treatments-xml/data/03/F1/08/03F10847FFD38F04FE9C492C3FB45037.xml

<document id="C9974456010BBF69EBB72A3412D48E9C" ID-DOI="10.5281/zenodo.273444" ID-GBIF-Dataset="12e36fec-83fa-4423-9cfc-027930025ecf" ID-ISSN="1175­5326" ID-Zenodo-Dep="273444" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459761510964" checkinUser="plazi" docAuthor="Santos, Franklin Noel Dos &amp; Absalão, Ricardo Silva" docDate="2006" docId="03F10847FFD38F04FE9C492C3FB45037" docLanguage="en" docName="zt01232p068.pdf" docOrigin="Zootaxa 1232" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Ketosia thomei Santos &amp; Absalão, 2006, new species" docType="treatment" docVersion="9" lastPageNumber="65" masterDocId="FFC8703FFFD08F02FF944F123B125571" masterDocTitle="A new genus of Barleeidae Gray, 1857 (Mollusca, Gastropoda) and the first occurrence of the genus Pseudodiala Ponder, 1967 off the Brazilian coast" masterLastPageNumber="68" masterPageNumber="59" pageNumber="62" updateTime="1698215278659" updateUser="plazi">
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<mods:title id="4E5919FEC7C6B81FF3B5B97290F9FFE7">A new genus of Barleeidae Gray, 1857 (Mollusca, Gastropoda) and the first occurrence of the genus Pseudodiala Ponder, 1967 off the Brazilian coast</mods:title>
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<mods:namePart id="C47B77276C40170ABED0CF9E5B91F267">Santos, Franklin Noel Dos</mods:namePart>
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<mods:namePart id="FDA4F42E0992664CF257764ACB4F8FE8">Absalão, Ricardo Silva</mods:namePart>
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<mods:date id="1BC3CE088ACC86EB245C590466857DB5">2006</mods:date>
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<mods:number id="FC05ADFF800B6D3A110EF95CC1C293CC">1232</mods:number>
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<treatment id="03F10847FFD38F04FE9C492C3FB45037" ID-DOI="http://doi.org/10.5281/zenodo.5690156" ID-GBIF-Taxon="119345028" ID-Zenodo-Dep="5690156" LSID="urn:lsid:plazi:treatment:03F10847FFD38F04FE9C492C3FB45037" httpUri="http://treatment.plazi.org/id/03F10847FFD38F04FE9C492C3FB45037" lastPageId="6" lastPageNumber="65" pageId="3" pageNumber="62">
<subSubSection id="C342EADAFFD38F01FE9C492C3A84530A" pageId="3" pageNumber="62" type="nomenclature">
<paragraph id="8BE7B951FFD38F01FE9C492C39415329" blockId="3.[264,595,1598,1659]" box="[264,595,1598,1624]" pageId="3" pageNumber="62">
<heading id="D0AF0E3DFFD38F01FE9C492C39415329" bold="true" box="[264,595,1598,1624]" fontSize="11" level="1" pageId="3" pageNumber="62" reason="1">
<emphasis id="B92C6543FFD38F01FE9C492C39415329" bold="true" box="[264,595,1598,1624]" pageId="3" pageNumber="62">
<taxonomicName id="4C58C2D2FFD38F01FE9C492C3AAE5329" ID-CoL="3R6RC" box="[264,444,1598,1624]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="62" phylum="Mollusca" rank="species" species="thomei" status="sp. nov.">
<emphasis id="B92C6543FFD38F01FE9C492C3AAE5329" bold="true" box="[264,444,1598,1624]" italics="true" pageId="3" pageNumber="62">Ketosia thomei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21FD838FFD38F01FE57492C39415329" box="[451,595,1598,1624]" pageId="3" pageNumber="62" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BE7B951FFD38F01FE9C49733A84530A" blockId="3.[264,595,1598,1659]" box="[264,406,1633,1659]" pageId="3" pageNumber="62">
(
<figureCitation id="1363A5D4FFD38F01FE8549733A9C530A" box="[273,398,1633,1659]" captionStart="FIGURES 6 – 11" captionStartId="4.[264,383,1765,1789]" captionTargetBox="[319,1281,830,1758]" captionTargetId="figure@4.[319,1281,829,1758]" captionTargetPageId="4" captionText="FIGURES 6 – 11. Ketosia thomei n. sp. ­ 6: holotype (MNRJ 10505), length 2.1 mm; 7: paratype (IBUFRJ 14696) length 1.8 mm; 8: detail of the teleoconch of holotype; 9: protoconch of paratype (IBUFRJ 14696); 10 – 11: detail of sculpture of the protoconch of paratype (IBUFRJ 14696). Scale bar: 6: 500 m; 7: 200 m; 8: 10 m; 9: 100 m; 10: 20 m; 11: 10 m." httpUri="https://zenodo.org/record/273446/files/figure.png" pageId="3" pageNumber="62">Figs. 6–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C342EADAFFD38F06FE9C49BE3AE85667" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62" type="description">
<paragraph id="8BE7B951FFD38F01FE9C49BE3A8053B4" blockId="3.[264,1325,1708,1894]" box="[264,402,1708,1733]" pageId="3" pageNumber="62">
<emphasis id="B92C6543FFD38F01FE9C49BE3A8053B4" box="[264,402,1708,1733]" italics="true" pageId="3" pageNumber="62">Description</emphasis>
</paragraph>
<paragraph id="8BE7B951FFD38F01FEAC49C63A915217" blockId="3.[264,1325,1708,1894]" pageId="3" pageNumber="62">Shell conical­pupoid, whorls convex, cream to orange, opaque. Protoconch paucispiral with deep pits. Teleoconch with 3.5 whorls, smooth; with deep pits. Suture impressed. Base rounded. Aperture oval to subcircular. Lip thin, projected backwards. Chink­like umbilicus.</paragraph>
<paragraph id="8BE7B951FFD48F06FE9C4E0E3A805444" blockId="4.[264,939,284,350]" box="[264,402,284,309]" pageId="4" pageNumber="63">
<emphasis id="B92C6543FFD48F06FE9C4E0E3A805444" box="[264,402,284,309]" italics="true" pageId="4" pageNumber="63">Dimensions</emphasis>
</paragraph>
<paragraph id="8BE7B951FFD48F06FEAC4E5638B9542F" blockId="4.[264,939,284,350]" box="[312,939,324,350]" pageId="4" pageNumber="63">
<typeStatus id="54E307F3FFD48F06FEAC4E563AB7542F" box="[312,421,324,350]" pageId="4" pageNumber="63" type="holotype">Holotype</typeStatus>
with 3.5 teleoconch whorls; length 2.1 mm.
</paragraph>
<paragraph id="8BE7B951FFD48F06FE9C4E863AB954DC" blockId="4.[264,1323,404,790]" box="[264,427,404,429]" pageId="4" pageNumber="63">
<emphasis id="B92C6543FFD48F06FE9C4E863AB954DC" box="[264,427,404,429]" italics="true" pageId="4" pageNumber="63">Type­material</emphasis>
</paragraph>
<paragraph id="8BE7B951FFD48F06FEAC4EAE3AE85667" blockId="4.[264,1323,404,790]" pageId="4" pageNumber="63">
<typeStatus id="54E307F3FFD48F06FEAC4EAE3ABB54A7" box="[312,425,444,470]" pageId="4" pageNumber="63" type="holotype">Holotype</typeStatus>
: MNRJ 10505. 
<typeStatus id="54E307F3FFD48F06FDE44EAE39CB54A7" box="[624,729,444,470]" pageId="4" pageNumber="63" type="paratype">Paratype</typeStatus>
, 1: IBUFRJ 14696 GEOMAR XVI #16 Am.18, 22º24’4 S 40º55’0 W, 
<date id="FFE69F91FFD48F06FD814EF639B4548F" box="[533,678,484,510]" pageId="4" pageNumber="63" value="1980-08-20">
20.viii. 
<quantity id="4CA014B4FFD48F06FDF34EF639B4548F" box="[615,678,484,510]" metricMagnitude="3" metricUnit="m" metricValue="1.98063" pageId="4" pageNumber="63" unit="m" value="1980.63">1980</quantity>
</date>
, 63 m, “NOAC Coll.”, [1]; 
<typeStatus id="54E307F3FFD48F06FC624EF63F49548F" box="[1014,1115,484,510]" pageId="4" pageNumber="63" type="paratype">paratype</typeStatus>
2: MZSP 55460, GEOMAR XVI, #9 Am.11, 22º06’8 S 40º31’2 S, 
<date id="FFE69F91FFD48F06FCCC4D1E38FA5757" box="[856,1000,524,550]" pageId="4" pageNumber="63" value="1980-08-20">
20.viii. 
<quantity id="4CA014B4FFD48F06FC3E4D1E38FA5757" box="[938,1000,524,550]" metricMagnitude="3" metricUnit="m" metricValue="1.98059" pageId="4" pageNumber="63" unit="m" value="1980.59">1980</quantity>
</date>
, 59 m, “NOAC Coll.”, [1]; 
<typeStatus id="54E307F3FFD48F06FE9C4D263A7F573F" box="[264,365,564,590]" pageId="4" pageNumber="63" type="paratype">paratype</typeStatus>
3: MORG 50447 GEOMAR XVI, #17 Am.19, 
<geoCoordinate id="EE6CDF96FFD48F06FC334D2638D5573F" box="[935,967,564,590]" direction="south" orientation="latitude" pageId="4" pageNumber="63" precision="925" value="-22.45">22</geoCoordinate>
º27’ S 
<geoCoordinate id="EE6CDF96FFD48F06FB824D263F9B573F" box="[1046,1161,564,590]" direction="west" orientation="longitude" pageId="4" pageNumber="63" precision="925" value="-40.966667">40º58’ W</geoCoordinate>
, 
<date id="FFE69F91FFD48F06FB014D263E34573F" box="[1173,1318,564,590]" pageId="4" pageNumber="63" value="1980-08-20">
20.viii. 
<quantity id="4CA014B4FFD48F06FB734D263E34573F" box="[1255,1318,564,590]" metricMagnitude="3" metricUnit="m" metricValue="1.9805599999999999" pageId="4" pageNumber="63" unit="m" value="1980.56">1980</quantity>
</date>
, 56 m, “NOAC Coll.”, [2]; 
<typeStatus id="54E307F3FFD48F06FDCD4D4E39AC5707" box="[601,702,604,630]" pageId="4" pageNumber="63" type="paratype">paratype</typeStatus>
4: MNHN GEOMAR XVI, #17 Am.19, 
<geoCoordinate id="EE6CDF96FFD48F06FB554D4E3FF35707" box="[1217,1249,604,630]" direction="south" orientation="latitude" pageId="4" pageNumber="63" precision="925" value="-22.45">22</geoCoordinate>
º27’ S 
<geoCoordinate id="EE6CDF96FFD48F06FE9C4D963A6E57EF" box="[264,380,644,670]" direction="west" orientation="longitude" pageId="4" pageNumber="63" precision="925" value="-40.966667">40º58’ W</geoCoordinate>
, 
<date id="FFE69F91FFD48F06FE1E4D96390857EF" box="[394,538,644,670]" pageId="4" pageNumber="63" value="1980-08-20">
20.viii. 
<quantity id="4CA014B4FFD48F06FE484D96390857EF" box="[476,538,644,670]" metricMagnitude="3" metricUnit="m" metricValue="1.9805599999999999" pageId="4" pageNumber="63" unit="m" value="1980.56">1980</quantity>
</date>
, 56 m, “NOAC Coll.”, [2]; IBUFRJ 7872: GEOMAR XII #102, 21º38’,8 S 40º14’5 W, 
<date id="FFE69F91FFD48F06FDB34DBE39AA57B7" box="[551,696,684,710]" pageId="4" pageNumber="63" value="1979-08-28">28.viii.1979</date>
, “NOAS Coll.” [2]; IBUFRJ 7659: GEOMAR XII #101, 
<date id="FFE69F91FFD48F06FEC54DC63A62579F" box="[337,368,724,750]" pageId="4" pageNumber="63" value="1979-08-29">29</date>
.viii.1979, “NOAS Coll.” [4]; IBUFRJ 13808 REVIZEE #D3, 
<geoCoordinate id="EE6CDF96FFD48F06FBDE4DC63F7B579F" box="[1098,1129,724,750]" direction="south" orientation="latitude" pageId="4" pageNumber="63" precision="925" value="-22.866667">22</geoCoordinate>
º52’ S 
<geoCoordinate id="EE6CDF96FFD48F06FB224DC63E34579F" box="[1206,1318,724,750]" direction="west" orientation="longitude" pageId="4" pageNumber="63" precision="925" value="-41.15">41º09’ W</geoCoordinate>
, 
<quantity id="4CA014B4FFD48F06FE9C4DEE3A545667" box="[264,326,764,790]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="63" unit="m" value="80.0">80 m</quantity>
, 
<date id="FFE69F91FFD48F06FEC64DEE3ADB5667" box="[338,457,764,790]" pageId="4" pageNumber="63" value="1996-02-23">23.ii.1996</date>
[1].
</paragraph>
</subSubSection>
<caption id="DF27E9D9FFD48F06FE9C49F73F1E5214" httpUri="https://zenodo.org/record/273446/files/figure.png" pageId="4" pageNumber="63" targetBox="[319,1281,830,1758]" targetPageId="4">
<paragraph id="8BE7B951FFD48F06FE9C49F73F1E5214" blockId="4.[264,1323,1765,1893]" pageId="4" pageNumber="63">
<emphasis id="B92C6543FFD48F06FE9C49F73AD3538C" bold="true" box="[264,449,1765,1789]" pageId="4" pageNumber="63">FIGURES 6–11.</emphasis>
<taxonomicName id="4C58C2D2FFD48F06FE5E49F4397E538C" box="[458,620,1766,1789]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="63" phylum="Mollusca" rank="species" species="thomei" status="sp. nov.">
<emphasis id="B92C6543FFD48F06FE5E49F4397E538C" box="[458,620,1766,1789]" italics="true" pageId="4" pageNumber="63">Ketosia thomei</emphasis>
</taxonomicName>
<emphasis id="B92C6543FFD48F06FDE149F739A1538C" bold="true" box="[629,691,1765,1789]" pageId="4" pageNumber="63">
<taxonomicNameLabel id="A21FD838FFD48F06FDE149F739A1538C" box="[629,691,1765,1789]" pageId="4" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
<emphasis id="B92C6543FFD48F06FD2849F439D7538C" box="[700,709,1766,1789]" italics="true" pageId="4" pageNumber="63">­</emphasis>
6: holotype (MNRJ 10505), length 2.1 mm; 7: paratype (IBUFRJ 14696) length 1.8 mm; 8: detail of the teleoconch of holotype; 9: protoconch of paratype (IBUFRJ 14696); 10–11: detail of sculpture of the protoconch of paratype (IBUFRJ 14696). Scale bar: 6: 500 m; 7: 200 m; 8: 10 m; 9: 100 m; 10: 20 m; 11: 10 m.
</paragraph>
</caption>
<paragraph id="8BE7B951FFD58F07FE9C4E0E3A4554F7" pageId="5" pageNumber="64">
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<th id="76B9D26FFFD570FDFE9C4E0E3AA55444" box="[264,439,284,309]" gridcol="0" gridrow="0" pageId="5" pageNumber="64">
<emphasis id="B92C6543FFD58F07FE9C4E0E3A8C5444" box="[264,414,284,309]" italics="true" pageId="5" pageNumber="64">Type locality</emphasis>
</th>
</tr>
<tr id="3568BB13FFD570FDFE9C4E563E38542F" box="[264,1322,324,350]" gridrow="1" pageId="5" pageNumber="64">
<th id="76B9D26FFFD570FDFE9C4E563AA5542F" box="[264,439,324,350]" gridcol="0" gridrow="1" pageId="5" pageNumber="64">GEOMAR</th>
<td id="76B9D26FFFD570FDFE564E563AE4542F" box="[450,502,324,350]" gridcol="1" gridrow="1" pageId="5" pageNumber="64">XVI</td>
<td id="76B9D26FFFD570FDFD954E56393F542F" box="[513,557,324,350]" gridcol="2" gridrow="1" pageId="5" pageNumber="64">#17</td>
<td id="76B9D26FFFD570FDFDAC4E563982542F" box="[568,656,324,350]" gridcol="3" gridrow="1" pageId="5" pageNumber="64">Am.19,</td>
<td id="76B9D26FFFD570FDFD0F4E5639FB542F" box="[667,745,324,350]" gridcol="4" gridrow="1" pageId="5" pageNumber="64">22º27’</td>
<td id="76B9D26FFFD570FDFD674E563811542F" box="[755,771,324,350]" gridcol="5" gridrow="1" pageId="5" pageNumber="64">S</td>
<td id="76B9D26FFFD570FDFC9B4E56384E542F" box="[783,860,324,350]" gridcol="6" gridrow="1" pageId="5" pageNumber="64">40º58’</td>
<td id="76B9D26FFFD570FDFCF34E563895542F" box="[871,903,324,350]" gridcol="7" gridrow="1" pageId="5" pageNumber="64">W,</td>
<td id="76B9D26FFFD570FDFC064E5638AE542F" box="[914,956,324,350]" gridcol="8" gridrow="1" pageId="5" pageNumber="64">Rio</td>
<td id="76B9D26FFFD570FDFC534E5638F1542F" box="[967,995,324,350]" gridcol="9" gridrow="1" pageId="5" pageNumber="64">de</td>
<td id="76B9D26FFFD570FDFC7A4E563F51542F" box="[1006,1091,324,350]" gridcol="10" gridrow="1" pageId="5" pageNumber="64">Janeiro</td>
<td id="76B9D26FFFD570FDFBD94E563F98542F" box="[1101,1162,324,350]" gridcol="11" gridrow="1" pageId="5" pageNumber="64">state,</td>
<td id="76B9D26FFFD570FDFB014E563E15542F" box="[1173,1287,324,350]" gridcol="12" gridrow="1" pageId="5" pageNumber="64">Southeast</td>
<td id="76B9D26FFFD570FDFA864E563E38542F" box="[1298,1322,324,350]" gridcol="13" gridrow="1" pageId="5" pageNumber="64">of</td>
</tr>
<tr id="3568BB13FFD570FDFE9C4E7E3E3854F7" box="[264,1322,364,390]" gridrow="2" pageId="5" pageNumber="64" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76B9D26FFFD570FDFE9C4E7E3AA554F7" box="[264,439,364,390]" gridcol="0" gridrow="2" pageId="5" pageNumber="64">Brazil.</th>
</tr>
</table>
</paragraph>
<subSubSection id="C342EADAFFD58F07FE9C4EAE38F05757" pageId="5" pageNumber="64" type="etymology">
<paragraph id="8BE7B951FFD58F07FE9C4EAE3A9754A4" blockId="5.[264,1322,444,550]" box="[264,389,444,469]" pageId="5" pageNumber="64">
<emphasis id="B92C6543FFD58F07FE9C4EAE3A9754A4" box="[264,389,444,469]" italics="true" pageId="5" pageNumber="64">Etymology</emphasis>
</paragraph>
<paragraph id="8BE7B951FFD58F07FEAC4EF638F05757" blockId="5.[264,1322,444,550]" pageId="5" pageNumber="64">
This species is named in honour of Dr. Wilibaldo Thomé, who has contributed to the knowledge of the 
<taxonomicName id="4C58C2D2FFD58F07FE4F4D1E39965757" box="[475,644,524,550]" class="Gastropoda" family="Veronicellidae" kingdom="Animalia" order="Systellommatophora" pageId="5" pageNumber="64" phylum="Mollusca" rank="family">Veronicellidae</taxonomicName>
in 
<collectingCountry id="F34FF9C1FFD58F07FD3D4D1E39E35757" box="[681,753,524,550]" name="Brazil" pageId="5" pageNumber="64">Brazil</collectingCountry>
and South 
<collectingCountry id="F34FF9C1FFD58F07FCE14D1E38CC5757" box="[885,990,524,550]" name="United States of America" pageId="5" pageNumber="64">America</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C342EADAFFD58F04FE9C4D4E3FB45037" lastPageId="6" lastPageNumber="65" pageId="5" pageNumber="64" type="discussion">
<paragraph id="8BE7B951FFD58F07FE9C4D4E3A9B5704" blockId="5.[264,1324,604,1905]" box="[264,393,604,629]" pageId="5" pageNumber="64">
<emphasis id="B92C6543FFD58F07FE9C4D4E3A9B5704" box="[264,393,604,629]" italics="true" pageId="5" pageNumber="64">Discussion</emphasis>
</paragraph>
<paragraph id="8BE7B951FFD58F07FEAC4D6D38C751D0" blockId="5.[264,1324,604,1905]" pageId="5" pageNumber="64">
<taxonomicName id="4C58C2D2FFD58F07FEAC4D6D3A8357E9" box="[312,401,639,664]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FEAC4D6D3A8357E9" box="[312,401,639,664]" italics="true" pageId="5" pageNumber="64">Ketosia</emphasis>
</taxonomicName>
, the new genus herein proposed, belongs to 
<taxonomicName id="4C58C2D2FFD58F07FC754D6D3F4C57E8" box="[993,1118,639,665]" class="Gastropoda" family="Barleeiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="family">Barleeidae</taxonomicName>
because of the presence of minute pits on the protoconch and teleoconch. The distribution of these pits can vary within the family. They may be present on the protoconch in spiral rows, as in 
<taxonomicName id="4C58C2D2FFD58F07FE9C4DE53A7F5661" box="[264,365,759,784]" class="Gastropoda" family="Barleeiidae" genus="Barleeia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FE9C4DE53A7F5661" box="[264,365,759,784]" italics="true" pageId="5" pageNumber="64">Barleeia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C58C2D2FFD58F07FEE84DE5390D5661" box="[380,543,759,784]" class="Gastropoda" family="Barleeiidae" genus="Protobarleeia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FEE84DE5390D5661" box="[380,543,759,784]" italics="true" pageId="5" pageNumber="64">Protobarleeia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C58C2D2FFD58F07FDBB4DE539BA5661" box="[559,680,759,784]" class="Gastropoda" family="Barleeiidae" genus="Fictonoba" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FDBB4DE539BA5661" box="[559,680,759,784]" italics="true" pageId="5" pageNumber="64">Fictonoba</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C58C2D2FFD58F07FD7E4DE538845661" box="[746,918,759,784]" class="Gastropoda" family="Barleeiidae" genus="Tropidorissoia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FD7E4DE538845661" box="[746,918,759,784]" italics="true" pageId="5" pageNumber="64">Tropidorissoia</emphasis>
</taxonomicName>
; or they are randomly distributed, as in 
<taxonomicName id="4C58C2D2FFD58F07FED14C0D3AAE5649" box="[325,444,799,824]" class="Gastropoda" family="Barleeiidae" genus="Caelatura" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FED14C0D3AAE5649" box="[325,444,799,824]" italics="true" pageId="5" pageNumber="64">Caelatura</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C58C2D2FFD58F07FE5F4C0D394C5649" box="[459,606,799,824]" class="Gastropoda" family="Barleeiidae" genus="Lirobarleeia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FE5F4C0D394C5649" box="[459,606,799,824]" italics="true" pageId="5" pageNumber="64">Lirobarleeia</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C58C2D2FFD58F07FD094C0D383D5649" box="[669,815,799,824]" class="Gastropoda" family="Barleeiidae" genus="Pseudodiala" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FD094C0D383D5649" box="[669,815,799,824]" italics="true" pageId="5" pageNumber="64">Pseudodiala</emphasis>
</taxonomicName>
. On the teleoconch, pits are present only in 
<taxonomicName id="4C58C2D2FFD58F07FE9C4C553A6D5611" box="[264,383,839,864]" class="Gastropoda" family="Barleeiidae" genus="Caelatura" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FE9C4C553A6D5611" box="[264,383,839,864]" italics="true" pageId="5" pageNumber="64">Caelatura</emphasis>
</taxonomicName>
(
<tableCitation id="C6DA8CEAFFD58F07FE044C553AFB5610" box="[400,489,839,865]" captionStart="TABLE 1" captionStartId="6.[264,343,289,310]" captionTargetBox="[264,1322,372,773]" captionTargetPageId="6" captionText="TABLE 1. Main diagnostic characters of the genera of the family Barleeidae. Sources: Ponder (1983) and" httpUri="http://table.plazi.org/id/DF27E9D9FFD68F04FE9C4E333E385447" pageId="5" pageNumber="64" tableUuid="DF27E9D9FFD68F04FE9C4E333E385447">Table 1</tableCitation>
) but in 
<taxonomicName id="4C58C2D2FFD58F07FDD94C5539B55611" box="[589,679,839,864]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FDD94C5539B55611" box="[589,679,839,864]" italics="true" pageId="5" pageNumber="64">Ketosia</emphasis>
</taxonomicName>
they are less abundant than on the protoconch. Thus, the presence and distribution of pits on both proto­ and teleoconch, besides the unique shell shape, resembling a 
<emphasis id="B92C6543FFD58F07FDA04C8539B956C1" box="[564,683,919,944]" italics="true" pageId="5" pageNumber="64">Halystilus</emphasis>
, produce a unique combination of shell characters that does not fit within any known genus of 
<taxonomicName id="4C58C2D2FFD58F07FD764CAD387156A8" box="[738,867,959,985]" class="Gastropoda" family="Barleeiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="family">Barleeidae</taxonomicName>
. The position of 
<taxonomicName id="4C58C2D2FFD58F07FBB84CAD3F9456A9" box="[1068,1158,959,984]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FBB84CAD3F9456A9" box="[1068,1158,959,984]" italics="true" pageId="5" pageNumber="64">Ketosia</emphasis>
</taxonomicName>
in 
<taxonomicName id="4C58C2D2FFD58F07FB394CAD3E3856A8" box="[1197,1322,959,985]" class="Gastropoda" family="Barleeiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="family">Barleeidae</taxonomicName>
could not be confirmed with soft part and opercular characters such as minute head with long cephalic tentacles, with eyes localized at the base; pedal sole short, simple; central tooth of radula with one pair of basal denticles, thick marginal teeth and short lateral teeth, and operculum corneus, oval, with or without peg, with excentric nucleus; but the presence of the shell pits makes a placement in 
<taxonomicName id="4C58C2D2FFD58F07FD5C4B95385751D0" box="[712,837,1159,1185]" class="Gastropoda" family="Barleeiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="family">Barleeidae</taxonomicName>
most likely.
</paragraph>
<paragraph id="8BE7B951FFD58F07FEAC4BBD3A8F5090" blockId="5.[264,1324,604,1905]" pageId="5" pageNumber="64">
Representatives of some genera of other families of Rissooidea also have pits on the protoconch or teleoconch: 
<taxonomicName id="4C58C2D2FFD58F07FDDC4BC5398A5181" box="[584,664,1239,1264]" class="Gastropoda" family="Rissoidae" genus="Onoba" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FDDC4BC5398A5181" box="[584,664,1239,1264]" italics="true" pageId="5" pageNumber="64">Onoba</emphasis>
</taxonomicName>
H. &amp; A. Adams, 1852; 
<taxonomicName id="4C58C2D2FFD58F07FC294BC53F1C5181" box="[957,1038,1239,1264]" class="Gastropoda" family="Rissoidae" genus="Zebina" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FC294BC53F1C5181" box="[957,1038,1239,1264]" italics="true" pageId="5" pageNumber="64">Zebina</emphasis>
</taxonomicName>
H. &amp; A. Adams, 1854; 
<taxonomicName id="4C58C2D2FFD58F07FE9C4BED39955068" authority="Cossmann, 1888" authorityName="Cossmann" authorityYear="1888" box="[264,647,1279,1305]" class="Gastropoda" family="Rissoinidae" genus="Pseudotaphrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FE9C4BED3AAA5069" box="[264,440,1279,1304]" italics="true" pageId="5" pageNumber="64">Pseudotaphrus</emphasis>
Cossmann, 1888
</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FD004BED3F9D5068" authority="Golikov and Kussakin, 1974" authorityName="Golikov and Kussakin" authorityYear="1974" box="[660,1167,1279,1305]" class="Gastropoda" family="Rissoidae" genus="Boreocingula" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FD004BED38215069" box="[660,819,1279,1304]" italics="true" pageId="5" pageNumber="64">Boreocingula</emphasis>
Golikov and Kussakin, 1974
</taxonomicName>
; and 
<taxonomicName id="4C58C2D2FFD58F07FB5A4BED39295030" authority="Bouchet &amp; Waren, 1993" authorityName="Bouchet &amp; Waren" authorityYear="1993" class="Gastropoda" family="Rissoidae" genus="Gofasia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FB5A4BED3E395069" box="[1230,1323,1279,1304]" italics="true" pageId="5" pageNumber="64">Gofasia</emphasis>
Bouchet &amp; Warén, 1993
</taxonomicName>
, all 
<taxonomicName id="4C58C2D2FFD58F07FDED4A3539FD5030" box="[633,751,1319,1345]" class="Gastropoda" family="Rissoidae" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="family">Rissoidae</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FC954A353F9F5030" authority="Carpenter, 1864" authorityName="Carpenter" authorityYear="1864" box="[769,1165,1319,1345]" class="Gastropoda" family="Barleeiidae" genus="Amphithalamus" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FC954A3538AB5031" box="[769,953,1319,1344]" italics="true" pageId="5" pageNumber="64">Amphithalamus</emphasis>
Carpenter, 1864
</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FB344A353AC55018" authority="Frauenfeld, 1867" authorityName="Frauenfeld" authorityYear="1867" class="Gastropoda" family="Anabathridae" genus="Anabathron" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FB344A353E395031" box="[1184,1323,1319,1344]" italics="true" pageId="5" pageNumber="64">Anabathron</emphasis>
Frauenfeld, 1867
</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FE7C4A5D38235018" authority="Monterosato, 1878" authorityName="Monterosato" authorityYear="1878" box="[488,817,1359,1385]" class="Gastropoda" family="Anabathridae" genus="Pisinna" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FE7C4A5D39505019" box="[488,578,1359,1384]" italics="true" pageId="5" pageNumber="64">Pisinna</emphasis>
Monterosato, 1878
</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FCD54A5D3F815018" authority="Monterosato, 1878" authorityName="Monterosato" authorityYear="1878" box="[833,1171,1359,1385]" class="Gastropoda" family="Anabathridae" genus="Nodulus" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FCD54A5D38B15019" box="[833,931,1359,1384]" italics="true" pageId="5" pageNumber="64">Nodulus</emphasis>
Monterosato, 1878
</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FB374A5D3AA950E0" authority="Laseron, 1950" authorityName="Laseron" authorityYear="1950" class="Gastropoda" family="Anabathridae" genus="Microdryas" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FB374A5D3E395019" box="[1187,1323,1359,1384]" italics="true" pageId="5" pageNumber="64">Microdryas</emphasis>
Laseron, 1950
</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FE5B4A65381E50E0" authority="Iredale, 1955" authorityName="Iredale" authorityYear="1955" box="[463,780,1399,1425]" class="Gastropoda" family="Anabathridae" genus="Badepigrus" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FE5B4A65394450E1" box="[463,598,1399,1424]" italics="true" pageId="5" pageNumber="64">Badepigrus</emphasis>
Iredale, 1955
</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FCB44A653F4850E0" authority="Ponder, 1967" authorityName="Ponder" authorityYear="1967" box="[800,1114,1399,1425]" class="Gastropoda" family="Anabathridae" genus="Pseudestea" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FCB44A6538B650E1" box="[800,932,1399,1424]" italics="true" pageId="5" pageNumber="64">Pseudestea</emphasis>
Ponder, 1967
</taxonomicName>
; and 
<emphasis id="B92C6543FFD58F07FB3C4A653E3850E1" box="[1192,1322,1399,1424]" italics="true" pageId="5" pageNumber="64">Afriscobris</emphasis>
<bibRefCitation id="EFC9C4A0FFD58F07FE9C4A8D3AB950C8" author="Ponder" box="[264,427,1439,1465]" pageId="5" pageNumber="68" refString="Ponder, W. F. (1983) Review of the genera of the Barleeidae (Mollusca: Gastropoda: Rissoacea). Records of the Australian Museum, 35, 231 - 281." type="journal article" year="1983">Ponder, 1983</bibRefCitation>
, all 
<taxonomicName id="4C58C2D2FFD58F07FE724A8D399650C8" box="[486,644,1439,1465]" class="Gastropoda" family="Anabathridae" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="family">Anabathridae</taxonomicName>
(
<bibRefCitation id="EFC9C4A0FFD58F07FD0D4A8D382550C8" author="Ponder" box="[665,823,1439,1465]" pageId="5" pageNumber="68" refString="Ponder, W. F. (1983) Review of the genera of the Barleeidae (Mollusca: Gastropoda: Rissoacea). Records of the Australian Museum, 35, 231 - 281." type="journal article" year="1983">Ponder 1983</bibRefCitation>
, 
<bibRefCitation id="EFC9C4A0FFD58F07FCDD4A8D389550C8" author="Ponder" box="[841,903,1439,1465]" pageId="5" pageNumber="68" refString="Ponder, W. F. (1984) A review of the genera of the Rissoidae (Mollusca: Mesogastropoda: Rissoacea). Records of the Australian Museum, 35, 231 - 281." type="journal article" year="1984">1984</bibRefCitation>
; 
<bibRefCitation id="EFC9C4A0FFD58F07FC0C4A8D3FC050C8" author="Ponder" box="[920,1234,1439,1465]" pageId="5" pageNumber="68" refString="Ponder, W. F. &amp; Worsfold, T. M. (1994) A Review of the Rissoiform Gastropods of Southwestern South America (Mollusca, Gastropoda). Contributions in Science, Natural History Museum of Los Angeles County, 445, 1 - 63." type="journal article" year="1994">Ponder &amp; Worsfold 1994</bibRefCitation>
; 
<bibRefCitation id="EFC9C4A0FFD58F07FB704A8D3A575090" author="Gofas" pageId="5" pageNumber="68" refString="Gofas, S. (1995) A remarkable specie richness of the Barleeidae (Gastropoda: Rissoacea) in the Eastern Atlantic. The Nautilus, 109, 14 - 37." type="journal article" year="1995">Gofas 1995</bibRefCitation>
, 
<bibRefCitation id="EFC9C4A0FFD58F07FEC54AD53A825090" author="Gofas" box="[337,400,1479,1505]" pageId="5" pageNumber="68" refString="Gofas, S. (1999). The west African Rissoidae (Gastropoda: Rissooidea) and their similarities to some European species. The Nautilus, 113, 78 - 101." type="journal article" year="1999">1999</bibRefCitation>
).
</paragraph>
<paragraph id="8BE7B951FFD58F07FEAC4AFD3E1D5328" blockId="5.[264,1324,604,1905]" pageId="5" pageNumber="64">
<taxonomicName id="4C58C2D2FFD58F07FEAC4AFD3A805379" box="[312,402,1519,1544]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FEAC4AFD3A805379" box="[312,402,1519,1544]" italics="true" pageId="5" pageNumber="64">Ketosia</emphasis>
</taxonomicName>
does not fit in 
<taxonomicName id="4C58C2D2FFD58F07FDC84AFD39EB5378" box="[604,761,1519,1545]" class="Gastropoda" family="Anabathridae" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="family">Anabathridae</taxonomicName>
because this family show a protoconch with spirally arranged pits, and show an umbilicus, and does not fit in 
<taxonomicName id="4C58C2D2FFD58F07FB9049053F645340" box="[1028,1142,1559,1585]" class="Gastropoda" family="Rissoidae" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="family">Rissoidae</taxonomicName>
because shows spiral and/or axial sculpture (e.g., 
<taxonomicName id="4C58C2D2FFD58F07FD0C492D389B5328" authority="Risso, 1826" authorityName="Risso" authorityYear="1826" box="[664,905,1599,1625]" class="Gastropoda" family="Rissoidae" genus="Alvania" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FD0C492D39E15329" box="[664,755,1599,1624]" italics="true" pageId="5" pageNumber="64">
<collectingCountry id="F34FF9C1FFD58F07FD0C492D39E15329" box="[664,755,1599,1624]" name="Albania" pageId="5" pageNumber="64">Alvania</collectingCountry>
</emphasis>
Risso, 1826
</taxonomicName>
), which is not present in 
<taxonomicName id="4C58C2D2FFD58F07FB3A492D3E1A5329" box="[1198,1288,1599,1624]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FB3A492D3E1A5329" box="[1198,1288,1599,1624]" italics="true" pageId="5" pageNumber="64">Ketosia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE7B951FFD58F07FEAC49753FBD5200" blockId="5.[264,1324,604,1905]" pageId="5" pageNumber="64">
Six species of 
<taxonomicName id="4C58C2D2FFD58F07FE664975397D53F0" box="[498,623,1639,1665]" class="Gastropoda" family="Barleeiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="family">Barleeidae</taxonomicName>
were previously known from Brazilian waters: 
<taxonomicName id="4C58C2D2FFD58F07FB514975384453D8" authority="Catellanos &amp; Fernandez, 1972" authorityName="Catellanos &amp; Fernandez" authorityYear="1972" class="Gastropoda" family="Barleeiidae" genus="Barleeia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="species" species="rubrooperculata">
<emphasis id="B92C6543FFD58F07FB5149753AD553D9" italics="true" pageId="5" pageNumber="64">Barleeia rubrooperculata</emphasis>
(Catellanos &amp; Fernandez, 1972)
</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FCF0499D3A5453A0" authority="Absalao &amp; Rios, 1995" authorityName="Absalao &amp; Rios" authorityYear="1995" class="Gastropoda" family="Barleeiidae" genus="Caelatura" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="species" species="barcellosi">
<emphasis id="B92C6543FFD58F07FCF0499D3F4A53D9" box="[868,1112,1679,1704]" italics="true" pageId="5" pageNumber="64">Caelatura barcellosi</emphasis>
<bibRefCitation id="EFC9C4A0FFD58F07FBF7499D3A5453A0" author="Absalao" pageId="5" pageNumber="68" refString="Absalao, R. S. &amp; Rios, E. C. (1995) Descriptions of two new species of Caelatura (Gastropoda, Risooidea, Barleeidae) from Brazil. Apex, 10 (2 / 3), 87 - 93." type="journal article" year="1995">Absalão &amp; Rios, 1995</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FEC049A5383253A0" authority="Absalao &amp; Rios, 1995" authorityName="Absalao &amp; Rios" authorityYear="1995" box="[340,800,1719,1745]" class="Gastropoda" family="Barleeiidae" genus="Caelatura" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="species" species="spirocordata">
<emphasis id="B92C6543FFD58F07FEC049A5391E53A1" box="[340,524,1719,1744]" italics="true" pageId="5" pageNumber="64">C. spirocordata</emphasis>
<bibRefCitation id="EFC9C4A0FFD58F07FD8049A5383253A0" author="Absalao" box="[532,800,1719,1745]" pageId="5" pageNumber="68" refString="Absalao, R. S. &amp; Rios, E. C. (1995) Descriptions of two new species of Caelatura (Gastropoda, Risooidea, Barleeidae) from Brazil. Apex, 10 (2 / 3), 87 - 93." type="journal article" year="1995">Absalão &amp; Rios, 1995</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FCB949A53F4553A0" authority="Absalao, 2002" authorityName="Absalao" authorityYear="2002" box="[813,1111,1719,1745]" class="Gastropoda" family="Barleeiidae" genus="Caelatura" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="species" species="tigrina">
<emphasis id="B92C6543FFD58F07FCB949A538B253A1" box="[813,928,1719,1744]" italics="true" pageId="5" pageNumber="64">C. tigrina</emphasis>
<bibRefCitation id="EFC9C4A0FFD58F07FC3D49A53F4553A0" author="Absalao" box="[937,1111,1719,1745]" pageId="5" pageNumber="68" refString="Absalao, R. S. (2002) Three new species of Barleeidae (Mollusca, Gastropoda, Rissooidea) from Brazil. Zootaxa, 56, 1 - 7." type="journal article" year="2002">Absalão, 2002</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C58C2D2FFD58F07FBF049A53AAE5388" authority="Absalao, 2002" authorityName="Absalao" authorityYear="2002" class="Gastropoda" family="Barleeiidae" genus="Caelatura" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="species" species="speculabunda">
<emphasis id="B92C6543FFD58F07FBF049A53E3953A1" box="[1124,1323,1719,1744]" italics="true" pageId="5" pageNumber="64">C. speculabunda</emphasis>
<bibRefCitation id="EFC9C4A0FFD58F07FE9C49CD3AAE5388" author="Absalao" box="[264,444,1759,1785]" pageId="5" pageNumber="68" refString="Absalao, R. S. (2002) Three new species of Barleeidae (Mollusca, Gastropoda, Rissooidea) from Brazil. Zootaxa, 56, 1 - 7." type="journal article" year="2002">Absalão, 2002</bibRefCitation>
</taxonomicName>
; and 
<taxonomicName id="4C58C2D2FFD58F07FD9D49CD3F145388" authority="Absalao, 2002" authorityName="Absalao" authorityYear="2002" box="[521,1030,1759,1785]" class="Gastropoda" family="Barleeiidae" genus="Protobarleeia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="species" species="pyrrocincta">
<emphasis id="B92C6543FFD58F07FD9D49CD38515389" box="[521,835,1759,1784]" italics="true" pageId="5" pageNumber="64">Protobarleeia pyrrocincta</emphasis>
<bibRefCitation id="EFC9C4A0FFD58F07FCC549CD3F145388" author="Absalao" box="[849,1030,1759,1785]" pageId="5" pageNumber="68" refString="Absalao, R. S. (2002) Three new species of Barleeidae (Mollusca, Gastropoda, Rissooidea) from Brazil. Zootaxa, 56, 1 - 7." type="journal article" year="2002">Absalão, 2002</bibRefCitation>
</taxonomicName>
. None of these can be confused with 
<taxonomicName id="4C58C2D2FFD58F07FE54481539085251" box="[448,538,1799,1824]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="64" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD58F07FE54481539085251" box="[448,538,1799,1824]" italics="true" pageId="5" pageNumber="64">Ketosia</emphasis>
</taxonomicName>
, because the new species belongs to a distinct genus with the surface of the protoconch and teleoconch covered with irregular pits, besides the depression separating the peripheral region of the teleoconch from the inner lip.
</paragraph>
<caption id="DF27E9D9FFD68F04FE9C4E333E385447" ID-Table-UUID="DF27E9D9FFD68F04FE9C4E333E385447" box="[264,1322,289,310]" httpUri="http://table.plazi.org/id/DF27E9D9FFD68F04FE9C4E333E385447" pageId="6" pageNumber="65" targetBox="[264,1322,372,773]" targetIsTable="true" targetPageId="6">
<paragraph id="8BE7B951FFD68F04FE9C4E333E385447" blockId="6.[264,1322,289,342]" box="[264,1322,289,310]" pageId="6" pageNumber="65">
<emphasis id="B92C6543FFD68F04FE9C4E333A615447" bold="true" box="[264,371,289,310]" pageId="6" pageNumber="65">TABLE 1.</emphasis>
Main diagnostic characters of the genera of the family 
<taxonomicName id="4C58C2D2FFD68F04FC364E333F1E5447" box="[930,1036,289,310]" class="Gastropoda" family="Barleeiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="family">Barleeidae</taxonomicName>
. Sources: Ponder (1983) and
</paragraph>
</caption>
<paragraph id="8BE7B951FFD68F04FE9C4E533A9C5427" blockId="6.[264,1322,289,342]" box="[264,398,321,342]" pageId="6" pageNumber="65">
<bibRefCitation id="EFC9C4A0FFD68F04FE9C4E533A985427" author="Gofas" box="[264,394,321,342]" pageId="6" pageNumber="68" refString="Gofas, S. (1995) A remarkable specie richness of the Barleeidae (Gastropoda: Rissoacea) in the Eastern Atlantic. The Nautilus, 109, 14 - 37." type="journal article" year="1995">Gofas (1995)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE7B951FFD68F04FE994E663F3E5674" pageId="6" pageNumber="65">
<table id="F9584BF1FFD670FDFE9C4E663E385674" box="[264,1322,372,773]" gridcols="4" gridrows="5" pageId="6" pageNumber="65">
<tr id="3568BB13FFD670FDFE9C4E663E3854FA" box="[264,1322,372,395]" gridrow="0" pageId="6" pageNumber="65">
<th id="76B9D26FFFD670FDFE9C4E663AB054FA" box="[264,418,372,395]" gridcol="0" gridrow="0" pageId="6" pageNumber="65">Genus</th>
<th id="76B9D26FFFD670FDFE234E66396654FA" box="[439,628,372,395]" gridcol="1" gridrow="0" pageId="6" pageNumber="65">Form of shell</th>
<th id="76B9D26FFFD670FDFD134E66389854FA" box="[647,906,372,395]" gridcol="2" gridrow="0" pageId="6" pageNumber="65">Protoconch</th>
<th id="76B9D26FFFD670FDFC094E663E3854FA" box="[925,1322,372,395]" gridcol="3" gridrow="0" pageId="6" pageNumber="65">Teleoconch</th>
</tr>
<tr id="3568BB13FFD670FDFE9C4E8C3E3854A9" box="[264,1322,414,472]" gridrow="1" pageId="6" pageNumber="65">
<th id="76B9D26FFFD670FDFE9C4E8C3AB054A9" box="[264,418,414,472]" gridcol="0" gridrow="1" pageId="6" pageNumber="65">
<taxonomicName id="4C58C2D2FFD68F04FE994E8C3A7B54C4" box="[269,361,414,437]" class="Gastropoda" family="Barleeiidae" genus="Barleeia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD68F04FE994E8C3A7B54C4" box="[269,361,414,437]" italics="true" pageId="6" pageNumber="65">Barleeia</emphasis>
</taxonomicName>
</th>
<td id="76B9D26FFFD670FDFE234E8C396654A9" box="[439,628,414,472]" gridcol="1" gridrow="1" pageId="6" pageNumber="65">conical to oval­ conical</td>
<td id="76B9D26FFFD670FDFD134E8C389854A9" box="[647,906,414,472]" gridcol="2" gridrow="1" pageId="6" pageNumber="65">smooth, with pits spirally arranged</td>
<td id="76B9D26FFFD670FDFC094E8C3E3854A9" box="[925,1322,414,472]" gridcol="3" gridrow="1" pageId="6" pageNumber="65">smooth or with weak spiral thread</td>
</tr>
<tr id="3568BB13FFD670FDFE9C4EFE3E385739" box="[264,1322,492,584]" gridrow="2" pageId="6" pageNumber="65">
<th id="76B9D26FFFD670FDFE9C4EFE3AB05739" box="[264,418,492,584]" gridcol="0" gridrow="2" pageId="6" pageNumber="65">
<taxonomicName id="4C58C2D2FFD68F04FE994EFE3A6B5772" box="[269,377,492,515]" class="Gastropoda" family="Barleeiidae" genus="Caelatura" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD68F04FE994EFE3A6B5772" box="[269,377,492,515]" italics="true" pageId="6" pageNumber="65">Caelatura</emphasis>
</taxonomicName>
</th>
<td id="76B9D26FFFD670FDFE234EFE39665739" box="[439,628,492,584]" gridcol="1" gridrow="2" pageId="6" pageNumber="65">conical­elongated</td>
<td id="76B9D26FFFD670FDFD134EFE38985739" box="[647,906,492,584]" gridcol="2" gridrow="2" pageId="6" pageNumber="65">smooth or with spiral lines, microsculpture of irregular pits</td>
<td id="76B9D26FFFD670FDFC094EFE3E385739" box="[925,1322,492,584]" gridcol="3" gridrow="2" pageId="6" pageNumber="65">smooth with pits, or with axial or spiral sculpturing</td>
</tr>
<tr id="3568BB13FFD670FDFE9C4D4E3E3857C9" box="[264,1322,604,696]" gridrow="3" pageId="6" pageNumber="65">
<th id="76B9D26FFFD670FDFE9C4D4E3AB057C9" box="[264,418,604,696]" gridcol="0" gridrow="3" pageId="6" pageNumber="65">
<taxonomicName id="4C58C2D2FFD68F04FE994D4E3A815702" box="[269,403,604,627]" class="Gastropoda" family="Barleeiidae" genus="Lirobarleeia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD68F04FE994D4E3A815702" box="[269,403,604,627]" italics="true" pageId="6" pageNumber="65">Lirobarleeia</emphasis>
</taxonomicName>
</th>
<td id="76B9D26FFFD670FDFE234D4E396657C9" box="[439,628,604,696]" gridcol="1" gridrow="3" pageId="6" pageNumber="65">conical­elongated</td>
<td id="76B9D26FFFD670FDFD134D4E389857C9" box="[647,906,604,696]" gridcol="2" gridrow="3" pageId="6" pageNumber="65">sometimes with weak spiral ridges, with irregular deep pits,</td>
<td id="76B9D26FFFD670FDFC094D4E3E3857C9" box="[925,1322,604,696]" gridcol="3" gridrow="3" pageId="6" pageNumber="65">axially and/or spirally sculptured by threads and nodulous, without pits</td>
</tr>
<tr id="3568BB13FFD670FDFE9C4DDE3E385674" box="[264,1322,716,773]" gridrow="4" pageId="6" pageNumber="65">
<th id="76B9D26FFFD670FDFE9C4DDE3AB05674" box="[264,418,716,773]" gridcol="0" gridrow="4" pageId="6" pageNumber="65">
<taxonomicName id="4C58C2D2FFD68F04FE994DDE3AB05792" box="[269,418,716,739]" class="Gastropoda" family="Barleeiidae" genus="Protobarleeia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD68F04FE994DDE3AB05792" box="[269,418,716,739]" italics="true" pageId="6" pageNumber="65">Protobarleeia</emphasis>
</taxonomicName>
</th>
<td id="76B9D26FFFD670FDFE234DDE39665674" box="[439,628,716,773]" gridcol="1" gridrow="4" pageId="6" pageNumber="65">o v a t e ­ c o n i c</td>
<td id="76B9D26FFFD670FDFD134DDE38985674" box="[647,906,716,773]" gridcol="2" gridrow="4" pageId="6" pageNumber="65">w i t h s p i r a l l i n e s a n d honey comb­shaped pits</td>
<td id="76B9D26FFFD670FDFC094DDE3E385674" box="[925,1322,716,773]" gridcol="3" gridrow="4" pageId="6" pageNumber="65">with fine, dense, rather irregular spiral threads</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE7B951FFD68F04FE994C0B3F99569C" pageId="6" pageNumber="65">
<table id="F9584BF1FFD670FDFE994C0B3E385161" box="[269,1322,793,1040]" gridcols="4" gridrows="3" pageId="6" pageNumber="65">
<tr id="3568BB13FFD670FDFE994C0B3E385622" box="[269,1322,793,851]" gridrow="0" pageId="6" pageNumber="65">
<th id="76B9D26FFFD670FDFE994C0B3ABB5622" box="[269,425,793,851]" gridcol="0" gridrow="0" pageId="6" pageNumber="65">
<taxonomicName id="4C58C2D2FFD68F04FE994C0B3A695641" box="[269,379,793,816]" class="Gastropoda" family="Barleeiidae" genus="Fictonoba" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD68F04FE994C0B3A695641" box="[269,379,793,816]" italics="true" pageId="6" pageNumber="65">Fictonoba</emphasis>
</taxonomicName>
</th>
<th id="76B9D26FFFD670FDFE234C0B396E5622" box="[439,636,793,851]" gridcol="1" gridrow="0" pageId="6" pageNumber="65">elongate­ovate</th>
<th id="76B9D26FFFD670FDFD134C0B38805622" box="[647,914,793,851]" gridcol="2" gridrow="0" pageId="6" pageNumber="65">with shallow pits and indistinct spiral elements</th>
<th id="76B9D26FFFD670FDFC094C0B3E385622" box="[925,1322,793,851]" gridcol="3" gridrow="0" pageId="6" pageNumber="65">with axial and sometimes spiral sculpture, rarely smooth, without pits</th>
</tr>
<tr id="3568BB13FFD670FDFE994C743E3856B2" box="[269,1322,870,963]" gridrow="1" pageId="6" pageNumber="65">
<th id="76B9D26FFFD670FDFE994C743ABB56B2" box="[269,425,870,963]" gridcol="0" gridrow="1" pageId="6" pageNumber="65">
<taxonomicName id="4C58C2D2FFD68F04FE994C743A80560C" box="[269,402,870,893]" class="Gastropoda" family="Barleeiidae" genus="Pseudodiala" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD68F04FE994C743A80560C" box="[269,402,870,893]" italics="true" pageId="6" pageNumber="65">Pseudodiala</emphasis>
</taxonomicName>
</th>
<td id="76B9D26FFFD670FDFE234C74396E56B2" box="[439,636,870,963]" gridcol="1" gridrow="1" pageId="6" pageNumber="65">conical­elongated, smooth</td>
<td id="76B9D26FFFD670FDFD134C74388056B2" box="[647,914,870,963]" gridcol="2" gridrow="1" pageId="6" pageNumber="65">with spiral lines and pits arranged non­linearly</td>
<td id="76B9D26FFFD670FDFC094C743E3856B2" box="[925,1322,870,963]" gridcol="3" gridrow="1" pageId="6" pageNumber="65">with small convex whorls and subsutural carina, whorl of body with peripheral keel</td>
</tr>
<tr id="3568BB13FFD670FDFE994CC43E385161" box="[269,1322,982,1040]" gridrow="2" pageId="6" pageNumber="65">
<th id="76B9D26FFFD670FDFE994CC43ABB5161" box="[269,425,982,1040]" gridcol="0" gridrow="2" pageId="6" pageNumber="65">
<taxonomicName id="4C58C2D2FFD68F04FE994CC43ABB569C" box="[269,425,982,1005]" class="Gastropoda" family="Barleeiidae" genus="Tropidorissoia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="genus">
<emphasis id="B92C6543FFD68F04FE994CC43ABB569C" box="[269,425,982,1005]" italics="true" pageId="6" pageNumber="65">Tropidorissoia</emphasis>
</taxonomicName>
</th>
<td id="76B9D26FFFD670FDFE234CC4396E5161" box="[439,636,982,1040]" gridcol="1" gridrow="2" pageId="6" pageNumber="65">c o n i c a l ­ o v a t e</td>
<td id="76B9D26FFFD670FDFD134CC438805161" box="[647,914,982,1040]" gridcol="2" gridrow="2" pageId="6" pageNumber="65">w i t h s p i r a l s e r i e s o f p i t s, or with spiral cords</td>
<td id="76B9D26FFFD670FDFC094CC43E385161" box="[925,1322,982,1040]" gridcol="3" gridrow="2" pageId="6" pageNumber="65">with strong spiral keel</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE7B951FFD68F04FEAC4B733FB45037" blockId="6.[264,1323,1118,1350]" pageId="6" pageNumber="65">
<taxonomicName id="4C58C2D2FFD68F04FEAC4B733AFB510B" box="[312,489,1121,1146]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="thomei">
<emphasis id="B92C6543FFD68F04FEAC4B733AFB510B" box="[312,489,1121,1146]" italics="true" pageId="6" pageNumber="65">Ketosia thomei</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C58C2D2FFD68F04FD114B7339CC510B" box="[645,734,1121,1146]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="riosi">
<emphasis id="B92C6543FFD68F04FD114B7339CC510B" box="[645,734,1121,1146]" italics="true" pageId="6" pageNumber="65">K. riosi</emphasis>
</taxonomicName>
in having less convex whorls (41.8o in 
<emphasis id="B92C6543FFD68F04FB254B733E38510B" box="[1201,1322,1121,1146]" italics="true" pageId="6" pageNumber="65">
<taxonomicName id="4C58C2D2FFD68F04FB254B733E35510B" box="[1201,1319,1121,1146]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="thomei">K. thomei</taxonomicName>
,
</emphasis>
46o in 
<taxonomicName id="4C58C2D2FFD68F04FEC34B9E3AA351D4" box="[343,433,1164,1189]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="riosi">
<emphasis id="B92C6543FFD68F04FEC34B9E3AA351D4" box="[343,433,1164,1189]" italics="true" pageId="6" pageNumber="65">K. riosi</emphasis>
</taxonomicName>
) and having a shorter shell (1.9 mm in 
<emphasis id="B92C6543FFD68F04FC054B9E3F1E51D4" box="[913,1036,1164,1189]" italics="true" pageId="6" pageNumber="65">
<taxonomicName id="4C58C2D2FFD68F04FC054B9E3F1A51D4" box="[913,1032,1164,1189]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="thomei">K. thomei</taxonomicName>
,
</emphasis>
2.2 mm in 
<taxonomicName id="4C58C2D2FFD68F04FB0F4B9E3FE751D4" box="[1179,1269,1164,1189]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="riosi">
<emphasis id="B92C6543FFD68F04FB0F4B9E3FE751D4" box="[1179,1269,1164,1189]" italics="true" pageId="6" pageNumber="65">K. riosi</emphasis>
</taxonomicName>
); 
<taxonomicName id="4C58C2D2FFD68F04FA844B9E3A4A51BC" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="thomei">
<emphasis id="B92C6543FFD68F04FA844B9E3A4A51BC" italics="true" pageId="6" pageNumber="65">K. thomei</emphasis>
</taxonomicName>
has a narrow umbilicus, which is absent in 
<taxonomicName id="4C58C2D2FFD68F04FC144BA638CF51BC" box="[896,989,1204,1230]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="riosi">
<emphasis id="B92C6543FFD68F04FC144BA6388151BC" box="[896,915,1204,1229]" italics="true" pageId="6" pageNumber="65">K</emphasis>
. 
<emphasis id="B92C6543FFD68F04FC334BA638CF51BC" box="[935,989,1204,1229]" italics="true" pageId="6" pageNumber="65">riosi</emphasis>
</taxonomicName>
. In 
<taxonomicName id="4C58C2D2FFD68F04FB804BA63F6251BC" box="[1044,1136,1204,1230]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="riosi">
<emphasis id="B92C6543FFD68F04FB804BA63F3551BC" box="[1044,1063,1204,1229]" italics="true" pageId="6" pageNumber="65">K</emphasis>
. 
<emphasis id="B92C6543FFD68F04FBAE4BA63F6251BC" box="[1082,1136,1204,1229]" italics="true" pageId="6" pageNumber="65">riosi</emphasis>
</taxonomicName>
the aperture is separated from the teleoconch by a depression not seen in 
<emphasis id="B92C6543FFD68F04FC5F4BCE3FB85184" box="[971,1194,1244,1269]" italics="true" pageId="6" pageNumber="65">
<taxonomicName id="4C58C2D2FFD68F04FC5F4BCE3F505184" box="[971,1090,1244,1269]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="thomei">K. thomei</taxonomicName>
. 
<taxonomicName id="4C58C2D2FFD68F04FBC44BCE3FB85184" box="[1104,1194,1244,1269]" class="Gastropoda" family="Barleeiidae" genus="Ketosia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="genus">Ketosia</taxonomicName>
</emphasis>
occurs off southeastern 
<collectingCountry id="F34FF9C1FFD68F04FE354A163AFE506F" box="[417,492,1284,1310]" name="Brazil" pageId="6" pageNumber="65">Brazil</collectingCountry>
, at shallow depths (10 to 80 meters). This agrees with the distributional patterns reported for most species of 
<taxonomicName id="4C58C2D2FFD68F04FD2E4A3E38255037" box="[698,823,1324,1350]" class="Gastropoda" family="Barleeiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="65" phylum="Mollusca" rank="family">Barleeidae</taxonomicName>
present off the Brazilian coast.
</paragraph>
</subSubSection>
</treatment>
</document>