258 lines
29 KiB
XML
258 lines
29 KiB
XML
<document id="06C4B2E2873802E2EC393A888385B79B" ID-DOI="10.11646/zootaxa.3681.4.1" ID-GBIF-Dataset="17afa4e8-b105-41f8-bf86-e856fc41a3a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="222798" ID-ZooBank="ABBE4175-C2E8-4BB5-9B61-599BD3D0F632" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460640219132" checkinUser="plazi" docAuthor="Colloff, Matthew J. & Cameron, Stephen L." docDate="2013" docId="03EF6064E46EFFE46AEA00B4FEDE7411" docLanguage="en" docName="zt03681p346.pdf" docOrigin="Zootaxa 3681 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malaconothrus Berlese 1904" docType="treatment" docVersion="10" lastPageNumber="307" masterDocId="FFD6181CE46BFFE26A7D0410FF8B764E" masterDocTitle="A phylogenetic analysis and taxonomic revision of the oribatid mite family Malaconothridae (Acari: Oribatida), with new species of Tyrphonothrus and Malaconothrus from Australia" masterLastPageNumber="346" masterPageNumber="301" pageNumber="306" updateTime="1698335724857" updateUser="plazi">
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<mods:title id="0809F063963FE7DCB94EF99FBA805AAB">A phylogenetic analysis and taxonomic revision of the oribatid mite family Malaconothridae (Acari: Oribatida), with new species of Tyrphonothrus and Malaconothrus from Australia</mods:title>
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<mods:namePart id="A4B8F5E3441F5D78120769797741BFA9">Colloff, Matthew J.</mods:namePart>
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<mods:namePart id="9BE6C3F780C553C1D0C407BD0F929155">Cameron, Stephen L.</mods:namePart>
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<mods:title id="D69B2B5EBA6383A8D3914A17FF712E56">Zootaxa</mods:title>
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<mods:date id="C38BF4853328205BACD8D2B6241F1A6E">2013</mods:date>
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<mods:number id="FDE47EE8B9FD29421D9549D267E7B57D">3681</mods:number>
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<treatment id="03EF6064E46EFFE46AEA00B4FEDE7411" ID-DOI="http://doi.org/10.5281/zenodo.6163946" ID-GBIF-Taxon="119571170" ID-Zenodo-Dep="6163946" LSID="urn:lsid:plazi:treatment:03EF6064E46EFFE46AEA00B4FEDE7411" httpUri="http://treatment.plazi.org/id/03EF6064E46EFFE46AEA00B4FEDE7411" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306">
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<subSubSection id="C35C82F9E46EFFE76AEA00B4FE7672F3" box="[151,509,1187,1214]" pageId="5" pageNumber="306" type="nomenclature">
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<paragraph id="8BF9D172E46EFFE76AEA00B4FE7672F3" blockId="5.[151,509,1187,1214]" box="[151,509,1187,1214]" pageId="5" pageNumber="306">
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<heading id="D0B1661EE46EFFE76AEA00B4FE7672F3" bold="true" box="[151,509,1187,1214]" fontSize="11" level="1" pageId="5" pageNumber="306" reason="1">
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<taxonomicName id="4C46AAF1E46EFFE76AEA00B4FE7672F3" ID-CoL="5KD4" authority="Berlese, 1904" authorityName="Berlese" authorityYear="1904" box="[151,509,1187,1214]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
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<emphasis id="B9320D60E46EFFE76AEA00B4FE7672F3" bold="true" box="[151,509,1187,1214]" pageId="5" pageNumber="306">
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<emphasis id="B9320D60E46EFFE76AEA00B4FEDA72F0" bold="true" box="[151,337,1188,1214]" italics="true" pageId="5" pageNumber="306">Malaconothrus</emphasis>
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<bibRefCitation id="EFD7AC83E46EFFE76B2500B3FE7672F3" author="Berlese" box="[344,509,1187,1213]" pageId="5" pageNumber="306" refString="Berlese, A. (1904) Acari nuovi. Manipulus III. Redia, 2, 10 - 32." type="journal article" year="1904">Berlese, 1904</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C82F9E46EFFE76AEA00FBFDC5734E" box="[151,590,1258,1281]" pageId="5" pageNumber="306" type="reference_group">
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<paragraph id="8BF9D172E46EFFE76AEA00FBFDC5734E" blockId="5.[151,1050,1258,1311]" box="[151,590,1258,1281]" pageId="5" pageNumber="306">
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||
<taxonomicName id="4C46AAF1E46EFFE76AEA00FBFDC5734E" authority="Berlese, 1904" authorityName="Berlese" authorityYear="1904" box="[151,590,1258,1281]" class="Arachnida" family="Lohmanniidae" genus="Lohmannia" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="subGenus" subGenus="Malaconothrus">
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<emphasis id="B9320D60E46EFFE76AEA00FBFE80734F" box="[151,267,1259,1281]" italics="true" pageId="5" pageNumber="306">Lohmannia</emphasis>
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||
(
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<emphasis id="B9320D60E46EFFE76B6400FBFE3F734F" box="[281,436,1259,1281]" italics="true" pageId="5" pageNumber="306">Malaconothrus</emphasis>
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||
)
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<bibRefCitation id="EFD7AC83E46EFFE76BBF00FAFDC5734E" author="Berlese" box="[450,590,1258,1280]" pageId="5" pageNumber="306" refString="Berlese, A. (1904) Acari nuovi. Manipulus III. Redia, 2, 10 - 32." type="journal article" year="1904">Berlese, 1904</bibRefCitation>
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||
</taxonomicName>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35C82F9E46EFFE76AEA011AFB917351" box="[151,1050,1289,1311]" pageId="5" pageNumber="306" type="materials_examined">
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||
<paragraph id="8BF9D172E46EFFE76AEA011AFB917351" blockId="5.[151,1050,1258,1311]" box="[151,1050,1289,1311]" pageId="5" pageNumber="306">
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||
<typeStatus id="54FD6FD0E46EFFE76AEA011AFF437351" box="[151,200,1290,1311]" pageId="5" pageNumber="306">Type</typeStatus>
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species:
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<taxonomicName id="4C46AAF1E46EFFE76B5B0119FD2E7351" authority="Michael, 1888" authorityName="Michael" authorityYear="1888" box="[294,677,1289,1311]" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="monodactylus">
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<emphasis id="B9320D60E46EFFE76B5B0119FD817351" box="[294,522,1289,1311]" italics="true" pageId="5" pageNumber="306">Nothrus monodactylus</emphasis>
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<bibRefCitation id="EFD7AC83E46EFFE7686C0119FD2E7351" author="Michael" box="[529,677,1289,1311]" pageId="5" pageNumber="306" refString="Michael, A. D. (1888) British Oribatidae. Vol. II. Ray Society, London." type="book" year="1888">Michael, 1888</bibRefCitation>
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</taxonomicName>
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, designated by
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<bibRefCitation id="EFD7AC83E46EFFE7693F0119FC5D7351" author="Sellnick" box="[834,982,1289,1311]" pageId="5" pageNumber="306" refString="Sellnick, M. (1928) Hornmilben, Oribatei. In: Brohmer, P., Ehrmann, P. & Ulmer, G. (Eds.), Die Tierwelt Mitteleuropeas, Band III Spinnentiere. Abt. 9. Quelle & Meyer, Leipzig. pp. 1 - 42." type="journal article" year="1928">Sellnick, 1928</bibRefCitation>
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, p. 18.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35C82F9E46EFFE76AEA0157FD187312" box="[151,659,1350,1373]" pageId="5" pageNumber="306" type="reference_group">
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<paragraph id="8BF9D172E46EFFE76AEA0157FD187312" blockId="5.[151,1163,1350,1403]" box="[151,659,1350,1373]" pageId="5" pageNumber="306">
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<taxonomicName id="4C46AAF1E46EFFE76AEA0157FE6B7312" ID-CoL="b8aa36c7-6696-441a-8b43-7105eee47948" authority="Berlese, 1916" authorityName="Berlese" authorityYear="1916" box="[151,480,1350,1373]" class="Arachnida" family="Malaconothridae" genus="Trimalaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
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<emphasis id="B9320D60E46EFFE76AEA0157FEC77313" box="[151,332,1351,1373]" italics="true" pageId="5" pageNumber="306">Trimalaconothrus</emphasis>
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||
<bibRefCitation id="EFD7AC83E46EFFE76B2E0156FE6B7312" author="Berlese" box="[339,480,1350,1372]" pageId="5" pageNumber="306" refString="Berlese, A. (1916) Centuria seconda di Acari nuovi. Redia, 12, 125 - 177." type="journal article" year="1916">Berlese, 1916</bibRefCitation>
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</taxonomicName>
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, p. 336,
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<emphasis id="B9320D60E46EFFE7684B0157FD187312" bold="true" box="[566,659,1351,1372]" pageId="5" pageNumber="306">syn. nov.</emphasis>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35C82F9E46EFFE76AEA0176FB007335" box="[151,1163,1381,1403]" pageId="5" pageNumber="306" type="materials_examined">
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||
<paragraph id="8BF9D172E46EFFE76AEA0176FB007335" blockId="5.[151,1163,1350,1403]" box="[151,1163,1381,1403]" pageId="5" pageNumber="306">
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||
<typeStatus id="54FD6FD0E46EFFE76AEA0176FF437335" box="[151,200,1382,1403]" pageId="5" pageNumber="306">Type</typeStatus>
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||
species:
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||
<taxonomicName id="4C46AAF1E46EFFE76B5B0175FC047335" authority="Berlese, 1916" authorityName="Berlese" authorityYear="1916" box="[294,911,1381,1403]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="indusiatus" subGenus="Trimalaconothrus">
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<emphasis id="B9320D60E46EFFE76B5B0175FE4A7335" box="[294,449,1381,1403]" italics="true" pageId="5" pageNumber="306">Malaconothrus</emphasis>
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(
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<emphasis id="B9320D60E46EFFE76BB20175FD0F7335" box="[463,644,1381,1403]" italics="true" pageId="5" pageNumber="306">Trimalaconothrus</emphasis>
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)
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<emphasis id="B9320D60E46EFFE768E90175FD777335" box="[660,764,1381,1403]" italics="true" pageId="5" pageNumber="306">indusiatus</emphasis>
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||
<bibRefCitation id="EFD7AC83E46EFFE7697F0175FC047335" author="Berlese" box="[770,911,1381,1403]" pageId="5" pageNumber="306" refString="Berlese, A. (1916) Centuria seconda di Acari nuovi. Redia, 12, 125 - 177." type="journal article" year="1916">Berlese, 1916</bibRefCitation>
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||
</taxonomicName>
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, by original designation.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35C82F9E46EFFE76AEA01B3FD6773F6" box="[151,748,1442,1465]" pageId="5" pageNumber="306" type="reference_group">
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||
<paragraph id="8BF9D172E46EFFE76AEA01B3FD6773F6" blockId="5.[151,1051,1442,1495]" box="[151,748,1442,1465]" pageId="5" pageNumber="306">
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||
<taxonomicName id="4C46AAF1E46EFFE76AEA01B3FE2273F6" authority="Hammer, 1966" authorityName="Hammer" authorityYear="1966" box="[151,425,1442,1465]" class="Arachnida" family="Malaconothridae" genus="Zeanothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
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<emphasis id="B9320D60E46EFFE76AEA01B3FE8073F7" box="[151,267,1443,1465]" italics="true" pageId="5" pageNumber="306">Zeanothrus</emphasis>
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||
<bibRefCitation id="EFD7AC83E46EFFE76B6D01B4FE2273F6" author="Hammer" box="[272,425,1442,1465]" pageId="5" pageNumber="306" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer, 1966</bibRefCitation>
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</taxonomicName>
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, p. 20; syn.
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<bibRefCitation id="EFD7AC83E46EFFE7685901B2FD2373F6" author="Subias" box="[548,680,1442,1464]" pageId="5" pageNumber="306" refString="Subias, L. S. (2004) Listado sistematico sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) del mundo (1758 - 2002). Graellsia, 60 (supplement), 3 - 305. http: // dx. doi. org / 10.3989 / graellsia. 2004. v 60. iExtra. 218" type="journal article" year="2004">Subías, 2004</bibRefCitation>
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, p. 61.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35C82F9E46EFFE46AEA01D2FA8C769A" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306" type="materials_examined">
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||
<paragraph id="8BF9D172E46EFFE76AEA01D2FB907399" blockId="5.[151,1051,1442,1495]" box="[151,1051,1473,1495]" pageId="5" pageNumber="306">
|
||
<typeStatus id="54FD6FD0E46EFFE76AEA01D2FF437399" box="[151,200,1474,1495]" pageId="5" pageNumber="306">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="4C46AAF1E46EFFE76B5B01D1FD077399" authority="Hammer, 1966" authorityName="Hammer" authorityYear="1966" box="[294,652,1473,1495]" class="Arachnida" family="Malaconothridae" genus="Zeanothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="elegans">
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||
<emphasis id="B9320D60E46EFFE76B5B01D1FE677399" box="[294,492,1473,1495]" italics="true" pageId="5" pageNumber="306">Zeanothrus elegans</emphasis>
|
||
<bibRefCitation id="EFD7AC83E46EFFE76B8E01D2FD077399" author="Hammer" box="[499,652,1473,1495]" pageId="5" pageNumber="306" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer, 1966</bibRefCitation>
|
||
</taxonomicName>
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||
, p. 20, Fig. 24, by original designation.
|
||
</paragraph>
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||
<paragraph id="8BF9D172E46EFFE76AEA01EFFDC2705A" blockId="5.[151,1094,1534,1587]" box="[151,585,1534,1557]" pageId="5" pageNumber="306">
|
||
<emphasis id="B9320D60E46EFFE76AEA01EFFEAF705B" box="[151,292,1535,1557]" italics="true" pageId="5" pageNumber="306">Cristonothrus</emphasis>
|
||
<bibRefCitation id="EFD7AC83E46EFFE76B5701EEFE24705A" author="Subias" box="[298,431,1534,1556]" pageId="5" pageNumber="306" refString="Subias, L. S. (2004) Listado sistematico sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) del mundo (1758 - 2002). Graellsia, 60 (supplement), 3 - 305. http: // dx. doi. org / 10.3989 / graellsia. 2004. v 60. iExtra. 218" type="journal article" year="2004">Subías, 2004</bibRefCitation>
|
||
, p. 7.
|
||
<emphasis id="B9320D60E46EFFE76B9101EFFDC2705A" bold="true" box="[492,585,1535,1556]" pageId="5" pageNumber="306">syn. nov.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF9D172E46EFFE76AEA020EFBCD707D" blockId="5.[151,1094,1534,1587]" box="[151,1094,1565,1587]" pageId="5" pageNumber="306">
|
||
<typeStatus id="54FD6FD0E46EFFE76AEA020EFF43707D" box="[151,200,1566,1587]" pageId="5" pageNumber="306">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="4C46AAF1E46EFFE76B5B020DFCC0707D" authority="Subias & Sarkar, 1982" authorityName="Subias & Sarkar" authorityYear="1982" box="[294,843,1565,1587]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="pauciareolatus">
|
||
<emphasis id="B9320D60E46EFFE76B5B020DFDD5707D" box="[294,606,1565,1587]" italics="true" pageId="5" pageNumber="306">Malaconothrus pauciareolatus</emphasis>
|
||
<bibRefCitation id="EFD7AC83E46EFFE76819020DFCC0707D" author="Subias" box="[612,843,1565,1587]" pageId="5" pageNumber="306" refString="Subias, L. S. & Sarkar, S. (1982) New representatives of Nothroides Oribatids (Acarida) from India (Thrypochthoniidae and Malaconothridae). Redia, 65, 39 - 49." type="journal article" year="1982">Subías & Sarkar, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
, by original designation.
|
||
</paragraph>
|
||
<paragraph id="8BF9D172E46EFFE76AEA024AFE7871AD" blockId="5.[151,1436,1626,2019]" pageId="5" pageNumber="306">
|
||
<emphasis id="B9320D60E46EFFE76AEA024AFE42703D" bold="true" box="[151,457,1626,1651]" pageId="5" pageNumber="306">Definition and diagnosis.</emphasis>
|
||
Malaconothrid mites with the following combination of characters. Carina narrow, straight or curved, extending as far as seta
|
||
<emphasis id="B9320D60E46EFFE768FE0290FD1170D9" box="[643,666,1664,1687]" italics="true" pageId="5" pageNumber="306">ro</emphasis>
|
||
, occasionally incurved posterior of
|
||
<emphasis id="B9320D60E46EFFE76E4B0290FBC670D9" box="[1078,1101,1664,1687]" italics="true" pageId="5" pageNumber="306">ro</emphasis>
|
||
, giving anterior prodorsum a truncated appearance; never enclosing seta
|
||
<emphasis id="B9320D60E46EFFE7680402B3FD0670F5" box="[633,653,1699,1723]" italics="true" pageId="5" pageNumber="306">le</emphasis>
|
||
. Seta
|
||
<emphasis id="B9320D60E46EFFE768AC02B4FD6C70F5" box="[721,743,1700,1723]" italics="true" pageId="5" pageNumber="306">in</emphasis>
|
||
long, typically 2–3 × longer than seta
|
||
<emphasis id="B9320D60E46EFFE76EE902B4FB2670F5" box="[1172,1197,1700,1723]" italics="true" pageId="5" pageNumber="306">ex</emphasis>
|
||
, or occasionally both
|
||
<emphasis id="B9320D60E46EFFE76AEA02D8FF267091" box="[151,173,1736,1759]" italics="true" pageId="5" pageNumber="306">in</emphasis>
|
||
and
|
||
<emphasis id="B9320D60E46EFFE76A9B02D8FF747091" box="[230,255,1736,1759]" italics="true" pageId="5" pageNumber="306">ex</emphasis>
|
||
1 short, sub-equal. Caudal region of hysterosoma typically U-shaped, as broad or narrower than anterior region; notogastral margins parallel. With centrodorsal ridges and M-shaped posterior ridges, or ridges secondarily lost, as indicated by vestigial structures. Anteriolateral margin of epimeral plates I rounded or with a pronounced projecting spur. Apodeme IV narrow, bow-shaped. Genital setae 4–9 pairs. Tarsi, particularly Tarsus I, short, broad, never elongated, typically no more than 1.5–2 × longer than maximum width. Notogastral setae sub-equal in length, or with the centrodorsal series (particularly
|
||
<emphasis id="B9320D60E46EFFE76880036FFC8271D8" box="[765,777,1919,1942]" italics="true" pageId="5" pageNumber="306">c</emphasis>
|
||
1,
|
||
<emphasis id="B9320D60E46EFFE76958036DFCB871DB" box="[805,819,1917,1941]" italics="true" pageId="5" pageNumber="306">d</emphasis>
|
||
1 and
|
||
<emphasis id="B9320D60E46EFFE76900036FFC0271D8" box="[893,905,1919,1942]" italics="true" pageId="5" pageNumber="306">e</emphasis>
|
||
1) longer than others, or with
|
||
<emphasis id="B9320D60E46EFFE76E89036FFA8B71D8" box="[1268,1280,1919,1942]" italics="true" pageId="5" pageNumber="306">e</emphasis>
|
||
2 and
|
||
<emphasis id="B9320D60E46EFFE76F37036DFAD371DB" box="[1354,1368,1917,1941]" italics="true" pageId="5" pageNumber="306">h</emphasis>
|
||
2, and occasionally
|
||
<emphasis id="B9320D60E46EFFE76B5603B4FEB271F2" box="[299,313,1956,1980]" italics="true" pageId="5" pageNumber="306">h</emphasis>
|
||
1, longer than others. Body setae typically smooth, occasionally barbed. Leg setae smooth. Pre-tarsi monodactylous or tridactylous.
|
||
</paragraph>
|
||
<paragraph id="8BF9D172E46DFFE46ABA0488FA8C769A" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="307">
|
||
<taxonomicName id="4C46AAF1E46DFFE46ABA0488FEE776FE" box="[199,364,152,176]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9320D60E46DFFE46ABA0488FEE776FE" box="[199,364,152,176]" italics="true" pageId="6" pageNumber="307">Tyrphonothrus</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="4C46AAF1E46DFFE4680D0488FC9076FE" box="[624,795,152,176]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9320D60E46DFFE4680D0488FC9076FE" box="[624,795,152,176]" italics="true" pageId="6" pageNumber="307">Malaconothrus</emphasis>
|
||
</taxonomicName>
|
||
by two consistent synapomorphies: an elongated tarsus I (compare
|
||
<figureCitation id="137DCDF7E46DFFE46B7A04ACFEDF769A" box="[263,340,188,212]" captionStart="FIGURE 6" captionStartId="11.[151,250,1723,1746]" captionTargetBox="[322,1271,204,1680]" captionTargetId="figure@11.[307,1287,194,1706]" captionTargetPageId="11" captionText="FIGURE 6. Tyrphonothrus gringai sp. nov., holotype female, left legs, antiaxial view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV." httpUri="https://zenodo.org/record/222804/files/figure.png" pageId="6" pageNumber="307">Figs. 6</figureCitation>
|
||
a and 14a) and the incurved, S-shaped prodorsal carinae (compare
|
||
<figureCitation id="137DCDF7E46DFFE46E4204ACFB01769A" box="[1087,1162,188,212]" captionStart="FIGURE 3" captionStartId="8.[151,250,1405,1428]" captionTargetBox="[207,1368,381,1377]" captionTargetId="figure@8.[200,1388,370,1382]" captionTargetPageId="8" captionText="FIGURE 3. Tyrphonothrus gnammaensis sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/222801/files/figure.png" pageId="6" pageNumber="307">Figs. 3</figureCitation>
|
||
a and 12a).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C82F9E46DFFE46ABA04F0FEDE7411" pageId="6" pageNumber="307" type="discussion">
|
||
<paragraph id="8BF9D172E46DFFE46ABA04F0FC347771" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="307">
|
||
<emphasis id="B9320D60E46DFFE46ABA04F0FEB076B6" bold="true" box="[199,315,224,248]" pageId="6" pageNumber="307">Remarks.</emphasis>
|
||
Because all of the species of
|
||
<taxonomicName id="4C46AAF1E46DFFE468DB04F0FCDA76B6" box="[678,849,224,248]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9320D60E46DFFE468DB04F0FCDA76B6" box="[678,849,224,248]" italics="true" pageId="6" pageNumber="307">Malaconothrus</emphasis>
|
||
</taxonomicName>
|
||
fall into one large clade and it is not possible to distinguish discrete subgenera, it is implicit that the genera and subgenera in the synonymic list above are synonyms at the subgenus level, i.e of
|
||
<taxonomicName id="4C46AAF1E46DFFE468350538FC327771" box="[584,953,295,320]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9320D60E46DFFE468350538FD78770E" box="[584,755,296,320]" italics="true" pageId="6" pageNumber="307">Malaconothrus</emphasis>
|
||
(
|
||
<emphasis id="B9320D60E46DFFE4697E0538FC25770E" box="[771,942,296,320]" italics="true" pageId="6" pageNumber="307">Malaconothrus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF9D172E46DFFE46ABA055CFEDE7411" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="307">
|
||
<emphasis id="B9320D60E46DFFE46ABA055CFEE8772A" box="[199,355,332,356]" italics="true" pageId="6" pageNumber="307">Cristonothrus</emphasis>
|
||
<bibRefCitation id="EFD7AC83E46DFFE46B10055CFD8A772A" author="Subias" box="[365,513,332,356]" pageId="6" pageNumber="307" refString="Subias, L. S. (2004) Listado sistematico sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) del mundo (1758 - 2002). Graellsia, 60 (supplement), 3 - 305. http: // dx. doi. org / 10.3989 / graellsia. 2004. v 60. iExtra. 218" type="journal article" year="2004">Subías, 2004</bibRefCitation>
|
||
was proposed as a subgenus of
|
||
<emphasis id="B9320D60E46DFFE469FD055CFBB8772A" box="[896,1075,332,356]" italics="true" pageId="6" pageNumber="307">
|
||
<taxonomicName id="4C46AAF1E46DFFE469FD055CFBA4772A" box="[896,1071,332,356]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">Malaconothrus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
It might be considered that its synonymy with
|
||
<taxonomicName id="4C46AAF1E46DFFE46B300560FE7377C6" box="[333,504,368,392]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9320D60E46DFFE46B300560FE7377C6" box="[333,504,368,392]" italics="true" pageId="6" pageNumber="307">Malaconothrus</emphasis>
|
||
</taxonomicName>
|
||
might be weakened because the
|
||
<typeStatus id="54FD6FD0E46DFFE469120560FC1477C6" box="[879,927,368,392]" pageId="6" pageNumber="307">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C46AAF1E46DFFE46E7A0561FB5F77C6" box="[1031,1236,368,392]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="pauciareolatus">
|
||
<emphasis id="B9320D60E46DFFE46E7A0561FB5F77C6" box="[1031,1236,368,392]" italics="true" pageId="6" pageNumber="307">M. pauciareolatus</emphasis>
|
||
</taxonomicName>
|
||
was not included in the phylogenetic analysis (15 characters could not be coded). However, the reasons for the synonymy are: 1)
|
||
<emphasis id="B9320D60E46DFFE46AEA05A8FEB8779E" box="[151,307,440,464]" italics="true" pageId="6" pageNumber="307">Cristonothrus</emphasis>
|
||
was created based on only one character, the notogastral ridges, and this is not a synapomorphy within
|
||
<taxonomicName id="4C46AAF1E46DFFE46A9505CCFE1877BA" box="[232,403,476,500]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9320D60E46DFFE46A9505CCFE1877BA" box="[232,403,476,500]" italics="true" pageId="6" pageNumber="307">Malaconothrus</emphasis>
|
||
</taxonomicName>
|
||
; 2)
|
||
<emphasis id="B9320D60E46DFFE46BB805CCFDEA77BA" box="[453,609,476,500]" italics="true" pageId="6" pageNumber="307">Cristonothrus</emphasis>
|
||
does not form a discrete phylogenetic unit. Even though all except one species are contained within one of the two major clades in
|
||
<taxonomicName id="4C46AAF1E46DFFE469520610FC517456" box="[815,986,512,536]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9320D60E46DFFE469520610FC517456" box="[815,986,512,536]" italics="true" pageId="6" pageNumber="307">Malaconothrus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="137DCDF7E46DFFE4699405EFFBA67459" box="[1001,1069,511,535]" captionStart="FIGURE 2" captionStartId="3.[151,250,1886,1909]" captionTargetBox="[268,1324,208,1836]" captionTargetId="figure@3.[247,1338,200,1846]" captionTargetPageId="3" captionText="FIGURE 2. Cladogram of selected Malaconothridae showing two major clades, Tyrphonothrus and Malaconothrus. Designations of species to nominate subgenera, Trimalaconothrus (Tyrphonothrus) and Malaconothrus (Cristonothrus) follows Subías (2004). Bold type = species with complete notogastral ridges; * = species with partial ridges; M (M), M (C) = monodactylous; T (T), T (Ty) = tridactylous. Percentage probabilities of each node are indicated if other than 100." httpUri="https://zenodo.org/record/222800/files/figure.png" pageId="6" pageNumber="307">Fig. 2</figureCitation>
|
||
), that clade also contains species formerly in
|
||
<taxonomicName id="4C46AAF1E46DFFE46B480634FD387472" box="[309,691,548,572]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9320D60E46DFFE46B480634FD237472" box="[309,680,548,572]" italics="true" pageId="6" pageNumber="307">Malaconothrus (Malaconothrus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C46AAF1E46DFFE468B10634FB0F7472" box="[716,1156,548,572]" class="Arachnida" family="Malaconothridae" genus="Trimalaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9320D60E46DFFE468B10634FC1E7472" box="[716,917,548,572]" italics="true" pageId="6" pageNumber="307">Trimalaconothrus</emphasis>
|
||
(
|
||
<emphasis id="B9320D60E46DFFE469CF0634FBF07472" box="[946,1147,548,572]" italics="true" pageId="6" pageNumber="307">Trimalaconothrus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C46AAF1E46DFFE46EAE0634FEC47411" authority="(Tyrphonothrus)" baseAuthorityName="Tyrphonothrus" class="Arachnida" family="Malaconothridae" genus="Trimalaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9320D60E46DFFE46EAE0634FA177472" box="[1235,1436,548,572]" italics="true" pageId="6" pageNumber="307">Trimalaconothrus</emphasis>
|
||
(
|
||
<emphasis id="B9320D60E46DFFE46ADD0658FECF742E" box="[160,324,584,608]" italics="true" pageId="6" pageNumber="307">Tyrphonothrus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |