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<document id="9D89CCC486A960C7387B6E2FB9A2A1BC" ID-DOI="10.5281/zenodo.176679" ID-GBIF-Dataset="8b730849-180f-4e01-98cf-2f12fdaaea97" ID-ISSN="1175-5326" ID-Zenodo-Dep="176679" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459837893502" checkinUser="plazi" docAuthor="Michalczyk, Łukasz &amp; Kaczmarek, Łukasz" docDate="2007" docId="03BCDE48FFDB6B403ED41B29C9E2149E" docLanguage="en" docName="zt01471p025.pdf" docOrigin="Zootaxa 1471" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Echiniscus ganczareki Michalczyk &amp; Kaczmarek, 2007, sp. nov." docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="FF85A630FFDA6B443E431F53CC691343" masterDocTitle="Echiniscus ganczareki, a new species of Tardigrada (Heterotardigrada: Echiniscidae, bigranulatus group) from Costa Rica" masterLastPageNumber="25" masterPageNumber="15" pageNumber="16" updateTime="1698223920605" updateUser="plazi">
<mods:mods id="006699A22515F5E22904A926FA9A77C1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B890A4772C007F4FFFFC1E7A9F362E9D">
<mods:title id="3AC38267C4DFAE4DEE3710D519C4F4B0">Echiniscus ganczareki, a new species of Tardigrada (Heterotardigrada: Echiniscidae, bigranulatus group) from Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="B70FB5CF567032C57C5079620375CD82" type="personal">
<mods:role id="4266C3C9DF346A303516F9E57AC9B85D">
<mods:roleTerm id="A48B0E32E2589D432606C23F303E69E7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="65203CFBEE92306A645ACF38F552DD08">Michalczyk, Łukasz</mods:namePart>
</mods:name>
<mods:name id="73FDC9E5897BCD935A64F4D8591F8598" type="personal">
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<mods:roleTerm id="3215A7EE5AFA2BE7776BACBE53DB9C0A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7EC6C4961A7D6A6308E9A8182A908271">Kaczmarek, Łukasz</mods:namePart>
</mods:name>
<mods:typeOfResource id="9B5F97366D4BCB7A44F5D625B61F54DB">text</mods:typeOfResource>
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<mods:title id="EA4A43725DE378C764A0D6FA7C95C8FC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="38607C5DFC20E6655C48F1622638C9E1">
<mods:date id="F968855A1A8FE49ED8B93EFEDF1452EA">2007</mods:date>
<mods:detail id="14FF8B87DAF9166B73F27FE5973BE09D" type="volume">
<mods:number id="1EECD49876FBD8674C06472816ECD1AB">1471</mods:number>
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<mods:classification id="DFB255F26A9843BE2E2234E31BDA0BF8">journal article</mods:classification>
<mods:identifier id="4DA5538FA7F589F2E738DA55504C43C3" type="DOI">10.5281/zenodo.176679</mods:identifier>
<mods:identifier id="AD675A76173192EDA1200C02DFA24C53" type="GBIF-Dataset">8b730849-180f-4e01-98cf-2f12fdaaea97</mods:identifier>
<mods:identifier id="AF8070E875D9CF47DB9628CB2E73DC32" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="077EAB5E3ED5E94C48394F921FAB3054" type="Zenodo-Dep">176679</mods:identifier>
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<treatment id="03BCDE48FFDB6B403ED41B29C9E2149E" ID-DOI="http://doi.org/10.5281/zenodo.5612646" ID-GBIF-Taxon="119359431" ID-Zenodo-Dep="5612646" LSID="urn:lsid:plazi:treatment:03BCDE48FFDB6B403ED41B29C9E2149E" httpUri="http://treatment.plazi.org/id/03BCDE48FFDB6B403ED41B29C9E2149E" lastPageId="4" lastPageNumber="19" pageId="1" pageNumber="16">
<subSubSection id="C30F3CD5FFDB6B453ED41B29CD3B17F4" pageId="1" pageNumber="16" type="nomenclature">
<paragraph id="8BAA6F5EFFDB6B453ED41B29CE6517D7" blockId="1.[151,524,1146,1207]" box="[151,524,1146,1172]" pageId="1" pageNumber="16">
<heading id="D0E2D832FFDB6B453ED41B29CE6517D7" bold="true" box="[151,524,1146,1172]" fontSize="11" level="1" pageId="1" pageNumber="16" reason="1">
<emphasis id="B961B34CFFDB6B453ED41B29CE6517D7" bold="true" box="[151,524,1146,1172]" pageId="1" pageNumber="16">
<taxonomicName id="4C1514DDFFDB6B453ED41B29CDCE17D7" box="[151,423,1146,1172]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="1" pageNumber="16" phylum="Tardigrada" rank="species" species="ganczareki" status="sp. nov.">
<emphasis id="B961B34CFFDB6B453ED41B29CDCE17D7" bold="true" box="[151,423,1146,1172]" italics="true" pageId="1" pageNumber="16">Echiniscus ganczareki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2520E37FFDB6B453FEC1B29CE6517D7" box="[431,524,1146,1172]" pageId="1" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA6F5EFFDB6B453ED41BCECD3B17F4" blockId="1.[151,524,1146,1207]" box="[151,338,1181,1207]" pageId="1" pageNumber="16">
(
<figureCitation id="132E73DBFFDB6B453EDC1BCECD7717F4" box="[159,286,1181,1207]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 10" captionStart-2="FIGURES 11 – 14" captionStart-3="FIGURES 15 – 20" captionStart-4="FIGURES 21 – 22" captionStartId-0="2.[151,269,1925,1949]" captionStartId-1="3.[151,269,1576,1600]" captionStartId-2="4.[151,269,1040,1064]" captionStartId-3="6.[151,269,1616,1640]" captionStartId-4="7.[151,269,1708,1732]" captionTargetBox-0="[223,1363,277,1899]" captionTargetBox-1="[164,1423,194,1548]" captionTargetBox-2="[166,1428,167,1015]" captionTargetBox-3="[168,1426,167,1602]" captionTargetBox-4="[154,1433,1192,1684]" captionTargetId-0="figure@2.[223,1363,277,1899]" captionTargetId-1="figure@3.[164,1424,194,1550]" captionTargetId-2="figure@4.[166,1429,167,1016]" captionTargetId-3="figure@6.[168,1427,167,1603]" captionTargetId-4="figure@7.[154,1434,1192,1684]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURES 1 – 4. Echiniscus ganczareki sp. nov. Habitus: 1 — dorsal view (holotype, PCM); 2 — dorso-lateral view (paratype, PCM); 3 — dorsal view (paratype, SEM); 4 — lateral view (paratype, SEM)." captionText-1="FIGURES 5 – 10. Echiniscus ganczareki sp. nov. 5 – 8 — dorsal plate cuticle; 5 — terminal plate with pores; 6 — the same fragment as shown on Fig. 5, but showing pillars that appear when focusing down through the cuticle (5 – 6 – holotype, PCM); 7 — pores in the scapular plate (paratype, SEM); 8 — structure of the dorsal plate cuticle (see also Fig. 25) (paratype, SEM); 9 – 10 — ventral head cuticle with pores (9 — holotype, PCM, 10 — paratype, SEM)." captionText-2="FIGURES 11 – 14. Echiniscus ganczareki sp. nov. 11 – 12 — second median plate, note that the anterior portion of the plate is devoid of pores and is covered with fine granulation (11 — paratype, PCM, 12 — paratype, SEM); 13 – 14 — third median plate (13 — paratype, PCM, 14 — paratype, SEM)." captionText-3="FIGURES 15 – 20. Echiniscus ganczareki sp. nov. Pores on legs: 15 – 16 — legs I with the spine (15 — holotype, PCM, 16 — paratype, SEM); 17 – 18 — legs II (17 — holotype, PCM, 18 — paratype, SEM); 19 – 20 — legs IV with the dentate collar and papilla (19 — holotype, PCM, 20 — paratype, SEM)." captionText-4="FIGURES 21 – 22. Echiniscus ganczareki sp. nov. Claws IV (21 — holotype, PCM, 22 — paratype, SEM)." httpUri-0="https://zenodo.org/record/176680/files/figure.png" httpUri-1="https://zenodo.org/record/176681/files/figure.png" httpUri-2="https://zenodo.org/record/176682/files/figure.png" httpUri-3="https://zenodo.org/record/176683/files/figure.png" httpUri-4="https://zenodo.org/record/176684/files/figure.png" pageId="1" pageNumber="16">Figs. 1–22</figureCitation>
,
<figureCitation id="132E73DBFFDB6B453F681BCECD2317F4" box="[299,330,1181,1207]" captionStart="FIGURES 23 – 26" captionStartId="9.[151,269,1899,1923]" captionTargetBox="[209,1383,1259,1863]" captionTargetId="figure@9.[199,1391,1252,1875]" captionTargetPageId="9" captionText="FIGURES 23 – 26. Structure of the dorsal plate cuticle in four species of the bigranulatus group (semi-schematic sections based on SEM observations). 23 — E. bigranulatus; 24 — E. ollantaytamboensis; 25 — E. ganczareki sp. nov. (see also Fig. 8); 26 — E. madonnae. Abbreviations: bc— body cavity, cs— cuticle surface, pi— pillar, po— pore, pp— pseudopore, sl— ‘ sponge’ layer, up— undeveloped pillar. Not scaled (23 – 24 and 26 from Michalczyk &amp; Kaczmarek 2006 c)." httpUri="https://zenodo.org/record/176685/files/figure.png" pageId="1" pageNumber="16">25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F3CD5FFDB6B453ED41BBECF461614" pageId="1" pageNumber="16" type="materials_examined">
<paragraph id="8BAA6F5EFFDB6B453ED41BBECF461614" blockId="1.[151,1437,1261,2007]" pageId="1" pageNumber="16">
<emphasis id="B961B34CFFDB6B453ED41BBECDE71644" bold="true" box="[151,398,1261,1287]" pageId="1" pageNumber="16">Material examined:</emphasis>
<typeStatus id="54AED1FCFFDB6B453FD51BBECE6A1644" box="[406,515,1261,1287]" pageId="1" pageNumber="16" type="holotype">Holotype</typeStatus>
(female) and
<specimenCount id="9D13A4D7FFDB6B453CEA1BBECF7D1644" box="[681,788,1261,1287]" pageId="1" pageNumber="16" type="adult">24 adults</specimenCount>
(
<specimenCount id="9D13A4D7FFDB6B453D661BBECFCF1644" box="[805,934,1261,1287]" pageId="1" pageNumber="16" type="female">18 females</specimenCount>
+
<specimenCount id="9D13A4D7FFDB6B453D841BBEC84A1644" box="[967,1059,1261,1287]" pageId="1" pageNumber="16" type="male">6 males</specimenCount>
) from the
<typeStatus id="54AED1FCFFDB6B453AE31BBEC8BA1644" box="[1184,1235,1261,1287]" pageId="1" pageNumber="16">type</typeStatus>
locality in
<collectingCountry id="F3022FCEFFDB6B453B1A1BBECCB8166C" name="Costa Rica" pageId="1" pageNumber="16">Costa Rica</collectingCountry>
. The
<typeStatus id="54AED1FCFFDB6B453F591A46CDE8166C" box="[282,385,1301,1327]" pageId="1" pageNumber="16" type="holotype">holotype</typeStatus>
and 14
<typeStatus id="54AED1FCFFDB6B453FAE1A46CE34166C" box="[493,605,1301,1327]" pageId="1" pageNumber="16" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D13A4D7FFDB6B453C301A46CE91166C" box="[627,760,1301,1327]" pageId="1" pageNumber="16" type="female">10 females</specimenCount>
+
<specimenCount id="9D13A4D7FFDB6B453D621A46CFEB166C" box="[801,898,1301,1327]" pageId="1" pageNumber="16" type="male">4 males</specimenCount>
) were mounted in Hoyer’s medium and 10
<typeStatus id="54AED1FCFFDB6B453ED41A6ECD6E1614" box="[151,263,1341,1367]" pageId="1" pageNumber="16" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D13A4D7FFDB6B453F5B1A6ECDE01614" box="[280,393,1341,1367]" pageId="1" pageNumber="16" type="female">8 females</specimenCount>
+
<specimenCount id="9D13A4D7FFDB6B453FEB1A6ECE6D1614" box="[424,516,1341,1367]" pageId="1" pageNumber="16" type="male">2 males</specimenCount>
) were prepared for SEM.
</paragraph>
</subSubSection>
<subSubSection id="C30F3CD5FFDB6B473E851A36CFAC149E" lastPageId="3" lastPageNumber="18" pageId="1" pageNumber="16" type="description">
<paragraph id="8BAA6F5EFFDB6B453E851A36C8C416B4" blockId="1.[151,1437,1261,2007]" pageId="1" pageNumber="16">
<emphasis id="B961B34CFFDB6B453E851A36CE79163C" bold="true" box="[198,528,1381,1407]" pageId="1" pageNumber="16">Description: Adult female</emphasis>
(measurements of the
<typeStatus id="54AED1FCFFDB6B453D661A36CFFB163C" box="[805,914,1381,1407]" pageId="1" pageNumber="16" type="holotype">holotype</typeStatus>
): Body length 185.0 (
<figureCitation id="132E73DBFFDB6B453AD91A36C967163C" box="[1178,1294,1381,1407]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1925,1949]" captionTargetBox="[223,1363,277,1899]" captionTargetId="figure@2.[223,1363,277,1899]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Echiniscus ganczareki sp. nov. Habitus: 1 — dorsal view (holotype, PCM); 2 — dorso-lateral view (paratype, PCM); 3 — dorsal view (paratype, SEM); 4 — lateral view (paratype, SEM)." httpUri="https://zenodo.org/record/176680/files/figure.png" pageId="1" pageNumber="16">Figs. 1–4</figureCitation>
). Body red. Red eyes visible in living specimens only. Internal cirrus 11.6 long, external cirrus 15.2 long; internal/external cirrus ratio = 0.76. Cephalic papilla 7.3 long (
<figureCitation id="132E73DBFFDB6B453CED1AE6CF6F168C" box="[686,774,1461,1487]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,269,1576,1600]" captionTargetBox="[164,1423,194,1548]" captionTargetId="figure@3.[164,1424,194,1550]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Echiniscus ganczareki sp. nov. 5 – 8 — dorsal plate cuticle; 5 — terminal plate with pores; 6 — the same fragment as shown on Fig. 5, but showing pillars that appear when focusing down through the cuticle (5 – 6 – holotype, PCM); 7 — pores in the scapular plate (paratype, SEM); 8 — structure of the dorsal plate cuticle (see also Fig. 25) (paratype, SEM); 9 – 10 — ventral head cuticle with pores (9 — holotype, PCM, 10 — paratype, SEM)." httpUri="https://zenodo.org/record/176681/files/figure.png" pageId="1" pageNumber="16">Fig. 10</figureCitation>
). Apart from head cirri and cirrus
<emphasis id="B961B34CFFDB6B453ADB1AE6C8C3168D" box="[1176,1194,1461,1486]" italics="true" pageId="1" pageNumber="16">A</emphasis>
no other appendices present. Cirrus
<emphasis id="B961B34CFFDB6B453F081A8ECD3416B5" box="[331,349,1501,1526]" italics="true" pageId="1" pageNumber="16">A</emphasis>
31.6 long, ending in a point. Cirrus
<emphasis id="B961B34CFFDB6B453D451A8ECF7116B5" box="[774,792,1501,1526]" italics="true" pageId="1" pageNumber="16">A</emphasis>
/body ratio = 0.17. Clava 5.6 long.
</paragraph>
<paragraph id="8BAA6F5EFFDB6B453E841956C9F5141C" blockId="1.[151,1437,1261,2007]" pageId="1" pageNumber="16">
Dorsal (plate) cuticle with true pores and with true, but very fine granulation (
<figureCitation id="132E73DBFFDB6B453A1E1956C8B6155C" box="[1117,1247,1541,1567]" captionStart-0="FIGURES 5 – 10" captionStart-1="FIGURES 11 – 14" captionStart-2="FIGURES 15 – 20" captionStartId-0="3.[151,269,1576,1600]" captionStartId-1="4.[151,269,1040,1064]" captionStartId-2="6.[151,269,1616,1640]" captionTargetBox-0="[164,1423,194,1548]" captionTargetBox-1="[166,1428,167,1015]" captionTargetBox-2="[168,1426,167,1602]" captionTargetId-0="figure@3.[164,1424,194,1550]" captionTargetId-1="figure@4.[166,1429,167,1016]" captionTargetId-2="figure@6.[168,1427,167,1603]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="FIGURES 5 – 10. Echiniscus ganczareki sp. nov. 5 – 8 — dorsal plate cuticle; 5 — terminal plate with pores; 6 — the same fragment as shown on Fig. 5, but showing pillars that appear when focusing down through the cuticle (5 – 6 – holotype, PCM); 7 — pores in the scapular plate (paratype, SEM); 8 — structure of the dorsal plate cuticle (see also Fig. 25) (paratype, SEM); 9 – 10 — ventral head cuticle with pores (9 — holotype, PCM, 10 — paratype, SEM)." captionText-1="FIGURES 11 – 14. Echiniscus ganczareki sp. nov. 11 – 12 — second median plate, note that the anterior portion of the plate is devoid of pores and is covered with fine granulation (11 — paratype, PCM, 12 — paratype, SEM); 13 – 14 — third median plate (13 — paratype, PCM, 14 — paratype, SEM)." captionText-2="FIGURES 15 – 20. Echiniscus ganczareki sp. nov. Pores on legs: 15 – 16 — legs I with the spine (15 — holotype, PCM, 16 — paratype, SEM); 17 – 18 — legs II (17 — holotype, PCM, 18 — paratype, SEM); 19 – 20 — legs IV with the dentate collar and papilla (19 — holotype, PCM, 20 — paratype, SEM)." httpUri-0="https://zenodo.org/record/176681/files/figure.png" httpUri-1="https://zenodo.org/record/176682/files/figure.png" httpUri-2="https://zenodo.org/record/176683/files/figure.png" pageId="1" pageNumber="16">Figs. 5–20</figureCitation>
). Granules (true granulation, on cuticle surface) present between scapular and first paired plates, on stripes in anterior portions of paired plates and in anterior part of second median plate (
<figureCitation id="132E73DBFFDB6B453D351906C86E152C" box="[886,1031,1621,1647]" captionStart="FIGURES 11 – 14" captionStartId="4.[151,269,1040,1064]" captionTargetBox="[166,1428,167,1015]" captionTargetId="figure@4.[166,1429,167,1016]" captionTargetPageId="4" captionText="FIGURES 11 – 14. Echiniscus ganczareki sp. nov. 11 – 12 — second median plate, note that the anterior portion of the plate is devoid of pores and is covered with fine granulation (11 — paratype, PCM, 12 — paratype, SEM); 13 – 14 — third median plate (13 — paratype, PCM, 14 — paratype, SEM)." httpUri="https://zenodo.org/record/176682/files/figure.png" pageId="1" pageNumber="16">Figs. 11–12</figureCitation>
). Granules very small (
<quantity id="4CEDC2BBFFDB6B453B611906C9F5152C" box="[1314,1436,1621,1647]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="1" pageNumber="16" unit="in" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 in</quantity>
diameter). Cuticle appearance in SEM: Pores small (
<quantity id="4CEDC2BBFFDB6B453D57192ECFE615D4" box="[788,911,1661,1687]" metricMagnitude="-2" metricUnit="m" metricValue="1.1429999999999998" metricValueMax="1.524" metricValueMin="0.762" pageId="1" pageNumber="16" unit="in" value="0.44999999999999996" valueMax="0.6" valueMin="0.3">0.3–0.6 in</quantity>
diameter) (
<figureCitation id="132E73DBFFDB6B453A5E192EC81A15D4" box="[1053,1139,1661,1687]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,269,1576,1600]" captionTargetBox="[164,1423,194,1548]" captionTargetId="figure@3.[164,1424,194,1550]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Echiniscus ganczareki sp. nov. 5 – 8 — dorsal plate cuticle; 5 — terminal plate with pores; 6 — the same fragment as shown on Fig. 5, but showing pillars that appear when focusing down through the cuticle (5 – 6 – holotype, PCM); 7 — pores in the scapular plate (paratype, SEM); 8 — structure of the dorsal plate cuticle (see also Fig. 25) (paratype, SEM); 9 – 10 — ventral head cuticle with pores (9 — holotype, PCM, 10 — paratype, SEM)." httpUri="https://zenodo.org/record/176681/files/figure.png" pageId="1" pageNumber="16">Figs. 7</figureCitation>
,
<figureCitation id="132E73DBFFDB6B453AC0192EC8BA15D4" box="[1155,1235,1661,1687]" captionStart="FIGURES 11 – 14" captionStartId="4.[151,269,1040,1064]" captionTargetBox="[166,1428,167,1015]" captionTargetId="figure@4.[166,1429,167,1016]" captionTargetPageId="4" captionText="FIGURES 11 – 14. Echiniscus ganczareki sp. nov. 11 – 12 — second median plate, note that the anterior portion of the plate is devoid of pores and is covered with fine granulation (11 — paratype, PCM, 12 — paratype, SEM); 13 – 14 — third median plate (13 — paratype, PCM, 14 — paratype, SEM)." httpUri="https://zenodo.org/record/176682/files/figure.png" pageId="1" pageNumber="16">12, 14</figureCitation>
). ‘Sponge layer’ very well developed, small pillars present (
<figureCitation id="132E73DBFFDB6B453CD719F6CE8E15FC" box="[660,743,1701,1727]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,269,1576,1600]" captionTargetBox="[164,1423,194,1548]" captionTargetId="figure@3.[164,1424,194,1550]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Echiniscus ganczareki sp. nov. 5 – 8 — dorsal plate cuticle; 5 — terminal plate with pores; 6 — the same fragment as shown on Fig. 5, but showing pillars that appear when focusing down through the cuticle (5 – 6 – holotype, PCM); 7 — pores in the scapular plate (paratype, SEM); 8 — structure of the dorsal plate cuticle (see also Fig. 25) (paratype, SEM); 9 – 10 — ventral head cuticle with pores (9 — holotype, PCM, 10 — paratype, SEM)." httpUri="https://zenodo.org/record/176681/files/figure.png" pageId="1" pageNumber="16">Figs. 8</figureCitation>
,
<figureCitation id="132E73DBFFDB6B453CB619F6CF7F15FC" box="[757,790,1701,1727]" captionStart="FIGURES 23 – 26" captionStartId="9.[151,269,1899,1923]" captionTargetBox="[209,1383,1259,1863]" captionTargetId="figure@9.[199,1391,1252,1875]" captionTargetPageId="9" captionText="FIGURES 23 – 26. Structure of the dorsal plate cuticle in four species of the bigranulatus group (semi-schematic sections based on SEM observations). 23 — E. bigranulatus; 24 — E. ollantaytamboensis; 25 — E. ganczareki sp. nov. (see also Fig. 8); 26 — E. madonnae. Abbreviations: bc— body cavity, cs— cuticle surface, pi— pillar, po— pore, pp— pseudopore, sl— ‘ sponge’ layer, up— undeveloped pillar. Not scaled (23 – 24 and 26 from Michalczyk &amp; Kaczmarek 2006 c)." httpUri="https://zenodo.org/record/176685/files/figure.png" pageId="1" pageNumber="16">25</figureCitation>
). Cuticle visible in PCM: Pores visible as bright/white dots; when focusing down through cuticle, cuticular pillars appear as faint dark dots (
<figureCitation id="132E73DBFFDB6B453ADF199EC97815A4" box="[1180,1297,1741,1767]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,269,1576,1600]" captionTargetBox="[164,1423,194,1548]" captionTargetId="figure@3.[164,1424,194,1550]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Echiniscus ganczareki sp. nov. 5 – 8 — dorsal plate cuticle; 5 — terminal plate with pores; 6 — the same fragment as shown on Fig. 5, but showing pillars that appear when focusing down through the cuticle (5 – 6 – holotype, PCM); 7 — pores in the scapular plate (paratype, SEM); 8 — structure of the dorsal plate cuticle (see also Fig. 25) (paratype, SEM); 9 – 10 — ventral head cuticle with pores (9 — holotype, PCM, 10 — paratype, SEM)." httpUri="https://zenodo.org/record/176681/files/figure.png" pageId="1" pageNumber="16">Figs. 5–6</figureCitation>
). Pores and cuticular pillars focusable. Ventral cuticle with very fine, dense and regular granulation. This granulation is caused by dense cuticular pillars. Several pores organised in 2 small irregular patches (left and right) present on ventral cuticle on head level (pores slightly smaller and shallower than those on dorsal plates) (
<figureCitation id="132E73DBFFDB6B453B4C1816C9E6141C" box="[1295,1423,1861,1887]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,269,1576,1600]" captionTargetBox="[164,1423,194,1548]" captionTargetId="figure@3.[164,1424,194,1550]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Echiniscus ganczareki sp. nov. 5 – 8 — dorsal plate cuticle; 5 — terminal plate with pores; 6 — the same fragment as shown on Fig. 5, but showing pillars that appear when focusing down through the cuticle (5 – 6 – holotype, PCM); 7 — pores in the scapular plate (paratype, SEM); 8 — structure of the dorsal plate cuticle (see also Fig. 25) (paratype, SEM); 9 – 10 — ventral head cuticle with pores (9 — holotype, PCM, 10 — paratype, SEM)." httpUri="https://zenodo.org/record/176681/files/figure.png" pageId="1" pageNumber="16">Figs. 9–10</figureCitation>
).
</paragraph>
<paragraph id="8BAA6F5EFFDB6B463E84183EC9591399" blockId="1.[151,1437,1261,2007]" lastBlockId="2.[151,1436,152,218]" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16">
Dorsal plates well developed. Head and scapular plates not faceted. In LM lateral portions of the scapular plate seem to be detached from the dorsal plate and form small shoulder plates (one on each side of the body) divided from the scapular plate by a thin bright stripe. This false division is caused by a bend of the plate where the cuticle is thinner; in SEM this pseudo division is not visible (
<figureCitation id="132E73DBFFD86B463D951FCBC82E13F1" box="[982,1095,152,178]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1925,1949]" captionTargetBox="[223,1363,277,1899]" captionTargetId="figure@2.[223,1363,277,1899]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Echiniscus ganczareki sp. nov. Habitus: 1 — dorsal view (holotype, PCM); 2 — dorso-lateral view (paratype, PCM); 3 — dorsal view (paratype, SEM); 4 — lateral view (paratype, SEM)." httpUri="https://zenodo.org/record/176680/files/figure.png" pageId="2" pageNumber="17">Figs. 2, 4</figureCitation>
). Third median plate present, but relatively narrow, developed anteriorly (
<figureCitation id="132E73DBFFD86B463CD81F93CF421399" box="[667,811,192,218]" captionStart="FIGURES 11 – 14" captionStartId="4.[151,269,1040,1064]" captionTargetBox="[166,1428,167,1015]" captionTargetId="figure@4.[166,1429,167,1016]" captionTargetPageId="4" captionText="FIGURES 11 – 14. Echiniscus ganczareki sp. nov. 11 – 12 — second median plate, note that the anterior portion of the plate is devoid of pores and is covered with fine granulation (11 — paratype, PCM, 12 — paratype, SEM); 13 – 14 — third median plate (13 — paratype, PCM, 14 — paratype, SEM)." httpUri="https://zenodo.org/record/176682/files/figure.png" pageId="2" pageNumber="17">Figs. 13–14</figureCitation>
). Terminal plate faceted. Notches 18.2 long.
</paragraph>
<caption id="DF6A3FD6FFD86B463ED418D6C84E14FC" httpUri="https://zenodo.org/record/176680/files/figure.png" pageId="2" pageNumber="17" targetBox="[223,1363,277,1899]" targetPageId="2">
<paragraph id="8BAA6F5EFFD86B463ED418D6C84E14FC" blockId="2.[151,1436,1925,1984]" pageId="2" pageNumber="17">
<emphasis id="B961B34CFFD86B463ED418D6CD2014DE" bold="true" box="[151,329,1925,1949]" pageId="2" pageNumber="17">FIGURES 1–4.</emphasis>
<taxonomicName id="4C1514DDFFD86B463F1518D5CE2314DE" box="[342,586,1926,1949]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="2" pageNumber="17" phylum="Tardigrada" rank="species" species="ganczareki" status="sp. nov.">
<emphasis id="B961B34CFFD86B463F1518D5CE2314DE" box="[342,586,1926,1949]" italics="true" pageId="2" pageNumber="17">Echiniscus ganczareki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2520E37FFD86B463C1418D6CEDA14DE" box="[599,691,1925,1949]" pageId="2" pageNumber="17" rank="species">
<emphasis id="B961B34CFFD86B463C1418D6CEDA14DE" bold="true" box="[599,691,1925,1949]" pageId="2" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
Habitus:
<emphasis id="B961B34CFFD86B463D6518D6CF2714DE" bold="true" box="[806,846,1925,1949]" pageId="2" pageNumber="17">1—</emphasis>
dorsal view (holotype, PCM);
<emphasis id="B961B34CFFD86B463AEF18D6C8BD14DE" bold="true" box="[1196,1236,1925,1949]" pageId="2" pageNumber="17">2—</emphasis>
dorso-lateral view (paratype, PCM);
<emphasis id="B961B34CFFD86B463F1B18FBCDE91483" bold="true" box="[344,384,1960,1984]" pageId="2" pageNumber="17">3—</emphasis>
dorsal view (paratype, SEM);
<emphasis id="B961B34CFFD86B463C8118FBCE801483" bold="true" box="[706,745,1960,1984]" pageId="2" pageNumber="17">4—</emphasis>
lateral view (paratype, SEM).
</paragraph>
</caption>
<caption id="DF6A3FD6FFD96B473ED4197BC8D215DC" httpUri="https://zenodo.org/record/176681/files/figure.png" pageId="3" pageNumber="18" targetBox="[164,1423,194,1548]" targetPageId="3">
<paragraph id="8BAA6F5EFFD96B473ED4197BC8D215DC" blockId="3.[151,1436,1576,1696]" pageId="3" pageNumber="18">
<emphasis id="B961B34CFFD96B473ED4197BCD381503" bold="true" box="[151,337,1576,1600]" pageId="3" pageNumber="18">FIGURES 5–10.</emphasis>
<taxonomicName id="4C1514DDFFD96B473F1B197ACE2E1503" box="[344,583,1577,1600]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="18" phylum="Tardigrada" rank="species" species="ganczareki" status="sp. nov.">
<emphasis id="B961B34CFFD96B473F1B197ACE2E1503" box="[344,583,1577,1600]" italics="true" pageId="3" pageNumber="18">Echiniscus ganczareki</emphasis>
</taxonomicName>
<emphasis id="B961B34CFFD96B473C0C197BCE9E1503" bold="true" box="[591,759,1576,1600]" pageId="3" pageNumber="18">
<taxonomicNameLabel id="A2520E37FFD96B473C0C197BCECF1503" box="[591,678,1576,1600]" pageId="3" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
5–8—
</emphasis>
dorsal plate cuticle;
<emphasis id="B961B34CFFD96B473D92197BCF931503" bold="true" box="[977,1018,1576,1600]" pageId="3" pageNumber="18">5—</emphasis>
terminal plate with pores;
<emphasis id="B961B34CFFD96B473B56197BC9541503" bold="true" box="[1301,1341,1576,1600]" pageId="3" pageNumber="18">6—</emphasis>
the same fragment as shown on Fig.
<emphasis id="B961B34CFFD96B473FFC191BCDA51523" bold="true" box="[447,460,1608,1632]" pageId="3" pageNumber="18">5</emphasis>
, but showing pillars that appear when focusing down through the cuticle (
<emphasis id="B961B34CFFD96B473AB0191BC9721523" bold="true" box="[1267,1307,1608,1632]" pageId="3" pageNumber="18">5–6</emphasis>
– holotype, PCM);
<emphasis id="B961B34CFFD96B473EA8193BCD7A15C3" bold="true" box="[235,275,1640,1664]" pageId="3" pageNumber="18">7—</emphasis>
pores in the scapular plate (paratype, SEM);
<emphasis id="B961B34CFFD96B473D56193BCF5415C3" bold="true" box="[789,829,1640,1664]" pageId="3" pageNumber="18">8—</emphasis>
structure of the dorsal plate cuticle (see also Fig.
<emphasis id="B961B34CFFD96B473B3B193BC9FA15C3" bold="true" box="[1400,1427,1640,1664]" pageId="3" pageNumber="18">25</emphasis>
) (paratype, SEM);
<emphasis id="B961B34CFFD96B473F1519DBCDCF15E3" bold="true" box="[342,422,1672,1696]" pageId="3" pageNumber="18">9–10—</emphasis>
ventral head cuticle with pores (
<emphasis id="B961B34CFFD96B473CBE19DBCF4C15E3" bold="true" box="[765,805,1672,1696]" pageId="3" pageNumber="18">9—</emphasis>
holotype, PCM,
<emphasis id="B961B34CFFD96B473D9619DBC86215E3" bold="true" box="[981,1035,1672,1696]" pageId="3" pageNumber="18">10—</emphasis>
paratype, SEM).
</paragraph>
</caption>
<paragraph id="8BAA6F5EFFD96B473E841983CF2B1421" blockId="3.[151,1436,1744,2013]" pageId="3" pageNumber="18">
Spine on legs I in shape of narrow triangle, 2.2 long (
<figureCitation id="132E73DBFFD96B473D761983CFAC15A9" box="[821,965,1744,1770]" captionStart="FIGURES 15 – 20" captionStartId="6.[151,269,1616,1640]" captionTargetBox="[168,1426,167,1602]" captionTargetId="figure@6.[168,1427,167,1603]" captionTargetPageId="6" captionText="FIGURES 15 – 20. Echiniscus ganczareki sp. nov. Pores on legs: 15 – 16 — legs I with the spine (15 — holotype, PCM, 16 — paratype, SEM); 17 – 18 — legs II (17 — holotype, PCM, 18 — paratype, SEM); 19 – 20 — legs IV with the dentate collar and papilla (19 — holotype, PCM, 20 — paratype, SEM)." httpUri="https://zenodo.org/record/176683/files/figure.png" pageId="3" pageNumber="18">Figs. 15–16</figureCitation>
). Papilla on legs IV finger-like, 4.0 long (
<figureCitation id="132E73DBFFD96B473EDC19ABCC911451" box="[159,248,1784,1810]" captionStart="FIGURES 15 – 20" captionStartId="6.[151,269,1616,1640]" captionTargetBox="[168,1426,167,1602]" captionTargetId="figure@6.[168,1427,167,1603]" captionTargetPageId="6" captionText="FIGURES 15 – 20. Echiniscus ganczareki sp. nov. Pores on legs: 15 – 16 — legs I with the spine (15 — holotype, PCM, 16 — paratype, SEM); 17 – 18 — legs II (17 — holotype, PCM, 18 — paratype, SEM); 19 – 20 — legs IV with the dentate collar and papilla (19 — holotype, PCM, 20 — paratype, SEM)." httpUri="https://zenodo.org/record/176683/files/figure.png" pageId="3" pageNumber="18">Fig. 20</figureCitation>
). Outer cuticle of all legs with sculpture same as on dorsal plates, but with slightly smaller and shallower pores (
<quantity id="4CEDC2BBFFD96B473F751873CDD81479" box="[310,433,1824,1850]" metricMagnitude="-3" metricUnit="m" metricValue="8.889999999999999" metricValueMax="15.24" metricValueMin="2.54" pageId="3" pageNumber="18" unit="in" value="0.35" valueMax="0.6" valueMin="0.1">0.1–0.6 in</quantity>
diameter) (
<figureCitation id="132E73DBFFD96B473C7D1873CEB91479" box="[574,720,1824,1850]" captionStart="FIGURES 15 – 20" captionStartId="6.[151,269,1616,1640]" captionTargetBox="[168,1426,167,1602]" captionTargetId="figure@6.[168,1427,167,1603]" captionTargetPageId="6" captionText="FIGURES 15 – 20. Echiniscus ganczareki sp. nov. Pores on legs: 15 – 16 — legs I with the spine (15 — holotype, PCM, 16 — paratype, SEM); 17 – 18 — legs II (17 — holotype, PCM, 18 — paratype, SEM); 19 – 20 — legs IV with the dentate collar and papilla (19 — holotype, PCM, 20 — paratype, SEM)." httpUri="https://zenodo.org/record/176683/files/figure.png" pageId="3" pageNumber="18">Figs. 15–20</figureCitation>
, see also
<figureCitation id="132E73DBFFD96B473D0B1873CFFA1479" box="[840,915,1824,1850]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1925,1949]" captionTargetBox="[223,1363,277,1899]" captionTargetId="figure@2.[223,1363,277,1899]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Echiniscus ganczareki sp. nov. Habitus: 1 — dorsal view (holotype, PCM); 2 — dorso-lateral view (paratype, PCM); 3 — dorsal view (paratype, SEM); 4 — lateral view (paratype, SEM)." httpUri="https://zenodo.org/record/176680/files/figure.png" pageId="3" pageNumber="18">Fig. 4</figureCitation>
). Pores on leg often distributed irregularly. Dentate collar with 8 sharp, triangular teeth (
<figureCitation id="132E73DBFFD96B473CE6181BCF5C1421" box="[677,821,1864,1890]" captionStart="FIGURES 15 – 20" captionStartId="6.[151,269,1616,1640]" captionTargetBox="[168,1426,167,1602]" captionTargetId="figure@6.[168,1427,167,1603]" captionTargetPageId="6" captionText="FIGURES 15 – 20. Echiniscus ganczareki sp. nov. Pores on legs: 15 – 16 — legs I with the spine (15 — holotype, PCM, 16 — paratype, SEM); 17 – 18 — legs II (17 — holotype, PCM, 18 — paratype, SEM); 19 – 20 — legs IV with the dentate collar and papilla (19 — holotype, PCM, 20 — paratype, SEM)." httpUri="https://zenodo.org/record/176683/files/figure.png" pageId="3" pageNumber="18">Figs. 19–20</figureCitation>
).
</paragraph>
<paragraph id="8BAA6F5EFFD96B473E841823CFAC149E" blockId="3.[151,1436,1744,2013]" pageId="3" pageNumber="18">
Claws of legs IV 12.6 long. External claws of all legs without spurs, internal claws of all legs with spur, directed downwards, growing out of claw at
<emphasis id="B961B34CFFD96B473CEE18C8CEB814F7" box="[685,721,1947,1972]" italics="true" pageId="3" pageNumber="18">ca.</emphasis>
45o (
<figureCitation id="132E73DBFFD96B473D5318C8CFCB14F6" box="[784,930,1947,1973]" captionStart="FIGURES 21 – 22" captionStartId="7.[151,269,1708,1732]" captionTargetBox="[154,1433,1192,1684]" captionTargetId="figure@7.[154,1434,1192,1684]" captionTargetPageId="7" captionText="FIGURES 21 – 22. Echiniscus ganczareki sp. nov. Claws IV (21 — holotype, PCM, 22 — paratype, SEM)." httpUri="https://zenodo.org/record/176684/files/figure.png" pageId="3" pageNumber="18">Figs. 21–22</figureCitation>
). Spur on claws of legs IV relatively large (3.2 long; spur/claw ratio = 0.25). Claws of legs I–III slightly shorter.
</paragraph>
</subSubSection>
<caption id="DF6A3FD6FFDE6B403ED41B43CE90172F" httpUri="https://zenodo.org/record/176682/files/figure.png" pageId="4" pageNumber="19" targetBox="[166,1428,167,1015]" targetPageId="4">
<paragraph id="8BAA6F5EFFDE6B403ED41B43CE90172F" blockId="4.[151,1436,1040,1133]" pageId="4" pageNumber="19">
<emphasis id="B961B34CFFDE6B403ED41B43CD36176B" bold="true" box="[151,351,1040,1064]" pageId="4" pageNumber="19">FIGURES 11–14.</emphasis>
<taxonomicName id="4C1514DDFFDE6B403F2B1B42CE31176B" box="[360,600,1041,1064]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="19" phylum="Tardigrada" rank="species" species="ganczareki" status="sp. nov.">
<emphasis id="B961B34CFFDE6B403F2B1B42CE31176B" box="[360,600,1041,1064]" italics="true" pageId="4" pageNumber="19">Echiniscus ganczareki</emphasis>
</taxonomicName>
<emphasis id="B961B34CFFDE6B403C211B43CF41176B" bold="true" box="[610,808,1040,1064]" pageId="4" pageNumber="19">
<taxonomicNameLabel id="A2520E37FFDE6B403C211B43CED0176B" box="[610,697,1040,1064]" pageId="4" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
11–12—
</emphasis>
second median plate, note that the anterior portion of the plate is devoid of pores and is covered with fine granulation (
<emphasis id="B961B34CFFDE6B403D741B60CF001708" bold="true" box="[823,873,1075,1099]" pageId="4" pageNumber="19">11—</emphasis>
paratype, PCM,
<emphasis id="B961B34CFFDE6B403A5E1B60C83B1708" bold="true" box="[1053,1106,1075,1099]" pageId="4" pageNumber="19">12—</emphasis>
paratype, SEM);
<emphasis id="B961B34CFFDE6B403B4F1B60C9001708" bold="true" box="[1292,1385,1075,1099]" pageId="4" pageNumber="19">13–14—</emphasis>
third median plate (
<emphasis id="B961B34CFFDE6B403F6C1B06CD0D172E" bold="true" box="[303,356,1109,1133]" pageId="4" pageNumber="19">13—</emphasis>
paratype, PCM,
<emphasis id="B961B34CFFDE6B403C501B06CE20172E" bold="true" box="[531,585,1109,1133]" pageId="4" pageNumber="19">14—</emphasis>
paratype, SEM).
</paragraph>
</caption>
<subSubSection id="C30F3CD5FFDE6B403E851BF3CFCE15D9" pageId="4" pageNumber="19" type="discussion">
<paragraph id="8BAA6F5EFFDE6B403E851BF3C8F416C1" blockId="4.[151,1436,1184,2013]" pageId="4" pageNumber="19">
<emphasis id="B961B34CFFDE6B403E851BF3CD2B17F9" bold="true" box="[198,322,1184,1210]" pageId="4" pageNumber="19">Remarks:</emphasis>
Extreme care is needed when determining the appearance of the anterior portion of the second median plate. When an animal is even slightly retracted, the porous posterior portions of the first paired plates (
<figureCitation id="132E73DBFFDE6B403EDC1BA3CC9C1649" box="[159,245,1264,1290]" captionStart="FIGURES 11 – 14" captionStartId="4.[151,269,1040,1064]" captionTargetBox="[166,1428,167,1015]" captionTargetId="figure@4.[166,1429,167,1016]" captionTargetPageId="4" captionText="FIGURES 11 – 14. Echiniscus ganczareki sp. nov. 11 – 12 — second median plate, note that the anterior portion of the plate is devoid of pores and is covered with fine granulation (11 — paratype, PCM, 12 — paratype, SEM); 13 – 14 — third median plate (13 — paratype, PCM, 14 — paratype, SEM)." httpUri="https://zenodo.org/record/176682/files/figure.png" pageId="4" pageNumber="19">Fig. 12</figureCitation>
) may cover the anterior portion of the first median plate and may lead to an incorrect conclusion that the median plate is porous in its anterior part. The scapular plate can sometimes be slightly faceted in bigger animals. Because pores on legs and especially pores on the ventral head cuticle are less visible than those on the dorsal plates, they can be overlooked if an animal is not examined carefully enough.
</paragraph>
<paragraph id="8BAA6F5EFFDE6B403E851AC3C8B81691" blockId="4.[151,1436,1184,2013]" pageId="4" pageNumber="19">
Results of simple statistical analysis of measurements of selected morphological structures for
<specimenCount id="9D13A4D7FFDE6B403B731AC3CCBC1691" pageId="4" pageNumber="19" type="generic">15 specimens</specimenCount>
(including the
<typeStatus id="54AED1FCFFDE6B403FC41AEBCD861691" box="[391,495,1464,1490]" pageId="4" pageNumber="19" type="holotype">holotype</typeStatus>
) are given in
<tableCitation id="C6975AE5FFDE6B403CD31AEBCE811691" box="[656,744,1464,1490]" captionStart="TABLE 1" captionStartId="5.[124,203,634,655]" captionTargetBox="[124,1878,723,1517]" captionTargetPageId="5" captionText="TABLE 1. Lengths [in µm] and length ratios of selected morphological structures of adults of Echiniscus ganczareki sp. nov. mounted in Hoyer’s medium (MIN and MAX refer to the smallest and the largest structure found among all measured specimens; SD – Standard Deviation, N – number of specimens / structures measured)." httpUri="http://table.plazi.org/id/DF6A3FD6FFDF6B413E3F1D29CA2111FA" pageId="4" pageNumber="19" tableUuid="DF6A3FD6FFDF6B413E3F1D29CA2111FA">Table 1</tableCitation>
. No juveniles, larvae or eggs were found.
</paragraph>
<paragraph id="8BAA6F5EFFDE6B403E851AB3CFCE15D9" blockId="4.[151,1436,1184,2013]" pageId="4" pageNumber="19">
Even though the number of measured males is low (
<specimenCount id="9D13A4D7FFDE6B403D691AB3CFA916B9" box="[810,960,1504,1530]" pageId="4" pageNumber="19" type="generic">4 specimens</specimenCount>
), and the statistics with such a low number are not conclusive, it is worth noting that males seem to be smaller than females (p=0.003,
<emphasis id="B961B34CFFDE6B403AB7195BC8951562" box="[1268,1276,1544,1569]" italics="true" pageId="4" pageNumber="19">t</emphasis>
= 3.606,
<emphasis id="B961B34CFFDE6B403B2E195BC9ED1562" box="[1389,1412,1544,1569]" italics="true" pageId="4" pageNumber="19">df</emphasis>
= 13), have a shorter cirrus internus (p=0.016,
<emphasis id="B961B34CFFDE6B403CEA1963CED8150A" box="[681,689,1584,1609]" italics="true" pageId="4" pageNumber="19">t</emphasis>
= 2.767,
<emphasis id="B961B34CFFDE6B403D671963CF52150A" box="[804,827,1584,1609]" italics="true" pageId="4" pageNumber="19">df</emphasis>
= 13), externus (p&lt;0.001,
<emphasis id="B961B34CFFDE6B403A391963C8EB150A" box="[1146,1154,1584,1609]" italics="true" pageId="4" pageNumber="19">t</emphasis>
= 4.936,
<emphasis id="B961B34CFFDE6B403AB61963C962150A" box="[1269,1291,1584,1609]" italics="true" pageId="4" pageNumber="19">df</emphasis>
= 11.8) and cirrus A (p=0.048,
<emphasis id="B961B34CFFDE6B403F3B190BCDE91532" box="[376,384,1624,1649]" italics="true" pageId="4" pageNumber="19">t</emphasis>
= 2.203,
<emphasis id="B961B34CFFDE6B403FB2190BCE611532" box="[497,520,1624,1649]" italics="true" pageId="4" pageNumber="19">df</emphasis>
= 12), also shorter claws (p=0.019,
<emphasis id="B961B34CFFDE6B403DF9190BCFAB1532" box="[954,962,1624,1649]" italics="true" pageId="4" pageNumber="19">t</emphasis>
= 3.038,
<emphasis id="B961B34CFFDE6B403A70190BC8201532" box="[1075,1097,1624,1649]" italics="true" pageId="4" pageNumber="19">df</emphasis>
= 7) and spurs (p=0.035,
<emphasis id="B961B34CFFDE6B403B3F190BC9ED1532" box="[1404,1412,1624,1649]" italics="true" pageId="4" pageNumber="19">t</emphasis>
= 2.706,
<emphasis id="B961B34CFFDE6B403EA419D3CC9715DA" box="[231,254,1664,1689]" italics="true" pageId="4" pageNumber="19">df</emphasis>
= 6). The remaining characters do not differ significantly.
</paragraph>
</subSubSection>
<subSubSection id="C30F3CD5FFDE6B403E8519FBCD8D14F6" pageId="4" pageNumber="19" type="materials_examined">
<paragraph id="8BAA6F5EFFDE6B403E8519FBCC811456" blockId="4.[151,1436,1184,2013]" pageId="4" pageNumber="19">
<emphasis id="B961B34CFFDE6B403E8519FBCD1D1581" bold="true" box="[198,372,1704,1730]" pageId="4" pageNumber="19">
<typeStatus id="54AED1FCFFDE6B403E8519FBCD6D1581" box="[198,260,1704,1730]" pageId="4" pageNumber="19">Type</typeStatus>
locality:
</emphasis>
Central
<collectingCountry id="F3022FCEFFDE6B403FA219FBCE201581" box="[481,585,1704,1730]" name="United States of America" pageId="4" pageNumber="19">America</collectingCountry>
,
<collectingCountry id="F3022FCEFFDE6B403C1A19FBCE891581" box="[601,736,1704,1730]" name="Costa Rica" pageId="4" pageNumber="19">Costa Rica</collectingCountry>
, Cartago Province, Volcán Irazú National Park, liverwort from tree trunk, near the top of volcano,
<emphasis id="B961B34CFFDE6B403C371980CEFE15AF" box="[628,663,1747,1772]" italics="true" pageId="4" pageNumber="19">ca.</emphasis>
<quantity id="4CEDC2BBFFDE6B403CDC1980CE9115AE" box="[671,760,1747,1773]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="19" unit="m" value="3400.0">3400 m</quantity>
asl,
<geoCoordinate id="EE210999FFDE6B403D6D1980CFEE15AE" box="[814,903,1744,1773]" direction="north" orientation="latitude" pageId="4" pageNumber="19" precision="925" value="9.983334">9o59' N</geoCoordinate>
,
<geoCoordinate id="EE210999FFDE6B403DD01980C86815AE" box="[915,1025,1744,1773]" direction="west" orientation="longitude" pageId="4" pageNumber="19" precision="925" value="-83.85">83o51' W</geoCoordinate>
;
<date id="FFAB499EFFDE6B403A4F1980C89F15AE" box="[1036,1270,1747,1773]" pageId="4" pageNumber="19" value="2002-12-17">17 December 2002</date>
; leg. Ł. Kaczmarek.
</paragraph>
<paragraph id="8BAA6F5EFFDE6B403E851870CFCB147E" blockId="4.[151,1436,1184,2013]" box="[198,930,1827,1853]" pageId="4" pageNumber="19">
<emphasis id="B961B34CFFDE6B403E851870CD97147E" bold="true" box="[198,510,1827,1853]" pageId="4" pageNumber="19">Geographic distribution:</emphasis>
Known only from the
<typeStatus id="54AED1FCFFDE6B403D491870CF54147E" box="[778,829,1827,1853]" pageId="4" pageNumber="19">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BAA6F5EFFDE6B403E851818CD8D14F6" blockId="4.[151,1436,1184,2013]" pageId="4" pageNumber="19">
<emphasis id="B961B34CFFDE6B403E851818CDC31426" bold="true" box="[198,426,1867,1893]" pageId="4" pageNumber="19">
<typeStatus id="54AED1FCFFDE6B403E851818CD6A1426" box="[198,259,1867,1893]" pageId="4" pageNumber="19">Type</typeStatus>
depositories:
</emphasis>
<typeStatus id="54AED1FCFFDE6B403FF21818CE771426" box="[433,542,1867,1893]" pageId="4" pageNumber="19" type="holotype">Holotype</typeStatus>
and 7
<typeStatus id="54AED1FCFFDE6B403C2E1818CEB41426" box="[621,733,1867,1893]" pageId="4" pageNumber="19" type="paratype">paratypes</typeStatus>
are preserved at the Natural Sciences Collection, Faculty of Biology, A. Mickiewicz University, Umultowska 89, 61-614 Poznań,
<collectingCountry id="F3022FCEFFDE6B403D8B1820C87414CE" box="[968,1053,1907,1933]" name="Poland" pageId="4" pageNumber="19">Poland</collectingCountry>
; 7
<typeStatus id="54AED1FCFFDE6B403A7D1820C8C714CE" box="[1086,1198,1907,1933]" pageId="4" pageNumber="19" type="paratype">paratypes</typeStatus>
are preserved in the collection of Ł. Michalczyk.
</paragraph>
</subSubSection>
<subSubSection id="C30F3CD5FFDE6B403E851890C9E2149E" box="[198,1419,1987,2013]" pageId="4" pageNumber="19" type="etymology">
<paragraph id="8BAA6F5EFFDE6B403E851890C9E2149E" blockId="4.[151,1436,1184,2013]" box="[198,1419,1987,2013]" pageId="4" pageNumber="19">
<emphasis id="B961B34CFFDE6B403E851890CD3E149E" bold="true" box="[198,343,1987,2013]" pageId="4" pageNumber="19">Etymology:</emphasis>
The new species is dedicated to the first author’s very dear friend Joanna ‘
<emphasis id="B961B34CFFDE6B403A861890C89D149F" box="[1221,1268,1987,2012]" italics="true" pageId="4" pageNumber="19">mzn</emphasis>
’ Ganczarek.
</paragraph>
</subSubSection>
</treatment>
</document>
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