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<mods:title id="5FEED5F23D3FD888A0B9A97D180A9140">Corvoheteromeyenia Ezcurra de Drago, 1979 (Spongillidae, Porifera): Genus review with proposal of neotype of Corvoheteromeyenia heterosclera (Ezcurra de Drago, 1974)</mods:title>
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<mods:namePart id="0CEFC5CE2A4851454355B7FB14DB8154">Calheira, Ludimila</mods:namePart>
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<mods:date id="E4F7B143943B3FBD0484BAB8BC055939">2016</mods:date>
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<treatment id="03EF6658F130FF9AFF05F8BEFEA48F86" ID-DOI="http://doi.org/10.5281/zenodo.5615566" ID-GBIF-Taxon="120829068" ID-Zenodo-Dep="5615566" LSID="urn:lsid:plazi:treatment:03EF6658F130FF9AFF05F8BEFEA48F86" httpUri="http://treatment.plazi.org/id/03EF6658F130FF9AFF05F8BEFEA48F86" lastPageId="2" lastPageNumber="353" pageId="1" pageNumber="352">
<subSubSection id="C35C84C5F130FF99FF05F8BEFCDE8B0E" box="[151,794,1830,1856]" pageId="1" pageNumber="352" type="nomenclature">
<paragraph id="8BF9D74EF130FF99FF05F8BEFCDE8B0E" blockId="1.[151,794,1830,1856]" box="[151,794,1830,1856]" pageId="1" pageNumber="352">
<heading id="D0B16022F130FF99FF05F8BEFCDE8B0E" bold="true" box="[151,794,1830,1856]" fontSize="11" level="1" pageId="1" pageNumber="352" reason="1">
<emphasis id="B9320B5CF130FF99FF05F8BEFCDE8B0E" bold="true" box="[151,794,1830,1856]" pageId="1" pageNumber="352">
Genus
<taxonomicName id="4C46ACCDF130FF99FF7CF8BEFCDE8B0E" authority="Ezcurra de Drago, 1979" authorityName="Ezcurra de Drago" authorityYear="1979" box="[238,794,1830,1856]" class="Demospongiae" family="Spongillidae" genus="Corvoheteromeyenia" kingdom="Animalia" order="Haplosclerida" pageId="1" pageNumber="352" phylum="Porifera" rank="genus">
<emphasis id="B9320B5CF130FF99FF7CF8BEFE2D8B0E" bold="true" box="[238,489,1830,1856]" italics="true" pageId="1" pageNumber="352">Corvoheteromeyenia</emphasis>
Ezcurra de Drago, 1979
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C84C5F130FF9AFF05F8F6FD1C8D0E" lastPageId="2" lastPageNumber="353" pageId="1" pageNumber="352" type="reference_group">
<paragraph id="8BF9D74EF130FF99FF05F8F6FD678BEC" blockId="1.[151,787,1901,1984]" pageId="1" pageNumber="352">
<treatmentCitationGroup id="AB56F060F130FF99FF05F8F6FD678BEC" pageId="1" pageNumber="352">
<taxonomicName id="4C46ACCDF130FF99FF05F8F6FD098BCD" authority="Bonetto &amp; Ezcurra de Drago, 1966: 129" authorityName="Bonetto &amp; Ezcurra de Drago" authorityPageNumber="129" authorityYear="1966" box="[151,717,1901,1923]" class="Demospongiae" family="Metaniidae" genus="Corvomeyenia" kingdom="Animalia" order="Haplosclerida" pageId="1" pageNumber="352" phylum="Porifera" rank="genus">
<emphasis id="B9320B5CF130FF99FF05F8F6FEEF8BCD" box="[151,299,1902,1923]" italics="true" pageId="1" pageNumber="352">Corvomeyenia</emphasis>
: Bonetto &amp; Ezcurra de
<treatmentCitation id="0AE7F15FF130FF99FD89F8F6FD098BCD" author="Drago" box="[539,717,1901,1923]" page="129" pageId="1" pageNumber="352" year="1966">
<bibRefCitation id="EFD7AABFF130FF99FD89F8F6FD598BCD" author="Drago" box="[539,669,1901,1923]" pageId="1" pageNumber="352" refString="Bonetto, A. A. &amp; Ezcurra de Drago, I. D. (1966) Nuevos aportes al conocimiento de las esponjas Argentinas. Physis, 26 (71), 129 - 140." type="journal article" year="1966">Drago, 1966</bibRefCitation>
: 129
</treatmentCitation>
</taxonomicName>
(part).
<taxonomicName id="4C46ACCDF130FF99FF05F814FD998BEC" authority="Ezcurra de Drago, 1974: 233" authorityName="Ezcurra de Drago" authorityPageNumber="233" authorityYear="1974" box="[151,605,1932,1954]" class="Demospongiae" family="Metaniidae" genus="Corvomeyenia" kingdom="Animalia" order="Haplosclerida" pageId="1" pageNumber="352" phylum="Porifera" rank="genus">
<emphasis id="B9320B5CF130FF99FF05F814FEEF8BEF" box="[151,299,1932,1953]" italics="true" pageId="1" pageNumber="352">Corvomeyenia</emphasis>
: Ezcurra de Drago, 1974: 233
</taxonomicName>
(part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF9D74EF130FF99FF05F832FD5F8B8E" blockId="1.[151,787,1901,1984]" box="[151,667,1962,1984]" pageId="1" pageNumber="352">
<taxonomicName id="4C46ACCDF130FF99FF05F832FD538B8E" authority="Ezcurra de Drago, 1979: 110" authorityName="Ezcurra de Drago" authorityPageNumber="110" authorityYear="1979" box="[151,663,1962,1984]" class="Demospongiae" family="Spongillidae" genus="Corvoheteromeyenia" kingdom="Animalia" order="Haplosclerida" pageId="1" pageNumber="352" phylum="Porifera" rank="genus">
<emphasis id="B9320B5CF130FF99FF05F832FEAF8B8E" box="[151,363,1962,1984]" italics="true" pageId="1" pageNumber="352">Corvoheteromeyenia</emphasis>
Ezcurra de Drago, 1979: 110
</taxonomicName>
.
</paragraph>
<paragraph id="8BF9D74EF133FF9AFF05FF0FFD1C8D0E" blockId="2.[151,1436,151,968]" pageId="2" pageNumber="353">
<emphasis id="B9320B5CF133FF9AFF05FF0FFE4D8CFE" bold="true" box="[151,393,151,176]" pageId="2" pageNumber="353">Emended diagnoses.</emphasis>
<taxonomicName id="4C46ACCDF133FF9AFE02FF0FFDD98CFE" box="[400,541,151,176]" class="Demospongiae" family="Spongillidae" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="353" phylum="Porifera" rank="family">Spongillidae</taxonomicName>
encrusting. Choanosomal skeleton irregularly alveolate. Megascleres are oxeas, slightly curved, generally smooth, sometimes irregularly microspined. Microscleres are pseudobirotules and/or acanthoxeas varying in shape and size. Gemmules adhered to the substrate or free in the sponge. Foramen single and circular. Gemmular theca tri-layered with gemmuloscleres radially embedded. Gemmuloscleres are birotules or sanidasters (modified from Pinheiro
<emphasis id="B9320B5CF133FF9AFDC2FEB1FD458D0E" box="[592,641,296,320]" italics="true" pageId="2" pageNumber="353">et al</emphasis>
. 2015).
</paragraph>
</subSubSection>
<subSubSection id="C35C84C5F133FF9AFF55FED4FB038ECB" pageId="2" pageNumber="353" type="discussion">
<paragraph id="8BF9D74EF133FF9AFF55FED4FB038ECB" blockId="2.[151,1436,151,968]" pageId="2" pageNumber="353">
<emphasis id="B9320B5CF133FF9AFF55FED4FEFF8D2B" bold="true" box="[199,315,332,357]" pageId="2" pageNumber="353">Remarks.</emphasis>
Ezcurra de Drago (1979) did not mention the presence of acanthoxea microscleres in the diagnosis of
<taxonomicName id="4C46ACCDF133FF9AFF29FEE8FE618DC6" box="[187,421,368,392]" class="Demospongiae" family="Spongillidae" genus="Corvoheteromeyenia" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="353" phylum="Porifera" rank="genus">
<emphasis id="B9320B5CF133FF9AFF29FEE8FE618DC6" box="[187,421,368,392]" italics="true" pageId="2" pageNumber="353">Corvoheteromeyenia</emphasis>
</taxonomicName>
, but in the description of
<taxonomicName id="4C46ACCDF133FF9AFD79FEE8FC598DC6" box="[747,925,368,392]" class="Demospongiae" family="Spongillidae" genus="Corvoheteromeyenia" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="353" phylum="Porifera" rank="species" species="heterosclera">
<emphasis id="B9320B5CF133FF9AFD79FEE8FC598DC6" box="[747,925,368,392]" italics="true" pageId="2" pageNumber="353">C. heterosclera</emphasis>
</taxonomicName>
the author describes these microscleres as &quot;spicules larger than the others, variously curved, with acute tip and surface covered with spikes and/or bifurcated or branched protuberances, which generally reach a greater development in the central portion of the spicule” (translated from Ezcurra de Drago 1974). Despite the acute tip, she considered the acanthoxea microscleres as a variation of the isochelae (
<figureCitation id="137DCBCBF133FF9AFE2DFE67FDC78E56" box="[447,515,511,536]" captionStart="FIGURE 2" captionStartId="4.[151,250,1910,1932]" captionTargetBox="[264,1335,199,1888]" captionTargetId="figure@4.[250,1337,193,1889]" captionTargetPageId="4" captionText="FIGURE 2. Corvoheteromeyenia heterosclera, spicules from original paper (modified from Ezcurra de Drago 1974). Acanthoxea microscleres circumscribed by red line. Holotype on white background and paratype on gray background." httpUri="https://zenodo.org/record/265913/files/figure.png" pageId="2" pageNumber="353">Fig. 2</figureCitation>
, modified from Ezcurra de Drago 1974; Fig.
<date id="FFF8F18EF133FF9AFC6EFD98FBFE8E56" box="[1020,1082,512,536]" pageId="2" pageNumber="353">I.47</date>
). Here, we emend the diagnosis of
<taxonomicName id="4C46ACCDF133FF9AFF2BFDBCFE678E72" box="[185,419,548,572]" class="Demospongiae" family="Spongillidae" genus="Corvoheteromeyenia" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="353" phylum="Porifera" rank="genus">
<emphasis id="B9320B5CF133FF9AFF2BFDBCFE678E72" box="[185,419,548,572]" italics="true" pageId="2" pageNumber="353">Corvoheteromeyenia</emphasis>
</taxonomicName>
to include the presence of acanthoxea microscleres. Additionally, we consider that the oxea, pseudobirotule and birotule spicules described by Bonetto &amp; Ezcurra de
<bibRefCitation id="EFD7AABFF133FF9AFB9AFDD0FB648E2E" author="Drago" box="[1032,1184,583,608]" pageId="2" pageNumber="353" refString="Bonetto, A. A. &amp; Ezcurra de Drago, I. D. (1966) Nuevos aportes al conocimiento de las esponjas Argentinas. Physis, 26 (71), 129 - 140." type="journal article" year="1966">Drago (1966)</bibRefCitation>
and Ezcurra de Drago (1974; 1979) for
<taxonomicName id="4C46ACCDF133FF9AFEC5FDF4FD858ECA" box="[343,577,620,644]" class="Demospongiae" family="Spongillidae" genus="Corvoheteromeyenia" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="353" phylum="Porifera" rank="genus">
<emphasis id="B9320B5CF133FF9AFEC5FDF4FD858ECA" box="[343,577,620,644]" italics="true" pageId="2" pageNumber="353">Corvoheteromeyenia</emphasis>
</taxonomicName>
to be amphioxea, isochelae and amphidiscs, respectively.
</paragraph>
</subSubSection>
<subSubSection id="C35C84C5F133FF9AFF55FD17FAE48EE6" box="[199,1312,655,680]" pageId="2" pageNumber="353" type="materials_examined">
<paragraph id="8BF9D74EF133FF9AFF55FD17FAE48EE6" blockId="2.[151,1436,151,968]" box="[199,1312,655,680]" pageId="2" pageNumber="353">
<emphasis id="B9320B5CF133FF9AFF55FD17FEA28EE6" bold="true" box="[199,358,655,680]" pageId="2" pageNumber="353">
<typeStatus id="54FD69ECF133FF9AFF55FD17FEC68EE6" box="[199,258,655,680]" pageId="2" pageNumber="353">Type</typeStatus>
Species.
</emphasis>
<taxonomicName id="4C46ACCDF133FF9AFEFFFD08FBCF8EE6" authority="Bonetto &amp; Ezcurra de Drago, 1966" authorityName="Bonetto &amp; Ezcurra de Drago" authorityYear="1966" box="[365,1035,655,680]" class="Demospongiae" family="Metaniidae" genus="Corvomeyenia" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="353" phylum="Porifera" rank="species" species="australis">
<emphasis id="B9320B5CF133FF9AFEFFFD08FDBE8EE6" box="[365,634,656,680]" italics="true" pageId="2" pageNumber="353">Corvomeyenia australis</emphasis>
Bonetto &amp; Ezcurra de
<bibRefCitation id="EFD7AABFF133FF9AFCEFFD08FBCF8EE6" author="Drago" box="[893,1035,655,680]" pageId="2" pageNumber="353" refString="Bonetto, A. A. &amp; Ezcurra de Drago, I. D. (1966) Nuevos aportes al conocimiento de las esponjas Argentinas. Physis, 26 (71), 129 - 140." type="journal article" year="1966">Drago, 1966</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C35C84C5F133FF9AFF55FD2CFEA48F86" pageId="2" pageNumber="353" type="discussion">
<paragraph id="8BF9D74EF133FF9AFF55FD2CFEA48F86" blockId="2.[151,1436,151,968]" pageId="2" pageNumber="353">
<emphasis id="B9320B5CF133FF9AFF55FD2CFEFF8E83" bold="true" box="[199,315,692,717]" pageId="2" pageNumber="353">Remarks.</emphasis>
The proposal of
<typeStatus id="54FD69ECF133FF9AFD9EFD2DFDB48E83" box="[524,624,693,717]" pageId="2" pageNumber="353" type="neotype">neotypes</typeStatus>
of
<taxonomicName id="4C46ACCDF133FF9AFD0FFD2CFBDA8E82" box="[669,1054,692,716]" class="Demospongiae" family="Spongillidae" genus="Corvoheteromeyenia" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="353" phylum="Porifera" rank="species" species="heterosclera">
<emphasis id="B9320B5CF133FF9AFD0FFD2CFBDA8E82" box="[669,1054,692,716]" italics="true" pageId="2" pageNumber="353">Corvoheteromeyenia heterosclera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C46ACCDF133FF9AFBCFFD2CFB208E82" box="[1117,1252,692,716]" class="Demospongiae" family="Metaniidae" genus="Corvomeyenia" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="353" phylum="Porifera" rank="species" species="australis">
<emphasis id="B9320B5CF133FF9AFBCFFD2CFB208E82" box="[1117,1252,692,716]" italics="true" pageId="2" pageNumber="353">C. australis</emphasis>
</taxonomicName>
is necessary to clarify the specific status of both species that has become confused due the loss of their
<typeStatus id="54FD69ECF133FF9AFB07FD40FB018EBE" box="[1173,1221,728,752]" pageId="2" pageNumber="353">type</typeStatus>
material. Since
<taxonomicName id="4C46ACCDF133FF9AFA10FD40FEE68F5A" class="Demospongiae" family="Spongillidae" genus="Corvoheteromeyenia" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="353" phylum="Porifera" rank="species" species="heterosclera">
<emphasis id="B9320B5CF133FF9AFA10FD40FEE68F5A" italics="true" pageId="2" pageNumber="353">C. heterosclera</emphasis>
</taxonomicName>
was originally described from Northeast
<collectingCountry id="F35197DEF133FF9AFD66FD64FCFF8F5B" box="[756,827,764,789]" name="Brazil" pageId="2" pageNumber="353">Brazil</collectingCountry>
, we choose UFPEPOR 975; UFPEPOR 1349 that was collected in Pernambuco (Northeast
<collectingCountry id="F35197DEF133FF9AFDA3FC87FDB38F76" box="[561,631,799,824]" name="Brazil" pageId="2" pageNumber="353">Brazil</collectingCountry>
) and we have the opportunity to follow the growth of the specimen. On the other hand, it was not possible to indicate the
<typeStatus id="54FD69ECF133FF9AFD44FCDDFCEB8F13" box="[726,815,837,861]" pageId="2" pageNumber="353" type="neotype">neotype</typeStatus>
of
<emphasis id="B9320B5CF133FF9AFCCAFCDCFC278F12" box="[856,995,836,860]" italics="true" pageId="2" pageNumber="353">
<taxonomicName id="4C46ACCDF133FF9AFCCAFCDCFC248F12" box="[856,992,836,860]" class="Demospongiae" family="Metaniidae" genus="Corvomeyenia" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="353" phylum="Porifera" rank="species" species="australis">C. australis</taxonomicName>
,
</emphasis>
since the loan of specimens that were collected in the
<typeStatus id="54FD69ECF133FF9AFED5FCF0FEB38FCE" box="[327,375,872,896]" pageId="2" pageNumber="353">type</typeStatus>
locality and are deposited at INALI (Instituto Nacional de Limnología) was denied. Therefore, we provide below the characterization of
<taxonomicName id="4C46ACCDF133FF9AFD1DFC14FCDC8FEA" box="[655,792,908,932]" class="Demospongiae" family="Metaniidae" genus="Corvomeyenia" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="353" phylum="Porifera" rank="species" species="australis">
<emphasis id="B9320B5CF133FF9AFD1DFC14FCDC8FEA" box="[655,792,908,932]" italics="true" pageId="2" pageNumber="353">C. australis</emphasis>
</taxonomicName>
based on material collected in the neighboring basin (
<collectingCountry id="F35197DEF133FF9AFF0CFC28FECC8F86" box="[158,264,944,968]" name="Paraguay" pageId="2" pageNumber="353">Paraguay</collectingCountry>
Basin).
</paragraph>
</subSubSection>
</treatment>
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