,297,323]" latitude="6.826111" longLatPrecision="21" longitude="80.52473" municipality="Delhausie" name="river Seethagangula" pageId="5" pageNumber="456" stateProvince="Central">river Seethagangula</location>
close to guesthouse «Vegetable Garden»,
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<geoCoordinate id="EE285073F202FF90E933FEE009F7FEFA" box="[1295,1432,297,323]" degrees="80" direction="east" minutes="31" orientation="longitude" pageId="5" pageNumber="456" precision="15" seconds="29" value="80.52473">80°3129”E</geoCoordinate>
,
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<collectingDate id="EFE6E99CF202FF90ECABFE850D73FEDE" box="[151,284,332,359]" pageId="5" pageNumber="456" value="2020-02-04" valueMax="2020-02-05" valueMin="2020-02-04">45.II.2020</collectingDate>
</date>
, coll.
<collectorName id="26E95362F202FF90ED5CFE840DA9FEDE" box="[352,454,333,359]" pageId="5" pageNumber="456">N. Kluge</collectorName>
&amp;
<collectorName id="26E95362F202FF90EDD6FE840E0EFEDE" box="[490,609,333,359]" pageId="5" pageNumber="456">L. Sheyko</collectorName>
: 1 L-S
<specimenCount id="9D1AFD3DF202FF90EE88FE840EACFEDE" box="[692,707,333,359]" pageId="5" pageNumber="456" type="female"></specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA336B4F202FF90ECFBFEB90D8EFC3A" blockId="5.[151,1437,152,1295]" pageId="5" pageNumber="456">
<emphasis id="B968EAA6F202FF90ECFBFEB90E5AFE32" bold="true" box="[199,565,368,395]" pageId="5" pageNumber="456">
Descriptions and notes.
<emphasis id="B968EAA6F202FF90EDD6FEB80E41FE32" bold="true" box="[490,558,369,395]" italics="true" pageId="5" pageNumber="456">Larva</emphasis>
.
</emphasis>
Labium with glossae, paraglossae and 2nd segment of labial palp relatively long (
<figureCitation id="13272A31F202FF90ECEBFE5C0D74FE16" box="[215,283,405,431]" captionStart="FIGURES 15" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 15. Larvae of Indocloeon s. str. 13, Indocloeon (Indocloeon) secundum sp. n.: 12, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 45, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="5" pageNumber="456">Fig. 4</figureCitation>
). Stout setae on outer side of femur non-pectinate, apically blunt (as in
<figureCitation id="13272A31F202FF90E810FE5C0812FE16" box="[1068,1149,405,431]" captionStart="FIGURES 7681" captionStartId="23.[151,264,1752,1777]" captionTargetBox="[151,1436,181,1727]" captionTargetId="figure@23.[151,1436,181,1727]" captionTargetPageId="23" captionText="FIGURES 7681. Larvae of Indocloeon (Hindocloeon) continentale sp. n. 7677, tibiae of fore and hind legs; 7879, fore and hind legs; 80, claw; 81, outer margin of femur (7679, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839363" httpUri="https://zenodo.org/record/3839363/files/figure.png" pageId="5" pageNumber="456">Fig. 81</figureCitation>
). Apex of femur normally with 2 (occasionally up to 8) stout, blunt setae. Tibiae of fore, middle and hind legs normally with one stout outerapical seta (occasionally with 2 or more such setae), without other stout setae on outer margin (as in
<figureCitation id="13272A31F202FF90E928FE1409FBFE4E" box="[1300,1428,477,503]" captionStart="FIGURES 1014" captionStartId="8.[151,264,1264,1289]" captionTargetBox="[151,1436,181,1239]" captionTargetId="figure@8.[151,1436,181,1239]" captionTargetPageId="8" captionText="FIGURES 1014. Larva of Indocloeon (Indocloeon) secundum sp. n. (holotype). 1011, claw; 12, two outer-apical setae on fore tibia; 13, tibia of fore leg; 14, tibia of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3839335" httpUri="https://zenodo.org/record/3839335/files/figure.png" pageId="5" pageNumber="456">Figs 1314</figureCitation>
) [see
<taxonomicName id="4C1C4D37F202FF90ECF7FDC80D26FDA2" authorityName="Muller-Liebenau" authorityYear="1982" box="[203,329,513,539]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F202FF90ECF7FDC80D26FDA2" box="[203,329,513,539]" italics="true" pageId="5" pageNumber="456">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF202FF90ED73FDC80DE6FDA2" box="[335,393,513,539]" pageId="5" pageNumber="456" sensu="stricto">s. str.</taxonomicNameLabel>
(8)]. Claw with 2 rows of denticles; in each row 27 (usually 34) most distal denticles larger than others. Hind margin of tergum I either lacks denticles or with few small denticles (
<figureCitation id="13272A31F202FF90E865FDEC08C3FD86" box="[1113,1196,548,575]" captionStart="FIGURES 1524" captionStartId="9.[151,264,1843,1868]" captionTargetBox="[151,1430,181,1817]" captionTargetId="figure@9.[151,1436,181,1817]" captionTargetPageId="9" captionText="FIGURES 1524. Larva of Indocloeon (Indocloeon) primum. 1520, posterior margins of abdominal terga I, II, IV, VII, IX and X (black triangle shows median line on tergum IX); 21, abdominal tergum VI; 22, abdominal sternum III; 23, cercus; 24, scales on abdominal sternum, enlarged." figureDoi="http://doi.org/10.5281/zenodo.3839337" httpUri="https://zenodo.org/record/3839337/files/figure.png" pageId="5" pageNumber="456">Fig. 15</figureCitation>
); hind margin of each tergum IIVIII bears regular row of long denticles fused at most part of their length (
<figureCitation id="13272A31F202FF90E875FD8008A4FDDD" box="[1097,1227,585,612]" captionStart="FIGURES 1524" captionStartId="9.[151,264,1843,1868]" captionTargetBox="[151,1430,181,1817]" captionTargetId="figure@9.[151,1436,181,1817]" captionTargetPageId="9" captionText="FIGURES 1524. Larva of Indocloeon (Indocloeon) primum. 1520, posterior margins of abdominal terga I, II, IV, VII, IX and X (black triangle shows median line on tergum IX); 21, abdominal tergum VI; 22, abdominal sternum III; 23, cercus; 24, scales on abdominal sternum, enlarged." figureDoi="http://doi.org/10.5281/zenodo.3839337" httpUri="https://zenodo.org/record/3839337/files/figure.png" pageId="5" pageNumber="456">Figs 1618</figureCitation>
); denticles on posterior margin of tergum IX also fused proximally; on middle of posterior margin of tergum IX (behind submedian pair of setae) denticles shorter (
<figureCitation id="13272A31F202FF90EE3CFD580E3AFD12" box="[512,597,657,683]" captionStart="FIGURES 1524" captionStartId="9.[151,264,1843,1868]" captionTargetBox="[151,1430,181,1817]" captionTargetId="figure@9.[151,1436,181,1817]" captionTargetPageId="9" captionText="FIGURES 1524. Larva of Indocloeon (Indocloeon) primum. 1520, posterior margins of abdominal terga I, II, IV, VII, IX and X (black triangle shows median line on tergum IX); 21, abdominal tergum VI; 22, abdominal sternum III; 23, cercus; 24, scales on abdominal sternum, enlarged." figureDoi="http://doi.org/10.5281/zenodo.3839337" httpUri="https://zenodo.org/record/3839337/files/figure.png" pageId="5" pageNumber="456">Fig. 19</figureCitation>
); denticles on posterior margin of tergum X partly fused proximally (
<figureCitation id="13272A31F202FF90E953FD580CD8FD76" captionStart="FIGURES 1524" captionStartId="9.[151,264,1843,1868]" captionTargetBox="[151,1430,181,1817]" captionTargetId="figure@9.[151,1436,181,1817]" captionTargetPageId="9" captionText="FIGURES 1524. Larva of Indocloeon (Indocloeon) primum. 1520, posterior margins of abdominal terga I, II, IV, VII, IX and X (black triangle shows median line on tergum IX); 21, abdominal tergum VI; 22, abdominal sternum III; 23, cercus; 24, scales on abdominal sternum, enlarged." figureDoi="http://doi.org/10.5281/zenodo.3839337" httpUri="https://zenodo.org/record/3839337/files/figure.png" pageId="5" pageNumber="456">Fig. 20</figureCitation>
). Hind margins of sterna III lack denticles; each sternum IIIIX with fused denticles (
<figureCitation id="13272A31F202FF90E8B3FD7C088BFD76" box="[1167,1252,693,719]" captionStart="FIGURES 1524" captionStartId="9.[151,264,1843,1868]" captionTargetBox="[151,1430,181,1817]" captionTargetId="figure@9.[151,1436,181,1817]" captionTargetPageId="9" captionText="FIGURES 1524. Larva of Indocloeon (Indocloeon) primum. 1520, posterior margins of abdominal terga I, II, IV, VII, IX and X (black triangle shows median line on tergum IX); 21, abdominal tergum VI; 22, abdominal sternum III; 23, cercus; 24, scales on abdominal sternum, enlarged." figureDoi="http://doi.org/10.5281/zenodo.3839337" httpUri="https://zenodo.org/record/3839337/files/figure.png" pageId="5" pageNumber="456">Fig. 22</figureCitation>
); on sternum IX of male fused denticles located only by sides of protogonostyli and between protogonostyli (
<bibRefCitation id="EF8D4B45F202FF90E891FD100959FD4A" author="Kluge, N. J." box="[1197,1334,729,755]" pageId="5" pageNumber="456" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
: fig. 35). Scales on abdominal terga and sterna blunt, forming irregular transverse rows (
<figureCitation id="13272A31F202FF90E822FD340816FCAE" box="[1054,1145,765,791]" captionStart="FIGURES 1524" captionStartId="9.[151,264,1843,1868]" captionTargetBox="[151,1430,181,1817]" captionTargetId="figure@9.[151,1436,181,1817]" captionTargetPageId="9" captionText="FIGURES 1524. Larva of Indocloeon (Indocloeon) primum. 1520, posterior margins of abdominal terga I, II, IV, VII, IX and X (black triangle shows median line on tergum IX); 21, abdominal tergum VI; 22, abdominal sternum III; 23, cercus; 24, scales on abdominal sternum, enlarged." figureDoi="http://doi.org/10.5281/zenodo.3839337" httpUri="https://zenodo.org/record/3839337/files/figure.png" pageId="5" pageNumber="456">Figs 24</figureCitation>
). Each segment of cercus with regular row of equal short denticles, without elongated denticles (
<figureCitation id="13272A31F202FF90EF89FCE80865FC82" box="[949,1034,801,827]" captionStart="FIGURES 1524" captionStartId="9.[151,264,1843,1868]" captionTargetBox="[151,1430,181,1817]" captionTargetId="figure@9.[151,1436,181,1817]" captionTargetPageId="9" captionText="FIGURES 1524. Larva of Indocloeon (Indocloeon) primum. 1520, posterior margins of abdominal terga I, II, IV, VII, IX and X (black triangle shows median line on tergum IX); 21, abdominal tergum VI; 22, abdominal sternum III; 23, cercus; 24, scales on abdominal sternum, enlarged." figureDoi="http://doi.org/10.5281/zenodo.3839337" httpUri="https://zenodo.org/record/3839337/files/figure.png" pageId="5" pageNumber="456">Fig. 23</figureCitation>
). For other characters see diagnosis of
<taxonomicName id="4C1C4D37F202FF90EC8AFC8C0D5BFCE6" authorityName="Muller-Liebenau" authorityYear="1982" box="[182,308,837,863]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F202FF90EC8AFC8C0D5BFCE6" box="[182,308,837,863]" italics="true" pageId="5" pageNumber="456">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF202FF90ED07FC8C0D18FCE6" box="[315,375,837,863]" pageId="5" pageNumber="456" sensu="stricto">s. str.</taxonomicNameLabel>
above, diagnosis of
<taxonomicName id="4C1C4D37F202FF90EE63FC8C0EB2FCE6" authorityName="Muller-Liebenau" authorityYear="1982" box="[607,733,837,863]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B968EAA6F202FF90EE63FC8C0EB2FCE6" box="[607,733,837,863]" italics="true" pageId="5" pageNumber="456">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF202FF90EED8FC8C0F62FCE6" box="[740,781,837,863]" pageId="5" pageNumber="456" sensu="lato">s. l.</taxonomicNameLabel>
(
<bibRefCitation id="EF8D4B45F202FF90EF27FC8C0FCEFCE6" author="Kluge, N. J." box="[795,929,837,863]" pageId="5" pageNumber="456" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
and above) and former descriptions (Müller- Liebenau 1982;
<bibRefCitation id="EF8D4B45F202FF90ED71FCA00DB9FC3A" author="Kluge, N. J." box="[333,470,873,899]" pageId="5" pageNumber="456" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8BA336B4F202FF90ECFBFC450917FBC6" blockId="5.[151,1437,152,1295]" pageId="5" pageNumber="456">
<emphasis id="B968EAA6F202FF90ECFBFC450DAEFC1E" bold="true" box="[199,449,908,935]" italics="true" pageId="5" pageNumber="456">Subimago and imago</emphasis>
. Described by
<bibRefCitation id="EF8D4B45F202FF90EE4CFC440F63FC1E" author="Kluge, N. J." box="[624,780,909,935]" pageId="5" pageNumber="456" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge (2012)</bibRefCitation>
. Statement that «pedicellus with basal half brown» is not quite correct, because only lateral and median sides of its basal half are brown (
<figureCitation id="13272A31F202FF90E82DFC78080DFC72" box="[1041,1122,944,971]" captionStart="FIGURES 3539" captionStartId="11.[151,264,1880,1905]" captionTargetBox="[151,1436,489,1856]" captionTargetId="figure@11.[151,1436,489,1856]" captionTargetPageId="11" captionText="FIGURES 3539. Imagines of Indocloeon (Indocloeon) primum. 35, antennae of female imago; 36, abdominal terga and sterna of male imago (genitals removed); 37, male imago; 38, abdominal terga and sterna of female imago; 39, female imago." figureDoi="http://doi.org/10.5281/zenodo.3839341" httpUri="https://zenodo.org/record/3839341/files/figure.png" pageId="5" pageNumber="456">Fig. 35</figureCitation>
) [see
<taxonomicName id="4C1C4D37F202FF90E899FC78094CFC72" authorityName="Muller-Liebenau" authorityYear="1982" box="[1189,1315,945,971]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F202FF90E899FC78094CFC72" box="[1189,1315,945,971]" italics="true" pageId="5" pageNumber="456">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF202FF90E916FC78090AFC72" box="[1322,1381,945,971]" pageId="5" pageNumber="456" sensu="stricto">s. str.</taxonomicNameLabel>
(9)]. In subimago of both sexes and in male imago cross veins colourless (
<figureCitation id="13272A31F202FF90EF8AFC1C0864FC56" box="[950,1035,981,1007]" captionStart="FIGURES 3539" captionStartId="11.[151,264,1880,1905]" captionTargetBox="[151,1436,489,1856]" captionTargetId="figure@11.[151,1436,489,1856]" captionTargetPageId="11" captionText="FIGURES 3539. Imagines of Indocloeon (Indocloeon) primum. 35, antennae of female imago; 36, abdominal terga and sterna of male imago (genitals removed); 37, male imago; 38, abdominal terga and sterna of female imago; 39, female imago." figureDoi="http://doi.org/10.5281/zenodo.3839341" httpUri="https://zenodo.org/record/3839341/files/figure.png" pageId="5" pageNumber="456">Fig. 37</figureCitation>
), only in female imago cross veins darkened (
<figureCitation id="13272A31F202FF90ED32FC300D0DFBAA" box="[270,354,1017,1043]" captionStart="FIGURES 3539" captionStartId="11.[151,264,1880,1905]" captionTargetBox="[151,1436,489,1856]" captionTargetId="figure@11.[151,1436,489,1856]" captionTargetPageId="11" captionText="FIGURES 3539. Imagines of Indocloeon (Indocloeon) primum. 35, antennae of female imago; 36, abdominal terga and sterna of male imago (genitals removed); 37, male imago; 38, abdominal terga and sterna of female imago; 39, female imago." figureDoi="http://doi.org/10.5281/zenodo.3839341" httpUri="https://zenodo.org/record/3839341/files/figure.png" pageId="5" pageNumber="456">Fig. 39</figureCitation>
). Abdominal terga IIIVI of both sexes always without median longitudinal stripe (
<figureCitation id="13272A31F202FF90E930FC3009FFFBAA" box="[1292,1424,1017,1043]" captionStart="FIGURES 3539" captionStartId="11.[151,264,1880,1905]" captionTargetBox="[151,1436,489,1856]" captionTargetId="figure@11.[151,1436,489,1856]" captionTargetPageId="11" captionText="FIGURES 3539. Imagines of Indocloeon (Indocloeon) primum. 35, antennae of female imago; 36, abdominal terga and sterna of male imago (genitals removed); 37, male imago; 38, abdominal terga and sterna of female imago; 39, female imago." figureDoi="http://doi.org/10.5281/zenodo.3839341" httpUri="https://zenodo.org/record/3839341/files/figure.png" pageId="5" pageNumber="456">Figs 36, 38</figureCitation>
). Proximal (1st+2nd) segment of gonostylus moderately widened at base; proximal 2/3 of its inner margin straight, at 2/3 length from base inner margin forms convex blunt angle (
<figureCitation id="13272A31F202FF90EF6CFB880FC9FBE2" box="[848,934,1089,1115]" captionStart="FIGURES 4649" captionStartId="14.[151,264,1772,1797]" captionTargetBox="[151,1431,460,1743]" captionTargetId="figure@14.[151,1436,453,1747]" captionTargetPageId="14" captionText="FIGURES 4649. Male genitals of Indocloeon s. str. 46, imaginal genitals of Indocloeon (Indocloeon) secundum sp. n. (holotype) (1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 47, their subimaginal exuviae; 48, gonostylus of Indocloeon (Indocloeon) primum; 49, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.3839347" httpUri="https://zenodo.org/record/3839347/files/figure.png" pageId="5" pageNumber="456">Fig. 48</figureCitation>
). For other characters see diagnosis of
<taxonomicName id="4C1C4D37F202FF90E95CFB880C8DFBC6" authorityName="Muller-Liebenau" authorityYear="1982" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F202FF90E95CFB880C8DFBC6" italics="true" pageId="5" pageNumber="456">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF202FF90ECD5FBAC0D4BFBC6" box="[233,292,1125,1151]" pageId="5" pageNumber="456" sensu="stricto">s. str.</taxonomicNameLabel>
above, diagnosis of
<taxonomicName id="4C1C4D37F202FF90EE36FBAC0EE7FBC6" authorityName="Muller-Liebenau" authorityYear="1982" box="[522,648,1125,1151]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B968EAA6F202FF90EE36FBAC0EE7FBC6" box="[522,648,1125,1151]" italics="true" pageId="5" pageNumber="456">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF202FF90EEB3FBAC0ED8FBC6" box="[655,695,1125,1151]" pageId="5" pageNumber="456" sensu="lato">s. l.</taxonomicNameLabel>
(
<bibRefCitation id="EF8D4B45F202FF90EEF9FBAC0F24FBC6" author="Kluge, N. J." box="[709,843,1125,1151]" pageId="5" pageNumber="456" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
and above) and former description (
<bibRefCitation id="EF8D4B45F202FF90E8DEFBAC0903FBC6" author="Kluge, N. J." box="[1250,1388,1125,1151]" pageId="5" pageNumber="456" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8BA336B4F202FF90ECFBFB400E75FB52" blockId="5.[151,1437,152,1295]" pageId="5" pageNumber="456">
<emphasis id="B968EAA6F202FF90ECFBFB400C92FB1A" bold="true" box="[199,253,1161,1187]" pageId="5" pageNumber="456">
<emphasis id="B968EAA6F202FF90ECFBFB400C99FB1A" bold="true" box="[199,246,1161,1187]" italics="true" pageId="5" pageNumber="456">Egg</emphasis>
.
</emphasis>
Oval, about
<quantity id="4CE49B51F202FF90EDB7FB410D81FB1A" box="[395,494,1160,1187]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="456" unit="mm" value="0.15">0.15 mm</quantity>
length. Polar area bearing net-like relief [see
<taxonomicName id="4C1C4D37F202FF90EFDAFB40080BFB1A" authorityName="Muller-Liebenau" authorityYear="1982" box="[998,1124,1161,1187]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F202FF90EFDAFB40080BFB1A" box="[998,1124,1161,1187]" italics="true" pageId="5" pageNumber="456">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF202FF90E856FB4008CBFB1A" box="[1130,1188,1161,1187]" pageId="5" pageNumber="456" sensu="stricto">s. str.</taxonomicNameLabel>
(15)] with cells small, lacking convexities on bottoms (
<figureCitation id="13272A31F202FF90EE35FB640E30FB7E" box="[521,607,1196,1223]" captionStart="FIGURES 5052" captionStartId="15.[151,264,1805,1830]" captionTargetBox="[151,1436,345,1780]" captionTargetId="figure@15.[151,1436,345,1780]" captionTargetPageId="15" captionText="FIGURES 5052. Egg of Indocloeon (Indocloeon) primum. 50, general view; 5152, enlarged areas marked by frames on Fig. 50." figureDoi="http://doi.org/10.5281/zenodo.3839349" httpUri="https://zenodo.org/record/3839349/files/figure.png" pageId="5" pageNumber="456">Fig. 52</figureCitation>
); remainder area of egg surface partly with sharply outlined cavities (see ibid.), partly smooth (
<figureCitation id="13272A31F202FF90EDB7FB180E61FB52" box="[395,526,1232,1259]" captionStart="FIGURES 5052" captionStartId="15.[151,264,1805,1830]" captionTargetBox="[151,1436,345,1780]" captionTargetId="figure@15.[151,1436,345,1780]" captionTargetPageId="15" captionText="FIGURES 5052. Egg of Indocloeon (Indocloeon) primum. 50, general view; 5152, enlarged areas marked by frames on Fig. 50." figureDoi="http://doi.org/10.5281/zenodo.3839349" httpUri="https://zenodo.org/record/3839349/files/figure.png" pageId="5" pageNumber="456">Figs 5051</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C306653FF202FF90ECFBFB3D0DB0FAB6" box="[199,479,1268,1295]" pageId="5" pageNumber="456" type="distribution">
<paragraph id="8BA336B4F202FF90ECFBFB3D0DB0FAB6" blockId="5.[151,1437,152,1295]" box="[199,479,1268,1295]" pageId="5" pageNumber="456">
<emphasis id="B968EAA6F202FF90ECFBFB3D0D0CFAB6" bold="true" box="[199,355,1268,1295]" pageId="5" pageNumber="456">Distribution:</emphasis>
<collectingCountry id="F30B7624F202FF90ED56FB3C0DB4FAB6" box="[362,475,1269,1295]" name="Sri Lanka" pageId="5" pageNumber="456">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>