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<document id="93F242CFCDA0E6FF59A5C8565AFBEBE7" ID-CLB-Dataset="168727" ID-DOI="10.11646/zootaxa.5195.6.3" ID-GBIF-Dataset="893c13e2-2785-463d-b114-a499cbe8a7c8" ID-ISSN="1175-5326" ID-Zenodo-Dep="7223884" ID-ZooBank="EB51F148-8EFC-4B22-9B74-1DBE4EB3A544" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666164886612" checkinUser="plazi" docAuthor="Bilton, David T. &amp; Shepard, William D." docDate="2022" docId="C74FFC13303EFF9BFF68544DDD82FA37" docLanguage="en" docName="zootaxa.5195.6.3.pdf" docOrigin="Zootaxa 5195 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhithrops capensis Bilton &amp; Shepard 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="546" masterDocId="3B76846B303CFF9CFFFF525FDD64FF81" masterDocTitle="Rhithrops capensis gen. et sp. nov., a new aquatic dryopid beetle from the Western Cape of South Africa (Coleoptera: Dryopoidea: Dryopidae)" masterLastPageNumber="553" masterPageNumber="539" pageNumber="541" updateTime="1699407958300" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="9C7332BDA4CF2948A5FEBFE9E855A253" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C9F56CFE361638DD045B20A080699A95">
<mods:title id="F9FA20B419F8FDC8CE0F49112C14B7FD">Rhithrops capensis gen. et sp. nov., a new aquatic dryopid beetle from the Western Cape of South Africa (Coleoptera: Dryopoidea: Dryopidae)</mods:title>
</mods:titleInfo>
<mods:name id="87F6FB906417579D7D8BF5DEAC92291C" type="personal">
<mods:role id="71D05D94A097E709903E44362621AB44">
<mods:roleTerm id="627EA49A484110C432433ECF4E6AC8A7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1C2A869FCBCDDDBCD4AA20F75491F72">Bilton, David T.</mods:namePart>
<mods:nameIdentifier id="ADDE19DBF466FE7AA9ABD989769CDFC0" type="ORCID">0000-0003-1136-0848</mods:nameIdentifier>
<mods:affiliation id="DA2B70F69DB3EFA00ABC865B0806DD60">School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth PL 4 8 AA, UK. d. bilton @ plymouth. ac. uk; https: // orcid. org / 0000 - 0003 - 1136 - 0848 &amp; Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, Johannesburg 2006, South Africa.</mods:affiliation>
<mods:nameIdentifier id="EBEEF716EA0A276D71DB931981E669C3" type="email">d.bilton@plymouth.ac.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="14D4437BE08A36EFFDB7F74436E5A98F" type="personal">
<mods:role id="58E631DB113403952DCBD18EA9D940B0">
<mods:roleTerm id="23FAF3CF415D95BBAC2D6D5475E6FEE7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="90CF5F3132727CEF220C1D1C8919AE24">Shepard, William D.</mods:namePart>
<mods:nameIdentifier id="0A7504D1B46075E2857C910365E34332" type="ORCID">0000-0003-4664-2597</mods:nameIdentifier>
<mods:affiliation id="BEA0EC0D4D31D128C17F75567C2F788A">Essig Museum of Entomology, 1101 Life Sciences Bldg. # 4780, University of California Berkeley, CA 94720 USA. William. Shepard @ csus. edu; http: // orcid. org / 0000 - 0003 - 4664 - 2597</mods:affiliation>
<mods:nameIdentifier id="AE06092B9FB715351364B2D94EFF4A45" type="email">hepard@csus.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="05F9A59C634D793FFADC9F6F07CDE188">text</mods:typeOfResource>
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<mods:titleInfo id="18882E6A92EB7C1D482ADAA1ADAA4A93">
<mods:title id="9EAEC355F9DCACBB90B10E1FB71C385A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="280E3D519796D11AC112A797D8CE74BF">
<mods:date id="CC6E7AD935B4B4EBF9F662188AC33BB2">2022</mods:date>
<mods:detail id="A60AC3460B9EB77DB11E8FFACA7390C4" type="pubDate">
<mods:number id="64280BEEECF4D47C28484390E4C837C5">2022-10-17</mods:number>
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<mods:number id="85814813F385D048E3E9E56DCF8856DF">5195</mods:number>
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<mods:detail id="D7817527D706C4D690C35C374DC188D6" type="issue">
<mods:number id="7DC2A108924211CE6D9F9982F081D12B">6</mods:number>
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<mods:classification id="0E25A742103BE7DCB7D031F524E7B888">journal article</mods:classification>
<mods:identifier id="F6F343D538BE002A0B6081EBB91E3599" type="CLB-Dataset">168727</mods:identifier>
<mods:identifier id="70478B660358B6A38CFABE1D4AC96890" type="DOI">10.11646/zootaxa.5195.6.3</mods:identifier>
<mods:identifier id="FD7F3527BB60B1CEE4DC4DEC18AFE16C" type="GBIF-Dataset">893c13e2-2785-463d-b114-a499cbe8a7c8</mods:identifier>
<mods:identifier id="B69750DD1C93654D2EFEC94B4FF14B70" type="ISSN">1175-5326</mods:identifier>
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<treatment id="C74FFC13303EFF9BFF68544DDD82FA37" ID-DOI="http://doi.org/10.5281/zenodo.7223892" ID-GBIF-Taxon="203115545" ID-Zenodo-Dep="7223892" LSID="urn:lsid:plazi:treatment:C74FFC13303EFF9BFF68544DDD82FA37" httpUri="http://treatment.plazi.org/id/C74FFC13303EFF9BFF68544DDD82FA37" lastPageId="7" lastPageNumber="546" pageId="2" pageNumber="541">
<subSubSection id="07FC1E8E303EFF9EFF68544DDC87F9AC" box="[151,483,1554,1581]" pageId="2" pageNumber="541" type="nomenclature">
<paragraph id="4F594D05303EFF9EFF68544DDC87F9AC" blockId="2.[151,483,1554,1617]" box="[151,483,1554,1581]" pageId="2" pageNumber="541">
<heading id="1411FA69303EFF9EFF68544DDC87F9AC" bold="true" box="[151,483,1554,1581]" fontSize="11" level="1" pageId="2" pageNumber="541" reason="1">
<emphasis id="7D929117303EFF9EFF68544DDC87F9AC" bold="true" box="[151,483,1554,1581]" pageId="2" pageNumber="541">
<taxonomicName id="88E63686303EFF9EFF68544DDC1AF9AC" authority="Bilton &amp; Shepard, 2022" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[151,382,1554,1581]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="capensis" status="sp. nov.">
<emphasis id="7D929117303EFF9EFF68544DDC1AF9AC" bold="true" box="[151,382,1554,1581]" italics="true" pageId="2" pageNumber="541">Rhithrops capensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="66A12C6C303EFF9EFE7A544CDC87F9AC" box="[389,483,1555,1581]" pageId="2" pageNumber="541" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07FC1E8E303EFF9EFF685468DC6FF9D0" box="[151,267,1591,1617]" pageId="2" pageNumber="541" type="description">
<paragraph id="4F594D05303EFF9EFF685468DC6FF9D0" blockId="2.[151,483,1554,1617]" box="[151,267,1591,1617]" pageId="2" pageNumber="541">
(
<figureCitation id="D7DD5180303EFF9EFF605468DC66F9D0" box="[159,258,1591,1617]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStart-6="FIG" captionStart-7="FIG" captionStart-8="FIG" captionStartId-0="3.[152,201,1943,1967]" captionStartId-1="4.[151,196,1943,1967]" captionStartId-2="5.[151,196,1643,1667]" captionStartId-3="6.[151,196,1614,1638]" captionStartId-4="7.[151,196,934,958]" captionStartId-5="8.[151,196,1842,1866]" captionStartId-6="9.[151,196,1643,1667]" captionStartId-7="10.[151,196,1643,1667]" captionStartId-8="11.[151,196,1943,1967]" captionTargetBox-0="[199,1378,203,1900]" captionTargetBox-1="[214,1373,181,1919]" captionTargetBox-2="[154,1434,181,1619]" captionTargetBox-3="[167,1421,181,1590]" captionTargetBox-4="[180,1407,209,861]" captionTargetBox-5="[154,1434,381,1819]" captionTargetBox-6="[154,1434,181,1619]" captionTargetBox-7="[154,1434,181,1619]" captionTargetBox-8="[211,1375,201,1919]" captionTargetId-0="figure-19@3.[187,1401,179,1918]" captionTargetId-1="figure-25@4.[214,1373,181,1919]" captionTargetId-2="figure-19@5.[154,1434,181,1619]" captionTargetId-3="figure-25@6.[167,1421,181,1590]" captionTargetId-4="figure-19@7.[154,1434,181,910]" captionTargetId-5="figure-126@8.[154,1434,381,1819]" captionTargetId-6="figure-19@9.[154,1434,181,1619]" captionTargetId-7="figure-25@10.[154,1434,181,1619]" captionTargetId-8="figure-19@11.[164,1423,181,1919]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionText-0="FIG. 1. Rhithrops capensis gen. et sp. nov., Holotype habitus. Scale bar = 1 mm." captionText-1="FIG. 2. South African endemic dryopid genera, dorsolateral view of pronotal margins. A) Rhithrops capensis gen. et sp. nov., holotype male; B) Rhithrops capensis gen. et sp. nov., paratype female; C) Rapnus sp., female; D) Strina sp., female. C) &amp; D) South Africa, Western Cape, Franschoek, Mont Rochelle. Scale bar = 0.5 mm." captionText-2="FIG. 3. Rhithrops capensis gen. et sp. nov. A) head, posterolateral view; B) compound eye – arrow indicates inter-ommatidial seta; C) detail of dorsolateral sculpture of head, inside compound eye; D) antennae; E) antennomeres 1–2; F) antennomeres 3–10." captionText-3="FIG. 4. Rhithrops capensis gen. et sp. nov. A) labrum, dorsal view; B) right mandible, ventral view; C) right maxilla, dorsal view; D) close-up of maxillary palp apex; E) female labrum and mentum, ventral view; F) male labrum and mentum, anteroventral view." captionText-4="FIG. 5. Rhithrops capensis gen. et sp. nov. A) right mandible, ventrolateral view – teeth numbered as in text; B) left mandible, ventral view – pt = prostheca, Lf = lateral flange. Scale bar = 100 μm." captionText-5="FIG. 6. Rhithrops capensis gen. et sp. nov. A) prothorax, dorsal view; B) close-up of sculpture of pronotal disc; C) Meso and metanotum, with scutellum; D) right elytron, dorsolateral view; E) detail of right elytral apex; F) prothorax, ventral view." captionText-6="FIG. 7. Rhithrops capensis gen. et sp. nov. A) meso and metaventrites, ventral view; B) male abdomen, ventral view; C) female abdomen, ventral view; D) close-up of plastron vestiture at junction of abdominal ventrites 2–3; E) male abdominal apex; F) female abdominal apex." captionText-7="FIG. 8. Rhithrops capensis gen. et sp. nov. A) male foreleg, ventral view; B) male foretarsus and tibial apex, ventral view; C) male hind tibia and tarsus, ventral view; D) aedeagus, ventral view; E) aedeagus, lateral view; F) aedeagal apex, ventral view." captionText-8="FIG. 9. Rhithrops capensis gen. et sp. nov. A) aedeagus, ventrolateral view; B) aedeagus, lateral view; C) male sternite 9; D) aedeagal apex, ventral view (not to scale); E) male tergite 8; F) male sternite 8. Scale bars = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7223888" figureDoi-1="http://doi.org/10.5281/zenodo.7223890" figureDoi-2="http://doi.org/10.5281/zenodo.7223894" figureDoi-3="http://doi.org/10.5281/zenodo.7223899" figureDoi-4="http://doi.org/10.5281/zenodo.7223903" figureDoi-5="http://doi.org/10.5281/zenodo.7223905" figureDoi-6="http://doi.org/10.5281/zenodo.7223907" figureDoi-7="http://doi.org/10.5281/zenodo.7223910" figureDoi-8="http://doi.org/10.5281/zenodo.7223912" httpUri-0="https://zenodo.org/record/7223888/files/figure.png" httpUri-1="https://zenodo.org/record/7223890/files/figure.png" httpUri-2="https://zenodo.org/record/7223894/files/figure.png" httpUri-3="https://zenodo.org/record/7223899/files/figure.png" httpUri-4="https://zenodo.org/record/7223903/files/figure.png" httpUri-5="https://zenodo.org/record/7223905/files/figure.png" httpUri-6="https://zenodo.org/record/7223907/files/figure.png" httpUri-7="https://zenodo.org/record/7223910/files/figure.png" httpUri-8="https://zenodo.org/record/7223912/files/figure.png" pageId="2" pageNumber="541">Figs 1–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="07FC1E8E303EFF9EFF685420DF97F838" pageId="2" pageNumber="541" type="materials_examined">
<paragraph id="4F594D05303EFF9EFF685420DC23F93C" blockId="2.[151,1437,1662,1977]" pageId="2" pageNumber="541">
<emphasis id="7D929117303EFF9EFF685420DC50F919" bold="true" box="[151,308,1662,1689]" pageId="2" pageNumber="541">Type locality.</emphasis>
<materialsCitation id="FF8E4758303EFF9EFEC55421DC27F93C" ID-GBIF-Occurrence="3926284302" country="South Africa" county="Cederberg" elevation="725" latitude="-32.518726" location="Matjiesrivier" longLatPrecision="1" longitude="19.350111" municipality="Matjies Rivier" pageId="2" pageNumber="541" specimenCount="725" stateProvince="Western Cape">
<emphasis id="7D929117303EFF9EFEC55421DCB4F919" bold="true" box="[314,464,1662,1688]" pageId="2" pageNumber="541">
<collectingCountry id="37F10D95303EFF9EFEC55421DCB4F919" box="[314,464,1662,1688]" name="South Africa" pageId="2" pageNumber="541">South Africa</collectingCountry>
</emphasis>
,
<collectingRegion id="8D2283E7303EFF9EFE225420DF1EF918" box="[477,634,1663,1689]" country="South Africa" name="Western Cape" pageId="2" pageNumber="541">Western Cape</collectingRegion>
,
<collectingCounty id="A6383589303EFF9EFD7B5420DF9FF918" box="[644,763,1663,1689]" pageId="2" pageNumber="541">Cederberg</collectingCounty>
,
<collectingMunicipality id="AF3DD77F303EFF9EFCFA5420DEC3F918" box="[773,935,1663,1689]" pageId="2" pageNumber="541">Matjies Rivier</collectingMunicipality>
,
<location id="4A391BDE303EFF9EFC4F5420D927F918" LSID="urn:lsid:plazi:treatment:C74FFC13303EFF9BFF68544DDD82FA37:4A391BDE303EFF9EFC4F5420D927F918" box="[944,1091,1663,1689]" country="South Africa" county="Cederberg" latitude="-32.518726" longLatPrecision="1" longitude="19.350111" municipality="Matjies Rivier" name="Matjiesrivier" pageId="2" pageNumber="541" stateProvince="Western Cape">Matjiesrivier</location>
,
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,
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(
<figureCitation id="D7DD5180303EFF9EFF0B54FCDC5DF93C" box="[244,313,1699,1725]" captionStart="FIG" captionStartId="11.[151,196,1943,1967]" captionTargetBox="[211,1375,201,1919]" captionTargetId="figure-19@11.[164,1423,181,1919]" captionTargetPageId="11" captionText="FIG. 9. Rhithrops capensis gen. et sp. nov. A) aedeagus, ventrolateral view; B) aedeagus, lateral view; C) male sternite 9; D) aedeagal apex, ventral view (not to scale); E) male tergite 8; F) male sternite 8. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7223912" httpUri="https://zenodo.org/record/7223912/files/figure.png" pageId="2" pageNumber="541">Fig. 9</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="4F594D05303EFF9EFF385498DE0FF884" blockId="2.[151,1437,1662,1977]" pageId="2" pageNumber="541">
<emphasis id="7D929117303EFF9EFF385498DC1CF961" bold="true" box="[199,376,1734,1761]" pageId="2" pageNumber="541">Type material.</emphasis>
<materialsCitation id="FF8E4758303EFF9EFE7C5498DE0FF884" ID-GBIF-Occurrence="3926284306" collectingDate="2011-09-22" collectionCode="AMG" collectorName="D T Bilton" country="South Africa" county="Matjiesrivier" elevation="725" latitude="-32.518726" location="Cederberg" longLatPrecision="1" longitude="19.350111" municipality="Matjies Rivier" pageId="2" pageNumber="541" specimenCount="725" typeStatus="holotype">
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<typeStatus id="905DF3A7303EFF9EFE7C5498DC94F960" box="[387,496,1735,1761]" pageId="2" pageNumber="541" type="holotype">Holotype</typeStatus>
</emphasis>
(male): “
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<collectingDate id="2B1C922D303EFF9EFDA05498DFBEF960" box="[607,730,1735,1761]" pageId="2" pageNumber="541" value="2011-09-22">22/ix/2011</collectingDate>
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<collectingCountry id="37F10D95303EFF9EFD1A5498DE13F960" box="[741,887,1735,1761]" name="South Africa" pageId="2" pageNumber="541">South Africa</collectingCountry>
WC//
<location id="4A391BDE303EFF9EFC375498D95BF960" LSID="urn:lsid:plazi:treatment:C74FFC13303EFF9BFF68544DDD82FA37:4A391BDE303EFF9EFC375498D95BF960" box="[968,1087,1735,1761]" country="South Africa" county="Matjiesrivier" latitude="-32.518726" longLatPrecision="1" longitude="19.350111" municipality="Matjies Rivier" name="Cederberg" pageId="2" pageNumber="541">Cederberg</location>
,
<collectingMunicipality id="AF3DD77F303EFF9EFBB25498D995F960" box="[1101,1265,1735,1761]" pageId="2" pageNumber="541">Matjies Rivier</collectingMunicipality>
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<collectingCounty id="A6383589303EFF9EFAF35498D8F9F960" box="[1292,1437,1735,1761]" pageId="2" pageNumber="541">Matjiesrivier</collectingCounty>
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//
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<specimenCount id="59E0868C303EFF9EFE0154B5DF28F884" box="[510,588,1770,1797]" pageId="2" pageNumber="541" type="generic">
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<collectorName id="E21328D3303EFF9EFDAC54B4DFAFF884" box="[595,715,1771,1797]" pageId="2" pageNumber="541">D T Bilton</collectorName>
leg.” (
<collectionCode id="29F7D5C0303EFF9EFCE854B4DE3BF884" box="[791,863,1771,1797]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="2" pageNumber="541">AMG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="4F594D05303EFF9EFF385550DF97F838" blockId="2.[151,1437,1662,1977]" pageId="2" pageNumber="541">
<materialsCitation id="FF8E4758303EFF9EFF385550DFB8F8A8" ID-GBIF-Occurrence="3926284303" box="[199,732,1806,1833]" collectingDate="2011-09-22" collectionCode="AMG" collectorName="D T Bilton" country="South Africa" county="Matjiesrivier" elevation="725" latitude="-32.518726" location="Cederberg" longLatPrecision="1" longitude="19.350111" municipality="Matjies Rivier" pageId="2" pageNumber="541" specimenCount="15" specimenCount-female="10" specimenCount-male="5" typeStatus="paratype">
<emphasis id="7D929117303EFF9EFF385550DC5BF8A8" bold="true" box="[199,319,1807,1833]" pageId="2" pageNumber="541">
<typeStatus id="905DF3A7303EFF9EFF385550DC5BF8A8" box="[199,319,1807,1833]" pageId="2" pageNumber="541" type="paratype">Paratypes</typeStatus>
</emphasis>
(23):
<specimenCount id="59E0868C303EFF9EFE7F5551DCC1F8A8" box="[384,421,1806,1833]" pageId="2" pageNumber="541" type="male">5♂</specimenCount>
,
<specimenCount id="59E0868C303EFF9EFE4F5550DC84F8A8" box="[432,480,1807,1833]" pageId="2" pageNumber="541" type="female">10♀</specimenCount>
same data as holotype
</materialsCitation>
;
<materialsCitation id="FF8E4758303EFF9EFD1A5550DEA5F8CC" ID-GBIF-Occurrence="3926284301" collectingDate="2010-09-21" collectorName="D T Bilton" country="South Africa" county="Cederberg" elevation="725" latitude="-32.518726" location="Matjiesrivier" longLatPrecision="1" longitude="19.350111" municipality="Matjies Rivier" pageId="2" pageNumber="541" specimenCount="726" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="59E0868C303EFF9EFD1A5550DE6CF8A8" box="[741,776,1807,1833]" pageId="2" pageNumber="541" type="female">1♀</specimenCount>
“
<date id="3B586BC5303EFF9EFCE75550DEF7F8A8" box="[792,915,1807,1833]" pageId="2" pageNumber="541" value="2010-09-21">
<collectingDate id="2B1C922D303EFF9EFCE75550DEF7F8A8" box="[792,915,1807,1833]" pageId="2" pageNumber="541" value="2010-09-21">21/ix/2010</collectingDate>
</date>
<collectingCountry id="37F10D95303EFF9EFC665550D942F8A8" box="[921,1062,1807,1833]" name="South Africa" pageId="2" pageNumber="541">South Africa</collectingCountry>
WC//
<collectingCounty id="A6383589303EFF9EFB925550D980F8A8" box="[1133,1252,1807,1833]" pageId="2" pageNumber="541">Cederberg</collectingCounty>
,
<collectingMunicipality id="AF3DD77F303EFF9EFB125550D8E8F8A8" box="[1261,1420,1807,1833]" pageId="2" pageNumber="541">Matjies Rivier</collectingMunicipality>
//
<location id="4A391BDE303EFF9EFF68556CDC4CF8CC" LSID="urn:lsid:plazi:treatment:C74FFC13303EFF9BFF68544DDD82FA37:4A391BDE303EFF9EFF68556CDC4CF8CC" box="[151,296,1843,1869]" country="South Africa" county="Cederberg" latitude="-32.518726" longLatPrecision="1" longitude="19.350111" municipality="Matjies Rivier" name="Matjiesrivier" pageId="2" pageNumber="541">Matjiesrivier</location>
<geoCoordinate id="2AD22BC2303EFF9EFECC556CDCB3F8CC" box="[307,471,1843,1869]" degrees="32" direction="south" minutes="31" orientation="latitude" pageId="2" pageNumber="541" precision="1" seconds="07.42" value="-32.518726">32°31’07.42”S</geoCoordinate>
//
<geoCoordinate id="2AD22BC2303EFF9EFE0A556CDFF6F8CC" box="[501,658,1843,1869]" degrees="19" direction="east" minutes="21" orientation="longitude" pageId="2" pageNumber="541" precision="1" seconds="00.4" value="19.350111">19°21’00.4”E</geoCoordinate>
,
<specimenCount id="59E0868C303EFF9EFD5E556DDF97F8CC" box="[673,755,1842,1869]" pageId="2" pageNumber="541" type="generic">
<quantity id="881EE0E0303EFF9EFD5E556DDF8BF8CC" box="[673,751,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="2" pageNumber="541" unit="m" value="725.0">
<elevation id="C4CBAA36303EFF9EFD5E556DDF8BF8CC" box="[673,751,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="2" pageNumber="541" unit="m" value="725.0">725 m</elevation>
</quantity>
.
</specimenCount>
<collectorName id="E21328D3303EFF9EFD01556CDE1AF8CC" box="[766,894,1843,1869]" pageId="2" pageNumber="541">D T Bilton</collectorName>
leg.”
</materialsCitation>
;
<materialsCitation id="FF8E4758303EFF9EFC2E556CD981F8F0" ID-GBIF-Occurrence="3926284305" collectingDate="2011-09-22" collectorName="D T Bilton" country="South Africa" county="Cederberg" elevation="725" latitude="-32.518726" location="Matjiesrivier" longLatPrecision="1" longitude="19.350111" municipality="Matjies Rivier" pageId="2" pageNumber="541" specimenCount="730" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="59E0868C303EFF9EFC2E556CDE92F8CC" box="[977,1014,1843,1869]" pageId="2" pageNumber="541" type="male">2♂</specimenCount>
,
<specimenCount id="59E0868C303EFF9EFBF9556CD94CF8CC" box="[1030,1064,1843,1869]" pageId="2" pageNumber="541" type="female">3♀</specimenCount>
“
<date id="3B586BC5303EFF9EFBC2556CD9DCF8CC" box="[1085,1208,1843,1869]" pageId="2" pageNumber="541" value="2011-09-22">
<collectingDate id="2B1C922D303EFF9EFBC2556CD9DCF8CC" box="[1085,1208,1843,1869]" pageId="2" pageNumber="541" value="2011-09-22">22/ix/2011</collectingDate>
</date>
<collectingCountry id="37F10D95303EFF9EFB3C556CD832F8CC" box="[1219,1366,1843,1869]" name="South Africa" pageId="2" pageNumber="541">South Africa</collectingCountry>
WC//
<collectingCounty id="A6383589303EFF9EFF685508DC6AF8F0" box="[151,270,1879,1905]" pageId="2" pageNumber="541">Cederberg</collectingCounty>
,
<collectingMunicipality id="AF3DD77F303EFF9EFEE55508DCDFF8F0" box="[282,443,1879,1905]" pageId="2" pageNumber="541">Matjies Rivier</collectingMunicipality>
//
<location id="4A391BDE303EFF9EFE2C5508DF00F8F0" LSID="urn:lsid:plazi:treatment:C74FFC13303EFF9BFF68544DDD82FA37:4A391BDE303EFF9EFE2C5508DF00F8F0" box="[467,612,1879,1905]" country="South Africa" county="Cederberg" latitude="-32.518726" longLatPrecision="1" longitude="19.350111" municipality="Matjies Rivier" name="Matjiesrivier" pageId="2" pageNumber="541">Matjiesrivier</location>
<geoCoordinate id="2AD22BC2303EFF9EFD935508DE74F8F0" box="[620,784,1879,1905]" degrees="32" direction="south" minutes="31" orientation="latitude" pageId="2" pageNumber="541" precision="1" seconds="07.42" value="-32.518726">32°31’07.42”S</geoCoordinate>
//
<geoCoordinate id="2AD22BC2303EFF9EFCD45508DEACF8F0" box="[811,968,1879,1905]" degrees="19" direction="east" minutes="21" orientation="longitude" pageId="2" pageNumber="541" precision="1" seconds="00.4" value="19.350111">19°21’00.4”E</geoCoordinate>
,
<specimenCount id="59E0868C303EFF9EFC2B5509D947F8F0" box="[980,1059,1878,1905]" pageId="2" pageNumber="541" type="generic">
<quantity id="881EE0E0303EFF9EFC2B5509D97BF8F0" box="[980,1055,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="2" pageNumber="541" unit="m" value="725.0">
<elevation id="C4CBAA36303EFF9EFC2B5509D97BF8F0" box="[980,1055,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="2" pageNumber="541" unit="m" value="725.0">725 m</elevation>
</quantity>
.
</specimenCount>
<collectorName id="E21328D3303EFF9EFBD45508D9C1F8F0" box="[1067,1189,1879,1905]" pageId="2" pageNumber="541">D T Bilton</collectorName>
leg.”
</materialsCitation>
;
<materialsCitation id="FF8E4758303EFF9EFB0D5508DF8BF838" ID-GBIF-Occurrence="3926284304" collectingDate="2004-12-28" collectingDateMax="2004-12-29" collectingDateMin="2004-12-28" collectionCode="AMG, CBP, DMSA, EMEC, NMW" collectorName="P. S. Cranston" country="South Africa" county="Cederberg" elevation="820" latitude="-32.494446" location="Matjiesrivier" longLatPrecision="20" longitude="19.280277" municipality="Maalgat" pageId="2" pageNumber="541" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="59E0868C303EFF9EFB0D5508D872F8F0" box="[1266,1302,1879,1905]" pageId="2" pageNumber="541" type="male">1♂</specimenCount>
,
<specimenCount id="59E0868C303EFF9EFADB5508D822F8F0" box="[1316,1350,1879,1905]" pageId="2" pageNumber="541" type="female">1♀</specimenCount>
“
<collectingCountry id="37F10D95303EFF9EFAA75508DD87F814" name="South Africa" pageId="2" pageNumber="541">South Africa</collectingCountry>
: W.
<collectingRegion id="8D2283E7303EFF9EFEEB5524DC28F814" box="[276,332,1915,1941]" country="South Africa" name="Western Cape" pageId="2" pageNumber="541">Cape</collectingRegion>
//
<collectingCounty id="A6383589303EFF9EFE9A5524DCB8F814" box="[357,476,1915,1941]" pageId="2" pageNumber="541">Cederberg</collectingCounty>
,
<collectingMunicipality id="AF3DD77F303EFF9EFE175524DF26F814" box="[488,578,1915,1941]" pageId="2" pageNumber="541">Maalgat</collectingMunicipality>
//
<location id="4A391BDE303EFF9EFDA45524DF8FF814" LSID="urn:lsid:plazi:treatment:C74FFC13303EFF9BFF68544DDD82FA37:4A391BDE303EFF9EFDA45524DF8FF814" box="[603,747,1915,1941]" country="South Africa" county="Cederberg" latitude="-32.494446" longLatPrecision="20" longitude="19.280277" municipality="Maalgat" name="Matjiesrivier" pageId="2" pageNumber="541" stateProvince="Western Cape">Matjiesrivier</location>
//
<date id="3B586BC5303EFF9EFCFC5524DED7F814" box="[771,947,1915,1941]" pageId="2" pageNumber="541" value="2004-12-28" valueMax="2004-12-29" valueMin="2004-12-28">
<collectingDate id="2B1C922D303EFF9EFCFC5524DED7F814" box="[771,947,1915,1941]" pageId="2" pageNumber="541" value="2004-12-28" valueMax="2004-12-29" valueMin="2004-12-28">28-29 XII 2004</collectingDate>
</date>
<elevation id="C4CBAA36303EFF9EFC445524D94CF814" box="[955,1064,1915,1941]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="541" unit="m" value="820.0">
<quantity id="881EE0E0303EFF9EFC445524D967F814" box="[955,1027,1915,1941]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="541" unit="m" value="820.0">820 m</quantity>
asl
</elevation>
//
<geoCoordinate id="2AD22BC2303EFF9EFBBE5524D9A1F814" box="[1089,1221,1915,1941]" degrees="32" direction="south" minutes="29" orientation="latitude" pageId="2" pageNumber="541" precision="15" seconds="40" value="-32.494446">32°29’40”S</geoCoordinate>
<geoCoordinate id="2AD22BC2303EFF9EFB325524D82BF814" box="[1229,1359,1915,1941]" degrees="19" direction="east" minutes="16" orientation="longitude" pageId="2" pageNumber="541" precision="15" seconds="49" value="19.280277">19°16’49”E</geoCoordinate>
//
<collectorName id="E21328D3303EFF9EFA955524DD9BF838" pageId="2" pageNumber="541">P. S. Cranston</collectorName>
, leg.” (
<collectionCode id="29F7D5C0303EFF9EFEB055C0DCF1F838" box="[335,405,1951,1977]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="2" pageNumber="541">AMG</collectionCode>
,
<collectionCode id="29F7D5C0303EFF9EFE5F55C0DCB1F838" box="[416,469,1951,1977]" pageId="2" pageNumber="541">CBP</collectionCode>
,
<collectionCode id="29F7D5C0303EFF9EFE2055C0DF57F838" box="[479,563,1951,1977]" country="South Africa" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="2" pageNumber="541">DMSA</collectionCode>
,
<collectionCode id="29F7D5C0303EFF9EFDC155C0DFEBF838" box="[574,655,1951,1977]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="2" pageNumber="541" type="University or college">EMEC</collectionCode>
,
<collectionCode id="29F7D5C0303EFF9EFD6555C0DF83F838" box="[666,743,1951,1977]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="541">NMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="1B991D8D303FFF9FFF6755C8DEB7F82E" ID-DOI="http://doi.org/10.5281/zenodo.7223888" ID-Zenodo-Dep="7223888" box="[152,979,1943,1968]" httpUri="https://zenodo.org/record/7223888/files/figure.png" pageId="3" pageNumber="542" startId="3.[152,201,1943,1967]" targetBox="[199,1378,203,1900]" targetPageId="3">
<paragraph id="4F594D05303FFF9FFF6755C8DEB7F82E" blockId="3.[152,979,1943,1968]" box="[152,979,1943,1968]" pageId="3" pageNumber="542">
<emphasis id="7D929117303FFF9FFF6755C8DD83F82E" bold="true" box="[152,231,1943,1967]" pageId="3" pageNumber="542">FIG. 1.</emphasis>
<taxonomicName id="88E63686303FFF9FFF1255C7DCC9F82E" authority="Bilton &amp; Shepard, 2022" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[237,429,1944,1968]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="capensis" status="gen. et sp. nov.">
<emphasis id="7D929117303FFF9FFF1255C7DCC9F82E" box="[237,429,1944,1968]" italics="true" pageId="3" pageNumber="542">Rhithrops capensis</emphasis>
</taxonomicName>
<emphasis id="7D929117303FFF9FFE4B55C7DF35F831" bold="true" box="[436,593,1944,1968]" pageId="3" pageNumber="542">
<taxonomicNameLabel id="66A12C6C303FFF9FFE4B55C7DF35F831" box="[436,593,1944,1968]" pageId="3" pageNumber="542" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, Holotype habitus. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="1B991D8D3038FF98FF6855C8DED4F876" ID-DOI="http://doi.org/10.5281/zenodo.7223890" ID-Zenodo-Dep="7223890" httpUri="https://zenodo.org/record/7223890/files/figure.png" pageId="4" pageNumber="543" startId="4.[151,196,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="4">
<paragraph id="4F594D053038FF98FF6855C8DED4F876" blockId="4.[151,1436,1943,2040]" pageId="4" pageNumber="543">
<emphasis id="7D9291173038FF98FF6855C8DD80F82E" bold="true" box="[151,228,1943,1968]" pageId="4" pageNumber="543">FIG. 2.</emphasis>
South African endemic dryopid genera, dorsolateral view of pronotal margins. A)
<taxonomicName id="88E636863038FF98FBD055C7D994F82E" authority="Bilton &amp; Shepard, 2022" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[1071,1264,1944,1968]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="capensis" status="gen. et sp. nov.">
<emphasis id="7D9291173038FF98FBD055C7D994F82E" box="[1071,1264,1944,1968]" italics="true" pageId="4" pageNumber="543">Rhithrops capensis</emphasis>
</taxonomicName>
<emphasis id="7D9291173038FF98FB0855C7D8F1F831" bold="true" box="[1271,1429,1944,1968]" pageId="4" pageNumber="543">
<taxonomicNameLabel id="66A12C6C3038FF98FB0855C7D8F1F831" box="[1271,1429,1944,1968]" pageId="4" pageNumber="543" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male; B)
<taxonomicName id="88E636863038FF98FEAA55E3DF72F852" authority="Bilton &amp; Shepard, 2022" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[341,534,1980,2004]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="capensis" status="gen. et sp. nov.">
<emphasis id="7D9291173038FF98FEAA55E3DF72F852" box="[341,534,1980,2004]" italics="true" pageId="4" pageNumber="543">Rhithrops capensis</emphasis>
</taxonomicName>
<emphasis id="7D9291173038FF98FDE355E3DFDEF855" bold="true" box="[540,698,1980,2004]" pageId="4" pageNumber="543">
<taxonomicNameLabel id="66A12C6C3038FF98FDE355E3DFDEF855" box="[540,698,1980,2004]" pageId="4" pageNumber="543" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype female; C)
<taxonomicName id="88E636863038FF98FC6855E3D96DF852" box="[919,1033,1980,2003]" class="Insecta" family="Dryopidae" genus="Rapnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="7D9291173038FF98FC6855E3DE87F852" box="[919,995,1980,2003]" italics="true" pageId="4" pageNumber="543">Rapnus</emphasis>
sp.
</taxonomicName>
, female; D)
<taxonomicName id="88E636863038FF98FB7855E3D98FF852" box="[1159,1259,1980,2003]" class="Insecta" family="Dryopidae" genus="Strina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="7D9291173038FF98FB7855E3D9A1F852" box="[1159,1221,1980,2003]" italics="true" pageId="4" pageNumber="543">Strina</emphasis>
sp.
</taxonomicName>
, female. C) &amp; D) South Africa, Western Cape, Franschoek, Mont Rochelle. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="1B991D8D3039FF99FF685434DDB5F94A" ID-DOI="http://doi.org/10.5281/zenodo.7223894" ID-Zenodo-Dep="7223894" httpUri="https://zenodo.org/record/7223894/files/figure.png" pageId="5" pageNumber="544" startId="5.[151,196,1643,1667]" targetBox="[154,1434,181,1619]" targetPageId="5">
<paragraph id="4F594D053039FF99FF685434DDB5F94A" blockId="5.[151,1437,1643,1739]" pageId="5" pageNumber="544">
<emphasis id="7D9291173039FF99FF685434DD80F902" bold="true" box="[151,228,1643,1667]" pageId="5" pageNumber="544">FIG. 3.</emphasis>
<taxonomicName id="88E636863039FF99FF145434DCCFF902" authority="Bilton &amp; Shepard, 2022" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[235,427,1643,1667]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="species" species="capensis" status="gen. et sp. nov.">
<emphasis id="7D9291173039FF99FF145434DCCFF902" box="[235,427,1643,1667]" italics="true" pageId="5" pageNumber="544">Rhithrops capensis</emphasis>
</taxonomicName>
<emphasis id="7D9291173039FF99FE4D5434DF34F902" bold="true" box="[434,592,1643,1667]" pageId="5" pageNumber="544">
<taxonomicNameLabel id="66A12C6C3039FF99FE4D5434DF34F902" box="[434,592,1643,1667]" pageId="5" pageNumber="544" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
A) head, posterolateral view; B) compound eye – arrow indicates inter-ommatidial seta; C) detail of dorsolateral sculpture of head, inside compound eye; D) antennae; E) antennomeres 1–2; F) antennomeres 3–10.
</paragraph>
</caption>
<subSubSection id="07FC1E8E3039FF9BFF3854A6DC6EFAA7" lastPageId="7" lastPageNumber="546" pageId="5" pageNumber="544" type="description">
<paragraph id="4F594D053039FF99FF3854A6DE5AF895" blockId="5.[151,1437,1785,2027]" box="[199,830,1785,1812]" pageId="5" pageNumber="544">
<emphasis id="7D9291173039FF99FF3854A6DC3FF892" bold="true" box="[199,347,1785,1811]" pageId="5" pageNumber="544">Description:</emphasis>
With all the above characters of the genus.
</paragraph>
<paragraph id="4F594D053039FF99FF385542D91DF801" blockId="5.[151,1437,1785,2027]" pageId="5" pageNumber="544">
Size:
<typeStatus id="905DF3A73039FF99FEF75542DC11F8B6" box="[264,373,1821,1847]" pageId="5" pageNumber="544" type="holotype">Holotype</typeStatus>
: BL
<quantity id="881EE0E03039FF99FE535542DF70F8B9" box="[428,532,1821,1848]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="5" pageNumber="544" unit="mm" value="4.1">4.10 mm</quantity>
; EL
<quantity id="881EE0E03039FF99FDB45542DFD0F8B9" box="[587,692,1821,1848]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" pageId="5" pageNumber="544" unit="mm" value="2.65">2.65 mm</quantity>
; EW
<quantity id="881EE0E03039FF99FD0A5542DE39F8B9" box="[757,861,1821,1848]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="5" pageNumber="544" unit="mm" value="1.35">1.35 mm</quantity>
; PL
<quantity id="881EE0E03039FF99FC6C5542DE9FF8B9" box="[915,1019,1821,1848]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="544" unit="mm" value="0.85">0.85 mm</quantity>
; PW
<quantity id="881EE0E03039FF99FBC55542D9C7F8B9" box="[1082,1187,1821,1848]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="544" unit="mm" value="1.2">1.20 mm</quantity>
.
<typeStatus id="905DF3A73039FF99FB505541D844F8B9" box="[1199,1312,1822,1848]" pageId="5" pageNumber="544" type="paratype">Paratypes</typeStatus>
: Males BL
<quantity id="881EE0E03039FF99FF68551EDC5AF8DD" box="[151,318,1857,1884]" metricMagnitude="-3" metricUnit="m" metricValue="4.575" metricValueMax="4.75" metricValueMin="4.4" pageId="5" pageNumber="544" unit="mm" value="4.575" valueMax="4.75" valueMin="4.4">4.40–4.75 mm</quantity>
; EL 2.60–3.00 mm; EW
<quantity id="881EE0E03039FF99FDA5551EDE65F8DD" box="[602,769,1857,1884]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.6" metricValueMin="1.45" pageId="5" pageNumber="544" unit="mm" value="1.525" valueMax="1.6" valueMin="1.45">1.45–1.60 mm</quantity>
; PL
<quantity id="881EE0E03039FF99FCCA551EDEB8F8DD" box="[821,988,1857,1884]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="10.5" metricValueMin="8.5" pageId="5" pageNumber="544" unit="mm" value="0.95" valueMax="1.05" valueMin="0.85">0.85–1.05 mm</quantity>
; PW
<quantity id="881EE0E03039FF99FBE5551ED9A5F8DD" box="[1050,1217,1857,1884]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.35" metricValueMin="1.2" pageId="5" pageNumber="544" unit="mm" value="1.275" valueMax="1.35" valueMin="1.2">1.20–1.35 mm</quantity>
. Females BL
<quantity id="881EE0E03039FF99FAA2551EDD9AF801" metricMagnitude="-3" metricUnit="m" metricValue="4.725" metricValueMax="4.8" metricValueMin="4.65" pageId="5" pageNumber="544" unit="mm" value="4.725" valueMax="4.8" valueMin="4.65">4.65– 4.80 mm</quantity>
; EL
<quantity id="881EE0E03039FF99FECD553ADCBCF801" box="[306,472,1893,1920]" metricMagnitude="-3" metricUnit="m" metricValue="2.825" metricValueMax="2.9" metricValueMin="2.75" pageId="5" pageNumber="544" unit="mm" value="2.825" valueMax="2.9" valueMin="2.75">2.75–2.90 mm</quantity>
; EW
<quantity id="881EE0E03039FF99FDEA553ADFDFF801" box="[533,699,1893,1920]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="5" pageNumber="544" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.40–1.60 mm</quantity>
; PL
<quantity id="881EE0E03039FF99FD12553ADEF7F801" box="[749,915,1893,1920]" metricMagnitude="-4" metricUnit="m" metricValue="9.75" metricValueMax="10.5" metricValueMin="9.0" pageId="5" pageNumber="544" unit="mm" value="0.975" valueMax="1.05" valueMin="0.9">0.90–1.05 mm</quantity>
; PW
<quantity id="881EE0E03039FF99FC30553AD911F801" box="[975,1141,1893,1920]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.35" metricValueMin="1.25" pageId="5" pageNumber="544" unit="mm" value="1.3" valueMax="1.35" valueMin="1.25">1.25–1.35 mm</quantity>
.
</paragraph>
<paragraph id="4F594D053039FF99FF3855D6DDBFF86A" blockId="5.[151,1437,1785,2027]" pageId="5" pageNumber="544">Head: Epicranium densely and shallowly punctate, with short decumbent setae, epicranial suture absent; frontal area twice as wide as one compound eye, antennal ridges absent, frontoclypeal suture straight between antennal bases.</paragraph>
<caption id="1B991D8D303AFF9AFF685411DDAAF92F" ID-DOI="http://doi.org/10.5281/zenodo.7223899" ID-Zenodo-Dep="7223899" httpUri="https://zenodo.org/record/7223899/files/figure.png" pageId="6" pageNumber="545" startId="6.[151,196,1614,1638]" targetBox="[167,1421,181,1590]" targetPageId="6">
<paragraph id="4F594D05303AFF9AFF685411DDAAF92F" blockId="6.[151,1437,1614,1710]" pageId="6" pageNumber="545">
<emphasis id="7D929117303AFF9AFF685411DD86F9E7" bold="true" box="[151,226,1614,1638]" pageId="6" pageNumber="545">FIG. 4.</emphasis>
<taxonomicName id="88E63686303AFF9AFF185411DCC1F9E7" authority="Bilton &amp; Shepard, 2022" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[231,421,1614,1638]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="capensis" status="gen. et sp. nov.">
<emphasis id="7D929117303AFF9AFF185411DCC1F9E7" box="[231,421,1614,1638]" italics="true" pageId="6" pageNumber="545">Rhithrops capensis</emphasis>
</taxonomicName>
<emphasis id="7D929117303AFF9AFE555411DF25F9E7" bold="true" box="[426,577,1614,1638]" pageId="6" pageNumber="545">
<taxonomicNameLabel id="66A12C6C303AFF9AFE555411DF25F9E7" box="[426,577,1614,1638]" pageId="6" pageNumber="545" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
A) labrum, dorsal view; B) right mandible, ventral view; C) right maxilla, dorsal view; D) close-up of maxillary palp apex; E) female labrum and mentum, ventral view; F) male labrum and mentum, anteroventral view.
</paragraph>
</caption>
<paragraph id="4F594D05303AFF9AFF385483D835F977" blockId="6.[151,1436,1756,2035]" box="[199,1361,1756,1783]" pageId="6" pageNumber="545">
Elytra: Conjointly twice as long as wide, parallel-sided, apices moderately protruding (
<figureCitation id="D7DD5180303AFF9AFB6B5483D9BFF976" box="[1172,1243,1756,1783]" captionStart="FIG" captionStartId="3.[152,201,1943,1967]" captionTargetBox="[199,1378,203,1900]" captionTargetId="figure-19@3.[187,1401,179,1918]" captionTargetPageId="3" captionText="FIG. 1. Rhithrops capensis gen. et sp. nov., Holotype habitus. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7223888" httpUri="https://zenodo.org/record/7223888/files/figure.png" pageId="6" pageNumber="545">Figs 1</figureCitation>
&amp;
<figureCitation id="D7DD5180303AFF9AFB005483D821F977" box="[1279,1349,1756,1782]" captionStart="FIG" captionStartId="8.[151,196,1842,1866]" captionTargetBox="[154,1434,381,1819]" captionTargetId="figure-126@8.[154,1434,381,1819]" captionTargetPageId="8" captionText="FIG. 6. Rhithrops capensis gen. et sp. nov. A) prothorax, dorsal view; B) close-up of sculpture of pronotal disc; C) Meso and metanotum, with scutellum; D) right elytron, dorsolateral view; E) detail of right elytral apex; F) prothorax, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7223905" httpUri="https://zenodo.org/record/7223905/files/figure.png" pageId="6" pageNumber="545">6D–E</figureCitation>
).
</paragraph>
<paragraph id="4F594D05303AFF9AFF38555ED9AAF89A" blockId="6.[151,1436,1756,2035]" box="[199,1230,1792,1819]" pageId="6" pageNumber="545">
Legs: Relatively long, elongate, with large, strongly-developed tarsi and claws (
<figureCitation id="D7DD5180303AFF9AFBB9555FD9A6F89A" box="[1094,1218,1792,1819]" captionStart="FIG" captionStartId="10.[151,196,1643,1667]" captionTargetBox="[154,1434,181,1619]" captionTargetId="figure-25@10.[154,1434,181,1619]" captionTargetPageId="10" captionText="FIG. 8. Rhithrops capensis gen. et sp. nov. A) male foreleg, ventral view; B) male foretarsus and tibial apex, ventral view; C) male hind tibia and tarsus, ventral view; D) aedeagus, ventral view; E) aedeagus, lateral view; F) aedeagal apex, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7223910" httpUri="https://zenodo.org/record/7223910/files/figure.png" pageId="6" pageNumber="545">Fig. 8A–C</figureCitation>
).
</paragraph>
<paragraph id="4F594D05303AFF9AFF38557BD9D8F806" blockId="6.[151,1436,1756,2035]" pageId="6" pageNumber="545">
Male: Last abdominal ventrite with broad, shallow apicomedial emargination (
<figureCitation id="D7DD5180303AFF9AFBCE557BD9ECF8BE" box="[1073,1160,1828,1855]" captionStart="FIG" captionStartId="9.[151,196,1643,1667]" captionTargetBox="[154,1434,181,1619]" captionTargetId="figure-19@9.[154,1434,181,1619]" captionTargetPageId="9" captionText="FIG. 7. Rhithrops capensis gen. et sp. nov. A) meso and metaventrites, ventral view; B) male abdomen, ventral view; C) female abdomen, ventral view; D) close-up of plastron vestiture at junction of abdominal ventrites 2–3; E) male abdominal apex; F) female abdominal apex." figureDoi="http://doi.org/10.5281/zenodo.7223907" httpUri="https://zenodo.org/record/7223907/files/figure.png" pageId="6" pageNumber="545">Fig. 7E</figureCitation>
). Labium with modified, spathulate setae on anteroventral margin (
<figureCitation id="D7DD5180303AFF9AFD895517DFAAF8E2" box="[630,718,1864,1891]" captionStart="FIG" captionStartId="6.[151,196,1614,1638]" captionTargetBox="[167,1421,181,1590]" captionTargetId="figure-25@6.[167,1421,181,1590]" captionTargetPageId="6" captionText="FIG. 4. Rhithrops capensis gen. et sp. nov. A) labrum, dorsal view; B) right mandible, ventral view; C) right maxilla, dorsal view; D) close-up of maxillary palp apex; E) female labrum and mentum, ventral view; F) male labrum and mentum, anteroventral view." figureDoi="http://doi.org/10.5281/zenodo.7223899" httpUri="https://zenodo.org/record/7223899/files/figure.png" pageId="6" pageNumber="545">Fig. 4F</figureCitation>
). Anterior edge of sternite 8 with short median strut (
<figureCitation id="D7DD5180303AFF9AFAC85517D8EAF8E2" box="[1335,1422,1864,1891]" captionStart="FIG" captionStartId="11.[151,196,1943,1967]" captionTargetBox="[211,1375,201,1919]" captionTargetId="figure-19@11.[164,1423,181,1919]" captionTargetPageId="11" captionText="FIG. 9. Rhithrops capensis gen. et sp. nov. A) aedeagus, ventrolateral view; B) aedeagus, lateral view; C) male sternite 9; D) aedeagal apex, ventral view (not to scale); E) male tergite 8; F) male sternite 8. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7223912" httpUri="https://zenodo.org/record/7223912/files/figure.png" pageId="6" pageNumber="545">Fig. 9F</figureCitation>
); sternite 9 as in
<figureCitation id="D7DD5180303AFF9AFEBE5533DCFCF806" box="[321,408,1900,1927]" captionStart="FIG" captionStartId="11.[151,196,1943,1967]" captionTargetBox="[211,1375,201,1919]" captionTargetId="figure-19@11.[164,1423,181,1919]" captionTargetPageId="11" captionText="FIG. 9. Rhithrops capensis gen. et sp. nov. A) aedeagus, ventrolateral view; B) aedeagus, lateral view; C) male sternite 9; D) aedeagal apex, ventral view (not to scale); E) male tergite 8; F) male sternite 8. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7223912" httpUri="https://zenodo.org/record/7223912/files/figure.png" pageId="6" pageNumber="545">Fig. 9C</figureCitation>
. Tergite 8 with numerous long setae close to posterior margin (
<figureCitation id="D7DD5180303AFF9AFBA65533D9D4F806" box="[1113,1200,1900,1927]" captionStart="FIG" captionStartId="11.[151,196,1943,1967]" captionTargetBox="[211,1375,201,1919]" captionTargetId="figure-19@11.[164,1423,181,1919]" captionTargetPageId="11" captionText="FIG. 9. Rhithrops capensis gen. et sp. nov. A) aedeagus, ventrolateral view; B) aedeagus, lateral view; C) male sternite 9; D) aedeagal apex, ventral view (not to scale); E) male tergite 8; F) male sternite 8. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7223912" httpUri="https://zenodo.org/record/7223912/files/figure.png" pageId="6" pageNumber="545">Fig. 9E</figureCitation>
).
</paragraph>
<paragraph id="4F594D05303AFF9AFF3855CFD9D9F872" blockId="6.[151,1436,1756,2035]" pageId="6" pageNumber="545">
Aedeagus: Long and tubular (
<figureCitation id="D7DD5180303AFF9AFDE955CFDFEBF82A" box="[534,655,1936,1963]" captionStart="FIG" captionStartId="10.[151,196,1643,1667]" captionTargetBox="[154,1434,181,1619]" captionTargetId="figure-25@10.[154,1434,181,1619]" captionTargetPageId="10" captionText="FIG. 8. Rhithrops capensis gen. et sp. nov. A) male foreleg, ventral view; B) male foretarsus and tibial apex, ventral view; C) male hind tibia and tarsus, ventral view; D) aedeagus, ventral view; E) aedeagus, lateral view; F) aedeagal apex, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7223910" httpUri="https://zenodo.org/record/7223910/files/figure.png" pageId="6" pageNumber="545">Figs 8D–F</figureCitation>
&amp;
<figureCitation id="D7DD5180303AFF9AFD4B55CEDE57F82A" box="[692,819,1937,1963]" captionStart="FIG" captionStartId="11.[151,196,1943,1967]" captionTargetBox="[211,1375,201,1919]" captionTargetId="figure-19@11.[164,1423,181,1919]" captionTargetPageId="11" captionText="FIG. 9. Rhithrops capensis gen. et sp. nov. A) aedeagus, ventrolateral view; B) aedeagus, lateral view; C) male sternite 9; D) aedeagal apex, ventral view (not to scale); E) male tergite 8; F) male sternite 8. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7223912" httpUri="https://zenodo.org/record/7223912/files/figure.png" pageId="6" pageNumber="545">9A, B &amp; D</figureCitation>
); basal piece:paramere ratio 0.6:0.4. Basal piece hookshaped basally, open on left side. Median lobe long, narrow, acute apically. Parameres parallel-sided, apically a little longer than median lobe, apices rounded on outer side, clasping apex of median lobe (
<figureCitation id="D7DD5180303AFF9AFBA55587D9D4F872" box="[1114,1200,2008,2035]" captionStart="FIG" captionStartId="10.[151,196,1643,1667]" captionTargetBox="[154,1434,181,1619]" captionTargetId="figure-25@10.[154,1434,181,1619]" captionTargetPageId="10" captionText="FIG. 8. Rhithrops capensis gen. et sp. nov. A) male foreleg, ventral view; B) male foretarsus and tibial apex, ventral view; C) male hind tibia and tarsus, ventral view; D) aedeagus, ventral view; E) aedeagus, lateral view; F) aedeagal apex, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7223910" httpUri="https://zenodo.org/record/7223910/files/figure.png" pageId="6" pageNumber="545">Fig. 8F</figureCitation>
).
</paragraph>
<caption id="1B991D8D303BFF9BFF6851F9DE39FC63" ID-DOI="http://doi.org/10.5281/zenodo.7223903" ID-Zenodo-Dep="7223903" httpUri="https://zenodo.org/record/7223903/files/figure.png" pageId="7" pageNumber="546" startId="7.[151,196,934,958]" targetBox="[180,1407,209,861]" targetPageId="7">
<paragraph id="4F594D05303BFF9BFF6851F9DE39FC63" blockId="7.[151,1437,934,994]" pageId="7" pageNumber="546">
<emphasis id="7D929117303BFF9BFF6851F9DD80FC3F" bold="true" box="[151,228,934,958]" pageId="7" pageNumber="546">FIG. 5.</emphasis>
<taxonomicName id="88E63686303BFF9BFF1551F9DCCEFC3F" authority="Bilton &amp; Shepard, 2022" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[234,426,934,958]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="546" phylum="Arthropoda" rank="species" species="capensis" status="gen. et sp. nov.">
<emphasis id="7D929117303BFF9BFF1551F9DCCEFC3F" box="[234,426,934,958]" italics="true" pageId="7" pageNumber="546">Rhithrops capensis</emphasis>
</taxonomicName>
<emphasis id="7D929117303BFF9BFE4E51F9DF29FC3F" bold="true" box="[433,589,934,958]" pageId="7" pageNumber="546">
<taxonomicNameLabel id="66A12C6C303BFF9BFE4E51F9DF29FC3F" box="[433,589,934,958]" pageId="7" pageNumber="546" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
A) right mandible, ventrolateral view – teeth numbered as in text; B) left mandible, ventral view – pt = prostheca, Lf = lateral flange. Scale bar = 100 μm.
</paragraph>
</caption>
<paragraph id="4F594D05303BFF9BFF38564FDC35FB17" blockId="7.[151,1437,1040,1462]" pageId="7" pageNumber="546">
Female: Few external differences to males. The sublateral furrows of the pronotum are deeper and more evident (
<figureCitation id="D7DD5180303BFF9BFF60566BDD9FFBCF" box="[159,251,1076,1102]" captionStart="FIG" captionStartId="4.[151,196,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-25@4.[214,1373,181,1919]" captionTargetPageId="4" captionText="FIG. 2. South African endemic dryopid genera, dorsolateral view of pronotal margins. A) Rhithrops capensis gen. et sp. nov., holotype male; B) Rhithrops capensis gen. et sp. nov., paratype female; C) Rapnus sp., female; D) Strina sp., female. C) &amp; D) South Africa, Western Cape, Franschoek, Mont Rochelle. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7223890" httpUri="https://zenodo.org/record/7223890/files/figure.png" pageId="7" pageNumber="546">Fig. 2B</figureCitation>
). The last abdominal ventrite has a conical apex, somewhat raised centrally and lacks an apicomedian emargination (
<figureCitation id="D7DD5180303BFF9BFEC65607DCE9FBF3" box="[313,397,1112,1138]" captionStart="FIG" captionStartId="9.[151,196,1643,1667]" captionTargetBox="[154,1434,181,1619]" captionTargetId="figure-19@9.[154,1434,181,1619]" captionTargetPageId="9" captionText="FIG. 7. Rhithrops capensis gen. et sp. nov. A) meso and metaventrites, ventral view; B) male abdomen, ventral view; C) female abdomen, ventral view; D) close-up of plastron vestiture at junction of abdominal ventrites 2–3; E) male abdominal apex; F) female abdominal apex." figureDoi="http://doi.org/10.5281/zenodo.7223907" httpUri="https://zenodo.org/record/7223907/files/figure.png" pageId="7" pageNumber="546">Fig. 7F</figureCitation>
). The labium has fewer, shorter modified setae on the anteroventral margin (
<figureCitation id="D7DD5180303BFF9BFB235607D857FBF3" box="[1244,1331,1112,1138]" captionStart="FIG" captionStartId="6.[151,196,1614,1638]" captionTargetBox="[167,1421,181,1590]" captionTargetId="figure-25@6.[167,1421,181,1590]" captionTargetPageId="6" captionText="FIG. 4. Rhithrops capensis gen. et sp. nov. A) labrum, dorsal view; B) right mandible, ventral view; C) right maxilla, dorsal view; D) close-up of maxillary palp apex; E) female labrum and mentum, ventral view; F) male labrum and mentum, anteroventral view." figureDoi="http://doi.org/10.5281/zenodo.7223899" httpUri="https://zenodo.org/record/7223899/files/figure.png" pageId="7" pageNumber="546">Fig. 4E</figureCitation>
). Sternite 8 as in
<figureCitation id="D7DD5180303BFF9BFF185623DC29FB17" box="[231,333,1148,1174]" captionStart="FIG" captionStartId="12.[151,196,1943,1967]" captionTargetBox="[180,1387,247,1903]" captionTargetId="figure-25@12.[164,1423,181,1919]" captionTargetPageId="12" captionText="FIG. 10. Rhithrops capensis gen. et sp. nov. A) ovipositor, lateral view; B) ovipositor apex, lateral view (not to scale); C) female sternite 8. Scale bar (A &amp; C) = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7223916" httpUri="https://zenodo.org/record/7223916/files/figure.png" pageId="7" pageNumber="546">Fig. 10C</figureCitation>
.
</paragraph>
<paragraph id="4F594D05303BFF9BFF3856FFDC6EFAA7" blockId="7.[151,1437,1040,1462]" pageId="7" pageNumber="546">
Ovipositor: Elongate (
<figureCitation id="D7DD5180303BFF9BFE3356FFDF33FB3B" box="[460,599,1184,1210]" captionStart="FIG" captionStartId="12.[151,196,1943,1967]" captionTargetBox="[180,1387,247,1903]" captionTargetId="figure-25@12.[164,1423,181,1919]" captionTargetPageId="12" captionText="FIG. 10. Rhithrops capensis gen. et sp. nov. A) ovipositor, lateral view; B) ovipositor apex, lateral view (not to scale); C) female sternite 8. Scale bar (A &amp; C) = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7223916" httpUri="https://zenodo.org/record/7223916/files/figure.png" pageId="7" pageNumber="546">Fig. 10A–B</figureCitation>
) and strongly sclerotized;
<typeStatus id="905DF3A7303BFF9BFC6856FEDEA3FB3A" box="[919,967,1185,1211]" pageId="7" pageNumber="546">type</typeStatus>
1 (sensu
<bibRefCitation id="2B7730F4303BFF9BFBBD56FFD84FFB3B" author="Kodada, J. &amp; Jach, M. A. &amp; Ciampor, F." box="[1090,1323,1184,1211]" pageId="7" pageNumber="546" pagination="590 - 602" refId="ref5043" refString="Kodada, J., Jach, M. A. &amp; Ciampor, F. (2016) 19.3 Dryopidae Bilberg, 1820 (1817). In: Beutel, R. G. &amp; Leschen, R. A. B. (Eds.), Coleoptera, Beetles. Vol. 1. Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). In: Beutel, R. G. &amp; Kristensen, N. P. (Eds.), Handbook of Zoology, Arthropoda: Insecta. 2 nd Edition. Walter de Gruyter, Berlin, pp. 590 - 602." type="book chapter" year="2016">Kodada et al. 2016</bibRefCitation>
); apically sharp-tipped. Valvifers ventral, asymmetrical, long, thin, forming a V-shaped sclerite. Coxites asymmetrical, laterally compressed, blade-shaped, apex very acute, dorsal side serrate, with basally directed long narrow dorsal extension.
</paragraph>
</subSubSection>
<subSubSection id="07FC1E8E303BFF9BFF38576FDE2AFAEF" pageId="7" pageNumber="546" type="distribution">
<paragraph id="4F594D05303BFF9BFF38576FDE2AFAEF" blockId="7.[151,1437,1040,1462]" pageId="7" pageNumber="546">
<emphasis id="7D929117303BFF9BFF38576FDC8AFACB" bold="true" box="[199,494,1328,1354]" pageId="7" pageNumber="546">Distribution and habitat.</emphasis>
Known only from
<typeStatus id="905DF3A7303BFF9BFD40576EDF8BFACA" box="[703,751,1329,1355]" pageId="7" pageNumber="546">type</typeStatus>
locality, the Matjies River (
<figureCitation id="D7DD5180303BFF9BFBDE576FD915FACB" box="[1057,1137,1328,1354]" captionStart="FIG" captionStartId="13.[151,196,1108,1132]" captionTargetBox="[191,1396,181,1085]" captionTargetId="figure-19@13.[191,1396,181,1085]" captionTargetPageId="13" captionText="FIG. 11. Rhithrops capensis gen. et sp. nov. Type locality, South Africa, Western Cape, Cederberg, Matjies Rivier, Matjiesrivier, 32°31’07.42”S 19°21’00.4”E, 725 m. Photo D T Bilton." figureDoi="http://doi.org/10.5281/zenodo.7223918" httpUri="https://zenodo.org/record/7223918/files/figure.png" pageId="7" pageNumber="546">Fig. 11</figureCitation>
), a tributary of the Olifants River in the Cederberg,
<collectingRegion id="8D2283E7303BFF9BFE5B570ADFC9FAEF" box="[420,685,1364,1391]" country="South Africa" name="Western Cape" pageId="7" pageNumber="546">Western Cape Province</collectingRegion>
,
<collectingCountry id="37F10D95303BFF9BFD47570BDE2EFAEF" box="[696,842,1364,1390]" name="South Africa" pageId="7" pageNumber="546">South Africa</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="07FC1E8E303BFF9BFF385727DD82FA37" pageId="7" pageNumber="546" type="etymology">
<paragraph id="4F594D05303BFF9BFF385727DD82FA37" blockId="7.[151,1437,1040,1462]" pageId="7" pageNumber="546">
<emphasis id="7D929117303BFF9BFF385727DC2AFA13" bold="true" box="[199,334,1400,1426]" pageId="7" pageNumber="546">Etymology.</emphasis>
The trivial name,
<taxonomicName id="88E63686303BFF9BFDE65726DF1DFA13" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[537,633,1401,1426]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="546" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="7D929117303BFF9BFDE65726DF1DFA13" box="[537,633,1401,1426]" italics="true" pageId="7" pageNumber="546">capensis</emphasis>
</taxonomicName>
, refers to location of this species in the
<collectingRegion id="8D2283E7303BFF9BFBCD5726D853FA13" box="[1074,1335,1400,1427]" country="South Africa" name="Western Cape" pageId="7" pageNumber="546">Western Cape Province</collectingRegion>
of
<collectingCountry id="37F10D95303BFF9BFAA45727DD86FA37" name="South Africa" pageId="7" pageNumber="546">South Africa</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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