treatments-xml/data/03/EF/87/03EF87FDFF9FB75FFF1010D3FB3BDE15.xml

<document id="002DDE7D372BEFF139472EC808E88EF5" ID-CLB-Dataset="27842" ID-DOI="10.11646/zootaxa.4527.3.9" ID-GBIF-Dataset="195c5a0b-ee6c-4865-ae6e-52403a5b0bc5" ID-ISSN="1175-5326" ID-Zenodo-Dep="2612371" ID-ZooBank="0BAB4687-ED83-416D-9573-8668E05F6E66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553698805300" checkinUser="plazi" docAuthor="Fiemapong, Armand Richard Nzoko &amp; Enghoff, Henrik" docDate="2018" docId="03EF87FDFF9FB75FFF1010D3FB3BDE15" docLanguage="en" docName="zootaxa.4527.3.9.pdf" docOrigin="Zootaxa 4527 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pelmatoiulus tectus Fiemapong &amp; Enghoff 2018, new combination" docType="treatment" docVersion="7" lastPageNumber="407" masterDocId="FFD6FF85FF9EB75BFF871528FFF2DB14" masterDocTitle="A giant African millipede re-discovered, re-located and re-classified after 120 years (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="413" masterPageNumber="403" pageNumber="404" updateTime="1698714523762" updateUser="ExternalLinkService">
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<mods:title id="D8C4F64C655E8B9AED66D95A4A9BC17A">A giant African millipede re-discovered, re-located and re-classified after 120 years (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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<mods:namePart id="A48DBC370044112659953C519B6B1014">Fiemapong, Armand Richard Nzoko</mods:namePart>
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<mods:namePart id="BB98511554989775D7F1E8E1A75FA448">Enghoff, Henrik</mods:namePart>
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<treatment id="03EF87FDFF9FB75FFF1010D3FB3BDE15" ID-DOI="http://doi.org/10.5281/zenodo.5990961" ID-GBIF-Taxon="154508267" ID-Zenodo-Dep="5990961" LSID="urn:lsid:plazi:treatment:03EF87FDFF9FB75FFF1010D3FB3BDE15" httpUri="http://treatment.plazi.org/id/03EF87FDFF9FB75FFF1010D3FB3BDE15" lastPageId="4" lastPageNumber="407" pageId="1" pageNumber="404">
<subSubSection id="C35C6560FF9FB75AFF1010D3FCE2DD02" box="[151,784,1531,1558]" pageId="1" pageNumber="404" type="nomenclature">
<paragraph id="8BF936EBFF9FB75AFF1010D3FCE2DD02" blockId="1.[151,784,1531,1591]" box="[151,784,1531,1558]" pageId="1" pageNumber="404">
<heading id="D0B18187FF9FB75AFF1010D3FCE2DD02" bold="true" box="[151,784,1531,1558]" fontSize="11" level="1" pageId="1" pageNumber="404" reason="1">
<emphasis id="B932EAF9FF9FB75AFF1010D3FCE2DD02" bold="true" box="[151,784,1531,1558]" pageId="1" pageNumber="404">
<taxonomicName id="4C464D68FF9FB75AFF1010D3FDDEDD02" authority="(Cook, 1897)" authorityName="Fiemapong &amp; Enghoff" authorityYear="2018" baseAuthorityName="Cook" baseAuthorityYear="1897" box="[151,556,1531,1558]" class="Diplopoda" family="Pachybolidae" genus="Pelmatoiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="tectus" status="comb. nov.">
<emphasis id="B932EAF9FF9FB75AFF1010D3FE76DD01" bold="true" box="[151,388,1531,1557]" italics="true" pageId="1" pageNumber="404">Pelmatoiulus tectus</emphasis>
(
<bibRefCitation id="EFD74B1AFF9FB75AFE1410D4FDD6DD02" author="Cook, O. F." box="[403,548,1532,1558]" pageId="1" pageNumber="404" pagination="73 - 75" refId="ref3291" refString="Cook, O. F. (1897) New relatives of Spirobolus giganteus. Brandtia, 18, 73 - 75." type="journal article" year="1897">Cook, 1897</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="A2015782FF9FB75AFDBE10D4FCE2DD02" box="[569,784,1532,1558]" pageId="1" pageNumber="404" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C6560FF9FB75AFF101336FF0BDD23" box="[151,249,1566,1591]" pageId="1" pageNumber="404" type="description">
<paragraph id="8BF936EBFF9FB75AFF101336FF0BDD23" blockId="1.[151,784,1531,1591]" box="[151,249,1566,1591]" pageId="1" pageNumber="404">
<figureCitation id="137D2A6EFF9FB75AFF101336FF0BDD23" box="[151,249,1566,1591]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="2.[151,250,1534,1556]" captionStartId-1="3.[151,250,1712,1734]" captionStartId-2="4.[151,250,1062,1084]" captionStartId-3="5.[151,250,1371,1393]" captionStartId-4="6.[151,250,1911,1933]" captionStartId-5="7.[151,250,1128,1150]" captionStartId-6="9.[151,250,1379,1401]" captionTargetBox-0="[152,1434,375,1510]" captionTargetBox-1="[150,1434,301,1690]" captionTargetBox-2="[234,1356,346,932]" captionTargetBox-3="[152,1434,303,1347]" captionTargetBox-4="[154,1435,520,1887]" captionTargetBox-5="[153,1432,194,1107]" captionTargetBox-6="[150,1436,193,1357]" captionTargetId-0="figure@2.[152,1434,375,1510]" captionTargetId-1="figure@3.[150,1436,301,1690]" captionTargetId-2="figure@4.[164,1424,193,1040]" captionTargetId-3="figure@5.[151,1435,303,1349]" captionTargetId-4="figure@6.[151,1435,517,1889]" captionTargetId-5="figure@7.[150,1436,193,1107]" captionTargetId-6="figure@9.[150,1436,193,1357]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="9" captionText-0="FIGURE 1. Pelmatojulus tectus (Cook, 1897), live specimen from Cameroon." captionText-1="FIGURE 2. Pelmatojulus tectus (Cook, 1897), male from Cameroon, A–D. Right antenna. A. Apical view. B. Close-up of a dyad of sensilla from apical (7th) antennomere. C. Tip of sensillum from apical antennomere. D. Apico-dorsal sensilla from 6th antennomere. E–F. Gnathochilarium. Abbreviations: ll = lamella lingualis, st = stipes. Scale bars = 0.5 mm (E–F), 0.1 mm (A), 0.01 mm (B, D), 0.001 mm (C)." captionText-2="FIGURE 3. Gonopods of Pelmatojulus tectus (Cook, 1897) (A–D) and P. togoensis (Cook, 1897) (E–G), after Cook (1899), rearranged. A, E. Anterior gonopods, anterior view. B: Anterior gonopods, posterior view. C, F. right posterior gonopod, anterior view. D, G. right posterior gonopod, posterior view. Not drawn to scale." captionText-3="FIGURE 4. Gonopods of Pelmatoiulus tectus (Cook, 1897), specimen from Cameroon (A–C) A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Left posterior gonopod, anterior view. Abbreviations: cx = coxa, lo = apical lobe of telopodite, sf = sigmoid flange of telopodite, st = sternum, tlp = telopodite. Scale bars = 1 mm." captionText-4="FIGURE 5. Pelmatoiulus spp., left posterior gonopod. A, B: P. tectus (Cook, 1897), male from Cameroon. C, D: P. togoensis (Cook, 1897), male from Ghana. A, C: anterior view. B, D: apical lobe with efferent canal. Abbreviations: al = apical lobe, cn = constriction, eg = efferent groove, pr = basal process, sa = striate area, sl = subapical lobe. Scale bars: 1 mm (A, B), 0.5 mm (B, D)." captionText-5="FIGURE 6. Right vulva of Pelmatojulus tectus (Cook, 1897), specimen from Cameroon. A. ventral view. B. lateral view. C. anterior-ventral view. D.posterior view. Abbreviations: lb = lobe of lateral valve, lv = lateral valve, mv = mesal valve, op = operculum. Scale bars = 0.5 cm (A, B), 0.2 mm (C, D)." captionText-6="FIGURE 7. Records of Pelmatojulus spp. from Cameroon." httpUri-0="https://zenodo.org/record/2612373/files/figure.png" httpUri-1="https://zenodo.org/record/2612375/files/figure.png" httpUri-2="https://zenodo.org/record/2612377/files/figure.png" httpUri-3="https://zenodo.org/record/2612379/files/figure.png" httpUri-4="https://zenodo.org/record/2612381/files/figure.png" httpUri-5="https://zenodo.org/record/2612383/files/figure.png" httpUri-6="https://zenodo.org/record/2612385/files/figure.png" pageId="1" pageNumber="404">Figs 1–7</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="8BF936EBFF9FB75AFF10134FFC24DD8F" blockId="1.[151,1436,1638,1691]" pageId="1" pageNumber="404">
<taxonomicName id="4C464D68FF9FB75AFF10134FFE3DDD68" ID-CoL="4BQ4T" authority="Cook, 1897" authorityName="Cook" authorityYear="1897" box="[151,463,1638,1660]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="tectus">
<emphasis id="B932EAF9FF9FB75AFF10134FFEBDDD68" box="[151,335,1639,1660]" italics="true" pageId="1" pageNumber="404">Pachybolus tectus</emphasis>
<bibRefCitation id="EFD74B1AFF9FB75AFED2134EFE3DDD68" author="Cook, O. F." box="[341,463,1638,1660]" pageId="1" pageNumber="404" pagination="73 - 75" refId="ref3291" refString="Cook, O. F. (1897) New relatives of Spirobolus giganteus. Brandtia, 18, 73 - 75." type="journal article" year="1897">Cook, 1897</bibRefCitation>
</taxonomicName>
. Type-locality: 
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. 
<typeStatus id="54FD8849FF9FB75AFD55134EFCC2DD68" box="[722,816,1638,1660]" pageId="1" pageNumber="404" type="holotype">Holotype</typeStatus>
(only specimen known until now) said by 
<bibRefCitation id="EFD74B1AFF9FB75AFB65134EFA91DD68" author="Cook, O. F." box="[1250,1379,1638,1660]" pageId="1" pageNumber="404" pagination="657 - 666" refId="ref3314" refString="Cook, O. F. (1899) African Diplopoda of the genus Pachybolus. Proceedings of the United States National Museum, 21, 657 - 666. https: // doi. org / 10.5479 / si. 00963801.21 - 1168.657" type="journal article" year="1899">Cook (1899)</bibRefCitation>
to be in the museum at Hamburg, but missing from that collection (
<bibRefCitation id="EFD74B1AFF9FB75AFCB013ADFC39DD8F" author="Weidner, H." box="[823,971,1669,1691]" pageId="1" pageNumber="404" pagination="57 - 104" refId="ref4266" refString="Weidner, H. (1960) Die entomologischen Sammlungen des zoologischen Staatsinstituts und zoologischen Museums Hamburg. III. Teil Chilopoda und Progoneata. Mitteilungen des hamburgischen zoologischen Museums und Instituts, 58, 57 - 104." type="journal article" year="1960">Weidner 1960</bibRefCitation>
).
</paragraph>
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<emphasis id="B932EAF9FF9FB75AFF1013EFFE9CDDF4" bold="true" box="[151,366,1735,1760]" pageId="1" pageNumber="404">Material studied.</emphasis>
<materialsCitation id="3B2E3CB6FF9FB75AFEF913E0FA97DC10" ID-GBIF-Occurrence="2029190995" collectingDate="2017-10-17" collectionCode="ZMUC" collectorName="A. R. Nzoko &amp; Fiemapong &amp; A. Yetchom Fondjo" country="Cameroon" latitude="4.3333335" location="Sanaga Maritime Division" municipality="Yansoki" pageId="1" pageNumber="404" specimenCount="1" specimenCount-male="1">
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, littoral evergreen forest, 
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, 
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, 
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, 009°83’E, secondary forest, 
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<date id="FFF8102BFF9FB75AFE7713C4FD91DC10" box="[496,611,1772,1796]" pageId="1" pageNumber="404" value="2017-10-17">17.x.2017</date>
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, 
<collectorName id="26B3533DFF9FB75AFDF013C4FC66DC10" box="[631,916,1772,1796]" pageId="1" pageNumber="404">A.R. Nzoko Fiemapong</collectorName>
&amp; 
<collectorName id="26B3533DFF9FB75AFC4E13C4FB44DC10" box="[969,1206,1772,1796]" pageId="1" pageNumber="404">A. Yetchom Fondjo</collectorName>
leg. (
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)
</materialsCitation>
; 
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<specimenCount id="9D40FD62FF9FB75AFAFE13C4FA6EDC10" box="[1401,1436,1772,1796]" pageId="1" pageNumber="404" type="female">1♀</specimenCount>
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, littoral evergreen forest, 
<collectionCode id="ED57AE2EFF9FB75AFDC11238FD69DC3C" box="[582,667,1808,1832]" pageId="1" pageNumber="404">NKAM</collectionCode>
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<materialsCitation id="3B2E3CB6FF9FB75AFD2F1227FE47DC58" ID-GBIF-Occurrence="2029191581" collectingDate="2017-09-10" collectionCode="ZMUC" collectorName="A. R. Nzoko Fiemapong" country="Cameroon" latitude="4.35" location="Division" municipality="Djawara" pageId="1" pageNumber="404" specimenCount="1">
<location id="8E996030FF9FB75AFD2F1227FCFEDC3C" LSID="urn:lsid:plazi:treatment:03EF87FDFF9FB75FFF1010D3FB3BDE15:8E996030FF9FB75AFD2F1227FCFEDC3C" box="[680,780,1807,1832]" country="Cameroon" latitude="4.35" municipality="Djawara" name="Division" pageId="1" pageNumber="404">Division</location>
, 
<collectingMunicipality id="6B9DAC91FF9FB75AFC9A1227FC8DDC3C" box="[797,895,1807,1832]" pageId="1" pageNumber="404">Djawara</collectingMunicipality>
, 
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, 009°76’E, 
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</collectingDate>
, 
<collectorName id="26B3533DFF9FB75AFA971238FEE7DC58" pageId="1" pageNumber="404">A.R. Nzoko Fiemapong</collectorName>
leg. (
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)
</materialsCitation>
; 
<materialsCitation id="3B2E3CB6FF9FB75AFE47121CFC2ADC64" ID-GBIF-Occurrence="2029193210" collectingDate="2017-01-27" collectionCode="ARNF" collectorName="A. R. Nzoko Fiemapong" country="Cameroon" location="Littoral Evergreen Forest Region" longitude="10.4" municipality="Yabassi" pageId="1" pageNumber="404" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D40FD62FF9FB75AFE47121CFE11DC58" box="[448,483,1844,1868]" pageId="1" pageNumber="404" type="male">1♂</specimenCount>
<collectingCountry id="F351767BFF9FB75AFE6C121CFD91DC58" box="[491,611,1844,1868]" name="Cameroon" pageId="1" pageNumber="404">Cameroon</collectingCountry>
, 
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, 
<collectingMunicipality id="6B9DAC91FF9FB75AFC76121CFBB8DC58" box="[1009,1098,1844,1868]" pageId="1" pageNumber="404">Yabassi</collectingMunicipality>
, 4°95′N, 
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, secondary forest, under dead leaves, 
<collectingDate id="EFBCE9C3FF9FB75AFE3C1270FDDBDC64" box="[443,553,1880,1904]" pageId="1" pageNumber="404" value="2017-01-27">
<date id="FFF8102BFF9FB75AFE3C1270FDDBDC64" box="[443,553,1880,1904]" pageId="1" pageNumber="404" value="2017-01-27">27.I.2017</date>
</collectingDate>
, 
<collectorName id="26B3533DFF9FB75AFDB21270FCB2DC64" box="[565,832,1879,1904]" pageId="1" pageNumber="404">A.R. Nzoko Fiemapong</collectorName>
leg. (
<collectionCode id="ED57AE2EFF9FB75AFCF81270FC22DC64" box="[895,976,1880,1904]" pageId="1" pageNumber="404">ARNF</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF936EBFF9FB75AFF401254FBBFDC81" blockId="1.[151,1437,1735,2013]" box="[199,1101,1916,1941]" pageId="1" pageNumber="404">
<emphasis id="B932EAF9FF9FB75AFF401254FE68DC81" bold="true" box="[199,410,1916,1941]" pageId="1" pageNumber="404">Descriptive notes.</emphasis>
The following notes are based on the newly studied material.
</paragraph>
<paragraph id="8BF936EBFF9FB75AFF4012B7FEFBDCC9" blockId="1.[151,1437,1735,2013]" pageId="1" pageNumber="404">
Male. Body length ca. 
<quantity id="4CBE9B0EFF9FB75AFE4012B7FDFDDCA3" box="[455,527,1951,1976]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="404" unit="cm" value="15.0">15 cm</quantity>
, diameter 1.5 cm, 53 podous body rings + telson (no apodous body rings in front of telson).
</paragraph>
<paragraph id="8BF936EBFF9CB759FF4015BFFC79DBEC" blockId="2.[151,1436,151,320]" pageId="2" pageNumber="405">
Colour of live animals (
<figureCitation id="137D2A6EFF9CB759FE5615BFFDE1DBA4" box="[465,531,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1534,1556]" captionTargetBox="[152,1434,375,1510]" captionTargetId="figure@2.[152,1434,375,1510]" captionTargetPageId="2" captionText="FIGURE 1. Pelmatojulus tectus (Cook, 1897), live specimen from Cameroon." httpUri="https://zenodo.org/record/2612373/files/figure.png" pageId="2" pageNumber="405">Fig. 1</figureCitation>
). Head, collum and telson brown; body ringed in black, orange and yellow, each body ring divided into a bright yellow anterior zone, an orange-red intermediate zone and a strongly contrasting, deep black posterior zone; antennae, legs and venter yellow-brown.
</paragraph>
<paragraph id="8BF936EBFF9CB759FF40142CFDD5DA54" blockId="2.[151,1436,151,320]" pageId="2" pageNumber="405">After one year in alcohol head, antennae, collum, legs and telson reddish brown; body rings with pro- and mesozona rust red, metazona black.</paragraph>
<caption id="DF396663FF9CB759FF1010D6FC46DD00" box="[151,948,1534,1556]" httpUri="https://zenodo.org/record/2612373/files/figure.png" pageId="2" pageNumber="405" startId="2.[151,250,1534,1556]" targetBox="[152,1434,375,1510]" targetPageId="2">
<paragraph id="8BF936EBFF9CB759FF1010D6FC46DD00" blockId="2.[151,948,1534,1556]" box="[151,948,1534,1556]" pageId="2" pageNumber="405">
<emphasis id="B932EAF9FF9CB759FF1010D6FEFFDD00" bold="true" box="[151,269,1534,1556]" pageId="2" pageNumber="405">FIGURE 1</emphasis>
. 
<taxonomicName id="4C464D68FF9CB759FE9E10D7FD82DD00" authority="(Cook, 1897)" authorityName="Cook" authorityYear="1897" baseAuthorityName="Cook" baseAuthorityYear="1897" box="[281,624,1534,1556]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="tectus">
<emphasis id="B932EAF9FF9CB759FE9E10D7FE12DD00" box="[281,480,1535,1556]" italics="true" pageId="2" pageNumber="405">Pelmatojulus tectus</emphasis>
(Cook, 1897)
</taxonomicName>
, live specimen from Cameroon.
</paragraph>
</caption>
<paragraph id="8BF936EBFF9CB759FF401314FAF8DDD5" blockId="2.[151,1436,1596,2017]" pageId="2" pageNumber="405">
Incisura lateralis closed. Antennae with ca. 30 apical sensilla (
<figureCitation id="137D2A6EFF9CB759FC301314FC0DDD40" box="[951,1023,1596,1621]" captionStart="FIGURE 2" captionStartId="3.[151,250,1712,1734]" captionTargetBox="[150,1434,301,1690]" captionTargetId="figure@3.[150,1436,301,1690]" captionTargetPageId="3" captionText="FIGURE 2. Pelmatojulus tectus (Cook, 1897), male from Cameroon, A–D. Right antenna. A. Apical view. B. Close-up of a dyad of sensilla from apical (7th) antennomere. C. Tip of sensillum from apical antennomere. D. Apico-dorsal sensilla from 6th antennomere. E–F. Gnathochilarium. Abbreviations: ll = lamella lingualis, st = stipes. Scale bars = 0.5 mm (E–F), 0.1 mm (A), 0.01 mm (B, D), 0.001 mm (C)." httpUri="https://zenodo.org/record/2612375/files/figure.png" pageId="2" pageNumber="405">Fig. 2</figureCitation>
A–D); tip of sensilla with a pore surrounded by finger-shaped processes (
<figureCitation id="137D2A6EFF9CB759FDDC1348FD49DD6C" box="[603,699,1632,1657]" captionStart="FIGURE 2" captionStartId="3.[151,250,1712,1734]" captionTargetBox="[150,1434,301,1690]" captionTargetId="figure@3.[150,1436,301,1690]" captionTargetPageId="3" captionText="FIGURE 2. Pelmatojulus tectus (Cook, 1897), male from Cameroon, A–D. Right antenna. A. Apical view. B. Close-up of a dyad of sensilla from apical (7th) antennomere. C. Tip of sensillum from apical antennomere. D. Apico-dorsal sensilla from 6th antennomere. E–F. Gnathochilarium. Abbreviations: ll = lamella lingualis, st = stipes. Scale bars = 0.5 mm (E–F), 0.1 mm (A), 0.01 mm (B, D), 0.001 mm (C)." httpUri="https://zenodo.org/record/2612375/files/figure.png" pageId="2" pageNumber="405">Fig. 2 C</figureCitation>
). Antennomeres 5 and 6 with dorso-apical fields of small sensilla (
<figureCitation id="137D2A6EFF9CB759FF1813ACFEF4DD88" box="[159,262,1668,1693]" captionStart="FIGURE 2" captionStartId="3.[151,250,1712,1734]" captionTargetBox="[150,1434,301,1690]" captionTargetId="figure@3.[150,1436,301,1690]" captionTargetPageId="3" captionText="FIGURE 2. Pelmatojulus tectus (Cook, 1897), male from Cameroon, A–D. Right antenna. A. Apical view. B. Close-up of a dyad of sensilla from apical (7th) antennomere. C. Tip of sensillum from apical antennomere. D. Apico-dorsal sensilla from 6th antennomere. E–F. Gnathochilarium. Abbreviations: ll = lamella lingualis, st = stipes. Scale bars = 0.5 mm (E–F), 0.1 mm (A), 0.01 mm (B, D), 0.001 mm (C)." httpUri="https://zenodo.org/record/2612375/files/figure.png" pageId="2" pageNumber="405">Fig. 2 D</figureCitation>
). Gnathochilarium: stipites (
<emphasis id="B932EAF9FF9CB759FDCA13ADFD92DD88" box="[589,608,1669,1692]" italics="true" pageId="2" pageNumber="405">st</emphasis>
) each with 4–6 apical setae; lamellae linguales (
<emphasis id="B932EAF9FF9CB759FB1213ACFB56DD88" box="[1173,1188,1668,1692]" italics="true" pageId="2" pageNumber="405">ll</emphasis>
) each with two setae arranged in a transverse subapical row (
<figureCitation id="137D2A6EFF9CB759FDD31380FD22DDD4" box="[596,720,1704,1729]" captionStart="FIGURE 2" captionStartId="3.[151,250,1712,1734]" captionTargetBox="[150,1434,301,1690]" captionTargetId="figure@3.[150,1436,301,1690]" captionTargetPageId="3" captionText="FIGURE 2. Pelmatojulus tectus (Cook, 1897), male from Cameroon, A–D. Right antenna. A. Apical view. B. Close-up of a dyad of sensilla from apical (7th) antennomere. C. Tip of sensillum from apical antennomere. D. Apico-dorsal sensilla from 6th antennomere. E–F. Gnathochilarium. Abbreviations: ll = lamella lingualis, st = stipes. Scale bars = 0.5 mm (E–F), 0.1 mm (A), 0.01 mm (B, D), 0.001 mm (C)." httpUri="https://zenodo.org/record/2612375/files/figure.png" pageId="2" pageNumber="405">Fig. 2 E, F</figureCitation>
). Legs from third to last pair with large tibial pads.
</paragraph>
<paragraph id="8BF936EBFF9CB759FF4013E4FC12DC61" blockId="2.[151,1436,1596,2017]" pageId="2" pageNumber="405">
Anterior gonopods (
<figureCitation id="137D2A6EFF9CB759FE3713E4FE09DDF0" box="[432,507,1740,1765]" captionStart="FIGURE 3" captionStartId="4.[151,250,1062,1084]" captionTargetBox="[234,1356,346,932]" captionTargetId="figure@4.[164,1424,193,1040]" captionTargetPageId="4" captionText="FIGURE 3. Gonopods of Pelmatojulus tectus (Cook, 1897) (A–D) and P. togoensis (Cook, 1897) (E–G), after Cook (1899), rearranged. A, E. Anterior gonopods, anterior view. B: Anterior gonopods, posterior view. C, F. right posterior gonopod, anterior view. D, G. right posterior gonopod, posterior view. Not drawn to scale." httpUri="https://zenodo.org/record/2612377/files/figure.png" pageId="2" pageNumber="405">Figs 3</figureCitation>
A–B, 4 A–B) exactly as illustrated by 
<bibRefCitation id="EFD74B1AFF9CB759FC4813E4FB94DDF0" author="Cook, O. F." box="[975,1126,1740,1765]" pageId="2" pageNumber="405" pagination="657 - 666" refId="ref3314" refString="Cook, O. F. (1899) African Diplopoda of the genus Pachybolus. Proceedings of the United States National Museum, 21, 657 - 666. https: // doi. org / 10.5479 / si. 00963801.21 - 1168.657" type="journal article" year="1899">Cook (1899)</bibRefCitation>
: sternum (
<emphasis id="B932EAF9FF9CB759FB6513E5FB07DDF0" box="[1250,1269,1741,1764]" italics="true" pageId="2" pageNumber="405">st</emphasis>
) with a large, tongue-shaped, broadly rounded median lobe; coxae (
<emphasis id="B932EAF9FF9CB759FC9113D9FCDCDC1C" box="[790,814,1777,1800]" italics="true" pageId="2" pageNumber="405">cx</emphasis>
) ca. 25% longer than sternal lobe, apically broadly rounded; telopodites (
<emphasis id="B932EAF9FF9CB759FE17123CFE5FDC38" box="[400,429,1812,1836]" italics="true" pageId="2" pageNumber="405">tlp</emphasis>
) longer than sternal lobe, but shorter than coxae, in posterior view strongly narrowed at ca. 2/3 of their length, apical part a rounded lobe (
<emphasis id="B932EAF9FF9CB759FD4A1210FD11DC44" box="[717,739,1848,1872]" italics="true" pageId="2" pageNumber="405">lo</emphasis>
, tip of 
<emphasis id="B932EAF9FF9CB759FCB21210FCB9DC44" box="[821,843,1848,1872]" italics="true" pageId="2" pageNumber="405">lo</emphasis>
visible in anterior view), mesal margin of telopodite broadly bent posteriad, forming a sigmoid flange (s 
<emphasis id="B932EAF9FF9CB759FD511274FD2CDC60" box="[726,734,1884,1908]" italics="true" pageId="2" pageNumber="405">f</emphasis>
) ending in apical lobe.
</paragraph>
<paragraph id="8BF936EBFF9CB758FF4012A8FE56DBEC" blockId="2.[151,1436,1596,2017]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" lastPageNumber="406" pageId="2" pageNumber="405">
Posterior gonopods (
<figureCitation id="137D2A6EFF9CB759FE3712A8FE05DC8C" box="[432,503,1920,1945]" captionStart="FIGURE 3" captionStartId="4.[151,250,1062,1084]" captionTargetBox="[234,1356,346,932]" captionTargetId="figure@4.[164,1424,193,1040]" captionTargetPageId="4" captionText="FIGURE 3. Gonopods of Pelmatojulus tectus (Cook, 1897) (A–D) and P. togoensis (Cook, 1897) (E–G), after Cook (1899), rearranged. A, E. Anterior gonopods, anterior view. B: Anterior gonopods, posterior view. C, F. right posterior gonopod, anterior view. D, G. right posterior gonopod, posterior view. Not drawn to scale." httpUri="https://zenodo.org/record/2612377/files/figure.png" pageId="2" pageNumber="405">Figs 3</figureCitation>
C–D, 4 C, 5 A–B) also exactly as illustrated by 
<bibRefCitation id="EFD74B1AFF9CB759FB9A12A8FB43DC8C" author="Cook, O. F." box="[1053,1201,1920,1945]" pageId="2" pageNumber="405" pagination="657 - 666" refId="ref3314" refString="Cook, O. F. (1899) African Diplopoda of the genus Pachybolus. Proceedings of the United States National Museum, 21, 657 - 666. https: // doi. org / 10.5479 / si. 00963801.21 - 1168.657" type="journal article" year="1899">Cook (1899)</bibRefCitation>
: a single, slender, Cshaped sclerite with an irregularly rounded process (
<emphasis id="B932EAF9FF9CB759FD61128DFD0DDCA8" box="[742,767,1957,1980]" italics="true" pageId="2" pageNumber="405">pr</emphasis>
) at base, close to articulation with tracheal apodeme; at ca. 2/3 from base with constriction (
<emphasis id="B932EAF9FF9CB759FD9B12E1FDC5DCF4" box="[540,567,1993,2016]" italics="true" pageId="2" pageNumber="405">cn</emphasis>
), tip folded mesad, forming a broadly rounded apical lobe (
<emphasis id="B932EAF9FF9CB759FA8612E0FAE5DCF4" box="[1281,1303,1992,2016]" italics="true" pageId="2" pageNumber="405">al</emphasis>
). A poorly sclerotized, obliquely striate area (
<emphasis id="B932EAF9FF9DB758FD9815B1FDCADBA4" box="[543,568,153,176]" italics="true" pageId="3" pageNumber="406">sa</emphasis>
) covering middle 1/3 and ending at constriction on external side of the &quot;C&quot;. Efferent groove (
<emphasis id="B932EAF9FF9DB758FEDD1595FE86DBC0" box="[346,372,189,212]" italics="true" pageId="3" pageNumber="406">eg</emphasis>
) (cf. 
<bibRefCitation id="EFD74B1AFF9DB758FE331594FDA2DBC0" author="Enghoff, H." box="[436,592,188,213]" pageId="3" pageNumber="406" pagination="1 - 41" refId="ref3692" refString="Enghoff, H. (2011) East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae), Zootaxa, 2753, 1 - 41." type="journal article" year="2011">Enghoff 2011</bibRefCitation>
) running along inner curvature of &quot;C&quot;, ending near gonopod apex at bentover side of apical lobe.
</paragraph>
</subSubSection>
<caption id="DF396663FF9DB758FF101398FDFBDC36" httpUri="https://zenodo.org/record/2612375/files/figure.png" pageId="3" pageNumber="406" startId="3.[151,250,1712,1734]" targetBox="[150,1434,301,1690]" targetPageId="3">
<paragraph id="8BF936EBFF9DB758FF101398FDFBDC36" blockId="3.[151,1436,1712,1826]" pageId="3" pageNumber="406">
<emphasis id="B932EAF9FF9DB758FF101398FEFCDDD1" bold="true" box="[151,270,1712,1734]" pageId="3" pageNumber="406">FIGURE 2</emphasis>
. 
<taxonomicName id="4C464D68FF9DB758FE9B1398FD85DDD2" authority="(Cook, 1897)" authorityName="Cook" authorityYear="1897" baseAuthorityName="Cook" baseAuthorityYear="1897" box="[284,631,1712,1734]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="tectus">
<emphasis id="B932EAF9FF9DB758FE9B1398FE16DDD2" box="[284,484,1712,1734]" italics="true" pageId="3" pageNumber="406">Pelmatojulus tectus</emphasis>
(Cook, 1897)
</taxonomicName>
, male from Cameroon, A–D. Right antenna. 
<emphasis id="B932EAF9FF9DB758FBCF1398FBA8DDD1" bold="true" box="[1096,1114,1712,1733]" pageId="3" pageNumber="406">A</emphasis>
. Apical view. 
<emphasis id="B932EAF9FF9DB758FB751398FAF1DDD1" bold="true" box="[1266,1283,1712,1733]" pageId="3" pageNumber="406">B</emphasis>
. Close-up of a dyad of sensilla from apical (7th) antennomere. 
<emphasis id="B932EAF9FF9DB758FDFD13E6FD7EDDF0" bold="true" box="[634,652,1742,1764]" pageId="3" pageNumber="406">C</emphasis>
. Tip of sensillum from apical antennomere. 
<emphasis id="B932EAF9FF9DB758FBCD13E7FBAEDDF0" bold="true" box="[1098,1116,1743,1764]" pageId="3" pageNumber="406">D</emphasis>
. Apico-dorsal sensilla from 6th antennomere. 
<emphasis id="B932EAF9FF9DB758FEAF13C6FEA1DC17" bold="true" box="[296,339,1774,1795]" pageId="3" pageNumber="406">E–F</emphasis>
. Gnathochilarium. 
<emphasis id="B932EAF9FF9DB758FD9C13C5FD46DC17" bold="true" box="[539,692,1773,1795]" pageId="3" pageNumber="406">Abbreviations</emphasis>
: 
<emphasis id="B932EAF9FF9DB758FD4513C5FD22DC16" box="[706,720,1773,1794]" italics="true" pageId="3" pageNumber="406">ll</emphasis>
= lamella lingualis, 
<emphasis id="B932EAF9FF9DB758FC2113C6FBEADC17" box="[934,1048,1774,1795]" italics="true" pageId="3" pageNumber="406">st = stipes.</emphasis>
<emphasis id="B932EAF9FF9DB758FB9813C4FB7DDC17" bold="true" box="[1055,1167,1772,1795]" pageId="3" pageNumber="406">Scale bars</emphasis>
= 0.5 mm (E–F), 0.1 mm (A), 0.01 mm (B, D), 0.001 mm (C).
</paragraph>
</caption>
<subSubSection id="C35C6560FF9DB75FFF401267FB3BDE15" lastPageId="4" lastPageNumber="407" pageId="3" pageNumber="406" type="description">
<paragraph id="8BF936EBFF9DB758FF401267FEB4DC9F" blockId="3.[151,1436,1870,2003]" pageId="3" pageNumber="406">
<emphasis id="B932EAF9FF9DB758FF401267FEE9DC73" box="[199,283,1871,1895]" italics="true" pageId="3" pageNumber="406">Female</emphasis>
. Body length ca. 
<quantity id="4CBE9B0EFF9DB758FE6F1266FDC1DC73" box="[488,563,1870,1895]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="406" unit="cm" value="15.0">15 cm</quantity>
, diameter 1.5 cm, 54 podous body rings + telson (no apodous body rings in front of telson).
</paragraph>
<paragraph id="8BF936EBFF9DB758FF4012BEFE53DCC6" blockId="3.[151,1436,1870,2003]" pageId="3" pageNumber="406">Non-sexual characters as in male, except antennae with 20–25 apical sensilla and gnathochilarial stipites each with 3 or 4 apical setae.</paragraph>
<caption id="DF396663FF9AB75FFF10110EFC9FDF6D" httpUri="https://zenodo.org/record/2612377/files/figure.png" pageId="4" pageNumber="407" startId="4.[151,250,1062,1084]" targetBox="[234,1356,346,932]" targetPageId="4">
<paragraph id="8BF936EBFF9AB75FFF10110EFC9FDF6D" blockId="4.[151,1436,1062,1145]" pageId="4" pageNumber="407">
<emphasis id="B932EAF9FF9AB75FFF10110EFEFEDF2F" bold="true" box="[151,268,1062,1084]" pageId="4" pageNumber="407">FIGURE 3</emphasis>
. Gonopods of 
<taxonomicName id="4C464D68FF9AB75FFE27110EFD01DF28" authority="(Cook, 1897)" authorityName="Cook" authorityYear="1897" baseAuthorityName="Cook" baseAuthorityYear="1897" box="[416,755,1062,1084]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="tectus">
<emphasis id="B932EAF9FF9AB75FFE27110EFD95DF28" box="[416,615,1062,1084]" italics="true" pageId="4" pageNumber="407">Pelmatojulus tectus</emphasis>
(Cook, 1897)
</taxonomicName>
(A–D) and 
<taxonomicName id="4C464D68FF9AB75FFCEC110FFB9CDF28" authority="(Cook, 1897)" authorityName="Cook" authorityYear="1897" baseAuthorityName="Cook" baseAuthorityYear="1897" box="[875,1134,1062,1084]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="togoensis">
<emphasis id="B932EAF9FF9AB75FFCEC110FFC11DF28" box="[875,995,1063,1084]" italics="true" pageId="4" pageNumber="407">P. togoensis</emphasis>
(Cook, 1897)
</taxonomicName>
(E–G), after Cook (1899), rearranged. 
<emphasis id="B932EAF9FF9AB75FFF7B116DFEDEDF4E" bold="true" box="[252,300,1093,1114]" pageId="4" pageNumber="407">A, E</emphasis>
. Anterior gonopods, anterior view. 
<emphasis id="B932EAF9FF9AB75FFD13116DFD57DF4E" bold="true" box="[660,677,1093,1114]" pageId="4" pageNumber="407">B</emphasis>
: Anterior gonopods, posterior view. 
<emphasis id="B932EAF9FF9AB75FFB9E116DFBB4DF4E" bold="true" box="[1049,1094,1093,1115]" pageId="4" pageNumber="407">C, F</emphasis>
. right posterior gonopod, anterior view. 
<emphasis id="B932EAF9FF9AB75FFF54114BFEF1DF6D" bold="true" box="[211,259,1123,1145]" pageId="4" pageNumber="407">D, G</emphasis>
. right posterior gonopod, posterior view. Not drawn to scale.
</paragraph>
</caption>
<paragraph id="8BF936EBFF9AB75FFF401188FB3BDE15" blockId="4.[151,1437,1184,1281]" pageId="4" pageNumber="407">
Vulvae (
<figureCitation id="137D2A6EFF9AB75FFEAC1188FE86DFAD" box="[299,372,1184,1209]" captionStart="FIGURE 6" captionStartId="7.[151,250,1128,1150]" captionTargetBox="[153,1432,194,1107]" captionTargetId="figure@7.[150,1436,193,1107]" captionTargetPageId="7" captionText="FIGURE 6. Right vulva of Pelmatojulus tectus (Cook, 1897), specimen from Cameroon. A. ventral view. B. lateral view. C. anterior-ventral view. D.posterior view. Abbreviations: lb = lobe of lateral valve, lv = lateral valve, mv = mesal valve, op = operculum. Scale bars = 0.5 cm (A, B), 0.2 mm (C, D)." httpUri="https://zenodo.org/record/2612383/files/figure.png" pageId="4" pageNumber="407">Fig. 6</figureCitation>
) kidney-shaped; valves in basal half meeting in almost straight line, mesal margins with numerous short setae from level of tip of small operculum (
<emphasis id="B932EAF9FF9AB75FFCAB11EDFCBADFC8" box="[812,840,1221,1244]" italics="true" pageId="4" pageNumber="407">op</emphasis>
) almost up to highest point of valves (
<figureCitation id="137D2A6EFF9AB75FFB7411ECFA80DFC9" box="[1267,1394,1220,1245]" captionStart="FIGURE 6" captionStartId="7.[151,250,1128,1150]" captionTargetBox="[153,1432,194,1107]" captionTargetId="figure@7.[150,1436,193,1107]" captionTargetPageId="7" captionText="FIGURE 6. Right vulva of Pelmatojulus tectus (Cook, 1897), specimen from Cameroon. A. ventral view. B. lateral view. C. anterior-ventral view. D.posterior view. Abbreviations: lb = lobe of lateral valve, lv = lateral valve, mv = mesal valve, op = operculum. Scale bars = 0.5 cm (A, B), 0.2 mm (C, D)." httpUri="https://zenodo.org/record/2612383/files/figure.png" pageId="4" pageNumber="407">Fig. 6 B, D</figureCitation>
), in apical half with poorly sclerotized margins, margin of lateral valve with large rounded lobe (
<emphasis id="B932EAF9FF9AB75FFB2411C1FB4BDE15" box="[1187,1209,1257,1281]" italics="true" pageId="4" pageNumber="407">lb</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>