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<mods:title id="1BC9CAF1FDFC5285DA1B6FB00BA85B47">Genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from Laos, with descriptions of five new species</mods:title>
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<treatment id="0381830DFFB3FFA2FF0BFDA302D8B54E" ID-DOI="http://doi.org/10.5281/zenodo.3812475" ID-GBIF-Taxon="163392074" ID-Zenodo-Dep="3812475" LSID="urn:lsid:plazi:treatment:0381830DFFB3FFA2FF0BFDA302D8B54E" httpUri="http://treatment.plazi.org/id/0381830DFFB3FFA2FF0BFDA302D8B54E" lastPageNumber="391" pageId="1" pageNumber="391">
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<taxonomicName id="4C284998FFB3FFA2FF0BFDA30262B0A1" authority="Wang" authorityName="Liu &amp; Wang" authorityYear="2020" box="[151,521,615,642]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="391" phylum="Arthropoda" rank="species" species="unidentalis" status="sp. nov.">
<emphasis id="B95CEE09FFB3FFA2FF0BFDA301DCB0A1" bold="true" box="[151,439,615,642]" italics="true" pageId="1" pageNumber="391">Promalactis unidentalis</emphasis>
Wang
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,
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<paragraph id="8B97321BFFB3FFA2FF0BFD4F0179B085" blockId="1.[151,627,615,678]" box="[151,274,651,678]" pageId="1" pageNumber="391">
(
<figureCitation id="13132E9EFFB3FFA2FF03FD4F0086B085" box="[159,237,651,678]" captionStart="FIGURES 18" captionStartId="5.[151,264,1881,1906]" captionTargetBox="[179,1435,179,1854]" captionTargetId="figure@5.[151,1435,179,1854]" captionTargetPageId="5" captionText="FIGURES 18. Adults of Promalactis spp. 1, P. unidentalis, holotype ♂; 2, P. tenuiclavifera, holotype ♂; 3, P. introflexa, holotype ♂; 4, P. bolikhamsaiana, holotype ♂; 5, P. serrulata, holotype ♂; 6, P. carinata Du, Li et Wang, 2011; 7, P. flavidia Kim, 2018; 8, P. apicuncata Wang, 2018. Scales=2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733723" httpUri="https://zenodo.org/record/3733723/files/figure.png" pageId="1" pageNumber="391">Figs. 1</figureCitation>
,
<figureCitation id="13132E9EFFB3FFA2FF67FD480161B085" box="[251,266,652,678]" captionStart="FIGURES 914" captionStartId="6.[151,264,1898,1923]" captionTargetBox="[248,1351,190,1853]" captionTargetId="figure@6.[226,1361,181,1872]" captionTargetPageId="6" captionText="FIGURES 914. Genitalia of Promalactis spp. 9, P. unidentalis, holotype ♂, slide No. LC19247; 10, P. tenuiclavifera, holotype ♂, slide No. LC19241; 11, P. introflexa, holotype ♂, slide No. LC19242; 12a, P. introflexa, paratype ♀, slide No. LC19302, 12b, enlarged signum, slide No. LC19246; 13, P. bolikhamsaiana, holotype ♂, slide No. LC19262; 14, P. serrulata, holotype ♂, slide No. LC19250. Scales=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733725" httpUri="https://zenodo.org/record/3733725/files/figure.png" pageId="1" pageNumber="391">9</figureCitation>
)
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<subSubSection id="C3326190FFB3FFA2FF0BFD1004FEB131" pageId="1" pageNumber="391" type="materials_examined">
<paragraph id="8B97321BFFB3FFA2FF0BFD1004FEB131" blockId="1.[151,1437,723,1902]" pageId="1" pageNumber="391">
<emphasis id="B95CEE09FFB3FFA2FF0BFD10012EB0CD" bold="true" box="[151,325,723,750]" pageId="1" pageNumber="391">Type material.</emphasis>
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,
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Conservation Area (
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,
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), Cham- pasak,
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,
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<collectingDate id="EFD2ED33FFB3FFA2FEE2FD3C0261B131" box="[382,522,760,786]" pageId="1" pageNumber="391" value="2019-07-30">30.VII.2019</collectingDate>
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, coll.
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, MJ Qi &amp;
<collectorName id="26DD57CDFFB3FFA2FC87FD3C0300B131" box="[795,875,760,786]" pageId="1" pageNumber="391">DJ Lee</collectorName>
<emphasis id="B95CEE09FFB3FFA2FCEEFD3C03C1B131" box="[882,938,760,786]" italics="true" pageId="1" pageNumber="391">et al.</emphasis>
, slide
<collectorName id="26DD57CDFFB3FFA2FC68FD3C0421B131" box="[1012,1098,760,786]" pageId="1" pageNumber="391">No. LC</collectorName>
19247.
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</paragraph>
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<subSubSection id="C3326190FFB3FFA2FF5BFCDF02E1B1CA" pageId="1" pageNumber="391" type="diagnosis">
<paragraph id="8B97321BFFB3FFA2FF5BFCDF02E1B1CA" blockId="1.[151,1437,723,1902]" pageId="1" pageNumber="391">
<emphasis id="B95CEE09FFB3FFA2FF5BFCDF0129B115" bold="true" box="[199,322,795,822]" pageId="1" pageNumber="391">Diagnosis.</emphasis>
This new species is similar to
<taxonomicName id="4C284998FFB3FFA2FD38FCD80409B115" authority="Wang, Du &amp; Li, 2013" authorityName="Wang, Du &amp; Li" authorityYear="2013" box="[676,1122,795,822]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="391" phylum="Arthropoda" rank="species" species="superimposita">
<emphasis id="B95CEE09FFB3FFA2FD38FCD8030BB116" box="[676,864,796,821]" italics="true" pageId="1" pageNumber="391">P. superimposita</emphasis>
<bibRefCitation id="EFB94FEAFFB3FFA2FCF4FCD80409B115" author="Wang, Du &amp; Li" box="[872,1122,795,822]" firstAuthor="Wang" pageId="1" pageNumber="391" pagination="401 - 455" refId="ref4932" refString="Wang, S. X., Du, Z. H. &amp; Li, H. H. (2013) Taxonomic study of the genus Promalactis Meyrick, 1908 from Southeast Asia (Lepidoptera: Oecophoridae). Zootaxa, 3669 (4), 401 - 455. https: // doi. org / 10.11646 / zootaxa. 3669.4.1" type="journal article" year="2013">Wang, Du &amp; Li, 2013</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C284998FFB3FFA2FB07FCD800A4B179" authority="Wang, 2018" authorityName="Wang" authorityYear="2018" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="391" phylum="Arthropoda" rank="species" species="trigonilobata">
<emphasis id="B95CEE09FFB3FFA2FB07FCD80527B115" box="[1179,1356,796,822]" italics="true" pageId="1" pageNumber="391">P. trigonilobata</emphasis>
<bibRefCitation id="EFB94FEAFFB3FFA2FAC9FCD800A4B179" author="Wang" firstAuthor="Wang" pageId="1" pageNumber="391" pagination="151 - 160" refId="ref5063" refString="Wang, S. X. &amp; Jia, Y. Y. (2018) Review of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) III. The trilineata species group, with descriptions of four new species. Zootaxa, 4433 (1), 151 - 160. https: // doi. org / 10.11646 / zootaxa. 4433.1.9" type="journal article" year="2018">
Wang,
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in the male genitalia. It can be distinguished from them by the dorsodistal part of the valva not extended ventrad but with a denticle ventroapically; in contrast, the dorsodistal part of the valva is produced ventrad to a large spine in
<taxonomicName id="4C284998FFB3FFA2FF64FC4C01D8B182" authorityName="Wang, Du &amp; Li" authorityYear="2013" box="[248,435,904,929]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="391" phylum="Arthropoda" rank="species" species="superimposita">
<emphasis id="B95CEE09FFB3FFA2FF64FC4C01D8B182" box="[248,435,904,929]" italics="true" pageId="1" pageNumber="391">P. superimposita</emphasis>
</taxonomicName>
, and enlarged ventrad triangularly and has two ventroapical denticles in
<taxonomicName id="4C284998FFB3FFA2FB70FC4C0105B1E5" authority="(Jia &amp; Wang 2018)" baseAuthorityName="Jia &amp; Wang" baseAuthorityYear="2018" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="391" phylum="Arthropoda" rank="species" species="trigonilobata">
<emphasis id="B95CEE09FFB3FFA2FB70FC4C05F7B181" box="[1260,1436,904,930]" italics="true" pageId="1" pageNumber="391">P. trigonilobata</emphasis>
(
<bibRefCitation id="EFB94FEAFFB3FFA2FF03FC6F010DB1E5" author="Jia &amp; Wang" box="[159,358,939,966]" firstAuthor="Jia" pageId="1" pageNumber="391" pagination="151 - 160" refId="ref5063" refString="Wang, S. X. &amp; Jia, Y. Y. (2018) Review of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) III. The trilineata species group, with descriptions of four new species. Zootaxa, 4433 (1), 151 - 160. https: // doi. org / 10.11646 / zootaxa. 4433.1.9" type="journal article" year="2018">Jia &amp; Wang 2018</bibRefCitation>
)
</taxonomicName>
. In addition, the new species has a much slenderer and longer cornutus, which is relatively short and stout spine-like in the latter two species.
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB3FFA2FF5BFC3701FAB522" pageId="1" pageNumber="391" type="description">
<paragraph id="8B97321BFFB3FFA2FF5BFC370282B62D" blockId="1.[151,1437,723,1902]" box="[199,745,1011,1038]" pageId="1" pageNumber="391">
<emphasis id="B95CEE09FFB3FFA2FF5BFC370132B62D" bold="true" box="[199,345,1011,1038]" pageId="1" pageNumber="391">Description.</emphasis>
Adult (
<figureCitation id="13132E9EFFB3FFA2FE31FC370184B62D" box="[429,495,1011,1038]" captionStart="FIGURES 18" captionStartId="5.[151,264,1881,1906]" captionTargetBox="[179,1435,179,1854]" captionTargetId="figure@5.[151,1435,179,1854]" captionTargetPageId="5" captionText="FIGURES 18. Adults of Promalactis spp. 1, P. unidentalis, holotype ♂; 2, P. tenuiclavifera, holotype ♂; 3, P. introflexa, holotype ♂; 4, P. bolikhamsaiana, holotype ♂; 5, P. serrulata, holotype ♂; 6, P. carinata Du, Li et Wang, 2011; 7, P. flavidia Kim, 2018; 8, P. apicuncata Wang, 2018. Scales=2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733723" httpUri="https://zenodo.org/record/3733723/files/figure.png" pageId="1" pageNumber="391">Fig. 1</figureCitation>
). Wingspan 10.0 mm.
</paragraph>
<paragraph id="8B97321BFFB3FFA2FF5BFBDC0135B6BE" blockId="1.[151,1437,723,1902]" pageId="1" pageNumber="391">
<emphasis id="B95CEE09FFB3FFA2FF5BFBDC016FB611" box="[199,260,1048,1074]" italics="true" pageId="1" pageNumber="391">Head</emphasis>
with vertex snowy white, frons greyish yellow, occiput pale rust brown. Labial palpus with second segment dim ochreous yellow on outer surface, yellowish white on inner surface; third segment pale ochreous brown, mixed with white on inner surface, white at apex. Antenna with scape white; flagellum black annulated with white on dorsal surface.
</paragraph>
<paragraph id="8B97321BFFB3FFA2FF5BFB6C02C3B756" blockId="1.[151,1437,723,1902]" pageId="1" pageNumber="391">
<emphasis id="B95CEE09FFB3FFA2FF5BFB6C0173B6E1" box="[199,280,1192,1218]" italics="true" pageId="1" pageNumber="391">Thorax</emphasis>
and tegula pale rust brown. Forewing pale yellowish brown, with grey and blackish brown scales, darkened between basal and antemedian fasciae; markings white edged with black scales: costal spot placed beyond distal 1/3, large, ovate, extending to above lower angle of cell; basal fascia slender; antemedian fascia from basal 1/4 of costal margin to distal 2/5 of dorsum, parallel with basal fascia, uniformly wide; tornal spot brown, ill-defined; apex with dense blackish brown scales; cilia orange yellow, grey along distal part of costal margin and from middle of termen to tornus. Hindwing and cilia brown.
</paragraph>
<paragraph id="8B97321BFFB3FFA2FF5BFA440396B4FE" blockId="1.[151,1437,723,1902]" pageId="1" pageNumber="391">
<emphasis id="B95CEE09FFB3FFA2FF5BFA440106B7B9" box="[199,365,1408,1434]" italics="true" pageId="1" pageNumber="391">Male genitalia</emphasis>
(
<figureCitation id="13132E9EFFB3FFA2FEE0FABB01AAB7B9" box="[380,449,1407,1434]" captionStart="FIGURES 914" captionStartId="6.[151,264,1898,1923]" captionTargetBox="[248,1351,190,1853]" captionTargetId="figure@6.[226,1361,181,1872]" captionTargetPageId="6" captionText="FIGURES 914. Genitalia of Promalactis spp. 9, P. unidentalis, holotype ♂, slide No. LC19247; 10, P. tenuiclavifera, holotype ♂, slide No. LC19241; 11, P. introflexa, holotype ♂, slide No. LC19242; 12a, P. introflexa, paratype ♀, slide No. LC19302, 12b, enlarged signum, slide No. LC19246; 13, P. bolikhamsaiana, holotype ♂, slide No. LC19262; 14, P. serrulata, holotype ♂, slide No. LC19250. Scales=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733725" httpUri="https://zenodo.org/record/3733725/files/figure.png" pageId="1" pageNumber="391">Fig. 9</figureCitation>
). Uncus elongate, wide at base, gradually narrowed to pointed apex, setose mediolaterally. Gnathos as long as uncus; gnathos plate narrow, scaly distally, rounded apically, with a large triangular membranous process from dorsal surface before apex, about 1/4 length of gnathos; lateral arm wide, band-shaped. Tegumen widened medially, bifurcate from posterior 1/4; lateral arm narrowed anteriorly. Valva narrow at base, widened to basal 3/5, then slightly narrowed and divided into two parts: costal part wider, not enlarged, setose distally, obtuse apically, with a ventroapical denticle, costa weakly convex at middle; saccular part slightly shorter than dorsal part, straight on dorsal margin, slightly narrowed to apex; sacculus slightly wide at base, smooth and narrowed to basal 3/5, thereafter densely setose and slightly widened to obtuse apex. Saccus slender, rod-like, approximately 1.5 times length of uncus, narrowly rounded at apex. Juxta small, rounded apically. Aedeagus straight, 1.5 times as long as valva; cornutus slender, straight, slightly shorter than half length of aedeagus.
</paragraph>
<paragraph id="8B97321BFFB3FFA2FF5BF92301FAB522" blockId="1.[151,1437,723,1902]" box="[199,401,1767,1793]" pageId="1" pageNumber="391">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB3FFA2FF5BF8CF0229B505" box="[199,578,1803,1830]" pageId="1" pageNumber="391" type="distribution">
<paragraph id="8B97321BFFB3FFA2FF5BF8CF0229B505" blockId="1.[151,1437,723,1902]" box="[199,578,1803,1830]" pageId="1" pageNumber="391">
<emphasis id="B95CEE09FFB3FFA2FF5BF8CF010AB505" bold="true" box="[199,353,1803,1830]" pageId="1" pageNumber="391">Distribution.</emphasis>
<collectingCountry id="F33F728BFFB3FFA2FEFBF8C801F6B505" box="[359,413,1804,1830]" name="Laos" pageId="1" pageNumber="391">Laos</collectingCountry>
(
<collectingRegion id="49ECFCF9FFB3FFA2FE31F8CF025CB505" box="[429,567,1803,1830]" country="Laos" name="Champasak" pageId="1" pageNumber="391">Champasak</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB3FFA2FF5BF8F402D8B54E" pageId="1" pageNumber="391" type="etymology">
<paragraph id="8B97321BFFB3FFA2FF5BF8F402D8B54E" blockId="1.[151,1437,723,1902]" pageId="1" pageNumber="391">
<emphasis id="B95CEE09FFB3FFA2FF5BF8F40125B569" bold="true" box="[199,334,1840,1866]" pageId="1" pageNumber="391">Etymology.</emphasis>
The epithet of the new species is derived from the Latin
<emphasis id="B95CEE09FFB3FFA2FC79F8F40479B56A" box="[997,1042,1840,1865]" italics="true" pageId="1" pageNumber="391">uni-</emphasis>
and
<emphasis id="B95CEE09FFB3FFA2FBD1F8F404CDB569" box="[1101,1190,1840,1866]" italics="true" pageId="1" pageNumber="391">dentalis</emphasis>
, referring to the ventroapical denticle on the costal part of the valva.
</paragraph>
</subSubSection>
</treatment>
</document>