treatments-xml/data/88/49/4F/88494F6BFFBC002BFEF3F9F91AD2FA8F.xml

<document id="8657E6E7D8B7343FC0EF2604F4E20FC7" ID-DOI="10.5281/zenodo.169967" ID-GBIF-Dataset="a3a633ea-0eff-4a69-bb1a-2f4926194820" ID-ISSN="1175­5326" ID-Zenodo-Dep="169967" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459749142542" checkinUser="plazi" docAuthor="Paulsen, M. J. &amp; Smith, Aaron D." docDate="2005" docId="88494F6BFFBC002BFEF3F9F91AD2FA8F" docLanguage="en" docName="zt01050p060.pdf" docOrigin="Zootaxa 1050" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Nicagus occultus Paulsen &amp; Smith, new species" docType="treatment" docVersion="9" lastPageNumber="55" masterDocId="74703713FFBF0020FFFBFFC71800FFC9" masterDocTitle="A new species of stag beetle from sand dunes in west Texas, and a synopsis of the genus Nicagus (Coleoptera: Lucanidae: Aesalinae: Nicagini)" masterLastPageNumber="60" masterPageNumber="45" pageNumber="48" updateTime="1698211318306" updateUser="plazi">
<mods:mods id="D17BE79EA73BDA9C19559337B63328A6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="665C2973A13C3D3ECE6A96859E4F91B1">
<mods:title id="700006A39B061F5D2A5888F90137D2D6">A new species of stag beetle from sand dunes in west Texas, and a synopsis of the genus Nicagus (Coleoptera: Lucanidae: Aesalinae: Nicagini)</mods:title>
</mods:titleInfo>
<mods:name id="DA52033CB3BA54D7D44815C469390901" type="personal">
<mods:role id="C504D64A94D51BF5CBFCBAAA3D5E77F8">
<mods:roleTerm id="FF2F34EDFFB9DBE8F3312B6E76F16189">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C11CF2A00F34E8B115CCCEBE00AFEAC0">Paulsen, M. J.</mods:namePart>
</mods:name>
<mods:name id="6ABC624CA6279F48C6FE273961626E31" type="personal">
<mods:role id="86B603EBBFBBE73536E2DF9F02F3B888">
<mods:roleTerm id="B32D72E2B8C9E8E559545D3FBE4852F9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5A79394F3562927453E8302B53612D3C">Smith, Aaron D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5797A4E6F8A256C42E5374DD1C9AAF1F">text</mods:typeOfResource>
<mods:relatedItem id="BB04FA38E0396CFF2DAF3F389B0277F6" type="host">
<mods:titleInfo id="E6346B63384CB8ED185773207127D862">
<mods:title id="1CFA078CC7A26EC0DD5E031C00CC6133">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7D7770AE47437730BFCD69A82769B871">
<mods:date id="0869859873F29317332F5638846B43B6">2005</mods:date>
<mods:detail id="04FF561A8B293FDB0C1EF3164B8C1D7C" type="volume">
<mods:number id="947CA7CCD08F685F84C7DD89E4C53904">1050</mods:number>
</mods:detail>
<mods:extent id="BB5309EDABFB0048DAACE60282A88EE7" unit="page">
<mods:start id="7E88F7330B83C1CCF815F239672BAA90">45</mods:start>
<mods:end id="F57ACCA1F8376855B0E5E76CC62EF720">60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="596B6CB0B777ECB897DA57391443B881">journal article</mods:classification>
<mods:identifier id="2D633E6265FF79C15668C5B9178BB3A1" type="DOI">10.5281/zenodo.169967</mods:identifier>
<mods:identifier id="9240D805B5320A52BEAEC3DC3FA42724" type="GBIF-Dataset">a3a633ea-0eff-4a69-bb1a-2f4926194820</mods:identifier>
<mods:identifier id="BC200DD25626E468843DC52DA101DC07" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="47634C95A9816EABA5A98C756F28C3F8" type="Zenodo-Dep">169967</mods:identifier>
</mods:mods>
<treatment id="88494F6BFFBC002BFEF3F9F91AD2FA8F" ID-DOI="http://doi.org/10.5281/zenodo.6266076" ID-GBIF-Taxon="119337302" ID-Zenodo-Dep="6266076" LSID="urn:lsid:plazi:treatment:88494F6BFFBC002BFEF3F9F91AD2FA8F" httpUri="http://treatment.plazi.org/id/88494F6BFFBC002BFEF3F9F91AD2FA8F" lastPageId="11" lastPageNumber="55" pageId="3" pageNumber="48">
<subSubSection id="48FAADF6FFBC0023FEF3F9F91C88F991" box="[264,1160,1598,1624]" pageId="3" pageNumber="48" type="nomenclature">
<paragraph id="005FFE7DFFBC0023FEF3F9F91C88F991" blockId="3.[264,1160,1598,1624]" box="[264,1160,1598,1624]" pageId="3" pageNumber="48">
<heading id="5B174911FFBC0023FEF3F9F91C88F991" bold="true" box="[264,1160,1598,1624]" fontSize="11" level="1" pageId="3" pageNumber="48" reason="1">
<emphasis id="3294226FFFBC0023FEF3F9F91C88F991" bold="true" box="[264,1160,1598,1624]" pageId="3" pageNumber="48">
<taxonomicName id="C7E085FEFFBC0023FEF3F9F91AB5F991" ID-CoL="73Y3R" authority="Paulsen &amp; Smith" authorityName="Paulsen &amp; Smith" box="[264,693,1598,1624]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="occultus" status="sp. nov.">
<emphasis id="3294226FFFBC0023FEF3F9F919D8F991" bold="true" box="[264,472,1598,1624]" italics="true" pageId="3" pageNumber="48">Nicagus occultus</emphasis>
Paulsen &amp; Smith
</taxonomicName>
, 
<taxonomicNameLabel id="29A79F14FFBC0023FD39F9F91B52F991" box="[706,850,1598,1624]" pageId="3" pageNumber="48" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="98DBE2F8FFBC0023FC99F9F91BD1F991" box="[866,977,1598,1624]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[264,368,1834,1858]" captionStartId-1="5.[264,368,709,733]" captionTargetBox-0="[315,1274,998,1779]" captionTargetBox-1="[264,1323,288,685]" captionTargetId-0="figure@4.[257,1316,990,1789]" captionTargetId-1="figure@5.[264,1323,288,685]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Dorsal habitus of N. occultus: a) male and b) female." captionText-1="FIGURE 2. Head and right antenna of N. occultus (ventral view) of: a) male and b) female." httpUri-0="https://zenodo.org/record/169968/files/figure.png" httpUri-1="https://zenodo.org/record/169969/files/figure.png" pageId="3" pageNumber="48">Figs. 1–2</figureCitation>
, 
<figureCitation id="98DBE2F8FFBC0023FC25F9F91BEDF991" box="[990,1005,1598,1624]" captionStart="FIGURE 3" captionStartId="6.[264,368,956,980]" captionTargetBox="[264,1323,526,932]" captionTargetId="figure@6.[264,1323,526,932]" captionTargetPageId="6" captionText="FIGURE 3. Vestiture of pronotum (lateral view) in male a) N. occultus and b) N. obscurus." httpUri="https://zenodo.org/record/169970/files/figure.png" pageId="3" pageNumber="48">3</figureCitation>
a, 4a, 5, 7–9)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="48FAADF6FFBC0024FEF3F94E1991FC7F" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48" type="materials_examined">
<paragraph id="005FFE7DFFBC0024FEF3F94E1991FC7F" blockId="3.[264,1323,1673,1899]" lastBlockId="4.[264,1323,284,950]" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
<emphasis id="3294226FFFBC0023FEF3F94E19D7F96A" bold="true" box="[264,471,1673,1699]" pageId="3" pageNumber="48">
<typeStatus id="DF5B40DFFFBC0023FEF3F94E1945F96A" box="[264,325,1673,1699]" pageId="3" pageNumber="48">Type</typeStatus>
specimens.
</emphasis>
Male 
<typeStatus id="DF5B40DFFFBC0023FDCCF94E1AA0F96A" box="[567,672,1673,1699]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
, female allotype, 
<specimenCount id="16E635F4FFBC0023FC86F94E1BEDF96A" box="[893,1005,1673,1699]" pageId="3" pageNumber="48" type="male">100 male</specimenCount>
and 
<specimenCount id="16E635F4FFBC0023FBD4F94E1CA8F96A" box="[1071,1192,1673,1699]" pageId="3" pageNumber="48" type="female">20 female</specimenCount>
<typeStatus id="DF5B40DFFFBC0023FB48F94E1D27F96A" box="[1203,1319,1673,1699]" pageId="3" pageNumber="48" type="paratype">paratypes</typeStatus>
. 
<typeStatus id="DF5B40DFFFBC0023FEF3F9761975F902" box="[264,373,1713,1739]" pageId="3" pageNumber="48" type="holotype">Holotype</typeStatus>
and allotype deposited at 
<collectionCode id="66F166B8FFBC0023FD5FF9761AFCF902" LSID="urn:lsid:biocol.org:col:34795" box="[676,764,1713,1739]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="48">FMNH</collectionCode>
, labeled: a) “ 
<collectingCountry id="78F7BEEDFFBC0023FC68F9761BD4F902" box="[915,980,1713,1739]" name="United States of America" pageId="3" pageNumber="48">USA</collectingCountry>
: 
<collectionCode id="66F166B8FFBC0023FC24F9761C0AF902" box="[991,1034,1713,1739]" pageId="3" pageNumber="48">TX</collectionCode>
: 
<collectionCode id="66F166B8FFBC0023FBEFF9761C6AF902" box="[1044,1130,1713,1739]" pageId="3" pageNumber="48">WARD</collectionCode>
Co. / Monahans Sandhills State / Park, N of campground on / active dunes 
<geoCoordinate id="65D498BAFFBC0023FC56F91E1C1AF93A" box="[941,1050,1753,1779]" direction="north" orientation="latitude" pageId="3" pageNumber="48" precision="55" value="31.637">N 31.637</geoCoordinate>
W / 102.817, morning, 
<quantity id="C7185398FFBC0023FEF3F8C61953F8D2" box="[264,339,1793,1819]" metricMagnitude="2" metricUnit="m" metricValue="8.55" pageId="3" pageNumber="48" unit="m" value="855.0">855 m</quantity>
/ 
<date id="745ED8BDFFBC0023FE91F8C61A17F8D2" box="[362,535,1793,1819]" pageId="3" pageNumber="48" value="2005-06-05" valueMax="2005-06-06" valueMin="2005-06-05">5–6 JUN 2005</date>
, M.J. Paulsen”; b) “ 
<taxonomicName id="C7E085FEFFBC0023FCFFF8C61BCDF8D3" box="[772,973,1793,1818]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="3294226FFFBC0023FCFFF8C61BCDF8D3" box="[772,973,1793,1818]" italics="true" pageId="3" pageNumber="48">Nicagus occultus</emphasis>
</taxonomicName>
/ PAULSEN &amp; 
<collectionCode id="66F166B8FFBC0023FB76F8C61CE8F8D2" box="[1165,1256,1793,1819]" pageId="3" pageNumber="48">SMITH</collectionCode>
2005 /”, “ 
<typeStatus id="DF5B40DFFFBC0023FEC3F8EE19D9F88A" box="[312,473,1833,1859]" pageId="3" pageNumber="48" type="holotype">HOLOTYPE</typeStatus>
” or “ALLOTYPE”. Male (63) and female (10) 
<typeStatus id="DF5B40DFFFBC0023FBC9F8EE1CA2F88A" box="[1074,1186,1833,1859]" pageId="3" pageNumber="48" type="paratype">paratypes</typeStatus>
labeled: a) labeled as 
<typeStatus id="DF5B40DFFFBC0023FE73F89619F2F8A2" box="[392,498,1873,1899]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
; b) 
<typeStatus id="DF5B40DFFFBC0023FDD9F8961A87F8A2" box="[546,647,1873,1899]" pageId="3" pageNumber="48" type="paratype">paratype</typeStatus>
label: “ 
<taxonomicName id="C7E085FEFFBC0023FD13F8961BB4F8A3" box="[744,948,1873,1898]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="3294226FFFBC0023FD13F8961BB4F8A3" box="[744,948,1873,1898]" italics="true" pageId="3" pageNumber="48">Nicagus occultus</emphasis>
</taxonomicName>
/ PAULSEN &amp; 
<collectionCode id="66F166B8FFBC0023FB82F8961CD4F8A2" box="[1145,1236,1873,1899]" pageId="3" pageNumber="48">SMITH</collectionCode>
2005 / 
<typeStatus id="DF5B40DFFFBB0024FEF3FEDB19A1FEFF" box="[264,417,284,310]" pageId="4" pageNumber="49" type="paratype">PARATYPE</typeStatus>
”. Male (21) and female (4) 
<typeStatus id="DF5B40DFFFBB0024FD0FFEDB1B64FEFF" box="[756,868,284,310]" pageId="4" pageNumber="49" type="paratype">paratypes</typeStatus>
labeled a) “ 
<collectingCountry id="78F7BEEDFFBB0024FC0FFEDB1C35FEFF" box="[1012,1077,284,310]" name="United States of America" pageId="4" pageNumber="49">USA</collectingCountry>
: 
<collectionCode id="66F166B8FFBB0024FBB9FEDB1C6EFEFF" box="[1090,1134,284,310]" pageId="4" pageNumber="49">TX</collectionCode>
: 
<collectionCode id="66F166B8FFBB0024FB81FEDB1CD0FEFF" box="[1146,1232,284,310]" pageId="4" pageNumber="49">WARD</collectionCode>
Co. / 
<quantity id="C7185398FFBB0024FAE7FEDB1927FE97" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="4" pageNumber="49" unit="mi" value="4.0">4 mi</quantity>
NE Monahans, / Monahans Sandhills SP / 
<geoCoordinate id="65D498BAFFBB0024FCDBFE831B9BFE97" box="[800,923,324,350]" direction="north" orientation="latitude" pageId="4" pageNumber="49" precision="5" value="31.5731">N 31.5731</geoCoordinate>
<geoCoordinate id="65D498BAFFBB0024FC59FE831C33FE97" box="[930,1075,324,350]" direction="west" orientation="longitude" pageId="4" pageNumber="49" precision="5" value="-102.8953">W 102.8953</geoCoordinate>
/ 
<date id="745ED8BDFFBB0024FBB2FE831CF4FE97" box="[1097,1268,324,350]" pageId="4" pageNumber="49" value="2005-05-30">30 MAY 2005</date>
; AD Smith”; b) 
<typeStatus id="DF5B40DFFFBB0024FE75FEAB19F3FE4F" box="[398,499,364,390]" pageId="4" pageNumber="49" type="paratype">paratype</typeStatus>
label as above. Male (5) and female (4) 
<typeStatus id="DF5B40DFFFBB0024FC27FEAB1C4CFE4F" box="[988,1100,364,390]" pageId="4" pageNumber="49" type="paratype">paratypes</typeStatus>
labeled: a) “ 
<collectingCountry id="78F7BEEDFFBB0024FB1DFEAB1D27FE4F" box="[1254,1319,364,390]" name="United States of America" pageId="4" pageNumber="49">USA</collectingCountry>
: 
<collectionCode id="66F166B8FFBB0024FEF3FE531933FE67" box="[264,307,404,430]" pageId="4" pageNumber="49">TX</collectionCode>
: Ward County / Monahans Sandhills; 
<quantity id="C7185398FFBB0024FD14FE531B3AFE67" box="[751,826,404,430]" metricMagnitude="3" metricUnit="m" metricValue="7.7248512" pageId="4" pageNumber="49" unit="mi" value="4.8">4.8 mi</quantity>
/ NE of Monahans 
<date id="745ED8BDFFBB0024FBDBFE531C98FE67" box="[1056,1176,404,430]" pageId="4" pageNumber="49" value="2005-05-30">30.V.2005</date>
/ 
<geoCoordinate id="65D498BAFFBB0024FB54FE531D2AFE67" box="[1199,1322,404,430]" direction="north" orientation="latitude" pageId="4" pageNumber="49" precision="5" value="31.5731">N 31.5731</geoCoordinate>
<geoCoordinate id="65D498BAFFBB0024FEF3FE7B19A0FE1F" box="[264,416,444,470]" direction="west" orientation="longitude" pageId="4" pageNumber="49" precision="5" value="-102.8953">W 102.8953</geoCoordinate>
, on / sand dunes, Glene Mynhardt”; b) 
<typeStatus id="DF5B40DFFFBB0024FC73FE7B1BEDFE1F" box="[904,1005,444,470]" pageId="4" pageNumber="49" type="paratype">paratype</typeStatus>
label as above. Male (3) 
<typeStatus id="DF5B40DFFFBB0024FEF3FE231978FE37" box="[264,376,484,510]" pageId="4" pageNumber="49" type="paratype">paratypes</typeStatus>
labeled: a) “ 
<collectingCountry id="78F7BEEDFFBB0024FDE2FE231A5AFE37" box="[537,602,484,510]" name="United States of America" pageId="4" pageNumber="49">USA</collectingCountry>
: 
<collectionCode id="66F166B8FFBB0024FD91FE231A95FE37" box="[618,661,484,510]" pageId="4" pageNumber="49">TX</collectionCode>
: 
<collectionCode id="66F166B8FFBB0024FD5EFE231AFBFE37" box="[677,763,484,510]" pageId="4" pageNumber="49">WARD</collectionCode>
Co. / Monahans Sandhills State / Park, N of campground on / active dunes 
<geoCoordinate id="65D498BAFFBB0024FD94FDCB1ADCFDEF" box="[623,732,524,550]" direction="north" orientation="latitude" pageId="4" pageNumber="49" precision="55" value="31.637">N 31.637</geoCoordinate>
W / 102.817, pre­dusk, 
<quantity id="C7185398FFBB0024FC02FDCB1C43FDEF" box="[1017,1091,524,550]" metricMagnitude="2" metricUnit="m" metricValue="8.55" pageId="4" pageNumber="49" unit="m" value="855.0">855 m</quantity>
/ 
<date id="745ED8BDFFBB0024FBA1FDCB1CEAFDEF" box="[1114,1258,524,550]" pageId="4" pageNumber="49" value="2005-06-04">4 JUN 2005</date>
, M.J. Paulsen”; b) 
<typeStatus id="DF5B40DFFFBB0024FE65FDF31A03FD87" box="[414,515,564,590]" pageId="4" pageNumber="49" type="paratype">paratype</typeStatus>
label as above. Male (6) and female (1) 
<typeStatus id="DF5B40DFFFBB0024FC1BFDF31C50FD87" box="[992,1104,564,590]" pageId="4" pageNumber="49" type="paratype">paratypes</typeStatus>
labeled: a) “ 
<collectingCountry id="78F7BEEDFFBB0024FB18FDF31D24FD87" box="[1251,1316,564,590]" name="United States of America" pageId="4" pageNumber="49">USA</collectingCountry>
: 
<collectionCode id="66F166B8FFBB0024FEF3FD9B1933FDBF" box="[264,307,604,630]" pageId="4" pageNumber="49">TX</collectionCode>
: 
<collectionCode id="66F166B8FFBB0024FEC4FD9B1995FDBF" box="[319,405,604,630]" pageId="4" pageNumber="49">WARD</collectionCode>
Co. / Monahans Sandhills State / Park, N of campground on / active dunes 
<geoCoordinate id="65D498BAFFBB0024FAEEFD9B1958FD57" direction="north" orientation="latitude" pageId="4" pageNumber="49" precision="55" value="31.637">N 31.637</geoCoordinate>
W / 102.817, pre­dusk, 
<quantity id="C7185398FFBB0024FD7BFD431ACCFD57" box="[640,716,644,670]" metricMagnitude="2" metricUnit="m" metricValue="8.55" pageId="4" pageNumber="49" unit="m" value="855.0">855 m</quantity>
/ 
<date id="745ED8BDFFBB0024FD1DFD431B7AFD57" box="[742,890,644,670]" pageId="4" pageNumber="49" value="2005-06-05">5 JUN 2005</date>
, M.J. Paulsen”; b) 
<typeStatus id="DF5B40DFFFBB0024FB99FD431CC7FD57" box="[1122,1223,644,670]" pageId="4" pageNumber="49" type="paratype">paratype</typeStatus>
label as above. Male (2) and female (1) 
<typeStatus id="DF5B40DFFFBB0024FD7EFD6B1AF5FD0F" box="[645,757,684,710]" pageId="4" pageNumber="49" type="paratype">paratypes</typeStatus>
labeled: “ 
<collectingCountry id="78F7BEEDFFBB0024FC97FD6B1BADFD0F" box="[876,941,684,710]" name="United States of America" pageId="4" pageNumber="49">USA</collectingCountry>
: 
<collectionCode id="66F166B8FFBB0024FC43FD6B1BE3FD0F" box="[952,995,684,710]" pageId="4" pageNumber="49">TX</collectionCode>
: WINKLER Co. / 
<quantity id="C7185398FFBB0024FB44FD6B1CF8FD0F" box="[1215,1272,684,710]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="4" pageNumber="49" unit="mi" value="8.0">8 mi</quantity>
. NE Kermit, 
<geoCoordinate id="65D498BAFFBB0024FE97FD1319D9FD27" box="[364,473,724,750]" direction="north" orientation="latitude" pageId="4" pageNumber="49" precision="925" value="31.933332">N 31 56’</geoCoordinate>
/ 24.6” 
<geoCoordinate id="65D498BAFFBB0024FDC4FD131ABEFD27" box="[575,702,724,750]" direction="west" orientation="longitude" pageId="4" pageNumber="49" precision="925" value="-102.96667">W 102 58’</geoCoordinate>
41.3”, elev. 
<quantity id="C7185398FFBB0024FCA9FD131B9FFD27" box="[850,927,724,750]" metricMagnitude="2" metricUnit="m" metricValue="9.28" pageId="4" pageNumber="49" unit="m" value="928.0">928 m</quantity>
/active dunes, morning; / 
<date id="745ED8BDFFBB0024FB26FD131946FCDF" pageId="4" pageNumber="49" value="2005-06-06">6 JUN 2005</date>
; M.J. Paulsen”; b) 
<typeStatus id="DF5B40DFFFBB0024FDB9FD3B1AA7FCDF" box="[578,679,764,790]" pageId="4" pageNumber="49" type="paratype">paratype</typeStatus>
label as above. 
<typeStatus id="DF5B40DFFFBB0024FC7AFD3B1BF3FCDF" box="[897,1011,764,790]" pageId="4" pageNumber="49" type="paratype">Paratypes</typeStatus>
deposited in the 
<collectionCode id="66F166B8FFBB0024FB37FD3B1D24FCDF" LSID="urn:lsid:biocol.org:col:34795" box="[1228,1316,764,790]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="49">FMNH</collectionCode>
, 
<collectionCode id="66F166B8FFBB0024FEF3FCE31961FCF7" box="[264,353,804,830]" pageId="4" pageNumber="49">CMNC</collectionCode>
, 
<collectionCode id="66F166B8FFBB0024FE8AFCE319BDFCF7" box="[369,445,804,830]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="49">FSCA</collectionCode>
, 
<collectionCode id="66F166B8FFBB0024FE35FCE31A26FCF7" LSID="urn:lsid:biocol.org:col:34871" box="[462,550,804,830]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="49">USNM</collectionCode>
, 
<collectionCode id="66F166B8FFBB0024FDCEFCE31A8AFCF7" LSID="urn:lsid:biocol.org:col:34868" box="[565,650,804,830]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="4" pageNumber="49">UNSM</collectionCode>
and in the collections of J.C. Abbott (Austin, 
<collectionCode id="66F166B8FFBB0024FB36FCE31CF8FCF7" box="[1229,1272,804,830]" pageId="4" pageNumber="49">TX</collectionCode>
), L. Bartolozzi (Firenze, 
<collectingCountry id="78F7BEEDFFBB0024FE06FC8B1A37FCAF" box="[509,567,844,870]" name="Italy" pageId="4" pageNumber="49">Italy</collectingCountry>
), D.C. Hawks (Riverside, CA), S. Kawai (Tokyo, 
<collectingCountry id="78F7BEEDFFBB0024FB60FC8B1CE2FCAF" box="[1179,1250,844,870]" name="Japan" pageId="4" pageNumber="49">Japan</collectingCountry>
), E.G. Riley &amp; Texas A&amp;M University (College Station, 
<collectionCode id="66F166B8FFBB0024FCAFFCB31B7FFC47" box="[852,895,884,910]" pageId="4" pageNumber="49">TX</collectionCode>
), P.E. Skelley (Gainesville, FL), and the authors.
</paragraph>
</subSubSection>
<caption id="549FAEF5FFBB0024FEF3F8ED1BCDF888" box="[264,973,1834,1858]" httpUri="https://zenodo.org/record/169968/files/figure.png" pageId="4" pageNumber="49" targetBox="[315,1274,998,1779]" targetPageId="4">
<paragraph id="005FFE7DFFBB0024FEF3F8ED1BCDF888" blockId="4.[264,973,1834,1858]" box="[264,973,1834,1858]" pageId="4" pageNumber="49">
<emphasis id="3294226FFFBB0024FEF3F8ED198AF88B" bold="true" box="[264,394,1834,1858]" pageId="4" pageNumber="49">FIGURE 1.</emphasis>
Dorsal habitus of 
<emphasis id="3294226FFFBB0024FDADF8ED1AD7F888" box="[598,727,1834,1857]" italics="true" pageId="4" pageNumber="49">
<taxonomicName id="C7E085FEFFBB0024FDADF8ED1AD1F888" box="[598,721,1834,1857]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="occultus">N. occultus</taxonomicName>
:
</emphasis>
a) male and b) female.
</paragraph>
</caption>
<caption id="549FAEF5FFBA0025FEF3FD021CE3FD14" box="[264,1251,709,733]" httpUri="https://zenodo.org/record/169969/files/figure.png" pageId="5" pageNumber="50" targetBox="[264,1323,288,685]" targetPageId="5">
<paragraph id="005FFE7DFFBA0025FEF3FD021CE3FD14" blockId="5.[264,1251,709,733]" box="[264,1251,709,733]" pageId="5" pageNumber="50">
<emphasis id="3294226FFFBA0025FEF3FD02198AFD14" bold="true" box="[264,394,709,733]" pageId="5" pageNumber="50">FIGURE 2.</emphasis>
Head and right antenna of 
<taxonomicName id="C7E085FEFFBA0025FD48FD011B2BFD14" box="[691,811,710,733]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="3294226FFFBA0025FD48FD011B2BFD14" box="[691,811,710,733]" italics="true" pageId="5" pageNumber="50">N. occultus</emphasis>
</taxonomicName>
(ventral view) of: a) male and b) female.
</paragraph>
</caption>
<subSubSection id="48FAADF6FFBA0027FEC3FCD71CB5FB9F" lastPageId="7" lastPageNumber="52" pageId="5" pageNumber="50" type="description">
<paragraph id="005FFE7DFFBA0026FEC3FCD71B31FE1F" blockId="5.[264,1323,784,1890]" lastBlockId="6.[264,1323,284,470]" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
<emphasis id="3294226FFFBA0025FEC3FCD71B04FCE3" bold="true" box="[312,772,784,810]" pageId="5" pageNumber="50">
Male 
<typeStatus id="DF5B40DFFFBA0025FE71FCD719F5FCE3" box="[394,501,784,810]" pageId="5" pageNumber="50" type="holotype">holotype</typeStatus>
description. 
<emphasis id="3294226FFFBA0025FD55FCD71B04FCE3" box="[686,772,784,810]" italics="true" pageId="5" pageNumber="50">Length</emphasis>
</emphasis>
: 
<quantity id="C7185398FFBA0025FCDAFCD71B8AFCE3" box="[801,906,784,810]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="5" pageNumber="50" unit="mm" value="7.8">7.8 mm</quantity>
. 
<emphasis id="3294226FFFBA0025FC5AFCD71BF1FCE3" bold="true" box="[929,1009,784,810]" italics="true" pageId="5" pageNumber="50">Width:</emphasis>
<quantity id="C7185398FFBA0025FBFEFCD71C6EFCE3" box="[1029,1134,784,810]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="50" unit="mm" value="3.5">3.5 mm</quantity>
. 
<emphasis id="3294226FFFBA0025FB7FFCD71CD3FCE3" box="[1156,1235,784,810]" italics="true" pageId="5" pageNumber="50">
<emphasis id="3294226FFFBA0025FB7FFCD71CC8FCE3" bold="true" box="[1156,1224,784,810]" italics="true" pageId="5" pageNumber="50">Color</emphasis>
:
</emphasis>
Head, pronotum, legs dark brown; elytra, tarsi, antennae slightly lighter, reddish brown. 
<emphasis id="3294226FFFBA0025FB1BFCFF1D2AFC9B" bold="true" box="[1248,1322,824,850]" italics="true" pageId="5" pageNumber="50">Head:</emphasis>
Eye large, entire. Anteocular margin with rounded process protruding in front of eye. Vertex tumid, in lateral view appearing subtuberculate. Frons weakly depressed behind anterior margin, margin rounded apically and produced near antennal insertions, not strongly reflexed. Clypeus (inter­mandibular projection) nearly vertical, one­third as long as broad, emarginate anteroventrally to receive labrum. Labrum semi­circular, projecting anteroventrally. Mandibles short, broad, densely setose internally, not obscured by setae, acute apically, apices prominent in dorsal view. Antennae not geniculate; scape short, not longer than combined length of segments 2–7; club closeable, elongate lamellae fitting together loosely when closed, lamellae distinctly longer than last segment of maxillary palpus (
<figureCitation id="98DBE2F8FFBA0025FE8AFB0F19C5FB2B" box="[369,453,1224,1250]" captionStart="FIGURE 2" captionStartId="5.[264,368,709,733]" captionTargetBox="[264,1323,288,685]" captionTargetId="figure@5.[264,1323,288,685]" captionTargetPageId="5" captionText="FIGURE 2. Head and right antenna of N. occultus (ventral view) of: a) male and b) female." httpUri="https://zenodo.org/record/169969/files/figure.png" pageId="5" pageNumber="50">Fig. 2</figureCitation>
a). Surface setose, densely punctate; punctures large, contiguous. 
<emphasis id="3294226FFFBA0025FEF3FB371982FAC3" bold="true" box="[264,386,1264,1290]" italics="true" pageId="5" pageNumber="50">Pronotum</emphasis>
: Lateral margins not explanate, weakly crenulate, subangulate behind middle, nearly straight from anterior margin to angulation, emarginate from angulation to acute posterior angle. Margins fimbriate with setae longer than scutellum. Surface densely punctate except at midline; punctures large, each bearing 1 long, fine seta; setae palecolored (almost colorless), erect (
<figureCitation id="98DBE2F8FFBA0025FD57FA571AF9FA63" box="[684,761,1424,1450]" captionStart="FIGURE 3" captionStartId="6.[264,368,956,980]" captionTargetBox="[264,1323,526,932]" captionTargetId="figure@6.[264,1323,526,932]" captionTargetPageId="6" captionText="FIGURE 3. Vestiture of pronotum (lateral view) in male a) N. occultus and b) N. obscurus." httpUri="https://zenodo.org/record/169970/files/figure.png" pageId="5" pageNumber="50">Fig. 3</figureCitation>
a). 
<emphasis id="3294226FFFBA0025FCDDFA571BA9FA63" bold="true" box="[806,937,1424,1450]" italics="true" pageId="5" pageNumber="50">Scutellum:</emphasis>
Surface medially and apically glabrous, punctate and setose basally. 
<emphasis id="3294226FFFBA0025FD28FA7F1B27FA1B" bold="true" box="[723,807,1464,1490]" italics="true" pageId="5" pageNumber="50">Elytra:</emphasis>
Lateral margins crenulate, fimbriate; setae longest at base, basal setae subequal in length to scutellum. Surface punctate; punctures large, each bearing 1 long, fine seta at anterior end; setae mostly more than 2 times longer than puncture diameter, pale­colored (almost colorless), erect. 
<emphasis id="3294226FFFBA0025FB8BF9F71CB3F983" bold="true" box="[1136,1203,1584,1610]" italics="true" pageId="5" pageNumber="50">Legs:</emphasis>
Protibia quadridentate 2 basal teeth small, apical 2 teeth strongly developed, acute; spur unciform, inwardly curved. Protarsus with last tarsomere subequal in length to tarsomeres 1–4 combined. Mesotibia with weak, oblique carina at basal third; spurs both longer than basal tarsomere, acute. Metatibia robust, in lateral view gradually widening towards apex (
<figureCitation id="98DBE2F8FFBA0025FB00F9171918F8DB" captionStart="FIGURES 5 – 8. 5" captionStartId="9.[264,382,1763,1787]" captionTargetBox="[264,1322,609,1753]" captionTargetId="figure@9.[264,1323,609,1761]" captionTargetPageId="9" captionText="FIGURES 5 – 8. 5. Lateral view of male N. occultus, illustrating diagnostic characters: short, conical metatibia; weakly curved claws; and dorsally curved parameres. 6. Lateral view of elongate male metatibia of N. obscurus. 7. Apex of metatibiae of male N. occultus. 8. Male genitalia of N. occultus (dorsal view), showing rounded apex of median lobe." httpUri="https://zenodo.org/record/169972/files/figure.png" pageId="5" pageNumber="50">Fig. 5</figureCitation>
), apex 1/3 as wide as long, oblique; posterior face expanded (
<figureCitation id="98DBE2F8FFBA0025FBD1F93F1C77F8DB" box="[1066,1143,1784,1810]" captionStart="FIGURES 5 – 8. 5" captionStartId="9.[264,382,1763,1787]" captionTargetBox="[264,1322,609,1753]" captionTargetId="figure@9.[264,1323,609,1761]" captionTargetPageId="9" captionText="FIGURES 5 – 8. 5. Lateral view of male N. occultus, illustrating diagnostic characters: short, conical metatibia; weakly curved claws; and dorsally curved parameres. 6. Lateral view of elongate male metatibia of N. obscurus. 7. Apex of metatibiae of male N. occultus. 8. Male genitalia of N. occultus (dorsal view), showing rounded apex of median lobe." httpUri="https://zenodo.org/record/169972/files/figure.png" pageId="5" pageNumber="50">Fig. 7</figureCitation>
); inferior spur subequal in length to basal tarsomere, longer superior spur subequal in length to tarsomeres 1 and 2 combined, spatulate after middle, both spurs with spatulate faces coplanar with oblique apex. External tooth of metatibia reduced to weak carina. Metatarsus plus claw length as long as metatibia. Claws long, weakly curved, subequal in length to last tarsal segment. 
<emphasis id="3294226FFFB90026FD9DFEAB1AE0FE4F" bold="true" box="[614,736,364,390]" italics="true" pageId="6" pageNumber="51">Genitalia:</emphasis>
(
<figureCitation id="98DBE2F8FFB90026FD0CFEAB1B3EFE4F" box="[759,830,364,390]" captionStart="FIGURES 5 – 8. 5" captionStartId="9.[264,382,1763,1787]" captionTargetBox="[264,1322,609,1753]" captionTargetId="figure@9.[264,1323,609,1761]" captionTargetPageId="9" captionText="FIGURES 5 – 8. 5. Lateral view of male N. occultus, illustrating diagnostic characters: short, conical metatibia; weakly curved claws; and dorsally curved parameres. 6. Lateral view of elongate male metatibia of N. obscurus. 7. Apex of metatibiae of male N. occultus. 8. Male genitalia of N. occultus (dorsal view), showing rounded apex of median lobe." httpUri="https://zenodo.org/record/169972/files/figure.png" pageId="6" pageNumber="51">Fig. 8</figureCitation>
). Median lobe short, not reaching apex of parameres, apex broadly rounded (not acuminate), apical margin continuous with ventral concavity. Parameres strongly curved dorsally.
</paragraph>
<caption id="549FAEF5FFB90026FEF3FC7B1CDEFC1D" box="[264,1246,956,980]" httpUri="https://zenodo.org/record/169970/files/figure.png" pageId="6" pageNumber="51" targetBox="[264,1323,526,932]" targetPageId="6">
<paragraph id="005FFE7DFFB90026FEF3FC7B1CDEFC1D" blockId="6.[264,1246,956,980]" box="[264,1246,956,980]" pageId="6" pageNumber="51">
<emphasis id="3294226FFFB90026FEF3FC7B198AFC1D" bold="true" box="[264,394,956,980]" pageId="6" pageNumber="51">FIGURE 3.</emphasis>
Vestiture of pronotum (lateral view) in male a) 
<taxonomicName id="C7E085FEFFB90026FC74FC7A1C07FC1D" box="[911,1031,957,980]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="3294226FFFB90026FC74FC7A1C07FC1D" box="[911,1031,957,980]" italics="true" pageId="6" pageNumber="51">N. occultus</emphasis>
</taxonomicName>
and b) 
<taxonomicName id="C7E085FEFFB90026FBA3FC7A1CDBFC1D" box="[1112,1243,957,980]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="obscurus">N. obscurus</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="005FFE7DFFB90026FEC3FBC01979F900" blockId="6.[264,1323,1031,1898]" pageId="6" pageNumber="51">
<emphasis id="3294226FFFB90026FEC3FBC01B0CFBE8" bold="true" box="[312,780,1031,1057]" pageId="6" pageNumber="51">
Female allotype description. 
<emphasis id="3294226FFFB90026FD57FBC01B0CFBE8" bold="true" box="[684,780,1031,1057]" italics="true" pageId="6" pageNumber="51">Length:</emphasis>
</emphasis>
<quantity id="C7185398FFB90026FCEEFBCF1B73FBEB" box="[789,883,1032,1058]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="51" unit="mm" value="7.8">7.8 mm</quantity>
. 
<emphasis id="3294226FFFB90026FC7BFBC01BD1FBE8" bold="true" box="[896,977,1031,1057]" italics="true" pageId="6" pageNumber="51">Width:</emphasis>
<quantity id="C7185398FFB90026FC21FBCF1C39FBEB" box="[986,1081,1032,1058]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="51" unit="mm" value="3.9">3.9 mm</quantity>
. As 
<typeStatus id="DF5B40DFFFB90026FB95FBCF1CD4FBEB" box="[1134,1236,1032,1058]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
except in the following respects: 
<emphasis id="3294226FFFB90026FDBFFBE81A8FFB80" bold="true" box="[580,655,1071,1097]" italics="true" pageId="6" pageNumber="51">Head:</emphasis>
Color darker, entirely dark brown except tarsi, antennae lighter reddish brown. Eye smaller than in male. Anteocular margin with subtriangular process protruding in front of eye. Vertex tumid, in lateral view appearing subtuberculate. Antennal club oval, small, lamellae subequal in length with last segment of maxillary palpus (
<figureCitation id="98DBE2F8FFB90026FE9DFB1719AEFB23" box="[358,430,1232,1258]" captionStart="FIGURE 2" captionStartId="5.[264,368,709,733]" captionTargetBox="[264,1323,288,685]" captionTargetId="figure@5.[264,1323,288,685]" captionTargetPageId="5" captionText="FIGURE 2. Head and right antenna of N. occultus (ventral view) of: a) male and b) female." httpUri="https://zenodo.org/record/169969/files/figure.png" pageId="6" pageNumber="51">Fig. 2</figureCitation>
b). 
<emphasis id="3294226FFFB90026FE2EFB081A4FFB20" bold="true" box="[469,591,1231,1257]" italics="true" pageId="6" pageNumber="51">Pronotum</emphasis>
: Lateral margin weakly crenulate, crenulation lacking behind median angulation. 
<emphasis id="3294226FFFB90026FE01FB301A4EFAD8" bold="true" box="[506,590,1271,1297]" italics="true" pageId="6" pageNumber="51">Elytra:</emphasis>
Surface setose, setae fine, short, most shorter than 2 puncture diameters. 
<emphasis id="3294226FFFB90026FE70FAD819CEFAF0" bold="true" box="[395,462,1311,1337]" italics="true" pageId="6" pageNumber="51">Legs:</emphasis>
Protibia quadridentate, 2 basal teeth small, 2 apical teeth more strongly developed than in male; spur simple, short, acute. All claws curved, shorter than in male. Mesotibiae with 1–2 weak external teeth; spurs both longer than basal tarsomere, acute. Metatibia robust, in lateral view gradually widening towards apex, at apex nearly 1/2 as wide as long, with external oblique carina at basal third, weakly developed; spurs broader than in male, inferior spur longer than basal tarsomere, superior spur abruptly expanded at middle, broadly rounded apically, spatulate. Metatarsus shortened, length including claw shorter than metatibia. 
<emphasis id="3294226FFFB90026FDDAF9F01A9AF998" bold="true" box="[545,666,1591,1617]" italics="true" pageId="6" pageNumber="51">Genitalia:</emphasis>
Sclerotized structures similar to those of 
<taxonomicName id="C7E085FEFFB90026FB60F9FF1D2BF998" box="[1179,1323,1592,1617]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB90026FB60F9FF1D2BF998" box="[1179,1323,1592,1617]" italics="true" pageId="6" pageNumber="51">N. obscurus</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="6471838CFFB90026FEBFF9A71A01F9B3" author="Holloway" box="[324,513,1632,1658]" pageId="6" pageNumber="59" refString="Holloway, B. A. (1969) Further studies on generic relationships in Lucanidae (Insecta: Coleoptera) with special reference to the ocular canthus. New Zealand Journal of Science, 12, 958 - 977." type="journal article" year="1969">Holloway 1969</bibRefCitation>
) except coxites with longer setae, hemisternite of ninth segment curved internally at base, and ninth pleurite approximately 2 times wider than in 
<emphasis id="3294226FFFB90026FAEBF94F1979F900" italics="true" pageId="6" pageNumber="51">
<taxonomicName id="C7E085FEFFB90026FAEBF94F1975F900" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="obscurus">N. obscurus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="005FFE7DFFB90027FEC3F9101957FE67" blockId="6.[264,1323,1031,1898]" lastBlockId="7.[264,1324,284,1870]" lastPageId="7" lastPageNumber="52" pageId="6" pageNumber="51">
<emphasis id="3294226FFFB90026FEC3F91019B1F938" bold="true" box="[312,433,1751,1777]" pageId="6" pageNumber="51">Variation.</emphasis>
Males (n=100). Male 
<typeStatus id="DF5B40DFFFB90026FD3EF91F1B35F93B" box="[709,821,1752,1778]" pageId="6" pageNumber="51" type="paratype">paratypes</typeStatus>
differ from the 
<typeStatus id="DF5B40DFFFB90026FC01F91F1C60F93B" box="[1018,1120,1752,1778]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
in the following characters: 
<emphasis id="3294226FFFB90026FE6EF93819F6F8D0" box="[405,502,1791,1817]" italics="true" pageId="6" pageNumber="51">
<emphasis id="3294226FFFB90026FE6EF93819EBF8D0" bold="true" box="[405,491,1791,1817]" italics="true" pageId="6" pageNumber="51">Length</emphasis>
:
</emphasis>
<quantity id="C7185398FFB90026FE06F8C71A8DF8D3" box="[509,653,1792,1818]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="7.9" metricValueMin="5.8" pageId="6" pageNumber="51" unit="mm" value="6.85" valueMax="7.9" valueMin="5.8">5.8–7.9 mm</quantity>
. 
<emphasis id="3294226FFFB90026FD6CF9381AE7F8D0" bold="true" box="[663,743,1791,1817]" italics="true" pageId="6" pageNumber="51">Width:</emphasis>
<quantity id="C7185398FFB90026FD0BF8C71B7EF8D3" box="[752,894,1792,1818]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="6" pageNumber="51" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 mm</quantity>
. 
<emphasis id="3294226FFFB90026FC72F9381BD8F8D0" bold="true" box="[905,984,1791,1817]" italics="true" pageId="6" pageNumber="51">Color:</emphasis>
light brown (teneral) to dark brown. 
<emphasis id="3294226FFFB90026FE9FF8E019AFF888" bold="true" box="[356,431,1831,1857]" italics="true" pageId="6" pageNumber="51">Head:</emphasis>
Anteocular process more or less strongly developed. Frons with weak to strong excavation behind anterior margin. Vertex with median elevation variably developed from simply tumid to tuberculate. 
<emphasis id="3294226FFFB80027FCB7FEDB1BC6FEFF" bold="true" box="[844,966,284,310]" italics="true" pageId="7" pageNumber="52">Pronotum</emphasis>
: Lateral margin rarely with crenulations reduced to absent behind middle. 
<emphasis id="3294226FFFB80027FCC6FE831B81FE97" bold="true" box="[829,897,324,350]" italics="true" pageId="7" pageNumber="52">Legs:</emphasis>
Anterior tibia with basal 1–2 teeth reduced or absent. Metatibia with external tooth usually absent or present as a reduced carina.
</paragraph>
<paragraph id="005FFE7DFFB80027FEC3FE7B1C1DFD0F" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="52">
Females (n= 20). Female 
<typeStatus id="DF5B40DFFFB80027FD6AFE7B1B01FE1F" box="[657,769,444,470]" pageId="7" pageNumber="52" type="paratype">paratypes</typeStatus>
differ from the allotype in the following characters: 
<emphasis id="3294226FFFB80027FE74FE2319E5FE37" bold="true" box="[399,485,484,510]" italics="true" pageId="7" pageNumber="52">Length</emphasis>
: 
<quantity id="C7185398FFB80027FE0DFE231A85FE37" box="[502,645,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.4" metricValueMin="6.6" pageId="7" pageNumber="52" unit="mm" value="8.0" valueMax="9.4" valueMin="6.6">6.6–9.4 mm</quantity>
. 
<emphasis id="3294226FFFB80027FD6AFE231AD8FE37" bold="true" box="[657,728,484,510]" italics="true" pageId="7" pageNumber="52">Width</emphasis>
: 
<quantity id="C7185398FFB80027FD12FE231B7AFE37" box="[745,890,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.7" metricValueMin="3.3" pageId="7" pageNumber="52" unit="mm" value="4.0" valueMax="4.7" valueMin="3.3">3.3–4.7 mm</quantity>
. 
<emphasis id="3294226FFFB80027FC7EFE231BD4FE37" bold="true" box="[901,980,484,510]" italics="true" pageId="7" pageNumber="52">Color:</emphasis>
light brown (teneral) to dark brown. 
<emphasis id="3294226FFFB80027FE91FDCB19B4FDEF" bold="true" box="[362,436,524,550]" italics="true" pageId="7" pageNumber="52">Head:</emphasis>
Anteocular margin with process subtriangular to rounded. 
<emphasis id="3294226FFFB80027FB53FDCB1D22FDEF" bold="true" box="[1192,1314,524,550]" italics="true" pageId="7" pageNumber="52">Pronotum</emphasis>
: Lateral margins with crenulation weak to absent. Posterior angle produced, subacute to acute. Surface with glabrous impunctate spot variably enlarged along midline. 
<emphasis id="3294226FFFB80027FB13FD9B1D2BFDBF" bold="true" box="[1256,1323,604,630]" italics="true" pageId="7" pageNumber="52">Legs:</emphasis>
Protibia tridentate or quadridentate, 1–2 basal teeth variably reduced or absent. External oblique carina at basal third variably produced, but never toothlike.
</paragraph>
<paragraph id="005FFE7DFFB80027FEC3FD131CB5FB9F" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="52">
<emphasis id="3294226FFFB80027FEC3FD131A39FD27" bold="true" box="[312,569,724,750]" pageId="7" pageNumber="52">Sexual Dimorphism.</emphasis>
Species of 
<taxonomicName id="C7E085FEFFB80027FD30FD131B2BFD24" box="[715,811,724,749]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="3294226FFFB80027FD30FD131B2BFD24" box="[715,811,724,749]" italics="true" pageId="7" pageNumber="52">Nicagus</emphasis>
</taxonomicName>
show a high degree of sexual dimorphism. Females are generally larger, darker, and more oval (
<figureCitation id="98DBE2F8FFB80027FC76FD3B1BD5FCDF" box="[909,981,764,790]" captionStart="FIGURE 1" captionStartId="4.[264,368,1834,1858]" captionTargetBox="[315,1274,998,1779]" captionTargetId="figure@4.[257,1316,990,1789]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of N. occultus: a) male and b) female." httpUri="https://zenodo.org/record/169968/files/figure.png" pageId="7" pageNumber="52">Fig. 1</figureCitation>
). This is accentuated in 
<taxonomicName id="C7E085FEFFB80027FAEBFD3B1972FCF4" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB80027FAEBFD3B1972FCF4" italics="true" pageId="7" pageNumber="52">N. obscurus</emphasis>
</taxonomicName>
and 
<taxonomicName id="C7E085FEFFB80027FE4BFCE31A47FCF4" box="[432,583,804,829]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="3294226FFFB80027FE4BFCE31A47FCF4" box="[432,583,804,829]" italics="true" pageId="7" pageNumber="52">N. japonicus</emphasis>
</taxonomicName>
because the dorsal punctures of males are often filled with a grayish exudate or debris. This coating was not found in any male specimens of 
<emphasis id="3294226FFFB80027FAEBFC8B1971FC44" italics="true" pageId="7" pageNumber="52">
<taxonomicName id="C7E085FEFFB80027FAEBFC8B196CFC44" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="occultus">N. occultus</taxonomicName>
,
</emphasis>
but males are generally still lighter in color than females. A more useful character for determining the sexes is the size of the antennal club. In females, the antennal club is small and round with segments about equal in length to the last segment of the maxillary palpus (
<figureCitation id="98DBE2F8FFB80027FDF6FC2B1A55FBCF" box="[525,597,1004,1030]" captionStart="FIGURE 2" captionStartId="5.[264,368,709,733]" captionTargetBox="[264,1323,288,685]" captionTargetId="figure@5.[264,1323,288,685]" captionTargetPageId="5" captionText="FIGURE 2. Head and right antenna of N. occultus (ventral view) of: a) male and b) female." httpUri="https://zenodo.org/record/169969/files/figure.png" pageId="7" pageNumber="52">Fig. 2</figureCitation>
b). Males have a much larger club with more elongate club segments, each distinctly longer than the last segment of the maxillary palpus (
<figureCitation id="98DBE2F8FFB80027FB3FFBD31D0EFBE7" box="[1220,1294,1044,1070]" captionStart="FIGURE 2" captionStartId="5.[264,368,709,733]" captionTargetBox="[264,1323,288,685]" captionTargetId="figure@5.[264,1323,288,685]" captionTargetPageId="5" captionText="FIGURE 2. Head and right antenna of N. occultus (ventral view) of: a) male and b) female." httpUri="https://zenodo.org/record/169969/files/figure.png" pageId="7" pageNumber="52">Fig. 2</figureCitation>
a). Females also have more robust metatibiae and shorter metatarsus than do males.
</paragraph>
</subSubSection>
<subSubSection id="48FAADF6FFB80028FEC3FBA31C09FB82" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52" type="diagnosis">
<paragraph id="005FFE7DFFB80027FEC3FBA31B44F964" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="52">
<emphasis id="3294226FFFB80027FEC3FBA319B7FBB7" bold="true" box="[312,439,1124,1150]" pageId="7" pageNumber="52">Diagnosis.</emphasis>
This species is readily separated from its only North American congener (
<taxonomicName id="C7E085FEFFB80027FEE9FB4B199FFB6C" box="[274,415,1164,1189]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB80027FEE9FB4B199FFB6C" box="[274,415,1164,1189]" italics="true" pageId="7" pageNumber="52">N. obscurus</emphasis>
</taxonomicName>
) by its dorsal vestiture, weakly crenulate lateral pronotal margins, and shape of the metatibiae. In 
<taxonomicName id="C7E085FEFFB80027FE01FB731A7DFB04" box="[506,637,1204,1229]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="3294226FFFB80027FE01FB731A7DFB04" box="[506,637,1204,1229]" italics="true" pageId="7" pageNumber="52">N. occultus</emphasis>
</taxonomicName>
the dorsum of males bears mostly long, pale­colored setae (
<figureCitation id="98DBE2F8FFB80027FEEAFB1B195AFB3F" box="[273,346,1244,1270]" captionStart="FIGURE 3" captionStartId="6.[264,368,956,980]" captionTargetBox="[264,1323,526,932]" captionTargetId="figure@6.[264,1323,526,932]" captionTargetPageId="6" captionText="FIGURE 3. Vestiture of pronotum (lateral view) in male a) N. occultus and b) N. obscurus." httpUri="https://zenodo.org/record/169970/files/figure.png" pageId="7" pageNumber="52">Fig. 3</figureCitation>
a). This differs from the short, flattened bristles of male 
<taxonomicName id="C7E085FEFFB80027FBE5FB1B1CAEFB3C" box="[1054,1198,1244,1269]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB80027FBE5FB1B1CAEFB3C" box="[1054,1198,1244,1269]" italics="true" pageId="7" pageNumber="52">N. obscurus</emphasis>
</taxonomicName>
(
<figureCitation id="98DBE2F8FFB80027FB3AFB1B1D0AFB3F" box="[1217,1290,1244,1270]" captionStart="FIGURE 3" captionStartId="6.[264,368,956,980]" captionTargetBox="[264,1323,526,932]" captionTargetId="figure@6.[264,1323,526,932]" captionTargetPageId="6" captionText="FIGURE 3. Vestiture of pronotum (lateral view) in male a) N. occultus and b) N. obscurus." httpUri="https://zenodo.org/record/169970/files/figure.png" pageId="7" pageNumber="52">Fig. 3</figureCitation>
b). Although the dorsal vestiture of females of both species consists of fine, pale­colored setae, that of 
<taxonomicName id="C7E085FEFFB80027FE3CFAEB1A57FA8C" box="[455,599,1324,1349]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="3294226FFFB80027FE3CFAEB1A57FA8C" box="[455,599,1324,1349]" italics="true" pageId="7" pageNumber="52">N. occultus</emphasis>
</taxonomicName>
females is relatively longer and more prominent. The crenulations of the lateral pronotal margins in 
<taxonomicName id="C7E085FEFFB80027FCBFFA931BCBFAA4" box="[836,971,1364,1389]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="3294226FFFB80027FCBFFA931BCBFAA4" box="[836,971,1364,1389]" italics="true" pageId="7" pageNumber="52">N. occultus</emphasis>
</taxonomicName>
are poorly developed, being weak in males and virtually absent in some females. In contrast, the crenulations are stronger and always apparent in 
<emphasis id="3294226FFFB80027FD7DFA631B1AFA74" box="[646,794,1444,1469]" italics="true" pageId="7" pageNumber="52">
<taxonomicName id="C7E085FEFFB80027FD7DFA631B15FA74" box="[646,789,1444,1469]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="obscurus">N. obscurus</taxonomicName>
.
</emphasis>
Both sexes of 
<taxonomicName id="C7E085FEFFB80027FC2BFA631C55FA74" box="[976,1109,1444,1469]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="3294226FFFB80027FC2BFA631C55FA74" box="[976,1109,1444,1469]" italics="true" pageId="7" pageNumber="52">N. occultus</emphasis>
</taxonomicName>
possess relatively short, conical metatibiae that gradually widen towards the apex (
<figureCitation id="98DBE2F8FFB80027FC03FA0B1C40FA2F" box="[1016,1088,1484,1510]" captionStart="FIGURES 5 – 8. 5" captionStartId="9.[264,382,1763,1787]" captionTargetBox="[264,1322,609,1753]" captionTargetId="figure@9.[264,1323,609,1761]" captionTargetPageId="9" captionText="FIGURES 5 – 8. 5. Lateral view of male N. occultus, illustrating diagnostic characters: short, conical metatibia; weakly curved claws; and dorsally curved parameres. 6. Lateral view of elongate male metatibia of N. obscurus. 7. Apex of metatibiae of male N. occultus. 8. Male genitalia of N. occultus (dorsal view), showing rounded apex of median lobe." httpUri="https://zenodo.org/record/169972/files/figure.png" pageId="7" pageNumber="52">Fig. 5</figureCitation>
). The metatibiae of 
<taxonomicName id="C7E085FEFFB80027FEF3FA331996F9C4" box="[264,406,1524,1549]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB80027FEF3FA331996F9C4" box="[264,406,1524,1549]" italics="true" pageId="7" pageNumber="52">N. obscurus</emphasis>
</taxonomicName>
are slender in males (
<figureCitation id="98DBE2F8FFB80027FD64FA331AE9F9C7" box="[671,745,1524,1550]" captionStart="FIGURES 5 – 8. 5" captionStartId="9.[264,382,1763,1787]" captionTargetBox="[264,1322,609,1753]" captionTargetId="figure@9.[264,1323,609,1761]" captionTargetPageId="9" captionText="FIGURES 5 – 8. 5. Lateral view of male N. occultus, illustrating diagnostic characters: short, conical metatibia; weakly curved claws; and dorsally curved parameres. 6. Lateral view of elongate male metatibia of N. obscurus. 7. Apex of metatibiae of male N. occultus. 8. Male genitalia of N. occultus (dorsal view), showing rounded apex of median lobe." httpUri="https://zenodo.org/record/169972/files/figure.png" pageId="7" pageNumber="52">Fig. 6</figureCitation>
), and in females are abruptly expanded after the distal third. In addition, both sexes of 
<taxonomicName id="C7E085FEFFB80027FD34F9DB1B53F9FC" box="[719,851,1564,1589]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="3294226FFFB80027FD34F9DB1B53F9FC" box="[719,851,1564,1589]" italics="true" pageId="7" pageNumber="52">N. occultus</emphasis>
</taxonomicName>
have more prominent, acute mandibles, and the apical protibial teeth are stronger and more acute (
<figureCitation id="98DBE2F8FFB80027FC40F9831C01F997" box="[955,1025,1604,1630]" captionStart="FIGURE 4" captionStartId="8.[264,367,883,907]" captionTargetBox="[536,1051,409,857]" captionTargetId="figure@8.[536,1051,409,857]" captionTargetPageId="8" captionText="FIGURE 4. Protibial dentation (dorsal view) of male: a) N. occultus, b) N. obscurus, and c) N. japonicus." httpUri="https://zenodo.org/record/169971/files/figure.png" pageId="7" pageNumber="52">Fig. 4</figureCitation>
a–b). The median lobe of the male genitalia is rounded (
<figureCitation id="98DBE2F8FFB80027FD88F9AB1ABAF94F" box="[627,698,1644,1670]" captionStart="FIGURES 5 – 8. 5" captionStartId="9.[264,382,1763,1787]" captionTargetBox="[264,1322,609,1753]" captionTargetId="figure@9.[264,1323,609,1761]" captionTargetPageId="9" captionText="FIGURES 5 – 8. 5. Lateral view of male N. occultus, illustrating diagnostic characters: short, conical metatibia; weakly curved claws; and dorsally curved parameres. 6. Lateral view of elongate male metatibia of N. obscurus. 7. Apex of metatibiae of male N. occultus. 8. Male genitalia of N. occultus (dorsal view), showing rounded apex of median lobe." httpUri="https://zenodo.org/record/169972/files/figure.png" pageId="7" pageNumber="52">Fig. 8</figureCitation>
) and not acuminate as in males of 
<taxonomicName id="C7E085FEFFB80027FB99F9AB1CEFF94C" box="[1122,1263,1644,1669]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB80027FB99F9AB1CEFF94C" box="[1122,1263,1644,1669]" italics="true" pageId="7" pageNumber="52">N. obscurus</emphasis>
</taxonomicName>
, and the parameres are more strongly curved dorsally.
</paragraph>
<paragraph id="005FFE7DFFB80028FEC3F97B19DBFE97" blockId="7.[264,1324,284,1870]" lastBlockId="8.[264,1323,284,350]" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52">
Both Nearctic species differ from 
<emphasis id="3294226FFFB80027FD29F97B1B6FF91C" box="[722,879,1724,1749]" italics="true" pageId="7" pageNumber="52">
<taxonomicName id="C7E085FEFFB80027FD29F97B1B6BF91C" box="[722,875,1724,1749]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="japonicus">N. japonicus</taxonomicName>
,
</emphasis>
which has an evenly rounded lateral margin of the pronotum (not angulate as in the Nearctic species) with stronger tooth­like crenulations. In addition, 
<taxonomicName id="C7E085FEFFB80027FDC2F8CB1ACDF8EC" box="[569,717,1804,1829]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="3294226FFFB80027FDC2F8CB1ACDF8EC" box="[569,717,1804,1829]" italics="true" pageId="7" pageNumber="52">N. japonicus</emphasis>
</taxonomicName>
has the anterior margin of the clypeus subtriangular and not rounded, the claws are relatively short and subequal in length to tarsomeres 3–4 together, and males have a bidentate protibial apex (
<figureCitation id="98DBE2F8FFB70028FC84FEDB1BC9FEFF" box="[895,969,284,310]" captionStart="FIGURE 4" captionStartId="8.[264,367,883,907]" captionTargetBox="[536,1051,409,857]" captionTargetId="figure@8.[536,1051,409,857]" captionTargetPageId="8" captionText="FIGURE 4. Protibial dentation (dorsal view) of male: a) N. occultus, b) N. obscurus, and c) N. japonicus." httpUri="https://zenodo.org/record/169971/files/figure.png" pageId="8" pageNumber="53">Fig. 4</figureCitation>
c), rather than an apex with a single large tooth.
</paragraph>
<caption id="549FAEF5FFB70028FEF3FCB41977FC67" httpUri="https://zenodo.org/record/169971/files/figure.png" pageId="8" pageNumber="53" targetBox="[536,1051,409,857]" targetPageId="8">
<paragraph id="005FFE7DFFB70028FEF3FCB41977FC67" blockId="8.[264,1323,883,942]" pageId="8" pageNumber="53">
<emphasis id="3294226FFFB70028FEF3FCB4198CFC42" bold="true" box="[264,396,883,907]" pageId="8" pageNumber="53">FIGURE 4.</emphasis>
Protibial dentation (dorsal view) of male: a) 
<emphasis id="3294226FFFB70028FC71FCB31C0BFC42" box="[906,1035,884,907]" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="C7E085FEFFB70028FC71FCB31C07FC42" box="[906,1031,884,907]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="occultus">N. occultus</taxonomicName>
,
</emphasis>
b) 
<emphasis id="3294226FFFB70028FBC8FCB31CBCFC42" box="[1075,1212,884,907]" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="C7E085FEFFB70028FBC8FCB31CB8FC42" box="[1075,1208,884,907]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="obscurus">N. obscurus</taxonomicName>
,
</emphasis>
and c) 
<taxonomicName id="C7E085FEFFB70028FAE9FCB31973FC67" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="japonicus">N. japonicus</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="005FFE7DFFB70028FEC3FC261C09FB82" blockId="8.[264,1323,993,1459]" pageId="8" pageNumber="53">
Species of 
<taxonomicName id="C7E085FEFFB70028FE43FC261A18FC33" box="[440,536,993,1018]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="3294226FFFB70028FE43FC261A18FC33" box="[440,536,993,1018]" italics="true" pageId="8" pageNumber="53">Nicagus</emphasis>
</taxonomicName>
can be immediately separated from members of the 
<taxonomicName id="C7E085FEFFB70028FB85FC261CEAFC32" box="[1150,1258,993,1019]" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
, with which they might be confused, by the presence in 
<taxonomicName id="C7E085FEFFB70028FCA4FBCE1BBFFBEB" box="[863,959,1033,1058]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="3294226FFFB70028FCA4FBCE1BBFFBEB" box="[863,959,1033,1058]" italics="true" pageId="8" pageNumber="53">Nicagus</emphasis>
</taxonomicName>
of a bisetose arolium between the claws. An arolium is not present in members of the 
<taxonomicName id="C7E085FEFFB70028FC63FBF61C05FB82" box="[920,1029,1073,1099]" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="48FAADF6FFB70028FEC3FB9E19DFFADA" pageId="8" pageNumber="53" type="etymology">
<paragraph id="005FFE7DFFB70028FEC3FB9E19DFFADA" blockId="8.[264,1323,993,1459]" pageId="8" pageNumber="53">
<emphasis id="3294226FFFB70028FEC3FB9E19C4FBBA" bold="true" box="[312,452,1113,1139]" pageId="8" pageNumber="53">Etymology.</emphasis>
The specific epithet is derived from the Latin “ 
<emphasis id="3294226FFFB70028FBC5FB9E1CB8FBBB" box="[1086,1208,1113,1138]" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="C7E085FEFFB70028FBC5FB9E1CA7FBBB" box="[1086,1191,1113,1138]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="occultus">occultus</taxonomicName>
”,
</emphasis>
meaning hidden, concealed, or secret, in reference to the elusive nature of the species that has evaded discovery by entomologists for decades in an ostensibly well­studied locality. In addition, many males were observed to remain hidden under the sand with only their antennae exposed.
</paragraph>
</subSubSection>
<subSubSection id="48FAADF6FFB70028FEC3FAE61C52FA42" pageId="8" pageNumber="53" type="distribution">
<paragraph id="005FFE7DFFB70028FEC3FAE61C4AFAF2" blockId="8.[264,1323,993,1459]" box="[312,1098,1313,1339]" pageId="8" pageNumber="53">
<emphasis id="3294226FFFB70028FEC3FAE619D8FAF2" bold="true" box="[312,472,1313,1339]" pageId="8" pageNumber="53">Distribution.</emphasis>
(
<figureCitation id="98DBE2F8FFB70028FE14FAE61A37FAF2" box="[495,567,1313,1339]" captionStart="FIGURE 9" captionStartId="10.[264,368,1637,1661]" captionTargetBox="[264,1323,649,1611]" captionTargetId="figure@10.[264,1323,649,1611]" captionTargetPageId="10" captionText="FIGURE 9. Distribution of Nearctic species of Nicagus, indicated by triangles (N. occultus), or circles (N. obscurus). An open circle indicates a state record without specific locality data." httpUri="https://zenodo.org/record/169973/files/figure.png" pageId="8" pageNumber="53">Fig. 9</figureCitation>
). 
<collectingCountry id="78F7BEEDFFB70028FDAEFAE61AF5FAF2" box="[597,757,1313,1339]" name="United States of America" pageId="8" pageNumber="53">United States</collectingCountry>
, Texas, Monahans Sandhills.
</paragraph>
<paragraph id="005FFE7DFFB70028FEC3FA8E1C52FA42" blockId="8.[264,1323,993,1459]" pageId="8" pageNumber="53">
<emphasis id="3294226FFFB70028FEC3FA8E1B74FAAA" bold="true" box="[312,884,1353,1379]" pageId="8" pageNumber="53">
Locality Data. 
<collectingCountry id="78F7BEEDFFB70028FDF3FA8E1AF7FAAA" box="[520,759,1353,1379]" name="United States of America" pageId="8" pageNumber="53">UNITED STATES</collectingCountry>
: TEXAS:
</emphasis>
WARD (113): Monahans Sandhills State Park (
<quantity id="C7185398FFB70028FE74FAB619C5FA42" box="[399,453,1393,1419]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="8" pageNumber="53" unit="mi" value="4.0">4 mi</quantity>
NE of Monahans); WINKLER (4): 
<quantity id="C7185398FFB70028FC96FAB61BA2FA42" box="[877,930,1393,1419]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="8" pageNumber="53" unit="mi" value="8.0">8 mi</quantity>
NE of Kermit.
</paragraph>
</subSubSection>
<subSubSection id="48FAADF6FFB70028FEC3FA5E1B46F9E2" pageId="8" pageNumber="53" type="description">
<paragraph id="005FFE7DFFB70028FEC3FA5E1AF4FA7A" blockId="8.[264,1323,993,1459]" box="[312,756,1433,1459]" pageId="8" pageNumber="53">
<emphasis id="3294226FFFB70028FEC3FA5E19FBFA7A" bold="true" box="[312,507,1433,1459]" pageId="8" pageNumber="53">Temporal Data.</emphasis>
May (28), June (89).
</paragraph>
<paragraph id="005FFE7DFFB70028FEF3F9D61B46F9E2" blockId="8.[264,838,1553,1579]" box="[264,838,1553,1579]" pageId="8" pageNumber="53">
<taxonomicName id="C7E085FEFFB70028FEF3F9D61A66F9E2" authority="LeConte" authorityName="LeConte" box="[264,614,1553,1579]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB70028FEF3F9D61A66F9E2" bold="true" box="[264,614,1553,1579]" pageId="8" pageNumber="53">
<emphasis id="3294226FFFB70028FEF3F9D619DFF9E2" bold="true" box="[264,479,1553,1579]" italics="true" pageId="8" pageNumber="53">Nicagus obscurus</emphasis>
(LeConte)
</emphasis>
</taxonomicName>
(
<figureCitation id="98DBE2F8FFB70028FD8DF9D61AC6F9E2" box="[630,710,1553,1579]" captionStart="FIGURE 3" captionStartId="6.[264,368,956,980]" captionTargetBox="[264,1323,526,932]" captionTargetId="figure@6.[264,1323,526,932]" captionTargetPageId="6" captionText="FIGURE 3. Vestiture of pronotum (lateral view) in male a) N. occultus and b) N. obscurus." httpUri="https://zenodo.org/record/169970/files/figure.png" pageId="8" pageNumber="53">Figs. 3</figureCitation>
b, 4b, 6, 9)
</paragraph>
</subSubSection>
<subSubSection id="48FAADF6FFB70028FEF3F9A51B3DF9B1" box="[264,829,1633,1657]" pageId="8" pageNumber="53" type="reference_group">
<paragraph id="005FFE7DFFB70028FEF3F9A51B3DF9B1" blockId="8.[264,829,1633,1657]" box="[264,829,1633,1657]" pageId="8" pageNumber="53">
<treatmentCitationGroup id="20F0D953FFB70028FEF3F9A51B3DF9B1" box="[264,829,1633,1657]" pageId="8" pageNumber="53">
<taxonomicName id="C7E085FEFFB70028FEF3F9A51B38F9B1" ID-CoL="73XS3" authority="(LeConte, 1847: 86) (Ochodaeus)" authorityName="LeConte" authorityPageNumber="86" authorityYear="1847" baseAuthorityName="LeConte" baseAuthorityPageNumber="86" baseAuthorityYear="1847" box="[264,824,1633,1657]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB70028FEF3F9A519C6F9B0" box="[264,454,1634,1657]" italics="true" pageId="8" pageNumber="53">Nicagus obscurus</emphasis>
(
<treatmentCitation id="8141D86CFFB70028FE2FF9A61A9FF9B1" author="LeConte" box="[468,671,1633,1656]" page="86" pageId="8" pageNumber="53" year="1847">
<bibRefCitation id="6471838CFFB70028FE2FF9A61A9FF9B1" author="LeConte" box="[468,671,1633,1656]" pageId="8" pageNumber="59" refString="LeConte, J. L. (1847) Fragmenta entomologica. Journal of the Academy of Natural Sciences of Philadelphia, series 2, 1, 71 - 93." type="journal article" year="1847">LeConte, 1847: 86</bibRefCitation>
</treatmentCitation>
) (
<emphasis id="3294226FFFB70028FD4EF9A51B2DF9B0" box="[693,813,1634,1657]" italics="true" pageId="8" pageNumber="53">Ochodaeus</emphasis>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="48FAADF6FFB70029FEF3F96E1ACBFDEC" lastPageId="9" lastPageNumber="54" pageId="8" pageNumber="53" type="diagnosis">
<paragraph id="005FFE7DFFB70029FEF3F96E1ACBFDEC" blockId="8.[264,1323,1705,1891]" lastBlockId="9.[264,1323,284,550]" lastPageId="9" lastPageNumber="54" pageId="8" pageNumber="53">
<emphasis id="3294226FFFB70028FEF3F96E1988F90A" bold="true" box="[264,392,1705,1731]" pageId="8" pageNumber="53">Diagnosis.</emphasis>
This species possesses several characters intermediate between 
<taxonomicName id="C7E085FEFFB70028FB6FF96E1D2BF90B" box="[1172,1323,1705,1730]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="3294226FFFB70028FB6FF96E1D2BF90B" box="[1172,1323,1705,1730]" italics="true" pageId="8" pageNumber="53">N. japonicus</emphasis>
</taxonomicName>
and 
<emphasis id="3294226FFFB70028FEC7F91619CAF923" box="[316,458,1745,1770]" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="C7E085FEFFB70028FEC7F91619C5F923" box="[316,453,1745,1770]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="occultus">N. occultus</taxonomicName>
.
</emphasis>
In males, the lateral margins of the pronotum are moderately crenulate and subangulate; not as strongly angulate as in 
<emphasis id="3294226FFFB70028FCD6F93E1BB8F8DB" box="[813,952,1785,1810]" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="C7E085FEFFB70028FCD6F93E1BB3F8DB" box="[813,947,1785,1810]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="occultus">N. occultus</taxonomicName>
,
</emphasis>
nor as broadly rounded as in 
<taxonomicName id="C7E085FEFFB70028FAEBF93E197AF8F3" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="3294226FFFB70028FAEBF93E197AF8F3" italics="true" pageId="8" pageNumber="53">N. japonicus</emphasis>
</taxonomicName>
. The apical protibial teeth (
<figureCitation id="98DBE2F8FFB70028FD34F8E61B18F8F2" box="[719,792,1825,1851]" captionStart="FIGURE 4" captionStartId="8.[264,367,883,907]" captionTargetBox="[536,1051,409,857]" captionTargetId="figure@8.[536,1051,409,857]" captionTargetPageId="8" captionText="FIGURE 4. Protibial dentation (dorsal view) of male: a) N. occultus, b) N. obscurus, and c) N. japonicus." httpUri="https://zenodo.org/record/169971/files/figure.png" pageId="8" pageNumber="53">Fig. 4</figureCitation>
b) are smaller and more rounded than in 
<emphasis id="3294226FFFB70028FAEBF8E61971F8AB" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="C7E085FEFFB70028FAEBF8E6196CF8AB" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="occultus">N. occultus</taxonomicName>
,
</emphasis>
but more strongly developed than in 
<emphasis id="3294226FFFB70028FCABF88E1BF2F8AB" box="[848,1010,1865,1890]" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="C7E085FEFFB70028FCABF88E1BEEF8AB" box="[848,1006,1865,1890]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="japonicus">N. japonicus</taxonomicName>
.
</emphasis>
However, the explanate lateral margins of the pronotum are more strongly developed in 
<taxonomicName id="C7E085FEFFB60029FC00FEDB1C88FEFC" box="[1019,1160,284,309]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB60029FC00FEDB1C88FEFC" box="[1019,1160,284,309]" italics="true" pageId="9" pageNumber="54">N. obscurus</emphasis>
</taxonomicName>
than in either of the remaining species. In male 
<taxonomicName id="C7E085FEFFB60029FD59FE831B30FE94" box="[674,816,324,349]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB60029FD59FE831B30FE94" box="[674,816,324,349]" italics="true" pageId="9" pageNumber="54">N. obscurus</emphasis>
</taxonomicName>
, the antennal club segments are relatively longer, and the dorsum bears short, flattened bristles (
<figureCitation id="98DBE2F8FFB60029FC56FEAB1BF9FE4F" box="[941,1017,364,390]" captionStart="FIGURE 3" captionStartId="6.[264,368,956,980]" captionTargetBox="[264,1323,526,932]" captionTargetId="figure@6.[264,1323,526,932]" captionTargetPageId="6" captionText="FIGURE 3. Vestiture of pronotum (lateral view) in male a) N. occultus and b) N. obscurus." httpUri="https://zenodo.org/record/169970/files/figure.png" pageId="9" pageNumber="54">Fig. 3</figureCitation>
b). The metatibiae of 
<taxonomicName id="C7E085FEFFB60029FAEBFEAB1972FE64" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB60029FAEBFEAB1972FE64" italics="true" pageId="9" pageNumber="54">N. obscurus</emphasis>
</taxonomicName>
are elongate in males (
<figureCitation id="98DBE2F8FFB60029FD6DFE531AE0FE67" box="[662,736,404,430]" captionStart="FIGURES 5 – 8. 5" captionStartId="9.[264,382,1763,1787]" captionTargetBox="[264,1322,609,1753]" captionTargetId="figure@9.[264,1323,609,1761]" captionTargetPageId="9" captionText="FIGURES 5 – 8. 5. Lateral view of male N. occultus, illustrating diagnostic characters: short, conical metatibia; weakly curved claws; and dorsally curved parameres. 6. Lateral view of elongate male metatibia of N. obscurus. 7. Apex of metatibiae of male N. occultus. 8. Male genitalia of N. occultus (dorsal view), showing rounded apex of median lobe." httpUri="https://zenodo.org/record/169972/files/figure.png" pageId="9" pageNumber="54">Fig. 6</figureCitation>
) and straight internally (in dorsal view), and in females are abruptly expanded after the distal third (lateral view). The median lobe of the male genitalia has a small acuminate process at the apex (
<figureCitation id="98DBE2F8FFB60029FC3EFE231C0EFE37" box="[965,1038,484,510]" captionStart="FIGURES 5 – 8. 5" captionStartId="9.[264,382,1763,1787]" captionTargetBox="[264,1322,609,1753]" captionTargetId="figure@9.[264,1323,609,1761]" captionTargetPageId="9" captionText="FIGURES 5 – 8. 5. Lateral view of male N. occultus, illustrating diagnostic characters: short, conical metatibia; weakly curved claws; and dorsally curved parameres. 6. Lateral view of elongate male metatibia of N. obscurus. 7. Apex of metatibiae of male N. occultus. 8. Male genitalia of N. occultus (dorsal view), showing rounded apex of median lobe." httpUri="https://zenodo.org/record/169972/files/figure.png" pageId="9" pageNumber="54">Fig. 8</figureCitation>
) not a broadly rounded apex as in 
<taxonomicName id="C7E085FEFFB60029FE7FFDCB1A09FDEC" box="[388,521,524,549]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="3294226FFFB60029FE7FFDCB1A09FDEC" box="[388,521,524,549]" italics="true" pageId="9" pageNumber="54">N. occultus</emphasis>
</taxonomicName>
or 
<emphasis id="3294226FFFB60029FDD4FDCB1ACBFDEC" box="[559,715,524,549]" italics="true" pageId="9" pageNumber="54">
<taxonomicName id="C7E085FEFFB60029FDD4FDCB1AC7FDEC" box="[559,711,524,549]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="japonicus">N. japonicus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<caption id="549FAEF5FFB60029FEF3F9241BA2F8AA" httpUri="https://zenodo.org/record/169972/files/figure.png" pageId="9" pageNumber="54" targetBox="[264,1322,609,1753]" targetPageId="9">
<paragraph id="005FFE7DFFB60029FEF3F9241BA2F8AA" blockId="9.[264,1323,1763,1891]" pageId="9" pageNumber="54">
<emphasis id="3294226FFFB60029FEF3F92419E4F932" bold="true" box="[264,484,1763,1787]" pageId="9" pageNumber="54">FIGURES 5–8. 5.</emphasis>
Lateral view of male 
<emphasis id="3294226FFFB60029FD13F9231B6BF932" box="[744,875,1764,1787]" italics="true" pageId="9" pageNumber="54">
<taxonomicName id="C7E085FEFFB60029FD13F9231B68F932" box="[744,872,1764,1787]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="occultus">N. occultus</taxonomicName>
,
</emphasis>
illustrating diagnostic characters: short, conical metatibia; weakly curved claws; and dorsally curved parameres. 
<emphasis id="3294226FFFB60029FBF2F8C11C1DF8D7" bold="true" box="[1033,1053,1798,1822]" pageId="9" pageNumber="54">6.</emphasis>
Lateral view of elongate male metatibia of 
<emphasis id="3294226FFFB60029FE32F8ED1A50F888" box="[457,592,1834,1857]" italics="true" pageId="9" pageNumber="54">
<taxonomicName id="C7E085FEFFB60029FE32F8ED1A4CF888" box="[457,588,1834,1857]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="obscurus">N. obscurus</taxonomicName>
.
</emphasis>
<emphasis id="3294226FFFB60029FDA6F8EE1A70F888" bold="true" box="[605,624,1833,1857]" pageId="9" pageNumber="54">7.</emphasis>
Apex of metatibiae of male 
<emphasis id="3294226FFFB60029FC5FF8ED1C24F888" box="[932,1060,1834,1857]" italics="true" pageId="9" pageNumber="54">
<taxonomicName id="C7E085FEFFB60029FC5FF8ED1C20F888" box="[932,1056,1834,1857]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="occultus">N. occultus</taxonomicName>
.
</emphasis>
<emphasis id="3294226FFFB60029FBC9F8EE1C46F888" bold="true" box="[1074,1094,1833,1857]" pageId="9" pageNumber="54">8.</emphasis>
Male genitalia of 
<taxonomicName id="C7E085FEFFB60029FAE9F8ED1961F8AA" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="3294226FFFB60029FAE9F8ED1961F8AA" italics="true" pageId="9" pageNumber="54">N. occultus</emphasis>
</taxonomicName>
(dorsal view), showing rounded apex of median lobe.
</paragraph>
</caption>
<subSubSection id="48FAADF6FFB5002AFEC3FEDC1954FDEF" pageId="10" pageNumber="55" type="distribution">
<paragraph id="005FFE7DFFB5002AFEC3FEDC1B5DFEFF" blockId="10.[264,1323,283,550]" box="[312,861,283,310]" pageId="10" pageNumber="55">
<emphasis id="3294226FFFB5002AFEC3FEDC19D8FEFC" bold="true" box="[312,472,283,309]" pageId="10" pageNumber="55">Distribution.</emphasis>
Eastern North 
<collectingCountry id="78F7BEEDFFB5002AFD69FEDB1AF7FEFF" box="[658,759,284,310]" name="United States of America" pageId="10" pageNumber="55">America</collectingCountry>
(
<figureCitation id="98DBE2F8FFB5002AFCFCFEDB1B4FFEFF" box="[775,847,284,310]" captionStart="FIGURE 9" captionStartId="10.[264,368,1637,1661]" captionTargetBox="[264,1323,649,1611]" captionTargetId="figure@10.[264,1323,649,1611]" captionTargetPageId="10" captionText="FIGURE 9. Distribution of Nearctic species of Nicagus, indicated by triangles (N. occultus), or circles (N. obscurus). An open circle indicates a state record without specific locality data." httpUri="https://zenodo.org/record/169973/files/figure.png" pageId="10" pageNumber="55">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="005FFE7DFFB5002AFEC3FE841954FDEF" blockId="10.[264,1323,283,550]" pageId="10" pageNumber="55">
<emphasis id="3294226FFFB5002AFEC3FE8419E9FE94" bold="true" box="[312,489,323,349]" pageId="10" pageNumber="55">Locality Data.</emphasis>
<specimenCount id="16E635F4FFB5002AFE03FE831AA5FE97" box="[504,677,324,350]" pageId="10" pageNumber="55" type="generic">338 specimens</specimenCount>
examined (
<specimenCount id="16E635F4FFB5002AFCD5FE831B9FFE97" box="[814,927,324,350]" pageId="10" pageNumber="55" type="male">294 male</specimenCount>
, 
<specimenCount id="16E635F4FFB5002AFC50FE831C24FE97" box="[939,1060,324,350]" pageId="10" pageNumber="55" type="female">44 female</specimenCount>
). 
<collectingCountry id="78F7BEEDFFB5002AFEF3FEAB198AFE4F" box="[264,394,364,390]" name="Canada" pageId="10" pageNumber="55">CANADA</collectingCountry>
: 
<emphasis id="3294226FFFB5002AFE5AFEAC1A2BFE4C" bold="true" box="[417,555,363,389]" pageId="10" pageNumber="55">ONTARIO</emphasis>
: GREATER TORONTO AREA: Toronto (CMNC, FMNH, UMMZ, USNM); LAMBTON: Grand Bend (CNC, FSCA); MANITOULIN: Billings (CNC); NIAGARA: Grimsby (MSUC); 
<collectingCountry id="78F7BEEDFFB5002AFCEDFE7B1BA5FE1F" box="[790,933,444,470]" name="Norfolk Island" pageId="10" pageNumber="55">NORFOLK</collectingCountry>
: Long Point (CMNC, CNC). 
<emphasis id="3294226FFFB5002AFEF3FE24198DFE34" bold="true" box="[264,397,483,509]" pageId="10" pageNumber="55">QUEBEC:</emphasis>
BROME­MISSISQUOI: Brome (CNC); no locality (
<bibRefCitation id="6471838CFFB5002AFBCCFE231948FDEF" author="Katovich" pageId="10" pageNumber="59" refString="Katovich, K. &amp; Kriska, N. L. (2002) Description of the larva of Nicagus obscurus (LeConte) (Coleoptera: Lucanidae: Nicaginae), with comments on its position in Lucanidae and notes on the larval and adult habitat. Coleopterists Bulletin, 56 (2), 253 - 258." type="journal article" year="2002">Katovich &amp; Kriska 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="549FAEF5FFB5002AFEF3F9A21CD3F96A" httpUri="https://zenodo.org/record/169973/files/figure.png" pageId="10" pageNumber="55" targetBox="[264,1323,649,1611]" targetPageId="10">
<paragraph id="005FFE7DFFB5002AFEF3F9A21CD3F96A" blockId="10.[264,1322,1637,1700]" pageId="10" pageNumber="55">
<emphasis id="3294226FFFB5002AFEF3F9A2198DF9B4" bold="true" box="[264,397,1637,1661]" pageId="10" pageNumber="55">FIGURE 9.</emphasis>
Distribution of Nearctic species of 
<emphasis id="3294226FFFB5002AFCE4F9A11B7DF9B4" box="[799,893,1638,1661]" italics="true" pageId="10" pageNumber="55">
<taxonomicName id="C7E085FEFFB5002AFCE4F9A11B79F9B4" box="[799,889,1638,1661]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="genus">Nicagus</taxonomicName>
,
</emphasis>
indicated by triangles (
<emphasis id="3294226FFFB5002AFB79F9A11D0BF9B4" box="[1154,1291,1638,1661]" italics="true" pageId="10" pageNumber="55">
<taxonomicName id="C7E085FEFFB5002AFB79F9A11CFFF9B4" box="[1154,1279,1638,1661]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="species" species="occultus">N. occultus</taxonomicName>
),
</emphasis>
or circles (
<taxonomicName id="C7E085FEFFB5002AFEA6F94A19DCF96D" box="[349,476,1677,1700]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="3294226FFFB5002AFEA6F94A19DCF96D" box="[349,476,1677,1700]" italics="true" pageId="10" pageNumber="55">N. obscurus</emphasis>
</taxonomicName>
). An open circle indicates a state record without specific locality data.
</paragraph>
</caption>
<subSubSection id="48FAADF6FFB5002BFEC3F8C51AD2FA8F" lastPageId="11" lastPageNumber="56" pageId="10" pageNumber="55" type="description">
<paragraph id="005FFE7DFFB5002BFEC3F8C51AD2FA8F" blockId="10.[264,1323,1794,1901]" lastBlockId="11.[264,1323,284,1350]" lastPageId="11" lastPageNumber="56" pageId="10" pageNumber="55">
<emphasis id="3294226FFFB5002AFEC3F8C51A2CF8D5" bold="true" box="[312,556,1794,1820]" pageId="10" pageNumber="55">
<collectingCountry id="78F7BEEDFFB5002AFEC3F8C51A2CF8D5" box="[312,556,1794,1820]" name="United States of America" pageId="10" pageNumber="55">UNITED STATES</collectingCountry>
</emphasis>
: 
<collectingCountry id="78F7BEEDFFB5002AFDB0F8C51AD9F8D5" box="[587,729,1794,1820]" name="Georgia" pageId="10" pageNumber="55">
<emphasis id="3294226FFFB5002AFDB0F8C51AD9F8D5" bold="true" box="[587,729,1794,1820]" pageId="10" pageNumber="55">GEORGIA</emphasis>
</collectingCountry>
: CLARKE: no locality (UGCA); FULTON/ DEKALB: Atlanta (FMNH, USNM); WHITE: Helen (UGCA). 
<emphasis id="3294226FFFB5002AFBE9F8ED1C97F88D" bold="true" box="[1042,1175,1834,1860]" pageId="10" pageNumber="55">ILLINOIS</emphasis>
: no locality (FMNH, MSUC). 
<emphasis id="3294226FFFB5002AFE08F8951A74F8A5" bold="true" box="[499,628,1874,1900]" pageId="10" pageNumber="55">INDIANA</emphasis>
: HENDRICKS: Stilesville (USNM); PARKE: Shades SP (FMNH, MJPC); TIPPECANOE: Wildcat Creek (FSCA), no locality (FMNH, MJPC, MSUC, UGCA, USNM); WARREN: no locality (FMNH). 
<emphasis id="3294226FFFB4002BFC3BFE841C11FE94" bold="true" box="[960,1041,323,349]" pageId="11" pageNumber="56">IOWA</emphasis>
: JOHNSON: Iowa City (USNM); LEE: Shimek SF (MJPC); LINN: Indian Creek Nature Center (MJPC); STORY: Ames (CMNC, FMNH). 
<emphasis id="3294226FFFB4002BFD87FE541B22FE64" bold="true" box="[636,802,403,429]" pageId="11" pageNumber="56">KENTUCKY</emphasis>
: LIVINGSTON: no locality (USNM). 
<emphasis id="3294226FFFB4002BFEF3FE7C19B5FE1C" bold="true" box="[264,437,443,469]" pageId="11" pageNumber="56">MARYLAND</emphasis>
: ALLEGANY: Green Ridge SF, Oldtown (CMNH); ANNE ARUNDEL: Crofton (CMNH); MONTGOMERY: Plummers Island (USNM); PRINCE GEORGE’S: Berwyn (USNM), College Park (CMNC, USNM), Laurel (CMNH), Riverdale (USNM). 
<emphasis id="3294226FFFB4002BFEF3FDF41A14FD84" bold="true" box="[264,532,563,589]" pageId="11" pageNumber="56">MASSACHUSETTS:</emphasis>
MIDDLESEX: Lowell (CNC); no locality (FMNH). 
<emphasis id="3294226FFFB4002BFEF3FD9C19A7FDBC" bold="true" box="[264,423,603,629]" pageId="11" pageNumber="56">MICHIGAN</emphasis>
: BERRIEN: Warren Dunes (FMNH); OAKLAND: no locality (MSUC, UMMZ); ST. CLAIR: Port Huron (USNM); VAN BUREN: South Haven (CUAC); WAYNE: Detroit (USNM). 
<emphasis id="3294226FFFB4002BFD84FD6B1B32FD0F" bold="true" box="[639,818,684,710]" pageId="11" pageNumber="56">MINNESOTA</emphasis>
: ST. LOUIS: Duluth (INHS). 
<emphasis id="3294226FFFB4002BFB1FFD6B19BCFD27" bold="true" pageId="11" pageNumber="56">NEW HAMPSHIRE</emphasis>
: GRAFTON: Plymouth (CNC, FMNH, FSCA, UMMZ). 
<emphasis id="3294226FFFB4002BFB9CFD131D22FD27" bold="true" box="[1127,1314,724,750]" pageId="11" pageNumber="56">
NEW 
<collectingCountry id="78F7BEEDFFB4002BFB4EFD131D22FD27" box="[1205,1314,724,750]" name="Jersey" pageId="11" pageNumber="56">JERSEY</collectingCountry>
</emphasis>
: ESSEX: Newark (USNM). 
<emphasis id="3294226FFFB4002BFD91FD3B1B15FCDF" bold="true" box="[618,789,764,790]" pageId="11" pageNumber="56">NEW YORK</emphasis>
: ERIE: Buffalo (INHS); ONEIDA: Sylvan Beach (CNC); OSWEGO: Selkirk Beach (FMNH, MSUC, UNSM); SUFFOLK: Montauk (FMNH, FSCA), Napeague (INHS), Riverhead (CMNC), Wading River (CMNC, FMNH, FSCA, USNM). 
<emphasis id="3294226FFFB4002BFE35FCB31ADFFC47" bold="true" box="[462,735,884,910]" pageId="11" pageNumber="56">NORTH CAROLINA</emphasis>
: 
<collectingCountry id="78F7BEEDFFB4002BFD14FCB31B49FC47" box="[751,841,884,910]" name="Wake Island" pageId="11" pageNumber="56">WAKE</collectingCountry>
: Raleigh (CMNC). 
<emphasis id="3294226FFFB4002BFBD4FCB31C7FFC47" bold="true" box="[1071,1151,884,910]" pageId="11" pageNumber="56">OHIO</emphasis>
: FAIRFIELD: Barnebey Center (CMNH, FSCA), Revenge (CMNH, MJPC); HOCKING: no locality (CMNC, FSCA, UNSM); PIKE: Liberty (KDKC, RAAC), Jackson Lake (CMNH); RICHLAND: Shelby (RAAC); VINTON: Lake Hope SP (CMNH, MLJC). 
<emphasis id="3294226FFFB4002BFB3FFC2B19A7FBE7" bold="true" pageId="11" pageNumber="56">SOUTH CAROLINA</emphasis>
: OCONEE: Salem (CUAC); PICKENS: Clemson (FMNH, USNM); SPARTANBURG: Spartanburg (CUAC). 
<emphasis id="3294226FFFB4002BFCFEFBFB1BE7FB9F" bold="true" box="[773,999,1084,1110]" pageId="11" pageNumber="56">PENNSYLVANIA</emphasis>
: ALLEGHENY: Guyasuta Run (CMNH), no locality (USNM); LYCOMING: Loyalsock Creek (CMNH); NORTHUMBERLAND: Milton, Milton SP (CMNH); WESTMORELAND: Jeannette (CMNH). 
<emphasis id="3294226FFFB4002BFE65FB731A2BFB07" bold="true" box="[414,555,1204,1230]" pageId="11" pageNumber="56">VIRGINIA</emphasis>
: FAIRFAX: Black Pond, Dead Run, Great Falls (USNM), Springfield (CMNH). 
<emphasis id="3294226FFFB4002BFDD2FB1B1B42FB3F" bold="true" box="[553,834,1244,1270]" pageId="11" pageNumber="56">WASHINGTON, D.C.</emphasis>
: Rock Creek (USNM). 
<emphasis id="3294226FFFB4002BFB8DFB1B1D2AFB3F" bold="true" box="[1142,1322,1244,1270]" pageId="11" pageNumber="56">WISCONSIN:</emphasis>
BAYFIELD, no locality (USNM), Quarry Point (
<bibRefCitation id="6471838CFFB4002BFCA9FAC31C7FFAD7" author="Katovich" box="[850,1151,1284,1310]" pageId="11" pageNumber="59" refString="Katovich, K. &amp; Kriska, N. L. (2002) Description of the larva of Nicagus obscurus (LeConte) (Coleoptera: Lucanidae: Nicaginae), with comments on its position in Lucanidae and notes on the larval and adult habitat. Coleopterists Bulletin, 56 (2), 253 - 258." type="journal article" year="2002">Katovich &amp; Kriska 2002</bibRefCitation>
); DOUGLAS: Brule Point (
<bibRefCitation id="6471838CFFB4002BFE66FAEB1AC5FA8F" author="Katovich" box="[413,709,1324,1350]" pageId="11" pageNumber="59" refString="Katovich, K. &amp; Kriska, N. L. (2002) Description of the larva of Nicagus obscurus (LeConte) (Coleoptera: Lucanidae: Nicaginae), with comments on its position in Lucanidae and notes on the larval and adult habitat. Coleopterists Bulletin, 56 (2), 253 - 258." type="journal article" year="2002">Katovich &amp; Kriska 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>