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<document id="5BB4AE862506782436650198AEA3D168" ID-CLB-Dataset="4918" ID-DOI="10.11646/zootaxa.4985.1.7" ID-GBIF-Dataset="6db5a7ca-bafb-40ba-89a1-350157ffb16f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4930427" ID-ZooBank="2CD17777-AA5B-45BC-A69D-B5C61C96FB84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623443952549" checkinUser="plazi" docAuthor="Matsumoto, Keita" docDate="2021" docId="03E8A1587751FFC81BB054855E33FC71" docLanguage="en" docName="zootaxa.4985.1.7.pdf" docOrigin="Zootaxa 4985 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Byrrhinus angolensis Matsumoto 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="117" masterDocId="FFD1D9207755FFCE1B2754125F4EFFF1" masterDocTitle="New species and records of Angolan Limnichidae (Coleoptera: Byrrhoidea)" masterLastPageNumber="117" masterPageNumber="111" pageNumber="115" updateTime="1699013244882" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="DD2DBB07B3170916A8525F18A9B0BC59" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CD8F0207F785729F0EF2D471B209BF94">
<mods:title id="168A0293BA9BEDD3707BA655E8AD951D">New species and records of Angolan Limnichidae (Coleoptera: Byrrhoidea)</mods:title>
</mods:titleInfo>
<mods:name id="331E3C840D6D03091D21EBA1A2F389F4" type="personal">
<mods:role id="AE0C399BE06722B0DD8AF27DB6CA35E8">
<mods:roleTerm id="82DA25008E23AB0364A8C697E12E7F62">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E8612A57E1F05304912EB496E004F993">Matsumoto, Keita</mods:namePart>
<mods:nameIdentifier id="4FDF79AE11368D1839A749C9265FC68F" type="ORCID">0000-0002-0527-7418</mods:nameIdentifier>
<mods:affiliation id="F5721E6246EAEEA53A8F35800FF22233">Department of Life Sciences, Natural History Museum, London, SW 7 5 BD, United Kingdom. &amp; National Geographic Okavango Wilderness Project, Wild Bird Trust, Parktown 2193, South Africa. https: // orcid. org / 0000 - 0002 - 0527 - 7418</mods:affiliation>
</mods:name>
<mods:typeOfResource id="15A392BF94185D3F9BCC4E42ABAAC790">text</mods:typeOfResource>
<mods:relatedItem id="4592C47B5E0D366D0DAB9869A7B79B95" type="host">
<mods:titleInfo id="C50A72D0298AE788F42A447CDEAC9874">
<mods:title id="9DADB8784270B422F651DB5F150DC0A0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AF825166B9C25607F67456F03BEEAFED">
<mods:date id="A1CD9AECB041BDCB68FA1B25B76F736A">2021</mods:date>
<mods:detail id="AD0A9D9830FF78EEB590403CEFD23243" type="pubDate">
<mods:number id="161FD243EFAA3A0BB48FD4FCBFC71718">2021-06-11</mods:number>
</mods:detail>
<mods:detail id="8DD631876D55CD0EA54C638A0B9EF0C4" type="volume">
<mods:number id="E455E60121517F1C2F0BDE834CA8B3CA">4985</mods:number>
</mods:detail>
<mods:detail id="2422D2DE9D1DCD6326FDC71C86869937" type="issue">
<mods:number id="172E9A5DDD175E3F0EA06BE7C96F140B">1</mods:number>
</mods:detail>
<mods:extent id="8C18A768CC72ADB6B3AEF6B9D4E0ED86" unit="page">
<mods:start id="ECE7172C1CD4E5EAFF20440D590F02DE">111</mods:start>
<mods:end id="A1644A82B1977A635B8647CB76D48AF1">117</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="34ACB74E1DBB6B089FC4A28A3E812DCB">journal article</mods:classification>
<mods:identifier id="5FE2FCF5C852791C47946566DCA57CB9" type="CLB-Dataset">4918</mods:identifier>
<mods:identifier id="FA7AF9835CE3B07B9AA4FDDD575F356C" type="DOI">10.11646/zootaxa.4985.1.7</mods:identifier>
<mods:identifier id="C4851D855E718B2032CA7C84B3BE7D0D" type="GBIF-Dataset">6db5a7ca-bafb-40ba-89a1-350157ffb16f</mods:identifier>
<mods:identifier id="994D8A171EFE7A609047DA023B7604D0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DCBA67F70351E0FCA4C5C0BDC0B19A89" type="Zenodo-Dep">4930427</mods:identifier>
<mods:identifier id="8B122DE88E4E4F50BBB11096B74F6D85" type="ZooBank">2CD17777-AA5B-45BC-A69D-B5C61C96FB84</mods:identifier>
</mods:mods>
<treatment id="03E8A1587751FFC81BB054855E33FC71" ID-DOI="http://doi.org/10.5281/zenodo.4930627" ID-GBIF-Taxon="184605074" ID-Zenodo-Dep="4930627" LSID="urn:lsid:plazi:treatment:03E8A1587751FFC81BB054855E33FC71" httpUri="http://treatment.plazi.org/id/03E8A1587751FFC81BB054855E33FC71" lastPageId="6" lastPageNumber="117" pageId="4" pageNumber="115">
<subSubSection id="C35B43C57751FFCA1BB054855C12FF40" box="[151,860,151,178]" pageId="4" pageNumber="115" type="nomenclature">
<paragraph id="8BFE104E7751FFCA1BB054855C12FF40" blockId="4.[151,860,151,178]" box="[151,860,151,178]" pageId="4" pageNumber="115">
<heading id="D0B6A7227751FFCA1BB054855C12FF40" bold="true" box="[151,860,151,178]" fontSize="11" level="1" pageId="4" pageNumber="115" reason="1">
<emphasis id="B935CC5C7751FFCA1BB054855C12FF40" bold="true" box="[151,860,151,178]" pageId="4" pageNumber="115">
<taxonomicName id="4C416BCD7751FFCA1BB054855D7CFF40" authority="Matsumoto" authorityName="Matsumoto" authorityYear="2021" box="[151,562,151,178]" class="Insecta" family="Limnichidae" genus="Byrrhinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="angolensis" status="sp. nov.">
<emphasis id="B935CC5C7751FFCA1BB054855ED5FF43" bold="true" box="[151,411,151,178]" italics="true" pageId="4" pageNumber="115">Byrrhinus angolensis</emphasis>
Matsumoto
</taxonomicName>
<taxonomicNameLabel id="A20671277751FFCA191E54855DD9FF40" box="[569,663,151,177]" pageId="4" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
[
<figureCitation id="137A0CCB7751FFCA198254855C1AFF40" box="[677,852,151,177]" captionStart="Figures 10‒12" captionStartId="4.[151,232,1636,1660]" captionTargetBox="[248,1338,643,1601]" captionTargetId="figure-171@4.[239,1349,633,1613]" captionText="Figures 10‒12. Byrrhinus angolensis Matsumoto sp. nov. 10, Dorsal habitus; 11, abdomen; 12, genitalia, ventral" figureDoi="http://doi.org/10.5281/zenodo.5224389" httpUri="https://zenodo.org/record/5224389/files/figure.png" pageId="4" pageNumber="115">Figures 10-12</figureCitation>
]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35B43C57751FFCA1BB054CD5D88FE04" pageId="4" pageNumber="115" type="materials_examined">
<paragraph id="8BFE104E7751FFCA1BB054CD5DD5FF08" blockId="4.[151,1436,223,610]" box="[151,667,223,249]" pageId="4" pageNumber="115">
<materialsCitation id="3B291A137751FFCA1BB054CD5DD9FF08" ID-GBIF-Occurrence="3336404304" box="[151,663,223,249]" country="Angola" location="Cassamba" municipality="Luio River" pageId="4" pageNumber="115" specimenCount="1" typeStatus="holotype">
<emphasis id="B935CC5C7751FFCA1BB054CD5E78FF08" bold="true" box="[151,310,223,249]" pageId="4" pageNumber="115">
<typeStatus id="54FAAEEC7751FFCA1BB054CD5F9CFF08" box="[151,210,223,249]" pageId="4" pageNumber="115">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F35650DE7751FFCA1A1C54CD5EDEFF08" box="[315,400,223,249]" name="Angola" pageId="4" pageNumber="115">Angola</collectingCountry>
,
<location id="8E9E46957751FFCA1ABC54CD5D41FF08" LSID="urn:lsid:plazi:treatment:03E8A1587751FFC81BB054855E33FC71:8E9E46957751FFCA1ABC54CD5D41FF08" box="[411,527,223,249]" country="Angola" municipality="Luio River" name="Cassamba" pageId="4" pageNumber="115">Cassamba</location>
,
<collectingMunicipality id="6B9A8A347751FFCA193C54CD5DD9FF08" box="[539,663,223,249]" pageId="4" pageNumber="115">Luio River</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE104E7751FFCA1BE055115D88FE04" blockId="4.[151,1436,223,610]" pageId="4" pageNumber="115">
<emphasis id="B935CC5C7751FFCA1BE055115EC0FEEC" bold="true" box="[199,398,259,285]" pageId="4" pageNumber="115">Type materials.</emphasis>
<materialsCitation id="3B291A137751FFCA1A8555115BF3FE78" ID-GBIF-Occurrence="3336404301" collectingDate="2019-11-30" collectingDateMax="2019-12-01" collectingDateMin="2019-11-30" collectionCode="BMNH" collectorName="Matsumoto, K. &amp; Okavango Wilderness Project" country="Angola" determinerName="K. Matsumoto" elevation="1187" latitude="-13.19398" location="Cassamba" longLatPrecision="5" longitude="20.2235" municipality="Luio River" pageId="4" pageNumber="115" specimenCount="2021" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54FAAEEC7751FFCA1A8555115D44FEEC" box="[418,522,259,285]" pageId="4" pageNumber="115" type="holotype">Holotype</typeStatus>
male, deposited at
<collectionCode id="ED50888B7751FFCA183D55165C20FEEF" box="[794,878,260,286]" pageId="4" pageNumber="115">BMNH</collectionCode>
“
<collectingCountry id="F35650DE7751FFCA18AB55115B4AFEEC" box="[908,1028,259,285]" name="Angola" pageId="4" pageNumber="115">ANGOLA</collectingCountry>
<quantity id="4CB9BDAB7751FFCA1F3E55115B26FEEC" box="[1049,1128,259,285]" metricMagnitude="3" metricUnit="m" metricValue="1.187" pageId="4" pageNumber="115" unit="m" value="1187.0">
<elevation id="006CF77D7751FFCA1F3E55115B26FEEC" box="[1049,1128,259,285]" metricMagnitude="3" metricUnit="m" metricValue="1.187" pageId="4" pageNumber="115" unit="m" value="1187.0">1187m</elevation>
</quantity>
,
<location id="8E9E46957751FFCA1FA655115BBBFEEC" LSID="urn:lsid:plazi:treatment:03E8A1587751FFC81BB054855E33FC71:8E9E46957751FFCA1FA655115BBBFEEC" box="[1153,1269,259,285]" country="Angola" latitude="-13.19398" longLatPrecision="5" longitude="20.2235" municipality="Luio River" name="Cassamba" pageId="4" pageNumber="115">Cassamba</location>
,
<collectingMunicipality id="6B9A8A347751FFCA1E2855115AD6FEEC" box="[1295,1432,259,285]" pageId="4" pageNumber="115">Luio River</collectingMunicipality>
,
<geoCoordinate id="EE7576897751FFCA1BB055355E5CFEB3" box="[151,274,295,322]" degrees="13.19398" direction="south" orientation="latitude" pageId="4" pageNumber="115" precision="1" value="-13.19398">13.19398S</geoCoordinate>
,
<geoCoordinate id="EE7576897751FFCA1A3255355EC9FEB3" box="[277,391,295,322]" degrees="20.2235" direction="east" orientation="longitude" pageId="4" pageNumber="115" precision="5" value="20.2235">20.2235E</geoCoordinate>
;
<date id="FFFF368E7751FFCA1AB255355D02FEB0" box="[405,588,295,321]" pageId="4" pageNumber="115" value="2019-11-30" valueMax="2019-12-01" valueMin="2019-11-30">
<collectingDate id="EFBBCF667751FFCA1AB255355D02FEB0" box="[405,588,295,321]" pageId="4" pageNumber="115" value="2019-11-30" valueMax="2019-12-01" valueMin="2019-11-30">30.xi-1.xii.2019</collectingDate>
</date>
, // General Coll. &amp; UV LED,
<collectorName id="26B475987751FFCA1891553A5B2DFEB3" box="[950,1123,296,322]" pageId="4" pageNumber="115">Matsumoto, K.</collectorName>
leg.
<collectorName id="26B475987751FFCA1F8555355FA7FE94" pageId="4" pageNumber="115">Okavango Wilderness Project</collectorName>
,
<collectionCode id="ED50888B7751FFCA1BDC555E5E01FE97" box="[251,335,332,358]" pageId="4" pageNumber="115">BMNH</collectionCode>
(E) 2020-30” with unique identification number “NHMUK014383825”. Additional label on red paper added “
<taxonomicName id="4C416BCD7751FFCA1A4C557D5D10FE78" authority="Matsumoto, 2021" authorityName="Matsumoto" authorityYear="2021" box="[363,606,367,393]" class="Insecta" family="Limnichidae" genus="Byrrhinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="angolensis" status="sp. nov.">
<emphasis id="B935CC5C7751FFCA1A4C557D5D10FE78" box="[363,606,367,393]" italics="true" pageId="4" pageNumber="115">Byrrhinus angolensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20671277751FFCA194D55625D8BFE7B" box="[618,709,368,394]" pageId="4" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
<determinerName id="7E22FC8A7751FFCA19F755625C39FE7B" box="[720,887,368,394]" pageId="4" pageNumber="115">K. Matsumoto</determinerName>
det.
<specimenCount id="9D47DBC77751FFCA1890557D5BFCFE78" box="[951,1202,367,393]" hasTypeStatus="true" pageId="4" pageNumber="115">
2020
<typeStatus id="54FAAEEC7751FFCA18DD557D5BC0FE78" box="[1018,1166,367,393]" pageId="4" pageNumber="115" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D47DBC77751FFCA1FBD557D5BFCFE78" box="[1178,1202,367,393]" pageId="4" pageNumber="115" type="male">♂</specimenCount>
</specimenCount>
”
</materialsCitation>
.
<materialsCitation id="3B291A137751FFCA1FE955625E06FE5C" ID-GBIF-Occurrence="3336404307" collectionCode="BMNH, AMG" country="Angola" pageId="4" pageNumber="115" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54FAAEEC7751FFCA1FE955625A7EFE7B" box="[1230,1328,368,394]" pageId="4" pageNumber="115" type="paratype">Paratype</typeStatus>
(
<specimenCount id="9D47DBC77751FFCA1E63557D5AD6FE78" box="[1348,1432,367,393]" pageId="4" pageNumber="115" type="male">2 male</specimenCount>
;
<collectionCode id="ED50888B7751FFCA1BB055865FA0FE5F" box="[151,238,404,430]" pageId="4" pageNumber="115">BMNH</collectionCode>
,
<collectionCode id="ED50888B7751FFCA1BDE55815E0EFE5C" box="[249,320,403,429]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="4" pageNumber="115">AMG</collectionCode>
)
</materialsCitation>
:
<materialsCitation id="3B291A137751FFCA1A7455865D88FE04" ID-GBIF-Occurrence="3336404302" collectionCode="BMNH, AMG" country="Angola" determinerName="K. Matsumoto" pageId="4" pageNumber="115" specimenCount="2021" specimenCount-male="1" typeStatus="holotype">
same locality data as the
<typeStatus id="54FAAEEC7751FFCA195655815D98FE5C" box="[625,726,403,429]" pageId="4" pageNumber="115" type="holotype">holotype</typeStatus>
, with unique identification number “NHMUK014383826” and “NHMUK014375572”.All
<typeStatus id="54FAAEEC7751FFCA1AEA55AA5D63FE23" box="[461,557,440,466]" pageId="4" pageNumber="115" type="paratype">paratype</typeStatus>
specimens have been labelled with a red
<typeStatus id="54FAAEEC7751FFCA18DF55AA5B16FE23" box="[1016,1112,440,466]" pageId="4" pageNumber="115" type="paratype">paratype</typeStatus>
label: “
<taxonomicName id="4C416BCD7751FFCA1F8855A55AD2FE20" authority="Matsumoto, 2021" authorityName="Matsumoto" authorityYear="2021" box="[1199,1436,439,465]" class="Insecta" family="Limnichidae" genus="Byrrhinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="angolensis" status="sp. nov.">
<emphasis id="B935CC5C7751FFCA1F8855A55AD2FE20" box="[1199,1436,439,465]" italics="true" pageId="4" pageNumber="115">Byrrhinus angolensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20671277751FFCA1BB055CE5FA0FE07" box="[151,238,476,502]" pageId="4" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
<determinerName id="7E22FC8A7751FFCA1BD355CE5ED8FE07" box="[244,406,476,502]" pageId="4" pageNumber="115">K. Matsumoto</determinerName>
det.
<specimenCount id="9D47DBC77751FFCA1AEA55C95DF8FE04" box="[461,694,475,501]" hasTypeStatus="true" pageId="4" pageNumber="115">
2020
<typeStatus id="54FAAEEC7751FFCA192B55C95DD9FE04" box="[524,663,475,501]" pageId="4" pageNumber="115" type="paratype">PARATYPE</typeStatus>
<specimenCount id="9D47DBC77751FFCA19B955C95DF8FE04" box="[670,694,475,501]" pageId="4" pageNumber="115" type="male">♂</specimenCount>
</specimenCount>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35B43C57751FFC81BE055ED5EECFD75" lastPageId="6" lastPageNumber="117" pageId="4" pageNumber="115" type="description">
<paragraph id="8BFE104E7751FFCA1BE055ED5B61FD93" blockId="4.[151,1436,223,610]" pageId="4" pageNumber="115">
<emphasis id="B935CC5C7751FFCA1BE055ED5D39FDE8" bold="true" box="[199,631,511,537]" pageId="4" pageNumber="115">
Description of
<typeStatus id="54FAAEEC7751FFCA1A5255ED5E93FDE8" box="[373,477,511,537]" pageId="4" pageNumber="115" type="holotype">holotype</typeStatus>
, male. Body:
</emphasis>
Elongate, oval, dorsum convex, BL:
<quantity id="4CB9BDAB7751FFCA1F3255ED5B22FDEB" box="[1045,1132,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="115" unit="mm" value="1.9">1.9 mm</quantity>
, EL:
<quantity id="4CB9BDAB7751FFCA1F8155ED5BB0FDEB" box="[1190,1278,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="115" unit="mm" value="1.5">1.5 mm</quantity>
, EW:
<quantity id="4CB9BDAB7751FFCA1E6655ED5AD6FDEB" box="[1345,1432,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="115" unit="mm" value="1.1">1.1 mm</quantity>
, PL:
<quantity id="4CB9BDAB7751FFCA1BE056315E51FDCF" box="[199,287,547,574]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="115" unit="mm" value="0.4">0.4 mm</quantity>
, PW: 1.0 mm and HW:
<quantity id="4CB9BDAB7751FFCA190A56315DC8FDCF" box="[557,646,547,574]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="115" unit="mm" value="0.5">0.5 mm</quantity>
.
<emphasis id="B935CC5C7751FFCA19B656315DA1FDCC" bold="true" box="[657,751,547,573]" pageId="4" pageNumber="115">Colour:</emphasis>
Dorsal side black; venter brown to dark reddish brown, tibia brown, tarsal claws brown. Dorsal and ventral surface densely covered with setae.
</paragraph>
<caption id="DF3E40C67751FFCA1BB052765A5DF98D" ID-DOI="http://doi.org/10.5281/zenodo.5224389" ID-Zenodo-Dep="5224389" box="[151,1299,1636,1661]" httpUri="https://zenodo.org/record/5224389/files/figure.png" pageId="4" pageNumber="115" startId="4.[151,232,1636,1660]" targetBox="[248,1338,643,1601]" targetPageId="4">
<paragraph id="8BFE104E7751FFCA1BB052765A5DF98D" blockId="4.[151,1299,1636,1661]" box="[151,1299,1636,1661]" pageId="4" pageNumber="115">
<emphasis id="B935CC5C7751FFCA1BB052765E7BF98D" bold="true" box="[151,309,1636,1660]" pageId="4" pageNumber="115">Figures 10‒12.</emphasis>
<taxonomicName id="4C416BCD7751FFCA1A1C52765DC2F98D" authority="Matsumoto" authorityName="Matsumoto" authorityYear="2021" box="[315,652,1636,1660]" class="Insecta" family="Limnichidae" genus="Byrrhinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="angolensis" status="sp. nov.">
<emphasis id="B935CC5C7751FFCA1A1C52765D5CF98D" box="[315,530,1636,1660]" italics="true" pageId="4" pageNumber="115">Byrrhinus angolensis</emphasis>
Matsumoto
</taxonomicName>
<emphasis id="B935CC5C7751FFCA19B552775DADF98C" bold="true" box="[658,739,1637,1661]" pageId="4" pageNumber="115">
<taxonomicNameLabel id="A20671277751FFCA19B552775DADF98C" box="[658,739,1637,1661]" pageId="4" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
10, Dorsal habitus; 11, abdomen; 12, genitalia, ventral
</paragraph>
</caption>
<paragraph id="8BFE104E7751FFCA1E3452765A59F98D" blockId="4.[1299,1303,1636,1660]" box="[1299,1303,1636,1660]" pageId="4" pageNumber="115">.</paragraph>
<paragraph id="8BFE104E7751FFCA1BE052B95A4EF814" blockId="4.[151,1436,1706,2021]" pageId="4" pageNumber="115">
<emphasis id="B935CC5C7751FFCA1BE052B95ECEF934" bold="true" box="[199,384,1707,1733]" pageId="4" pageNumber="115">Dorsum. Head:</emphasis>
Partially retracted into pronotum; overall surface smooth, no punctation across surface.
<emphasis id="B935CC5C7751FFCA1E4852B85FBEF918" bold="true" pageId="4" pageNumber="115">Antennae:</emphasis>
11 segmented, filiform, fully pubescent, antennomeres long and slender, 1st
<collectingCountry id="F35650DE7751FFCA1F4352DD5BC6F918" box="[1124,1160,1743,1769]" name="American Samoa" pageId="4" pageNumber="115">AS</collectingCountry>
widest, 2nd
<collectingCountry id="F35650DE7751FFCA1E3C52DD5A71F918" box="[1307,1343,1743,1769]" name="American Samoa" pageId="4" pageNumber="115">AS</collectingCountry>
slightly narrower than 1st
<collectingCountry id="F35650DE7751FFCA1A4352E15EC4F8FC" box="[356,394,1779,1805]" name="American Samoa" pageId="4" pageNumber="115">AS</collectingCountry>
, 3rd to 6th
<collectingCountry id="F35650DE7751FFCA192B52E15D7EF8FC" box="[524,560,1779,1805]" name="American Samoa" pageId="4" pageNumber="115">AS</collectingCountry>
long and thin, 7th to 11th
<collectingCountry id="F35650DE7751FFCA187A52E15CCFF8FC" box="[861,897,1779,1805]" name="American Samoa" pageId="4" pageNumber="115">AS</collectingCountry>
skewed oval.
<emphasis id="B935CC5C7751FFCA1F0352E15BE9F8FC" bold="true" box="[1060,1191,1779,1805]" pageId="4" pageNumber="115">Pronotum:</emphasis>
Transverse, widest at posterior, narrowest at anterior, narrower than elytra; anterior margin concave, posterior margin strongly sinuate; lateral margin nearly straight; surface covered with moderately dense punctation, surface between punctures smooth and shiny.
<emphasis id="B935CC5C7751FFCA1A28534C5EDFF889" bold="true" box="[271,401,1886,1912]" pageId="4" pageNumber="115">Scutellum:</emphasis>
Subtriangular, large, lateral sides slightly convex. Elytra: Longer than wide; anterior margin sinuate; anterior lateral angle obtuse; lateral margins gently curve from anterior end to anterior 2/3, increase in curvature towards the apex; surface with small and moderately dense punctuation; punctuation decreases in depth towards the elytral tip; elytral striae not clearly visible; surface between punctures smooth and shiny.
</paragraph>
<caption id="DF3E40C67750FFCB1BB053855B94F841" ID-DOI="http://doi.org/10.5281/zenodo.5224391" ID-Zenodo-Dep="5224391" box="[151,1242,1943,1968]" httpUri="https://zenodo.org/record/5224391/files/figure.png" pageId="5" pageNumber="116" startId="5.[151,232,1943,1967]" targetBox="[223,1365,184,1919]" targetPageId="5">
<paragraph id="8BFE104E7750FFCB1BB053855B94F841" blockId="5.[151,1242,1943,1968]" box="[151,1242,1943,1968]" pageId="5" pageNumber="116">
<emphasis id="B935CC5C7750FFCB1BB053855E7BF85E" bold="true" box="[151,309,1943,1967]" pageId="5" pageNumber="116">Figures 13‒14.</emphasis>
<taxonomicName id="4C416BCD7750FFCB1A1C538A5EF5F841" authorityName="Matsumoto" authorityYear="2021" box="[315,443,1944,1968]" class="Insecta" family="Limnichidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="116" phylum="Arthropoda" rank="family">Limnichidae</taxonomicName>
habitat. 13, Kangumbe, Lungué-Bungo River; 14, Cassamba, near Luio River.
</paragraph>
</caption>
<paragraph id="8BFE104E7753FFC81BE054845EC4FE5C" blockId="6.[151,1437,150,896]" pageId="6" pageNumber="117">
<emphasis id="B935CC5C7753FFC81BE054845E52FF41" bold="true" box="[199,284,150,176]" pageId="6" pageNumber="117">Venter.</emphasis>
Overall surface with moderately dense micro-punctures with setae emerging from the punctures.
<emphasis id="B935CC5C7753FFC81E4F54845E4AFF25" bold="true" pageId="6" pageNumber="117">Prosternum:</emphasis>
Concave on anterior margin; prosternal process as long than wide, lateral margin concave, posterior margin semi-oval.
<emphasis id="B935CC5C7753FFC81A4F54CC5D40FF09" bold="true" box="[360,526,222,248]" pageId="6" pageNumber="117">Mesoventrite:</emphasis>
Wider than long, narrows towards posterior; anterior margin concave where prosternal process fits, discrimen very small; lateral sub-parallel and gently curved, posterior nearly flat, anterior trisinuate.
<emphasis id="B935CC5C7753FFC81BD255345ED4FEB1" bold="true" box="[245,410,294,320]" pageId="6" pageNumber="117">Metaventrite:</emphasis>
Wider than long, flat in the middle; discrimen thin and not clearly visible.
<emphasis id="B935CC5C7753FFC81FC255345A2DFEB1" bold="true" box="[1253,1379,294,320]" pageId="6" pageNumber="117">Abdomen:</emphasis>
Five clearly visible ventrites. Intercoxal process sinuate with pointed apex. Posterior margin of 1st and 2nd ventrites convex, 3rd to 4th ventrites slightly concave, 5th ventrite nearly semi-circle. Surface covered with long, light yellow, recumbent setae.
</paragraph>
<paragraph id="8BFE104E7753FFC81BE055A45DF8FE05" blockId="6.[151,1437,150,896]" pageId="6" pageNumber="117">
<emphasis id="B935CC5C7753FFC81BE055A45E2DFE21" bold="true" box="[199,355,438,464]" pageId="6" pageNumber="117">Legs. Tibiae:</emphasis>
Outer and inner lateral margin convex.
<emphasis id="B935CC5C7753FFC8180C55A45C3CFE21" bold="true" box="[811,882,438,464]" pageId="6" pageNumber="117">Tarsi:</emphasis>
Simple, segments increase in length towards the apex.
<emphasis id="B935CC5C7753FFC81BFD55C85E39FE05" bold="true" box="[218,375,474,500]" pageId="6" pageNumber="117">Tarsal claws:</emphasis>
Short, narrow, symmetrical.
</paragraph>
<paragraph id="8BFE104E7753FFC81BE055EC5AC6FD90" blockId="6.[151,1437,150,896]" pageId="6" pageNumber="117">
<emphasis id="B935CC5C7753FFC81BE055EC5E73FDE9" bold="true" box="[199,317,510,536]" pageId="6" pageNumber="117">Genitalia.</emphasis>
Parameres twice as long as median lobe, long and slender, symmetrical, widest as base, apex acute, inner side near the apical end straight. Median lobe apex acute and slightly rounded, base to anterior 0.25 relatively straight, gentle convex curve near the apex. Phallobase strongly asymmetrical. Relatively sclerotised (
<figureCitation id="137A0CCB7753FFC81E2A56545A34FD90" box="[1293,1402,582,609]" captionStart="Figures 6‒9" captionStartId="3.[151,232,1306,1330]" captionTargetBox="[223,1361,196,1270]" captionTargetId="figure-16@3.[215,1371,181,1282]" captionText="Figures 6‒9. Simplocarina angolensis Matsumoto sp. nov. 6, Dorsal habitus; 7, abdomen; 8, genitalia, ventral; 9, genitalia, lateral." figureDoi="http://doi.org/10.5281/zenodo.5224387" httpUri="https://zenodo.org/record/5224387/files/figure.png" pageId="6" pageNumber="117">Figures 7</figureCitation>
).
</paragraph>
<paragraph id="8BFE104E7753FFC81BE056785EECFD75" blockId="6.[151,1437,150,896]" box="[199,418,618,644]" pageId="6" pageNumber="117">
<emphasis id="B935CC5C7753FFC81BE056785E6BFD75" bold="true" box="[199,293,618,644]" pageId="6" pageNumber="117">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35B43C57753FFC81BE0569C5FBAFD3C" pageId="6" pageNumber="117" type="distribution">
<paragraph id="8BFE104E7753FFC81BE0569C5FBAFD3C" blockId="6.[151,1437,150,896]" pageId="6" pageNumber="117">
<emphasis id="B935CC5C7753FFC81BE0569C5E2FFD59" bold="true" box="[199,353,654,680]" pageId="6" pageNumber="117">Distribution.</emphasis>
<taxonomicName id="4C416BCD7753FFC81A4E569C5DACFD58" authority="Matsumoto" authorityName="Matsumoto" authorityYear="2021" box="[361,738,654,681]" class="Insecta" family="Limnichidae" genus="Byrrhinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="angolensis" status="sp. nov.">
<emphasis id="B935CC5C7753FFC81A4E569C5D14FD59" box="[361,602,654,680]" italics="true" pageId="6" pageNumber="117">Byrrhinus angolensis</emphasis>
Matsumoto
</taxonomicName>
<emphasis id="B935CC5C7753FFC819CC569C5C08FD59" bold="true" box="[747,838,654,680]" pageId="6" pageNumber="117">
<taxonomicNameLabel id="A20671277753FFC819CC569C5C08FD59" box="[747,838,654,680]" pageId="6" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="54FAAEEC7753FFC81F42569D5BDBFD58" box="[1125,1173,655,681]" pageId="6" pageNumber="117">type</typeStatus>
locality (
<figureCitation id="137A0CCB7753FFC81E24569C5A3DFD59" box="[1283,1395,654,680]" captionStart="Figures 13‒14" captionStartId="5.[151,232,1943,1967]" captionTargetBox="[223,1365,184,1919]" captionTargetId="figure-16@5.[222,1365,181,1919]" captionText="Figures 13‒14. Limnichidae habitat. 13, Kangumbe, Lungué-Bungo River; 14, Cassamba, near Luio River." figureDoi="http://doi.org/10.5281/zenodo.5224391" httpUri="https://zenodo.org/record/5224391/files/figure.png" pageId="6" pageNumber="117">Figure 14</figureCitation>
for habitat).
</paragraph>
</subSubSection>
<subSubSection id="C35B43C57753FFC81BE056C45C85FD01" box="[199,971,726,753]" pageId="6" pageNumber="117" type="etymology">
<paragraph id="8BFE104E7753FFC81BE056C45C85FD01" blockId="6.[151,1437,150,896]" box="[199,971,726,753]" pageId="6" pageNumber="117">
<emphasis id="B935CC5C7753FFC81BE056C45E00FD01" bold="true" box="[199,334,726,752]" pageId="6" pageNumber="117">Etymology.</emphasis>
The species named after
<collectingCountry id="F35650DE7753FFC8194E56C45DF0FD01" box="[617,702,726,752]" name="Angola" pageId="6" pageNumber="117">Angola</collectingCountry>
, where it was collected.
</paragraph>
</subSubSection>
<subSubSection id="C35B43C57753FFC81BE056E85E33FC71" pageId="6" pageNumber="117" type="diagnosis">
<paragraph id="8BFE104E7753FFC81BE056E85E33FC71" blockId="6.[151,1437,150,896]" pageId="6" pageNumber="117">
<emphasis id="B935CC5C7753FFC81BE056E85E9FFCE5" bold="true" box="[199,465,762,788]" pageId="6" pageNumber="117">Differential diagnosis.</emphasis>
According to the grouping system of
<bibRefCitation id="EFD06DBF7753FFC818B456E85B77FCE4" author="Deleve, J." box="[915,1081,762,789]" pageId="6" pageNumber="117" pagination="212 - 274" refId="ref2624" refString="Deleve, J. (1968) Contribution a l'etude des &quot; Dryopoidea &quot;. Les Limnichinae d'Afrique (Coleoptera Limnichidae). Bulletin and Annales de la Societe Royale d'Entomologie de Belgique, 104, 212 - 274." type="journal article" year="1968">Delève (1968)</bibRefCitation>
, this new species would be in ‘group one’, which has a triangular median lobe and symmetrical parameres. This new species can be distinguished by the inner side near the apical end of the parameres straight, apex of parameres acute, majority of the median lobe straight and parallel.
</paragraph>
</subSubSection>
</treatment>
</document>
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