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<document id="F2271A5B9C1617F46935219F710FD1DC" ID-CLB-Dataset="168727" ID-DOI="10.11646/zootaxa.5195.6.3" ID-GBIF-Dataset="893c13e2-2785-463d-b114-a499cbe8a7c8" ID-ISSN="1175-5326" ID-Zenodo-Dep="7223884" ID-ZooBank="EB51F148-8EFC-4B22-9B74-1DBE4EB3A544" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666164886612" checkinUser="plazi" docAuthor="Bilton, David T. &amp; Shepard, William D." docDate="2022" docId="C74FFC13303DFF9DFF685645D8F6FA40" docLanguage="en" docName="zootaxa.5195.6.3.pdf" docOrigin="Zootaxa 5195 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhithrops Bilton &amp; Shepard 2022, gen. nov." docType="treatment" docVersion="5" lastPageNumber="540" masterDocId="3B76846B303CFF9CFFFF525FDD64FF81" masterDocTitle="Rhithrops capensis gen. et sp. nov., a new aquatic dryopid beetle from the Western Cape of South Africa (Coleoptera: Dryopoidea: Dryopidae)" masterLastPageNumber="553" masterPageNumber="539" pageNumber="540" updateTime="1699407958300" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="BF9EBB9E0300E4A93FB700CF0A04D5CE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2FE36F9DDA0E456A8B0EFB603A32B7C0">
<mods:title id="F9792B998D343DDFA170068114AD5EA3">Rhithrops capensis gen. et sp. nov., a new aquatic dryopid beetle from the Western Cape of South Africa (Coleoptera: Dryopoidea: Dryopidae)</mods:title>
</mods:titleInfo>
<mods:name id="DC6213A32E29AD711BCD45854298BCD2" type="personal">
<mods:role id="E02282810FB8B2C2479D5E5DAE17370C">
<mods:roleTerm id="9A51912AB55E73EA257BD9D8DFD9161E">Author</mods:roleTerm>
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<mods:namePart id="69175CC8CE137EF9E9719DD80FCDFF41">Bilton, David T.</mods:namePart>
<mods:nameIdentifier id="21D3F367695DF95FFF719127B73B2A6D" type="ORCID">0000-0003-1136-0848</mods:nameIdentifier>
<mods:affiliation id="BA61F89EA5EE4035C08EF5FD46B7ED3C">School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth PL 4 8 AA, UK. d. bilton @ plymouth. ac. uk; https: // orcid. org / 0000 - 0003 - 1136 - 0848 &amp; Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, Johannesburg 2006, South Africa.</mods:affiliation>
<mods:nameIdentifier id="01DF24D1F1712E7E37D6ED58BE98B812" type="email">d.bilton@plymouth.ac.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="93CCBAEA914FF9DF8FFAE2B94CF0D29D" type="personal">
<mods:role id="CEECF19C947D7965051C54E6C024FC04">
<mods:roleTerm id="BF4B168F7EA7B54A80698DC491AD9233">Author</mods:roleTerm>
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<mods:namePart id="9944E30CF54BD1ED5563606E88A60B29">Shepard, William D.</mods:namePart>
<mods:nameIdentifier id="997813057798C350F2F6F213781A5A44" type="ORCID">0000-0003-4664-2597</mods:nameIdentifier>
<mods:affiliation id="DBFAD094B217B9E0BCC3E74E694752D4">Essig Museum of Entomology, 1101 Life Sciences Bldg. # 4780, University of California Berkeley, CA 94720 USA. William. Shepard @ csus. edu; http: // orcid. org / 0000 - 0003 - 4664 - 2597</mods:affiliation>
<mods:nameIdentifier id="033FEF484F85E461AB3B0CEA96473887" type="email">hepard@csus.edu</mods:nameIdentifier>
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<mods:typeOfResource id="558343D8405CF628C373B861F5D27575">text</mods:typeOfResource>
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<mods:title id="F0D318ABB05C3517901E940712049F63">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="76E8A092B04FD4B25085FAAE5474788F">
<mods:date id="1EEFC45B948D3AA9C6C441CEC4752025">2022</mods:date>
<mods:detail id="E4FF277E6B4D04527DF70A768F34589F" type="pubDate">
<mods:number id="8491692431C16C769F1E3A46264E1347">2022-10-17</mods:number>
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<mods:detail id="BB104ACE17E02C0A646917807B5F21A4" type="volume">
<mods:number id="C70175152C48F1EF3963CF18322210D4">5195</mods:number>
</mods:detail>
<mods:detail id="10F02B905453B963C76B6D689C600737" type="issue">
<mods:number id="91BB8A43249ED219F816EFA6C7257BDA">6</mods:number>
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<mods:classification id="900EE05792C4325E129804316AD416AC">journal article</mods:classification>
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<mods:identifier id="9B25FB21E67CF6D77549A13D334BD439" type="DOI">10.11646/zootaxa.5195.6.3</mods:identifier>
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<mods:identifier id="72824706E87B3D85FECAD4377AA35F09" type="ISSN">1175-5326</mods:identifier>
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<treatment id="C74FFC13303DFF9DFF685645D8F6FA40" ID-DOI="http://doi.org/10.5281/zenodo.7223886" ID-GBIF-Taxon="203115544" ID-Zenodo-Dep="7223886" LSID="urn:lsid:plazi:treatment:C74FFC13303DFF9DFF685645D8F6FA40" httpUri="http://treatment.plazi.org/id/C74FFC13303DFF9DFF685645D8F6FA40" lastPageNumber="540" pageId="1" pageNumber="540">
<subSubSection id="07FC1E8E303DFF9DFF685645DCE1FBB4" box="[151,389,1050,1077]" pageId="1" pageNumber="540" type="nomenclature">
<paragraph id="4F594D05303DFF9DFF685645DCE1FBB4" blockId="1.[151,389,1050,1113]" box="[151,389,1050,1077]" pageId="1" pageNumber="540">
<heading id="1411FA69303DFF9DFF685645DCE1FBB4" bold="true" box="[151,389,1050,1077]" fontSize="11" level="1" pageId="1" pageNumber="540" reason="1">
<emphasis id="7D929117303DFF9DFF685645DCE1FBB4" bold="true" box="[151,389,1050,1077]" pageId="1" pageNumber="540">
<taxonomicName id="88E63686303DFF9DFF685645DC74FBB4" authority="Bilton &amp; Shepard, 2022" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[151,272,1050,1077]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="7D929117303DFF9DFF685645DC74FBB4" bold="true" box="[151,272,1050,1077]" italics="true" pageId="1" pageNumber="540">Rhithrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="66A12C6C303DFF9DFEE85644DCE1FBB4" box="[279,389,1051,1077]" pageId="1" pageNumber="540" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07FC1E8E303DFF9DFF685660DC7CFBD8" box="[151,280,1087,1113]" pageId="1" pageNumber="540" type="description">
<paragraph id="4F594D05303DFF9DFF685660DC7CFBD8" blockId="1.[151,389,1050,1113]" box="[151,280,1087,1113]" pageId="1" pageNumber="540">
(
<figureCitation id="D7DD5180303DFF9DFF605660DC6BFBD8" box="[159,271,1087,1113]" captionStart-0="FIG" captionStart-1="FIG" captionStart-10="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStart-6="FIG" captionStart-7="FIG" captionStart-8="FIG" captionStart-9="FIG" captionStartId-0="3.[152,201,1943,1967]" captionStartId-1="4.[151,196,1943,1967]" captionStartId-10="13.[151,196,1108,1132]" captionStartId-2="5.[151,196,1643,1667]" captionStartId-3="6.[151,196,1614,1638]" captionStartId-4="7.[151,196,934,958]" captionStartId-5="8.[151,196,1842,1866]" captionStartId-6="9.[151,196,1643,1667]" captionStartId-7="10.[151,196,1643,1667]" captionStartId-8="11.[151,196,1943,1967]" captionStartId-9="12.[151,196,1943,1967]" captionTargetBox-0="[199,1378,203,1900]" captionTargetBox-1="[214,1373,181,1919]" captionTargetBox-10="[191,1396,181,1085]" captionTargetBox-2="[154,1434,181,1619]" captionTargetBox-3="[167,1421,181,1590]" captionTargetBox-4="[180,1407,209,861]" captionTargetBox-5="[154,1434,381,1819]" captionTargetBox-6="[154,1434,181,1619]" captionTargetBox-7="[154,1434,181,1619]" captionTargetBox-8="[211,1375,201,1919]" captionTargetBox-9="[180,1387,247,1903]" captionTargetId-0="figure-19@3.[187,1401,179,1918]" captionTargetId-1="figure-25@4.[214,1373,181,1919]" captionTargetId-10="figure-19@13.[191,1396,181,1085]" captionTargetId-2="figure-19@5.[154,1434,181,1619]" captionTargetId-3="figure-25@6.[167,1421,181,1590]" captionTargetId-4="figure-19@7.[154,1434,181,910]" captionTargetId-5="figure-126@8.[154,1434,381,1819]" captionTargetId-6="figure-19@9.[154,1434,181,1619]" captionTargetId-7="figure-25@10.[154,1434,181,1619]" captionTargetId-8="figure-19@11.[164,1423,181,1919]" captionTargetId-9="figure-25@12.[164,1423,181,1919]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="13" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="FIG. 1. Rhithrops capensis gen. et sp. nov., Holotype habitus. Scale bar = 1 mm." captionText-1="FIG. 2. South African endemic dryopid genera, dorsolateral view of pronotal margins. A) Rhithrops capensis gen. et sp. nov., holotype male; B) Rhithrops capensis gen. et sp. nov., paratype female; C) Rapnus sp., female; D) Strina sp., female. C) &amp; D) South Africa, Western Cape, Franschoek, Mont Rochelle. Scale bar = 0.5 mm." captionText-10="FIG. 11. Rhithrops capensis gen. et sp. nov. Type locality, South Africa, Western Cape, Cederberg, Matjies Rivier, Matjiesrivier, 32°31’07.42”S 19°21’00.4”E, 725 m. Photo D T Bilton." captionText-2="FIG. 3. Rhithrops capensis gen. et sp. nov. A) head, posterolateral view; B) compound eye – arrow indicates inter-ommatidial seta; C) detail of dorsolateral sculpture of head, inside compound eye; D) antennae; E) antennomeres 1–2; F) antennomeres 3–10." captionText-3="FIG. 4. Rhithrops capensis gen. et sp. nov. A) labrum, dorsal view; B) right mandible, ventral view; C) right maxilla, dorsal view; D) close-up of maxillary palp apex; E) female labrum and mentum, ventral view; F) male labrum and mentum, anteroventral view." captionText-4="FIG. 5. Rhithrops capensis gen. et sp. nov. A) right mandible, ventrolateral view – teeth numbered as in text; B) left mandible, ventral view – pt = prostheca, Lf = lateral flange. Scale bar = 100 μm." captionText-5="FIG. 6. Rhithrops capensis gen. et sp. nov. A) prothorax, dorsal view; B) close-up of sculpture of pronotal disc; C) Meso and metanotum, with scutellum; D) right elytron, dorsolateral view; E) detail of right elytral apex; F) prothorax, ventral view." captionText-6="FIG. 7. Rhithrops capensis gen. et sp. nov. A) meso and metaventrites, ventral view; B) male abdomen, ventral view; C) female abdomen, ventral view; D) close-up of plastron vestiture at junction of abdominal ventrites 2–3; E) male abdominal apex; F) female abdominal apex." captionText-7="FIG. 8. Rhithrops capensis gen. et sp. nov. A) male foreleg, ventral view; B) male foretarsus and tibial apex, ventral view; C) male hind tibia and tarsus, ventral view; D) aedeagus, ventral view; E) aedeagus, lateral view; F) aedeagal apex, ventral view." captionText-8="FIG. 9. Rhithrops capensis gen. et sp. nov. A) aedeagus, ventrolateral view; B) aedeagus, lateral view; C) male sternite 9; D) aedeagal apex, ventral view (not to scale); E) male tergite 8; F) male sternite 8. Scale bars = 0.5 mm." captionText-9="FIG. 10. Rhithrops capensis gen. et sp. nov. A) ovipositor, lateral view; B) ovipositor apex, lateral view (not to scale); C) female sternite 8. Scale bar (A &amp; C) = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7223888" figureDoi-1="http://doi.org/10.5281/zenodo.7223890" figureDoi-10="http://doi.org/10.5281/zenodo.7223918" figureDoi-2="http://doi.org/10.5281/zenodo.7223894" figureDoi-3="http://doi.org/10.5281/zenodo.7223899" figureDoi-4="http://doi.org/10.5281/zenodo.7223903" figureDoi-5="http://doi.org/10.5281/zenodo.7223905" figureDoi-6="http://doi.org/10.5281/zenodo.7223907" figureDoi-7="http://doi.org/10.5281/zenodo.7223910" figureDoi-8="http://doi.org/10.5281/zenodo.7223912" figureDoi-9="http://doi.org/10.5281/zenodo.7223916" httpUri-0="https://zenodo.org/record/7223888/files/figure.png" httpUri-1="https://zenodo.org/record/7223890/files/figure.png" httpUri-10="https://zenodo.org/record/7223918/files/figure.png" httpUri-2="https://zenodo.org/record/7223894/files/figure.png" httpUri-3="https://zenodo.org/record/7223899/files/figure.png" httpUri-4="https://zenodo.org/record/7223903/files/figure.png" httpUri-5="https://zenodo.org/record/7223905/files/figure.png" httpUri-6="https://zenodo.org/record/7223907/files/figure.png" httpUri-7="https://zenodo.org/record/7223910/files/figure.png" httpUri-8="https://zenodo.org/record/7223912/files/figure.png" httpUri-9="https://zenodo.org/record/7223916/files/figure.png" pageId="1" pageNumber="540">Figs 1–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="07FC1E8E303DFF9DFF6856D8DE73FB20" box="[151,791,1158,1185]" pageId="1" pageNumber="540" type="type_taxon">
<paragraph id="4F594D05303DFF9DFF6856D8DE73FB20" blockId="1.[151,791,1158,1185]" box="[151,791,1158,1185]" pageId="1" pageNumber="540">
<emphasis id="7D929117303DFF9DFF6856D8DC56FB21" bold="true" box="[151,306,1158,1185]" pageId="1" pageNumber="540">
<typeStatus id="905DF3A7303DFF9DFF6856D8DDB6FB20" box="[151,210,1159,1185]" pageId="1" pageNumber="540">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="88E63686303DFF9DFEC656D8DF6AFB21" authority="Bilton &amp; Shepard, 2022" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[313,526,1159,1185]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="species" species="capensis" status="sp. nov.">
<emphasis id="7D929117303DFF9DFEC656D8DF6AFB21" box="[313,526,1159,1185]" italics="true" pageId="1" pageNumber="540">Rhithrops capensis</emphasis>
</taxonomicName>
<emphasis id="7D929117303DFF9DFDEA56D8DF0BFB20" bold="true" box="[533,623,1159,1185]" pageId="1" pageNumber="540">
<taxonomicNameLabel id="66A12C6C303DFF9DFDEA56D8DF0BFB20" box="[533,623,1159,1185]" pageId="1" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="07FC1E8E303DFF9DFF685691D8F6FA40" pageId="1" pageNumber="540" type="diagnosis">
<paragraph id="4F594D05303DFF9DFF685691D8F6FA40" blockId="1.[151,1437,1230,1473]" pageId="1" pageNumber="540">
<emphasis id="7D929117303DFF9DFF685691DC76FB69" bold="true" box="[151,274,1230,1256]" pageId="1" pageNumber="540">Diagnosis.</emphasis>
<taxonomicName id="88E63686303DFF9DFEE75690DCE1FB68" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[280,389,1231,1257]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFEE75690DCE1FB68" box="[280,389,1231,1257]" italics="true" pageId="1" pageNumber="540">Rhithrops</emphasis>
</taxonomicName>
is unique amongst aquatic dryopid genera in that the labrum is transversely upturned apically. In
<collectingCountry id="37F10D95303DFF9DFF4A56ACDC22FA8C" box="[181,326,1267,1293]" name="South Africa" pageId="1" pageNumber="540">South Africa</collectingCountry>
,
<taxonomicName id="88E63686303DFF9DFEAE56ACDCDAFA8C" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[337,446,1267,1293]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFEAE56ACDCDAFA8C" box="[337,446,1267,1293]" italics="true" pageId="1" pageNumber="540">Rhithrops</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="88E63686303DFF9DFD3C56ACDE5FFA8C" authorityName="Bollow" authorityYear="1938" box="[707,827,1267,1293]" class="Insecta" family="Dryopidae" genus="Ahaggaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFD3C56ACDE5FFA8C" box="[707,827,1267,1293]" italics="true" pageId="1" pageNumber="540">Ahaggaria</emphasis>
</taxonomicName>
and
<taxonomicName id="88E63686303DFF9DFC8E56ACDEA7FA8D" authorityName="Olivier" authorityYear="1791" box="[881,963,1267,1292]" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFC8E56ACDEA7FA8D" box="[881,963,1267,1292]" italics="true" pageId="1" pageNumber="540">Dryops</emphasis>
</taxonomicName>
by the lack of long erect setae on the eyes and dorsum of the body.
<taxonomicName id="88E63686303DFF9DFE5B5748DF75FAB0" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[420,529,1303,1329]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFE5B5748DF75FAB0" box="[420,529,1303,1329]" italics="true" pageId="1" pageNumber="540">Rhithrops</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="88E63686303DFF9DFCF25748DE35FAB0" authorityName="Redtenbacher" authorityYear="1867" box="[781,849,1303,1329]" class="Insecta" family="Dryopidae" genus="Strina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFCF25748DE35FAB0" box="[781,849,1303,1329]" italics="true" pageId="1" pageNumber="540">Strina</emphasis>
</taxonomicName>
and
<taxonomicName id="88E63686303DFF9DFC7B5748DEBCFAB1" authorityName="Grouvelle" authorityYear="1899" box="[900,984,1303,1328]" class="Insecta" family="Dryopidae" genus="Rapnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFC7B5748DEBCFAB1" box="[900,984,1303,1328]" italics="true" pageId="1" pageNumber="540">Rapnus</emphasis>
</taxonomicName>
by the antennae having 10 antennomeres as opposed to nine in
<taxonomicName id="88E63686303DFF9DFE715764DCB6FAD4" authorityName="Redtenbacher" authorityYear="1867" box="[398,466,1339,1365]" class="Insecta" family="Dryopidae" genus="Strina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFE715764DCB6FAD4" box="[398,466,1339,1365]" italics="true" pageId="1" pageNumber="540">Strina</emphasis>
</taxonomicName>
and seven in
<taxonomicName id="88E63686303DFF9DFD905764DFA7FAD5" authorityName="Grouvelle" authorityYear="1899" box="[623,707,1339,1364]" class="Insecta" family="Dryopidae" genus="Rapnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFD905764DFA7FAD5" box="[623,707,1339,1364]" italics="true" pageId="1" pageNumber="540">Rapnus</emphasis>
</taxonomicName>
. Additionally,
<taxonomicName id="88E63686303DFF9DFC945764DEBCFAD4" authorityName="Bilton &amp; Shepard" authorityYear="2022" box="[875,984,1339,1365]" class="Insecta" family="Dryopidae" genus="Rhithrops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFC945764DEBCFAD4" box="[875,984,1339,1365]" italics="true" pageId="1" pageNumber="540">Rhithrops</emphasis>
</taxonomicName>
has shallow sublateral pronotal furrows rather than sublateral step-like carinae, with discrete gibbosities on the pronotum outside of the sublateral furrows. The dorsally serrate coxites of the ovipositor are only shared with
<taxonomicName id="88E63686303DFF9DFC6657DCDEB9FA1C" authorityName="Redtenbacher" authorityYear="1867" box="[921,989,1411,1437]" class="Insecta" family="Dryopidae" genus="Strina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFC6657DCDEB9FA1C" box="[921,989,1411,1437]" italics="true" pageId="1" pageNumber="540">Strina</emphasis>
</taxonomicName>
, another South African dryopid genus. Abdominal ventrite 5 lacking a plastron is a character state commonly seen in the genus
<taxonomicName id="88E63686303DFF9DFB8857F8D8EAFA40" authority="Erichson, 1847" authorityName="Erichson" authorityYear="1847" box="[1143,1422,1447,1473]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="7D929117303DFF9DFB8857F8D9BFFA40" box="[1143,1243,1447,1473]" italics="true" pageId="1" pageNumber="540">Helichus</emphasis>
<bibRefCitation id="2B7730F4303DFF9DFB1E57F8D8EAFA40" author="Erichson, W. H." box="[1249,1422,1447,1473]" pageId="1" pageNumber="540" pagination="481 - 640" refId="ref4832" refString="Erichson, W. H. (1847) Naturgeschichte der Insekten Deutschlands. Abteilung 1. Coleoptera, 3, 481 - 640." type="journal article" year="1847">Erichson, 1847</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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