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<document id="BCD57ABD4006D6C984F0A0D882AC6523" ID-DOI="10.11646/zootaxa.5374.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10148159" ID-ZooBank="072341AB-7693-4DF3-9286-D69D0BBD8FAD" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700206341287" checkinUser="plazi" docAuthor="MACFARLAN, DONALD ALEXANDER BANKIER" docDate="2023" docId="073F87ADFFA95A75FF4BFE23FEC5FAB8" docLanguage="en" docName="zootaxa.5374.1.1.pdf" docOrigin="Zootaxa 5374 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5374.1.1/52274" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tibetothyris johnstoni MACFARLAN 2023, new species" docType="treatment" docVersion="5" lastPageNumber="9" masterDocId="FB06FFD5FFAF5A7DFFDCFFF9FFCCFFF6" masterDocTitle="Otapirian (Rhaetian) Terebratulida (Brachiopoda) of Zealandia" masterLastPageNumber="34" masterPageNumber="1" pageNumber="7" updateTime="1706308076791" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="2D269A505F6A28BCF0C6994E379403F4">Otapirian (Rhaetian) Terebratulida (Brachiopoda) of Zealandia</mods:title>
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<treatment id="073F87ADFFA95A75FF4BFE23FEC5FAB8" ID-DOI="http://doi.org/10.5281/zenodo.10168046" ID-Zenodo-Dep="10168046" LSID="urn:lsid:plazi:treatment:073F87ADFFA95A75FF4BFE23FEC5FAB8" httpUri="http://treatment.plazi.org/id/073F87ADFFA95A75FF4BFE23FEC5FAB8" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="C78C6530FFA95A7BFF4BFE23FDF4FE02" box="[151,568,474,501]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8F2936BBFFA95A7BFF4BFE23FDF4FE02" blockId="6.[151,568,474,537]" box="[151,568,474,501]" pageId="6" pageNumber="7">
<heading id="D46181D7FFA95A7BFF4BFE23FDF4FE02" bold="true" box="[151,568,474,501]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="BDE2EAA9FFA95A7BFF4BFE23FDF4FE02" bold="true" box="[151,568,474,501]" pageId="6" pageNumber="7">
<taxonomicName id="48964D38FFA95A7BFF4BFE23FE6CFE03" ID-CoL="BW7CQ" authority="MACFARLAN, 2023" authorityName="MACFARLAN" authorityYear="2023" box="[151,416,474,501]" class="Rhynchonellata" family="Dielasmatidae" genus="Tibetothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="7" phylum="Brachiopoda" rank="species" species="johnstoni" status="sp. nov.">
<emphasis id="BDE2EAA9FFA95A7BFF4BFE23FE6CFE03" bold="true" box="[151,416,474,501]" italics="true" pageId="6" pageNumber="7">Tibetothyris johnstoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6D157D2FFA95A7BFE74FE22FDF4FE02" box="[424,568,474,501]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C78C6530FFA95A7BFF4BFE06FE6EFDEF" box="[151,418,510,537]" pageId="6" pageNumber="7" type="description">
<paragraph id="8F2936BBFFA95A7BFF4BFE06FE6EFDEF" blockId="6.[151,568,474,537]" box="[151,418,510,537]" pageId="6" pageNumber="7">
<figureCitation id="17AD2A3EFFA95A7BFF4BFE06FF10FDEF" box="[151,220,510,537]" captionStart="FIGURE 5" captionStartId="7.[151,250,1823,1847]" captionTargetBox="[233,1361,193,1800]" captionTargetId="figure-20@7.[222,1365,181,1800]" captionTargetPageId="7" captionText="FIGURE 5. Tibetothyris johnstoni n. sp. 1 Holotype BR 3420 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 2 BR 3419 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior. 3 B890 (NC/f0109B) shelly specimen, ventral. 4 C1963 (E45/f6635) partly shelly specimen (a) dorsal (b) ventral (c) lateral (d) anterior. 5 C1965 (E45/f6635) internal mould, distorted, dorsal. 6 C1967 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 7 C1970 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 C1971 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral." figureDoi="http://doi.org/10.5281/zenodo.10148169" httpUri="https://zenodo.org/record/10148169/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
: 18.
<figureCitation id="17AD2A3EFFA95A7BFEC3FE06FE52FDEF" box="[287,414,511,537]" captionStart="FIGURE 6" captionStartId="8.[151,250,974,998]" captionTargetBox="[151,1435,184,950]" captionTargetId="figure-34@8.[151,1435,181,950]" captionTargetPageId="8" captionText="FIGURE 6. Detailed view of beak area, Tibetothyris johnstoni n. sp. A Holotype BR 3420 (N28/f7454), dorsal view of internal mould showing pedicle collar, ridged septalium, broad median septum and short, widely divergent dental plates. B C1970 (E45/f6635) dorsal view of internal mould, showing small cardinal process." figureDoi="http://doi.org/10.5281/zenodo.10148171" httpUri="https://zenodo.org/record/10148171/files/figure.png" pageId="6" pageNumber="7">Fig. 6 AB</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C78C6530FFA95A7BFF4BFDBEFCC3FCCF" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8F2936BBFFA95A7BFF4BFDBEFE1AFD73" blockId="6.[151,1437,582,1509]" pageId="6" pageNumber="7">
<materialsCitation id="3FFE3CE6FFA95A7BFF4BFDBEFE1AFD73" collectingDate="1898-03-19" collectorName="A. McKay" location="Eighty-Eight valley" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Bolu" typeStatus="holotype">
<emphasis id="BDE2EAA9FFA95A7BFF4BFDBEFEC8FD97" bold="true" box="[151,260,583,609]" pageId="6" pageNumber="7">
<typeStatus id="502D8819FFA95A7BFF4BFDBEFEC8FD97" box="[151,260,583,609]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingRegion id="4D52F859FFA95A7BFEC9FDBEFEF6FD97" box="[277,314,583,609]" country="Turkey" name="Bolu" pageId="6" pageNumber="7">BR</collectingRegion>
3420 a double-valved internal mould from N28/f7454, GS 196,
<location id="8A496060FFA95A7BFBF4FDBEFACAFD97" LSID="urn:lsid:plazi:treatment:073F87ADFFA95A75FF4BFE23FEC5FAB8:8A496060FFA95A7BFBF4FDBEFACAFD97" box="[1064,1286,583,609]" name="Eighty-Eight valley" pageId="6" pageNumber="7" stateProvince="Bolu">Eighty-Eight valley</location>
<collectingRegion id="4D52F859FFA95A7BFACCFDBEFAAFFD97" box="[1296,1379,583,609]" country="New Zealand" name="Nelson" pageId="6" pageNumber="7">Nelson</collectingRegion>
; collected
<collectorName id="2263536DFFA95A7BFF03FD92FE9AFD73" box="[223,342,619,645]" pageId="6" pageNumber="7">A. McKay</collectorName>
<date id="FB28107BFFA95A7BFE81FD92FE1EFD73" box="[349,466,619,645]" pageId="6" pageNumber="7" value="1898-03-19">
<collectingDate id="EB6CE993FFA95A7BFE81FD92FE1EFD73" box="[349,466,619,645]" pageId="6" pageNumber="7" value="1898-03-19">19/3/1898</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8F2936BBFFA95A7BFF1BFD77FB6DFD5F" blockId="6.[151,1437,582,1509]" box="[199,1185,654,681]" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFF1BFD77FE61FD5F" bold="true" box="[199,429,654,681]" pageId="6" pageNumber="7">Derivation of name</emphasis>
. This species is named for Dr Mike Johnston, geologist, of
<collectingRegion id="4D52F859FFA95A7BFB96FD76FB51FD5F" box="[1098,1181,655,681]" country="New Zealand" name="Nelson" pageId="6" pageNumber="7">Nelson</collectingRegion>
.
</paragraph>
<paragraph id="8F2936BBFFA95A7BFF1BFD4BFA8FFD3B" blockId="6.[151,1437,582,1509]" box="[199,1347,690,717]" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFF1BFD4BFEE3FD3A" bold="true" box="[199,303,690,716]" pageId="6" pageNumber="7">Material</emphasis>
. The data series consists of
<specimenCount id="9990FD32FFA95A7BFDB6FD4AFCF2FD3B" box="[618,830,691,717]" count="13" pageId="6" pageNumber="7" type="generic">thirteen specimens</specimenCount>
, eleven of which yielded valid measurements.
</paragraph>
<paragraph id="8F2936BBFFA95A7BFF1BFD2EFD34FD07" blockId="6.[151,1437,582,1509]" box="[199,760,726,753]" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFF1BFD2EFEB0FD06" bold="true" box="[199,380,726,753]" pageId="6" pageNumber="7">
<collectingCountry id="F781762BFFA95A7BFF1BFD2EFEB0FD06" box="[199,380,726,753]" name="New Caledonia" pageId="6" pageNumber="7">New Caledonia</collectingCountry>
</emphasis>
. île Hugon NC/f109a (AU 7165).
</paragraph>
<paragraph id="8F2936BBFFA95A7BFF1BFD02FB4CFCE3" blockId="6.[151,1437,582,1509]" box="[199,1152,762,789]" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFF1BFD02FED4FCE3" bold="true" box="[199,280,763,789]" pageId="6" pageNumber="7">
<collectingRegion id="4D52F859FFA95A7BFF1BFD02FED4FCE3" box="[199,280,763,789]" country="New Zealand" name="Nelson" pageId="6" pageNumber="7">Nelson</collectingRegion>
</emphasis>
. Eighty-Eight Valley N28/f7454 (
<collectionCode id="E987AE7EFFA95A7BFD48FD02FD75FCE3" box="[660,697,763,789]" pageId="6" pageNumber="7">GS</collectionCode>
196), Ram Creek; N28/f0045 (
<collectionCode id="E987AE7EFFA95A7BFBCBFD02FB80FCE3" box="[1047,1100,763,789]" pageId="6" pageNumber="7">PGS</collectionCode>
40).
</paragraph>
<paragraph id="8F2936BBFFA95A7BFF1BFCE7FCC3FCCF" blockId="6.[151,1437,582,1509]" box="[199,783,798,825]" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFF1BFCE7FE85FCCE" bold="true" box="[199,329,798,824]" pageId="6" pageNumber="7">
<collectingRegion id="4D52F859FFA95A7BFF1BFCE7FE88FCCE" box="[199,324,798,824]" country="New Zealand" name="Southland" pageId="6" pageNumber="7">Southland</collectingRegion>
.
</emphasis>
Taringatura Hills E45/f6635 (
<collectionCode id="E987AE7EFFA95A7BFD46FCE6FD02FCCF" box="[666,718,799,825]" pageId="6" pageNumber="7">JDC</collectionCode>
241).
</paragraph>
</subSubSection>
<subSubSection id="C78C6530FFA95A7BFF1BFCBBFDF7FA13" pageId="6" pageNumber="7" type="description">
<paragraph id="8F2936BBFFA95A7BFF1BFCBBFB8FFC77" blockId="6.[151,1437,582,1509]" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFF1BFCBBFE9DFCAA" bold="true" box="[199,337,834,860]" pageId="6" pageNumber="7">Description</emphasis>
. Small to medium sized terebratulide with rounded-pentagonal outline. The dorsal valve is moderately convex posteriorly and has a distinct shallow, rounded sulcus anteriorly.
</paragraph>
<paragraph id="8F2936BBFFA95A7BFF1BFC72FC97FBE7" blockId="6.[151,1437,582,1509]" pageId="6" pageNumber="7">The ventral valve is strongly convex posteriorly, with two close-spaced rounded plicae which diverge behind the beak, separated by a shallow rounded central sulcus. The flanks are broad and slightly concave. The anterior part has a high fold and a narrow, shallowly concave central part. The anterior commissure is strongly sulcate. The shell surface is smooth apart from a few strong, stepped concentrics.</paragraph>
<paragraph id="8F2936BBFFA95A7BFF1BFBE2FF79FB57" blockId="6.[151,1437,582,1509]" pageId="6" pageNumber="7">
The beak is large and triangular with a bluntly rounded tip. The foramen is large, circular, and mesothyrid, with a short pedicle collar (
<figureCitation id="17AD2A3EFFA95A7BFE4CFBC6FE3BFBAF" box="[400,503,1086,1113]" captionStart="FIGURE 5" captionStartId="7.[151,250,1823,1847]" captionTargetBox="[233,1361,193,1800]" captionTargetId="figure-20@7.[222,1365,181,1800]" captionTargetPageId="7" captionText="FIGURE 5. Tibetothyris johnstoni n. sp. 1 Holotype BR 3420 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 2 BR 3419 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior. 3 B890 (NC/f0109B) shelly specimen, ventral. 4 C1963 (E45/f6635) partly shelly specimen (a) dorsal (b) ventral (c) lateral (d) anterior. 5 C1965 (E45/f6635) internal mould, distorted, dorsal. 6 C1967 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 7 C1970 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 C1971 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral." figureDoi="http://doi.org/10.5281/zenodo.10148169" httpUri="https://zenodo.org/record/10148169/files/figure.png" pageId="6" pageNumber="7">Fig. 5.1d</figureCitation>
,
<figureCitation id="17AD2A3EFFA95A7BFDDFFBC6FDE5FBAF" box="[515,553,1087,1113]" captionStart="FIGURE 6" captionStartId="8.[151,250,974,998]" captionTargetBox="[151,1435,184,950]" captionTargetId="figure-34@8.[151,1435,181,950]" captionTargetPageId="8" captionText="FIGURE 6. Detailed view of beak area, Tibetothyris johnstoni n. sp. A Holotype BR 3420 (N28/f7454), dorsal view of internal mould showing pedicle collar, ridged septalium, broad median septum and short, widely divergent dental plates. B C1970 (E45/f6635) dorsal view of internal mould, showing small cardinal process." figureDoi="http://doi.org/10.5281/zenodo.10148171" httpUri="https://zenodo.org/record/10148171/files/figure.png" pageId="6" pageNumber="7">6A</figureCitation>
). The beak ridges are strong and bluntly rounded. The deltidial plates are small and widely disjunct (
<figureCitation id="17AD2A3EFFA95A7BFE5BFB9AFE23FB8B" box="[391,495,1122,1149]" captionStart="FIGURE 5" captionStartId="7.[151,250,1823,1847]" captionTargetBox="[233,1361,193,1800]" captionTargetId="figure-20@7.[222,1365,181,1800]" captionTargetPageId="7" captionText="FIGURE 5. Tibetothyris johnstoni n. sp. 1 Holotype BR 3420 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 2 BR 3419 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior. 3 B890 (NC/f0109B) shelly specimen, ventral. 4 C1963 (E45/f6635) partly shelly specimen (a) dorsal (b) ventral (c) lateral (d) anterior. 5 C1965 (E45/f6635) internal mould, distorted, dorsal. 6 C1967 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 7 C1970 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 C1971 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral." figureDoi="http://doi.org/10.5281/zenodo.10148169" httpUri="https://zenodo.org/record/10148169/files/figure.png" pageId="6" pageNumber="7">Fig. 5.1a</figureCitation>
,
<figureCitation id="17AD2A3EFFA95A7BFE21FB9AFDEEFB8B" box="[509,546,1123,1149]" captionStart="FIGURE 6" captionStartId="8.[151,250,974,998]" captionTargetBox="[151,1435,184,950]" captionTargetId="figure-34@8.[151,1435,181,950]" captionTargetPageId="8" captionText="FIGURE 6. Detailed view of beak area, Tibetothyris johnstoni n. sp. A Holotype BR 3420 (N28/f7454), dorsal view of internal mould showing pedicle collar, ridged septalium, broad median septum and short, widely divergent dental plates. B C1970 (E45/f6635) dorsal view of internal mould, showing small cardinal process." figureDoi="http://doi.org/10.5281/zenodo.10148171" httpUri="https://zenodo.org/record/10148171/files/figure.png" pageId="6" pageNumber="7">6A</figureCitation>
). The dental plates are short, divergent and widely separated. (
<figureCitation id="17AD2A3EFFA95A7BFB2CFB9AFF64FB57" captionStart="FIGURE 5" captionStartId="7.[151,250,1823,1847]" captionTargetBox="[233,1361,193,1800]" captionTargetId="figure-20@7.[222,1365,181,1800]" captionTargetPageId="7" captionText="FIGURE 5. Tibetothyris johnstoni n. sp. 1 Holotype BR 3420 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 2 BR 3419 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior. 3 B890 (NC/f0109B) shelly specimen, ventral. 4 C1963 (E45/f6635) partly shelly specimen (a) dorsal (b) ventral (c) lateral (d) anterior. 5 C1965 (E45/f6635) internal mould, distorted, dorsal. 6 C1967 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 7 C1970 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 C1971 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral." figureDoi="http://doi.org/10.5281/zenodo.10148169" httpUri="https://zenodo.org/record/10148169/files/figure.png" pageId="6" pageNumber="7">Fig. 5.2a, 5.7a, b</figureCitation>
).
</paragraph>
<paragraph id="8F2936BBFFA95A7BFF1BFB52FB9DFAFB" blockId="6.[151,1437,582,1509]" pageId="6" pageNumber="7">
The dorsal valve has a hingeplate with a deep U-shaped septalium (Fig, 5.1a, 6A, B), a broad, shallow median septum with longitudinal grooves which appear to be muscle attachments (
<figureCitation id="17AD2A3EFFA95A7BFC34FB36FB5CFB1F" box="[1000,1168,1230,1257]" captionStart="FIGURE 5" captionStartId="7.[151,250,1823,1847]" captionTargetBox="[233,1361,193,1800]" captionTargetId="figure-20@7.[222,1365,181,1800]" captionTargetPageId="7" captionText="FIGURE 5. Tibetothyris johnstoni n. sp. 1 Holotype BR 3420 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 2 BR 3419 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior. 3 B890 (NC/f0109B) shelly specimen, ventral. 4 C1963 (E45/f6635) partly shelly specimen (a) dorsal (b) ventral (c) lateral (d) anterior. 5 C1965 (E45/f6635) internal mould, distorted, dorsal. 6 C1967 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 7 C1970 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 C1971 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral." figureDoi="http://doi.org/10.5281/zenodo.10148169" httpUri="https://zenodo.org/record/10148169/files/figure.png" pageId="6" pageNumber="7">Fig. 5.1a, 5.2a</figureCitation>
). A small, high, striated cardinal process is present, and is close to the anterior of the foramen (
<figureCitation id="17AD2A3EFFA95A7BFC72FB0AFBD8FAFB" box="[942,1044,1266,1293]" captionStart="FIGURE 5" captionStartId="7.[151,250,1823,1847]" captionTargetBox="[233,1361,193,1800]" captionTargetId="figure-20@7.[222,1365,181,1800]" captionTargetPageId="7" captionText="FIGURE 5. Tibetothyris johnstoni n. sp. 1 Holotype BR 3420 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 2 BR 3419 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior. 3 B890 (NC/f0109B) shelly specimen, ventral. 4 C1963 (E45/f6635) partly shelly specimen (a) dorsal (b) ventral (c) lateral (d) anterior. 5 C1965 (E45/f6635) internal mould, distorted, dorsal. 6 C1967 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 7 C1970 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 C1971 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral." figureDoi="http://doi.org/10.5281/zenodo.10148169" httpUri="https://zenodo.org/record/10148169/files/figure.png" pageId="6" pageNumber="7">Fig. 5.2a</figureCitation>
,
<figureCitation id="17AD2A3EFFA95A7BFBFCFB0AFB88FAFB" box="[1056,1092,1267,1293]" captionStart="FIGURE 6" captionStartId="8.[151,250,974,998]" captionTargetBox="[151,1435,184,950]" captionTargetId="figure-34@8.[151,1435,181,950]" captionTargetPageId="8" captionText="FIGURE 6. Detailed view of beak area, Tibetothyris johnstoni n. sp. A Holotype BR 3420 (N28/f7454), dorsal view of internal mould showing pedicle collar, ridged septalium, broad median septum and short, widely divergent dental plates. B C1970 (E45/f6635) dorsal view of internal mould, showing small cardinal process." figureDoi="http://doi.org/10.5281/zenodo.10148171" httpUri="https://zenodo.org/record/10148171/files/figure.png" pageId="6" pageNumber="7">6B</figureCitation>
).
</paragraph>
<paragraph id="8F2936BBFFA95A7BFF1BFAEEFD6AFAC7" blockId="6.[151,1437,582,1509]" box="[199,678,1303,1329]" pageId="6" pageNumber="7">The shell is coarsely and densely punctate.</paragraph>
<paragraph id="8F2936BBFFA95A7BFF1BFAC3FC33FA8F" blockId="6.[151,1437,582,1509]" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFF1BFAC3FE9DFAA2" bold="true" box="[199,337,1338,1364]" pageId="6" pageNumber="7">Dimensions</emphasis>
. Dimensions of the
<typeStatus id="502D8819FFA95A7BFD90FAC2FD62FAA3" box="[588,686,1339,1365]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
and representative specimens, and statistics for all measurable specimens are shown in
<tableCitation id="C2140300FFA95A7BFE74FAA6FDD7FA8F" box="[424,539,1375,1401]" captionStart="TABLE 1" captionStartId="6.[151,239,1556,1580]" captionText="TABLE 1. Dimensions, Tibetothyris johnstoni n. sp." pageId="6" pageNumber="7">Table 1. A</tableCitation>
length vs width graph is shown in
<figureCitation id="17AD2A3EFFA95A7BFC7EFAA6FC37FA8F" box="[930,1019,1375,1401]" captionStart="FIGURE 9" captionStartId="11.[151,250,1489,1513]" captionTargetBox="[153,1434,181,1465]" captionTargetId="figure-20@11.[153,1434,181,1465]" captionTargetPageId="11" captionText="FIGURE 9. Length vs width graphs for ventral valve. A Tibetothyris johnstoni and T. hamishi. B Rhaetina rainei, New Caledonia and Nelson. C Lobothyris richardsi, (E44/f6635 South Taringatura Hills and other localities). D Zeilleria minima (E44/ f6635 and other localities), Zeilleria spiculata and Zeilleria n. sp. Z. spiculata data from MacFarlan (2021)." figureDoi="http://doi.org/10.5281/zenodo.10148177" httpUri="https://zenodo.org/record/10148177/files/figure.png" pageId="6" pageNumber="7">Fig. 9A</figureCitation>
.
</paragraph>
<paragraph id="8F2936BBFFA95A7BFF1BFA7AFDF7FA13" blockId="6.[151,1437,582,1509]" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFF1BFA7AFE13FA6A" bold="true" box="[199,479,1410,1437]" pageId="6" pageNumber="7">Range and Distribution</emphasis>
.
<materialsCitation id="3FFE3CE6FFA95A7BFE30FA7AFDF7FA13" collectionCode="JDC" country="New Caledonia" county="western Southland." location="Otapirian in Nelson. Of" municipality="South Taringatura Hills" pageId="6" pageNumber="7" specimenCode="JDC 241" specimenCount="13">
Otapirian, early Otapirian in
<collectingCountry id="F781762BFFA95A7BFCF0FA7AFC15FA6B" box="[812,985,1411,1437]" name="New Caledonia" pageId="6" pageNumber="7">New Caledonia</collectingCountry>
and Southland, late
<location id="8A496060FFA95A7BFB66FA7AFF79FA37" LSID="urn:lsid:plazi:treatment:073F87ADFFA95A75FF4BFE23FEC5FAB8:8A496060FFA95A7BFB66FA7AFF79FA37" country="New Caledonia" county="western Southland." municipality="South Taringatura Hills" name="Otapirian in Nelson. Of" pageId="6" pageNumber="7">Otapirian in Nelson. Of</location>
the
<specimenCount id="9990FD32FFA95A7BFF35FA5EFE4FFA37" box="[233,387,1447,1473]" count="13" pageId="6" pageNumber="7" type="generic">13 specimens</specimenCount>
measured, nine come from one early Otapirian locality (E45/f6635 (
<specimenCode id="DF309EC0FFA95A7BFB44FA5EFB33FA37" box="[1176,1279,1447,1473]" collectionCode="JDC" pageId="6" pageNumber="7">JDC 241</specimenCode>
) in the
<collectingMunicipality id="6F4DACC1FFA95A7BFA87FA5EFE94FA13" pageId="6" pageNumber="7">South Taringatura Hills</collectingMunicipality>
,
<collectingCounty id="66484E37FFA95A7BFEBFFA32FDF7FA13" box="[355,571,1483,1509]" pageId="6" pageNumber="7">western Southland.</collectingCounty>
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DBE96633FFA95A7BFF4BF9EDFD06F9DB" box="[151,714,1556,1581]" pageId="6" pageNumber="7" startId="6.[151,239,1556,1580]" targetType="table">
<paragraph id="8F2936BBFFA95A7BFF4BF9EDFD06F9DB" blockId="6.[151,714,1556,1581]" box="[151,714,1556,1581]" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFF4BF9EDFEC6F9DA" bold="true" box="[151,266,1556,1580]" pageId="6" pageNumber="7">TABLE 1.</emphasis>
Dimensions,
<taxonomicName id="48964D38FFA95A7BFE42F9EDFD4BF9DA" authority="MACFARLAN, 2023" authorityName="MACFARLAN" authorityYear="2023" box="[414,647,1556,1580]" class="Rhynchonellata" family="Dielasmatidae" genus="Tibetothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="7" phylum="Brachiopoda" rank="species" species="johnstoni" status="sp. nov.">
<emphasis id="BDE2EAA9FFA95A7BFE42F9EDFD4BF9DA" box="[414,647,1556,1580]" italics="true" pageId="6" pageNumber="7">Tibetothyris johnstoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6D157D2FFA95A7BFD52F9ECFD06F9DB" box="[654,714,1557,1581]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8F2936BBFFA95A7BFF7FF9C4FB3BF899" pageId="6" pageNumber="7">
<table id="FD96C41BFFA9A582FF7FF9C5FABAF875" box="[163,1398,1596,1923]" colsContinueIn="22.[163,1383,1126,1392]" gridcols="10" gridrows="7" pageId="6" pageNumber="7">
<tr id="31A634F9FFA9A582FF7FF9C5FABAF9A3" box="[163,1398,1596,1621]" gridrow="0" pageId="6" pageNumber="7">
<th id="72775D85FFA9A582FF7FF9C5FEF9F9A3" box="[163,309,1596,1621]" gridcol="0" gridrow="0" pageId="6" pageNumber="7">FR no.</th>
<th id="72775D85FFA9A582FE90F9C5FE60F9A3" box="[332,428,1596,1621]" gridcol="1" gridrow="0" pageId="6" pageNumber="7">specimen</th>
<th id="72775D85FFA9A582FE05F9C5FDD8F9A3" box="[473,532,1596,1621]" gridcol="2" gridrow="0" pageId="6" pageNumber="7">Lv</th>
<th id="72775D85FFA9A582FD97F9C5FD4AF9A3" box="[587,646,1596,1621]" gridcol="3" gridrow="0" pageId="6" pageNumber="7">Lp</th>
<th id="72775D85FFA9A582FD60F9C5FD24F9A3" box="[700,744,1596,1621]" gridcol="4" gridrow="0" pageId="6" pageNumber="7">W</th>
<th id="72775D85FFA9A582FCF2F9C5FC96F9A3" box="[814,858,1596,1621]" gridcol="5" gridrow="0" pageId="6" pageNumber="7">H</th>
<th id="72775D85FFA9A582FC43F9C5FC14F9A3" box="[927,984,1596,1621]" gridcol="6" gridrow="0" pageId="6" pageNumber="7">b&lt;</th>
<th id="72775D85FFA9A582FBCDF9C5FB82F9A3" box="[1041,1102,1596,1621]" gridcol="7" gridrow="0" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFBCDF9C5FB82F9A2" bold="true" box="[1041,1102,1596,1620]" pageId="6" pageNumber="7">MS/L</emphasis>
</th>
<th id="72775D85FFA9A582FBB2F9C5FB3BF9A3" box="[1134,1271,1596,1621]" gridcol="8" gridrow="0" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFBB2F9C5FB05F9A2" bold="true" box="[1134,1225,1596,1620]" pageId="6" pageNumber="7">material</emphasis>
</th>
<th id="72775D85FFA9A582FAC4F9C5FABAF9A3" box="[1304,1398,1596,1621]" gridcol="9" gridrow="0" pageId="6" pageNumber="7">
<emphasis id="BDE2EAA9FFA95A7BFAC4F9C5FA9CF9A2" bold="true" box="[1304,1360,1596,1620]" pageId="6" pageNumber="7">notes</emphasis>
</th>
</tr>
<tr id="31A634F9FFA9A582FF7FF997FABAF97E" box="[163,1398,1646,1672]" gridrow="1" pageId="6" pageNumber="7">
<th id="72775D85FFA9A582FF7FF997FEF9F97E" box="[163,309,1646,1672]" gridcol="0" gridrow="1" pageId="6" pageNumber="7">N28/f7454</th>
<td id="72775D85FFA9A582FE90F997FE60F97E" box="[332,428,1646,1672]" gridcol="1" gridrow="1" pageId="6" pageNumber="7">BR 3420</td>
<td id="72775D85FFA9A582FE05F997FDD8F97E" box="[473,532,1646,1672]" gridcol="2" gridrow="1" pageId="6" pageNumber="7">19.7</td>
<td id="72775D85FFA9A582FD97F997FD4AF97E" box="[587,646,1646,1672]" gridcol="3" gridrow="1" pageId="6" pageNumber="7">16.7</td>
<td id="72775D85FFA9A582FD60F997FD24F97E" box="[700,744,1646,1672]" gridcol="4" gridrow="1" pageId="6" pageNumber="7">16.3</td>
<td id="72775D85FFA9A582FCF2F997FC96F97E" box="[814,858,1646,1672]" gridcol="5" gridrow="1" pageId="6" pageNumber="7">10.2</td>
<td id="72775D85FFA9A582FC43F997FC14F97E" box="[927,984,1646,1672]" gridcol="6" gridrow="1" pageId="6" pageNumber="7">79</td>
<td id="72775D85FFA9A582FBCDF997FB82F97E" box="[1041,1102,1646,1672]" gridcol="7" gridrow="1" pageId="6" pageNumber="7">0.5</td>
<td id="72775D85FFA9A582FBB2F997FB3BF97E" box="[1134,1271,1646,1672]" gridcol="8" gridrow="1" pageId="6" pageNumber="7">b int</td>
<td id="72775D85FFA9A582FAC4F997FABAF97E" box="[1304,1398,1646,1672]" gridcol="9" gridrow="1" pageId="6" pageNumber="7">Holotype</td>
</tr>
<tr id="31A634F9FFA9A582FF7FF961FABAF946" box="[163,1398,1688,1712]" gridrow="2" pageId="6" pageNumber="7" rowspan-9="1">
<th id="72775D85FFA9A582FF7FF961FEF9F946" box="[163,309,1688,1712]" gridcol="0" gridrow="2" pageId="6" pageNumber="7">N28/f7454</th>
<td id="72775D85FFA9A582FE90F961FE60F946" box="[332,428,1688,1712]" gridcol="1" gridrow="2" pageId="6" pageNumber="7">BR 3419</td>
<td id="72775D85FFA9A582FE05F961FDD8F946" box="[473,532,1688,1712]" gridcol="2" gridrow="2" pageId="6" pageNumber="7">26.3+</td>
<td id="72775D85FFA9A582FD97F961FD4AF946" box="[587,646,1688,1712]" gridcol="3" gridrow="2" pageId="6" pageNumber="7">22.5+</td>
<td id="72775D85FFA9A582FD60F961FD24F946" box="[700,744,1688,1712]" gridcol="4" gridrow="2" pageId="6" pageNumber="7">20.5</td>
<td id="72775D85FFA9A582FCF2F961FC96F946" box="[814,858,1688,1712]" gridcol="5" gridrow="2" pageId="6" pageNumber="7">10.1</td>
<td id="72775D85FFA9A582FC43F961FC14F946" box="[927,984,1688,1712]" gridcol="6" gridrow="2" pageId="6" pageNumber="7">77</td>
<td id="72775D85FFA9A582FBCDF961FB82F946" box="[1041,1102,1688,1712]" gridcol="7" gridrow="2" pageId="6" pageNumber="7">0.5</td>
<td id="72775D85FFA9A582FBB2F961FB3BF946" box="[1134,1271,1688,1712]" gridcol="8" gridrow="2" pageId="6" pageNumber="7">b int</td>
</tr>
<tr id="31A634F9FFA9A582FF7FF93CFABAF914" box="[163,1398,1733,1762]" gridrow="3" pageId="6" pageNumber="7" rowspan-7="1" rowspan-9="1">
<th id="72775D85FFA9A582FF7FF93CFEF9F914" box="[163,309,1733,1762]" gridcol="0" gridrow="3" pageId="6" pageNumber="7">E45/f6635</th>
<td id="72775D85FFA9A582FE90F93CFE60F914" box="[332,428,1733,1762]" gridcol="1" gridrow="3" pageId="6" pageNumber="7">C1963</td>
<td id="72775D85FFA9A582FE05F93CFDD8F914" box="[473,532,1733,1762]" gridcol="2" gridrow="3" pageId="6" pageNumber="7">20.4</td>
<td id="72775D85FFA9A582FD97F93CFD4AF914" box="[587,646,1733,1762]" gridcol="3" gridrow="3" pageId="6" pageNumber="7">18.8</td>
<td id="72775D85FFA9A582FD60F93CFD24F914" box="[700,744,1733,1762]" gridcol="4" gridrow="3" pageId="6" pageNumber="7">16.4</td>
<td id="72775D85FFA9A582FCF2F93CFC96F914" box="[814,858,1733,1762]" gridcol="5" gridrow="3" pageId="6" pageNumber="7">9.8</td>
<td id="72775D85FFA9A582FC43F93CFC14F914" box="[927,984,1733,1762]" gridcol="6" gridrow="3" pageId="6" pageNumber="7">60</td>
<td id="72775D85FFA9A582FBB2F93CFB3BF914" box="[1134,1271,1733,1762]" gridcol="8" gridrow="3" pageId="6" pageNumber="7">b pt shelly</td>
</tr>
<tr id="31A634F9FFA9A582FF7FF90BFABAF8FC" box="[163,1398,1778,1802]" gridrow="4" pageId="6" pageNumber="7" rowspan-9="1">
<th id="72775D85FFA9A582FF7FF90BFEF9F8FC" box="[163,309,1778,1802]" gridcol="0" gridrow="4" pageId="6" pageNumber="7">E45/f6635</th>
<td id="72775D85FFA9A582FE90F90BFE60F8FC" box="[332,428,1778,1802]" gridcol="1" gridrow="4" pageId="6" pageNumber="7">C1967</td>
<td id="72775D85FFA9A582FE05F90BFDD8F8FC" box="[473,532,1778,1802]" gridcol="2" gridrow="4" pageId="6" pageNumber="7">21.9</td>
<td id="72775D85FFA9A582FD97F90BFD4AF8FC" box="[587,646,1778,1802]" gridcol="3" gridrow="4" pageId="6" pageNumber="7">18.9</td>
<td id="72775D85FFA9A582FD60F90BFD24F8FC" box="[700,744,1778,1802]" gridcol="4" gridrow="4" pageId="6" pageNumber="7">17.5</td>
<td id="72775D85FFA9A582FCF2F90BFC96F8FC" box="[814,858,1778,1802]" gridcol="5" gridrow="4" pageId="6" pageNumber="7">8.9</td>
<td id="72775D85FFA9A582FC43F90BFC14F8FC" box="[927,984,1778,1802]" gridcol="6" gridrow="4" pageId="6" pageNumber="7">78</td>
<td id="72775D85FFA9A582FBCDF90BFB82F8FC" box="[1041,1102,1778,1802]" gridcol="7" gridrow="4" pageId="6" pageNumber="7">0.6</td>
<td id="72775D85FFA9A582FBB2F90BFB3BF8FC" box="[1134,1271,1778,1802]" gridcol="8" gridrow="4" pageId="6" pageNumber="7">b int</td>
</tr>
<tr id="31A634F9FFA9A582FF7FF8E3FABAF8C5" box="[163,1398,1818,1843]" gridrow="5" pageId="6" pageNumber="7" rowspan-9="1">
<th id="72775D85FFA9A582FF7FF8E3FEF9F8C5" box="[163,309,1818,1843]" gridcol="0" gridrow="5" pageId="6" pageNumber="7">E45/f6635</th>
<td id="72775D85FFA9A582FE90F8E3FE60F8C5" box="[332,428,1818,1843]" gridcol="1" gridrow="5" pageId="6" pageNumber="7">C1970</td>
<td id="72775D85FFA9A582FE05F8E3FDD8F8C5" box="[473,532,1818,1843]" gridcol="2" gridrow="5" pageId="6" pageNumber="7">19.3</td>
<td id="72775D85FFA9A582FD97F8E3FD4AF8C5" box="[587,646,1818,1843]" gridcol="3" gridrow="5" pageId="6" pageNumber="7">15.7</td>
<td id="72775D85FFA9A582FD60F8E3FD24F8C5" box="[700,744,1818,1843]" gridcol="4" gridrow="5" pageId="6" pageNumber="7">16.2</td>
<td id="72775D85FFA9A582FCF2F8E3FC96F8C5" box="[814,858,1818,1843]" gridcol="5" gridrow="5" pageId="6" pageNumber="7">7.2</td>
<td id="72775D85FFA9A582FC43F8E3FC14F8C5" box="[927,984,1818,1843]" gridcol="6" gridrow="5" pageId="6" pageNumber="7">72</td>
<td id="72775D85FFA9A582FBCDF8E3FB82F8C5" box="[1041,1102,1818,1843]" gridcol="7" gridrow="5" pageId="6" pageNumber="7">0.5</td>
<td id="72775D85FFA9A582FBB2F8E3FB3BF8C5" box="[1134,1271,1818,1843]" gridcol="8" gridrow="5" pageId="6" pageNumber="7">b int</td>
</tr>
<tr id="31A634F9FFA9A582FF7FF8BAFABAF875" box="[163,1398,1859,1923]" gridrow="6" pageId="6" pageNumber="7" rowspan-7="1" rowspan-9="1">
<th id="72775D85FFA9A582FF7FF8BAFEF9F875" box="[163,309,1859,1923]" gridcol="0" gridrow="6" pageId="6" pageNumber="7">All valid spec- imens</th>
<td id="72775D85FFA9A582FE90F8BAFE60F875" box="[332,428,1859,1923]" gridcol="1" gridrow="6" pageId="6" pageNumber="7">mean S.D.</td>
<td id="72775D85FFA9A582FE05F8BAFDD8F875" box="[473,532,1859,1923]" gridcol="2" gridrow="6" pageId="6" pageNumber="7">18.9 8.96</td>
<td id="72775D85FFA9A582FD97F8BAFD4AF875" box="[587,646,1859,1923]" gridcol="3" gridrow="6" pageId="6" pageNumber="7">17.3 8.30</td>
<td id="72775D85FFA9A582FD60F8BAFD24F875" box="[700,744,1859,1923]" gridcol="4" gridrow="6" pageId="6" pageNumber="7">15.4 7.33</td>
<td id="72775D85FFA9A582FCF2F8BAFC96F875" box="[814,858,1859,1923]" gridcol="5" gridrow="6" pageId="6" pageNumber="7">8.4 4.07</td>
<td id="72775D85FFA9A582FC43F8BAFC14F875" box="[927,984,1859,1923]" gridcol="6" gridrow="6" pageId="6" pageNumber="7">71.8 30.91</td>
<td id="72775D85FFA9A582FBB2F8BAFB3BF875" box="[1134,1271,1859,1923]" gridcol="8" gridrow="6" pageId="6" pageNumber="7">11 specimens</td>
</tr>
</table>
</paragraph>
<caption id="DBE96633FFA85A7AFF4BF8E6FB54F81A" ID-DOI="http://doi.org/10.5281/zenodo.10148169" ID-Zenodo-Dep="10148169" httpUri="https://zenodo.org/record/10148169/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1823,1847]" targetBox="[233,1361,193,1800]" targetPageId="7" targetType="figure">
<paragraph id="8F2936BBFFA85A7AFF4BF8E6FDFBF8CE" blockId="7.[151,1437,1823,2028]" box="[151,567,1823,1848]" pageId="7" pageNumber="8">
<emphasis id="BDE2EAA9FFA85A7AFF4BF8E6FEDFF8CE" bold="true" box="[151,275,1823,1848]" pageId="7" pageNumber="8">FIGURE 5.</emphasis>
<taxonomicName id="48964D38FFA85A7AFEC5F8D9FE3BF8CE" authority="MACFARLAN, 2023" authorityName="MACFARLAN" authorityYear="2023" box="[281,503,1824,1848]" class="Rhynchonellata" family="Dielasmatidae" genus="Tibetothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="7" pageNumber="8" phylum="Brachiopoda" rank="species" species="johnstoni" status="sp. nov.">
<emphasis id="BDE2EAA9FFA85A7AFEC5F8D9FE3BF8CE" box="[281,503,1824,1848]" italics="true" pageId="7" pageNumber="8">Tibetothyris johnstoni</emphasis>
</taxonomicName>
<emphasis id="BDE2EAA9FFA85A7AFE21F8D9FDFBF8CE" bold="true" box="[509,567,1824,1848]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A6D157D2FFA85A7AFE21F8D9FDFBF8CE" box="[509,567,1824,1848]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8F2936BBFFA85A7AFF4BF8BAFB54F81A" blockId="7.[151,1437,1823,2028]" pageId="7" pageNumber="8">
<emphasis id="BDE2EAA9FFA85A7AFF4BF8BAFF68F8AD" bold="true" box="[151,164,1859,1883]" pageId="7" pageNumber="8">1</emphasis>
Holotype BR 3420 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior.
<emphasis id="BDE2EAA9FFA85A7AFBE3F8BAFB80F8AD" bold="true" box="[1087,1100,1859,1883]" pageId="7" pageNumber="8">2</emphasis>
BR 3419 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior.
<emphasis id="BDE2EAA9FFA85A7AFE25F89EFDCAF889" bold="true" box="[505,518,1895,1919]" pageId="7" pageNumber="8">3</emphasis>
B890 (NC/f0109B) shelly specimen, ventral.
<emphasis id="BDE2EAA9FFA85A7AFC05F89EFC2AF889" bold="true" box="[985,998,1895,1919]" pageId="7" pageNumber="8">4</emphasis>
C1963 (E45/f6635) partly shelly specimen (a) dorsal (b) ventral (c) lateral (d) anterior.
<emphasis id="BDE2EAA9FFA85A7AFD97F875FD94F852" bold="true" box="[587,600,1932,1956]" pageId="7" pageNumber="8">5</emphasis>
C1965 (E45/f6635) internal mould, distorted, dorsal.
<emphasis id="BDE2EAA9FFA85A7AFBAFF872FB4CF855" bold="true" box="[1139,1152,1931,1955]" pageId="7" pageNumber="8">6</emphasis>
C1967 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior.
<emphasis id="BDE2EAA9FFA85A7AFCF4F849FCF9F83E" bold="true" box="[808,821,1968,1992]" pageId="7" pageNumber="8">7</emphasis>
C1970 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior.
<emphasis id="BDE2EAA9FFA85A7AFE02F82AFE27F81D" bold="true" box="[478,491,2003,2027]" pageId="7" pageNumber="8">8</emphasis>
C1971 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral.
</paragraph>
</caption>
<caption id="DBE96633FFA75A75FF4BFC37FC5AFBD8" ID-DOI="http://doi.org/10.5281/zenodo.10148171" ID-Zenodo-Dep="10148171" httpUri="https://zenodo.org/record/10148171/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,974,998]" targetBox="[151,1435,184,950]" targetPageId="8" targetType="figure">
<paragraph id="8F2936BBFFA75A75FF4BFC37FC5AFBD8" blockId="8.[151,1436,974,1070]" pageId="8" pageNumber="9">
<emphasis id="BDE2EAA9FFA75A75FF4BFC37FEDFFC10" bold="true" box="[151,275,974,998]" pageId="8" pageNumber="9">FIGURE 6.</emphasis>
Detailed view of beak area,
<taxonomicName id="48964D38FFA75A75FDE5FC37FCDAFC10" authority="MACFARLAN, 2023" authorityName="MACFARLAN" authorityYear="2023" box="[569,790,974,998]" class="Rhynchonellata" family="Dielasmatidae" genus="Tibetothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="8" pageNumber="9" phylum="Brachiopoda" rank="species" species="johnstoni" status="sp. nov.">
<emphasis id="BDE2EAA9FFA75A75FDE5FC37FCDAFC10" box="[569,790,974,998]" italics="true" pageId="8" pageNumber="9">Tibetothyris johnstoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6D157D2FFA75A75FCC1FC36FC98FC10" box="[797,852,975,998]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="BDE2EAA9FFA75A75FCBEFC37FCB8FC10" bold="true" box="[866,884,974,998]" pageId="8" pageNumber="9">A</emphasis>
Holotype BR 3420 (N28/f7454), dorsal view of internal mould showing pedicle collar, ridged septalium, broad median septum and short, widely divergent dental plates.
<emphasis id="BDE2EAA9FFA75A75FA9DFC0BFA9EFBFC" bold="true" box="[1345,1362,1010,1034]" pageId="8" pageNumber="9">B</emphasis>
C1970 (E45/f6635) dorsal view of internal mould, showing small cardinal process.
</paragraph>
</caption>
<subSubSection id="C78C6530FFA75A75FF1BFBA4FEC5FAB8" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8F2936BBFFA75A75FF1BFBA4FEC5FAB8" blockId="8.[151,1437,1116,1358]" pageId="8" pageNumber="9">
<emphasis id="BDE2EAA9FFA75A75FF1BFBA4FEF8FB81" bold="true" box="[199,308,1117,1143]" pageId="8" pageNumber="9">Remarks</emphasis>
. This species is close to
<taxonomicName id="48964D38FFA75A75FD9BFBA5FCFAFB80" authorityName="Jin and Sun" authorityYear="1976" box="[583,822,1116,1142]" class="Rhynchonellata" family="Dielasmatidae" genus="Tibetothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="8" pageNumber="9" phylum="Brachiopoda" rank="species" species="depressa">
<emphasis id="BDE2EAA9FFA75A75FD9BFBA5FCFAFB80" box="[583,822,1116,1142]" italics="true" pageId="8" pageNumber="9">Tibetothyris depressa</emphasis>
</taxonomicName>
in shape, but the folding differs.
<taxonomicName id="48964D38FFA75A75FB7AFBA4FAE8FB80" authorityName="Jin and Sun" authorityYear="1976" box="[1190,1316,1116,1142]" class="Rhynchonellata" family="Dielasmatidae" genus="Tibetothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="8" pageNumber="9" phylum="Brachiopoda" rank="species" species="depressa">
<emphasis id="BDE2EAA9FFA75A75FB7AFBA4FAE8FB80" box="[1190,1316,1116,1142]" italics="true" pageId="8" pageNumber="9">T. depressa</emphasis>
</taxonomicName>
has strong rounded lateral folds and two weak central folds on the dorsal valve, and a weak central fold on the ventral fold. The anterior commissure is sulciplicate to bisulcate (
<bibRefCitation id="EB074B4AFFA75A75FD73FB5DFC83FB48" author="Jin, Y. G. &amp; Sun, D. L. &amp; Rong, J. Y." box="[687,847,1188,1215]" pageId="8" pageNumber="9" pagination="271 - 346" refId="ref13996" refString="Jin, Y. G., Sun, D. L. &amp; Rong, J. Y. (1976) Mesozoic and Cenozoic Brachiopods from the Mount Jolmo Lungma Region. In: Xizang Scientific Expedition Team of Chinese Academy of Sciences (Ed.), A Report of Scientific Expedition in the Mount Jolmo Lungma Region (1966 - 1968). (Palaeontology). Fasc. II. Science Press, Beijing, pp. 271 - 346, pls. 1 - 10. [in Chinese]" type="book chapter" year="1976">Jin et al. 1976</bibRefCitation>
,
<bibRefCitation id="EB074B4AFFA75A75FC86FB5DFC34FB48" author="Jin, Y-G. &amp; Lee, D. E. &amp; Sun D-L &amp; Smirnova, T. N. &amp; Dagys, A. S. &amp; Sandy, M. R." box="[858,1016,1188,1215]" pageId="8" pageNumber="9" refId="ref14092" refString="Jin, Y-G., Lee, D. E., Sun D-L, Smirnova, T. N., Dagys, A. S. &amp; Sandy, M. R. (2006) Dielasmatoidea. In: Kaesler, R. L. (Ed.), Treatise on Invertebrate Paleontology. Part H, Brachiopoda. Vol. 5. Revised. Geological Society of America, Boulder, Colorado and Paleontological Institute, Lawrence, Kansas, pp. 2029 - 2053." type="book" year="2006">Jin et al. 2006</bibRefCitation>
,
<bibRefCitation id="EB074B4AFFA75A75FBDFFB5CFB62FB49" author="Sun, D. L. &amp; Xu, G. R. &amp; Qiao, L." box="[1027,1198,1188,1215]" pageId="8" pageNumber="9" pagination="883 - 1011" refId="ref17064" refString="Sun, D. L., Xu, G. R. &amp; Qiao, L. (2017) Triassic brachiopod genera on type species from China. In: Rong, J. Y., Jin Y. G., Shen S. Z. &amp; Zhan R. B. (Eds.), Phanerozoic Brachiopod Genera of China. Science Press, Beijing, pp. 883 - 1011." type="book chapter" year="2017">Sun et al. 2017</bibRefCitation>
).
<taxonomicName id="48964D38FFA75A75FB1CFB5CFA8FFB48" authorityName="MACFARLAN" authorityYear="2023" box="[1216,1347,1188,1214]" class="Rhynchonellata" family="Dielasmatidae" genus="Tibetothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="8" pageNumber="9" phylum="Brachiopoda" rank="species" species="johnstoni">
<emphasis id="BDE2EAA9FFA75A75FB1CFB5CFA8FFB48" box="[1216,1347,1188,1214]" italics="true" pageId="8" pageNumber="9">T. johnstoni</emphasis>
</taxonomicName>
is more strongly folded. The dorsal valve has similar strong rounded lateral folds and a central sulcus. The ventral valve is strongly convex with two low, incipient or poorly developed rounded folds anteriorly. The anterior commissure is unisulcate.
<taxonomicName id="48964D38FFA75A75FEE4FAE8FE76FADC" authorityName="Jin and Sun" authorityYear="1976" box="[312,442,1296,1322]" class="Rhynchonellata" family="Dielasmatidae" genus="Tibetothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="8" pageNumber="9" phylum="Brachiopoda" rank="species" species="depressa">
<emphasis id="BDE2EAA9FFA75A75FEE4FAE8FE76FADC" box="[312,442,1296,1322]" italics="true" pageId="8" pageNumber="9">T. depressa</emphasis>
</taxonomicName>
has long dental plates whereas in
<taxonomicName id="48964D38FFA75A75FC8FFAE8FC16FADC" authorityName="MACFARLAN" authorityYear="2023" box="[851,986,1296,1322]" class="Rhynchonellata" family="Dielasmatidae" genus="Tibetothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="8" pageNumber="9" phylum="Brachiopoda" rank="species" species="johnstoni">
<emphasis id="BDE2EAA9FFA75A75FC8FFAE8FC16FADC" box="[851,986,1296,1322]" italics="true" pageId="8" pageNumber="9">T. johnstoni</emphasis>
</taxonomicName>
the dental plates are short and widely separated.
</paragraph>
</subSubSection>
</treatment>
</document>