treatments-xml/data/03/EF/62/03EF6217BF21FFD80AC0FE779814ABA3.xml
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<document id="9B34A7A19DAE28BF29D9ED15F6681666" ID-CLB-Dataset="20762" ID-DOI="10.11646/zootaxa.3365.1.1" ID-GBIF-Dataset="1db9bc74-46dd-4263-9784-88a59bd05d59" ID-ISSN="1175­5334" ID-Zenodo-Dep="6038900" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629787146833" checkinUser="felipe" docAuthor="Garcia, Francisco Hita &amp; Fisher, Brian L." docDate="2012" docId="03EF6217BF21FFD80AC0FE779814ABA3" docLanguage="en" docName="zt03365p123.pdf" docOrigin="Zootaxa 3365" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Tetramorium ryanphelanae Hita Garcia &amp; Fisher 2012, sp. n." docType="treatment" docVersion="9" lastPageNumber="26" masterDocId="FFD61A6FBF39FFC20A57FF889C5BAB77" masterDocTitle="The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region — taxonomy of the T. bessonii, T. bonibony, T. dysalum, T. marginatum, T. tsingy, and T. weitzeckeri species groups" masterLastPageNumber="123" masterPageNumber="1" pageNumber="25" updateTime="1699142484625" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E4FEEAF35E83B1AE8BB6BAB39000CD4E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="30F065436BC8FF9BC8669E2DE214A694">
<mods:title id="748D420AA5010A82344AC219FD7C46DD">The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region — taxonomy of the T. bessonii, T. bonibony, T. dysalum, T. marginatum, T. tsingy, and T. weitzeckeri species groups</mods:title>
</mods:titleInfo>
<mods:name id="C952AD1C7D2E4BB91D8D10F8B324F8A7" type="personal">
<mods:role id="8D7D2F3F2D3DFC698359434373F707A3">
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<mods:namePart id="49B8D4EE2DAF8A6C7D511FC31449D288">Garcia, Francisco Hita</mods:namePart>
</mods:name>
<mods:name id="04F38CECDE768F1BE389065974EA581B" type="personal">
<mods:role id="E44884168C6C8448AA73453FB4D2005C">
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<mods:namePart id="519B7DE7AAFE7652548EFFD01C59FF7C">Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource id="B32120E3132C892A89EDE1B2C3C59DD9">text</mods:typeOfResource>
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<mods:title id="716EFEFE511BEB6D8E5D8548C21EEF06">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3397B83BBDE39B811D4AFF2050F97EB5">
<mods:date id="2F9A6889C9699ADE93E99E71ECA43D98">2012</mods:date>
<mods:detail id="8DDCDCEC3212B9B337EF5ECC66191F05" type="pubDate">
<mods:number id="BB479ADE159AABE2F32C302B27FAFB9C">2012-07-04</mods:number>
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<mods:number id="BFE305DC56796274355ED38E875F5064">3365</mods:number>
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<mods:classification id="E28D423063352D8F26525D292E743909">journal article</mods:classification>
<mods:identifier id="4EDD4404CEA24D350DBBDEED93AEF691" type="CLB-Dataset">20762</mods:identifier>
<mods:identifier id="089192E99E457EF926743DC7E805A53E" type="DOI">10.11646/zootaxa.3365.1.1</mods:identifier>
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<mods:identifier id="5B8FE2FCA264409AF4C1D50C44F515CA" type="Zenodo-Dep">6038900</mods:identifier>
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<treatment id="03EF6217BF21FFD80AC0FE779814ABA3" ID-DOI="http://doi.org/10.5281/zenodo.5253628" ID-GBIF-Taxon="193281060" ID-Zenodo-Dep="5253628" LSID="urn:lsid:plazi:treatment:03EF6217BF21FFD80AC0FE779814ABA3" httpUri="http://treatment.plazi.org/id/03EF6217BF21FFD80AC0FE779814ABA3" lastPageId="26" lastPageNumber="26" pageId="24" pageNumber="25">
<subSubSection id="C35C808ABF21FFDA0AC0FE779F65A96E" box="[151,830,511,537]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8BF9D301BF21FFDA0AC0FE779F65A96E" blockId="24.[151,830,511,570]" box="[151,830,511,537]" pageId="24" pageNumber="25">
<heading id="D0B1646DBF21FFDA0AC0FE779F65A96E" bold="true" box="[151,830,511,537]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="B9320F13BF21FFDA0AC0FE779F65A96E" bold="true" box="[151,830,511,537]" pageId="24" pageNumber="25">
<taxonomicName id="4C46A882BF21FFDA0AC0FE779EAEA96E" authority="Hita Garcia &amp; Fisher" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[151,757,511,537]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="ryanphelanae" status="sp. nov.">
<emphasis id="B9320F13BF21FFDA0AC0FE779DB8A96E" bold="true" box="[151,483,511,537]" italics="true" pageId="24" pageNumber="25">Tetramorium ryanphelanae</emphasis>
Hita Garcia &amp; Fisher
</taxonomicName>
<taxonomicNameLabel id="A201B268BF21FFDA08ABFE779F65A96E" box="[764,830,511,537]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF21FFDA0AC0FDAA9D2FA94D" box="[151,372,546,570]" pageId="24" pageNumber="25" type="description">
<paragraph id="8BF9D301BF21FFDA0AC0FDAA9D2FA94D" blockId="24.[151,830,511,570]" box="[151,372,546,570]" pageId="24" pageNumber="25">(figs 10, 23, 24, 25)</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF21FFDA0AC0FDE29FA2AF0D" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8BF9D301BF21FFDA0AC0FDE29FA2AF0D" blockId="24.[151,1436,618,1146]" pageId="24" pageNumber="25">
<materialsCitation id="3B2ED95CBF21FFDA0AC0FDE29D80A9BD" ID-GBIF-Occurrence="3462706107" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0454495" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<typeStatus id="54FD6DA3BF21FFDA0AC0FDE29CA4A9F5" box="[151,255,618,642]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
worker,
<collectingCountry id="F3519391BF21FFDA0B31FDE29E6BA9F5" box="[358,560,618,642]" name="Madagascar" pageId="24" pageNumber="25">MADAGASCAR</collectingCountry>
,
<collectingRegion id="49821DE3BF21FFDA086AFDE29E9CA9F5" box="[573,711,618,642]" country="Madagascar" name="Antsiranana" pageId="24" pageNumber="25">Antsiranana</collectingRegion>
,
<location id="8E9985DABF21FFDA0883FDE2981BA9F5" LSID="urn:lsid:plazi:treatment:03EF6217BF21FFD80AC0FE779814ABA3:8E9985DABF21FFDA0883FDE2981BA9F5" box="[724,1088,618,642]" country="Madagascar" latitude="-12.90889" longLatPrecision="1" longitude="49.10983" name="Reserve Speciale de l'Ankarana" pageId="24" pageNumber="25" stateProvince="Antsiranana">Réserve Spéciale de l'Ankarana</location>
,
<locationDeviation id="9A4BCA23BF21FFDA0E1AFDE29C8FA9D1" location="Anivorano Nord" pageId="24" pageNumber="25">
<quantity id="4CBE7EE4BF21FFDA0E1AFDE298F2A9F5" box="[1101,1193,618,642]" metricMagnitude="4" metricUnit="m" metricValue="2.29" pageId="24" pageNumber="25" unit="km" value="22.9">22.9 km</quantity>
224° SW Anivorano Nord
</locationDeviation>
,
<geoCoordinate id="EE72B5C6BF21FFDA0A88FD069D3BA9D1" box="[223,352,654,678]" degrees="12.90889" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="1" value="-12.90889">12.90889 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF21FFDA0B3AFD069DAAA9D1" box="[365,497,654,678]" degrees="49.10983" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="1" value="49.10983">49.10983 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF21FFDA0BAAFD069E63A9D1" box="[509,568,654,678]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="24" pageNumber="25" unit="m" value="80.0">
<elevation id="006B3432BF21FFDA0BAAFD069E63A9D1" box="[509,568,654,678]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="24" pageNumber="25" unit="m" value="80.0">80 m</elevation>
</quantity>
, sifted litter, tropical dry forest, BLF2972,
<date id="FFF8F5C1BF21FFDA0E4AFD069898A9D1" box="[1053,1219,654,678]" pageId="24" pageNumber="25" value="2001-02-10" valueMax="2001-02-16" valueMin="2001-02-10">
<collectingDate id="EFBC0C29BF21FFDA0E4AFD069898A9D1" box="[1053,1219,654,678]" pageId="24" pageNumber="25" value="2001-02-10" valueMax="2001-02-16" valueMin="2001-02-10">10.16.II.2001</collectingDate>
</date>
(
<emphasis id="B9320F13BF21FFDA0E83FD0799C8A9D1" box="[1236,1427,654,678]" italics="true" pageId="24" pageNumber="25">
<collectorName id="26B3B6D7BF21FFDA0E83FD079909A9D1" box="[1236,1362,654,678]" pageId="24" pageNumber="25">B.L. Fisher</collectorName>
et al.
</emphasis>
) (
<collectionCode id="ED574BC4BF21FFDA0AC8FD3A9CB5A9BD" box="[159,238,690,714]" pageId="24" pageNumber="25">CASC</collectionCode>
:
<specimenCode id="DBE07B7ABF21FFDA0AA9FD3A9D89A9BD" box="[254,466,690,714]" collectionCode="CASC" pageId="24" pageNumber="25">CASENT0454495</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B2ED95CBF21FFDA0BBCFD3B99C7A9BD" ID-GBIF-Occurrence="3462706155" box="[491,1436,690,715]" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0454488" specimenCount="1" specimenCount-worker="1" stateProvince="Antsiranana" typeStatus="paratype">
<typeStatus id="54FD6DA3BF21FFDA0BBCFD3B9E03A9BC" box="[491,600,691,715]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D401888BF21FFDA0832FD3A9EA7A9BD" box="[613,764,690,714]" pageId="24" pageNumber="25" type="worker">nine workers</specimenCount>
with same data as holotype (
<collectionCode id="ED574BC4BF21FFDA0E31FD3A98EEA9BD" box="[1126,1205,690,714]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="24" pageNumber="25">CASC</collectionCode>
:
<specimenCode id="DBE07B7ABF21FFDA0E92FD3A99CCA9BD" box="[1221,1431,690,714]" collectionCode="CASC" pageId="24" pageNumber="25">CASENT0454488</specimenCode>
;
</materialsCitation>
<materialsCitation id="3B2ED95CBF21FFDA0AC0FD5E9D36A999" ID-GBIF-Occurrence="3462706303" box="[151,365,726,750]" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0454488" specimenCount="1" specimenCount-worker="1" stateProvince="Antsiranana" typeStatus="paratype">CASENT0454492;</materialsCitation>
<materialsCitation id="3B2ED95CBF21FFDA0BF5FD5E9E22A999" ID-GBIF-Occurrence="3462706178" box="[418,633,726,750]" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0454488" specimenCount="1" specimenCount-worker="1" stateProvince="Antsiranana" typeStatus="paratype">CASENT0454498;</materialsCitation>
<materialsCitation id="3B2ED95CBF21FFDA08F9FD5E9FDFA999" ID-GBIF-Occurrence="3462705618" box="[686,900,726,750]" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0454488" specimenCount="1" specimenCount-worker="1" stateProvince="Antsiranana" typeStatus="paratype">CASENT0454506;</materialsCitation>
<materialsCitation id="3B2ED95CBF21FFDA09EEFD5E98CBA999" ID-GBIF-Occurrence="3462705639" box="[953,1168,726,750]" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0454488" specimenCount="1" specimenCount-worker="1" stateProvince="Antsiranana" typeStatus="paratype">CASENT0454508;</materialsCitation>
<materialsCitation id="3B2ED95CBF21FFDA0E92FD5E9D36A865" ID-GBIF-Occurrence="3462705655" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0454488" specimenCount="1" specimenCount-worker="1" stateProvince="Antsiranana" typeStatus="paratype">CASENT0454510; CASENT0454512;</materialsCitation>
<materialsCitation id="3B2ED95CBF21FFDA0B2FFD729E15A865" ID-GBIF-Occurrence="3462705377" box="[376,590,762,786]" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0454488" specimenCount="1" specimenCount-worker="1" stateProvince="Antsiranana" typeStatus="paratype">CASENT0454514;</materialsCitation>
<materialsCitation id="3B2ED95CBF21FFDA080FFD729F6FA865" ID-GBIF-Occurrence="3462706264" box="[600,820,762,786]" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0454488" specimenCount="1" specimenCount-worker="1" stateProvince="Antsiranana" typeStatus="paratype">CASENT0454515)</materialsCitation>
;
<materialsCitation id="3B2ED95CBF21FFDA0915FD739FF2A841" ID-GBIF-Occurrence="3462705409" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0427964" specimenCount="1" specimenCount-worker="1" stateProvince="Antsiranana" typeStatus="paratype">
<specimenCount id="9D401888BF21FFDA0915FD739FBDA865" box="[834,998,762,786]" pageId="24" pageNumber="25" type="worker">seven workers</specimenCount>
with same data as holotype but from ground nest and collection code BLF2992 (
<collectionCode id="ED574BC4BF21FFDA082DFC969E92A841" box="[634,713,798,822]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="24" pageNumber="25">CASC</collectionCode>
:
<specimenCode id="DBE07B7ABF21FFDA0884FC969FFEA841" box="[723,933,798,822]" collectionCode="CASC" pageId="24" pageNumber="25">CASENT0427964</specimenCode>
;
</materialsCitation>
<materialsCitation id="3B2ED95CBF21FFDA09E7FC9698DDA841" ID-GBIF-Occurrence="3462705430" box="[944,1158,798,822]" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0427964" specimenCount="1" specimenCount-worker="1" stateProvince="Antsiranana" typeStatus="paratype">CASENT0427965;</materialsCitation>
<materialsCitation id="3B2ED95CBF21FFDA0EDAFC9699C7A841" ID-GBIF-Occurrence="3462705731" box="[1165,1436,798,822]" collectingDate="2001-02-10" collectingDateMax="2001-02-16" collectingDateMin="2001-02-10" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="80" latitude="-12.90889" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.10983" pageId="24" pageNumber="25" specimenCode="CASENT0427964" specimenCount="1" specimenCount-worker="1" stateProvince="Antsiranana" typeStatus="paratype">CASENT0427966); and</materialsCitation>
<materialsCitation id="3B2ED95CBF21FFDA0AC0FCCA9ED5A8D5" ID-GBIF-Occurrence="3462705577" collectingDate="2001-02-16" collectingDateMax="2001-02-21" collectingDateMin="2001-02-16" collectionCode="BMNH" collectorName="B. L. Fisher" country="Madagascar" elevation="210" latitude="-12.86361" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.22583" pageId="24" pageNumber="25" specimenCode="CASENT0441698" specimenCount="31" specimenCount-worker="31" stateProvince="Antsiranana" typeStatus="paratype">
<specimenCount id="9D401888BF21FFDA0AC0FCCA9D4EA82D" box="[151,277,834,858]" pageId="24" pageNumber="25" type="worker">31 workers</specimenCount>
from
<collectingRegion id="49821DE3BF21FFDA0B0BFCCA9DBDA82D" box="[348,486,834,858]" country="Madagascar" name="Antsiranana" pageId="24" pageNumber="25">Antsiranana</collectingRegion>
,
<location id="8E9985DABF21FFDA0BA5FCCA9F07A82D" LSID="urn:lsid:plazi:treatment:03EF6217BF21FFD80AC0FE779814ABA3:8E9985DABF21FFDA0BA5FCCA9F07A82D" box="[498,860,834,858]" country="Madagascar" latitude="-12.86361" longLatPrecision="1" longitude="49.22583" name="Reserve Speciale de l'Ankarana" pageId="24" pageNumber="25" stateProvince="Antsiranana">Réserve Spéciale de l'Ankarana</location>
,
<locationDeviation id="9A4BCA23BF21FFDA093FFCCA9953A82D" box="[872,1288,834,858]" location="Anivorano Nord" pageId="24" pageNumber="25">
<quantity id="4CBE7EE4BF21FFDA093FFCCA9F9EA82D" box="[872,965,834,858]" metricMagnitude="4" metricUnit="m" metricValue="1.3599999999999999" pageId="24" pageNumber="25" unit="km" value="13.6">13.6 km</quantity>
192° SSW Anivorano Nord
</locationDeviation>
,
<geoCoordinate id="EE72B5C6BF21FFDA0F43FCCA99CDA82D" box="[1300,1430,834,858]" degrees="12.86361" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="1" value="-12.86361">12.86361 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF21FFDA0AC0FCEE9D41A809" box="[151,282,870,894]" degrees="49.22583" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="1" value="49.22583">49.22583 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF21FFDA0B70FCEE9D2BA809" box="[295,368,870,894]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="24" pageNumber="25" unit="m" value="210.0">
<elevation id="006B3432BF21FFDA0B70FCEE9D2BA809" box="[295,368,870,894]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="24" pageNumber="25" unit="m" value="210.0">210 m</elevation>
</quantity>
, sifted litter, tropical dry forest, collection codes BLF3010 and BLF3012,
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<materialsCitation id="3B2ED95CBF21FFDA08E1FBEB9FAEAF0D" ID-GBIF-Occurrence="3462706038" box="[694,1013,1122,1146]" collectingDate="2001-02-16" collectingDateMax="2001-02-21" collectingDateMin="2001-02-16" collectionCode="NHMB" collectorName="B. L. Fisher" country="Madagascar" elevation="210" latitude="-12.86361" location="Reserve Speciale de l'Ankarana" longLatPrecision="1" longitude="49.22583" pageId="24" pageNumber="25" specimenCode="CASENT0441581" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<collectionCode id="ED574BC4BF21FFDA08E1FBEB9F55AF0D" box="[694,782,1123,1146]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="24" pageNumber="25">NHMB</collectionCode>
:
<specimenCode id="DBE07B7ABF21FFDA094EFBEA9FB5AF0D" box="[793,1006,1122,1146]" collectionCode="NHMB" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="24" pageNumber="25">CASENT0441581</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSection id="E2C9CAEABF21FFDA0AC0FB22983BAE01" pageId="24" pageNumber="25" type="multiple">
<subSubSection id="C35C808ABF21FFDA0AC0FB22983BAE01" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8BF9D301BF21FFDA0AC0FB229D51AFB4" blockId="24.[151,1436,1194,1398]" box="[151,266,1194,1219]" pageId="24" pageNumber="25">
<heading id="D0B1646DBF21FFDA0AC0FB229D51AFB4" bold="true" box="[151,266,1194,1219]" fontSize="10" level="2" pageId="24" pageNumber="25" reason="2">
<emphasis id="B9320F13BF21FFDA0AC0FB229D51AFB4" bold="true" box="[151,266,1194,1219]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF21FFDA0AC0FB46983BAE01" blockId="24.[151,1436,1194,1398]" pageId="24" pageNumber="25">
The following character combination renders
<taxonomicName id="4C46A882BF21FFDA08FBFB479F3DAF91" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[684,870,1230,1254]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="ryanphelanae">
<emphasis id="B9320F13BF21FFDA08FBFB479F3DAF91" box="[684,870,1230,1254]" italics="true" pageId="24" pageNumber="25">T. ryanphelanae</emphasis>
</taxonomicName>
recognisable within the
<taxonomicName id="4C46A882BF21FFDA0EDEFB47995EAF91" baseAuthorityName="Forel" baseAuthorityYear="1895" box="[1161,1285,1230,1254]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B9320F13BF21FFDA0EDEFB47995EAF91" box="[1161,1285,1230,1254]" italics="true" pageId="24" pageNumber="25">T. bessonii</emphasis>
</taxonomicName>
group: short antennal scapes (SI 7678); PSLI 2833; absence of promesonotal suture; high nodiform petiolar node in profile with approximately parallel anterior and posterior faces, anterodorsal and posterodorsal angles at about same height, dorsum not tapering backwards posteriorly; presence of standing hairs on waist segments and first gastral tergite; and relatively long, appressed to decumbent pubescence on first gastral tergite.
</paragraph>
</subSubSection>
</subSection>
<subSection id="E2C9CAEABF21FFDB0AC0FA2E9DB8A960" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="multiple">
<subSubSection id="C35C808ABF21FFDB0AC0FA2E9DB8A960" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
<paragraph id="8BF9D301BF21FFDA0AC0FA2E9D7AAEC8" blockId="24.[151,1436,1446,2010]" box="[151,289,1446,1471]" pageId="24" pageNumber="25">
<heading id="D0B1646DBF21FFDA0AC0FA2E9D7AAEC8" bold="true" box="[151,289,1446,1471]" fontSize="10" level="2" pageId="24" pageNumber="25" reason="2">
<emphasis id="B9320F13BF21FFDA0AC0FA2E9D7AAEC8" bold="true" box="[151,289,1446,1471]" pageId="24" pageNumber="25">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF21FFDB0AC0FA439DB8A960" blockId="24.[151,1436,1446,2010]" lastBlockId="25.[151,1436,151,535]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
HL 0.680.88 (0.81); HW 0.700.90 (0.83); SL 0.540.68 (0.63); EL 0.150.21 (0.18);
<collectingCountry id="F3519391BF21FFDA0E2CFA4398C4AE95" box="[1147,1183,1483,1506]" name="Philippines" pageId="24" pageNumber="25">PH</collectingCountry>
0.410.56 (0.49); PW 0.530.68 (0.62); WL 0.911.22 (1.09); PSL 0.210.28 (0.25); PTL 0.170.23 (0.20); PTH 0.310.37 (0.34); PTW 0.250.32 (0.28); PPL 0.220.27 (0.25); PPH 0.290.37 (0.33); PPW 0.310.42 (0.36); CI 101104 (102); SI 76 78 (77); OI 2023 (22); DMI 5659 (57); LMI 4347 (45); PSLI 2833 (30); PeNI 4150 (46); LPeI 5262 (59); DPeI 133150 (142); PpNI 5262 (58); LPpI 7080 (74); DPpI 135155 (146); PPI 122132 (126) (15 measured). Head wider than long (CI 101104). Anterior clypeal margin with distinct median impression. Frontal carinae well-developed, becoming weaker and fading out shortly after level of posterior eye margin. Antennal scrobes faint to absent, posterior and ventral margins never differentiated. Antennal scapes short, not reaching posterior head margin (SI 7678). Eyes small to moderate (OI 2023). Mesosomal outline in profile convex, dorsally transversely rounded, weakly to moderately marginate from lateral to dorsal mesosoma, promesonotal suture and metanotal groove absent; mesosoma comparatively stout (LMI 4347). Propodeal spines elongate-triangular to spinose, long, and acute (PSLI 2833). Propodeal lobes very small and broadly triangular. Petiolar node in profile high nodiform, anterior and posterior faces approximately parallel, anterodorsal and posterodorsal angles at about same height, dorsum not tapering backwards posteriorly; node in dorsal view not strongly transverse, between 1.3 to 1.5 times wider than long (DPeI 133150), in lateral view between 1.6 to 1.9 times higher than long (LPeI 5262). Postpetiole in profile rounded and weakly anteroposteriorly compressed, approximately 1.2 to 1.4 times higher than long (LPpI 7080), in dorsal view between 1.3 to 1.6 wider than long (DPpI 135155). Postpetiole in profile a bit less voluminous than petiolar node, in dorsal view approximately 1.2 to 1.3 times wider than petiolar node (PPI 122132). Mandibles distinctly longitudinally striate; clypeus always with strong median longitudinal ruga and generally one weaker ruga at each side; remainder of head mostly unsculptured, cephalic dorsum between frontal carinae only with one well-developed longitudinal median ruga, remainder unsculptured, median ruga of same length as frontal carinae and diverging approximately at eye level into two rugae running to posterior clypeal margin; scrobal area, lateral and ventral head mostly unsculptured. Ground sculpture faint. Mesosoma, waist segments, and gaster unsculptured and smooth. Ground sculpture on whole body faint to absent. All dorsal surfaces of body generally with long, erect pilosity in combination with comparatively long appressed pubescence. Body colouration uniformly brown.
</paragraph>
</subSubSection>
</subSection>
<subSection id="E2C9CAEABF20FFDB0AC0FDC09E46AF9F" pageId="25" pageNumber="26" type="multiple">
<subSubSection id="C35C808ABF20FFDB0AC0FDC09900A9BB" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8BF9D301BF20FFDB0AC0FDC09C81A917" blockId="25.[151,1438,584,1256]" box="[151,218,584,608]" pageId="25" pageNumber="26">
<heading id="D0B1646DBF20FFDB0AC0FDC09C81A917" bold="true" box="[151,218,584,608]" fontSize="10" level="2" pageId="25" pageNumber="26" reason="2">
<emphasis id="B9320F13BF20FFDB0AC0FDC09C81A917" bold="true" box="[151,218,584,608]" pageId="25" pageNumber="26">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF20FFDB0AC0FDE49900A9BB" blockId="25.[151,1438,584,1256]" pageId="25" pageNumber="26">
The distribution of the new species seems to be restricted to the northernmost tip of
<collectingCountry id="F3519391BF20FFDB0ED1FDE49954A9F3" box="[1158,1295,620,644]" name="Madagascar" pageId="25" pageNumber="26">Madagascar</collectingCountry>
, where it is generally found in tropical dry forests at elevations of
<quantity id="4CBE7EE4BF20FFDB0977FD079F05A9D0" box="[800,862,655,679]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="26" unit="m" value="60.0">60 m</quantity>
to
<specimenCount id="9D401888BF20FFDB09DCFD079F86A9D0" box="[907,989,655,679]" pageId="25" pageNumber="26" type="generic">
<quantity id="4CBE7EE4BF20FFDB09DCFD079F82A9D0" box="[907,985,655,679]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="25" pageNumber="26" unit="m" value="425.0">425 m</quantity>
.
</specimenCount>
The southernmost localities are Foret d'Andavakoera and Foret de Binara, while the northernmost are Montagne des Français and Foret d'Orangea.
</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF20FFDB0A90FD509E46AF9F" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BF9D301BF20FFDB0A90FD509E46AF9F" blockId="25.[151,1438,584,1256]" pageId="25" pageNumber="26">
Due to its high nodiform petiolar node with anterodorsal and posterodorsal margins at approximately the same height,
<taxonomicName id="4C46A882BF20FFDB0AA0FD759DE2A863" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[247,441,764,788]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ryanphelanae">
<emphasis id="B9320F13BF20FFDB0AA0FD759DE2A863" box="[247,441,764,788]" italics="true" pageId="25" pageNumber="26">T. ryanphelanae</emphasis>
</taxonomicName>
cannot be misidentified with the three species that possess a much stronger anteroposteriorly compressed petiolar node in which the dorsum tapers backwards posteriorly (
<taxonomicName id="4C46A882BF20FFDB0EA8FCA9992FA840" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[1279,1396,800,824]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="B9320F13BF20FFDB0EA8FCA9992FA840" box="[1279,1396,800,824]" italics="true" pageId="25" pageNumber="26">T. artemis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C46A882BF20FFDB0FD0FCA99CA8A82B" baseAuthorityName="Forel" baseAuthorityYear="1895" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B9320F13BF20FFDB0FD0FCA99CA8A82B" italics="true" pageId="25" pageNumber="26">T. bessonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C46A882BF20FFDB0B54FCCD9D3AA82B" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[259,353,836,860]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="B9320F13BF20FFDB0B54FCCD9D3AA82B" box="[259,353,836,860]" italics="true" pageId="25" pageNumber="26">T. wardi</emphasis>
</taxonomicName>
). The remaining two species are morphologically much closer to
<taxonomicName id="4C46A882BF20FFDB0E0CFCCD9948A82B" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[1115,1299,836,860]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ryanphelanae">
<emphasis id="B9320F13BF20FFDB0E0CFCCD9948A82B" box="[1115,1299,836,860]" italics="true" pageId="25" pageNumber="26">T. ryanphelanae</emphasis>
</taxonomicName>
, but can be clearly separated from each other.
<taxonomicName id="4C46A882BF20FFDB0879FCE09F6BA8F7" authorityName="Forel" authorityYear="1895" box="[558,816,872,896]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="orientale">
<emphasis id="B9320F13BF20FFDB0879FCE09F6BA8F7" box="[558,816,872,896]" italics="true" pageId="25" pageNumber="26">Tetramorium orientale</emphasis>
</taxonomicName>
, which is only known from the
<typeStatus id="54FD6DA3BF20FFDB0EE3FCEF9942A808" box="[1204,1305,871,895]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
, displays a distinct promesonotal suture, whereas this character is absent in
<taxonomicName id="4C46A882BF20FFDB0939FC059FAEA8D3" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[878,1013,908,932]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malagasy">
<emphasis id="B9320F13BF20FFDB0939FC059FAEA8D3" box="[878,1013,908,932]" italics="true" pageId="25" pageNumber="26">T. malagasy</emphasis>
</taxonomicName>
and
<taxonomicName id="4C46A882BF20FFDB0E7AFC0598BFA8D3" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[1069,1252,908,932]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ryanphelanae">
<emphasis id="B9320F13BF20FFDB0E7AFC0598BFA8D3" box="[1069,1252,908,932]" italics="true" pageId="25" pageNumber="26">T. ryanphelanae</emphasis>
</taxonomicName>
. The distinction between these two morphologically comparatively similar species relies on antennal scape and propodeal spine lengths and gastral pilosity and pubescence. First,
<taxonomicName id="4C46A882BF20FFDB0884FC5D9F00A89B" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[723,859,980,1004]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malagasy">
<emphasis id="B9320F13BF20FFDB0884FC5D9F00A89B" box="[723,859,980,1004]" italics="true" pageId="25" pageNumber="26">T. malagasy</emphasis>
</taxonomicName>
has comparatively shorter antennal scapes (SI 71 74) and longer propodeal spines (PSLI 3337) than
<taxonomicName id="4C46A882BF20FFDB08B9FC719FFDAF67" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[750,934,1016,1040]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ryanphelanae">
<emphasis id="B9320F13BF20FFDB08B9FC719FFDAF67" box="[750,934,1016,1040]" italics="true" pageId="25" pageNumber="26">T. ryanphelanae</emphasis>
</taxonomicName>
(SI 7678; PSLI 2833). More importantly though,
<taxonomicName id="4C46A882BF20FFDB0AA3FB959D27AF43" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[244,380,1052,1076]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malagasy">
<emphasis id="B9320F13BF20FFDB0AA3FB959D27AF43" box="[244,380,1052,1076]" italics="true" pageId="25" pageNumber="26">T. malagasy</emphasis>
</taxonomicName>
has only very short and strongly appressed pubescence on the first gastral tergite while in
<taxonomicName id="4C46A882BF20FFDB0FD0FB959D6AAF2F" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ryanphelanae">
<emphasis id="B9320F13BF20FFDB0FD0FB959D6AAF2F" italics="true" pageId="25" pageNumber="26">T. ryanphelanae</emphasis>
</taxonomicName>
the gastral pubescence is comparatively long and appressed, and combined with few long erect hairs. Additional information about the differentiation between both species is provided above in the species description of
<taxonomicName id="4C46A882BF20FFDB0AE0FB019D66AFD7" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[183,317,1160,1184]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malagasy">
<emphasis id="B9320F13BF20FFDB0AE0FB019D66AFD7" box="[183,317,1160,1184]" italics="true" pageId="25" pageNumber="26">T. malagasy</emphasis>
</taxonomicName>
. Caution is demanded in some specimens, which do not possess standing hairs on the first gastral tergite, since they could be misidentified as
<taxonomicName id="4C46A882BF20FFDB08CDFB259F79AFB3" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[666,802,1196,1220]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malagasy">
<emphasis id="B9320F13BF20FFDB08CDFB259F79AFB3" box="[666,802,1196,1220]" italics="true" pageId="25" pageNumber="26">T. malagasy</emphasis>
</taxonomicName>
. However, this can be avoided by comparing antennal scape and propodeal spine lengths.
</paragraph>
</subSubSection>
</subSection>
<subSection id="E2C9CAEABF20FFDB0AC0FA90997DAE23" box="[151,1318,1304,1364]" pageId="25" pageNumber="26" type="multiple">
<subSubSection id="C35C808ABF20FFDB0AC0FA90997DAE23" box="[151,1318,1304,1364]" pageId="25" pageNumber="26" type="etymology">
<paragraph id="8BF9D301BF20FFDB0AC0FA909D43AE47" blockId="25.[151,1318,1304,1364]" box="[151,280,1304,1328]" pageId="25" pageNumber="26">
<heading id="D0B1646DBF20FFDB0AC0FA909D43AE47" bold="true" box="[151,280,1304,1328]" fontSize="10" level="2" pageId="25" pageNumber="26" reason="2">
<emphasis id="B9320F13BF20FFDB0AC0FA909D43AE47" bold="true" box="[151,280,1304,1328]" pageId="25" pageNumber="26">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF20FFDB0AC0FAB4997DAE23" blockId="25.[151,1318,1304,1364]" box="[151,1318,1340,1364]" pageId="25" pageNumber="26">The new species is named in honor of Ryan Phelan for her support to discover and identify life on earth.</paragraph>
</subSubSection>
</subSection>
<subSection id="E2C9CAEABF20FFD80AC0FA0B9814ABA3" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="multiple">
<subSubSection id="C35C808ABF20FFD80AC0FA0B9814ABA3" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BF9D301BF20FFDB0AC0FA0B9D22AEEB" blockId="25.[151,1436,1411,2012]" box="[151,377,1411,1436]" pageId="25" pageNumber="26">
<heading id="D0B1646DBF20FFDB0AC0FA0B9D22AEEB" bold="true" box="[151,377,1411,1436]" fontSize="10" level="2" pageId="25" pageNumber="26" reason="2">
<emphasis id="B9320F13BF20FFDB0AC0FA0B9D22AEEB" bold="true" box="[151,377,1411,1436]" pageId="25" pageNumber="26">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF20FFD80AC0FA2F9814ABA3" blockId="25.[151,1436,1411,2012]" lastBlockId="26.[151,1436,151,212]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<materialsCitation id="3B2ED95CBF20FFDB0AC0FA2F9FDFAE93" ID-GBIF-Occurrence="3462705441" pageId="25" pageNumber="26">
<emphasis id="B9320F13BF20FFDB0AC0FA2F9D3FAEB7" bold="true" box="[151,356,1447,1472]" pageId="25" pageNumber="26">
<collectingCountry id="F3519391BF20FFDB0AC0FA2F9D3FAEB7" box="[151,356,1447,1472]" name="Madagascar" pageId="25" pageNumber="26">MADAGASCAR</collectingCountry>
</emphasis>
:
<collectingRegion id="49821DE3BF20FFDB0B2FFA2F9E59AEC8" box="[376,514,1447,1471]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Forêt d' Andavakoera,
<quantity id="4CBE7EE4BF20FFDB0974FA2F9F2CAEC8" box="[803,887,1447,1471]" metricMagnitude="4" metricUnit="m" metricValue="2.1399999999999997" pageId="25" pageNumber="26" unit="km" value="21.4">21.4km</quantity>
75° ENE Ambilobe,
<quantity id="4CBE7EE4BF20FFDB0E28FA2F989EAEC8" box="[1151,1221,1447,1471]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="25" pageNumber="26" unit="km" value="4.6">4.6km</quantity>
356° N Betsiaka,
<geoCoordinate id="EE72B5C6BF20FFDB0AC0FA449D4CAE93" box="[151,279,1484,1508]" degrees="13.11833" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-13.11833">13.11833 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF20FFDB0B73FA449D25AE94" box="[292,382,1484,1508]" degrees="49.23" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="555" value="49.23">49.23 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF20FFDB0BDDFA449D8FAE94" box="[394,468,1484,1508]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="25" pageNumber="26" unit="m" value="425.0">425 m</quantity>
, rainforest,
<date id="FFF8F5C1BF20FFDB080DFA449EB9AE93" box="[602,738,1484,1508]" pageId="25" pageNumber="26" value="2003-12-15">15.XII.2003</date>
(
<emphasis id="B9320F13BF20FFDB08A3FA459F28AE93" box="[756,883,1484,1508]" italics="true" pageId="25" pageNumber="26">B.L. Fisher</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF20FFDB09DBFA44981EAD70" ID-GBIF-Occurrence="3462705844" pageId="25" pageNumber="26">
<collectingRegion id="49821DE3BF20FFDB09DBFA44984DAE93" box="[908,1046,1484,1508]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Forêt de Binara,
<quantity id="4CBE7EE4BF20FFDB0E88FA449975AE93" box="[1247,1326,1484,1508]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="25" pageNumber="26" unit="km" value="7.5">7.5 km</quantity>
230° SW Daraina,
<geoCoordinate id="EE72B5C6BF20FFDB0AAAFA679D39AD70" box="[253,354,1519,1543]" degrees="13.255" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="55" value="-13.255">13.255 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF20FFDB0B38FA679DA9AD70" box="[367,498,1519,1543]" degrees="49.61667" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="49.61667">49.61667 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF20FFDB0BA9FA679E1CAD70" box="[510,583,1519,1543]" metricMagnitude="2" metricUnit="m" metricValue="3.75" pageId="25" pageNumber="26" unit="m" value="375.0">375 m</quantity>
, tropical dry forest,
<date id="FFF8F5C1BF20FFDB097DFA679FFEAD70" box="[810,933,1519,1543]" pageId="25" pageNumber="26" value="2003-12-02">2.XII.2003</date>
(
<emphasis id="B9320F13BF20FFDB09E2FA79986FAD7F" box="[949,1076,1520,1544]" italics="true" pageId="25" pageNumber="26">B.L. Fisher</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF20FFDB0E1BFA6799C7AD5B" ID-GBIF-Occurrence="3462705664" pageId="25" pageNumber="26">
<collectingRegion id="49821DE3BF20FFDB0E1BFA67988DAD70" box="[1100,1238,1519,1543]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Forêt d'Orangea,
<quantity id="4CBE7EE4BF20FFDB0AC0F99C9CB3AD5B" box="[151,232,1556,1580]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="25" pageNumber="26" unit="km" value="3.6">3.6 km</quantity>
128° SE Remena,
<geoCoordinate id="EE72B5C6BF20FFDB0B90F99C9E10AD5B" box="[455,587,1556,1580]" degrees="12.25889" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-12.25889">12.25889 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF20FFDB080DF99C9EBBAD5C" box="[602,736,1556,1580]" degrees="49.37467" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="49.37467">49.37467 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF20FFDB08B8F99C9F76AD5C" box="[751,813,1556,1580]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="25" pageNumber="26" unit="m" value="90.0">90 m</quantity>
, littoral rainforest,
<date id="FFF8F5C1BF20FFDB0E5BF99C98E9AD5B" box="[1036,1202,1556,1580]" pageId="25" pageNumber="26" value="2001-02-22" valueMax="2001-02-28" valueMin="2001-02-22">22.28.II.2001</date>
(
<emphasis id="B9320F13BF20FFDB0E93F99D99D0AD5B" box="[1220,1419,1556,1580]" italics="true" pageId="25" pageNumber="26">B.L. Fisher et al.</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF20FFDB0AC0F9BF9E81AD03" ID-GBIF-Occurrence="3462706302" pageId="25" pageNumber="26">
<collectingRegion id="49821DE3BF20FFDB0AC0F9BF9D7AAD38" box="[151,289,1591,1615]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Montagne des Français,
<quantity id="4CBE7EE4BF20FFDB0817F9BF9ED4AD38" box="[576,655,1591,1615]" metricMagnitude="3" metricUnit="m" metricValue="7.2" pageId="25" pageNumber="26" unit="km" value="7.2">7.2 km</quantity>
142° SE
<collectingRegion id="49821DE3BF20FFDB08ACF9BF9FD9AD38" box="[763,898,1591,1615]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
(=Diego Suarez),
<geoCoordinate id="EE72B5C6BF20FFDB0E04F9BF988EAD38" box="[1107,1237,1591,1615]" degrees="12.32278" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-12.32278">12.32278 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF20FFDB0EB5F9BF993EAD38" box="[1250,1381,1591,1615]" degrees="49.33817" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="49.33817">49.33817 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF20FFDB0F25F9BF9CF4AD04" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="25" pageNumber="26" unit="m" value="180.0">180 m</quantity>
, tropical dry forest,
<date id="FFF8F5C1BF20FFDB0BC5F9D49E63AD03" box="[402,568,1628,1652]" pageId="25" pageNumber="26" value="2001-02-22" valueMax="2001-02-25" valueMin="2001-02-22">22.25.II.2001</date>
(
<emphasis id="B9320F13BF20FFDB081FF9D49E93AD03" box="[584,712,1628,1652]" italics="true" pageId="25" pageNumber="26">G.D. Alpert</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF20FFDB08B6F9D498CCADE0" ID-GBIF-Occurrence="3462706089" pageId="25" pageNumber="26">
<collectingRegion id="49821DE3BF20FFDB08B6F9D49F30AD03" box="[737,875,1628,1652]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Réserve Analamerana,
<quantity id="4CBE7EE4BF20FFDB0E2CF9D4988CAD03" box="[1147,1239,1628,1652]" metricMagnitude="4" metricUnit="m" metricValue="1.67" pageId="25" pageNumber="26" unit="km" value="16.7">16.7 km</quantity>
123° AnivoranoNord,
<geoCoordinate id="EE72B5C6BF20FFDB0AB1F9F79D36ADE0" box="[230,365,1663,1687]" degrees="12.80467" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-12.80467">12.80467 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF20FFDB0BD6F9F79E50ADE0" box="[385,523,1663,1687]" degrees="49.37383" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="49.37383">49.37383 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF20FFDB0848F9F79E34ADE0" box="[543,623,1663,1687]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="25" pageNumber="26" unit="m" value="225.0">225 m</quantity>
, tropical dry forest,
<date id="FFF8F5C1BF20FFDB0939F9F79FB3ADE0" box="[878,1000,1663,1687]" pageId="25" pageNumber="26" value="2004-12-03">3.XII.2004</date>
(
<emphasis id="B9320F13BF20FFDB0E57F90998DDADEF" box="[1024,1158,1664,1688]" italics="true" pageId="25" pageNumber="26">B.L. Fisher</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF20FFDB0EF2F9F79CAAADA8" ID-GBIF-Occurrence="3462705397" pageId="25" pageNumber="26">
<collectingRegion id="49821DE3BF20FFDB0EF2F9F79974ADE0" box="[1189,1327,1663,1687]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Réserve Analamerana,
<quantity id="4CBE7EE4BF20FFDB0B6BF92C9DC2ADCB" box="[316,409,1700,1724]" metricMagnitude="4" metricUnit="m" metricValue="2.84" pageId="25" pageNumber="26" unit="km" value="28.4">28.4 km</quantity>
99° Anivorano-Nord,
<geoCoordinate id="EE72B5C6BF20FFDB08CCF92C9F46ADCB" box="[667,797,1700,1724]" degrees="12.74667" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-12.74667">12.74667 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF20FFDB097CF92C9FF4ADCC" box="[811,943,1700,1724]" degrees="49.49483" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="49.49483">49.49483 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF20FFDB09EBF92C9FA2ADCC" box="[956,1017,1700,1724]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="26" unit="m" value="60.0">60 m</quantity>
, tropical dry forest,
<date id="FFF8F5C1BF20FFDB0EB6F92C9907ADCB" box="[1249,1372,1700,1724]" pageId="25" pageNumber="26" value="2004-12-05">5.XII.2004</date>
(
<emphasis id="B9320F13BF20FFDB0F3AF92D9CBBAD97" italics="true" pageId="25" pageNumber="26">B.L. Fisher</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF20FFDB0AACF94F9EB4AC73" ID-GBIF-Occurrence="3462705308" pageId="25" pageNumber="26">
<collectingRegion id="49821DE3BF20FFDB0AACF94F9DDEADA8" box="[251,389,1735,1759]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Réserve Spéciale d'Ambre,
<quantity id="4CBE7EE4BF20FFDB0884F94F9F7FADA8" box="[723,804,1735,1759]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="26" unit="km" value="3.5">3.5 km</quantity>
235° SW Sakaramy,
<geoCoordinate id="EE72B5C6BF20FFDB0E76F94F98FEADA8" box="[1057,1189,1735,1759]" degrees="12.46889" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-12.46889">12.46889 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF20FFDB0EE2F94F9960ADA8" box="[1205,1339,1735,1759]" degrees="49.24217" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="49.24217">49.24217 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF20FFDB0F1CF94F99C3ADA8" box="[1355,1432,1735,1759]" metricMagnitude="2" metricUnit="m" metricValue="3.25" pageId="25" pageNumber="26" unit="m" value="325.0">325 m</quantity>
, tropical dry forest,
<date id="FFF8F5C1BF20FFDB0BD2F9649DABAC73" box="[389,496,1772,1796]" pageId="25" pageNumber="26" value="2001-01-26">26.I.2001</date>
(
<emphasis id="B9320F13BF20FFDB085EF9659E84AC73" box="[521,735,1772,1796]" italics="true" pageId="25" pageNumber="26">B.L. Fisher et al.</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF20FFDB08A8F96499C0AC50" ID-GBIF-Occurrence="3462705629" pageId="25" pageNumber="26">
<collectingRegion id="49821DE3BF20FFDB08A8F9649FD2AC73" box="[767,905,1772,1796]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Réserve Spéciale de l'Ankarana,
<quantity id="4CBE7EE4BF20FFDB0F7BF9649937AC73" box="[1324,1388,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="26" unit="km" value="7.0">7 km</quantity>
SE Matsaborimanga,
<geoCoordinate id="EE72B5C6BF20FFDB0B31F8879D82AC50" box="[358,473,1807,1831]" degrees="12" direction="south" minutes="54" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="-12.9">12° 54' S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF20FFDB0BBCF8879E04AC50" box="[491,607,1807,1831]" degrees="49" direction="east" minutes="07" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="49.116665">49° 07' E</geoCoordinate>
,
<quantity id="4CBE7EE4BF20FFDB0826F8879EE4AC50" box="[625,703,1807,1831]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="26" unit="m" value="150.0">150 m</quantity>
, rainforest &amp; tropical dry forest,
<date id="FFF8F5C1BF20FFDB0E05F8879959AC50" box="[1106,1282,1807,1831]" pageId="25" pageNumber="26" value="1990-11-27" valueMax="1990-11-28" valueMin="1990-11-27">27.28.XI.1990</date>
(P.S. Ward);
</materialsCitation>
<materialsCitation id="3B2ED95CBF20FFDB0AC0F8BC9FD7AC07" ID-GBIF-Occurrence="3462705693" pageId="25" pageNumber="26">
<collectingRegion id="49821DE3BF20FFDB0AC0F8BC9D7AAC3B" box="[151,289,1844,1868]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Réserve Spéciale de l'Ankarana,
<quantity id="4CBE7EE4BF20FFDB08F7F8BC9EA6AC3B" box="[672,765,1844,1868]" metricMagnitude="4" metricUnit="m" metricValue="2.29" pageId="25" pageNumber="26" unit="km" value="22.9">22.9 km</quantity>
224° SW Anivorano Nord,
<geoCoordinate id="EE72B5C6BF20FFDB0E6BF8BC98E6AC3B" box="[1084,1213,1844,1868]" degrees="12.90889" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-12.90889">12.90889 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF20FFDB0E9CF8BC9914AC3C" box="[1227,1359,1844,1868]" degrees="49.10983" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="49.10983">49.10983 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF20FFDB0F0CF8BC99CCAC3C" box="[1371,1431,1844,1868]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="26" unit="m" value="80.0">80 m</quantity>
, sifted litter, tropical dry forest,
<date id="FFF8F5C1BF20FFDB0BA8F8D09EFFAC07" box="[511,676,1880,1904]" pageId="25" pageNumber="26" value="2001-02-10" valueMax="2001-02-16" valueMin="2001-02-10">10.16.II.2001</date>
(
<emphasis id="B9320F13BF20FFDB08E0F8D19F20AC07" box="[695,891,1880,1904]" italics="true" pageId="25" pageNumber="26">B.L. Fisher et al.</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF20FFDB09C2F8D099C7ACE3" ID-GBIF-Occurrence="3462706270" pageId="25" pageNumber="26">
<collectingRegion id="49821DE3BF20FFDB09C2F8D09844AC07" box="[917,1055,1880,1904]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Réserve Spéciale de l'Ankarana,
<quantity id="4CBE7EE4BF20FFDB0AC0F8F49CAEACE3" box="[151,245,1916,1940]" metricMagnitude="4" metricUnit="m" metricValue="2.29" pageId="25" pageNumber="26" unit="km" value="22.9">22.9 km</quantity>
224° SW Anivorano Nord,
<geoCoordinate id="EE72B5C6BF20FFDB086FF8F49EE0ACE3" box="[568,699,1916,1940]" degrees="12.90889" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-12.90889">12.90889 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF20FFDB089EF8F49F15ACE4" box="[713,846,1916,1940]" degrees="49.10983" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="49.10983">49.10983 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF20FFDB090BF8F49FC3ACE4" box="[860,920,1916,1940]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="26" unit="m" value="80.0">80 m</quantity>
, tropical dry forest,
<date id="FFF8F5C1BF20FFDB0ED5F8F498ACACE3" box="[1154,1271,1916,1940]" pageId="25" pageNumber="26" value="2001-02-12">12.II.2001</date>
(
<emphasis id="B9320F13BF20FFDB0F5EF8F499D0ACE3" box="[1289,1419,1916,1940]" italics="true" pageId="25" pageNumber="26">G.D. Alpert</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF20FFDB0AC0F8289F7DACAB" ID-GBIF-Occurrence="3462706279" pageId="25" pageNumber="26">
<collectingRegion id="49821DE3BF20FFDB0AC0F8289D7AACCF" box="[151,289,1952,1976]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Réserve Spéciale de l'Ankarana,
<quantity id="4CBE7EE4BF20FFDB08F4F8289F5BACCF" box="[675,768,1952,1976]" metricMagnitude="4" metricUnit="m" metricValue="1.3599999999999999" pageId="25" pageNumber="26" unit="km" value="13.6">13.6 km</quantity>
192° SSW Anivorano Nord,
<geoCoordinate id="EE72B5C6BF20FFDB0E07F8289889ACCF" box="[1104,1234,1952,1976]" degrees="12.86361" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-12.86361">12.86361 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF20FFDB0EB7F828993EACCF" box="[1248,1381,1952,1976]" degrees="49.22583" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="49.22583">49.22583 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF20FFDB0F25F8289CF4ACAC" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="25" pageNumber="26" unit="m" value="210.0">210 m</quantity>
, tropical dry forest,
<date id="FFF8F5C1BF20FFDB0BCEF84C9E65ACAB" box="[409,574,1988,2012]" pageId="25" pageNumber="26" value="2001-02-16" valueMax="2001-02-21" valueMin="2001-02-16">16.21.II.2001</date>
(
<emphasis id="B9320F13BF20FFDB0806F84D9F4EACAB" box="[593,789,1988,2012]" italics="true" pageId="25" pageNumber="26">B.L. Fisher et al.</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF20FFD80978F84C9814ABA3" ID-GBIF-Occurrence="3462705725" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<collectingRegion id="49821DE3BF20FFDB0978F84C9FE2ACAB" box="[815,953,1988,2012]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
, Réserve Spéciale de l'Ankarana,
<quantity id="4CBE7EE4BF20FFDB0F69F84C99C7ACAB" box="[1342,1436,1988,2012]" metricMagnitude="4" metricUnit="m" metricValue="1.3599999999999999" pageId="25" pageNumber="26" unit="km" value="13.6">13.6 km</quantity>
192° SSW Anivorano Nord,
<geoCoordinate id="EE72B5C6BF23FFD80BB0FF1F9E37ABD8" box="[487,620,151,175]" degrees="12.86361" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="1" value="-12.86361">12.86361 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF23FFD8082BFF1F9F59ABD8" box="[636,770,151,175]" degrees="49.22583" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="1" value="49.22583">49.22583 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF23FFD80946FF1F9F05ABD8" box="[785,862,151,175]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="26" pageNumber="27" unit="m" value="210.0">210 m</quantity>
, tropical dry forest,
<date id="FFF8F5C1BF23FFD80E19FF1F98A8ABD8" box="[1102,1267,151,175]" pageId="26" pageNumber="27" value="2001-02-19" valueMax="2001-02-20" valueMin="2001-02-19">19.20.II.2001</date>
(
<emphasis id="B9320F13BF23FFD80F50FF1099D1ABC7" box="[1287,1418,152,176]" italics="true" pageId="26" pageNumber="27">G.D. Alpert</emphasis>
);
<collectingRegion id="49821DE3BF23FFD80AC0FF349D7AABA3" box="[151,289,188,212]" country="Madagascar" name="Antsiranana" pageId="26" pageNumber="27">Antsiranana</collectingRegion>
, Diego Surez, above Sakaramy,
<quantity id="4CBE7EE4BF23FFD808DCFF349E8FABA4" box="[651,724,188,212]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="m" value="500.0">500 m</quantity>
,
<date id="FFF8F5C1BF23FFD808B7FF349F0FABA3" box="[736,852,188,212]" pageId="26" pageNumber="27" value="1972-02-12">12.II.1972</date>
(
<emphasis id="B9320F13BF23FFD80932FF359865ABA3" box="[869,1086,188,212]" italics="true" pageId="26" pageNumber="27">W.L. &amp; D.E. Brown</emphasis>
).
</materialsCitation>
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>