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<document id="2F6ED39C057B69C9773255B06FA0343B" ID-DOI="10.5281/zenodo.157708" ID-GBIF-Dataset="aabec420-0763-4ca5-b4bb-e0528c17f814" ID-ISSN="1175­5326" ID-Zenodo-Dep="157708" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461018528122" checkinUser="plazi" docAuthor="Rafael, José Albertino &amp; Ale-Rocha, Rosaly" docDate="2004" docId="C779878C9A482225FECECA93C808FADD" docLanguage="en" docName="zt00529.pdf" docOrigin="Zootaxa 529" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Tomosvaryella crassa Ale-Rocha, n. sp." docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="3B40FFF49A45222BFFC6CF5FCC45FFC3" masterDocTitle="Nicaraguan Pipunculidae (Diptera): new records and description of new species" masterLastPageNumber="18" masterPageNumber="1" pageNumber="14" updateTime="1698569576216" updateUser="plazi">
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<mods:titleInfo id="ED0F54E403F84CB985B9B2D060601B85">
<mods:title id="87505CFEA77E36222CC388C34DDF2BEE">Nicaraguan Pipunculidae (Diptera): new records and description of new species</mods:title>
</mods:titleInfo>
<mods:name id="07083D82730270DC347E12240FDA7239" type="personal">
<mods:role id="D03C7FC53592D64944638B796632CED9">
<mods:roleTerm id="95CD39600951EF3DE295767AF2AA5DDC">Author</mods:roleTerm>
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<mods:namePart id="335036F5FC3D74F3E7A51A878B30EDD0">Rafael, José Albertino</mods:namePart>
</mods:name>
<mods:name id="A55D51B2FB10C172ECA6F423F74AC1FA" type="personal">
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<mods:namePart id="87E90AB73A22B21C8F7FDFF6367D02EE">Ale-Rocha, Rosaly</mods:namePart>
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<mods:title id="1AE9A2A15CE3D551FAD37104A60BC55F">Zootaxa</mods:title>
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<mods:date id="58784960F67DD7307C44298517F48D69">2004</mods:date>
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<mods:number id="921EEB055D80BCD3554C703A664D1D31">529</mods:number>
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<mods:classification id="A27E530A7E7FCE1DA791D7B4CFBD2A33">journal article</mods:classification>
<mods:identifier id="2A6DFE2EE64B45A50108F4B55B10ABE8" type="DOI">10.5281/zenodo.157708</mods:identifier>
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<mods:identifier id="466C547E9D73D1A790ACCA66354764D8" type="ISSN">1175­5326</mods:identifier>
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<treatment id="C779878C9A482225FECECA93C808FADD" ID-DOI="http://doi.org/10.5281/zenodo.6269524" ID-GBIF-Taxon="119602132" ID-Zenodo-Dep="6269524" LSID="urn:lsid:plazi:treatment:C779878C9A482225FECECA93C808FADD" httpUri="http://treatment.plazi.org/id/C779878C9A482225FECECA93C808FADD" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="07CA65119A482226FECECA93CDE1F9CA" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="4F6F369A9A482226FECECA93CEA2FA25" blockId="13.[264,743,1484,1545]" box="[264,743,1484,1510]" pageId="13" pageNumber="14">
<heading id="142781F69A482226FECECA93CEA2FA25" bold="true" box="[264,743,1484,1510]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="7DA4EA889A482226FECECA93CEA2FA25" bold="true" box="[264,743,1484,1510]" pageId="13" pageNumber="14">
<taxonomicName id="88D04D199A482226FECECA93CEDDFA25" ID-CoL="7CH4C" authority="Ale-Rocha" authorityName="Ale-Rocha" box="[264,664,1484,1510]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="crassa" status="sp. nov.">
<emphasis id="7DA4EA889A482226FECECA93CE4EFA25" bold="true" box="[264,523,1484,1510]" italics="true" pageId="13" pageNumber="14">Tomosvaryella crassa</emphasis>
Ale­Rocha
</taxonomicName>
,
<taxonomicNameLabel id="669757F39A482226FD63CA93CEA2FA25" box="[677,743,1484,1510]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4F6F369A9A482226FECECAB0CDE1F9CA" blockId="13.[264,743,1484,1545]" box="[264,420,1519,1545]" pageId="13" pageNumber="14">
(
<figureCitation id="D7EB2A1F9A482226FED7CAB0CDD9F9CA" box="[273,412,1519,1545]" captionStart="FIGURES 33 – 43. 33 – 37" captionStartId="12.[264,383,1312,1336]" captionTargetBox="[275,1303,412,1287]" captionTargetId="figure@12.[264,1323,388,1307]" captionTargetPageId="12" captionText="FIGURES 33 – 43. 33 – 37 Dasydorylas vulcanus, n. sp., ht ɗ: 33. abdomen, dv; 34. syntergosternite 8, epandrium and surstyli, vv; 35. outer surstylus, lv; 36. apex of phallic guide and phallus, lv; 37. ejaculatory apodeme. 38 – 43 Tomosvaryella crassa, sp. n., ht ɗ: 38. hind trochanter, 39. syntergosternite 8, epandrium and surstyli, vv; 40. inner surstylus, lv; 41. outer surstylus, lv; 42. apex of phal­" httpUri="https://zenodo.org/record/157711/files/figure.png" pageId="13" pageNumber="14">Figs. 38–43</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="07CA65119A482225FECEC960C808FADD" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="4F6F369A9A482226FECEC960CF5DF96A" blockId="13.[264,1323,1599,1905]" pageId="13" pageNumber="14">
Diagnosis. Wing anal lobe widened.
<collectingCountry id="37C7760A9A482226FD07C960CEB9F99A" box="[705,764,1599,1625]" name="India" pageId="13" pageNumber="14">Hind</collectingCountry>
trochanter with rounded ventral process.
<collectingCountry id="37C7760A9A482226FB29C960C96FF99A" box="[1263,1322,1599,1625]" name="India" pageId="13" pageNumber="14">Hind</collectingCountry>
tarsomeres slender; first hind tarsomere as long as the three following segments together. Surstyli very wide, as long as the epandrium.
</paragraph>
<paragraph id="4F6F369A9A482225FEFEC9E8C88DFD05" blockId="13.[264,1323,1599,1905]" lastBlockId="14.[264,1325,284,1310]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Description (ɗ). Body length 3.5 mm; wings 3.3 mm. Eyes contiguous on the frons for about 10 facets; ocellar triangle as long as half of length of the contiguous area of eyes; postcranium black, silvery pruinose; frontal triangle silvery pruinose; antennae brown, anterior margin of flagellum pale yellow. Thorax: scutum and scutellum black, mesopleuron dark brown; postsutural region brown pruinose; presutural and postalar region, scutellum, subscutellum, laterotergite and notopleuron silvery­gray pruinose; mesopleuron sparsely silvery­gray pruinose; setae short, inconspicuous. Wing hyaline; third costal section as long as one third of the fourth; anal lobe widened. Legs dark brown except apex of femora, basal one fourth of the tibiae and ventral surface of the tarsi yellow; tibiae apex and dorsal surface of the tarsi pale brown, first and fifth tarsi slightly darkened; forefemur with an inconspicuous pair of sub­basal spines; hind trochanter (
<figureCitation id="D7EB2A1F9A4B2225FBD8CEBBC832FE3D" box="[1054,1143,484,510]" captionStart="FIGURES 33 – 43. 33 – 37" captionStartId="12.[264,383,1312,1336]" captionTargetBox="[275,1303,412,1287]" captionTargetId="figure@12.[264,1323,388,1307]" captionTargetPageId="12" captionText="FIGURES 33 – 43. 33 – 37 Dasydorylas vulcanus, n. sp., ht ɗ: 33. abdomen, dv; 34. syntergosternite 8, epandrium and surstyli, vv; 35. outer surstylus, lv; 36. apex of phallic guide and phallus, lv; 37. ejaculatory apodeme. 38 – 43 Tomosvaryella crassa, sp. n., ht ɗ: 38. hind trochanter, 39. syntergosternite 8, epandrium and surstyli, vv; 40. inner surstylus, lv; 41. outer surstylus, lv; 42. apex of phal­" httpUri="https://zenodo.org/record/157711/files/figure.png" pageId="14" pageNumber="15">Fig. 38</figureCitation>
) with rounded ventral process; hind tarsomeres slender, first tarsomere as long as the three following segments together. Abdomen wide, black, brown pruinose dorsally, silvery­gray pruinose laterally on tergites 1, 4 and 5. Syntergoesternite 8 short, narrow, about half of length of tergite 5, setae inconspicuous. Terminalia as in
<figureCitation id="D7EB2A1F9A4B2225FCFFCDDBCFD5FD5D" box="[825,912,644,670]" captionStart="FIGURES 33 – 43. 33 – 37" captionStartId="12.[264,383,1312,1336]" captionTargetBox="[275,1303,412,1287]" captionTargetId="figure@12.[264,1323,388,1307]" captionTargetPageId="12" captionText="FIGURES 33 – 43. 33 – 37 Dasydorylas vulcanus, n. sp., ht ɗ: 33. abdomen, dv; 34. syntergosternite 8, epandrium and surstyli, vv; 35. outer surstylus, lv; 36. apex of phallic guide and phallus, lv; 37. ejaculatory apodeme. 38 – 43 Tomosvaryella crassa, sp. n., ht ɗ: 38. hind trochanter, 39. syntergosternite 8, epandrium and surstyli, vv; 40. inner surstylus, lv; 41. outer surstylus, lv; 42. apex of phal­" httpUri="https://zenodo.org/record/157711/files/figure.png" pageId="14" pageNumber="15">Fig. 39</figureCitation>
. Surstyli in lateral view as in
<figureCitation id="D7EB2A1F9A4B2225FB37CDDBCD10FD05" captionStart="FIGURES 33 – 43. 33 – 37" captionStartId="12.[264,383,1312,1336]" captionTargetBox="[275,1303,412,1287]" captionTargetId="figure@12.[264,1323,388,1307]" captionTargetPageId="12" captionText="FIGURES 33 – 43. 33 – 37 Dasydorylas vulcanus, n. sp., ht ɗ: 33. abdomen, dv; 34. syntergosternite 8, epandrium and surstyli, vv; 35. outer surstylus, lv; 36. apex of phallic guide and phallus, lv; 37. ejaculatory apodeme. 38 – 43 Tomosvaryella crassa, sp. n., ht ɗ: 38. hind trochanter, 39. syntergosternite 8, epandrium and surstyli, vv; 40. inner surstylus, lv; 41. outer surstylus, lv; 42. apex of phal­" httpUri="https://zenodo.org/record/157711/files/figure.png" pageId="14" pageNumber="15">Figs. 40, 41</figureCitation>
. Phallic guide and phallus as in
<figureCitation id="D7EB2A1F9A4B2225FD0BCDF3CF61FD05" box="[717,804,684,710]" captionStart="FIGURES 33 – 43. 33 – 37" captionStartId="12.[264,383,1312,1336]" captionTargetBox="[275,1303,412,1287]" captionTargetId="figure@12.[264,1323,388,1307]" captionTargetPageId="12" captionText="FIGURES 33 – 43. 33 – 37 Dasydorylas vulcanus, n. sp., ht ɗ: 33. abdomen, dv; 34. syntergosternite 8, epandrium and surstyli, vv; 35. outer surstylus, lv; 36. apex of phallic guide and phallus, lv; 37. ejaculatory apodeme. 38 – 43 Tomosvaryella crassa, sp. n., ht ɗ: 38. hind trochanter, 39. syntergosternite 8, epandrium and surstyli, vv; 40. inner surstylus, lv; 41. outer surstylus, lv; 42. apex of phal­" httpUri="https://zenodo.org/record/157711/files/figure.png" pageId="14" pageNumber="15">Fig. 43</figureCitation>
. Ejaculatory apodeme as in
<figureCitation id="D7EB2A1F9A4B2225FBABCDF3C881FD05" box="[1133,1220,684,710]" captionStart="FIGURES 33 – 43. 33 – 37" captionStartId="12.[264,383,1312,1336]" captionTargetBox="[275,1303,412,1287]" captionTargetId="figure@12.[264,1323,388,1307]" captionTargetPageId="12" captionText="FIGURES 33 – 43. 33 – 37 Dasydorylas vulcanus, n. sp., ht ɗ: 33. abdomen, dv; 34. syntergosternite 8, epandrium and surstyli, vv; 35. outer surstylus, lv; 36. apex of phallic guide and phallus, lv; 37. ejaculatory apodeme. 38 – 43 Tomosvaryella crassa, sp. n., ht ɗ: 38. hind trochanter, 39. syntergosternite 8, epandrium and surstyli, vv; 40. inner surstylus, lv; 41. outer surstylus, lv; 42. apex of phal­" httpUri="https://zenodo.org/record/157711/files/figure.png" pageId="14" pageNumber="15">Fig. 43</figureCitation>
.
</paragraph>
<paragraph id="4F6F369A9A4B2225FEFECD8BCE4EFD2D" blockId="14.[264,1325,284,1310]" box="[312,523,724,750]" pageId="14" pageNumber="15">Female unknown.</paragraph>
<paragraph id="4F6F369A9A4B2225FEFECDA3CEC0FC4D" blockId="14.[264,1325,284,1310]" pageId="14" pageNumber="15">
Material.
<typeStatus id="906B88389A4B2225FE6CCDA3CE52FCD5" box="[426,535,764,790]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
ɗ (CAS). Verbatim text of two pin labels:
<collectingCountry id="37C7760A9A4B2225FBD7CDA3C8CFFCD5" box="[1041,1162,764,790]" name="Mexico" pageId="14" pageNumber="15">MEXICO</collectingCountry>
: Sonora, Alamos,
<date id="3B6E105A9A4B2225FE8FCC7BCDEEFCFD" box="[329,427,804,830]" pageId="14" pageNumber="15" value="1971-01-05">5.i.1971</date>
, P.H. &amp; M. Arnaud [white paper] /
<typeStatus id="906B88389A4B2225FC9ECC7BCF80FCFD" box="[856,965,804,830]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
ɗ,
<emphasis id="7DA4EA889A4B2225FC36CC7BCD17FCA6" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="88D04D199A4B2225FC36CC7BC8B5FCFE" box="[1008,1264,804,829]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="crassa">Tomosvaryella crassa</taxonomicName>
Ale­ Rocha
</emphasis>
[red paper]. Terminalia and left hind leg placed in a microvial with glycerine and pinned along with the specimen.
</paragraph>
<paragraph id="4F6F369A9A4B2225FEFECCC3CD39FBED" blockId="14.[264,1325,284,1310]" pageId="14" pageNumber="15">
<typeStatus id="906B88389A4B2225FEFECCC3CDEAFC75" box="[312,431,924,950]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="37C7760A9A4B2225FE7DCCC3CE71FC75" box="[443,564,924,950]" name="Mexico" pageId="14" pageNumber="15">MEXICO</collectingCountry>
: Sonora, 134 road Km, south Sonoyta,
<date id="3B6E105A9A4B2225FBCACCC3C8C0FC75" box="[1036,1157,924,950]" pageId="14" pageNumber="15" value="1963-02-15">15.ii.1963</date>
, P.H. Arnaud, Jr. (1ɗ CAS);
<collectingCountry id="37C7760A9A4B2225FE78CC9BCE34FC1D" box="[446,625,964,990]" name="Nicaragua" pageId="14" pageNumber="15">NICARAGUA</collectingCountry>
, Masaya, Masatepe,
<date id="3B6E105A9A4B2225FCB5CC9BCF8DFC1D" box="[883,968,964,990]" pageId="14" pageNumber="15" value="1995-02">ii.1995</date>
, s/ citrus + Malezas, Projeto NIM (2ɗ INPA, MEL).
<typeStatus id="906B88389A4B2225FDEFCCB3CE26FBC5" box="[553,611,1004,1030]" pageId="14" pageNumber="15">Type</typeStatus>
conditions: abdomen and right hindleg of
<typeStatus id="906B88389A4B2225FBA6CCB3C8D7FBC5" box="[1120,1170,1004,1030]" pageId="14" pageNumber="15">type</typeStatus>
in tube with glycerine.
</paragraph>
<paragraph id="4F6F369A9A4B2225FEFECB63C959FB95" blockId="14.[264,1325,284,1310]" box="[312,1308,1084,1110]" pageId="14" pageNumber="15">
Etymology. The specific name refers to the broad surstyli. From Latin
<emphasis id="7DA4EA889A4B2225FBB5CB63C896FB96" box="[1139,1235,1084,1109]" italics="true" pageId="14" pageNumber="15">crassus,</emphasis>
thick.
</paragraph>
<paragraph id="4F6F369A9A4B2225FEFECB3BC808FADD" blockId="14.[264,1325,284,1310]" pageId="14" pageNumber="15">
Remarks.
<taxonomicName id="88D04D199A4B2225FE72CB3BCE5AFBBE" box="[436,543,1124,1149]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="7DA4EA889A4B2225FE72CB3BCE5AFBBE" box="[436,543,1124,1149]" italics="true" pageId="14" pageNumber="15">T. crassa</emphasis>
</taxonomicName>
can be included in dichotomy 9 of the key of Ale­Rocha and Rafael (1995). The large size and form of the hind trochanter suggest that it is closely related to
<taxonomicName id="88D04D199A4B2225FEBBCBEBCF7DFB0D" authority="Ale-Rocha &amp; Rafael" authorityName="Ale-Rocha &amp; Rafael" box="[381,824,1204,1230]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="membranacea">
<emphasis id="7DA4EA889A4B2225FEBBCBEBCE05FB0E" box="[381,576,1204,1229]" italics="true" pageId="14" pageNumber="15">T. membranacea</emphasis>
Ale­Rocha &amp; Rafael
</taxonomicName>
and
<taxonomicName id="88D04D199A4B2225FCB4CBEBCD5FFB35" authority="Ale-Rocha &amp; Rafael. It" authorityName="Ale-Rocha &amp; Rafael. It" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mexicanensis">
<emphasis id="7DA4EA889A4B2225FCB4CBEBC86EFB0E" box="[882,1067,1204,1229]" italics="true" pageId="14" pageNumber="15">T. mexicanensis</emphasis>
Ale­Rocha &amp; Rafael. It
</taxonomicName>
differs from both species by the lack of long bristles on the hind femur, short and narrow epandrium, wide, large surstyli and phallic guide without dorsal thorns.
</paragraph>
</subSubSection>
</treatment>
</document>
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