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<document id="8985B545B353592CAFE6459FCCE0D563" ID-DOI="10.11646/zootaxa.3990.3.1" ID-GBIF-Dataset="cf747765-b9bf-42ed-8479-35355e712db0" ID-ISSN="1175-5326" ID-Zenodo-Dep="244301" ID-ZooBank="378C0359-E0E4-4CCC-821D-649144E37A63" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461268581225" checkinUser="plazi" docAuthor="Toledo, Mario &amp; Michat, Mariano C." docDate="2015" docId="03EE87FEFFF8FFF891A6FC31FC91A56B" docLanguage="en" docName="zt03990p354.pdf" docOrigin="Zootaxa 3990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccomimus malkini Toledo &amp; Michat, 2015, sp. n." docType="treatment" docVersion="9" lastPageNumber="313" masterDocId="FFD7FF86FFF3FFF49131FFBDFFF8A03E" masterDocTitle="Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)" masterLastPageNumber="354" masterPageNumber="301" pageNumber="312" updateTime="1698611165842" updateUser="plazi">
<mods:mods id="5E0709B53DD09AF8AA7FE7F2F0BD8042" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1B83E1A2C24CD995B2D357A375B458FD">
<mods:title id="56DE3203745F7B33E95AFA80A655373F">Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)</mods:title>
</mods:titleInfo>
<mods:name id="790E10A6E625F1F590B5DC27E5EC9242" type="personal">
<mods:role id="2293D9C890897B5C5F10069DE3CE2656">
<mods:roleTerm id="7FA61F271B76DA1545AA61C98A345EDD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="61C6A8DF439A8E56E3BF4640680BDA0B">Toledo, Mario</mods:namePart>
</mods:name>
<mods:name id="1E8EBAB978C2A54579703E32264AF402" type="personal">
<mods:role id="86F4C2D7F50514187663457552ABA70F">
<mods:roleTerm id="C57390CB1A34B7DFAF3037F06AE83728">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="94AE369A584E7F24DBDE03EB9CC10EF7">Michat, Mariano C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0B89BF26BCF2F6E140FFA3FE2087ECFF">text</mods:typeOfResource>
<mods:relatedItem id="2ED7F807910470724C32DAAB6CDEFFB8" type="host">
<mods:titleInfo id="9415A6D517063A797C1327E2C3F3D7AC">
<mods:title id="A3C7A46291D7F2E722EE76ECC36D26E6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7576826B38DACC9E72AB9180B1B31D4C">
<mods:date id="D26C92B5A3BBBF52C7D47B113C6CD1B3">2015</mods:date>
<mods:detail id="A79B60B28547FCAF58AB75803D478FB7" type="volume">
<mods:number id="125BF10C4222B1E544A996CAE36E12B9">3990</mods:number>
</mods:detail>
<mods:detail id="3EEA64BE0D3B11107FEA85655137AE87" type="issue">
<mods:number id="82D2EF99B94EDA383D47275756F72685">3</mods:number>
</mods:detail>
<mods:extent id="78A66C5D5BEB1FCB493CCE4274FC7C69" unit="page">
<mods:start id="65B83ABB33ACCED03ED3B0F0B82B4FE1">301</mods:start>
<mods:end id="DDBE7DF400A7973D9F25511455B280A8">354</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6B8244E8234D2D289B37C25223D1A4EB">journal article</mods:classification>
<mods:identifier id="01149EBC63327FBFCF48A5BF2EAD8E06" type="DOI">10.11646/zootaxa.3990.3.1</mods:identifier>
<mods:identifier id="151B217E34736F3E45C5136DB1AA85EC" type="GBIF-Dataset">cf747765-b9bf-42ed-8479-35355e712db0</mods:identifier>
<mods:identifier id="9E3BF261AFBEBF53D78699B1AED127D2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="90B5F175362B7FA2660DF03B371BB6ED" type="Zenodo-Dep">244301</mods:identifier>
<mods:identifier id="1DA54506E76BE85C7BD4FB2937CE9E48" type="ZooBank">378C0359-E0E4-4CCC-821D-649144E37A63</mods:identifier>
</mods:mods>
<treatment id="03EE87FEFFF8FFF891A6FC31FC91A56B" ID-DOI="http://doi.org/10.5281/zenodo.5692042" ID-GBIF-Taxon="119638438" ID-Zenodo-Dep="5692042" LSID="urn:lsid:plazi:treatment:03EE87FEFFF8FFF891A6FC31FC91A56B" httpUri="http://treatment.plazi.org/id/03EE87FEFFF8FFF891A6FC31FC91A56B" lastPageId="12" lastPageNumber="313" pageId="11" pageNumber="312">
<subSubSection id="C35D6563FFF8FFFF91A6FC31FD5EA3F9" pageId="11" pageNumber="312" type="nomenclature">
<paragraph id="8BF836E8FFF8FFFF91A6FC31FE27A398" blockId="11.[151,678,908,967]" box="[151,479,908,934]" pageId="11" pageNumber="312">
<heading id="D0B08184FFF8FFFF91A6FC31FE27A398" bold="true" box="[151,479,908,934]" fontSize="11" level="1" pageId="11" pageNumber="312" reason="1">
<emphasis id="B933EAFAFFF8FFFF91A6FC31FE27A398" bold="true" box="[151,479,908,934]" pageId="11" pageNumber="312">
<taxonomicName id="4C474D6BFFF8FFFF91A6FC31FE6EA398" ID-CoL="8LXJY" box="[151,406,908,934]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="malkini" status="sp. nov.">
<emphasis id="B933EAFAFFF8FFFF91A6FC31FE6EA398" bold="true" box="[151,406,908,934]" italics="true" pageId="11" pageNumber="312">Laccomimus malkini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2005781FFF8FFFF90ACFC31FE27A398" box="[413,479,908,934]" pageId="11" pageNumber="312" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF836E8FFF8FFFF91A6FC13FD5EA3F9" blockId="11.[151,678,908,967]" box="[151,678,942,967]" pageId="11" pageNumber="312">
(
<figureCitation id="137C2A6DFFF8FFFF91AEFC13FF1FA3F9" box="[159,231,942,967]" captionStart="FIGURES 8 – 13" captionStartId="30.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@30.[151,1436,193,1932]" captionTargetPageId="30" captionText="FIGURES 8 – 13. SEMs of Laccomimus species, ventral view: 8) L. distinctus (male); 9) L. malkini (female); 10) L. amazonas (male); 11) L. alvarengi (female); 12) L. bordoni (male); 13) L. bolivari (female)." httpUri="https://zenodo.org/record/244303/files/figure.png" pageId="11" pageNumber="312">Figs 9</figureCitation>
; 23; 37; 60; 71; 85; 108; 117; 123; 135)
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFF8FFFF91A6FC4BFC02A685" pageId="11" pageNumber="312" type="materials_examined">
<paragraph id="8BF836E8FFF8FFFF91A6FC4BFD6AA431" blockId="11.[151,1436,1014,2011]" box="[151,658,1014,1039]" pageId="11" pageNumber="312">
<emphasis id="B933EAFAFFF8FFFF91A6FC4BFECDA431" bold="true" box="[151,309,1014,1039]" pageId="11" pageNumber="312">
<typeStatus id="54FC884AFFF8FFFF91A6FC4BFF2AA431" box="[151,210,1014,1039]" pageId="11" pageNumber="312">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3507678FFF8FFFF900DFC4BFE55A431" box="[316,429,1014,1039]" name="Colombia" pageId="11" pageNumber="312">Colombia</collectingCountry>
,
<taxonomicName id="4C474D6BFFF8FFFF9089FC4AFDC9A431" box="[440,561,1015,1039]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="354" phylum="Arthropoda" rank="species" species="amazonas">Amazonas</taxonomicName>
, Leticia.
</paragraph>
<paragraph id="8BF836E8FFF8FFFF91F6FBA7FE47A4A1" blockId="11.[151,1436,1014,2011]" pageId="11" pageNumber="312">
<emphasis id="B933EAFAFFF8FFFF91F6FBA7FE8CA40D" bold="true" box="[199,372,1050,1075]" pageId="11" pageNumber="312">
<typeStatus id="54FC884AFFF8FFFF91F6FBA7FEFAA40D" box="[199,258,1050,1075]" pageId="11" pageNumber="312">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54FC884AFFF8FFFF904BFBA7FE1BA40D" box="[378,483,1050,1075]" pageId="11" pageNumber="312" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED56AE2DFFF8FFFF931FFBA6FD7EA40D" LSID="urn:lsid:biocol.org:col:34871" box="[558,646,1051,1075]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="312">USNM</collectionCode>
): with genitalia dissected probably by Young; median lobe of aedeagus and parameres included in
<collectionCode id="ED56AE2DFFF8FFFF90FFFB82FDE7A468" box="[462,543,1087,1110]" pageId="11" pageNumber="312">DMHF</collectionCode>
on plastic label pinned under the
<typeStatus id="54FC884AFFF8FFFF929BFB83FBF4A469" box="[938,1036,1086,1111]" pageId="11" pageNumber="312" type="holotype">holotype</typeStatus>
“
<collectingCountry id="F3507678FFF8FFFF952EFB83FB6BA469" box="[1055,1171,1086,1111]" name="Colombia" pageId="11" pageNumber="312">Colombia</collectingCountry>
, Amaz. [=
<taxonomicName id="4C474D6BFFF8FFFF9427FB82FA6AA469" box="[1302,1426,1087,1111]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="354" phylum="Arthropoda" rank="species" species="amazonas">Amazonas</taxonomicName>
], Leticia,
<date id="FFF91028FFF8FFFF91C4FBDEFECBA445" box="[245,307,1123,1147]" pageId="11" pageNumber="312" value="2012-03">III.12</date>
–15.1969, P. &amp; P. Spangler” [printed], “
<typeStatus id="54FC884AFFF8FFFF93CBFBDEFC68A445" box="[762,912,1123,1147]" pageId="11" pageNumber="312" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C474D6BFFF8FFFF92ABFBDEFB7CA445" box="[922,1156,1123,1147]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="malkini">
<emphasis id="B933EAFAFFF8FFFF92ABFBDEFB7CA445" box="[922,1156,1123,1147]" italics="true" pageId="11" pageNumber="312">Laccomimus malkini</emphasis>
</taxonomicName>
Toledo &amp; Michat” [red, rectangular, printed label].
</paragraph>
<paragraph id="8BF836E8FFF8FFFF91F6FB16FC02A685" blockId="11.[151,1436,1014,2011]" pageId="11" pageNumber="312">
<typeStatus id="54FC884AFFF8FFFF91F6FB16FECCA4FD" box="[199,308,1195,1219]" pageId="11" pageNumber="312" type="paratype">Paratypes</typeStatus>
(68 exs, all bearing red, printed label “
<typeStatus id="54FC884AFFF8FFFF922BFB16FC50A4FD" box="[794,936,1195,1219]" pageId="11" pageNumber="312" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C474D6BFFF8FFFF9284FB16FB5CA4FD" box="[949,1188,1195,1219]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="malkini">
<emphasis id="B933EAFAFFF8FFFF9284FB16FB5CA4FD" box="[949,1188,1195,1219]" italics="true" pageId="11" pageNumber="312">Laccomimus malkini</emphasis>
</taxonomicName>
Toledo &amp; Michat”):
<emphasis id="B933EAFAFFF8FFFF91A6FB73FEEDA4D9" bold="true" box="[151,277,1230,1255]" pageId="11" pageNumber="312">
<collectingCountry id="F3507678FFF8FFFF91A6FB73FEE8A4D9" box="[151,272,1230,1255]" name="Colombia" pageId="11" pageNumber="312">Colombia</collectingCountry>
:
</emphasis>
same data as
<typeStatus id="54FC884AFFF8FFFF9083FB73FDECA4D9" box="[434,532,1230,1255]" pageId="11" pageNumber="312" type="holotype">holotype</typeStatus>
(28 exs
<collectionCode id="ED56AE2DFFF8FFFF9344FB72FD31A4D9" LSID="urn:lsid:biocol.org:col:34871" box="[629,713,1231,1255]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="312">USNM</collectionCode>
,
<collectionCode id="ED56AE2DFFF8FFFF93E4FB72FCF7A4D8" box="[725,783,1231,1254]" pageId="11" pageNumber="312">MTP</collectionCode>
).
<emphasis id="B933EAFAFFF8FFFF921CFB73FC73A4D9" bold="true" box="[813,907,1230,1255]" pageId="11" pageNumber="312">
<collectingCountry id="F3507678FFF8FFFF921CFB73FC7EA4D9" box="[813,902,1230,1255]" name="Bolivia" pageId="11" pageNumber="312">Bolivia</collectingCountry>
:
</emphasis>
“Beni,
<quantity id="4CBF9B0DFFF8FFFF92D3FB72FBD1A4D9" box="[994,1065,1230,1255]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="312" unit="km" value="40.0">40 km</quantity>
E San Borja, Estacion Biologica Beni, Estancia El Polvenir,
<date id="FFF91028FFF8FFFF90E6FB4FFD9FA535" box="[471,615,1266,1291]" pageId="11" pageNumber="312" value="1987-09-06" valueMax="1987-09-08" valueMin="1987-09-06">6–8.IX.1987</date>
, W.E. Steiner”, “At black light: open grass savanna and marsh” (2 exs
<collectionCode id="ED56AE2DFFF8FFFF91A6FAAAFF15A511" LSID="urn:lsid:biocol.org:col:34871" box="[151,237,1303,1327]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="312">USNM</collectionCode>
).
<emphasis id="B933EAFAFFF8FFFF9030FAABFEACA511" bold="true" box="[257,340,1302,1327]" pageId="11" pageNumber="312">
<collectingCountry id="F3507678FFF8FFFF9030FAABFEB7A511" box="[257,335,1302,1327]" name="Brazil" pageId="11" pageNumber="312">Brazil</collectingCountry>
:
</emphasis>
“Sao Paulo, Piracicaba, blacklight,
<date id="FFF91028FFF8FFFF93C2FAABFC92A511" box="[755,874,1302,1327]" pageId="11" pageNumber="312" value="1966-02-12">12.II.1966</date>
, C.A. Triplehorn” (4 exs
<collectionCode id="ED56AE2DFFF8FFFF95A3FAAAFB10A511" LSID="urn:lsid:biocol.org:col:34871" box="[1170,1256,1303,1327]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="312">USNM</collectionCode>
); “Para, Belem,
<date id="FFF91028FFF8FFFF91A6FA87FEDAA56D" box="[151,290,1338,1363]" pageId="11" pageNumber="312" value="1969-07-10">
10.
<collectionCode id="ED56AE2DFFF8FFFF9186FA87FF1BA56D" box="[183,227,1338,1363]" pageId="11" pageNumber="312">VII</collectionCode>
.1969
</date>
, P. &amp; P. Spangler” (1 ex.
<collectionCode id="ED56AE2DFFF8FFFF936FFA86FD4CA56D" LSID="urn:lsid:biocol.org:col:34871" box="[606,692,1339,1363]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="312">USNM</collectionCode>
).
<emphasis id="B933EAFAFFF8FFFF93FAFA87FCC3A56D" bold="true" box="[715,827,1338,1363]" pageId="11" pageNumber="312">
<collectingCountry id="F3507678FFF8FFFF93FAFA87FCCEA56D" box="[715,822,1338,1363]" name="Ecuador" pageId="11" pageNumber="312">Ecuador</collectingCountry>
:
</emphasis>
“Napo, Lago Agrio (
<quantity id="4CBF9B0DFFF8FFFF950CFA86FB84A56D" box="[1085,1148,1338,1363]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="312" unit="km" value="2.0">2 km</quantity>
N),
<date id="FFF91028FFF8FFFF9586FA87FAC1A56D" box="[1207,1337,1338,1363]" pageId="11" pageNumber="312" value="1975-09-26">26.IX.1975</date>
, Andrea Langley” (3 exs
<collectionCode id="ED56AE2DFFF8FFFF9066FAE2FE55A549" LSID="urn:lsid:biocol.org:col:34871" box="[343,429,1375,1399]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="312">USNM</collectionCode>
); idem except for “
<date id="FFF91028FFF8FFFF93BBFAE3FCD8A549" box="[650,800,1374,1399]" pageId="11" pageNumber="312" value="1975-08-16">16.VIII.1975</date>
, blacklight” (1 ex.
<collectionCode id="ED56AE2DFFF8FFFF92CFFAE2FBACA549" LSID="urn:lsid:biocol.org:col:34871" box="[1022,1108,1375,1399]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="312">USNM</collectionCode>
).
<emphasis id="B933EAFAFFF8FFFF9559FAE3FB1CA549" bold="true" box="[1128,1252,1374,1399]" pageId="11" pageNumber="312">
<collectingCountry id="F3507678FFF8FFFF9559FAE3FB27A549" box="[1128,1247,1374,1399]" name="Paraguay" pageId="11" pageNumber="312">Paraguay</collectingCountry>
:
</emphasis>
“Jejuí-Mí, light trap,
<date id="FFF91028FFF8FFFF91E1FA3EFEA3A5A5" box="[208,347,1411,1435]" pageId="11" pageNumber="312" value="2003-12-19">19.XII.2003</date>
, leg. M.C. Michat” (5 exs
<collectionCode id="ED56AE2DFFF8FFFF93B2FA3EFD27A5A5" box="[643,735,1411,1435]" httpUri="http://grbio.org/cool/654h-avhc" name="Museo de Historia Natural de la Ciudad de Mexico" pageId="11" pageNumber="312">MCMC</collectionCode>
); idem “
<date id="FFF91028FFF8FFFF920AFA3EFC3FA5A5" box="[827,967,1411,1435]" pageId="11" pageNumber="312" value="2003-12-20">20.XII.2003</date>
” (2 exs
<collectionCode id="ED56AE2DFFF8FFFF9513FA3EFB87A5A5" box="[1058,1151,1411,1435]" httpUri="http://grbio.org/cool/654h-avhc" name="Museo de Historia Natural de la Ciudad de Mexico" pageId="11" pageNumber="312">MCMC</collectionCode>
).
<emphasis id="B933EAFAFFF8FFFF95A1FA3FFB24A5A5" bold="true" box="[1168,1244,1410,1435]" pageId="11" pageNumber="312">
<collectingCountry id="F3507678FFF8FFFF95A1FA3FFB28A5A5" box="[1168,1232,1410,1435]" name="Peru" pageId="11" pageNumber="312">Perú</collectingCountry>
:
</emphasis>
“Madre de Dios, Rio Tambopata Res, 30 air km SW Pto-Maldonado,
<quantity id="4CBF9B0DFFF8FFFF93ECFA1AFCDDA580" box="[733,805,1447,1471]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="11" pageNumber="312" unit="m" value="290.0">290 m</quantity>
,
<date id="FFF91028FFF8FFFF9200FA1BFC39A581" box="[817,961,1446,1471]" pageId="11" pageNumber="312" value="1979-11-02" valueMax="1979-11-05" valueMin="1979-11-02">2–5.XI.1979</date>
, J.B. Heppner, subtropical moist forest” (8 exs
<collectionCode id="ED56AE2DFFF8FFFF91FAFA76FED9A5DD" LSID="urn:lsid:biocol.org:col:34871" box="[203,289,1483,1507]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="312">USNM</collectionCode>
).
<emphasis id="B933EAFAFFF8FFFF900AFA77FE41A5DD" bold="true" box="[315,441,1482,1507]" pageId="11" pageNumber="312">
<collectingCountry id="F3507678FFF8FFFF900AFA77FE4CA5DD" box="[315,436,1482,1507]" name="Suriname" pageId="11" pageNumber="312">Suriname</collectingCountry>
:
</emphasis>
“Langaman, Kondre-Marowijne distr.,
<date id="FFF91028FFF8FFFF92A4FA77FBF6A5DD" box="[917,1038,1482,1507]" pageId="11" pageNumber="312" value="1963-10-25">25.X.1963</date>
, Borys Malkin” (5 exs
<collectionCode id="ED56AE2DFFF8FFFF940BFA76FA68A5DD" LSID="urn:lsid:biocol.org:col:34871" box="[1338,1424,1483,1507]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="312">USNM</collectionCode>
); “Zanderijsavanne, 1st tributary of Colakreek,
<geoCoordinate id="EE73502FFFF8FFFF93A7FA53FD00A639" box="[662,760,1518,1543]" direction="west" orientation="longitude" pageId="11" pageNumber="312" precision="925" value="-55.233334">55°14'W</geoCoordinate>
,
<geoCoordinate id="EE73502FFFF8FFFF9232FA53FC9AA639" box="[771,866,1518,1543]" direction="north" orientation="latitude" pageId="11" pageNumber="312" precision="925" value="5.45">05°27'N</geoCoordinate>
,
<date id="FFF91028FFF8FFFF925CFA53FC19A639" box="[877,993,1518,1543]" pageId="11" pageNumber="312" value="1969-09-04">4.IX.1969</date>
, leg N. Nieser (SN 114)” (1 ex.
<collectionCode id="ED56AE2DFFF8FFFF9472FA52FA68A639" box="[1347,1424,1519,1543]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="312">NMW</collectionCode>
); “Paramaribo, Guest House of Museum, at light,
<date id="FFF91028FFF8FFFF93F6F9AFFC52A615" box="[711,938,1554,1579]" pageId="11" pageNumber="312" value="1969-08-20" valueMax="1969-09-23" valueMin="1969-08-20">20.VIII–23.IX.1969</date>
, leg N. Nieser” (2 exs
<collectionCode id="ED56AE2DFFF8FFFF9588F9AEFAFEA615" box="[1209,1286,1555,1579]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="312">NMW</collectionCode>
).
<emphasis id="B933EAFAFFF8FFFF942BF9AFFA62A615" bold="true" box="[1306,1434,1554,1579]" pageId="11" pageNumber="312">
<collectingCountry id="F3507678FFF8FFFF942BF9AFFA68A615" box="[1306,1424,1554,1579]" name="Venezuela" pageId="11" pageNumber="312">Venezuela</collectingCountry>
:
</emphasis>
“Guarico,
<quantity id="4CBF9B0DFFF8FFFF903EF98AFEA1A671" box="[271,345,1590,1615]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="312" unit="km" value="12.0">12 km</quantity>
S Calabozo,
<date id="FFF91028FFF8FFFF90C9F98BFDE4A671" box="[504,540,1590,1615]" pageId="11" pageNumber="312">II.6</date>
–12.1969, P. &amp; P. Spangler” (3 exs
<collectionCode id="ED56AE2DFFF8FFFF92F3F98AFBE0A671" LSID="urn:lsid:biocol.org:col:34871" box="[962,1048,1591,1615]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="312">USNM</collectionCode>
); idem “
<quantity id="4CBF9B0DFFF8FFFF954DF98BFB3FA671" box="[1148,1223,1590,1615]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="312" unit="km" value="15.0">15 km</quantity>
S Calabozo,
<date id="FFF91028FFF8FFFF9457F98AFA74A671" box="[1382,1420,1591,1615]" pageId="11" pageNumber="312">II.9</date>
– 13.1969” (1 ex.
<collectionCode id="ED56AE2DFFF8FFFF907FF9E6FE5CA64D" LSID="urn:lsid:biocol.org:col:34871" box="[334,420,1627,1651]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="312">USNM</collectionCode>
); “Bolivar State,
<geoCoordinate id="EE73502FFFF8FFFF9356F9E6FD0FA64D" box="[615,759,1627,1651]" direction="north" orientation="latitude" pageId="11" pageNumber="312" precision="1" value="4.6979666">04°41.878'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFF8FFFF9232F9E7FC6FA64D" box="[771,919,1626,1651]" direction="west" orientation="longitude" pageId="11" pageNumber="312" precision="1" value="-61.070766">61°04.246'W</geoCoordinate>
,
<quantity id="4CBF9B0DFFF8FFFF9292F9E7FC15A64C" box="[931,1005,1626,1651]" metricMagnitude="2" metricUnit="m" metricValue="8.15" pageId="11" pageNumber="312" unit="m" value="815.0">815 m</quantity>
, Gran Sabana, N
<collectingCountry id="F3507678FFF8FFFF9584F9E6FAC6A64D" box="[1205,1342,1626,1651]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="11" pageNumber="312">Santa Elena</collectingCountry>
, marshy area long Rt. 10,
<date id="FFF91028FFF8FFFF906BF9C2FE1EA6A9" box="[346,486,1663,1687]" pageId="11" pageNumber="312" value="2010-07-17">
17.
<collectionCode id="ED56AE2DFFF8FFFF9049F9C2FE5BA6A9" box="[376,419,1663,1687]" pageId="11" pageNumber="312">VII</collectionCode>
.2010
</date>
, leg. Short, Tellez &amp; Camacho, VZ10-0717-03A” (4 exs
<collectionCode id="ED56AE2DFFF8FFFF955AF9C2FB51A6A9" box="[1131,1193,1663,1687]" pageId="11" pageNumber="312">VAIP</collectionCode>
– each specimen with additional label
<collectionCode id="ED56AE2DFFF8FFFF907AF91EFE0DA684" box="[331,501,1699,1722]" pageId="11" pageNumber="312">KUNHM-ENT</collectionCode>
, with its own barcode and catalogue number).
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFF8FFFF91F6F97BFD83A719" pageId="11" pageNumber="312" type="diagnosis">
<paragraph id="8BF836E8FFF8FFFF91F6F97BFD83A719" blockId="11.[151,1436,1014,2011]" pageId="11" pageNumber="312">
<emphasis id="B933EAFAFFF8FFFF91F6F97BFEB9A6E1" bold="true" box="[199,321,1734,1759]" pageId="11" pageNumber="312">Diagnosis.</emphasis>
Easily recognised by the combination of (usually) uniformly brown elytra and bright pale pronotum, along with male (scimitar-shaped outer foreclaw, characteristic bisinuate left paramere) and female (two-toothed gonocoxal blades) characters.
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFF8FFF891F6F88FFE7FA2CE" lastPageId="12" lastPageNumber="313" pageId="11" pageNumber="312" type="description">
<paragraph id="8BF836E8FFF8FFFF91F6F88FFBE7A7AD" blockId="11.[151,1436,1014,2011]" pageId="11" pageNumber="312">
<emphasis id="B933EAFAFFF8FFFF91F6F88FFEA1A775" bold="true" box="[199,345,1842,1867]" pageId="11" pageNumber="312">Description.</emphasis>
<emphasis id="B933EAFAFFF8FFFF9051F88EFE41A775" box="[352,441,1843,1867]" italics="true" pageId="11" pageNumber="312">Habitus</emphasis>
(
<figureCitation id="137C2A6DFFF8FFFF90FBF88FFDE7A775" box="[458,543,1842,1867]" captionStart="FIGURES 59 – 66" captionStartId="38.[151,264,1953,1975]" captionTargetBox="[161,1426,197,1927]" captionTargetId="figure@38.[151,1435,193,1932]" captionTargetPageId="38" captionText="FIGURES 59 – 66. Subschematic body shape and colour patterns in Laccomimus species: 59) L. distinctus; 60) L. malkini, a) typical specimen from Colombia, b) specimen from Piracicaba, Brazil; 61) L. variegatus (Perú); 62) L. amazonas; 63) L. pumilio (Florida, USA); 64) L. alvarengi, a – b) variation in colouration patterns; 65) L. bordoni; 66) L. spangleri." httpUri="https://zenodo.org/record/244311/files/figure.png" pageId="11" pageNumber="312">Fig. 60</figureCitation>
). TL: 2.0–
<quantity id="4CBF9B0DFFF8FFFF93A9F88EFD09A774" box="[664,753,1843,1867]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="11" pageNumber="312" unit="mm" value="2.3">2.3 mm</quantity>
; MW: 1.0–
<quantity id="4CBF9B0DFFF8FFFF9243F88EFC30A774" box="[882,968,1843,1867]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="312" unit="mm" value="1.2">1.2 mm</quantity>
(
<typeStatus id="54FC884AFFF8FFFF92E9F88FFBB8A775" box="[984,1088,1842,1867]" pageId="11" pageNumber="312" type="holotype">holotype</typeStatus>
: TL:
<quantity id="4CBF9B0DFFF8FFFF954CF88FFB1EA774" box="[1149,1254,1842,1867]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="11" pageNumber="312" unit="mm" value="2.05">2.05 mm</quantity>
; MW:
<quantity id="4CBF9B0DFFF8FFFF9405F88EFA68A775" box="[1332,1424,1843,1867]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="312" unit="mm" value="1.1">1.1 mm</quantity>
); A-TL:
<quantity id="4CBF9B0DFFF8FFFF91D6F8EAFEB9A750" box="[231,321,1879,1903]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="312" unit="mm" value="2.1">2.1 mm</quantity>
; A-MW:
<quantity id="4CBF9B0DFFF8FFFF909FF8EAFDFEA750" box="[430,518,1879,1903]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="312" unit="mm" value="1.2">1.2 mm</quantity>
(NUM: 36 exs); almost regularly oval, gradually narrowed backward; elytral tip almost truncate. Glossy to almost dull; iridescence on elytra more or less visible.
</paragraph>
<paragraph id="8BF836E8FFF8FFF891F6F823FD15A0EB" blockId="11.[151,1436,1014,2011]" lastBlockId="12.[151,1437,151,1365]" lastPageId="12" lastPageNumber="313" pageId="11" pageNumber="312">
<emphasis id="B933EAFAFFF8FFFF91F6F823FEEDA788" box="[199,277,1950,1974]" italics="true" pageId="11" pageNumber="312">Colour</emphasis>
. Head chestnut brown, gradually paler on front and clypeus; pronotum testaceous, with darker band along hind margin. Elytra uniformly chestnut brown, rarely with more or less defined paler markings on subbasal and preapical position. Underside yellowish, at most slightly darkened on sutures. Antennae and mouthparts testaceous. Legs testaceous, metatarsi slightly darker.
</paragraph>
<paragraph id="8BF836E8FFFFFFF891F6FF5DFAD6A1B6" blockId="12.[151,1437,151,1365]" pageId="12" pageNumber="313">
<emphasis id="B933EAFAFFFFFFF891F6FF5DFE1DA0C6" box="[199,485,224,248]" italics="true" pageId="12" pageNumber="313">Structures and sculpture</emphasis>
. Head smooth, shiny, with quite impressed microreticulation of rounded cells. Pronotum shiny, often with slight iridescence. Few scattered punctures only along sides. Elytra with no or hardly visible subapical sinuation; shiny to almost dull, with iridescence of variable intensity; microreticulation quite impressed, accompanied by subtle scratches and scattered, poorly visible punctures. Prosternal process (
<figureCitation id="137C2A6DFFFFFFF89471FEF1FA6CA15A" box="[1344,1428,332,357]" captionStart="FIGURES 20 – 25" captionStartId="32.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@32.[151,1436,193,1932]" captionTargetPageId="32" captionText="FIGURES 20 – 25. SEMs of head, prosternum and prosternal process in Laccomimus species: 20) L. distinctus (male), ventral view, 21) idem, at higher magnification; 22) idem, oblique lateral view; 23) L. malkini (female) ventral view; 24) L. amazonas (male), ventral view (arrow indicates protruding prosternum); 25) idem, at higher magnification." httpUri="https://zenodo.org/record/244305/files/figure.png" pageId="12" pageNumber="313">Fig. 23</figureCitation>
) rounded at tip, slightly elevated. Lobes of metacoxal processes regularly rounded in both sexes (
<figureCitation id="137C2A6DFFFFFFF895FCFED2FADAA1B6" box="[1229,1314,367,392]" captionStart="FIGURES 32 – 37" captionStartId="34.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@34.[151,1436,193,1932]" captionTargetPageId="34" captionText="FIGURES 32 – 37. SEMs of metacoxal processes in Laccomimus species, ventral view: 32) L. distinctus (male); 33 – 36) L. distinctus (females), variability in development of apical angulation in different specimens; 37) L. malkini (female)." httpUri="https://zenodo.org/record/244307/files/figure.png" pageId="12" pageNumber="313">Fig. 37</figureCitation>
).
</paragraph>
<paragraph id="8BF836E8FFFFFFF891F6FE29FE41A25E" blockId="12.[151,1437,151,1365]" pageId="12" pageNumber="313">
<emphasis id="B933EAFAFFFFFFF891F6FE29FEF9A192" box="[199,257,404,428]" italics="true" pageId="12" pageNumber="313">Male</emphasis>
. Last abdominal ventrite tectiform, with antero-lateral sides visibly sinuate. Outer claw of forelegs scimitar-shaped (
<figureCitation id="137C2A6DFFFFFFF89066FE0AFE54A1EE" box="[343,428,439,464]" captionStart="FIGURES 67 – 83" captionStartId="39.[151,264,1798,1820]" captionTargetBox="[159,1430,201,1748]" captionTargetId="figure@39.[151,1436,193,1777]" captionTargetPageId="39" captionText="FIGURES 67 – 83. Prosternal process and claws in Laccomimus species. 67 – 69) P rosternal process: 67) L. pumilio (Florida, USA); 68) L. pumilio (Cuba); 69) L. alvarengi (Perú). 70 – 83) M ale left protarsal claws: 70) L. bordoni (Paraguay); 71) L. malkini (Madre de Dios, Perú); 72) L. spangleri (holotype); 73) L. distinctus (holotype); 74) L. bolivari (Trinidad); 75) L. amazonas (holotype); 76) L. variegatus (Colonia Calleria, Perú); 77) L. alvarengi (holotype); 78) L. alvarengi (Loreto, Perú); 79) L. alvarengi (Guayaramerin, Bolivia); 80) L. alvarengi (Esmeraldas, Ecuador); 81) L. alvarengi (Albrook, Panamá); 82) L. pumilio (Cayo District, Belize); 83) L. pumilio (Cayamas, Cuba)." httpUri="https://zenodo.org/record/244312/files/figure.png" pageId="12" pageNumber="313">Fig. 71</figureCitation>
). Median lobe of aedeagus, in left lateral view (
<figureCitation id="137C2A6DFFFFFFF892F9FE0AFBE3A1EE" box="[968,1051,439,464]" captionStart="FIGURES 84 – 87" captionStartId="40.[151,264,1915,1937]" captionTargetBox="[169,1412,201,1889]" captionTargetId="figure@40.[160,1420,193,1898]" captionTargetPageId="40" captionText="FIGURES 84 – 87. Male genitalia in Laccomimus species, left lateral view: 84) L. distinctus, a) median lobe, b) left paramere, c) right paramere; 85) L. malkini (Loreto, Perú), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere, e) magnification of apex of median lobe (specimen from Piracicaba, Brazil); 86) L. variegatus, a) median lobe, b) left paramere, c) right paramere; 87) L. spinosus, a) median lobe, b) left paramere." httpUri="https://zenodo.org/record/244313/files/figure.png" pageId="12" pageNumber="313">Fig. 85</figureCitation>
a), slender; base and distal portion forming an angle of about 160°; distal portion straight, gradually narrowed to apex, ending in small button-like expansion, quite variable in shape. In dorsal view (
<figureCitation id="137C2A6DFFFFFFF893FAFE42FCD2A226" box="[715,810,511,536]" captionStart="FIGURES 107 – 115" captionStartId="46.[151,264,1951,1973]" captionTargetBox="[184,1390,196,1925]" captionTargetId="figure@46.[170,1417,193,1930]" captionTargetPageId="46" captionText="FIGURES 107 – 115. Median lobe of aedeagus in Laccomimus species, dorsal view: 107) L. distinctus; 108) L. malkini; 109) L. variegatus; 110) L. amazonas; 111) L. alvarengi; 112) L. improvidus; 113) L. bordoni; 114) L. spangleri; 115) L. bolivari." httpUri="https://zenodo.org/record/244319/files/figure.png" pageId="12" pageNumber="313">Fig. 108</figureCitation>
) slender and acuminate, gradually tapering from base to tip; dorsal lamina strongly sinuate close to base; ejaculator groove narrow. Left paramere (
<figureCitation id="137C2A6DFFFFFFF895B2FD99FB2CA203" box="[1155,1236,548,573]" captionStart="FIGURES 84 – 87" captionStartId="40.[151,264,1915,1937]" captionTargetBox="[169,1412,201,1889]" captionTargetId="figure@40.[160,1420,193,1898]" captionTargetPageId="40" captionText="FIGURES 84 – 87. Male genitalia in Laccomimus species, left lateral view: 84) L. distinctus, a) median lobe, b) left paramere, c) right paramere; 85) L. malkini (Loreto, Perú), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere, e) magnification of apex of median lobe (specimen from Piracicaba, Brazil); 86) L. variegatus, a) median lobe, b) left paramere, c) right paramere; 87) L. spinosus, a) median lobe, b) left paramere." httpUri="https://zenodo.org/record/244313/files/figure.png" pageId="12" pageNumber="313">Fig. 85</figureCitation>
c) straight, visibly bi-sinuate on ventral side.
</paragraph>
<paragraph id="8BF836E8FFFFFFF891F6FDD1FE7FA2CE" blockId="12.[151,1437,151,1365]" pageId="12" pageNumber="313">
<emphasis id="B933EAFAFFFFFFF891F6FDD1FEE3A2BA" box="[199,283,620,644]" italics="true" pageId="12" pageNumber="313">Female</emphasis>
. Elytra and pronotum often slightly duller than in males. Last abdominal ventrite triangular, narrowly rounded at apex. Lateral tergites similar to
<taxonomicName id="4C474D6BFFFFFFF893B0FD2CFCF3A296" box="[641,779,656,680]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B933EAFAFFFFFFF893B0FD2CFCF3A296" box="[641,779,656,680]" italics="true" pageId="12" pageNumber="313">L. distinctus</emphasis>
</taxonomicName>
: wide and sclerotized. Gonocoxal blades (
<figureCitation id="137C2A6DFFFFFFF895DFFD32FAB5A296" box="[1262,1357,655,680]" captionStart="FIGURES 116 – 121" captionStartId="47.[151,264,1829,1851]" captionTargetBox="[229,1344,199,1783]" captionTargetId="figure@47.[198,1370,193,1789]" captionTargetPageId="47" captionText="FIGURES 116 – 121. Female genitalia and cladogram. 116 – 120) Female genitalia in Laccomimus species, left lateral view: 116) L. distinctus (Paraguay), a) left gonocoxa, b) ramus; 117) L. malkini (Perú), a) left gonocoxa, b) ramus; 118) L. alvarengi (Brazil), a) left gonocoxa, b) ramus; 119) L. variegatus (Perú), a) left gonocoxa, b) ramus; 120) L. bordoni (Bolivia), ramus (arrow indicates lateral proximal process of ramus). 121) Single most parsimonious cladogram rooted between ingroup and outgroup, with character changes mapped for each clade. Solid rectangles indicate unique character state transformations; open rectangles indicate homoplasious character state transformations. Support values are indicated on each branch as Bremer / Bootstrap." httpUri="https://zenodo.org/record/244320/files/figure.png" pageId="12" pageNumber="313">Fig. 117</figureCitation>
a) with second denticle at about distal 4/5. Ramus also distinctive (
<figureCitation id="137C2A6DFFFFFFF89275FD09FC5EA2F3" box="[836,934,692,717]" captionStart="FIGURES 116 – 121" captionStartId="47.[151,264,1829,1851]" captionTargetBox="[229,1344,199,1783]" captionTargetId="figure@47.[198,1370,193,1789]" captionTargetPageId="47" captionText="FIGURES 116 – 121. Female genitalia and cladogram. 116 – 120) Female genitalia in Laccomimus species, left lateral view: 116) L. distinctus (Paraguay), a) left gonocoxa, b) ramus; 117) L. malkini (Perú), a) left gonocoxa, b) ramus; 118) L. alvarengi (Brazil), a) left gonocoxa, b) ramus; 119) L. variegatus (Perú), a) left gonocoxa, b) ramus; 120) L. bordoni (Bolivia), ramus (arrow indicates lateral proximal process of ramus). 121) Single most parsimonious cladogram rooted between ingroup and outgroup, with character changes mapped for each clade. Solid rectangles indicate unique character state transformations; open rectangles indicate homoplasious character state transformations. Support values are indicated on each branch as Bremer / Bootstrap." httpUri="https://zenodo.org/record/244320/files/figure.png" pageId="12" pageNumber="313">Fig. 117</figureCitation>
b), with numerous densely distributed small teeth of uniform size.
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFFFFFF891F6FD41FE97A363" pageId="12" pageNumber="313" type="distribution">
<paragraph id="8BF836E8FFFFFFF891F6FD41FAD9A32A" blockId="12.[151,1437,151,1365]" box="[199,1313,764,789]" pageId="12" pageNumber="313">
<emphasis id="B933EAFAFFFFFFF891F6FD41FE98A32B" bold="true" box="[199,352,764,789]" pageId="12" pageNumber="313">Distribution.</emphasis>
<collectingCountry id="F3507678FFFFFFF89056FD41FE44A32B" box="[359,444,764,789]" name="Bolivia" pageId="12" pageNumber="313">Bolivia</collectingCountry>
,
<collectingCountry id="F3507678FFFFFFF890F6FD41FDF6A32B" box="[455,526,764,789]" name="Brazil" pageId="12" pageNumber="313">Brazil</collectingCountry>
,
<collectingCountry id="F3507678FFFFFFF8932BFD41FD73A32B" box="[538,651,764,789]" name="Colombia" pageId="12" pageNumber="313">Colombia</collectingCountry>
,
<collectingCountry id="F3507678FFFFFFF893A7FD41FD0DA32B" box="[662,757,764,789]" name="Ecuador" pageId="12" pageNumber="313">Ecuador</collectingCountry>
,
<collectingCountry id="F3507678FFFFFFF89231FD40FC91A32B" box="[768,873,765,789]" name="Paraguay" pageId="12" pageNumber="313">Paraguay</collectingCountry>
,
<collectingCountry id="F3507678FFFFFFF89244FD41FC53A32A" box="[885,939,764,788]" name="Peru" pageId="12" pageNumber="313">Perú</collectingCountry>
,
<collectingCountry id="F3507678FFFFFFF89287FD41FBDCA32B" box="[950,1060,764,789]" name="Suriname" pageId="12" pageNumber="313">Suriname</collectingCountry>
,
<collectingCountry id="F3507678FFFFFFF8951EFD41FB5AA32B" box="[1071,1186,764,789]" name="Venezuela" pageId="12" pageNumber="313">Venezuela</collectingCountry>
(
<figureCitation id="137C2A6DFFFFFFF89583FD41FAECA32A" box="[1202,1300,764,789]" captionStart="FIGURES 122 – 125" captionStartId="48.[151,264,1871,1893]" captionTargetBox="[157,1432,197,1844]" captionTargetId="figure@48.[151,1436,193,1850]" captionTargetPageId="48" captionText="FIGURES 122 – 125. Distribution maps of Laccomimus species: 122) L. distinctus; 123) L. malkini; 124) L. variegatus (open circles), L. spinosus (single pentagon in Brazil), L. amazonas (single star in Venezuela); 125) L. alvarengi in South America." httpUri="https://zenodo.org/record/244321/files/figure.png" pageId="12" pageNumber="313">Fig. 123</figureCitation>
).
</paragraph>
<paragraph id="8BF836E8FFFFFFF891F6FCA2FE97A363" blockId="12.[151,1437,151,1365]" pageId="12" pageNumber="313">
<emphasis id="B933EAFAFFFFFFF891F6FCA2FED2A306" bold="true" box="[199,298,799,824]" pageId="12" pageNumber="313">Habitat.</emphasis>
In
<collectingCountry id="F3507678FFFFFFF89068FC9DFE1CA306" box="[345,484,799,824]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="12" pageNumber="313">Santa Elena</collectingCountry>
(
<collectingCountry id="F3507678FFFFFFF890C6FCA2FD96A306" box="[503,622,799,824]" name="Venezuela" pageId="12" pageNumber="313">Venezuela</collectingCountry>
) some specimens were collected in a marshy area exposed to direct sunlight (Fig. 135).
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFFFFFF891F6FCDAFD65A49E" pageId="12" pageNumber="313" type="etymology">
<paragraph id="8BF836E8FFFFFFF891F6FCDAFBDBA39A" blockId="12.[151,1437,151,1365]" pageId="12" pageNumber="313">
<emphasis id="B933EAFAFFFFFFF891F6FCDAFEB6A3BE" bold="true" box="[199,334,871,896]" pageId="12" pageNumber="313">Etymology.</emphasis>
Name given
<emphasis id="B933EAFAFFFFFFF890D6FCD5FDB6A3BE" box="[487,590,872,896]" italics="true" pageId="12" pageNumber="313">in litteris</emphasis>
by Frank N. Young and Paul J. Spangler, in honour of the collector Borys Malkin (cf.
<bibRefCitation id="EFD64B19FFFFFFF8902DFC30FE50A39A" author="Young" box="[284,424,908,933]" pageId="12" pageNumber="313" refString="Young, F. N. (1974) Review of the predaceous water beetles of genus Anodocheilus (Coleoptera: Dytiscidae: Hydroporinae). Occasional Papers of the Museum of Zoology of the University of Michigan, 670, 1 - 28." type="journal article" year="1974">Young 1974</bibRefCitation>
: 2, 3). The specific epithet is a noun in the genitive case.
</paragraph>
<paragraph id="8BF836E8FFFFFFF891F6FC12FD65A49E" blockId="12.[151,1437,151,1365]" pageId="12" pageNumber="313">
<emphasis id="B933EAFAFFFFFFF891F6FC12FE42A3F6" bold="true" box="[199,442,943,968]" pageId="12" pageNumber="313">Notes on variability.</emphasis>
Specimens from most localities have uniformly brown elytra. Paler markings, variably developed but never sharply defined, have been observed in most specimens from
<collectingCountry id="F3507678FFFFFFF8957AFC68FB4DA3D3" box="[1099,1205,981,1005]" name="Paraguay" pageId="12" pageNumber="313">Paraguay</collectingCountry>
. The few exemplars seen from
<collectingCountry id="F3507678FFFFFFF89020FC4AFEADA42E" box="[273,341,1015,1040]" name="Brazil" pageId="12" pageNumber="313">Brazil</collectingCountry>
(Piracicaba) share peculiar characters: larger size (TL:
<quantity id="4CBF9B0DFFFFFFF892E4FC4AFB99A431" box="[981,1121,1015,1040]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="12" pageNumber="313" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
; MW:
<quantity id="4CBF9B0DFFFFFFF89583FC4AFAB5A42E" box="[1202,1357,1015,1040]" metricMagnitude="-3" metricUnit="m" metricValue="1.225" metricValueMax="1.25" metricValueMin="1.2" pageId="12" pageNumber="313" unit="mm" value="1.225" valueMax="1.25" valueMin="1.2">1.2–1.25 mm</quantity>
), more elongate body shape (with apical sinuation of elytra missing or almost so), pale patterns on the elytra well developed and clear (
<figureCitation id="137C2A6DFFFFFFF890B9FB82FE23A466" box="[392,475,1087,1112]" captionStart="FIGURES 59 – 66" captionStartId="38.[151,264,1953,1975]" captionTargetBox="[161,1426,197,1927]" captionTargetId="figure@38.[151,1435,193,1932]" captionTargetPageId="38" captionText="FIGURES 59 – 66. Subschematic body shape and colour patterns in Laccomimus species: 59) L. distinctus; 60) L. malkini, a) typical specimen from Colombia, b) specimen from Piracicaba, Brazil; 61) L. variegatus (Perú); 62) L. amazonas; 63) L. pumilio (Florida, USA); 64) L. alvarengi, a – b) variation in colouration patterns; 65) L. bordoni; 66) L. spangleri." httpUri="https://zenodo.org/record/244311/files/figure.png" pageId="12" pageNumber="313">Fig. 60</figureCitation>
b), and a button-like apical expansion of the median lobe projecting posteriorly into a small, acute process (
<figureCitation id="137C2A6DFFFFFFF890B9FBD9FE23A443" box="[392,475,1124,1149]" captionStart="FIGURES 84 – 87" captionStartId="40.[151,264,1915,1937]" captionTargetBox="[169,1412,201,1889]" captionTargetId="figure@40.[160,1420,193,1898]" captionTargetPageId="40" captionText="FIGURES 84 – 87. Male genitalia in Laccomimus species, left lateral view: 84) L. distinctus, a) median lobe, b) left paramere, c) right paramere; 85) L. malkini (Loreto, Perú), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere, e) magnification of apex of median lobe (specimen from Piracicaba, Brazil); 86) L. variegatus, a) median lobe, b) left paramere, c) right paramere; 87) L. spinosus, a) median lobe, b) left paramere." httpUri="https://zenodo.org/record/244313/files/figure.png" pageId="12" pageNumber="313">Fig. 85</figureCitation>
e). Such specimens might represent a distinct species, but at present this is difficult to determine due to the few available specimens.
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFFFFFF891F6FB11FC91A56B" pageId="12" pageNumber="313" type="discussion">
<paragraph id="8BF836E8FFFFFFF891F6FB11FC91A56B" blockId="12.[151,1437,151,1365]" pageId="12" pageNumber="313">
<emphasis id="B933EAFAFFFFFFF891F6FB11FEC3A4FB" bold="true" box="[199,315,1196,1221]" pageId="12" pageNumber="313">Remarks.</emphasis>
This species is close to
<taxonomicName id="4C474D6BFFFFFFF89369FB10FD36A4FA" box="[600,718,1196,1220]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B933EAFAFFFFFFF89369FB10FD36A4FA" box="[600,718,1196,1220]" italics="true" pageId="12" pageNumber="313">L. pumilio</emphasis>
</taxonomicName>
and related species (see below), sharing with them a series of characters: scimitar-shaped male outer foreclaws, the lobes of the metacoxal processes rounded in both sexes, and the prosternal process broad and short, barely reaching the metaventrite. However, it is one of only two species, along with
<taxonomicName id="4C474D6BFFFFFFF89027FAA4FE58A50E" box="[278,416,1304,1328]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B933EAFAFFFFFFF89027FAA4FE58A50E" box="[278,416,1304,1328]" italics="true" pageId="12" pageNumber="313">L. distinctus</emphasis>
</taxonomicName>
, that exhibits female gonocoxal blades with two teeth. The peculiar ramus, bearing dense, numerous and uniformly small teeth, is unique within the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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