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<document id="95BD2E1E47B38CC352A00C50807A9508" ID-DOI="10.5281/zenodo.157708" ID-GBIF-Dataset="aabec420-0763-4ca5-b4bb-e0528c17f814" ID-ISSN="1175­5326" ID-Zenodo-Dep="157708" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461018528122" checkinUser="plazi" docAuthor="Rafael, José Albertino &amp; Ale-Rocha, Rosaly" docDate="2004" docId="C779878C9A43222DFECECE43CF76F8E0" docLanguage="en" docName="zt00529.pdf" docOrigin="Zootaxa 529" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Eudorylas maesi Rafael, n. sp." docType="treatment" docVersion="9" lastPageNumber="7" masterDocId="3B40FFF49A45222BFFC6CF5FCC45FFC3" masterDocTitle="Nicaraguan Pipunculidae (Diptera): new records and description of new species" masterLastPageNumber="18" masterPageNumber="1" pageNumber="7" updateTime="1698569576216" updateUser="plazi">
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<mods:title id="C6F56E1C7C35046A226A04BF61E06FF7">Nicaraguan Pipunculidae (Diptera): new records and description of new species</mods:title>
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<mods:namePart id="40B57F0FA6E7302A4956B19AB9958E56">Rafael, José Albertino</mods:namePart>
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<mods:namePart id="CF719B0503A3C7F6942D603C7471CF7F">Ale-Rocha, Rosaly</mods:namePart>
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<mods:date id="C55C75F6E2D81116FC87AD4E52DA9244">2004</mods:date>
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<treatment id="C779878C9A43222DFECECE43CF76F8E0" ID-DOI="http://doi.org/10.5281/zenodo.6269504" ID-GBIF-Taxon="119602153" ID-Zenodo-Dep="6269504" LSID="urn:lsid:plazi:treatment:C779878C9A43222DFECECE43CF76F8E0" httpUri="http://treatment.plazi.org/id/C779878C9A43222DFECECE43CF76F8E0" lastPageNumber="7" pageId="6" pageNumber="7">
<subSubSection id="07CA65119A43222DFECECE43CDD3FE9A" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="4F6F369A9A43222DFECECE43CE3EFEF6" blockId="6.[264,635,283,345]" box="[264,635,283,310]" pageId="6" pageNumber="7">
<heading id="142781F69A43222DFECECE43CE3EFEF6" bold="true" box="[264,635,283,310]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="7DA4EA889A43222DFECECE43CE3EFEF6" bold="true" box="[264,635,283,310]" pageId="6" pageNumber="7">
<taxonomicName id="88D04D199A43222DFECECE43CE69FEF6" ID-CoL="3C2X8" authority="Rafael" authorityName="Rafael" box="[264,556,283,310]" class="Insecta" family="Pipunculidae" genus="Eudorylas" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maesi" status="sp. nov.">
<emphasis id="7DA4EA889A43222DFECECE43CD94FEF5" bold="true" box="[264,465,284,310]" italics="true" pageId="6" pageNumber="7">Eudorylas maesi</emphasis>
Rafael
</taxonomicName>
,
<taxonomicNameLabel id="669757F39A43222DFDFFCE44CE3EFEF6" box="[569,635,283,309]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4F6F369A9A43222DFECECE60CDD3FE9A" blockId="6.[264,635,283,345]" box="[264,406,319,345]" pageId="6" pageNumber="7">
(
<figureCitation id="D7EB2A1F9A43222DFED7CE60CDC8FE9A" box="[273,397,319,345]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="6" pageNumber="7">Figs. 8–12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="07CA65119A43222DFECECED0CF76F8E0" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="4F6F369A9A43222DFECECED0C88AFE3A" blockId="6.[264,1325,399,1827]" pageId="6" pageNumber="7">Diagnosis. Antennae with scape and pedicel brown, flagellum yellow, acute. Third costal section slightly shorter than fourth. Legs yellow, except coxae and base of femora black. Syntergosternite 8 with large membranous area ventrally. Surstyli subsymmetrical.</paragraph>
<paragraph id="4F6F369A9A43222DFEFECD58CE57FB60" blockId="6.[264,1325,399,1827]" pageId="6" pageNumber="7">
Description (male). Body length 3.5 mm; wing 3.6 mm. Eyes contiguous on frons for a distance greater than length of frontal triangle. Frontal triangle and face gray pruinose. Antennae with scape and pedicel brown, flagellum yellow, acute. Thorax dark brown to black with scutum, scutellum, postpronotal lobe and notopleuron brown pruinose. Mesopleuron, subscutellum and laterotergite gray pruinose. Legs yellow, except coxae black and anterior face of fore and midfemora and posterior face of hind femur mainly brown basally. Distal tarsomeres brown. All femora with ventral ctenidia. Tibia without developed bristles, either distally or medially. Wing with third costal section slightly shorter than fourth; pterostigma incomplete; vein r­m placed near basal third of cell dm; vein M1 slightly sinuous; vein dm­cu straight, about as long as vein CuA1. Halter with stem light yellow, knob brown. Abdomen (
<figureCitation id="D7EB2A1F9A43222DFCCCCCC6CF17FC70" box="[778,850,921,947]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
) black, brown pruinose with gray pruinosity on all of tergite 1, base of tergite 2 and posterolaterally on tergites 3–5. Syntergosternite 8 black, brown pruinose, as long as tergite 5, with large membranous area ventrally. Epandrium and surstyli (
<figureCitation id="D7EB2A1F9A43222DFDEACB4ECE37FBE8" box="[556,626,1041,1067]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
) yellow to brown. Surstyli subsymmetrical; in lateral view as in
<figureCitation id="D7EB2A1F9A43222DFE8ECB66CDDAFB90" box="[328,415,1081,1107]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="6" pageNumber="7">Fig. 10</figureCitation>
, showing the base projecting downward. Phallic guide and phallus as in
<figureCitation id="D7EB2A1F9A43222DFB3BCB66CD62FBB8" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
. Phallus trifid, short, ramificated, with subapical membranous expansion. Ejaculatory apodeme as in
<figureCitation id="D7EB2A1F9A43222DFE71CBD6CE4BFB60" box="[439,526,1161,1187]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="6" pageNumber="7">Fig. 12</figureCitation>
.
</paragraph>
<paragraph id="4F6F369A9A43222DFEFECBEEC944FAD8" blockId="6.[264,1325,399,1827]" pageId="6" pageNumber="7">Female. Body length 3.3 mm; wing 3.4 mm. Frons gray pruinose. Syntergosternite 7+8 and ovipositor (Fig. 13) black, brown pruinose dorsally and gray laterally; ovipositor straight, longer than syntergosternite. In other respects identical to description of male.</paragraph>
<paragraph id="4F6F369A9A43222DFEFECA76C813F998" blockId="6.[264,1325,399,1827]" pageId="6" pageNumber="7">
Material.
<typeStatus id="906B88389A43222DFE6CCA76CE52FA80" box="[426,535,1321,1347]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
ɗ (INPA). Verbatim text of three pin labels:
<collectingCountry id="37C7760A9A43222DFBE1CA76C89FFA80" box="[1063,1242,1321,1347]" name="Nicaragua" pageId="6" pageNumber="7">NICARAGUA</collectingCountry>
,
<collectingCountry id="37C7760A9A43222DFB20CA76CD76FAA8" name="Grenada" pageId="6" pageNumber="7">Granada</collectingCountry>
, Volcán Mombacho, Santa Ana #2 [white paper] /
<date id="3B6E105A9A43222DFC44CA0EC841FAA8" box="[898,1028,1361,1387]" pageId="6" pageNumber="7" value="1998-03-16">16.iii.1998</date>
, J.M. Maes, Malaise trap [white paper] /
<typeStatus id="906B88389A43222DFE0ECA26CE70FA50" box="[456,565,1401,1427]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
ɗ,
<taxonomicName id="88D04D199A43222DFDA1CA26CFCDFA50" authority="Rafael" authorityName="Rafael" box="[615,904,1401,1427]" class="Insecta" family="Pipunculidae" genus="Eudorylas" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maesi">
<emphasis id="7DA4EA889A43222DFDA1CA26CF75FA51" box="[615,816,1401,1426]" italics="true" pageId="6" pageNumber="7">Eudorylas maesi</emphasis>
Rafael
</taxonomicName>
[red paper].
<typeStatus id="906B88389A43222DFBEACA26C8E6FA50" box="[1068,1187,1401,1427]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
: NICARA­
<collectingCountry id="37C7760A9A43222DFECECAFECD0EFA78" box="[264,331,1441,1467]" name="India" pageId="6" pageNumber="7">GUA</collectingCountry>
,
<collectingCountry id="37C7760A9A43222DFE9ECAFECDFBFA78" box="[344,446,1441,1467]" name="Grenada" pageId="6" pageNumber="7">Granada</collectingCountry>
, Volcán Mombacho, San Joaquin #2,
<date id="3B6E105A9A43222DFC42CAFECFE1FA78" box="[900,932,1441,1467]" pageId="6" pageNumber="7" value="1998-02-28">28</date>
.ii.1998, J.M. Maes, Malaise trap (1ɗ 1Ψ INPA); Santa Ana #2,
<date id="3B6E105A9A43222DFDB7CA96CECBFA20" box="[625,654,1481,1507]" pageId="6" pageNumber="7" value="1998-03-16">16</date>
.iii.1998 (4ɗ 1Ψ INPA); El Progreso, #1,
<date id="3B6E105A9A43222DFBBECA96C8DDFA20" box="[1144,1176,1481,1507]" pageId="6" pageNumber="7" value="1998-03-02">02</date>
.iii.1998 (1ɗ MEL); Bosque Nuboso,
<date id="3B6E105A9A43222DFDE1CAAECEE3F9C8" box="[551,678,1521,1547]" pageId="6" pageNumber="7" value="1998-03-02">02.iii.1998</date>
(1Ψ MEL); San Joaquin #1,
<date id="3B6E105A9A43222DFC3ECAAEC850F9C8" box="[1016,1045,1521,1547]" pageId="6" pageNumber="7" value="1998-03-31">31</date>
.iii.1998 (1ɗ 1Ψ MEL); Santa Ana #1,
<date id="3B6E105A9A43222DFE0FC946CDA3F9F0" box="[457,486,1561,1587]" pageId="6" pageNumber="7" value="1998-03-31">31</date>
.iii.1998 (1Ψ MEL); Santa Ana,
<quantity id="88289B7F9A43222DFC43C946CF9CF9F0" box="[901,985,1561,1587]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="7" unit="m" value="700.0">700 m</quantity>
,
<date id="3B6E105A9A43222DFC2DC946C827F9F0" box="[1003,1122,1561,1587]" pageId="6" pageNumber="7" value="1998-02-15">15.ii.1998</date>
(2ɗ 4Ψ MEL); Masaya, Masatepe,
<date id="3B6E105A9A43222DFE37C91ECE00F998" box="[497,581,1601,1627]" pageId="6" pageNumber="7" value="1995-02">ii.1995</date>
, s/ citrus+malezas, projecto NIM (1Ψ MEL).
</paragraph>
<paragraph id="4F6F369A9A43222DFEFEC936CEB4F968" blockId="6.[264,1325,399,1827]" pageId="6" pageNumber="7">Etymology. The specific name is in honour of Dr. Jean Michel Maes, who collected almost all specimens treated in this paper.</paragraph>
<paragraph id="4F6F369A9A43222DFEFEC9E6CF76F8E0" blockId="6.[264,1325,399,1827]" pageId="6" pageNumber="7">
Remarks. This species differs from other species of
<taxonomicName id="88D04D199A43222DFC65C9E6C859F911" box="[931,1052,1721,1746]" class="Insecta" family="Pipunculidae" genus="Eudorylas" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7DA4EA889A43222DFC65C9E6C859F911" box="[931,1052,1721,1746]" italics="true" pageId="6" pageNumber="7">Eudorylas</emphasis>
</taxonomicName>
by the combination of the following characters: incomplete pterostigma, surstyli base projecting downward and phallus with subapical membranous expansion.
</paragraph>
</subSubSection>
</treatment>
</document>
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