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<mods:title id="3238F15250FD90CE4F6207046FCFCA37">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:namePart id="E1D732A54FD71E3FFD8F96CBB85739DD">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="CC20562A80F7C2C2C1E255119E1CD24C">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
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<mods:namePart id="189D79E6EA6C9B2AFBE1F4DB30133767">Coca-Abia, María Milagro</mods:namePart>
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<paragraph id="8BA3D655A61A0605FD95F931FC24E282" blockId="47.[622,966,1688,1750]" box="[622,966,1688,1715]" pageId="47" pageNumber="48">
<heading id="D0EB6139A61A0605FD95F931FC24E282" box="[622,966,1688,1715]" centered="true" fontSize="11" level="2" pageId="47" pageNumber="48" reason="2">
<taxonomicName id="4C1CADD6A61A0605FD95F931FC24E282" authority="(Frey, 1971)" baseAuthorityName="Frey" baseAuthorityYear="1971" box="[622,966,1688,1715]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A61A0605FD95F931FCCFE282" bold="true" box="[622,813,1688,1715]" italics="true" pageId="47" pageNumber="48">Miridiba herteli</emphasis>
(
<bibRefCitation id="EF8DABA4A61A0605FCC6F930FC5FE282" author="Frey G." box="[829,957,1689,1715]" pageId="47" pageNumber="48" pagination="206 - 225" refId="ref43999" refString="Frey G. 1971. Bestimmungstabelle der Indischen und Ceylonesischen arten der Gattung Holotrichia Hope. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 22: 206 - 225." type="journal article" year="1971">Frey, 1971</bibRefCitation>
)
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</heading>
</paragraph>
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<subSubSection id="C30685DEA61A0605FD23F915FCB9E2E7" box="[728,859,1724,1750]" pageId="47" pageNumber="48" type="description">
<paragraph id="8BA3D655A61A0605FD23F915FCB9E2E7" blockId="47.[622,966,1688,1750]" box="[728,859,1724,1750]" pageId="47" pageNumber="48">
<figureCitation id="1327CAD0A61A0605FD23F915FCB9E2E7" box="[728,859,1724,1750]" captionStart="Figs 65–68. 65" captionStartId="48.[189,241,1885,1911]" captionTargetBox="[281,1310,526,1835]" captionTargetId="figure-76@48.[264,1323,509,1848]" captionTargetPageId="48" captionText="Figs 65–68. 65. Miridiba gressitti (Frey, 1970), paratype, ♀ (MHNB). Habitus in dorsal view. 66– 68. M. herteli (Frey, 1971) holotype, ♂ (SNSD 10629). 66. Habitus in dorsal view. 67. Parameres in dorsal (A), ventral (B) and lateral (C) views, 68. Endophallus and temones in lateral view. Scale bars: 65–66 = 5.0 mm; 67–68 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770380" httpUri="https://zenodo.org/record/4770380/files/figure.png" pageId="47" pageNumber="48">Figs 66–68</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA61A0605FF46F954FE94E391" pageId="47" pageNumber="48" type="reference_group">
<paragraph id="8BA3D655A61A0605FF46F954FAF0E329" blockId="47.[189,1298,1789,1816]" box="[189,1298,1789,1816]" pageId="47" pageNumber="48">
<treatmentCitationGroup id="AB0CF17BA61A0605FF46F954FAF0E329" box="[189,1298,1789,1816]" pageId="47" pageNumber="48">
<treatmentCitation id="0ABDF044A61A0605FF46F954FD75E326" author="Frey G." box="[189,663,1789,1816]" page="221" pageId="47" pageNumber="48" year="1971">
<taxonomicName id="4C1CADD6A61A0605FF46F954FD75E326" authority="Frey, 1971: 220 - 221" authorityName="Frey" authorityPageNumber="220 - 221" authorityYear="1971" box="[189,663,1789,1816]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A61A0605FF46F954FE78E326" box="[189,410,1789,1815]" italics="true" pageId="47" pageNumber="48">Holotrichia herteli</emphasis>
<bibRefCitation id="EF8DABA4A61A0605FE5AF957FD75E326" author="Frey G." box="[417,663,1789,1816]" pageId="47" pageNumber="48" pagination="206 - 225" refId="ref43999" refString="Frey G. 1971. Bestimmungstabelle der Indischen und Ceylonesischen arten der Gattung Holotrichia Hope. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 22: 206 - 225." type="journal article" year="1971">Frey, 1971: 220–221</bibRefCitation>
</taxonomicName>
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, fig. 29b (
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loc.: Saharanpur, 
<collectingRegion id="49D818B7A61A0605FBEEF957FB5BE326" box="[1045,1209,1789,1816]" country="India" name="Uttar Pradesh" pageId="47" pageNumber="48">Uttar Pradesh</collectingRegion>
, 
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).
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<treatmentCitationGroup id="AB0CF17BA61A0605FF46F896FC64E36B" box="[189,902,1854,1882]" pageId="47" pageNumber="48">
<taxonomicName id="4C1CADD6A61A0605FF46F896FE78E368" authorityName="Frey" authorityYear="1971" box="[189,410,1855,1881]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A61A0605FF46F896FE78E368" box="[189,410,1855,1881]" italics="true" pageId="47" pageNumber="48">Holotrichia herteli</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0ABDF044A61A0605FE4CF896FD11E368" author="Smetana A. &amp; Kral D." box="[439,755,1854,1881]" page="220" pageId="47" pageNumber="48" year="2006">
<bibRefCitation id="EF8DABA4A61A0605FE4CF896FD11E368" author="Smetana A. &amp; Kral D." box="[439,755,1854,1881]" pageId="47" pageNumber="48" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 220</bibRefCitation>
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(catalogue).
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<paragraph id="8BA3D655A61A0605FF46F8CBFE94E391" blockId="47.[189,1398,1854,1952]" pageId="47" pageNumber="48">
<treatmentCitationGroup id="AB0CF17BA61A0605FF46F8CBFE94E391" pageId="47" pageNumber="48">
<taxonomicName id="4C1CADD6A61A0605FF46F8CBFE9DE34D" baseAuthorityName="Frey" baseAuthorityYear="1971" box="[189,383,1890,1916]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A61A0605FF46F8CBFE9DE34D" box="[189,383,1890,1916]" italics="true" pageId="47" pageNumber="48">Miridiba herteli</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0ABDF044A61A0605FE5DF8CBFD83E34D" author="Keith D." box="[422,609,1890,1916]" page="81" pageId="47" pageNumber="48" year="2004">
<bibRefCitation id="EF8DABA4A61A0605FE5DF8CBFD83E34D" author="Keith D." box="[422,609,1890,1916]" pageId="47" pageNumber="48" pagination="81 - 83" refId="ref44533" refString="Keith D. 2004. Un nouveau Melolonthidae du nord-est de la Thailande (Coleoptera: Scarabaeoidea). Bulletin de l'Institut royal des Sciences naturelles de Belgique. Entomologie 74: 81 - 83." type="journal article" year="2004">Keith 2004: 81</bibRefCitation>
</treatmentCitation>
(combination). — 
<treatmentCitation id="0ABDF044A61A0605FCAAF8CBFBBFE34D" author="Coca-Abia M. M." box="[849,1117,1890,1916]" page="680" pageId="47" pageNumber="48" year="2008">
<bibRefCitation id="EF8DABA4A61A0605FCAAF8CBFBBFE34D" author="Coca-Abia M. M." box="[849,1117,1890,1916]" pageId="47" pageNumber="48" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008: 680</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0ABDF044A61A0605FB6DF8CBFA94E34D" author="Bezdek A." box="[1174,1398,1890,1916]" page="271" pageId="47" pageNumber="48" year="2016">
<bibRefCitation id="EF8DABA4A61A0605FB6DF8CBFA94E34D" author="Bezdek A." box="[1174,1398,1890,1916]" pageId="47" pageNumber="48" pagination="249 - 280" refId="ref43171" refString="Bezdek A. 2016. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 249 - 280. Koninklijke Brill NV, Leiden, The Netherlands." type="book chapter" year="2016">Bezděk 2016: 271</bibRefCitation>
</treatmentCitation>
(catalogue).
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</paragraph>
</subSubSection>
<subSubSection id="C30685DEA605061BFF46FEA1FED4E70E" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="8BA3D655A605061AFF46FEA1FED4E513" blockId="48.[189,310,264,290]" box="[189,310,264,290]" pageId="48" pageNumber="49">
<heading id="D0EB6139A605061AFF46FEA1FED4E513" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B9680A47A605061AFF46FEA1FED4E513" bold="true" box="[189,310,264,290]" pageId="48" pageNumber="49">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A605061BFF46FE91FED4E70E" blockId="48.[189,1400,312,446]" lastBlockId="49.[188,1400,265,831]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
External morphology of adult (
<figureCitation id="1327CAD0A605061AFDC2FE91FD77E562" box="[569,661,312,339]" captionStart="Figs 65–68. 65" captionStartId="48.[189,241,1885,1911]" captionTargetBox="[281,1310,526,1835]" captionTargetId="figure-76@48.[264,1323,509,1848]" captionTargetPageId="48" captionText="Figs 65–68. 65. Miridiba gressitti (Frey, 1970), paratype, ♀ (MHNB). Habitus in dorsal view. 66– 68. M. herteli (Frey, 1971) holotype, ♂ (SNSD 10629). 66. Habitus in dorsal view. 67. Parameres in dorsal (A), ventral (B) and lateral (C) views, 68. Endophallus and temones in lateral view. Scale bars: 65–66 = 5.0 mm; 67–68 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770380" httpUri="https://zenodo.org/record/4770380/files/figure.png" pageId="48" pageNumber="49">Fig. 66</figureCitation>
). Body size 
<quantity id="4CE47BB0A605061AFCD4FE91FC7DE563" box="[815,927,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="48" pageNumber="49" unit="mm" value="17.2">17.2 mm</quantity>
. Dorsal surface glabrous, at most with tiny setae in each puncture hardly visible; matt appearance. Clypeus shorter than frons, flat, oblique sides, anterior edge strongly emarginate at middle. Frons with carina moderately developed. Pronotal anterior margin glabrous, regularly curved; posterior margin glabrous, finely flanged with a row of punctures except at middle; lateral margins with posterior half sinuate, moderately serrated and with short setae; anterior angles obtuse, not projected forward, posterior angles almost right and marked. Prosternal process cone-shaped. Scutellum glabrous, with scattered punctures. Epipleuron with few short setae at basal part. Foretibia with dorsal carina sharp; insertion of inner spur almost equidistant between second and third outer tooth. Meso- and metatibia with transverse carina complete, inner margin of dorsal surface with few strong spines. Prepygidium with scattered punctured regularly distributed. Pygidium regularly flanged, surface with scattered punctures irregularly distributed, tiny setae in each puncture hardly visible, apical margin pubescent. Ventrites 3 and 4 glabrous, with inconspicuous punctures at middle, more conspicuous at sides. Ventrite 5 moderately depressed at posterior half, with pubescence of different length, with conspicuous scattered punctures. Ventrite 6 with a transversal anterior keel, with long pubescence near posterior margin. Male genitalia matches morphotype II “ 
<emphasis id="B9680A47A604061BFE85FD3CFE06E69E" box="[382,484,661,687]" italics="true" pageId="49" pageNumber="50">Gressitti</emphasis>
” above described. Dorsal complex of parameres with intermediate branches longer than ventral ones, bending toward distal end; apices of ventral branches with a ventral sharp tip (
<figureCitation id="1327CAD0A604061BFF3EFD74FEFCE6C9" box="[197,286,733,760]" captionStart="Figs 65–68. 65" captionStartId="48.[189,241,1885,1911]" captionTargetBox="[281,1310,526,1835]" captionTargetId="figure-76@48.[264,1323,509,1848]" captionTargetPageId="48" captionText="Figs 65–68. 65. Miridiba gressitti (Frey, 1970), paratype, ♀ (MHNB). Habitus in dorsal view. 66– 68. M. herteli (Frey, 1971) holotype, ♂ (SNSD 10629). 66. Habitus in dorsal view. 67. Parameres in dorsal (A), ventral (B) and lateral (C) views, 68. Endophallus and temones in lateral view. Scale bars: 65–66 = 5.0 mm; 67–68 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770380" httpUri="https://zenodo.org/record/4770380/files/figure.png" pageId="49" pageNumber="50">Fig. 67</figureCitation>
). Endophallus with a soft raspula with scattered small spines (
<figureCitation id="1327CAD0A604061BFBFBFD74FBBBE6C9" box="[1024,1113,733,760]" captionStart="Figs 65–68. 65" captionStartId="48.[189,241,1885,1911]" captionTargetBox="[281,1310,526,1835]" captionTargetId="figure-76@48.[264,1323,509,1848]" captionTargetPageId="48" captionText="Figs 65–68. 65. Miridiba gressitti (Frey, 1970), paratype, ♀ (MHNB). Habitus in dorsal view. 66– 68. M. herteli (Frey, 1971) holotype, ♂ (SNSD 10629). 66. Habitus in dorsal view. 67. Parameres in dorsal (A), ventral (B) and lateral (C) views, 68. Endophallus and temones in lateral view. Scale bars: 65–66 = 5.0 mm; 67–68 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770380" httpUri="https://zenodo.org/record/4770380/files/figure.png" pageId="49" pageNumber="50">Fig. 68</figureCitation>
). Apophysis of temones with each distal end prolonged by an oval plate each weakly sclerotized (
<figureCitation id="1327CAD0A604061BFBC0FCA8FB75E72D" box="[1083,1175,769,796]" captionStart="Figs 65–68. 65" captionStartId="48.[189,241,1885,1911]" captionTargetBox="[281,1310,526,1835]" captionTargetId="figure-76@48.[264,1323,509,1848]" captionTargetPageId="48" captionText="Figs 65–68. 65. Miridiba gressitti (Frey, 1970), paratype, ♀ (MHNB). Habitus in dorsal view. 66– 68. M. herteli (Frey, 1971) holotype, ♂ (SNSD 10629). 66. Habitus in dorsal view. 67. Parameres in dorsal (A), ventral (B) and lateral (C) views, 68. Endophallus and temones in lateral view. Scale bars: 65–66 = 5.0 mm; 67–68 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770380" httpUri="https://zenodo.org/record/4770380/files/figure.png" pageId="49" pageNumber="50">Fig. 68</figureCitation>
). Female genitalia unknown.
</paragraph>
</subSubSection>
<caption id="DF6386DDA605061AFF46F8F4FDB8E3D3" ID-DOI="http://doi.org/10.5281/zenodo.4770380" ID-Zenodo-Dep="4770380" httpUri="https://zenodo.org/record/4770380/files/figure.png" pageId="48" pageNumber="49" startId="48.[189,241,1885,1911]" targetBox="[281,1310,526,1835]" targetPageId="48">
<paragraph id="8BA3D655A605061AFF46F8F4FDB8E3D3" blockId="48.[189,1399,1885,2018]" pageId="48" pageNumber="49">
<emphasis id="B9680A47A605061AFF46F8F4FE94E346" bold="true" box="[189,374,1885,1911]" pageId="48" pageNumber="49">Figs 65–68. 65</emphasis>
. 
<taxonomicName id="4C1CADD6A605061AFE72F8F4FD1FE349" authority="(Frey, 1970)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[393,765,1885,1912]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="gressitti">
<emphasis id="B9680A47A605061AFE72F8F4FDB8E346" box="[393,602,1885,1911]" italics="true" pageId="48" pageNumber="49">Miridiba gressitti</emphasis>
(
<bibRefCitation id="EF8DABA4A605061AFD96F8F7FD17E349" author="Frey G." box="[621,757,1886,1912]" pageId="48" pageNumber="49" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey, 1970</bibRefCitation>
)
</taxonomicName>
, paratype, ♀ (MHNB). Habitus in dorsal view. 
<emphasis id="B9680A47A605061AFAB1F8F4FF38E3AB" bold="true" pageId="48" pageNumber="49">66– 68</emphasis>
. 
<taxonomicName id="4C1CADD6A605061AFF10F828FE1CE3AA" authority="(Frey, 1971)" baseAuthorityName="Frey" baseAuthorityYear="1971" box="[235,510,1921,1947]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A605061AFF10F828FE80E3AA" box="[235,354,1921,1947]" italics="true" pageId="48" pageNumber="49">M. herteli</emphasis>
(
<bibRefCitation id="EF8DABA4A605061AFE88F828FE17E3AA" author="Frey G." box="[371,501,1921,1947]" pageId="48" pageNumber="49" pagination="206 - 225" refId="ref43999" refString="Frey G. 1971. Bestimmungstabelle der Indischen und Ceylonesischen arten der Gattung Holotrichia Hope. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 22: 206 - 225." type="journal article" year="1971">Frey, 1971</bibRefCitation>
)
</taxonomicName>
holotype, ♂ (SNSD 10629). 
<emphasis id="B9680A47A605061AFC9AF829FC9CE3AB" bold="true" box="[865,894,1920,1946]" pageId="48" pageNumber="49">66</emphasis>
. Habitus in dorsal view. 
<emphasis id="B9680A47A605061AFB4BF829FB29E3AB" bold="true" box="[1200,1227,1920,1946]" pageId="48" pageNumber="49">67</emphasis>
. Parameres in dorsal (A), ventral (B) and lateral (C) views, 
<emphasis id="B9680A47A605061AFD22F80DFD17E38F" bold="true" box="[729,757,1956,1982]" pageId="48" pageNumber="49">68</emphasis>
. Endophallus and temones in lateral view. Scale bars: 65–66 = 5.0 mm; 67–68 = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C30685DEA604061BFF46FCCEFE3DE0DE" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="8BA3D655A604061BFF46FCCEFE48E7B0" blockId="49.[189,426,871,897]" box="[189,426,871,897]" pageId="49" pageNumber="50">
<heading id="D0EB6139A604061BFF46FCCEFE48E7B0" bold="true" box="[189,426,871,897]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B9680A47A604061BFF46FCCEFE48E7B0" bold="true" box="[189,426,871,897]" pageId="49" pageNumber="50">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B74DC08A604061BFF11FC3EFE8BE070" ID-GBIF-Occurrence="3114133301" country="India" determinerName="G. Frey" location="INDIA" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="8BA3D655A604061BFF11FC3EFA95E02D" blockId="49.[189,1399,919,1089]" pageId="49" pageNumber="50">
<emphasis id="B9680A47A604061BFF11FC3EFEBEE780" bold="true" box="[234,348,919,945]" pageId="49" pageNumber="50">
<typeStatus id="54A768F7A604061BFF11FC3EFEBEE780" box="[234,348,919,945]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F30B96C5A604061BFF46FC12FEECE7E4" box="[189,270,955,981]" name="India" pageId="49" pageNumber="50">INDIA</collectingCountry>
• 
<specimenCount id="9D1A1DDCA604061BFEDEFC12FEDCE7E4" box="[293,318,955,981]" pageId="49" pageNumber="50" type="male">♂</specimenCount>
; “Saharanpur, N.W. 
<collectingCountry id="F30B96C5A604061BFDD0FC13FD9EE7E5" box="[555,636,954,980]" name="India" pageId="49" pageNumber="50">Indien</collectingCountry>
”; “10629”; “Staatl. Museum für Tierkunde, Dresden” [Staatliches Museum für Tierkunde Dresden]; “ 
<taxonomicName id="4C1CADD6A604061BFDA0FC77FD06E7C9" authority="Gao &amp; Coca-Abia, 2021" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[603,740,990,1016]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B9680A47A604061BFDA0FC77FD06E7C9" box="[603,740,990,1016]" italics="true" pageId="49" pageNumber="50">Holotrichia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BB73CA604061BFD10FC76FCC8E7C8" box="[747,810,991,1017]" pageId="49" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C1CADD6A604061BFCC9FC76FC6DE7C9" baseAuthorityName="Hope" baseAuthorityYear="1842" box="[818,911,991,1016]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B9680A47A604061BFCC9FC76FC6DE7C9" box="[818,911,991,1016]" italics="true" pageId="49" pageNumber="50">sinensis</emphasis>
</taxonomicName>
?”; “ 
<typeStatus id="54A768F7A604061BFC32FC76FBE4E7C8" box="[969,1030,991,1017]" pageId="49" pageNumber="50">Type</typeStatus>
<taxonomicName id="4C1CADD6A604061BFBF5FC77FB14E7C9" authority="Gao &amp; Coca-Abia, 2021" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[1038,1270,990,1016]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="herterli" status="sp. nov.">
<emphasis id="B9680A47A604061BFBF5FC77FB14E7C9" box="[1038,1270,990,1016]" italics="true" pageId="49" pageNumber="50">Holotrichia herterli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BB73CA604061BFB05FC76FADEE7C8" box="[1278,1340,991,1017]" pageId="49" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
det. 
<determinerName id="7E7F3A91A604061BFF46FBAAFEFBE02C" box="[189,281,1027,1053]" pageId="49" pageNumber="50">
G. 
<bibRefCitation id="EF8DABA4A604061BFF18FBAAFEFBE02C" author="Frey G." box="[227,281,1027,1053]" pageId="49" pageNumber="50" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey</bibRefCitation>
</determinerName>
1970”; “TYPE”; “ 
<typeStatus id="54A768F7A604061BFE01FBABFD88E02D" box="[506,618,1026,1052]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C1CADD6A604061BFD8DFBABFAA0E02D" authority="(Frey, 1970) CH. Bu Gao &amp; M. Coca-Abia" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[630,1346,1026,1053]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A604061BFD8DFBABFCD7E02D" box="[630,821,1026,1052]" italics="true" pageId="49" pageNumber="50">Miridiba herteli</emphasis>
(
<bibRefCitation id="EF8DABA4A604061BFCBFFBAAFC24E02C" author="Frey G." box="[836,966,1027,1053]" pageId="49" pageNumber="50" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey, 1970</bibRefCitation>
) 
<collectingCountry id="F30B96C5A604061BFC23FBAAFBE6E02C" box="[984,1028,1027,1053]" name="Switzerland" pageId="49" pageNumber="50">CH</collectingCountry>
. Bu Gao &amp; M. Coca-Abia
</taxonomicName>
det.
</paragraph>
<paragraph id="8BA3D655A604061BFF44FB8EFE8BE070" blockId="49.[189,1399,919,1089]" box="[191,361,1063,1089]" pageId="49" pageNumber="50">
2017”; 
<collectionCode id="ED0D4E90A604061BFEEDFB8EFE87E070" box="[278,357,1063,1089]" collectionName="SNSD" pageId="49" pageNumber="50">SNSD</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8BA3D655A604061BFF11FBC0FE3DE0DE" blockId="49.[189,1399,1129,1263]" pageId="49" pageNumber="50">
<materialsCitation id="3B74DC08A604061BFF11FBC0FE3DE0DE" ID-GBIF-Occurrence="3114133303" collectionCode="NHMB" country="India" determinerName="Coca- Abia &amp; Chuan-bu Gao" location="Sylhet region in India" municipality="Sylheti Ind Prov. Nagpur" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B9680A47A604061BFF11FBC0FEBEE0B2" bold="true" box="[234,348,1129,1155]" pageId="49" pageNumber="50">
<typeStatus id="54A768F7A604061BFF11FBC0FEBEE0B2" box="[234,348,1129,1155]" pageId="49" pageNumber="50" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="F30B96C5A604061BFF46FB24FEECE096" box="[189,270,1165,1191]" name="India" pageId="49" pageNumber="50">INDIA</collectingCountry>
• 
<specimenCount id="9D1A1DDCA604061BFEC8FB24FE8AE096" box="[307,360,1165,1191]" pageId="49" pageNumber="50" type="male">1 ♂</specimenCount>
; “ 
<collectingMunicipality id="6BC74C2FA604061BFE7FFB24FD23E096" box="[388,705,1165,1191]" pageId="49" pageNumber="50">Sylheti Ind Prov. Nagpur</collectingMunicipality>
” [
<location id="8EC3808EA604061BFD25FB24FC17E096" LSID="urn:lsid:plazi:treatment:03B56743A61A0618FD95F931FD2DE2B4:8EC3808EA604061BFD25FB24FC17E096" box="[734,1013,1165,1191]" country="India" municipality="Sylheti Ind Prov. Nagpur" name="Sylhet region in India" pageId="49" pageNumber="50">Sylhet region in India</location>
, 
<location id="8EC3808EA604061BFBFDFB25FB87E096" LSID="urn:lsid:plazi:treatment:03B56743A61A0618FD95F931FD2DE2B4:8EC3808EA604061BFBFDFB25FB87E096" box="[1030,1125,1164,1191]" country="India" municipality="Sylheti Ind Prov. Nagpur" name="Nagpur" pageId="49" pageNumber="50">Nagpur</location>
]; “♀”; “PARATYPE”; “ 
<taxonomicName id="4C1CADD6A604061BFF31FB18FE45E0FA" authority="Gao &amp; Coca-Abia, 2021" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[202,423,1201,1227]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="herteli" status="sp. nov.">
<emphasis id="B9680A47A604061BFF31FB18FE45E0FA" box="[202,423,1201,1227]" italics="true" pageId="49" pageNumber="50">Holotrichia herteli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BB73CA604061BFE55FB18FE0FE0FA" box="[430,493,1201,1227]" pageId="49" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
det. G. 
<bibRefCitation id="EF8DABA4A604061BFDB0FB18FD2AE0FA" author="Frey G." box="[587,712,1201,1227]" pageId="49" pageNumber="50" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey 1970</bibRefCitation>
”; “ 
<taxonomicName id="4C1CADD6A604061BFD17FB18FBB3E0FA" authority="(Frey, 1970)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[748,1105,1201,1227]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="gressitti">
<emphasis id="B9680A47A604061BFD17FB18FC5BE0FA" box="[748,953,1201,1227]" italics="true" pageId="49" pageNumber="50">Miridiba gressitti</emphasis>
(
<bibRefCitation id="EF8DABA4A604061BFC33FB18FBABE0FA" author="Frey G." box="[968,1097,1201,1227]" pageId="49" pageNumber="50" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey, 1970</bibRefCitation>
)
</taxonomicName>
<determinerName id="7E7F3A91A604061BFBA3FB18FB3FE0FA" box="[1112,1245,1201,1227]" pageId="49" pageNumber="50">Coca- Abia</determinerName>
&amp; 
<determinerName id="7E7F3A91A604061BFAF8FB18FF0CE0DE" pageId="49" pageNumber="50">Chuan-bu Gao</determinerName>
det. 2018”; 
<collectionCode id="ED0D4E90A604061BFE84FB7CFE39E0DE" box="[383,475,1237,1263]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="49" pageNumber="50">NHMB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA604061BFF46FABEFED8E305" pageId="49" pageNumber="50" type="discussion">
<paragraph id="8BA3D655A604061BFF46FABEFECDE100" blockId="49.[189,303,1303,1329]" box="[189,303,1303,1329]" pageId="49" pageNumber="50">
<heading id="D0EB6139A604061BFF46FABEFECDE100" bold="true" box="[189,303,1303,1329]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B9680A47A604061BFF46FABEFECDE100" bold="true" box="[189,303,1303,1329]" pageId="49" pageNumber="50">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A604061BFF46FAEEFED8E305" blockId="49.[186,1401,1350,1844]" pageId="49" pageNumber="50">
<bibRefCitation id="EF8DABA4A604061BFF46FAEEFEAAE151" author="Frey G." box="[189,328,1350,1377]" pageId="49" pageNumber="50" pagination="206 - 225" refId="ref43999" refString="Frey G. 1971. Bestimmungstabelle der Indischen und Ceylonesischen arten der Gattung Holotrichia Hope. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 22: 206 - 225." type="journal article" year="1971">Frey (1971)</bibRefCitation>
described 
<taxonomicName id="4C1CADD6A604061BFE3AFAEFFD7DE151" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[449,671,1350,1376]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A604061BFE3AFAEFFD7DE151" box="[449,671,1350,1376]" italics="true" pageId="49" pageNumber="50">Holotrichia herteli</emphasis>
</taxonomicName>
from 
<specimenCount id="9D1A1DDCA604061BFD1DFAEEFC75E151" box="[742,919,1350,1377]" pageId="49" pageNumber="50" type="generic">two specimens</specimenCount>
, male and female, designated as 
<typeStatus id="54A768F7A604061BFAF5FAEFFA94E151" box="[1294,1398,1350,1376]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
(SNSD) and 
<typeStatus id="54A768F7A604061BFEAFFAC2FE59E1B4" box="[340,443,1387,1413]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
(NHMB), respectively. Later, 
<bibRefCitation id="EF8DABA4A604061BFCD2FAC3FC20E1B5" author="Keith D." box="[809,962,1386,1412]" pageId="49" pageNumber="50" pagination="81 - 83" refId="ref44533" refString="Keith D. 2004. Un nouveau Melolonthidae du nord-est de la Thailande (Coleoptera: Scarabaeoidea). Bulletin de l'Institut royal des Sciences naturelles de Belgique. Entomologie 74: 81 - 83." type="journal article" year="2004">Keith (2004)</bibRefCitation>
transferred this species to 
<taxonomicName id="4C1CADD6A604061BFAFFFAC3FA8CE1B5" baseAuthorityName="Nomura" baseAuthorityYear="1977" box="[1284,1390,1386,1412]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A604061BFAFFFAC3FA8CE1B5" box="[1284,1390,1386,1412]" italics="true" pageId="49" pageNumber="50">Miridiba</emphasis>
</taxonomicName>
. We have studied both type specimens and realized that the 
<typeStatus id="54A768F7A604061BFC8BFA26FC35E198" box="[880,983,1423,1449]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
is not female but male. In addition, after comparing both types, we have found that they differ by the following characters (
<tableCitation id="C69EE3EEA604061BFB3CFA1BFAC0E1FD" box="[1223,1314,1458,1484]" captionStart="Table 3" captionStartId="50.[189,258,265,291]" captionTargetPageId="50" captionText="Table 3. Differences between holotype and paratype of M. herteli (Frey, 1971) and paratype of M. gressitti (Frey, 1970)." httpUri="http://table.plazi.org/id/DF6386DDA6070618FF46FEA0FE03E577" pageId="49" pageNumber="50" tableUuid="DF6386DDA6070618FF46FEA0FE03E577">Table 3</tableCitation>
): body (size, appearance and pubescence), pronotum (anterior and lateral margins), foretibia (carina, inner spur), mesotibia (lateral carina), pygidium (apical margin), ventrites 5 and 6 (appearance) and male genitalia (parameres shape and internal structures of endophallus). These differences between the 
<typeStatus id="54A768F7A604061BFF46F9EBFEC4E26D" box="[189,294,1602,1628]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54A768F7A604061BFE9BF9EAFE25E26C" box="[352,455,1603,1629]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C1CADD6A604061BFE0BF9EAFD84E26D" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[496,614,1602,1628]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A604061BFE0BF9EAFD84E26D" box="[496,614,1602,1628]" italics="true" pageId="49" pageNumber="50">M. herteli</emphasis>
</taxonomicName>
raise questions about the identity of the 
<typeStatus id="54A768F7A604061BFBB5F9EAFB5BE26C" box="[1102,1209,1603,1629]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
. However, after studying and comparing the 
<typeStatus id="54A768F7A604061BFDECF9CEFD68E2B0" box="[535,650,1639,1665]" pageId="49" pageNumber="50" type="paratype">paratypes</typeStatus>
of 
<taxonomicName id="4C1CADD6A604061BFD4DF9CEFCCFE2B1" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[694,813,1638,1664]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A604061BFD4DF9CEFCCFE2B1" box="[694,813,1638,1664]" italics="true" pageId="49" pageNumber="50">M. herteli</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1CADD6A604061BFC91F9CEFC10E2B1" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[874,1010,1639,1664]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="gressitti">
<emphasis id="B9680A47A604061BFC91F9CEFC10E2B1" box="[874,1010,1639,1664]" italics="true" pageId="49" pageNumber="50">M. gressitti</emphasis>
</taxonomicName>
, they turned out to be identical. Consequently, the 
<typeStatus id="54A768F7A604061BFE67F922FDE1E294" box="[412,515,1675,1701]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C1CADD6A604061BFDD0F922FD42E295" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[555,672,1674,1700]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A604061BFDD0F922FD42E295" box="[555,672,1674,1700]" italics="true" pageId="49" pageNumber="50">M. herteli</emphasis>
</taxonomicName>
is identified as 
<taxonomicName id="4C1CADD6A604061BFCA5F922FC07E295" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[862,997,1675,1700]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="gressitti">
<emphasis id="B9680A47A604061BFCA5F922FC07E295" box="[862,997,1675,1700]" italics="true" pageId="49" pageNumber="50">M. gressitti</emphasis>
</taxonomicName>
; features of external morphology confirm it (
<tableCitation id="C69EE3EEA604061BFEBFF907FE41E2F9" box="[324,419,1710,1736]" captionStart="Table 3" captionStartId="50.[189,258,265,291]" captionTargetPageId="50" captionText="Table 3. Differences between holotype and paratype of M. herteli (Frey, 1971) and paratype of M. gressitti (Frey, 1970)." httpUri="http://table.plazi.org/id/DF6386DDA6070618FF46FEA0FE03E577" pageId="49" pageNumber="50" tableUuid="DF6386DDA6070618FF46FEA0FE03E577">Table 3</tableCitation>
). Therefore, the male genitalia of 
<taxonomicName id="4C1CADD6A604061BFCBFF906FC2FE2F9" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[836,973,1711,1736]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="gressitti">
<emphasis id="B9680A47A604061BFCBFF906FC2FE2F9" box="[836,973,1711,1736]" italics="true" pageId="49" pageNumber="50">M. gressitti</emphasis>
</taxonomicName>
is described based on the 
<typeStatus id="54A768F7A604061BFAF4F906FA94E2F8" box="[1295,1398,1711,1737]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C1CADD6A604061BFF1BF97AFEBAE2DD" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[224,344,1746,1772]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A604061BFF1BF97AFEBAE2DD" box="[224,344,1746,1772]" italics="true" pageId="49" pageNumber="50">M. herteli</emphasis>
</taxonomicName>
, detailed above (morphotype II “ 
<emphasis id="B9680A47A604061BFD0DF97AFCBEE2DC" box="[758,860,1747,1773]" italics="true" pageId="49" pageNumber="50">Gressitti</emphasis>
”). The 
<typeStatus id="54A768F7A604061BFC46F97BFBC4E2DD" box="[957,1062,1746,1772]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C1CADD6A604061BFBA8F97AFB2EE2DD" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[1107,1228,1746,1772]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A604061BFBA8F97AFB2EE2DD" box="[1107,1228,1746,1772]" italics="true" pageId="49" pageNumber="50">M. herteli</emphasis>
</taxonomicName>
is considered to belonging to genital morphotype II “ 
<emphasis id="B9680A47A604061BFD51F95EFCF2E320" box="[682,784,1783,1809]" italics="true" pageId="49" pageNumber="50">Gressitti</emphasis>
”. The features of male genitalia described below confirm it.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA604061BFF46F8F5FE5EE396" box="[189,444,1884,1959]" pageId="49" pageNumber="50" type="distribution">
<paragraph id="8BA3D655A604061BFF46F8F5FEB4E347" blockId="49.[189,342,1884,1910]" box="[189,342,1884,1910]" pageId="49" pageNumber="50">
<heading id="D0EB6139A604061BFF46F8F5FEB4E347" bold="true" box="[189,342,1884,1910]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B9680A47A604061BFF46F8F5FEB4E347" bold="true" box="[189,342,1884,1910]" pageId="49" pageNumber="50">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A604061BFF46F825FE5EE396" blockId="49.[189,444,1932,1959]" box="[189,444,1932,1959]" pageId="49" pageNumber="50">
<collectingCountry id="F30B96C5A604061BFF46F825FF1BE397" box="[189,249,1932,1958]" name="India" pageId="49" pageNumber="50">India</collectingCountry>
(
<collectingRegion id="49D818B7A604061BFEF3F825FE52E396" box="[264,432,1932,1959]" country="India" name="Uttar Pradesh" pageId="49" pageNumber="50">Uttar Pradesh</collectingRegion>
).
</paragraph>
</subSubSection>
<caption id="DF6386DDA6070618FF46FEA0FE03E577" ID-Table-UUID="DF6386DDA6070618FF46FEA0FE03E577" httpUri="http://table.plazi.org/id/DF6386DDA6070618FF46FEA0FE03E577" pageId="50" pageNumber="51" startId="50.[189,258,265,291]" targetBox="[200,1378,395,1431]" targetIsTable="true" targetPageId="50">
<paragraph id="8BA3D655A6070618FF46FEA0FE03E577" blockId="50.[189,1398,265,326]" pageId="50" pageNumber="51">
<emphasis id="B9680A47A6070618FF46FEA0FEC4E512" bold="true" box="[189,294,265,291]" pageId="50" pageNumber="51">Table 3.</emphasis>
Differences between holotype and paratype of 
<taxonomicName id="4C1CADD6A6070618FC78FEA3FB46E512" authority="(Frey, 1971)" baseAuthorityName="Frey" baseAuthorityYear="1971" box="[899,1188,265,291]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A6070618FC78FEA3FC1CE512" box="[899,1022,265,291]" italics="true" pageId="50" pageNumber="51">M. herteli</emphasis>
(
<bibRefCitation id="EF8DABA4A6070618FBEFFEA0FB79E512" author="Frey G." box="[1044,1179,265,291]" pageId="50" pageNumber="51" pagination="206 - 225" refId="ref43999" refString="Frey G. 1971. Bestimmungstabelle der Indischen und Ceylonesischen arten der Gattung Holotrichia Hope. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 22: 206 - 225." type="journal article" year="1971">Frey, 1971</bibRefCitation>
)
</taxonomicName>
and paratype of 
<taxonomicName id="4C1CADD6A6070618FF46FE84FE3BE577" authority="(Frey, 1970)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[189,473,300,326]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="gressitti">
<emphasis id="B9680A47A6070618FF46FE84FEA3E577" box="[189,321,301,326]" italics="true" pageId="50" pageNumber="51">M. gressitti</emphasis>
(
<bibRefCitation id="EF8DABA4A6070618FEAAFE85FE33E577" author="Frey G." box="[337,465,300,326]" pageId="50" pageNumber="51" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey, 1970</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA3D655A6070618FF33FE22FBD4E1A7" pageId="50" pageNumber="51">
<table id="F91C24F5A607F9D5FF33FE22FA80E1A6" box="[200,1378,395,1431]" gridcols="4" gridrows="14" pageId="50" pageNumber="51">
<tr id="352CD417A607F9D5FF33FE22FA80E597" box="[200,1378,395,422]" gridrow="0" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FE22FE35E597" box="[200,471,395,422]" gridcol="0" gridrow="0" pageId="50" pageNumber="51">
<emphasis id="B9680A47A6070618FF33FE22FEDFE592" bold="true" box="[200,317,395,419]" pageId="50" pageNumber="51">Character</emphasis>
</th>
<th id="76FDBD6BA607F9D5FDF2FE22FCF3E597" box="[521,785,395,422]" gridcol="1" gridrow="0" pageId="50" pageNumber="51">
<emphasis id="B9680A47A6070618FDF2FE25FCE8E597" bold="true" box="[521,778,395,422]" pageId="50" pageNumber="51">
<taxonomicName id="4C1CADD6A6070618FDF2FE25FD97E592" baseAuthorityName="Frey" baseAuthorityYear="1971" box="[521,629,395,419]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A6070618FDF2FE25FD97E592" bold="true" box="[521,629,395,419]" italics="true" pageId="50" pageNumber="51">M. herteli</emphasis>
</taxonomicName>
(holotype) ♂
</emphasis>
</th>
<th id="76FDBD6BA607F9D5FCC9FE22FBD5E597" box="[818,1079,395,422]" gridcol="2" gridrow="0" pageId="50" pageNumber="51">
<emphasis id="B9680A47A6070618FCC9FE25FBD5E597" bold="true" box="[818,1079,395,422]" pageId="50" pageNumber="51">
<taxonomicName id="4C1CADD6A6070618FCC9FE25FC7CE592" baseAuthorityName="Frey" baseAuthorityYear="1971" box="[818,926,395,419]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A6070618FCC9FE25FC7CE592" bold="true" box="[818,926,395,419]" italics="true" pageId="50" pageNumber="51">M. herteli</emphasis>
</taxonomicName>
(paratype) ♂
</emphasis>
</th>
<th id="76FDBD6BA607F9D5FBB5FE22FA80E597" box="[1102,1378,395,422]" gridcol="3" gridrow="0" pageId="50" pageNumber="51">
<emphasis id="B9680A47A6070618FBB5FE25FA80E597" bold="true" box="[1102,1378,395,422]" pageId="50" pageNumber="51">
<taxonomicName id="4C1CADD6A6070618FBB5FE25FB2BE592" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[1102,1225,395,419]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="gressitti">
<emphasis id="B9680A47A6070618FBB5FE25FB2BE592" bold="true" box="[1102,1225,395,419]" italics="true" pageId="50" pageNumber="51">M. gressitti</emphasis>
</taxonomicName>
(paratype) ♀
</emphasis>
</th>
</tr>
<tr id="352CD417A607F9D5FF33FE7EFA80E5DE" box="[200,1378,471,495]" gridrow="1" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FE7EFE35E5DE" box="[200,471,471,495]" gridcol="0" gridrow="1" pageId="50" pageNumber="51">Body size</th>
<td id="76FDBD6BA607F9D5FDF2FE7EFCF3E5DE" box="[521,785,471,495]" gridcol="1" gridrow="1" pageId="50" pageNumber="51">&lt;18 mm</td>
<td id="76FDBD6BA607F9D5FCC9FE7EFBD5E5DE" box="[818,1079,471,495]" gridcol="2" gridrow="1" pageId="50" pageNumber="51">≥ 18 mm</td>
<td id="76FDBD6BA607F9D5FBB5FE7EFA80E5DE" box="[1102,1378,471,495]" gridcol="3" gridrow="1" pageId="50" pageNumber="51">≥ 18 mm</td>
</tr>
<tr id="352CD417A607F9D5FF33FD8AFA80E60A" box="[200,1378,547,571]" gridrow="2" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FD8AFE35E60A" box="[200,471,547,571]" gridcol="0" gridrow="2" pageId="50" pageNumber="51">Body appearance</th>
<td id="76FDBD6BA607F9D5FDF2FD8AFCF3E60A" box="[521,785,547,571]" gridcol="1" gridrow="2" pageId="50" pageNumber="51">matte</td>
<td id="76FDBD6BA607F9D5FCC9FD8AFBD5E60A" box="[818,1079,547,571]" gridcol="2" gridrow="2" pageId="50" pageNumber="51">bright</td>
<td id="76FDBD6BA607F9D5FBB5FD8AFA80E60A" box="[1102,1378,547,571]" gridcol="3" gridrow="2" pageId="50" pageNumber="51">bright</td>
</tr>
<tr id="352CD417A607F9D5FF33FDC7FA80E6B7" box="[200,1378,622,646]" gridrow="3" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FDC7FE35E6B7" box="[200,471,622,646]" gridcol="0" gridrow="3" pageId="50" pageNumber="51">Body surface pubescence</th>
<td id="76FDBD6BA607F9D5FDF2FDC7FCF3E6B7" box="[521,785,622,646]" gridcol="1" gridrow="3" pageId="50" pageNumber="51">tiny hardly visible</td>
<td id="76FDBD6BA607F9D5FCC9FDC7FBD5E6B7" box="[818,1079,622,646]" gridcol="2" gridrow="3" pageId="50" pageNumber="51">without pubescence</td>
<td id="76FDBD6BA607F9D5FBB5FDC7FA80E6B7" box="[1102,1378,622,646]" gridcol="3" gridrow="3" pageId="50" pageNumber="51">without pubescence</td>
</tr>
<tr id="352CD417A607F9D5FF33FD03FA80E6D2" box="[200,1378,682,739]" gridrow="4" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FD03FE35E6D2" box="[200,471,682,739]" gridcol="0" gridrow="4" pageId="50" pageNumber="51">Anterior margin of pronotum</th>
<td id="76FDBD6BA607F9D5FDF2FD03FCF3E6D2" box="[521,785,682,739]" gridcol="1" gridrow="4" pageId="50" pageNumber="51">regularly curved</td>
<td id="76FDBD6BA607F9D5FCC9FD03FBD5E6D2" box="[818,1079,682,739]" gridcol="2" gridrow="4" pageId="50" pageNumber="51">wavy forward just behind eyes</td>
<td id="76FDBD6BA607F9D5FBB5FD03FA80E6D2" box="[1102,1378,682,739]" gridcol="3" gridrow="4" pageId="50" pageNumber="51">wavy forward just behind eyes</td>
</tr>
<tr id="352CD417A607F9D5FF33FD5FFA80E71C" box="[200,1378,758,813]" gridrow="5" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FD5FFE35E71C" box="[200,471,758,813]" gridcol="0" gridrow="5" pageId="50" pageNumber="51">Lateral margins of pronotum</th>
<td id="76FDBD6BA607F9D5FDF2FD5FFCF3E71C" box="[521,785,758,813]" gridcol="1" gridrow="5" pageId="50" pageNumber="51">posterior half sinuate</td>
<td id="76FDBD6BA607F9D5FCC9FD5FFBD5E71C" box="[818,1079,758,813]" gridcol="2" gridrow="5" pageId="50" pageNumber="51">posterior half straight</td>
<td id="76FDBD6BA607F9D5FBB5FD5FFA80E71C" box="[1102,1378,758,813]" gridcol="3" gridrow="5" pageId="50" pageNumber="51">posterior half straight</td>
</tr>
<tr id="352CD417A607F9D5FF33FCF8FA80E758" box="[200,1378,849,873]" gridrow="6" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FCF8FE35E758" box="[200,471,849,873]" gridcol="0" gridrow="6" pageId="50" pageNumber="51">Carina on foretibia</th>
<td id="76FDBD6BA607F9D5FDF2FCF8FCF3E758" box="[521,785,849,873]" gridcol="1" gridrow="6" pageId="50" pageNumber="51">sharp</td>
<td id="76FDBD6BA607F9D5FCC9FCF8FBD5E758" box="[818,1079,849,873]" gridcol="2" gridrow="6" pageId="50" pageNumber="51">blunt</td>
<td id="76FDBD6BA607F9D5FBB5FCF8FA80E758" box="[1102,1378,849,873]" gridcol="3" gridrow="6" pageId="50" pageNumber="51">blunt</td>
</tr>
<tr id="352CD417A607F9D5FF33FC32FA80E784" box="[200,1378,923,949]" gridrow="7" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FC32FE35E784" box="[200,471,923,949]" gridcol="0" gridrow="7" pageId="50" pageNumber="51">Inner spur of foretibia</th>
<td id="76FDBD6BA607F9D5FDF2FC32FCF3E784" box="[521,785,923,949]" gridcol="1" gridrow="7" pageId="50" pageNumber="51">between 2nd and 3rd tooth</td>
<td id="76FDBD6BA607F9D5FCC9FC32FBD5E784" box="[818,1079,923,949]" gridcol="2" gridrow="7" pageId="50" pageNumber="51">closer to 3rd tooth</td>
<td id="76FDBD6BA607F9D5FBB5FC32FA80E784" box="[1102,1378,923,949]" gridcol="3" gridrow="7" pageId="50" pageNumber="51">closer to 3rd tooth</td>
</tr>
<tr id="352CD417A607F9D5FF33FC41FA80E031" box="[200,1378,1000,1024]" gridrow="8" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FC41FE35E031" box="[200,471,1000,1024]" gridcol="0" gridrow="8" pageId="50" pageNumber="51">Lateral mesotibial carina</th>
<td id="76FDBD6BA607F9D5FDF2FC41FCF3E031" box="[521,785,1000,1024]" gridcol="1" gridrow="8" pageId="50" pageNumber="51">complete</td>
<td id="76FDBD6BA607F9D5FCC9FC41FBD5E031" box="[818,1079,1000,1024]" gridcol="2" gridrow="8" pageId="50" pageNumber="51">interrupted</td>
<td id="76FDBD6BA607F9D5FBB5FC41FA80E031" box="[1102,1378,1000,1024]" gridcol="3" gridrow="8" pageId="50" pageNumber="51">interrupted</td>
</tr>
<tr id="352CD417A607F9D5FF33FB8DFA80E06D" box="[200,1378,1060,1116]" gridrow="9" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FB8DFE35E06D" box="[200,471,1060,1116]" gridcol="0" gridrow="9" pageId="50" pageNumber="51">Apex of pygidium</th>
<td id="76FDBD6BA607F9D5FDF2FB8DFCF3E06D" box="[521,785,1060,1116]" gridcol="1" gridrow="9" pageId="50" pageNumber="51">regularly marginate</td>
<td id="76FDBD6BA607F9D5FCC9FB8DFBD5E06D" box="[818,1079,1060,1116]" gridcol="2" gridrow="9" pageId="50" pageNumber="51">margin widened and gently raised</td>
<td id="76FDBD6BA607F9D5FBB5FB8DFA80E06D" box="[1102,1378,1060,1116]" gridcol="3" gridrow="9" pageId="50" pageNumber="51">margin widened and gently raised</td>
</tr>
<tr id="352CD417A607F9D5FF33FBC7FA80E098" box="[200,1378,1134,1193]" gridrow="10" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FBC7FE35E098" box="[200,471,1134,1193]" gridcol="0" gridrow="10" pageId="50" pageNumber="51">Fifth ventrite</th>
<td id="76FDBD6BA607F9D5FDF2FBC7FCF3E098" box="[521,785,1134,1193]" gridcol="1" gridrow="10" pageId="50" pageNumber="51">gently depressed at posterior half</td>
<td id="76FDBD6BA607F9D5FCC9FBC7FBD5E098" box="[818,1079,1134,1193]" gridcol="2" gridrow="10" pageId="50" pageNumber="51">not depressed</td>
<td id="76FDBD6BA607F9D5FBB5FBC7FA80E098" box="[1102,1378,1134,1193]" gridcol="3" gridrow="10" pageId="50" pageNumber="51">not depressed</td>
</tr>
<tr id="352CD417A607F9D5FF33FB12FA80E0C2" box="[200,1378,1211,1267]" gridrow="11" pageId="50" pageNumber="51">
<th id="76FDBD6BA607F9D5FF33FB12FE35E0C2" box="[200,471,1211,1267]" gridcol="0" gridrow="11" pageId="50" pageNumber="51">Sixth ventrite</th>
<td id="76FDBD6BA607F9D5FDF2FB12FCF3E0C2" box="[521,785,1211,1267]" gridcol="1" gridrow="11" pageId="50" pageNumber="51">with a transversal anterior keel</td>
<td id="76FDBD6BA607F9D5FCC9FB12FBD5E0C2" box="[818,1079,1211,1267]" gridcol="2" gridrow="11" pageId="50" pageNumber="51">without keel only gently bulging</td>
<td id="76FDBD6BA607F9D5FBB5FB12FA80E0C2" box="[1102,1378,1211,1267]" gridcol="3" gridrow="11" pageId="50" pageNumber="51">without keel only gently bulging</td>
</tr>
<tr id="352CD417A607F9D5FF33FAACFA80E16C" box="[200,1378,1285,1373]" gridrow="12" pageId="50" pageNumber="51" rowspan-3="1">
<th id="76FDBD6BA607F9D5FF33FAACFE35E16C" box="[200,471,1285,1373]" gridcol="0" gridrow="12" pageId="50" pageNumber="51">Intermediate branches of dorsal structure of the parameres</th>
<td id="76FDBD6BA607F9D5FDF2FAACFCF3E16C" box="[521,785,1285,1373]" gridcol="1" gridrow="12" pageId="50" pageNumber="51">longer than ventral branches</td>
<td id="76FDBD6BA607F9D5FCC9FAACFBD5E16C" box="[818,1079,1285,1373]" gridcol="2" gridrow="12" pageId="50" pageNumber="51">shorter than ventral branches</td>
</tr>
<tr id="352CD417A607F9D5FF33FAD6FA80E1A6" box="[200,1378,1407,1431]" gridrow="13" pageId="50" pageNumber="51" rowspan-3="1">
<th id="76FDBD6BA607F9D5FF33FAD6FE35E1A6" box="[200,471,1407,1431]" gridcol="0" gridrow="13" pageId="50" pageNumber="51">Raspula in endophallum</th>
<td id="76FDBD6BA607F9D5FDF2FAD6FCF3E1A6" box="[521,785,1407,1431]" gridcol="1" gridrow="13" pageId="50" pageNumber="51">soft, with scattered setae</td>
<td id="76FDBD6BA607F9D5FCC9FAD6FBD5E1A6" box="[818,1079,1407,1431]" gridcol="2" gridrow="13" pageId="50" pageNumber="51">strong, with plenty setae</td>
</tr>
</table>
</paragraph>
<subSubSection id="C30685DEA6070618FF46FA58FD2DE2B4" pageId="50" pageNumber="51" type="description">
<paragraph id="8BA3D655A6070618FF46FA58FC6BE23C" blockId="50.[189,905,1520,1549]" box="[189,905,1520,1549]" pageId="50" pageNumber="51">
<emphasis id="B9680A47A6070618FF46FA58FC6BE23C" bold="true" box="[189,905,1520,1549]" italics="true" pageId="50" pageNumber="51">Species included in Morphotype III “Leucophthalma”</emphasis>
</paragraph>
<paragraph id="8BA3D655A6070618FF46F98CFD2DE2B4" blockId="50.[189,1399,1573,1670]" pageId="50" pageNumber="51">The group is composed of three species, which are characterized by antenna 10-segmented, pronotal surface densely punctate (distance between punctures equal or smaller than diameter of a puncture) and male and female genitalia as specified above.</paragraph>
</subSubSection>
</treatment>
</document>