treatments-xml/data/03/EE/87/03EE87FEFFF1FFFC91A6F887FF18A1CA.xml

<document id="9EDDC2630D341EA61F73A36B68FECC6D" ID-DOI="10.11646/zootaxa.3990.3.1" ID-GBIF-Dataset="cf747765-b9bf-42ed-8479-35355e712db0" ID-ISSN="1175-5326" ID-Zenodo-Dep="244301" ID-ZooBank="378C0359-E0E4-4CCC-821D-649144E37A63" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461268581225" checkinUser="plazi" docAuthor="Toledo, Mario &amp; Michat, Mariano C." docDate="2015" docId="03EE87FEFFF1FFFC91A6F887FF18A1CA" docLanguage="en" docName="zt03990p354.pdf" docOrigin="Zootaxa 3990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccomimus Toledo &amp; Michat, 2015, gen. n." docType="treatment" docVersion="9" lastPageNumber="308" masterDocId="FFD7FF86FFF3FFF49131FFBDFFF8A03E" masterDocTitle="Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)" masterLastPageNumber="354" masterPageNumber="301" pageNumber="303" updateTime="1698611165842" updateUser="plazi">
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<mods:title id="158ADAF208415992159F71665A906915">Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)</mods:title>
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<mods:namePart id="C33AFD7EA1D9542F76FFE8D886BC8351">Toledo, Mario</mods:namePart>
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<mods:namePart id="7B76970B310F7F6FFB89DFDA52341C14">Michat, Mariano C.</mods:namePart>
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<treatment id="03EE87FEFFF1FFFC91A6F887FF18A1CA" ID-DOI="http://doi.org/10.5281/zenodo.5692036" ID-GBIF-Taxon="119638440" ID-Zenodo-Dep="5692036" LSID="urn:lsid:plazi:treatment:03EE87FEFFF1FFFC91A6F887FF18A1CA" httpUri="http://treatment.plazi.org/id/03EE87FEFFF1FFFC91A6F887FF18A1CA" lastPageId="8" lastPageNumber="308" pageId="2" pageNumber="303">
<subSubSection id="C35D6563FFF1FFF691A6F887FE73A76B" box="[151,395,1850,1877]" pageId="2" pageNumber="303" type="nomenclature">
<paragraph id="8BF836E8FFF1FFF691A6F887FE73A76B" blockId="2.[151,395,1850,1877]" box="[151,395,1850,1877]" pageId="2" pageNumber="303">
<heading id="D0B08184FFF1FFF691A6F887FE73A76B" bold="true" box="[151,395,1850,1877]" fontSize="11" level="1" pageId="2" pageNumber="303" reason="1">
<emphasis id="B933EAFAFFF1FFF691A6F887FE73A76B" bold="true" box="[151,395,1850,1877]" pageId="2" pageNumber="303">
<taxonomicName id="4C474D6BFFF1FFF691A6F887FECAA76A" ID-CoL="8LY8Y" box="[151,306,1850,1876]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF1FFF691A6F887FECAA76A" bold="true" box="[151,306,1850,1876]" italics="true" pageId="2" pageNumber="303">Laccomimus</emphasis>
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<taxonomicNameLabel id="A2005781FFF1FFF69008F886FE73A76B" box="[313,395,1851,1877]" pageId="2" pageNumber="303" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BF836E8FFF1FFF691A6F8C7FCB2A7AE" blockId="2.[151,842,1914,1936]" box="[151,842,1914,1936]" pageId="2" pageNumber="303">
<emphasis id="B933EAFAFFF1FFF691A6F8C7FEDEA7AE" bold="true" box="[151,294,1914,1936]" pageId="2" pageNumber="303">
<typeStatus id="54FC884AFFF1FFF691A6F8C7FF34A7AE" box="[151,204,1914,1936]" pageId="2" pageNumber="303">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C474D6BFFF1FFF6901AF8C7FD62A7AE" authority="LeConte, 1878" authorityName="LeConte" authorityYear="1878" box="[299,666,1914,1936]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B933EAFAFFF1FFF6901AF8C7FE02A7AE" box="[299,506,1914,1936]" italics="true" pageId="2" pageNumber="303">Laccophilus pumilio</emphasis>
LeConte, 1878
</taxonomicName>
, here designated.
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<subSubSection id="C35D6563FFF1FFF791A6F80BFBCBA716" lastPageId="3" lastPageNumber="304" pageId="2" pageNumber="303" type="description">
<paragraph id="8BF836E8FFF1FFF791A6F80BFCE2A123" blockId="2.[151,1436,1974,2035]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="304" pageId="2" pageNumber="303">
<emphasis id="B933EAFAFFF1FFF691A6F80BFED1A7F1" bold="true" box="[151,297,1974,1999]" pageId="2" pageNumber="303">Description.</emphasis>
Habitus (
<figureCitation id="137C2A6DFFF1FFF690A4F80BFE25A7F1" box="[405,477,1974,1999]" captionStart="FIGURES 1 – 7. 1" captionStartId="29.[151,264,1905,1927]" captionTargetBox="[169,1412,219,1874]" captionTargetId="figure@29.[169,1413,193,1876]" captionTargetPageId="29" captionText="FIGURES 1 – 7. 1) Habitus of Laccomimus bolivari, dorsal view (Photo A. E. Z. Short); 2) habitus of Laccodytes apalodes, dorsal view (Photo A. E. Z. Short); 3 – 7) SEMs of Laccomimus species (females): 3) habitus of L. alvarengi, dorsal view; 4) habitus of L. bordoni, dorsal view; 5) pronotum of L. alvarengi, dorsal view; 6) pronotum of L. bordoni, dorsal view; 7) pronotum of L. bordoni, dorsolateral view." httpUri="https://zenodo.org/record/244302/files/figure.png" pageId="2" pageNumber="303">Figs 1</figureCitation>
; 3–4). Small 
<taxonomicName id="4C474D6BFFF1FFF69345F80BFCF9A7F1" box="[628,769,1974,1999]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
(TL: 
<quantity id="4CBF9B0DFFF1FFF69272F80BFC35A7F0" box="[835,973,1974,1999]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.8" pageId="2" pageNumber="303" unit="mm" value="2.15" valueMax="2.5" valueMin="1.8">1.8–2.5 mm</quantity>
; MW: 
<quantity id="4CBF9B0DFFF1FFF6952BF80AFB5DA7F1" box="[1050,1189,1975,1999]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="2" pageNumber="303" unit="mm" value="1.1" valueMax="1.3" valueMin="0.9">0.9–1.3 mm</quantity>
), oval or oblong-oval, teardrop-shaped, outline between head, pronotum and elytra continuously curved. Maximum width at about proximal 1/5 of elytra. Elytral tip pointed, gradually to abruptly narrowed. Colouration testaceous or reddishyellow, darker on elytra, which often bear pale markings; in one species elytra marmorate. Body surface smooth, without strongly impressed punctures. Elytra with characteristic iridescence, also occurring (although sometimes less evident) on pronotum, ventral surface, and metatarsi.
</paragraph>
<paragraph id="8BF836E8FFF0FFF791F6FE95FD8AA15A" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B933EAFAFFF0FFF791F6FE95FEF3A17E" box="[199,267,296,320]" italics="true" pageId="3" pageNumber="304">Head.</emphasis>
Broad, smooth, shiny, never iridescent, with fine and poorly impressed microreticulation, sometimes with few, scattered and shallow punctures.
</paragraph>
<paragraph id="8BF836E8FFF0FFF791F6FECCFB81A1EE" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B933EAFAFFF0FFF791F6FECCFEC5A1B6" box="[199,317,369,392]" italics="true" pageId="3" pageNumber="304">Pronotum.</emphasis>
Short, with angled projection at middle of base (
<figureCitation id="137C2A6DFFF0FFF7926AFED2FC39A1B6" box="[859,961,367,392]" captionStart="FIGURES 1 – 7. 1" captionStartId="29.[151,264,1905,1927]" captionTargetBox="[169,1412,219,1874]" captionTargetId="figure@29.[169,1413,193,1876]" captionTargetPageId="29" captionText="FIGURES 1 – 7. 1) Habitus of Laccomimus bolivari, dorsal view (Photo A. E. Z. Short); 2) habitus of Laccodytes apalodes, dorsal view (Photo A. E. Z. Short); 3 – 7) SEMs of Laccomimus species (females): 3) habitus of L. alvarengi, dorsal view; 4) habitus of L. bordoni, dorsal view; 5) pronotum of L. alvarengi, dorsal view; 6) pronotum of L. bordoni, dorsal view; 7) pronotum of L. bordoni, dorsolateral view." httpUri="https://zenodo.org/record/244302/files/figure.png" pageId="3" pageNumber="304">Figs 5–6</figureCitation>
), lateral margins not bordered, hind angles always rounded (
<figureCitation id="137C2A6DFFF0FFF7906DFE29FE5AA192" box="[348,418,404,429]" captionStart="FIGURES 1 – 7. 1" captionStartId="29.[151,264,1905,1927]" captionTargetBox="[169,1412,219,1874]" captionTargetId="figure@29.[169,1413,193,1876]" captionTargetPageId="29" captionText="FIGURES 1 – 7. 1) Habitus of Laccomimus bolivari, dorsal view (Photo A. E. Z. Short); 2) habitus of Laccodytes apalodes, dorsal view (Photo A. E. Z. Short); 3 – 7) SEMs of Laccomimus species (females): 3) habitus of L. alvarengi, dorsal view; 4) habitus of L. bordoni, dorsal view; 5) pronotum of L. alvarengi, dorsal view; 6) pronotum of L. bordoni, dorsal view; 7) pronotum of L. bordoni, dorsolateral view." httpUri="https://zenodo.org/record/244302/files/figure.png" pageId="3" pageNumber="304">Fig. 7</figureCitation>
). Surface smooth, with fine microreticulation; often with a subtle iridescence. Punctures mostly along lateral sides, hind and fore margins, arranged in more or less regular series.
</paragraph>
<paragraph id="8BF836E8FFF0FFF791F6FE61FC2DA2BA" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B933EAFAFFF0FFF791F6FE61FEF5A1CA" box="[199,269,476,500]" italics="true" pageId="3" pageNumber="304">Elytra</emphasis>
. Elongate, apically attenuate and often strongly narrowed. Sutural line between elytron and epipleuron not visible in dorsal view, therefore sides of elytra apparently not bordered. Surface smooth, iridescent, with fine microreticulation of elongate cells (
<figureCitation id="137C2A6DFFF0FFF79301FD99FD4EA202" box="[560,694,548,573]" captionStart="FIGURES 14 – 19" captionStartId="31.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@31.[151,1436,193,1932]" captionTargetPageId="31" captionText="FIGURES 14 – 19. SEMs of microsculpture in Laccomimus species (females): 14) L. alvarengi, elytral reticulation; 15) idem, at higher magnification; 16) L. bordoni, elytral reticulation; 17) idem, at higher magnification; 18) L. bordoni, metacoxal reticulation; 19) idem, at higher magnification." httpUri="https://zenodo.org/record/244304/files/figure.png" pageId="3" pageNumber="304">Figs 14–17</figureCitation>
). Small and superficial punctures visible in most species, sparse and/or arranged in irregular, longitudinal series. Epipleuron broad to metacoxae/first abdominal ventrite, then strongly narrowed, distinctly attenuate just before elytral tip (
<figureCitation id="137C2A6DFFF0FFF79277FDD1FC30A2BA" box="[838,968,620,645]" captionStart="FIGURES 8 – 13" captionStartId="30.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@30.[151,1436,193,1932]" captionTargetPageId="30" captionText="FIGURES 8 – 13. SEMs of Laccomimus species, ventral view: 8) L. distinctus (male); 9) L. malkini (female); 10) L. amazonas (male); 11) L. alvarengi (female); 12) L. bordoni (male); 13) L. bolivari (female)." httpUri="https://zenodo.org/record/244303/files/figure.png" pageId="3" pageNumber="304">Figs 10–13</figureCitation>
).
</paragraph>
<paragraph id="8BF836E8FFF0FFF791F6FD2DFCA1A3F6" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B933EAFAFFF0FFF791F6FD2DFEBBA296" box="[199,323,656,680]" italics="true" pageId="3" pageNumber="304">Underside.</emphasis>
(
<figureCitation id="137C2A6DFFF0FFF79064FD32FE32A296" box="[341,458,655,680]" captionStart="FIGURES 8 – 13" captionStartId="30.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@30.[151,1436,193,1932]" captionTargetPageId="30" captionText="FIGURES 8 – 13. SEMs of Laccomimus species, ventral view: 8) L. distinctus (male); 9) L. malkini (female); 10) L. amazonas (male); 11) L. alvarengi (female); 12) L. bordoni (male); 13) L. bolivari (female)." httpUri="https://zenodo.org/record/244303/files/figure.png" pageId="3" pageNumber="304">Figs 8–13</figureCitation>
) Smooth, more or less iridescent, with microreticulation as on elytra (
<figureCitation id="137C2A6DFFF0FFF79435FD32FA98A296" box="[1284,1376,655,680]" captionStart="FIGURES 14 – 19" captionStartId="31.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@31.[151,1436,193,1932]" captionTargetPageId="31" captionText="FIGURES 14 – 19. SEMs of microsculpture in Laccomimus species (females): 14) L. alvarengi, elytral reticulation; 15) idem, at higher magnification; 16) L. bordoni, elytral reticulation; 17) idem, at higher magnification; 18) L. bordoni, metacoxal reticulation; 19) idem, at higher magnification." httpUri="https://zenodo.org/record/244304/files/figure.png" pageId="3" pageNumber="304">Figs 18</figureCitation>
; 19), usually more impressed, especially in females. Lateral expansions of metaventrite (“metasternal wings”) thin and arched. Prosternum and head separated by a more or less pronounced step, (
<figureCitation id="137C2A6DFFF0FFF792C4FD6AFB81A2CE" box="[1013,1145,727,752]" captionStart-0="FIGURES 20 – 25" captionStart-1="FIGURES 26 – 31" captionStartId-0="32.[151,264,1953,1975]" captionStartId-1="33.[151,264,1953,1975]" captionTargetBox-0="[151,1436,193,1930]" captionTargetBox-1="[151,1436,193,1930]" captionTargetId-0="figure@32.[151,1436,193,1932]" captionTargetId-1="figure@33.[151,1436,193,1932]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionText-0="FIGURES 20 – 25. SEMs of head, prosternum and prosternal process in Laccomimus species: 20) L. distinctus (male), ventral view, 21) idem, at higher magnification; 22) idem, oblique lateral view; 23) L. malkini (female) ventral view; 24) L. amazonas (male), ventral view (arrow indicates protruding prosternum); 25) idem, at higher magnification." captionText-1="FIGURES 26 – 31. SEMs of head, prosternum and prosternal process in Laccomimus species: 26) L. amazonas (male), oblique lateral view; 27) idem, at higher magnification; 28) L. alvarengi (female) ventral view; 29) L. bordoni (female), ventral view; 30) idem, oblique lateral view; 31) idem, at higher magnification." httpUri-0="https://zenodo.org/record/244305/files/figure.png" httpUri-1="https://zenodo.org/record/244306/files/figure.png" pageId="3" pageNumber="304">Figs 20–31</figureCitation>
). Prosternal process more or less blunt (
<figureCitation id="137C2A6DFFF0FFF79003FD41FE4EA32A" box="[306,438,764,789]" captionStart-0="FIGURES 20 – 25" captionStart-1="FIGURES 26 – 31" captionStartId-0="32.[151,264,1953,1975]" captionStartId-1="33.[151,264,1953,1975]" captionTargetBox-0="[151,1436,193,1930]" captionTargetBox-1="[151,1436,193,1930]" captionTargetId-0="figure@32.[151,1436,193,1932]" captionTargetId-1="figure@33.[151,1436,193,1932]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionText-0="FIGURES 20 – 25. SEMs of head, prosternum and prosternal process in Laccomimus species: 20) L. distinctus (male), ventral view, 21) idem, at higher magnification; 22) idem, oblique lateral view; 23) L. malkini (female) ventral view; 24) L. amazonas (male), ventral view (arrow indicates protruding prosternum); 25) idem, at higher magnification." captionText-1="FIGURES 26 – 31. SEMs of head, prosternum and prosternal process in Laccomimus species: 26) L. amazonas (male), oblique lateral view; 27) idem, at higher magnification; 28) L. alvarengi (female) ventral view; 29) L. bordoni (female), ventral view; 30) idem, oblique lateral view; 31) idem, at higher magnification." httpUri-0="https://zenodo.org/record/244305/files/figure.png" httpUri-1="https://zenodo.org/record/244306/files/figure.png" pageId="3" pageNumber="304">Figs 20–31</figureCitation>
), in lateral view keeled to saddle shaped, depending on species; in ventral view broadly lanceolate to cordiform, bordered, ending in short, pointed to broadly rounded tip, reaching or barely reaching metaventrite. Metacoxal lines sinuate, strongly convergent anteriorly (
<figureCitation id="137C2A6DFFF0FFF792F8FCF9FBA8A362" box="[969,1104,836,861]" captionStart-0="FIGURES 32 – 37" captionStart-1="FIGURES 38 – 43" captionStartId-0="34.[151,264,1953,1975]" captionStartId-1="35.[151,264,1953,1975]" captionTargetBox-0="[151,1436,193,1930]" captionTargetBox-1="[151,1436,193,1930]" captionTargetId-0="figure@34.[151,1436,193,1932]" captionTargetId-1="figure@35.[151,1436,193,1932]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionText-0="FIGURES 32 – 37. SEMs of metacoxal processes in Laccomimus species, ventral view: 32) L. distinctus (male); 33 – 36) L. distinctus (females), variability in development of apical angulation in different specimens; 37) L. malkini (female)." captionText-1="FIGURES 38 – 43. SEMs of Laccomimus species, ventral view. 38 – 41) metacoxal processes: 38) L. amazonas (male); 39) L. alvarengi (female); 40) L. bordoni (female); 41) L. bolivari (female); 42 – 43) abdomen on L. bordoni, showing sexual difference in last ventrite: 42) male (arrow indicates slightly sinuate caudal margins); 43) female." httpUri-0="https://zenodo.org/record/244307/files/figure.png" httpUri-1="https://zenodo.org/record/244308/files/figure.png" pageId="3" pageNumber="304">Figs 32–41</figureCitation>
); fore laminae of metacoxal processes divided by deep V-shaped incision, either broadly rounded to angulate or ending in sharp spine. Abdominal ventrites lacking curved scratchlike sculpture. Last ventrite as long as rest of abdomen, or almost so (
<figureCitation id="137C2A6DFFF0FFF791AEFC12FF0BA3F6" box="[159,243,943,968]" captionStart="FIGURES 38 – 43" captionStartId="35.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@35.[151,1436,193,1932]" captionTargetPageId="35" captionText="FIGURES 38 – 43. SEMs of Laccomimus species, ventral view. 38 – 41) metacoxal processes: 38) L. amazonas (male); 39) L. alvarengi (female); 40) L. bordoni (female); 41) L. bolivari (female); 42 – 43) abdomen on L. bordoni, showing sexual difference in last ventrite: 42) male (arrow indicates slightly sinuate caudal margins); 43) female." httpUri="https://zenodo.org/record/244308/files/figure.png" pageId="3" pageNumber="304">Fig. 42</figureCitation>
), with large spiculiferous punctures at about distal 2/3.
</paragraph>
<paragraph id="8BF836E8FFF0FFF791F6FC68FE5DA4FB" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B933EAFAFFF0FFF791F6FC68FF04A3D2" box="[199,252,981,1004]" italics="true" pageId="3" pageNumber="304">Legs</emphasis>
. Pro- and mesothoracic legs short, tarsomeres with width greater than length except for fifth almost the length of third and fourth together. Pro- and mesotarsal claws as long as or slightly longer than fifth tarsomere. Spurs of mesotibiae very long, almost as long as entire mesotarsus (
<figureCitation id="137C2A6DFFF0FFF792AEFBA1FC00A40B" box="[927,1016,1052,1077]" captionStart="FIGURES 52 – 58" captionStartId="37.[151,264,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@37.[151,1436,193,1856]" captionTargetPageId="37" captionText="FIGURES 52 – 58. SEMs of legs in Laccomimus species: 52) L. bordoni (female), left mesothoracic leg, dorsal view; 53) L. bordoni (male), left mesotarsus (arrow indicates mesotibial spurs); 54) L. bordoni (female), right metathoracic leg, ventral view; 55) L. bordoni (male), left metathoracic leg, dorsal view; 56) L. bordoni (female), right metatarsus, ventral view (arrow indicates combs of flat spines on posterolateral lobes); 57) L. alvarengi (female), metatarsi, ventral view; 58) L. bolivari (female), metathoracic leg, dorsal view (arrow indicates posterolateral lobes of metatarsus)." httpUri="https://zenodo.org/record/244310/files/figure.png" pageId="3" pageNumber="304">Figs 52</figureCitation>
; 53). Swimming setae on mesotibiae present only apically. Posterior surface of mesofemora always with a series of 3–4 long, stiff setae (
<figureCitation id="137C2A6DFFF0FFF795C1FB82FAC0A466" box="[1264,1336,1087,1112]" captionStart="FIGURES 8 – 13" captionStartId="30.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@30.[151,1436,193,1932]" captionTargetPageId="30" captionText="FIGURES 8 – 13. SEMs of Laccomimus species, ventral view: 8) L. distinctus (male); 9) L. malkini (female); 10) L. amazonas (male); 11) L. alvarengi (female); 12) L. bordoni (male); 13) L. bolivari (female)." httpUri="https://zenodo.org/record/244303/files/figure.png" pageId="3" pageNumber="304">Figs 8</figureCitation>
; 12; 52). Spurs of metatibiae simple, not bifid (
<figureCitation id="137C2A6DFFF0FFF7930DFBD9FD47A443" box="[572,703,1124,1149]" captionStart="FIGURES 52 – 58" captionStartId="37.[151,264,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@37.[151,1436,193,1856]" captionTargetPageId="37" captionText="FIGURES 52 – 58. SEMs of legs in Laccomimus species: 52) L. bordoni (female), left mesothoracic leg, dorsal view; 53) L. bordoni (male), left mesotarsus (arrow indicates mesotibial spurs); 54) L. bordoni (female), right metathoracic leg, ventral view; 55) L. bordoni (male), left metathoracic leg, dorsal view; 56) L. bordoni (female), right metatarsus, ventral view (arrow indicates combs of flat spines on posterolateral lobes); 57) L. alvarengi (female), metatarsi, ventral view; 58) L. bolivari (female), metathoracic leg, dorsal view (arrow indicates posterolateral lobes of metatarsus)." httpUri="https://zenodo.org/record/244310/files/figure.png" pageId="3" pageNumber="304">Figs 54–55</figureCitation>
). Metatarsi with iridescence as on elytra; segments 1–4 with short posterolateral lobes and small combs of flat spines. Last metatarsomere with inner claw smaller, outer claw more developed, spur shaped.
</paragraph>
<paragraph id="8BF836E8FFF0FFF791F6FB6DFEBAA636" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B933EAFAFFF0FFF791F6FB6DFEF3A4D6" box="[199,267,1232,1256]" italics="true" pageId="3" pageNumber="304">Males</emphasis>
. Last abdominal ventrite tectiform, distally slightly sinuate to almost straight at sides, ending in more or less pointed medial tip (
<figureCitation id="137C2A6DFFF0FFF79094FB49FE02A532" box="[421,506,1268,1293]" captionStart="FIGURES 38 – 43" captionStartId="35.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@35.[151,1436,193,1932]" captionTargetPageId="35" captionText="FIGURES 38 – 43. SEMs of Laccomimus species, ventral view. 38 – 41) metacoxal processes: 38) L. amazonas (male); 39) L. alvarengi (female); 40) L. bordoni (female); 41) L. bolivari (female); 42 – 43) abdomen on L. bordoni, showing sexual difference in last ventrite: 42) male (arrow indicates slightly sinuate caudal margins); 43) female." httpUri="https://zenodo.org/record/244308/files/figure.png" pageId="3" pageNumber="304">Fig. 42</figureCitation>
). Pro- and mesotarsomeres 1–3 weakly but visibly dilated, each one bearing a pair of stalked suction palettes (
<figureCitation id="137C2A6DFFF0FFF790E0FAAAFDAEA50E" box="[465,598,1303,1328]" captionStart="FIGURES 44 – 51" captionStartId="36.[151,264,1877,1899]" captionTargetBox="[182,1398,193,1837]" captionTargetId="figure@36.[182,1398,193,1839]" captionTargetPageId="36" captionText="FIGURES 44 – 51. SEMs of left protarsus in Laccomimus species (males): 44) L. distinctus, general view; 45) idem, detail at higher magnification of last protarsal segment and claws; 46) idem, detail at higher magnification of stalked suction palettes; 47) L. amazonas, general view; 48) idem, detail at higher magnification of stalked suction palettes; 49) L. bordoni, general view; 50) idem, detail at higher magnification of last protarsal segment and claws; 51) idem, detail at higher magnification of stalked suction palettes." httpUri="https://zenodo.org/record/244309/files/figure.png" pageId="3" pageNumber="304">Figs 44–51</figureCitation>
). In all but one species, outer claw of forelegs modified, scimitar-shaped (
<figureCitation id="137C2A6DFFF0FFF791AEFA81FF0EA56A" box="[159,246,1340,1365]" captionStart="FIGURES 44 – 51" captionStartId="36.[151,264,1877,1899]" captionTargetBox="[182,1398,193,1837]" captionTargetId="figure@36.[182,1398,193,1839]" captionTargetPageId="36" captionText="FIGURES 44 – 51. SEMs of left protarsus in Laccomimus species (males): 44) L. distinctus, general view; 45) idem, detail at higher magnification of last protarsal segment and claws; 46) idem, detail at higher magnification of stalked suction palettes; 47) L. amazonas, general view; 48) idem, detail at higher magnification of stalked suction palettes; 49) L. bordoni, general view; 50) idem, detail at higher magnification of last protarsal segment and claws; 51) idem, detail at higher magnification of stalked suction palettes." httpUri="https://zenodo.org/record/244309/files/figure.png" pageId="3" pageNumber="304">Figs 49</figureCitation>
; 70–76), in some cases with few small denticles on ventral margin. Aedeagus (
<figureCitation id="137C2A6DFFF0FFF79558FA81FB0FA56B" box="[1129,1271,1340,1365]" captionStart-0="FIGURES 84 – 87" captionStart-1="FIGURES 88 – 90" captionStart-2="FIGURES 91 – 93" captionStart-3="FIGURES 94 – 97" captionStart-4="FIGURES 98 – 102" captionStart-5="FIGURES 103 – 106" captionStart-6="FIGURES 107 – 115" captionStartId-0="40.[151,264,1915,1937]" captionStartId-1="41.[151,264,1919,1941]" captionStartId-2="42.[151,264,1921,1943]" captionStartId-3="43.[151,264,1919,1941]" captionStartId-4="44.[151,264,1877,1899]" captionStartId-5="45.[151,264,1913,1935]" captionStartId-6="46.[151,264,1951,1973]" captionTargetBox-0="[169,1412,201,1889]" captionTargetBox-1="[225,1366,198,1894]" captionTargetBox-2="[168,1414,201,1892]" captionTargetBox-3="[174,1408,200,1884]" captionTargetBox-4="[162,1422,201,1834]" captionTargetBox-5="[158,1429,198,1873]" captionTargetBox-6="[184,1390,196,1925]" captionTargetId-0="figure@40.[160,1420,193,1898]" captionTargetId-1="figure@41.[204,1370,193,1898]" captionTargetId-2="figure@42.[162,1422,193,1898]" captionTargetId-3="figure@43.[164,1418,193,1889]" captionTargetId-4="figure@44.[151,1436,193,1856]" captionTargetId-5="figure@45.[151,1436,193,1892]" captionTargetId-6="figure@46.[170,1417,193,1930]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionTargetPageId-2="42" captionTargetPageId-3="43" captionTargetPageId-4="44" captionTargetPageId-5="45" captionTargetPageId-6="46" captionText-0="FIGURES 84 – 87. Male genitalia in Laccomimus species, left lateral view: 84) L. distinctus, a) median lobe, b) left paramere, c) right paramere; 85) L. malkini (Loreto, Perú), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere, e) magnification of apex of median lobe (specimen from Piracicaba, Brazil); 86) L. variegatus, a) median lobe, b) left paramere, c) right paramere; 87) L. spinosus, a) median lobe, b) left paramere." captionText-1="FIGURES 88 – 90. Male genitalia in Laccomimus species, left lateral view: 88) L. amazonas, a) median lobe, b) left paramere, c) right paramere; 89) L. improvidus, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 90) L. youngi, a) median lobe (holotype), b) magnification of apex of median lobe (holotype), c) magnification of apex of median lobe (different specimen from type locality), d) left paramere (holotype), e) left paramere (different specimen from type locality)." captionText-2="FIGURES 91 – 93. Male genitalia in Laccomimus pumilio from different localities, left lateral view: 91) Belize, a) median lobe (showing also the three general parts of median lobe), b) magnification of apex of median lobe, c) left paramere, d) right paramere; 92) Florida, USA, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 93) Cayamas, Cuba, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere." captionText-3="FIGURES 94 – 97. Male genitalia in Laccomimus alvarengi from different localities, left lateral view: 94) holotype, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 95) likely aberrant specimen from type locality, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 96) Loreto, Perú, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 97) Mbrucuyá National Park, Argentina, a) median lobe, b) left paramere." captionText-4="FIGURES 98 – 102. Male genitalia in Laccomimus alvarengi from different localities, left lateral view: 98) Mbrucuyá National Park, Argentina, a) median lobe, b) left paramere; 99) Beni, Bolivia, a) median lobe, b) magnification of apex of median lobe; 100) Santa Rosa, Bolivia, median lobe; 101) Esmeraldas, Ecuador, a) median lobe, b) left paramere; 102) Albrook, Panamá, a) median lobe, b) magnification of apex of median lobe, c) left paramere." captionText-5="FIGURES 103 – 106. Male genitalia in Laccomimus species, left lateral view: 103) L. bordoni, a) median lobe, b) magnification of apex of median lobe, c) left paramere, showing apex in different positions, d) right paramere; 104) L. spangleri, a) median lobe, b) left paramere; c) right paramere; 105) L. bolivari (holotype), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 106) L. bolivari (Suriname), a) median lobe, b) magnification of apex of median lobe." captionText-6="FIGURES 107 – 115. Median lobe of aedeagus in Laccomimus species, dorsal view: 107) L. distinctus; 108) L. malkini; 109) L. variegatus; 110) L. amazonas; 111) L. alvarengi; 112) L. improvidus; 113) L. bordoni; 114) L. spangleri; 115) L. bolivari." httpUri-0="https://zenodo.org/record/244313/files/figure.png" httpUri-1="https://zenodo.org/record/244314/files/figure.png" httpUri-2="https://zenodo.org/record/244315/files/figure.png" httpUri-3="https://zenodo.org/record/244316/files/figure.png" httpUri-4="https://zenodo.org/record/244317/files/figure.png" httpUri-5="https://zenodo.org/record/244318/files/figure.png" httpUri-6="https://zenodo.org/record/244319/files/figure.png" pageId="3" pageNumber="304">Figs 84–115</figureCitation>
) asymmetrical as in other genera of 
<taxonomicName id="4C474D6BFFF0FFF790B9FAE2FDE1A546" box="[392,537,1375,1400]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
; median lobe elongate, generally divided into three parts, variable in shape and proportions depending on species: i) with a thicker base, ii) with distal portion gradually narrowed to apex, or iii) ending in apical expansion (
<figureCitation id="137C2A6DFFF0FFF790FEFA1AFDDAA5FE" box="[463,546,1447,1472]" captionStart="FIGURES 91 – 93" captionStartId="42.[151,264,1921,1943]" captionTargetBox="[168,1414,201,1892]" captionTargetId="figure@42.[162,1422,193,1898]" captionTargetPageId="42" captionText="FIGURES 91 – 93. Male genitalia in Laccomimus pumilio from different localities, left lateral view: 91) Belize, a) median lobe (showing also the three general parts of median lobe), b) magnification of apex of median lobe, c) left paramere, d) right paramere; 92) Florida, USA, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 93) Cayamas, Cuba, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere." httpUri="https://zenodo.org/record/244315/files/figure.png" pageId="3" pageNumber="304">Fig. 91</figureCitation>
). Left paramere long and thin, arched, either sinuate or straight, reaching half or more than half length of median lobe; right paramere smaller, subtriangular or oval. Both parameres with single long seta at tip.
</paragraph>
<paragraph id="8BF836E8FFF0FFF791F6F9A9FEDEA6DE" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B933EAFAFFF0FFF791F6F9A9FEDEA612" box="[199,294,1556,1580]" italics="true" pageId="3" pageNumber="304">Females</emphasis>
. Often duller than males due to more impressed reticulation; iridescence on elytra, pronotum and metacoxae more intense than in males in most cases. Last abdominal ventrite triangular, with almost pointed to almost truncate apex (
<figureCitation id="137C2A6DFFF0FFF790ACF9E1FE0EA64A" box="[413,502,1628,1653]" captionStart="FIGURES 38 – 43" captionStartId="35.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@35.[151,1436,193,1932]" captionTargetPageId="35" captionText="FIGURES 38 – 43. SEMs of Laccomimus species, ventral view. 38 – 41) metacoxal processes: 38) L. amazonas (male); 39) L. alvarengi (female); 40) L. bordoni (female); 41) L. bolivari (female); 42 – 43) abdomen on L. bordoni, showing sexual difference in last ventrite: 42) male (arrow indicates slightly sinuate caudal margins); 43) female." httpUri="https://zenodo.org/record/244308/files/figure.png" pageId="3" pageNumber="304">Fig. 43</figureCitation>
). Female genitalia (
<figureCitation id="137C2A6DFFF0FFF793D0F9E1FC7CA64B" box="[737,900,1628,1653]" captionStart="FIGURES 116 – 121" captionStartId="47.[151,264,1829,1851]" captionTargetBox="[229,1344,199,1783]" captionTargetId="figure@47.[198,1370,193,1789]" captionTargetPageId="47" captionText="FIGURES 116 – 121. Female genitalia and cladogram. 116 – 120) Female genitalia in Laccomimus species, left lateral view: 116) L. distinctus (Paraguay), a) left gonocoxa, b) ramus; 117) L. malkini (Perú), a) left gonocoxa, b) ramus; 118) L. alvarengi (Brazil), a) left gonocoxa, b) ramus; 119) L. variegatus (Perú), a) left gonocoxa, b) ramus; 120) L. bordoni (Bolivia), ramus (arrow indicates lateral proximal process of ramus). 121) Single most parsimonious cladogram rooted between ingroup and outgroup, with character changes mapped for each clade. Solid rectangles indicate unique character state transformations; open rectangles indicate homoplasious character state transformations. Support values are indicated on each branch as Bremer / Bootstrap." httpUri="https://zenodo.org/record/244320/files/figure.png" pageId="3" pageNumber="304">Figs 116–120</figureCitation>
): distal segments of gonocoxae blade-shaped, sharply acute at tip, with small median tooth-like process; an additional subapical denticle present in two species. Ramus short, with more or less densely dentate “saw”, depending on species, and lateroproximal processes rudimentary.
</paragraph>
<paragraph id="8BF836E8FFF0FFF791F6F950FBCBA716" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
In general, the species of 
<taxonomicName id="4C474D6BFFF0FFF790C7F950FD7CA73A" box="[502,644,1773,1796]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF0FFF790C7F950FD7CA73A" box="[502,644,1773,1796]" italics="true" pageId="3" pageNumber="304">Laccomimus</emphasis>
</taxonomicName>
are hard to identify without examination of male sexual characters. External features are often homogeneous or very variable and difficult to evaluate.
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFF0FFF791F6F889FA64A7E2" pageId="3" pageNumber="304" type="distribution">
<paragraph id="8BF836E8FFF0FFF791F6F889FA64A7E2" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B933EAFAFFF0FFF791F6F889FE02A773" bold="true" box="[199,506,1844,1869]" pageId="3" pageNumber="304">Distribution and ecology.</emphasis>
A mainly Neotropical genus, widespread in Central and South 
<collectingCountry id="F3507678FFF0FFF795D9F889FAB4A773" box="[1256,1356,1844,1869]" name="United States of America" pageId="3" pageNumber="304">America</collectingCountry>
. Three species reach 
<collectingCountry id="F3507678FFF0FFF79002F8EAFE5DA74E" box="[307,421,1879,1904]" name="Argentina" pageId="3" pageNumber="304">Argentina</collectingCountry>
, and another one reaches Florida in the 
<collectingCountry id="F3507678FFF0FFF79255F8E5FC73A74E" box="[868,907,1880,1904]" name="United States of America" pageId="3" pageNumber="304">US</collectingCountry>
. Little ecological information is known for this genus. A large portion of the studied material was collected at light. Available field data (see Figs 130–138) suggest a preference for lentic or stagnant waters that are rich in debris and vegetation. Apparently, a few species live in running waters or have a wider ecological range. 
<bibRefCitation id="EFD64B19FFF0FFF7938CF878FCAFA7E3" author="Young" box="[701,855,1988,2013]" pageId="3" pageNumber="304" refString="Young, F. N. (1954) The water beetles of Florida. University of Florida Studies, Biological Science Series, 5, 1 - 238." type="journal article" year="1954">Young (1954)</bibRefCitation>
gave a quite detailed description of the habitat of 
<emphasis id="B933EAFAFFF0FFF794B4F878FA64A7E2" box="[1413,1436,1989,2012]" italics="true" pageId="3" pageNumber="304">L.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFF7FFF091A6FF25FB0DA123" pageId="4" pageNumber="305" type="reference_group">
<paragraph id="8BF836E8FFF7FFF091A6FF25FB0DA123" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="305">
<taxonomicName id="4C474D6BFFF7FFF091A6FF25FF15A08E" box="[151,237,152,176]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B933EAFAFFF7FFF091A6FF25FF15A08E" box="[151,237,152,176]" italics="true" pageId="4" pageNumber="305">pumilio</emphasis>
</taxonomicName>
in Florida (Alachua County), where specimens were collected from the heavily shaded edges of a large, permanent woodland pond near Gainesville, in association with 
<taxonomicName id="4C474D6BFFF7FFF092B6FF01FBD1A0EB" ID-CoL="8VR3G" box="[903,1065,188,213]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
(
<taxonomicName id="4C474D6BFFF7FFF0950CFF01FF20A0C6" authority="LeConte, 1855" authorityName="LeConte" authorityYear="1855" class="Insecta" family="Hydrophilidae" genus="Helocombus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="B933EAFAFFF7FFF0950CFF01FAD8A0EA" box="[1085,1312,188,212]" italics="true" pageId="4" pageNumber="305">Helocombus bifidus</emphasis>
(LeConte, 1855)
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF7FFF09024FF5DFE78A0C6" ID-CoL="8VRNH" box="[277,384,224,248]" class="Insecta" family="Hydrophilidae" genus="Enochrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF09024FF5DFE78A0C6" box="[277,384,224,248]" italics="true" pageId="4" pageNumber="305">Enochrus</emphasis>
</taxonomicName>
spp.) and 
<taxonomicName id="4C474D6BFFF7FFF090CEFF62FD8FA0C6" ID-CoL="9JJ" box="[511,631,223,248]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
(
<taxonomicName id="4C474D6BFFF7FFF093BAFF5DFC69A0C6" ID-CoL="6CNJC" box="[651,913,224,248]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="granum">
<emphasis id="B933EAFAFFF7FFF093BAFF5DFC69A0C6" box="[651,913,224,248]" italics="true" pageId="4" pageNumber="305">Desmopachria granum</emphasis>
</taxonomicName>
-complex, listed by 
<bibRefCitation id="EFD64B19FFF7FFF09548FF5DFAE0A0C6" author="Young" box="[1145,1304,223,248]" pageId="4" pageNumber="305" refString="Young, F. N. (1954) The water beetles of Florida. University of Florida Studies, Biological Science Series, 5, 1 - 238." type="journal article" year="1954">Young (1954)</bibRefCitation>
as &quot; 
<emphasis id="B933EAFAFFF7FFF09461FF5CFA6BA0C6" box="[1360,1427,225,248]" italics="true" pageId="4" pageNumber="305">grana</emphasis>
- complex&quot;). The occurrence of adults of this genus is reported to be very seasonal (see 
<bibRefCitation id="EFD64B19FFF7FFF0956DFEB8FB10A123" author="Young" box="[1116,1256,260,285]" pageId="4" pageNumber="305" refString="Young, F. N. (1954) The water beetles of Florida. University of Florida Studies, Biological Science Series, 5, 1 - 238." type="journal article" year="1954">Young 1954</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFF7FFF391F6FE9AFEDAA669" lastPageId="7" lastPageNumber="308" pageId="4" pageNumber="305" type="etymology">
<paragraph id="8BF836E8FFF7FFF091F6FE9AFD53A1B6" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="305">
<emphasis id="B933EAFAFFF7FFF091F6FE9AFEB6A17E" bold="true" box="[199,334,295,320]" pageId="4" pageNumber="305">Etymology.</emphasis>
<taxonomicName id="4C474D6BFFF7FFF09068FE95FE1EA101" box="[345,486,296,319]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF7FFF09068FE95FE1EA101" box="[345,486,296,319]" italics="true" pageId="4" pageNumber="305">Laccomimus</emphasis>
</taxonomicName>
is derived from the words 
<emphasis id="B933EAFAFFF7FFF09200FE95FC95A17E" box="[817,877,296,320]" italics="true" pageId="4" pageNumber="305">lacco</emphasis>
, meaning “lake”, and 
<emphasis id="B933EAFAFFF7FFF0954DFE95FB3DA101" box="[1148,1221,296,319]" italics="true" pageId="4" pageNumber="305">mimus</emphasis>
, meaning “actor”. Maybe “water actor” for their animated and aquatic habits. Name given 
<emphasis id="B933EAFAFFF7FFF092FCFEF0FBCCA15A" box="[973,1076,332,356]" italics="true" pageId="4" pageNumber="305">in litteris</emphasis>
by Frank N. Young and Paul J. Spangler. The gender of the name is masculine.
</paragraph>
<paragraph id="8BF836E8FFF7FFF091A6FE0AFAB9A1EE" blockId="4.[151,1345,439,464]" box="[151,1345,439,464]" pageId="4" pageNumber="305">
<emphasis id="B933EAFAFFF7FFF091A6FE0AFD68A1EE" bold="true" box="[151,656,439,464]" pageId="4" pageNumber="305">
Phylogenetic relationships of 
<taxonomicName id="4C474D6BFFF7FFF090C4FE05FD70A1EE" box="[501,648,440,464]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF7FFF090C4FE05FD70A1EE" bold="true" box="[501,648,440,464]" italics="true" pageId="4" pageNumber="305">Laccomimus</emphasis>
</taxonomicName>
.
</emphasis>
The phylogenetic analysis included the following characters:
</paragraph>
<paragraph id="8BF836E8FFF7FFF091A6FDBDFAE9A296" blockId="4.[151,1437,511,680]" pageId="4" pageNumber="305">
<emphasis id="B933EAFAFFF7FFF091A6FDBDFEDCA226" box="[151,292,512,536]" italics="true" pageId="4" pageNumber="305">Character 0</emphasis>
. Prosternum + head: (0) divided by a shallow step; (1) divided by a deep step; (2) prosternum protruding anteriorly. In 
<taxonomicName id="4C474D6BFFF7FFF0909FFD98FDC3A202" box="[430,571,549,572]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF7FFF0909FFD98FDC3A202" box="[430,571,549,572]" italics="true" pageId="4" pageNumber="305">Laccomimus</emphasis>
</taxonomicName>
the anterior side of the prosternum, just posterior to the head, is quite thick. Pronotum and head are therefore separated by a kind of step (
<emphasis id="B933EAFAFFF7FFF092BAFDF5FC00A25E" box="[907,1016,584,608]" italics="true" pageId="4" pageNumber="305">character</emphasis>
0.1; 
<figureCitation id="137C2A6DFFF7FFF0950DFDFAFB3BA25E" box="[1084,1219,583,608]" captionStart="FIGURES 20 – 25" captionStartId="32.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@32.[151,1436,193,1932]" captionTargetPageId="32" captionText="FIGURES 20 – 25. SEMs of head, prosternum and prosternal process in Laccomimus species: 20) L. distinctus (male), ventral view, 21) idem, at higher magnification; 22) idem, oblique lateral view; 23) L. malkini (female) ventral view; 24) L. amazonas (male), ventral view (arrow indicates protruding prosternum); 25) idem, at higher magnification." httpUri="https://zenodo.org/record/244305/files/figure.png" pageId="4" pageNumber="305">Figs 20–22</figureCitation>
; 29–31), which is particularly visible in those species with a protruding prosternum (
<emphasis id="B933EAFAFFF7FFF0924EFDD1FBEEA2BA" box="[895,1046,620,644]" italics="true" pageId="4" pageNumber="305">character 0.2</emphasis>
; 
<figureCitation id="137C2A6DFFF7FFF09515FDD1FB5EA2BA" box="[1060,1190,620,645]" captionStart-0="FIGURES 20 – 25" captionStart-1="FIGURES 26 – 31" captionStartId-0="32.[151,264,1953,1975]" captionStartId-1="33.[151,264,1953,1975]" captionTargetBox-0="[151,1436,193,1930]" captionTargetBox-1="[151,1436,193,1930]" captionTargetId-0="figure@32.[151,1436,193,1932]" captionTargetId-1="figure@33.[151,1436,193,1932]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionText-0="FIGURES 20 – 25. SEMs of head, prosternum and prosternal process in Laccomimus species: 20) L. distinctus (male), ventral view, 21) idem, at higher magnification; 22) idem, oblique lateral view; 23) L. malkini (female) ventral view; 24) L. amazonas (male), ventral view (arrow indicates protruding prosternum); 25) idem, at higher magnification." captionText-1="FIGURES 26 – 31. SEMs of head, prosternum and prosternal process in Laccomimus species: 26) L. amazonas (male), oblique lateral view; 27) idem, at higher magnification; 28) L. alvarengi (female) ventral view; 29) L. bordoni (female), ventral view; 30) idem, oblique lateral view; 31) idem, at higher magnification." httpUri-0="https://zenodo.org/record/244305/files/figure.png" httpUri-1="https://zenodo.org/record/244306/files/figure.png" pageId="4" pageNumber="305">Figs 24–27</figureCitation>
). In other genera such as 
<taxonomicName id="4C474D6BFFF7FFF09184FD2DFECBA296" box="[181,307,656,680]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF09184FD2DFECBA296" box="[181,307,656,680]" italics="true" pageId="4" pageNumber="305">Laccodytes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C474D6BFFF7FFF09070FD2CFE2DA296" box="[321,469,657,680]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF09070FD2CFE2DA296" box="[321,469,657,680]" italics="true" pageId="4" pageNumber="305">Neptosternus</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C474D6BFFF7FFF090CBFD2DFD7BA296" box="[506,643,656,680]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF090CBFD2DFD7BA296" box="[506,643,656,680]" italics="true" pageId="4" pageNumber="305">Laccophilus</emphasis>
</taxonomicName>
, the anterior rim of the prosternum is thin (
<emphasis id="B933EAFAFFF7FFF0955BFD2DFAF8A296" box="[1130,1280,656,680]" italics="true" pageId="4" pageNumber="305">character 0.0</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF7FFF091A6FD65FB9AA3BE" blockId="4.[151,1436,727,896]" pageId="4" pageNumber="305">
<emphasis id="B933EAFAFFF7FFF091A6FD65FED9A2CE" box="[151,289,728,752]" italics="true" pageId="4" pageNumber="305">Character 1</emphasis>
. Base of pronotum: (0) with median angular projection; (1) straight or almost so. In most genera of 
<taxonomicName id="4C474D6BFFF7FFF091A6FD41FEDFA32B" box="[151,295,764,789]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, including 
<taxonomicName id="4C474D6BFFF7FFF09095FD40FDCAA32A" box="[420,562,765,788]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF7FFF09095FD40FDCAA32A" box="[420,562,765,788]" italics="true" pageId="4" pageNumber="305">Laccomimus</emphasis>
</taxonomicName>
and the 
<taxonomicName id="4C474D6BFFF7FFF093A5FD40FCBCA32A" box="[660,836,764,788]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="phalacroides">
<emphasis id="B933EAFAFFF7FFF093A5FD40FCBCA32A" box="[660,836,764,788]" italics="true" pageId="4" pageNumber="305">L. phalacroides</emphasis>
</taxonomicName>
-group of 
<taxonomicName id="4C474D6BFFF7FFF09284FD41FBCBA32A" box="[949,1075,764,788]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF09284FD41FBCBA32A" box="[949,1075,764,788]" italics="true" pageId="4" pageNumber="305">Laccodytes</emphasis>
</taxonomicName>
, a median, posterior lobe of the hind margin of the pronotum projects between the elytra covering the scutellum (
<emphasis id="B933EAFAFFF7FFF09515FC9DFB42A306" box="[1060,1210,800,824]" italics="true" pageId="4" pageNumber="305">character 1.0</emphasis>
; 
<figureCitation id="137C2A6DFFF7FFF095F8FCA2FAE8A306" box="[1225,1296,799,824]" captionStart="FIGURES 1 – 7. 1" captionStartId="29.[151,264,1905,1927]" captionTargetBox="[169,1412,219,1874]" captionTargetId="figure@29.[169,1413,193,1876]" captionTargetPageId="29" captionText="FIGURES 1 – 7. 1) Habitus of Laccomimus bolivari, dorsal view (Photo A. E. Z. Short); 2) habitus of Laccodytes apalodes, dorsal view (Photo A. E. Z. Short); 3 – 7) SEMs of Laccomimus species (females): 3) habitus of L. alvarengi, dorsal view; 4) habitus of L. bordoni, dorsal view; 5) pronotum of L. alvarengi, dorsal view; 6) pronotum of L. bordoni, dorsal view; 7) pronotum of L. bordoni, dorsolateral view." httpUri="https://zenodo.org/record/244302/files/figure.png" pageId="4" pageNumber="305">Figs 5</figureCitation>
; 6). In some genera (e.g. 
<taxonomicName id="4C474D6BFFF7FFF0901AFCF8FE47A362" box="[299,447,837,860]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF0901AFCF8FE47A362" box="[299,447,837,860]" italics="true" pageId="4" pageNumber="305">Neptosternus</emphasis>
</taxonomicName>
, the 
<emphasis id="B933EAFAFFF7FFF090CEFCF8FD77A362" box="[511,655,836,860]" italics="true" pageId="4" pageNumber="305">L. apalodes-</emphasis>
group of 
<taxonomicName id="4C474D6BFFF7FFF09231FCF9FC86A362" box="[768,894,836,860]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF09231FCF9FC86A362" box="[768,894,836,860]" italics="true" pageId="4" pageNumber="305">Laccodytes</emphasis>
</taxonomicName>
) the hind margin of the pronotum is straight, lacking a visible projection, although the scutellum remains concealed (
<emphasis id="B933EAFAFFF7FFF0928AFCD5FBAAA3BE" box="[955,1106,872,896]" italics="true" pageId="4" pageNumber="305">character 1.1</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF7FFF091A6FC0DFAA0A466" blockId="4.[151,1436,943,1112]" pageId="4" pageNumber="305">
<emphasis id="B933EAFAFFF7FFF091A6FC0DFEDEA3F6" box="[151,294,944,968]" italics="true" pageId="4" pageNumber="305">Character 2</emphasis>
. Posterior angles of pronotum: (0) rounded or angulate; (1) ending in a spiniform process. In 
<taxonomicName id="4C474D6BFFF7FFF091A6FC68FED3A3D2" box="[151,299,981,1004]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF091A6FC68FED3A3D2" box="[151,299,981,1004]" italics="true" pageId="4" pageNumber="305">Neptosternus</emphasis>
</taxonomicName>
and in the 
<taxonomicName id="4C474D6BFFF7FFF09097FC68FDD0A3D2" box="[422,552,980,1004]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="B933EAFAFFF7FFF09097FC68FDD0A3D2" box="[422,552,980,1004]" italics="true" pageId="4" pageNumber="305">L. apalodes</emphasis>
</taxonomicName>
-group of 
<taxonomicName id="4C474D6BFFF7FFF093A6FC69FCEDA3D2" box="[663,789,980,1004]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF093A6FC69FCEDA3D2" box="[663,789,980,1004]" italics="true" pageId="4" pageNumber="305">Laccodytes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD64B19FFF7FFF09215FC69FC0CA3D2" author="Toledo" box="[804,1012,980,1005]" pageId="4" pageNumber="305" refString="Toledo, M., Spangler, P. J. &amp; Balke, M. (2010) Taxonomic revision of the Neotropical diving beetles genus Laccodytes Regimbart, 1895 (Coleoptera: Dytiscidae). Zootaxa, 2347, 37 - 58." type="journal article" year="2010">
Toledo 
<emphasis id="B933EAFAFFF7FFF09248FC68FC52A3D2" box="[889,938,980,1004]" italics="true" pageId="4" pageNumber="305">et al</emphasis>
. 2010
</bibRefCitation>
), the posterior angles of the pronotum end in a spiniform process, projected backward (
<emphasis id="B933EAFAFFF7FFF0938DFC45FCACA42E" box="[700,852,1016,1040]" italics="true" pageId="4" pageNumber="305">character 2.1</emphasis>
; see fig. 
<quantity id="4CBF9B0DFFF7FFF0928DFC4AFC0EA42E" box="[956,1014,1015,1040]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="4" pageNumber="305" unit="in" value="26.0">26 in</quantity>
<bibRefCitation id="EFD64B19FFF7FFF092CCFC4AFB37A42E" author="Toledo" box="[1021,1231,1015,1040]" pageId="4" pageNumber="305" refString="Toledo, M., Spangler, P. J. &amp; Balke, M. (2010) Taxonomic revision of the Neotropical diving beetles genus Laccodytes Regimbart, 1895 (Coleoptera: Dytiscidae). Zootaxa, 2347, 37 - 58." type="journal article" year="2010">
Toledo 
<emphasis id="B933EAFAFFF7FFF09563FC44FB7AA42E" box="[1106,1154,1016,1040]" italics="true" pageId="4" pageNumber="305">et al</emphasis>
. 2010
</bibRefCitation>
). In the remaining 
<taxonomicName id="4C474D6BFFF7FFF091A6FBA1FEDCA40B" box="[151,292,1052,1077]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
genera (including 
<taxonomicName id="4C474D6BFFF7FFF090CBFBA0FD7FA40A" box="[506,647,1053,1076]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF7FFF090CBFBA0FD7FA40A" box="[506,647,1053,1076]" italics="true" pageId="4" pageNumber="305">Laccomimus</emphasis>
</taxonomicName>
) and in the 
<taxonomicName id="4C474D6BFFF7FFF09220FBA0FC38A40A" box="[785,960,1052,1076]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="phalacroides">
<emphasis id="B933EAFAFFF7FFF09220FBA0FC38A40A" box="[785,960,1052,1076]" italics="true" pageId="4" pageNumber="305">L. phalacroides</emphasis>
</taxonomicName>
-group of 
<taxonomicName id="4C474D6BFFF7FFF09503FBA1FB48A40A" box="[1074,1200,1052,1076]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF09503FBA1FB48A40A" box="[1074,1200,1052,1076]" italics="true" pageId="4" pageNumber="305">Laccodytes</emphasis>
</taxonomicName>
, the posterior angles of the pronotum are rounded or slightly angulate (
<emphasis id="B933EAFAFFF7FFF093F6FBFDFCA6A466" box="[711,862,1088,1112]" italics="true" pageId="4" pageNumber="305">character 2.0</emphasis>
; 
<figureCitation id="137C2A6DFFF7FFF0925DFB82FC56A469" box="[876,942,1087,1112]" captionStart="FIGURES 1 – 7. 1" captionStartId="29.[151,264,1905,1927]" captionTargetBox="[169,1412,219,1874]" captionTargetId="figure@29.[169,1413,193,1876]" captionTargetPageId="29" captionText="FIGURES 1 – 7. 1) Habitus of Laccomimus bolivari, dorsal view (Photo A. E. Z. Short); 2) habitus of Laccodytes apalodes, dorsal view (Photo A. E. Z. Short); 3 – 7) SEMs of Laccomimus species (females): 3) habitus of L. alvarengi, dorsal view; 4) habitus of L. bordoni, dorsal view; 5) pronotum of L. alvarengi, dorsal view; 6) pronotum of L. bordoni, dorsal view; 7) pronotum of L. bordoni, dorsolateral view." httpUri="https://zenodo.org/record/244302/files/figure.png" pageId="4" pageNumber="305">Fig. 7</figureCitation>
and see fig. 
<quantity id="4CBF9B0DFFF7FFF0950DFBFDFB8DA466" box="[1084,1141,1087,1112]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="4" pageNumber="305" unit="in" value="27.0">27 in</quantity>
<bibRefCitation id="EFD64B19FFF7FFF0954DFB82FAB4A466" author="Toledo" box="[1148,1356,1087,1112]" pageId="4" pageNumber="305" refString="Toledo, M., Spangler, P. J. &amp; Balke, M. (2010) Taxonomic revision of the Neotropical diving beetles genus Laccodytes Regimbart, 1895 (Coleoptera: Dytiscidae). Zootaxa, 2347, 37 - 58." type="journal article" year="2010">
Toledo 
<emphasis id="B933EAFAFFF7FFF095E1FBFCFAF9A466" box="[1232,1281,1088,1112]" italics="true" pageId="4" pageNumber="305">et al</emphasis>
. 2010
</bibRefCitation>
).
</paragraph>
<paragraph id="8BF836E8FFF7FFF091A6FB35FD29A636" blockId="4.[151,1436,1159,1544]" pageId="4" pageNumber="305">
<emphasis id="B933EAFAFFF7FFF091A6FB35FED9A49E" box="[151,289,1160,1184]" italics="true" pageId="4" pageNumber="305">Character 3</emphasis>
. Prosternal process (ventral view): (0) elongate, reaching mesocoxae but not extending beyond their hind margins; (1) very long, extending beyond hind margins of mesocoxae; (2) broad, not or barely reaching mesocoxae. In 
<taxonomicName id="4C474D6BFFF7FFF09062FB6DFE24A4D6" box="[339,476,1232,1256]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF09062FB6DFE24A4D6" box="[339,476,1232,1256]" italics="true" pageId="4" pageNumber="305">Laccophilus</emphasis>
</taxonomicName>
, the 
<taxonomicName id="4C474D6BFFF7FFF09316FB6DFD40A4D6" box="[551,696,1232,1256]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="phalacroides">
<emphasis id="B933EAFAFFF7FFF09316FB6DFD40A4D6" box="[551,696,1232,1256]" italics="true" pageId="4" pageNumber="305">phalacroides</emphasis>
</taxonomicName>
-group of 
<taxonomicName id="4C474D6BFFF7FFF0920AFB6DFC41A4D6" box="[827,953,1232,1256]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF0920AFB6DFC41A4D6" box="[827,953,1232,1256]" italics="true" pageId="4" pageNumber="305">Laccodytes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD64B19FFF7FFF092E3FB72FB38A4D6" author="Toledo" box="[978,1216,1231,1256]" pageId="4" pageNumber="305" refString="Toledo, M., Spangler, P. J. &amp; Balke, M. (2010) Taxonomic revision of the Neotropical diving beetles genus Laccodytes Regimbart, 1895 (Coleoptera: Dytiscidae). Zootaxa, 2347, 37 - 58." type="journal article" year="2010">
Toledo 
<emphasis id="B933EAFAFFF7FFF09503FB6CFB94A4D6" box="[1074,1132,1232,1256]" italics="true" pageId="4" pageNumber="305">et al</emphasis>
. 2010
</bibRefCitation>
), and some other 
<taxonomicName id="4C474D6BFFF7FFF091A6FB49FEDCA533" box="[151,292,1268,1293]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
genera, the prosternal process is elongate, narrow, with its apex extending between the mesocoxae (at most extending a little over the hind margins of the mesocoxae as in some species of 
<taxonomicName id="4C474D6BFFF7FFF0956BFAA5FB1BA50E" box="[1114,1251,1304,1328]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF0956BFAA5FB1BA50E" box="[1114,1251,1304,1328]" italics="true" pageId="4" pageNumber="305">Laccophilus</emphasis>
</taxonomicName>
) (
<emphasis id="B933EAFAFFF7FFF095CCFAA5FA6CA50E" box="[1277,1428,1304,1328]" italics="true" pageId="4" pageNumber="305">character 3.0</emphasis>
; see figs 6, 8 in 
<bibRefCitation id="EFD64B19FFF7FFF09074FA81FDECA56A" author="Toledo" box="[325,532,1340,1365]" pageId="4" pageNumber="305" refString="Toledo, M., Spangler, P. J. &amp; Balke, M. (2010) Taxonomic revision of the Neotropical diving beetles genus Laccodytes Regimbart, 1895 (Coleoptera: Dytiscidae). Zootaxa, 2347, 37 - 58." type="journal article" year="2010">
Toledo 
<emphasis id="B933EAFAFFF7FFF090AAFA80FE34A56A" box="[411,460,1340,1364]" italics="true" pageId="4" pageNumber="305">et al</emphasis>
. 2010
</bibRefCitation>
and figs 
<quantity id="4CBF9B0DFFF7FFF0934EFA81FD07A56B" box="[639,767,1340,1365]" metricMagnitude="0" metricUnit="m" metricValue="2.5907999999999998" metricValueMax="2.6162" metricValueMin="2.5654" pageId="4" pageNumber="305" unit="in" value="102.0" valueMax="103.0" valueMin="101.0">101–103 in</quantity>
<bibRefCitation id="EFD64B19FFF7FFF09236FA81FC32A56B" author="Brancucci" box="[775,970,1340,1365]" pageId="4" pageNumber="305" refString="Brancucci, M. (1983 b) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col., Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen, 31 - 32, 241 - 426." type="book chapter" year="1983" yearSuffix="b">Brancucci 1983b</bibRefCitation>
). In the 
<taxonomicName id="4C474D6BFFF7FFF09519FA80FB54A56A" box="[1064,1196,1340,1364]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="B933EAFAFFF7FFF09519FA80FB54A56A" box="[1064,1196,1340,1364]" italics="true" pageId="4" pageNumber="305">L. apalodes</emphasis>
</taxonomicName>
-group of 
<taxonomicName id="4C474D6BFFF7FFF0942FFA81FA64A56A" box="[1310,1436,1340,1364]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF0942FFA81FA64A56A" box="[1310,1436,1340,1364]" italics="true" pageId="4" pageNumber="305">Laccodytes</emphasis>
</taxonomicName>
and in 
<taxonomicName id="4C474D6BFFF7FFF091D5FADCFE80A546" box="[228,376,1377,1400]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF091D5FADCFE80A546" box="[228,376,1377,1400]" italics="true" pageId="4" pageNumber="305">Neptosternus</emphasis>
</taxonomicName>
, the prosternal process ends in a long, almost needle-like apex, visibly going beyond the hind margins of the mesocoxae (
<emphasis id="B933EAFAFFF7FFF090D0FA39FD86A5A2" box="[481,638,1412,1436]" italics="true" pageId="4" pageNumber="305">character 3.1</emphasis>
; see figs 4, 7 in 
<bibRefCitation id="EFD64B19FFF7FFF09267FA39FBCFA5A2" author="Toledo" box="[854,1079,1412,1437]" pageId="4" pageNumber="305" refString="Toledo, M., Spangler, P. J. &amp; Balke, M. (2010) Taxonomic revision of the Neotropical diving beetles genus Laccodytes Regimbart, 1895 (Coleoptera: Dytiscidae). Zootaxa, 2347, 37 - 58." type="journal article" year="2010">
Toledo 
<emphasis id="B933EAFAFFF7FFF09281FA38FC1FA5A2" box="[944,999,1412,1436]" italics="true" pageId="4" pageNumber="305">et al</emphasis>
. 2010
</bibRefCitation>
). Finally, in 
<taxonomicName id="4C474D6BFFF7FFF095E3FA38FA9FA5A2" box="[1234,1383,1413,1436]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF095E3FA38FA9FA5A2" box="[1234,1383,1413,1436]" italics="true" pageId="4" pageNumber="305">Laccosternus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF7FFF091A6FA15FEDDA581" box="[151,293,1448,1471]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF7FFF091A6FA15FEDDA581" box="[151,293,1448,1471]" italics="true" pageId="4" pageNumber="305">Laccomimus</emphasis>
</taxonomicName>
the prosternal process is short and broad, with its apex not or barely reaching the mesocoxae (
<emphasis id="B933EAFAFFF7FFF09191FA71FECFA5DA" box="[160,311,1484,1508]" italics="true" pageId="4" pageNumber="305">character 3.2</emphasis>
; 
<figureCitation id="137C2A6DFFF7FFF09074FA71FE3EA5DA" box="[325,454,1484,1509]" captionStart-0="FIGURES 20 – 25" captionStart-1="FIGURES 26 – 31" captionStartId-0="32.[151,264,1953,1975]" captionStartId-1="33.[151,264,1953,1975]" captionTargetBox-0="[151,1436,193,1930]" captionTargetBox-1="[151,1436,193,1930]" captionTargetId-0="figure@32.[151,1436,193,1932]" captionTargetId-1="figure@33.[151,1436,193,1932]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionText-0="FIGURES 20 – 25. SEMs of head, prosternum and prosternal process in Laccomimus species: 20) L. distinctus (male), ventral view, 21) idem, at higher magnification; 22) idem, oblique lateral view; 23) L. malkini (female) ventral view; 24) L. amazonas (male), ventral view (arrow indicates protruding prosternum); 25) idem, at higher magnification." captionText-1="FIGURES 26 – 31. SEMs of head, prosternum and prosternal process in Laccomimus species: 26) L. amazonas (male), oblique lateral view; 27) idem, at higher magnification; 28) L. alvarengi (female) ventral view; 29) L. bordoni (female), ventral view; 30) idem, oblique lateral view; 31) idem, at higher magnification." httpUri-0="https://zenodo.org/record/244305/files/figure.png" httpUri-1="https://zenodo.org/record/244306/files/figure.png" pageId="4" pageNumber="305">Figs 23–31</figureCitation>
); the only exception occurs in 
<taxonomicName id="4C474D6BFFF7FFF0922AFA70FBE2A5DA" box="[795,1050,1484,1508]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B933EAFAFFF7FFF0922AFA70FBE2A5DA" box="[795,1050,1484,1508]" italics="true" pageId="4" pageNumber="305">Laccomimus distinctus</emphasis>
</taxonomicName>
, in which the prosternal process is almost similar to that of 
<taxonomicName id="4C474D6BFFF7FFF09098FA4DFDCAA636" box="[425,562,1520,1544]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF09098FA4DFDCAA636" box="[425,562,1520,1544]" italics="true" pageId="4" pageNumber="305">Laccophilus</emphasis>
</taxonomicName>
(
<figureCitation id="137C2A6DFFF7FFF09370FA52FD3DA636" box="[577,709,1519,1544]" captionStart="FIGURES 20 – 25" captionStartId="32.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@32.[151,1436,193,1932]" captionTargetPageId="32" captionText="FIGURES 20 – 25. SEMs of head, prosternum and prosternal process in Laccomimus species: 20) L. distinctus (male), ventral view, 21) idem, at higher magnification; 22) idem, oblique lateral view; 23) L. malkini (female) ventral view; 24) L. amazonas (male), ventral view (arrow indicates protruding prosternum); 25) idem, at higher magnification." httpUri="https://zenodo.org/record/244305/files/figure.png" pageId="4" pageNumber="305">Figs 20–22</figureCitation>
).
</paragraph>
<paragraph id="8BF836E8FFF7FFF091A6F985FCF9A6A6" blockId="4.[151,1436,1591,1688]" pageId="4" pageNumber="305">
<emphasis id="B933EAFAFFF7FFF091A6F985FED9A66E" box="[151,289,1592,1616]" italics="true" pageId="4" pageNumber="305">Character 4</emphasis>
. Prosternal process (ventral view): (0) simple; (1) trifid. Members of the genus 
<taxonomicName id="4C474D6BFFF7FFF09588F984FAB5A66E" box="[1209,1357,1593,1616]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF09588F984FAB5A66E" box="[1209,1357,1593,1616]" italics="true" pageId="4" pageNumber="305">Neptosternus</emphasis>
</taxonomicName>
bear a peculiar three pointed prosternal process, resembling a fork (
<emphasis id="B933EAFAFFF7FFF09278F9E1FC18A64A" box="[841,992,1628,1652]" italics="true" pageId="4" pageNumber="305">character 4.1</emphasis>
). In the remaining 
<taxonomicName id="4C474D6BFFF7FFF0958DF9E1FAB1A64B" box="[1212,1353,1628,1653]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
genera the prosternal process is single pointed (
<emphasis id="B933EAFAFFF7FFF0936BF93DFD08A6A6" box="[602,752,1664,1688]" italics="true" pageId="4" pageNumber="305">character 4.0</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF7FFF091A6F975FA64A7E2" blockId="4.[151,1436,1735,2013]" pageId="4" pageNumber="305">
<emphasis id="B933EAFAFFF7FFF091A6F975FED9A6E1" box="[151,289,1736,1760]" italics="true" pageId="4" pageNumber="305">Character 5</emphasis>
. Prosternal process (lateral view): (0) elevated or carinate, visibly reaching and positioned on same plane as metaventrite; (1) elevated or carinate, barely reaching and positioned on same plane as metaventrite; (2) saddle-shaped, barely reaching and positioned below metaventrite. In most genera of 
<taxonomicName id="4C474D6BFFF7FFF0955BF8B2FB0FA716" box="[1130,1271,1807,1832]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
the prosternal process in lateral view is elevated or even carinate, and its apex clearly reaches the metaventrite between the mesocoxae. It is also positioned on the same frontal plane as the mesocoxae (
<emphasis id="B933EAFAFFF7FFF092C0F8E5FB7FA74E" box="[1009,1159,1880,1904]" italics="true" pageId="4" pageNumber="305">character 5.0</emphasis>
). In 
<taxonomicName id="4C474D6BFFF7FFF0958DF8E4FAA9A74E" box="[1212,1361,1881,1904]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF7FFF0958DF8E4FAA9A74E" box="[1212,1361,1881,1904]" italics="true" pageId="4" pageNumber="305">Laccosternus</emphasis>
</taxonomicName>
and in most species of 
<taxonomicName id="4C474D6BFFF7FFF09060F8C0FE27A7AA" box="[337,479,1917,1940]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF7FFF09060F8C0FE27A7AA" box="[337,479,1917,1940]" italics="true" pageId="4" pageNumber="305">Laccomimus</emphasis>
</taxonomicName>
the situation is similar but the apex barely reaches the metaventrite (
<emphasis id="B933EAFAFFF7FFF095CAF8C1FA6CA7AA" box="[1275,1428,1916,1940]" italics="true" pageId="4" pageNumber="305">character 5.1</emphasis>
; 
<figureCitation id="137C2A6DFFF7FFF091A6F81DFF17A786" box="[151,239,1952,1977]" captionStart="FIGURES 20 – 25" captionStartId="32.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@32.[151,1436,193,1932]" captionTargetPageId="32" captionText="FIGURES 20 – 25. SEMs of head, prosternum and prosternal process in Laccomimus species: 20) L. distinctus (male), ventral view, 21) idem, at higher magnification; 22) idem, oblique lateral view; 23) L. malkini (female) ventral view; 24) L. amazonas (male), ventral view (arrow indicates protruding prosternum); 25) idem, at higher magnification." httpUri="https://zenodo.org/record/244305/files/figure.png" pageId="4" pageNumber="305">Figs 23</figureCitation>
; 28; 29–31), with the exception of 
<taxonomicName id="4C474D6BFFF7FFF093B3F81CFC7BA786" box="[642,899,1952,1976]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B933EAFAFFF7FFF093B3F81CFC7BA786" box="[642,899,1952,1976]" italics="true" pageId="4" pageNumber="305">Laccomimus distinctus</emphasis>
</taxonomicName>
in which the prosternal process in lateral view looks more like that described for 
<emphasis id="B933EAFAFFF7FFF09313F879FD42A7E2" box="[546,698,1988,2012]" italics="true" pageId="4" pageNumber="305">character 5.0</emphasis>
(
<figureCitation id="137C2A6DFFF7FFF093FAF879FCA9A7E2" box="[715,849,1988,2013]" captionStart="FIGURES 20 – 25" captionStartId="32.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@32.[151,1436,193,1932]" captionTargetPageId="32" captionText="FIGURES 20 – 25. SEMs of head, prosternum and prosternal process in Laccomimus species: 20) L. distinctus (male), ventral view, 21) idem, at higher magnification; 22) idem, oblique lateral view; 23) L. malkini (female) ventral view; 24) L. amazonas (male), ventral view (arrow indicates protruding prosternum); 25) idem, at higher magnification." httpUri="https://zenodo.org/record/244305/files/figure.png" pageId="4" pageNumber="305">Figs 20–22</figureCitation>
). In a few species of 
<taxonomicName id="4C474D6BFFF7FFF09577F878FB2CA7E2" box="[1094,1236,1989,2012]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF7FFF09577F878FB2CA7E2" box="[1094,1236,1989,2012]" italics="true" pageId="4" pageNumber="305">Laccomimus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C474D6BFFF7FFF095D7F878FA8DA7E2" box="[1254,1397,1989,2012]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="amazonas">
<emphasis id="B933EAFAFFF7FFF095D7F878FA8DA7E2" box="[1254,1397,1989,2012]" italics="true" pageId="4" pageNumber="305">L. amazonas</emphasis>
</taxonomicName>
, 
<emphasis id="B933EAFAFFF7FFF094B4F878FA64A7E2" box="[1413,1436,1989,2012]" italics="true" pageId="4" pageNumber="305">L.</emphasis>
</paragraph>
<paragraph id="8BF836E8FFF6FFF191A6FF25FF0FA0C6" blockId="5.[151,1436,151,248]" pageId="5" pageNumber="306">
<taxonomicName id="4C474D6BFFF6FFF191A6FF25FF0FA091" box="[151,247,152,175]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B933EAFAFFF6FFF191A6FF25FF0FA091" box="[151,247,152,175]" italics="true" pageId="5" pageNumber="306">spinosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C474D6BFFF6FFF19036FF24FE64A091" box="[263,412,152,176]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="B933EAFAFFF6FFF19036FF24FE64A091" box="[263,412,152,176]" italics="true" pageId="5" pageNumber="306">L. variegatus</emphasis>
</taxonomicName>
), the prosternal process can be thought as saddle-shaped, i.e. the outline of the process is strongly sinuate in lateral view and the apex is bent upward, not or barely reaching the metaventrite (
<emphasis id="B933EAFAFFF6FFF195CCFF01FA6CA0EA" box="[1277,1428,188,212]" italics="true" pageId="5" pageNumber="306">character 5.2</emphasis>
; 
<figureCitation id="137C2A6DFFF6FFF191A6FF62FF12A0C6" box="[151,234,223,248]" captionStart="FIGURES 26 – 31" captionStartId="33.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@33.[151,1436,193,1932]" captionTargetPageId="33" captionText="FIGURES 26 – 31. SEMs of head, prosternum and prosternal process in Laccomimus species: 26) L. amazonas (male), oblique lateral view; 27) idem, at higher magnification; 28) L. alvarengi (female) ventral view; 29) L. bordoni (female), ventral view; 30) idem, oblique lateral view; 31) idem, at higher magnification." httpUri="https://zenodo.org/record/244306/files/figure.png" pageId="5" pageNumber="306">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="8BF836E8FFF6FFF191A6FE95FB0DA1EE" blockId="5.[151,1436,295,464]" pageId="5" pageNumber="306">
<emphasis id="B933EAFAFFF6FFF191A6FE95FED9A17E" box="[151,289,296,320]" italics="true" pageId="5" pageNumber="306">Character 6</emphasis>
. Strong and sparse punctures on elytra: (0) absent; (1) present. Both species of 
<taxonomicName id="4C474D6BFFF6FFF1958BFE94FAB7A17E" box="[1210,1359,297,320]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF1958BFE94FAB7A17E" box="[1210,1359,297,320]" italics="true" pageId="5" pageNumber="306">Laccosternus</emphasis>
</taxonomicName>
(more evident in 
<taxonomicName id="4C474D6BFFF6FFF1903DFEF0FD8FA15B" authority="Regimbart, 1895" authorityName="Regimbart" authorityYear="1895" box="[268,631,332,357]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="grouvellei">
<emphasis id="B933EAFAFFF6FFF1903DFEF0FE61A15A" box="[268,409,332,356]" italics="true" pageId="5" pageNumber="306">L. grouvellei</emphasis>
<bibRefCitation id="EFD64B19FFF6FFF19091FEF1FD8FA15B" author="Regimbart" box="[416,631,332,357]" pageId="5" pageNumber="306" refString="Regimbart, M. (1895) Dytiscides trouves dans les tabacs par les soins de M. Antoine Grouvelle. Annales de la Societe Entomologique de France, 64, 321 - 348, 1 pl." type="journal article" year="1895">(Régimbart, 1895))</bibRefCitation>
</taxonomicName>
share a strong punctation of sparsely distributed punctures on the whole elytral surface (
<emphasis id="B933EAFAFFF6FFF19069FECDFE00A1B6" box="[344,504,368,392]" italics="true" pageId="5" pageNumber="306">character 6.1</emphasis>
). A strong and diffuse punctation on the elytra was not observed in other 
<taxonomicName id="4C474D6BFFF6FFF191A6FE29FEDFA193" box="[151,295,404,429]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, with the exception of the longitudinal series of punctures (more or less clearly impressed, and usually less regular and somewhat sparse in posterior half) commonly present in all species (
<emphasis id="B933EAFAFFF6FFF1957FFE05FB1DA1EE" box="[1102,1253,440,464]" italics="true" pageId="5" pageNumber="306">character 6.0</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF6FFF191A6FDBDFA64A2BA" blockId="5.[151,1436,511,645]" pageId="5" pageNumber="306">
<emphasis id="B933EAFAFFF6FFF191A6FDBDFEDCA229" box="[151,292,512,536]" italics="true" pageId="5" pageNumber="306">Character 7</emphasis>
. Epipleural carina: (0) not visible dorsally; (1) visible dorsally. In 
<taxonomicName id="4C474D6BFFF6FFF19572FDBCFB2FA226" box="[1091,1239,513,536]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF19572FDBCFB2FA226" box="[1091,1239,513,536]" italics="true" pageId="5" pageNumber="306">Neptosternus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C474D6BFFF6FFF195DBFDBDFA90A226" box="[1258,1384,512,536]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF195DBFDBDFA90A226" box="[1258,1384,512,536]" italics="true" pageId="5" pageNumber="306">Laccodytes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF6FFF191A6FD99FEF5A202" box="[151,269,548,572]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF191A6FD99FEF5A202" box="[151,269,548,572]" italics="true" pageId="5" pageNumber="306">Napodytes</emphasis>
</taxonomicName>
the carina between the elytron and epipleuron is visible in dorsal view. This gives the appearance of a border running along the lateral side of each elytron (
<emphasis id="B933EAFAFFF6FFF193C3FDF5FC71A25E" box="[754,905,584,608]" italics="true" pageId="5" pageNumber="306">character 7.1</emphasis>
). In all other genera of 
<taxonomicName id="4C474D6BFFF6FFF195A7FDFAFADEA25E" box="[1174,1318,583,608]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, including 
<taxonomicName id="4C474D6BFFF6FFF191A6FDD0FEDCA2BA" box="[151,292,621,644]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF6FFF191A6FDD0FEDCA2BA" box="[151,292,621,644]" italics="true" pageId="5" pageNumber="306">Laccomimus</emphasis>
</taxonomicName>
, this carina is not visible and the elytra do not appear laterally bordered in dorsal view (
<emphasis id="B933EAFAFFF6FFF195C4FDD1FA74A2BA" box="[1269,1420,620,644]" italics="true" pageId="5" pageNumber="306">character 7.0</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF6FFF191A6FD09FA80A32A" blockId="5.[151,1436,692,789]" pageId="5" pageNumber="306">
<emphasis id="B933EAFAFFF6FFF191A6FD09FEDAA2F2" box="[151,290,692,716]" italics="true" pageId="5" pageNumber="306">Character 8</emphasis>
. Silky sheen, mainly on elytra and metatarsi: (0) absent; (1) present. Members of 
<taxonomicName id="4C474D6BFFF6FFF195D3FD08FA88A2F2" box="[1250,1392,693,716]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF6FFF195D3FD08FA88A2F2" box="[1250,1392,693,716]" italics="true" pageId="5" pageNumber="306">Laccomimus</emphasis>
</taxonomicName>
are unique within the 
<taxonomicName id="4C474D6BFFF6FFF19053FD6AFE17A2CE" box="[354,495,727,752]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
genera studied in the presence of a characteristic silky sheen which is more evident on the elytra and metatarsi (
<emphasis id="B933EAFAFFF6FFF190E1FD41FD9FA32A" box="[464,615,764,788]" italics="true" pageId="5" pageNumber="306">character 8.1</emphasis>
). This sheen is absent in other genera of 
<taxonomicName id="4C474D6BFFF6FFF19502FD41FB38A32B" box="[1075,1216,764,789]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
(
<emphasis id="B933EAFAFFF6FFF195E0FD41FA90A32A" box="[1233,1384,764,788]" italics="true" pageId="5" pageNumber="306">character 8.0</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF6FFF191A6FCF9FF3BA3D2" blockId="5.[151,1436,836,1004]" pageId="5" pageNumber="306">
<emphasis id="B933EAFAFFF6FFF191A6FCF9FEDCA362" box="[151,292,836,860]" italics="true" pageId="5" pageNumber="306">Character 9</emphasis>
. Metacoxal lines: (0) sinuate, converging anteriorly; (1) straight, subparallel. In some genera of 
<taxonomicName id="4C474D6BFFF6FFF191A6FCDAFEDFA3BE" box="[151,295,871,896]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, including 
<taxonomicName id="4C474D6BFFF6FFF19099FCD4FDC4A3BE" box="[424,572,873,896]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF19099FCD4FDC4A3BE" box="[424,572,873,896]" italics="true" pageId="5" pageNumber="306">Neptosternus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF6FFF19349FCD5FD0EA3BE" box="[632,758,872,896]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF19349FCD5FD0EA3BE" box="[632,758,872,896]" italics="true" pageId="5" pageNumber="306">Laccodytes</emphasis>
</taxonomicName>
, the metacoxal lines are straight and parallel, or almost so (
<emphasis id="B933EAFAFFF6FFF19191FC31FEC0A39A" box="[160,312,908,932]" italics="true" pageId="5" pageNumber="306">character 9.1</emphasis>
; see figs 
<quantity id="4CBF9B0DFFF6FFF19096FC31FE04A39B" box="[423,508,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="5" pageNumber="306" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">9–11 in</quantity>
<bibRefCitation id="EFD64B19FFF6FFF19335FC31FD20A39A" author="Toledo" box="[516,728,908,933]" pageId="5" pageNumber="306" refString="Toledo, M., Spangler, P. J. &amp; Balke, M. (2010) Taxonomic revision of the Neotropical diving beetles genus Laccodytes Regimbart, 1895 (Coleoptera: Dytiscidae). Zootaxa, 2347, 37 - 58." type="journal article" year="2010">
Toledo 
<emphasis id="B933EAFAFFF6FFF1936BFC30FD74A39A" box="[602,652,908,932]" italics="true" pageId="5" pageNumber="306">et al</emphasis>
. 2010
</bibRefCitation>
). In other genera the metacoxal lines are sinuate and converge anteriorly at the level of the suture separating the metacoxae and the metaventrite (
<emphasis id="B933EAFAFFF6FFF19578FC0DFB19A3F6" box="[1097,1249,944,968]" italics="true" pageId="5" pageNumber="306">character 9.0</emphasis>
; 
<figureCitation id="137C2A6DFFF6FFF195C0FC12FA9CA3F6" box="[1265,1380,943,968]" captionStart="FIGURES 8 – 13" captionStartId="30.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@30.[151,1436,193,1932]" captionTargetPageId="30" captionText="FIGURES 8 – 13. SEMs of Laccomimus species, ventral view: 8) L. distinctus (male); 9) L. malkini (female); 10) L. amazonas (male); 11) L. alvarengi (female); 12) L. bordoni (male); 13) L. bolivari (female)." httpUri="https://zenodo.org/record/244303/files/figure.png" pageId="5" pageNumber="306">Figs 8–13</figureCitation>
; 32– 41).
</paragraph>
<paragraph id="8BF836E8FFF6FFF191A6FBA1FCC5A49E" blockId="5.[151,1436,1052,1184]" pageId="5" pageNumber="306">
<emphasis id="B933EAFAFFF6FFF191A6FBA1FEC9A40A" box="[151,305,1052,1076]" italics="true" pageId="5" pageNumber="306">Character 10</emphasis>
. Lobes of metacoxal processes: (0) separated by a deep notch; (1) notch almost absent. In most genera of 
<taxonomicName id="4C474D6BFFF6FFF19034FB82FE6DA466" box="[261,405,1087,1112]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, including 
<taxonomicName id="4C474D6BFFF6FFF19321FBFDFD65A469" box="[528,669,1088,1111]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF6FFF19321FBFDFD65A469" box="[528,669,1088,1111]" italics="true" pageId="5" pageNumber="306">Laccomimus</emphasis>
</taxonomicName>
, the lobes of the metacoxal processes are separated by a more or less deep notch (
<emphasis id="B933EAFAFFF6FFF19014FBD9FE34A442" box="[293,460,1124,1148]" italics="true" pageId="5" pageNumber="306">character 10.0</emphasis>
; 
<figureCitation id="137C2A6DFFF6FFF190EDFBD9FD98A442" box="[476,608,1124,1149]" captionStart-0="FIGURES 32 – 37" captionStart-1="FIGURES 38 – 43" captionStartId-0="34.[151,264,1953,1975]" captionStartId-1="35.[151,264,1953,1975]" captionTargetBox-0="[151,1436,193,1930]" captionTargetBox-1="[151,1436,193,1930]" captionTargetId-0="figure@34.[151,1436,193,1932]" captionTargetId-1="figure@35.[151,1436,193,1932]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionText-0="FIGURES 32 – 37. SEMs of metacoxal processes in Laccomimus species, ventral view: 32) L. distinctus (male); 33 – 36) L. distinctus (females), variability in development of apical angulation in different specimens; 37) L. malkini (female)." captionText-1="FIGURES 38 – 43. SEMs of Laccomimus species, ventral view. 38 – 41) metacoxal processes: 38) L. amazonas (male); 39) L. alvarengi (female); 40) L. bordoni (female); 41) L. bolivari (female); 42 – 43) abdomen on L. bordoni, showing sexual difference in last ventrite: 42) male (arrow indicates slightly sinuate caudal margins); 43) female." httpUri-0="https://zenodo.org/record/244307/files/figure.png" httpUri-1="https://zenodo.org/record/244308/files/figure.png" pageId="5" pageNumber="306">Figs 32–41</figureCitation>
). In 
<taxonomicName id="4C474D6BFFF6FFF193A7FBD9FCE7A442" box="[662,799,1124,1148]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF193A7FBD9FCE7A442" box="[662,799,1124,1148]" italics="true" pageId="5" pageNumber="306">Laccophilus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF6FFF19268FBD9FC37A442" box="[857,975,1124,1148]" class="Insecta" family="Dytiscidae" genus="Philodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF19268FBD9FC37A442" box="[857,975,1124,1148]" italics="true" pageId="5" pageNumber="306">Philodytes</emphasis>
</taxonomicName>
these lobes are very close to each other (
<emphasis id="B933EAFAFFF6FFF19191FB35FEBDA49E" box="[160,325,1160,1184]" italics="true" pageId="5" pageNumber="306">character 10.1</emphasis>
; see figs 
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<bibRefCitation id="EFD64B19FFF6FFF1935CFB3AFCC8A49E" author="Brancucci" box="[621,816,1159,1184]" pageId="5" pageNumber="306" refString="Brancucci, M. (1983 b) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col., Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen, 31 - 32, 241 - 426." type="book chapter" year="1983" yearSuffix="b">Brancucci 1983b</bibRefCitation>
).
</paragraph>
<paragraph id="8BF836E8FFF6FFF191A6FB6DFC63A5A3" blockId="5.[151,1436,1231,1437]" pageId="5" pageNumber="306">
<emphasis id="B933EAFAFFF6FFF191A6FB6DFED4A4D6" box="[151,300,1232,1256]" italics="true" pageId="5" pageNumber="306">Character 11</emphasis>
. Lobes of metacoxal processes: (0) rounded to slightly angulate; (1) strongly angulate to acuminate; (2) truncate. Within the studied genera of 
<taxonomicName id="4C474D6BFFF6FFF193BFFB49FCE6A533" box="[654,798,1268,1293]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, 
<taxonomicName id="4C474D6BFFF6FFF1921EFB48FC3CA532" box="[815,964,1269,1292]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF1921EFB48FC3CA532" box="[815,964,1269,1292]" italics="true" pageId="5" pageNumber="306">Laccosternus</emphasis>
</taxonomicName>
and most species of 
<taxonomicName id="4C474D6BFFF6FFF195F6FB48FAACA532" box="[1223,1364,1269,1292]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF6FFF195F6FB48FAACA532" box="[1223,1364,1269,1292]" italics="true" pageId="5" pageNumber="306">Laccomimus</emphasis>
</taxonomicName>
share rounded or slightly angulate lobes of the metacoxal processes (
<emphasis id="B933EAFAFFF6FFF1926BFAA5FC04A50E" box="[858,1020,1304,1328]" italics="true" pageId="5" pageNumber="306">character 11.0</emphasis>
; 
<figureCitation id="137C2A6DFFF6FFF1953DFAAAFB9AA50E" box="[1036,1122,1303,1328]" captionStart="FIGURES 32 – 37" captionStartId="34.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@34.[151,1436,193,1932]" captionTargetPageId="34" captionText="FIGURES 32 – 37. SEMs of metacoxal processes in Laccomimus species, ventral view: 32) L. distinctus (male); 33 – 36) L. distinctus (females), variability in development of apical angulation in different specimens; 37) L. malkini (female)." httpUri="https://zenodo.org/record/244307/files/figure.png" pageId="5" pageNumber="306">Figs 37</figureCitation>
; 39–41). In some species of 
<taxonomicName id="4C474D6BFFF6FFF191A6FA80FEDDA56A" box="[151,293,1341,1364]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF6FFF191A6FA80FEDDA56A" box="[151,293,1341,1364]" italics="true" pageId="5" pageNumber="306">Laccomimus</emphasis>
</taxonomicName>
the lobes end in a sharp spine in both sexes or in a sort of spine in males and in a more or less marked angulation in females (
<emphasis id="B933EAFAFFF6FFF190AEFADDFDBCA546" box="[415,580,1376,1400]" italics="true" pageId="5" pageNumber="306">character 11.1</emphasis>
; 
<figureCitation id="137C2A6DFFF6FFF19367FAE2FD56A546" box="[598,686,1375,1400]" captionStart="FIGURES 32 – 37" captionStartId="34.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@34.[151,1436,193,1932]" captionTargetPageId="34" captionText="FIGURES 32 – 37. SEMs of metacoxal processes in Laccomimus species, ventral view: 32) L. distinctus (male); 33 – 36) L. distinctus (females), variability in development of apical angulation in different specimens; 37) L. malkini (female)." httpUri="https://zenodo.org/record/244307/files/figure.png" pageId="5" pageNumber="306">Figs 32</figureCitation>
; 33–36; 38). In 
<taxonomicName id="4C474D6BFFF6FFF1925BFADDFC0BA546" box="[874,1011,1376,1400]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF1925BFADDFC0BA546" box="[874,1011,1376,1400]" italics="true" pageId="5" pageNumber="306">Laccophilus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF6FFF1951FFADDFB5DA546" box="[1070,1189,1376,1400]" class="Insecta" family="Dytiscidae" genus="Philodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF1951FFADDFB5DA546" box="[1070,1189,1376,1400]" italics="true" pageId="5" pageNumber="306">Philodytes</emphasis>
</taxonomicName>
the lobes are distally truncate (
<emphasis id="B933EAFAFFF6FFF19030FA39FE5CA5A2" box="[257,420,1412,1436]" italics="true" pageId="5" pageNumber="306">character 11.2</emphasis>
; see figs 
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<bibRefCitation id="EFD64B19FFF6FFF193FDFA39FC77A5A3" author="Brancucci" box="[716,911,1412,1437]" pageId="5" pageNumber="306" refString="Brancucci, M. (1983 b) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col., Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen, 31 - 32, 241 - 426." type="book chapter" year="1983" yearSuffix="b">Brancucci 1983b</bibRefCitation>
).
</paragraph>
<paragraph id="8BF836E8FFF6FFF191A6FA71FF20A64A" blockId="5.[151,1436,1484,1652]" pageId="5" pageNumber="306">
<emphasis id="B933EAFAFFF6FFF191A6FA71FECAA5DA" box="[151,306,1484,1508]" italics="true" pageId="5" pageNumber="306">Character 12</emphasis>
. Larger mesotibial spur: (0) not longer than first two tarsomeres; (1) almost as long as whole mesotarsus. In 
<taxonomicName id="4C474D6BFFF6FFF19079FA4DFE2EA639" box="[328,470,1520,1543]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF6FFF19079FA4DFE2EA639" box="[328,470,1520,1543]" italics="true" pageId="5" pageNumber="306">Laccomimus</emphasis>
</taxonomicName>
the mesotibial spurs are very long, the larger one is almost as long as the whole mesotarsus, excluding the claws (
<emphasis id="B933EAFAFFF6FFF1931AF9A9FD2EA612" box="[555,726,1556,1580]" italics="true" pageId="5" pageNumber="306">character 12.1</emphasis>
; 
<figureCitation id="137C2A6DFFF6FFF193DDF9A9FCB1A613" box="[748,841,1556,1581]" captionStart="FIGURES 52 – 58" captionStartId="37.[151,264,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@37.[151,1436,193,1856]" captionTargetPageId="37" captionText="FIGURES 52 – 58. SEMs of legs in Laccomimus species: 52) L. bordoni (female), left mesothoracic leg, dorsal view; 53) L. bordoni (male), left mesotarsus (arrow indicates mesotibial spurs); 54) L. bordoni (female), right metathoracic leg, ventral view; 55) L. bordoni (male), left metathoracic leg, dorsal view; 56) L. bordoni (female), right metatarsus, ventral view (arrow indicates combs of flat spines on posterolateral lobes); 57) L. alvarengi (female), metatarsi, ventral view; 58) L. bolivari (female), metathoracic leg, dorsal view (arrow indicates posterolateral lobes of metatarsus)." httpUri="https://zenodo.org/record/244310/files/figure.png" pageId="5" pageNumber="306">Figs 52</figureCitation>
; 53). In all other genera of 
<taxonomicName id="4C474D6BFFF6FFF19598F9A9FACFA613" box="[1193,1335,1556,1581]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
studied, including 
<taxonomicName id="4C474D6BFFF6FFF1903BF984FE67A66E" box="[266,415,1593,1616]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF1903BF984FE67A66E" box="[266,415,1593,1616]" italics="true" pageId="5" pageNumber="306">Laccosternus</emphasis>
</taxonomicName>
, the mesotibial spurs are shorter, reaching at most the second mesotarsomere (
<emphasis id="B933EAFAFFF6FFF1941EF985FF30A64A" italics="true" pageId="5" pageNumber="306">character 12.0</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF6FFF191A6F919FEADA716" blockId="5.[151,1436,1700,1832]" pageId="5" pageNumber="306">
<emphasis id="B933EAFAFFF6FFF191A6F919FED7A682" box="[151,303,1700,1724]" italics="true" pageId="5" pageNumber="306">Character 13</emphasis>
. Apex of metatibial spurs: (0) simple; (1) bifid. The apically bifid condition of the metatibial spurs (
<emphasis id="B933EAFAFFF6FFF19191F975FEB0A6DE" box="[160,328,1736,1760]" italics="true" pageId="5" pageNumber="306">character 13.1</emphasis>
) is typical of all known species of 
<taxonomicName id="4C474D6BFFF6FFF193DDF975FC8DA6DE" box="[748,885,1736,1760]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF193DDF975FC8DA6DE" box="[748,885,1736,1760]" italics="true" pageId="5" pageNumber="306">Laccophilus</emphasis>
</taxonomicName>
except one (see key to genera of 
<taxonomicName id="4C474D6BFFF6FFF19436F97AFA6FA6DE" box="[1287,1431,1735,1760]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, below). In all other genera of 
<taxonomicName id="4C474D6BFFF6FFF19330F951FD69A73B" box="[513,657,1772,1797]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, including 
<taxonomicName id="4C474D6BFFF6FFF19226F951FC75A73A" box="[791,909,1772,1796]" class="Insecta" family="Dytiscidae" genus="Philodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF6FFF19226F951FC75A73A" box="[791,909,1772,1796]" italics="true" pageId="5" pageNumber="306">Philodytes</emphasis>
</taxonomicName>
, the apices of the metatibial spurs are acute (
<emphasis id="B933EAFAFFF6FFF19191F8ADFEBDA716" box="[160,325,1808,1832]" italics="true" pageId="5" pageNumber="306">character 13.0</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF6FFF191A6F8E5FA96A7E2" blockId="5.[151,1436,1879,2013]" pageId="5" pageNumber="306">
<emphasis id="B933EAFAFFF6FFF191A6F8E5FED5A74E" box="[151,301,1880,1904]" italics="true" pageId="5" pageNumber="306">Character 14</emphasis>
. Claws of male protarsus: (0) equal in shape, unmodified; (1) different in shape, outer claw scimitarshaped. In all species of 
<taxonomicName id="4C474D6BFFF6FFF190F7F8C0FDABA7AA" box="[454,595,1917,1940]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF6FFF190F7F8C0FDABA7AA" box="[454,595,1917,1940]" italics="true" pageId="5" pageNumber="306">Laccomimus</emphasis>
</taxonomicName>
, except 
<taxonomicName id="4C474D6BFFF6FFF1938CF8C0FCB3A7AA" box="[701,843,1916,1940]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B933EAFAFFF6FFF1938CF8C0FCB3A7AA" box="[701,843,1916,1940]" italics="true" pageId="5" pageNumber="306">L. distinctus</emphasis>
</taxonomicName>
, the outer protarsal claws of males are modified, typically in the shape of a scimitar, different from the inner claw which has a normal sabre shape (
<emphasis id="B933EAFAFFF6FFF195DFF81DFA6CA786" box="[1262,1428,1952,1976]" italics="true" pageId="5" pageNumber="306">character 14.1</emphasis>
; 
<figureCitation id="137C2A6DFFF6FFF191A6F879FF15A7E2" box="[151,237,1988,2013]" captionStart="FIGURES 44 – 51" captionStartId="36.[151,264,1877,1899]" captionTargetBox="[182,1398,193,1837]" captionTargetId="figure@36.[182,1398,193,1839]" captionTargetPageId="36" captionText="FIGURES 44 – 51. SEMs of left protarsus in Laccomimus species (males): 44) L. distinctus, general view; 45) idem, detail at higher magnification of last protarsal segment and claws; 46) idem, detail at higher magnification of stalked suction palettes; 47) L. amazonas, general view; 48) idem, detail at higher magnification of stalked suction palettes; 49) L. bordoni, general view; 50) idem, detail at higher magnification of last protarsal segment and claws; 51) idem, detail at higher magnification of stalked suction palettes." httpUri="https://zenodo.org/record/244309/files/figure.png" pageId="5" pageNumber="306">Figs 47</figureCitation>
; 49–50). In all other genera of 
<taxonomicName id="4C474D6BFFF6FFF19379F879FD20A7E3" box="[584,728,1988,2013]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, both male anterior claws are unmodified (
<emphasis id="B933EAFAFFF6FFF19588F879FAA6A7E2" box="[1209,1374,1988,2012]" italics="true" pageId="5" pageNumber="306">character 14.0</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF5FFF291A6FF25FCDEA122" blockId="6.[151,1436,151,285]" pageId="6" pageNumber="307">
<emphasis id="B933EAFAFFF5FFF291A6FF25FECEA08E" box="[151,310,152,176]" italics="true" pageId="6" pageNumber="307">Character 15</emphasis>
. Median lobe of aedeagus: (0) without a lateral membranous expansion; (1) with a lateral membranous expansion. A lateral membranous expansion is present on the median lobe of the aedeagus of species of the 
<taxonomicName id="4C474D6BFFF5FFF291DDFF5CFE8EA0C6" box="[236,374,224,248]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="B933EAFAFFF5FFF291DDFF5CFE8EA0C6" box="[236,374,224,248]" italics="true" pageId="6" pageNumber="307">L. apalodes</emphasis>
</taxonomicName>
-group of 
<taxonomicName id="4C474D6BFFF5FFF290C5FF5DFD8AA0C6" box="[500,626,224,248]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF290C5FF5DFD8AA0C6" box="[500,626,224,248]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
(
<emphasis id="B933EAFAFFF5FFF293B8FF5DFCCDA0C6" box="[649,821,224,248]" italics="true" pageId="6" pageNumber="307">character 15.1</emphasis>
; see figs 44, 45 in 
<bibRefCitation id="EFD64B19FFF5FFF29505FF62FAE1A0C6" author="Toledo" box="[1076,1305,223,248]" pageId="6" pageNumber="307" refString="Toledo, M., Spangler, P. J. &amp; Balke, M. (2010) Taxonomic revision of the Neotropical diving beetles genus Laccodytes Regimbart, 1895 (Coleoptera: Dytiscidae). Zootaxa, 2347, 37 - 58." type="journal article" year="2010">
Toledo 
<emphasis id="B933EAFAFFF5FFF295BEFF5CFB3FA0C6" box="[1167,1223,224,248]" italics="true" pageId="6" pageNumber="307">et al</emphasis>
. 2010
</bibRefCitation>
). No other 
<taxonomicName id="4C474D6BFFF5FFF291A6FEB9FEDCA123" box="[151,292,260,285]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
studied exhibits this feature (
<emphasis id="B933EAFAFFF5FFF29341FEB9FCEDA122" box="[624,789,260,284]" italics="true" pageId="6" pageNumber="307">character 15.0</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF5FFF291A6FEF1FCCFA226" blockId="6.[151,1436,332,536]" pageId="6" pageNumber="307">
<emphasis id="B933EAFAFFF5FFF291A6FEF1FED6A15A" box="[151,302,332,356]" italics="true" pageId="6" pageNumber="307">Character 16</emphasis>
. Left paramere: (0) short and broad, shorter than half of length of median lobe; (1) elongate, at least half length of median lobe. In 
<taxonomicName id="4C474D6BFFF5FFF290DEFECDFD85A1B9" box="[495,637,368,391]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF290DEFECDFD85A1B9" box="[495,637,368,391]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
the left paramere is elongate and very long, at least as long as half the length of the median lobe, usually longer (
<emphasis id="B933EAFAFFF5FFF2934EFE29FCDEA192" box="[639,806,404,428]" italics="true" pageId="6" pageNumber="307">character 16.1</emphasis>
; 
<figureCitation id="137C2A6DFFF5FFF29207FE29FC40A192" box="[822,952,404,429]" captionStart-0="FIGURES 84 – 87" captionStart-1="FIGURES 88 – 90" captionStart-2="FIGURES 91 – 93" captionStart-3="FIGURES 94 – 97" captionStart-4="FIGURES 98 – 102" captionStartId-0="40.[151,264,1915,1937]" captionStartId-1="41.[151,264,1919,1941]" captionStartId-2="42.[151,264,1921,1943]" captionStartId-3="43.[151,264,1919,1941]" captionStartId-4="44.[151,264,1877,1899]" captionTargetBox-0="[169,1412,201,1889]" captionTargetBox-1="[225,1366,198,1894]" captionTargetBox-2="[168,1414,201,1892]" captionTargetBox-3="[174,1408,200,1884]" captionTargetBox-4="[162,1422,201,1834]" captionTargetId-0="figure@40.[160,1420,193,1898]" captionTargetId-1="figure@41.[204,1370,193,1898]" captionTargetId-2="figure@42.[162,1422,193,1898]" captionTargetId-3="figure@43.[164,1418,193,1889]" captionTargetId-4="figure@44.[151,1436,193,1856]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionTargetPageId-2="42" captionTargetPageId-3="43" captionTargetPageId-4="44" captionText-0="FIGURES 84 – 87. Male genitalia in Laccomimus species, left lateral view: 84) L. distinctus, a) median lobe, b) left paramere, c) right paramere; 85) L. malkini (Loreto, Perú), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere, e) magnification of apex of median lobe (specimen from Piracicaba, Brazil); 86) L. variegatus, a) median lobe, b) left paramere, c) right paramere; 87) L. spinosus, a) median lobe, b) left paramere." captionText-1="FIGURES 88 – 90. Male genitalia in Laccomimus species, left lateral view: 88) L. amazonas, a) median lobe, b) left paramere, c) right paramere; 89) L. improvidus, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 90) L. youngi, a) median lobe (holotype), b) magnification of apex of median lobe (holotype), c) magnification of apex of median lobe (different specimen from type locality), d) left paramere (holotype), e) left paramere (different specimen from type locality)." captionText-2="FIGURES 91 – 93. Male genitalia in Laccomimus pumilio from different localities, left lateral view: 91) Belize, a) median lobe (showing also the three general parts of median lobe), b) magnification of apex of median lobe, c) left paramere, d) right paramere; 92) Florida, USA, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 93) Cayamas, Cuba, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere." captionText-3="FIGURES 94 – 97. Male genitalia in Laccomimus alvarengi from different localities, left lateral view: 94) holotype, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 95) likely aberrant specimen from type locality, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 96) Loreto, Perú, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 97) Mbrucuyá National Park, Argentina, a) median lobe, b) left paramere." captionText-4="FIGURES 98 – 102. Male genitalia in Laccomimus alvarengi from different localities, left lateral view: 98) Mbrucuyá National Park, Argentina, a) median lobe, b) left paramere; 99) Beni, Bolivia, a) median lobe, b) magnification of apex of median lobe; 100) Santa Rosa, Bolivia, median lobe; 101) Esmeraldas, Ecuador, a) median lobe, b) left paramere; 102) Albrook, Panamá, a) median lobe, b) magnification of apex of median lobe, c) left paramere." httpUri-0="https://zenodo.org/record/244313/files/figure.png" httpUri-1="https://zenodo.org/record/244314/files/figure.png" httpUri-2="https://zenodo.org/record/244315/files/figure.png" httpUri-3="https://zenodo.org/record/244316/files/figure.png" httpUri-4="https://zenodo.org/record/244317/files/figure.png" pageId="6" pageNumber="307">Figs 84–98</figureCitation>
; 101–105). The right paramere is, instead, very small. In all other genera of 
<taxonomicName id="4C474D6BFFF5FFF29311FE0AFD49A1EE" box="[544,689,439,464]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, the left paramere is at most as long as half of the length of the median lobe, usually shorter although generally broad; the right paramere is normally shorter then the left paramere, but the difference is not marked (
<emphasis id="B933EAFAFFF5FFF293B3FDBDFCDEA226" box="[642,806,512,536]" italics="true" pageId="6" pageNumber="307">character 16.0</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF5FFF291A6FDF5FAB1A2CE" blockId="6.[151,1436,583,752]" pageId="6" pageNumber="307">
<emphasis id="B933EAFAFFF5FFF291A6FDF5FEC8A25E" box="[151,304,584,608]" italics="true" pageId="6" pageNumber="307">Character 17</emphasis>
. Gonocoxal blades: (0) with two denticles; (1) smooth or with one denticle. In 
<taxonomicName id="4C474D6BFFF5FFF295E5FDF4FA91A25E" box="[1236,1385,585,608]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF295E5FDF4FA91A25E" box="[1236,1385,585,608]" italics="true" pageId="6" pageNumber="307">Laccosternus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF5FFF291A6FDD0FEDDA2BA" box="[151,293,621,644]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF291A6FDD0FEDDA2BA" box="[151,293,621,644]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
(with the exception of 
<taxonomicName id="4C474D6BFFF5FFF2930CFDD0FD32A2BA" box="[573,714,620,644]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B933EAFAFFF5FFF2930CFDD0FD32A2BA" box="[573,714,620,644]" italics="true" pageId="6" pageNumber="307">L. distinctus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF5FFF29238FDD0FC86A2BA" box="[777,894,620,644]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="malkini">
<emphasis id="B933EAFAFFF5FFF29238FDD0FC86A2BA" box="[777,894,620,644]" italics="true" pageId="6" pageNumber="307">L. malkini</emphasis>
</taxonomicName>
), the gonocoxal blades are smooth or at most exhibit a single small rounded tooth at about half the length on the dorsal side (
<emphasis id="B933EAFAFFF5FFF2952CFD2DFB3CA296" box="[1053,1220,656,680]" italics="true" pageId="6" pageNumber="307">character 17.1</emphasis>
; 
<figureCitation id="137C2A6DFFF5FFF295E2FD32FA88A296" box="[1235,1392,655,680]" captionStart="FIGURES 116 – 121" captionStartId="47.[151,264,1829,1851]" captionTargetBox="[229,1344,199,1783]" captionTargetId="figure@47.[198,1370,193,1789]" captionTargetPageId="47" captionText="FIGURES 116 – 121. Female genitalia and cladogram. 116 – 120) Female genitalia in Laccomimus species, left lateral view: 116) L. distinctus (Paraguay), a) left gonocoxa, b) ramus; 117) L. malkini (Perú), a) left gonocoxa, b) ramus; 118) L. alvarengi (Brazil), a) left gonocoxa, b) ramus; 119) L. variegatus (Perú), a) left gonocoxa, b) ramus; 120) L. bordoni (Bolivia), ramus (arrow indicates lateral proximal process of ramus). 121) Single most parsimonious cladogram rooted between ingroup and outgroup, with character changes mapped for each clade. Solid rectangles indicate unique character state transformations; open rectangles indicate homoplasious character state transformations. Support values are indicated on each branch as Bremer / Bootstrap." httpUri="https://zenodo.org/record/244320/files/figure.png" pageId="6" pageNumber="307">Figs 118–119</figureCitation>
); In all other genera of 
<taxonomicName id="4C474D6BFFF5FFF2905DFD09FE01A2F3" box="[364,505,692,717]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
studied and in two species of 
<taxonomicName id="4C474D6BFFF5FFF2927EFD08FC24A2F2" box="[847,988,693,716]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF2927EFD08FC24A2F2" box="[847,988,693,716]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
, the gonocoxal blades bear a subapical, sharp tooth on dorsal side, together with the median tooth described before (
<emphasis id="B933EAFAFFF5FFF292DCFD65FB6AA2CE" box="[1005,1170,728,752]" italics="true" pageId="6" pageNumber="307">character 17.0</emphasis>
; 
<figureCitation id="137C2A6DFFF5FFF29590FD6AFAC5A2CE" box="[1185,1341,727,752]" captionStart="FIGURES 116 – 121" captionStartId="47.[151,264,1829,1851]" captionTargetBox="[229,1344,199,1783]" captionTargetId="figure@47.[198,1370,193,1789]" captionTargetPageId="47" captionText="FIGURES 116 – 121. Female genitalia and cladogram. 116 – 120) Female genitalia in Laccomimus species, left lateral view: 116) L. distinctus (Paraguay), a) left gonocoxa, b) ramus; 117) L. malkini (Perú), a) left gonocoxa, b) ramus; 118) L. alvarengi (Brazil), a) left gonocoxa, b) ramus; 119) L. variegatus (Perú), a) left gonocoxa, b) ramus; 120) L. bordoni (Bolivia), ramus (arrow indicates lateral proximal process of ramus). 121) Single most parsimonious cladogram rooted between ingroup and outgroup, with character changes mapped for each clade. Solid rectangles indicate unique character state transformations; open rectangles indicate homoplasious character state transformations. Support values are indicated on each branch as Bremer / Bootstrap." httpUri="https://zenodo.org/record/244320/files/figure.png" pageId="6" pageNumber="307">Figs 116–117</figureCitation>
).
</paragraph>
<paragraph id="8BF836E8FFF5FFF291A6FC9DFC65A39A" blockId="6.[151,1436,799,933]" pageId="6" pageNumber="307">
<emphasis id="B933EAFAFFF5FFF291A6FC9DFED6A306" box="[151,302,800,824]" italics="true" pageId="6" pageNumber="307">Character 18</emphasis>
. Lateral proximal processes of ramus: (0) well developed; (1) short, almost rudimentary. In females of 
<taxonomicName id="4C474D6BFFF5FFF2918CFCF8FEAAA362" box="[189,338,837,860]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF2918CFCF8FEAAA362" box="[189,338,837,860]" italics="true" pageId="6" pageNumber="307">Laccosternus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF5FFF290AAFCF8FDD0A362" box="[411,552,837,860]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF290AAFCF8FDD0A362" box="[411,552,837,860]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
the lateral proximal process of the female ramus is very short, almost rudimentary (
<emphasis id="B933EAFAFFF5FFF29003FCD5FE21A3BE" box="[306,473,872,896]" italics="true" pageId="6" pageNumber="307">character 18.1</emphasis>
; 
<figureCitation id="137C2A6DFFF5FFF290DBFCDAFD71A3BE" box="[490,649,871,896]" captionStart="FIGURES 116 – 121" captionStartId="47.[151,264,1829,1851]" captionTargetBox="[229,1344,199,1783]" captionTargetId="figure@47.[198,1370,193,1789]" captionTargetPageId="47" captionText="FIGURES 116 – 121. Female genitalia and cladogram. 116 – 120) Female genitalia in Laccomimus species, left lateral view: 116) L. distinctus (Paraguay), a) left gonocoxa, b) ramus; 117) L. malkini (Perú), a) left gonocoxa, b) ramus; 118) L. alvarengi (Brazil), a) left gonocoxa, b) ramus; 119) L. variegatus (Perú), a) left gonocoxa, b) ramus; 120) L. bordoni (Bolivia), ramus (arrow indicates lateral proximal process of ramus). 121) Single most parsimonious cladogram rooted between ingroup and outgroup, with character changes mapped for each clade. Solid rectangles indicate unique character state transformations; open rectangles indicate homoplasious character state transformations. Support values are indicated on each branch as Bremer / Bootstrap." httpUri="https://zenodo.org/record/244320/files/figure.png" pageId="6" pageNumber="307">Figs 116–120</figureCitation>
). In 
<taxonomicName id="4C474D6BFFF5FFF2938FFCD5FCBFA3BE" box="[702,839,872,896]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF2938FFCD5FCBFA3BE" box="[702,839,872,896]" italics="true" pageId="6" pageNumber="307">Laccophilus</emphasis>
</taxonomicName>
and in all other studied genera, the lateral proximal processes of the ramus are long and well developed (
<emphasis id="B933EAFAFFF5FFF293D9FC31FC75A39A" box="[744,909,908,932]" italics="true" pageId="6" pageNumber="307">character 18.0</emphasis>
).
</paragraph>
<paragraph id="8BF836E8FFF5FFF291F6FC69FD23A442" blockId="6.[151,1436,980,2013]" pageId="6" pageNumber="307">
The final data matrix includes 20 taxa and 19 characters (15 binary and four multistate) (
<tableCitation id="C6C50353FFF5FFF29595FC69FB0FA3D2" box="[1188,1271,980,1005]" captionStart="TABLE 1" captionStartId="7.[151,239,332,355]" captionTargetBox="[160,1148,424,1314]" captionTargetPageId="7" captionText="TABLE 1. Data matrix used for the cladistic analysis." httpUri="http://table.plazi.org/id/DF386660FFF4FFF391A6FEF1FD2FA15D" pageId="6" pageNumber="307" tableUuid="DF386660FFF4FFF391A6FEF1FD2FA15D">Table 1</tableCitation>
). The analysis with TNT yielded a single most parsimonious tree of 26 steps (CI = 0.92; RI = 0.97), which supports the monophyly of 
<taxonomicName id="4C474D6BFFF5FFF29074FBA0FE2AA40A" box="[325,466,1053,1076]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF29074FBA0FE2AA40A" box="[325,466,1053,1076]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
and the paraphyly of 
<taxonomicName id="4C474D6BFFF5FFF293D0FBA1FCA7A40A" box="[737,863,1052,1076]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF293D0FBA1FCA7A40A" box="[737,863,1052,1076]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
as presently defined. It also demonstrated weak support for the internal nodes of 
<taxonomicName id="4C474D6BFFF5FFF2932EFBFDFD54A469" box="[543,684,1088,1111]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF2932EFBFDFD54A469" box="[543,684,1088,1111]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
, with the exception of a robustly supported clade formed by 
<taxonomicName id="4C474D6BFFF5FFF294B4FBFCFEF5A442" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="B933EAFAFFF5FFF294B4FBFCFEF5A442" italics="true" pageId="6" pageNumber="307">L. variegatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C474D6BFFF5FFF2902AFBD8FE62A442" box="[283,410,1125,1148]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B933EAFAFFF5FFF2902AFBD8FE62A442" box="[283,410,1125,1148]" italics="true" pageId="6" pageNumber="307">L. spinosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF5FFF290FEFBD8FDA5A442" box="[463,605,1125,1148]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="amazonas">
<emphasis id="B933EAFAFFF5FFF290FEFBD8FDA5A442" box="[463,605,1125,1148]" italics="true" pageId="6" pageNumber="307">L. amazonas</emphasis>
</taxonomicName>
(
<figureCitation id="137C2A6DFFF5FFF2935DFBD9FD35A442" box="[620,717,1124,1149]" captionStart="FIGURES 116 – 121" captionStartId="47.[151,264,1829,1851]" captionTargetBox="[229,1344,199,1783]" captionTargetId="figure@47.[198,1370,193,1789]" captionTargetPageId="47" captionText="FIGURES 116 – 121. Female genitalia and cladogram. 116 – 120) Female genitalia in Laccomimus species, left lateral view: 116) L. distinctus (Paraguay), a) left gonocoxa, b) ramus; 117) L. malkini (Perú), a) left gonocoxa, b) ramus; 118) L. alvarengi (Brazil), a) left gonocoxa, b) ramus; 119) L. variegatus (Perú), a) left gonocoxa, b) ramus; 120) L. bordoni (Bolivia), ramus (arrow indicates lateral proximal process of ramus). 121) Single most parsimonious cladogram rooted between ingroup and outgroup, with character changes mapped for each clade. Solid rectangles indicate unique character state transformations; open rectangles indicate homoplasious character state transformations. Support values are indicated on each branch as Bremer / Bootstrap." httpUri="https://zenodo.org/record/244320/files/figure.png" pageId="6" pageNumber="307">Fig. 121</figureCitation>
).
</paragraph>
<paragraph id="8BF836E8FFF5FFF291F6FB35FDCBA64A" blockId="6.[151,1436,980,2013]" pageId="6" pageNumber="307">
Among the New World genera of 
<taxonomicName id="4C474D6BFFF5FFF29352FB3AFD0BA49E" box="[611,755,1159,1184]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, 
<taxonomicName id="4C474D6BFFF5FFF29235FB35FC69A4A1" box="[772,913,1160,1183]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF29235FB35FC69A4A1" box="[772,913,1160,1183]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
has been confounded with 
<taxonomicName id="4C474D6BFFF5FFF295EEFB35FAA5A49E" box="[1247,1373,1160,1184]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF295EEFB35FAA5A49E" box="[1247,1373,1160,1184]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
. The confusion may have arisen from the small size of members of both genera and the scarce knowledge of 
<taxonomicName id="4C474D6BFFF5FFF2942FFB11FA64A4FA" box="[1310,1436,1196,1220]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF2942FFB11FA64A4FA" box="[1310,1436,1196,1220]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
until recently. In spite of this, 
<taxonomicName id="4C474D6BFFF5FFF290D6FB6DFD9DA4D6" box="[487,613,1232,1256]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF290D6FB6DFD9DA4D6" box="[487,613,1232,1256]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF5FFF293ADFB6DFCD2A4D9" box="[668,810,1232,1255]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF293ADFB6DFCD2A4D9" box="[668,810,1232,1255]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
are very different morphologically. They have different body shapes, very flat with short to truncate elytral tips and lateral sides of the elytra finely bordered in dorsal view in 
<taxonomicName id="4C474D6BFFF5FFF29184FAA5FECBA50E" box="[181,307,1304,1328]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF29184FAA5FECBA50E" box="[181,307,1304,1328]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
(character 7.1, 
<figureCitation id="137C2A6DFFF5FFF290D4FAAAFDD2A50E" box="[485,554,1303,1328]" captionStart="FIGURES 1 – 7. 1" captionStartId="29.[151,264,1905,1927]" captionTargetBox="[169,1412,219,1874]" captionTargetId="figure@29.[169,1413,193,1876]" captionTargetPageId="29" captionText="FIGURES 1 – 7. 1) Habitus of Laccomimus bolivari, dorsal view (Photo A. E. Z. Short); 2) habitus of Laccodytes apalodes, dorsal view (Photo A. E. Z. Short); 3 – 7) SEMs of Laccomimus species (females): 3) habitus of L. alvarengi, dorsal view; 4) habitus of L. bordoni, dorsal view; 5) pronotum of L. alvarengi, dorsal view; 6) pronotum of L. bordoni, dorsal view; 7) pronotum of L. bordoni, dorsolateral view." httpUri="https://zenodo.org/record/244302/files/figure.png" pageId="6" pageNumber="307">Fig. 2</figureCitation>
), less flat, with strongly attenuate elytral tip and elytra not bordered in dorsal view in 
<taxonomicName id="4C474D6BFFF5FFF291C0FA80FE87A56A" box="[241,383,1341,1364]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF291C0FA80FE87A56A" box="[241,383,1341,1364]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
(character 7.0). In 
<taxonomicName id="4C474D6BFFF5FFF29367FA81FD2CA56A" box="[598,724,1340,1364]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF29367FA81FD2CA56A" box="[598,724,1340,1364]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
the metacoxal lines are straight and subparallel (character 9.1), whereas in 
<taxonomicName id="4C474D6BFFF5FFF29026FADDFE5CA549" box="[279,420,1376,1399]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF29026FADDFE5CA549" box="[279,420,1376,1399]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
they are sinuate and strongly converging anteriorly (character 9.0). In 
<taxonomicName id="4C474D6BFFF5FFF2958FFADDFAC4A546" box="[1214,1340,1376,1400]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF2958FFADDFAC4A546" box="[1214,1340,1376,1400]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
the pro- and mesothoracic legs are long and slender, whereas in 
<taxonomicName id="4C474D6BFFF5FFF2920DFA38FC31A5A2" box="[828,969,1413,1436]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF2920DFA38FC31A5A2" box="[828,969,1413,1436]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
they are short and robust. The distant relationship of 
<taxonomicName id="4C474D6BFFF5FFF2900EFA15FE45A5FE" box="[319,445,1448,1472]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF2900EFA15FE45A5FE" box="[319,445,1448,1472]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF5FFF290C3FA15FD87A581" box="[498,639,1448,1471]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF290C3FA15FD87A581" box="[498,639,1448,1471]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
suggested by our analysis (
<figureCitation id="137C2A6DFFF5FFF29280FA1AFBE8A5FE" box="[945,1040,1447,1472]" captionStart="FIGURES 116 – 121" captionStartId="47.[151,264,1829,1851]" captionTargetBox="[229,1344,199,1783]" captionTargetId="figure@47.[198,1370,193,1789]" captionTargetPageId="47" captionText="FIGURES 116 – 121. Female genitalia and cladogram. 116 – 120) Female genitalia in Laccomimus species, left lateral view: 116) L. distinctus (Paraguay), a) left gonocoxa, b) ramus; 117) L. malkini (Perú), a) left gonocoxa, b) ramus; 118) L. alvarengi (Brazil), a) left gonocoxa, b) ramus; 119) L. variegatus (Perú), a) left gonocoxa, b) ramus; 120) L. bordoni (Bolivia), ramus (arrow indicates lateral proximal process of ramus). 121) Single most parsimonious cladogram rooted between ingroup and outgroup, with character changes mapped for each clade. Solid rectangles indicate unique character state transformations; open rectangles indicate homoplasious character state transformations. Support values are indicated on each branch as Bremer / Bootstrap." httpUri="https://zenodo.org/record/244320/files/figure.png" pageId="6" pageNumber="307">Fig. 121</figureCitation>
) is supported by previous results of and analysis by 
<bibRefCitation id="EFD64B19FFF5FFF29061FA71FDCCA5DA" box="[336,564,1484,1509]" pageId="6" pageNumber="307" refString="Ribera, I., Vogler, A. P. &amp; Balke, M. (2008) Phylogeny and diversification of diving beetles (Coleoptera: Dytiscidae). Cladistics, 24, 563 - 590. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2007.00192. x" type="journal article">
Ribera 
<emphasis id="B933EAFAFFF5FFF29095FA70FE26A5DA" box="[420,478,1484,1508]" italics="true" pageId="6" pageNumber="307">et al.</emphasis>
(2008)
</bibRefCitation>
. In fact, the specimens identified as &quot; 
<taxonomicName id="4C474D6BFFF5FFF292D5FA71FB9AA5DA" box="[996,1122,1484,1508]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF292D5FA71FB9AA5DA" box="[996,1122,1484,1508]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
sp1&quot; and “ 
<taxonomicName id="4C474D6BFFF5FFF295D0FA71FA8BA5DB" box="[1249,1395,1484,1509]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
” in fig. 3 of that paper actually belong to 
<taxonomicName id="4C474D6BFFF5FFF2930BFA4DFD3FA639" box="[570,711,1520,1543]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF2930BFA4DFD3FA639" box="[570,711,1520,1543]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
, whereas the specimen identified as &quot; 
<taxonomicName id="4C474D6BFFF5FFF2955DFA4DFB12A636" box="[1132,1258,1520,1544]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF2955DFA4DFB12A636" box="[1132,1258,1520,1544]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
sp2&quot; is actually 
<taxonomicName id="4C474D6BFFF5FFF291A6F9A9FEEDA612" box="[151,277,1556,1580]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF291A6F9A9FEEDA612" box="[151,277,1556,1580]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
(Ignacio Ribera, 2015, personal communication). 
<taxonomicName id="4C474D6BFFF5FFF2925DF9A8FC01A612" box="[876,1017,1557,1580]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF2925DF9A8FC01A612" box="[876,1017,1557,1580]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
is therefore related to 
<taxonomicName id="4C474D6BFFF5FFF29420F9A9FEC2A66E" authority="Guignot, 1948" authorityName="Guignot" authorityYear="1948" class="Insecta" family="Dytiscidae" genus="Africophilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF29420F9A9FA64A612" box="[1297,1436,1556,1580]" italics="true" pageId="6" pageNumber="307">Africophilus</emphasis>
Guignot, 1948
</taxonomicName>
based on results by 
<bibRefCitation id="EFD64B19FFF5FFF29316F98AFCF2A66E" box="[551,778,1591,1616]" pageId="6" pageNumber="307" refString="Ribera, I., Vogler, A. P. &amp; Balke, M. (2008) Phylogeny and diversification of diving beetles (Coleoptera: Dytiscidae). Cladistics, 24, 563 - 590. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2007.00192. x" type="journal article">
Ribera 
<emphasis id="B933EAFAFFF5FFF2934AF984FD4DA66E" box="[635,693,1592,1616]" italics="true" pageId="6" pageNumber="307">et al.</emphasis>
(2008)
</bibRefCitation>
, with a rather isolated position within 
<taxonomicName id="4C474D6BFFF5FFF295F4F98AFA9EA66E" box="[1221,1382,1591,1616]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="subFamily" subFamily="Laccophilinae">Laccophilinae</taxonomicName>
, and more distantly related to 
<taxonomicName id="4C474D6BFFF5FFF2909EF9E1FDD5A64A" box="[431,557,1628,1652]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF2909EF9E1FDD5A64A" box="[431,557,1628,1652]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF836E8FFF5FFF391F6F9C2FB7DA123" blockId="6.[151,1436,980,2013]" lastBlockId="7.[151,1437,151,285]" lastPageId="7" lastPageNumber="308" pageId="6" pageNumber="307">
Apomorphies of 
<taxonomicName id="4C474D6BFFF5FFF290BDF93DFDE2A6A9" box="[396,538,1664,1687]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF290BDF93DFDE2A6A9" box="[396,538,1664,1687]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
are: i) the prosternum and head divided by a deep step (character 0) (this is particularly marked in species with a protruding prosternum); ii) prosternal process short and blunt, cordiform or almost so, not or barely reaching mesocoxae (character 3); iii) aedeagus with very long left paramere, normally longer than half the length of the median lobe (character 16); iv) larger mesotibial spur almost as long as mesotarsus (character 12); v) surface of body, especially of elytra and metatarsi, with a characteristic silky sheen (character 8); vi) male outer claw of protarsi modified (except in one species) (character 14); vii) gonocoxae with one tooth (with two teeth in two species) (character 17); viii) lateral proximal process of ramus short, almost rudimentary (character 18). None of these character states occurs in 
<taxonomicName id="4C474D6BFFF5FFF292A0F8C1FBF7A7AA" box="[913,1039,1916,1940]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF292A0F8C1FBF7A7AA" box="[913,1039,1916,1940]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
in its current concept (
<bibRefCitation id="EFD64B19FFF5FFF2943EF8C1FF2CA786" author="Toledo" pageId="6" pageNumber="307" refString="Toledo, M., Spangler, P. J. &amp; Balke, M. (2010) Taxonomic revision of the Neotropical diving beetles genus Laccodytes Regimbart, 1895 (Coleoptera: Dytiscidae). Zootaxa, 2347, 37 - 58." type="journal article" year="2010">Toledo et al. 2010</bibRefCitation>
). On the other hand, all species of 
<taxonomicName id="4C474D6BFFF5FFF29364F81DFD2BA786" box="[597,723,1952,1976]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF5FFF29364F81DFD2BA786" box="[597,723,1952,1976]" italics="true" pageId="6" pageNumber="307">Laccodytes</emphasis>
</taxonomicName>
are stream dwellers as may be indicated by the variegate dorsal colour patterns, a colouration generally exhibited by reophilic 
<taxonomicName id="4C474D6BFFF5FFF2924AF879FBF0A7E3" box="[891,1032,1988,2013]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
(Balke et al. 2000). 
<taxonomicName id="4C474D6BFFF5FFF29436F878FA6CA7E2" box="[1287,1428,1989,2012]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF5FFF29436F878FA6CA7E2" box="[1287,1428,1989,2012]" italics="true" pageId="6" pageNumber="307">Laccomimus</emphasis>
</taxonomicName>
, instead, apparently inhabits mainly stagnant or slowly flowing waters, although two or three species appear to be found mainly in running waters. The third genus of American 
<taxonomicName id="4C474D6BFFF4FFF39259FF01FC00A0EB" box="[872,1016,188,213]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, 
<taxonomicName id="4C474D6BFFF4FFF39534FF01FB83A0EA" box="[1029,1147,188,212]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF4FFF39534FF01FB83A0EA" box="[1029,1147,188,212]" italics="true" pageId="7" pageNumber="308">Napodytes</emphasis>
</taxonomicName>
, is morphologically very close to 
<taxonomicName id="4C474D6BFFF4FFF391CDFF5DFE82A0C6" box="[252,378,224,248]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF4FFF391CDFF5DFE82A0C6" box="[252,378,224,248]" italics="true" pageId="7" pageNumber="308">Laccodytes</emphasis>
</taxonomicName>
, and apparently also similar in ecology, differing only in few, very specialised characters. 
<taxonomicName id="4C474D6BFFF4FFF391A6FEB8FEDDA122" box="[151,293,261,284]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF4FFF391A6FEB8FEDDA122" box="[151,293,261,284]" italics="true" pageId="7" pageNumber="308">Laccomimus</emphasis>
</taxonomicName>
is a quite homogeneous genus, although some species groups are distinctive.
</paragraph>
<caption id="DF386660FFF4FFF391A6FEF1FD2FA15D" ID-Table-UUID="DF386660FFF4FFF391A6FEF1FD2FA15D" box="[151,727,332,355]" httpUri="http://table.plazi.org/id/DF386660FFF4FFF391A6FEF1FD2FA15D" pageId="7" pageNumber="308" targetBox="[160,1148,424,1314]" targetIsTable="true" targetPageId="7">
<paragraph id="8BF836E8FFF4FFF391A6FEF1FD2FA15D" blockId="7.[151,727,332,355]" box="[151,727,332,355]" pageId="7" pageNumber="308">
<emphasis id="B933EAFAFFF4FFF391A6FEF1FEF2A15D" bold="true" box="[151,266,332,355]" pageId="7" pageNumber="308">TABLE 1.</emphasis>
Data matrix used for the cladistic analysis.
</paragraph>
</caption>
<paragraph id="8BF836E8FFF4FFF39191FEC3FEDAA669" blockId="7.[160,967,382,404]" lastBlockId="7.[151,1436,1382,2019]" pageId="7" pageNumber="308">
Species Character 
<taxonomicName id="4C474D6BFFF4FFF391F6FADBFEBDA540" box="[199,325,1382,1406]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF4FFF391F6FADBFEBDA540" box="[199,325,1382,1406]" italics="true" pageId="7" pageNumber="308">Laccodytes</emphasis>
</taxonomicName>
is presently divided into two very distinct groups: the 
<taxonomicName id="4C474D6BFFF4FFF392F2FADAFBB1A540" box="[963,1097,1382,1406]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="B933EAFAFFF4FFF392F2FADAFBB1A540" box="[963,1097,1382,1406]" italics="true" pageId="7" pageNumber="308">L. apalodes</emphasis>
</taxonomicName>
-group, including two species that appear similar to 
<taxonomicName id="4C474D6BFFF4FFF390BEFA36FDDBA59C" box="[399,547,1419,1442]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF4FFF390BEFA36FDDBA59C" box="[399,547,1419,1442]" italics="true" pageId="7" pageNumber="308">Neptosternus</emphasis>
</taxonomicName>
, and the 
<taxonomicName id="4C474D6BFFF4FFF393BCFA36FCC5A59D" box="[653,829,1419,1443]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="phalacroides">
<emphasis id="B933EAFAFFF4FFF393BCFA36FCC5A59D" box="[653,829,1419,1443]" italics="true" pageId="7" pageNumber="308">L. phalacroides</emphasis>
</taxonomicName>
-group, including the remaining species (
<bibRefCitation id="EFD64B19FFF4FFF3943CFA37FF2CA5F9" author="Toledo" pageId="7" pageNumber="308" refString="Toledo, M., Spangler, P. J. &amp; Balke, M. (2010) Taxonomic revision of the Neotropical diving beetles genus Laccodytes Regimbart, 1895 (Coleoptera: Dytiscidae). Zootaxa, 2347, 37 - 58." type="journal article" year="2010">Toledo et al. 2010</bibRefCitation>
). Characters 1, 2, 3 and 15 define these two groups, with all the derived conditions present in the 
<taxonomicName id="4C474D6BFFF4FFF39421FA12FA6AA5F8" box="[1296,1426,1454,1478]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="B933EAFAFFF4FFF39421FA12FA6AA5F8" box="[1296,1426,1454,1478]" italics="true" pageId="7" pageNumber="308">L. apalodes</emphasis>
</taxonomicName>
- group, and the primitive states present in the 
<taxonomicName id="4C474D6BFFF4FFF393ACFA6EFCB5A5D5" box="[669,845,1491,1515]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="phalacroides">
<emphasis id="B933EAFAFFF4FFF393ACFA6EFCB5A5D5" box="[669,845,1491,1515]" italics="true" pageId="7" pageNumber="308">L. phalacroides</emphasis>
</taxonomicName>
-group (
<tableCitation id="C6C50353FFF4FFF39299FA6FFC05A5D5" box="[936,1021,1490,1515]" captionStart="TABLE 1" captionStartId="7.[151,239,332,355]" captionTargetBox="[160,1148,424,1314]" captionTargetPageId="7" captionText="TABLE 1. Data matrix used for the cladistic analysis." httpUri="http://table.plazi.org/id/DF386660FFF4FFF391A6FEF1FD2FA15D" pageId="7" pageNumber="308" tableUuid="DF386660FFF4FFF391A6FEF1FD2FA15D">Table 1</tableCitation>
). The strong differentiation of these two groups, supported by our phylogenetic results (
<figureCitation id="137C2A6DFFF4FFF393D7FA4BFCB1A631" box="[742,841,1526,1551]" captionStart="FIGURES 116 – 121" captionStartId="47.[151,264,1829,1851]" captionTargetBox="[229,1344,199,1783]" captionTargetId="figure@47.[198,1370,193,1789]" captionTargetPageId="47" captionText="FIGURES 116 – 121. Female genitalia and cladogram. 116 – 120) Female genitalia in Laccomimus species, left lateral view: 116) L. distinctus (Paraguay), a) left gonocoxa, b) ramus; 117) L. malkini (Perú), a) left gonocoxa, b) ramus; 118) L. alvarengi (Brazil), a) left gonocoxa, b) ramus; 119) L. variegatus (Perú), a) left gonocoxa, b) ramus; 120) L. bordoni (Bolivia), ramus (arrow indicates lateral proximal process of ramus). 121) Single most parsimonious cladogram rooted between ingroup and outgroup, with character changes mapped for each clade. Solid rectangles indicate unique character state transformations; open rectangles indicate homoplasious character state transformations. Support values are indicated on each branch as Bremer / Bootstrap." httpUri="https://zenodo.org/record/244320/files/figure.png" pageId="7" pageNumber="308">Fig. 121</figureCitation>
), suggests that 
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<emphasis id="B933EAFAFFF4FFF392CAFA4BFB81A630" box="[1019,1145,1526,1550]" italics="true" pageId="7" pageNumber="308">Laccodytes</emphasis>
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in its current form is not monophyletic. Since this study is focused on 
<taxonomicName id="4C474D6BFFF4FFF393A4F9A6FCDAA60C" box="[661,802,1563,1586]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF4FFF393A4F9A6FCDAA60C" box="[661,802,1563,1586]" italics="true" pageId="7" pageNumber="308">Laccomimus</emphasis>
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, however, we prefer not to make changes to 
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<emphasis id="B933EAFAFFF4FFF3942FF9A6FA64A60D" box="[1310,1436,1563,1587]" italics="true" pageId="7" pageNumber="308">Laccodytes</emphasis>
</taxonomicName>
at this stage.
</paragraph>
</subSubSection>
<paragraph id="8BF836E8FFF4FFF39254FE15FB82A51C" pageId="7" pageNumber="308">
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<taxonomicName id="4C474D6BFFF4FFF39191FE46FE6FA22E" box="[160,407,506,528]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B933EAFAFFF4FFF39191FE46FE6FA22E" box="[160,407,506,528]" italics="true" pageId="7" pageNumber="308">Neptosternus ceylonicus</emphasis>
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<taxonomicName id="4C474D6BFFF4FFF39191FD99FE76A204" box="[160,398,548,570]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B933EAFAFFF4FFF39191FD99FE76A204" box="[160,398,548,570]" italics="true" pageId="7" pageNumber="308">Laccodytes americanus</emphasis>
</taxonomicName>
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<taxonomicName id="4C474D6BFFF4FFF39191FDF0FE8CA25D" box="[160,372,589,611]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="B933EAFAFFF4FFF39191FDF0FE8CA25D" box="[160,372,589,611]" italics="true" pageId="7" pageNumber="308">Laccodytes apalodes</emphasis>
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<taxonomicName id="4C474D6BFFF4FFF39191FDCBFE83A2B2" box="[160,379,630,652]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B933EAFAFFF4FFF39191FDCBFE83A2B2" box="[160,379,630,652]" italics="true" pageId="7" pageNumber="308">Laccophilus chilensis</emphasis>
</taxonomicName>
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<td id="76A65DD6FFF4000B92CBFDCBFBC5A2B2" box="[1018,1085,630,652]" gridcol="3" gridrow="5" pageId="7" pageNumber="308">12010</td>
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<taxonomicName id="4C474D6BFFF4FFF39191FD1DFE8AA288" box="[160,370,672,694]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B933EAFAFFF4FFF39191FD1DFE8AA288" box="[160,370,672,694]" italics="true" pageId="7" pageNumber="308">Laccophilus minutus</emphasis>
</taxonomicName>
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<taxonomicName id="4C474D6BFFF4FFF39191FD77FE6BA2E1" box="[160,403,713,735]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="grouvellei">
<emphasis id="B933EAFAFFF4FFF39191FD77FE6BA2E1" box="[160,403,713,735]" italics="true" pageId="7" pageNumber="308">Laccosternus grouvellei</emphasis>
</taxonomicName>
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<taxonomicName id="4C474D6BFFF4FFF39191FD4EFE94A336" box="[160,364,754,776]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="krausi">
<emphasis id="B933EAFAFFF4FFF39191FD4EFE94A336" box="[160,364,754,776]" italics="true" pageId="7" pageNumber="308">Laccosternus krausi</emphasis>
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<taxonomicName id="4C474D6BFFF4FFF39191FCA1FE89A30C" box="[160,369,796,818]" class="Insecta" family="Dytiscidae" genus="Philodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B933EAFAFFF4FFF39191FCA1FE89A30C" box="[160,369,796,818]" italics="true" pageId="7" pageNumber="308">Philodytes umbrinus</emphasis>
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<td id="76A65DD6FFF4000B92CBFCA1FBC5A30C" box="[1018,1085,796,818]" gridcol="3" gridrow="9" pageId="7" pageNumber="308">12000</td>
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<taxonomicName id="4C474D6BFFF4FFF39191FCFBFE7EA365" box="[160,390,837,859]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="alvarengi">
<emphasis id="B933EAFAFFF4FFF39191FCFBFE7EA365" box="[160,390,837,859]" italics="true" pageId="7" pageNumber="308">Laccomimus alvarengi</emphasis>
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<emphasis id="B933EAFAFFF4FFF39191FCCDFE73A3BB" box="[160,395,880,901]" italics="true" pageId="7" pageNumber="308">Laccomimus amazonas</emphasis>
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<taxonomicName id="4C474D6BFFF4FFF39191FC24FE8EA390" box="[160,374,920,942]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B933EAFAFFF4FFF39191FC24FE8EA390" box="[160,374,920,942]" italics="true" pageId="7" pageNumber="308">Laccomimus bolivari</emphasis>
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<th id="76A65DD6FFF4000B9191FC7CFE6FA3E9" box="[160,407,961,983]" gridcol="0" gridrow="13" pageId="7" pageNumber="308">
<taxonomicName id="4C474D6BFFF4FFF39191FC7FFE8DA3E9" box="[160,373,961,983]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="bordoni">
<emphasis id="B933EAFAFFF4FFF39191FC7FFE8DA3E9" box="[160,373,961,983]" italics="true" pageId="7" pageNumber="308">Laccomimus bordoni</emphasis>
</taxonomicName>
</th>
<td id="76A65DD6FFF4000B9254FC7CFC5DA3E9" box="[869,933,961,983]" gridcol="1" gridrow="13" pageId="7" pageNumber="308">10020</td>
<td id="76A65DD6FFF4000B929FFC7CFC09A3E9" box="[942,1009,961,983]" gridcol="2" gridrow="13" pageId="7" pageNumber="308">0 0 0 10</td>
<td id="76A65DD6FFF4000B92CBFC7CFBC5A3E9" box="[1018,1085,961,983]" gridcol="3" gridrow="13" pageId="7" pageNumber="308">0 0 101</td>
<td id="76A65DD6FFF4000B9574FC7CFB84A3E9" box="[1093,1148,961,983]" gridcol="4" gridrow="13" pageId="7" pageNumber="308">0 111</td>
</tr>
<tr id="357734AAFFF4000B9191FC57FB84A43E" box="[160,1148,1002,1024]" gridrow="14" pageId="7" pageNumber="308">
<th id="76A65DD6FFF4000B9191FC57FE6FA43E" box="[160,407,1002,1024]" gridcol="0" gridrow="14" pageId="7" pageNumber="308">
<taxonomicName id="4C474D6BFFF4FFF39191FC56FE7FA43E" box="[160,391,1002,1024]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B933EAFAFFF4FFF39191FC56FE7FA43E" box="[160,391,1002,1024]" italics="true" pageId="7" pageNumber="308">Laccomimus distinctus</emphasis>
</taxonomicName>
</th>
<td id="76A65DD6FFF4000B9254FC57FC5DA43E" box="[869,933,1002,1024]" gridcol="1" gridrow="14" pageId="7" pageNumber="308">10020</td>
<td id="76A65DD6FFF4000B929FFC57FC09A43E" box="[942,1009,1002,1024]" gridcol="2" gridrow="14" pageId="7" pageNumber="308">0 0 0 10</td>
<td id="76A65DD6FFF4000B92CBFC57FBC5A43E" box="[1018,1085,1002,1024]" gridcol="3" gridrow="14" pageId="7" pageNumber="308">0 1100</td>
<td id="76A65DD6FFF4000B9574FC57FB84A43E" box="[1093,1148,1002,1024]" gridcol="4" gridrow="14" pageId="7" pageNumber="308">0 101</td>
</tr>
<tr id="357734AAFFF4000B9191FBA9FB84A414" box="[160,1148,1044,1066]" gridrow="15" pageId="7" pageNumber="308">
<th id="76A65DD6FFF4000B9191FBA9FE6FA414" box="[160,407,1044,1066]" gridcol="0" gridrow="15" pageId="7" pageNumber="308">
<taxonomicName id="4C474D6BFFF4FFF39191FBA8FE6FA414" box="[160,407,1044,1066]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="improvidus">
<emphasis id="B933EAFAFFF4FFF39191FBA8FE6FA414" box="[160,407,1044,1066]" italics="true" pageId="7" pageNumber="308">Laccomimus improvidus</emphasis>
</taxonomicName>
</th>
<td id="76A65DD6FFF4000B9254FBA9FC5DA414" box="[869,933,1044,1066]" gridcol="1" gridrow="15" pageId="7" pageNumber="308">10020</td>
<td id="76A65DD6FFF4000B929FFBA9FC09A414" box="[942,1009,1044,1066]" gridcol="2" gridrow="15" pageId="7" pageNumber="308">0 0 0 10</td>
<td id="76A65DD6FFF4000B92CBFBA9FBC5A414" box="[1018,1085,1044,1066]" gridcol="3" gridrow="15" pageId="7" pageNumber="308">0 0 101</td>
<td id="76A65DD6FFF4000B9574FBA9FB84A414" box="[1093,1148,1044,1066]" gridcol="4" gridrow="15" pageId="7" pageNumber="308">0 111</td>
</tr>
<tr id="357734AAFFF4000B9191FB80FB84A46D" box="[160,1148,1085,1107]" gridrow="16" pageId="7" pageNumber="308">
<th id="76A65DD6FFF4000B9191FB80FE6FA46D" box="[160,407,1085,1107]" gridcol="0" gridrow="16" pageId="7" pageNumber="308">
<taxonomicName id="4C474D6BFFF4FFF39191FB83FE8AA46D" box="[160,370,1085,1107]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="malkini">
<emphasis id="B933EAFAFFF4FFF39191FB83FE8AA46D" box="[160,370,1085,1107]" italics="true" pageId="7" pageNumber="308">Laccomimus malkini</emphasis>
</taxonomicName>
</th>
<td id="76A65DD6FFF4000B9254FB80FC5DA46D" box="[869,933,1085,1107]" gridcol="1" gridrow="16" pageId="7" pageNumber="308">10020</td>
<td id="76A65DD6FFF4000B929FFB80FC09A46D" box="[942,1009,1085,1107]" gridcol="2" gridrow="16" pageId="7" pageNumber="308">0 0 0 10</td>
<td id="76A65DD6FFF4000B92CBFB80FBC5A46D" box="[1018,1085,1085,1107]" gridcol="3" gridrow="16" pageId="7" pageNumber="308">0 0 101</td>
<td id="76A65DD6FFF4000B9574FB80FB84A46D" box="[1093,1148,1085,1107]" gridcol="4" gridrow="16" pageId="7" pageNumber="308">0 101</td>
</tr>
<tr id="357734AAFFF4000B9191FBDBFB84A443" box="[160,1148,1126,1149]" gridrow="17" pageId="7" pageNumber="308">
<th id="76A65DD6FFF4000B9191FBDBFE6FA443" box="[160,407,1126,1149]" gridcol="0" gridrow="17" pageId="7" pageNumber="308">
<taxonomicName id="4C474D6BFFF4FFF39191FBD5FE8CA442" box="[160,372,1126,1149]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B933EAFAFFF4FFF39191FBD5FE8CA442" box="[160,372,1126,1149]" italics="true" pageId="7" pageNumber="308">Laccomimus pumilio</emphasis>
</taxonomicName>
</th>
<td id="76A65DD6FFF4000B9254FBDBFC5DA443" box="[869,933,1126,1149]" gridcol="1" gridrow="17" pageId="7" pageNumber="308">10020</td>
<td id="76A65DD6FFF4000B929FFBDBFC09A443" box="[942,1009,1126,1149]" gridcol="2" gridrow="17" pageId="7" pageNumber="308">0 0 0 10</td>
<td id="76A65DD6FFF4000B92CBFBDBFBC5A443" box="[1018,1085,1126,1149]" gridcol="3" gridrow="17" pageId="7" pageNumber="308">0 0 101</td>
<td id="76A65DD6FFF4000B9574FBDBFB84A443" box="[1093,1148,1126,1149]" gridcol="4" gridrow="17" pageId="7" pageNumber="308">0 111</td>
</tr>
<tr id="357734AAFFF4000B9191FB2DFB84A498" box="[160,1148,1168,1190]" gridrow="18" pageId="7" pageNumber="308">
<th id="76A65DD6FFF4000B9191FB2DFE6FA498" box="[160,407,1168,1190]" gridcol="0" gridrow="18" pageId="7" pageNumber="308">
<taxonomicName id="4C474D6BFFF4FFF39191FB2CFE7EA498" box="[160,390,1168,1190]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="spangleri">
<emphasis id="B933EAFAFFF4FFF39191FB2CFE7EA498" box="[160,390,1168,1190]" italics="true" pageId="7" pageNumber="308">Laccomimus spangleri</emphasis>
</taxonomicName>
</th>
<td id="76A65DD6FFF4000B9254FB2DFC5DA498" box="[869,933,1168,1190]" gridcol="1" gridrow="18" pageId="7" pageNumber="308">10020</td>
<td id="76A65DD6FFF4000B929FFB2DFC09A498" box="[942,1009,1168,1190]" gridcol="2" gridrow="18" pageId="7" pageNumber="308">0 0 0 10</td>
<td id="76A65DD6FFF4000B92CBFB2DFBC5A498" box="[1018,1085,1168,1190]" gridcol="3" gridrow="18" pageId="7" pageNumber="308">0 0 101</td>
<td id="76A65DD6FFF4000B9574FB2DFB84A498" box="[1093,1148,1168,1190]" gridcol="4" gridrow="18" pageId="7" pageNumber="308">0 111</td>
</tr>
<tr id="357734AAFFF4000B9191FB04FB84A4F1" box="[160,1148,1209,1231]" gridrow="19" pageId="7" pageNumber="308">
<th id="76A65DD6FFF4000B9191FB04FE6FA4F1" box="[160,407,1209,1231]" gridcol="0" gridrow="19" pageId="7" pageNumber="308">
<taxonomicName id="4C474D6BFFF4FFF39191FB07FE85A4F1" box="[160,381,1210,1231]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B933EAFAFFF4FFF39191FB07FE85A4F1" box="[160,381,1210,1231]" italics="true" pageId="7" pageNumber="308">Laccomimus spinosus</emphasis>
</taxonomicName>
</th>
<td id="76A65DD6FFF4000B9254FB04FC5DA4F1" box="[869,933,1209,1231]" gridcol="1" gridrow="19" pageId="7" pageNumber="308">20020</td>
<td id="76A65DD6FFF4000B929FFB04FC09A4F1" box="[942,1009,1209,1231]" gridcol="2" gridrow="19" pageId="7" pageNumber="308">20010</td>
<td id="76A65DD6FFF4000B92CBFB04FBC5A4F1" box="[1018,1085,1209,1231]" gridcol="3" gridrow="19" pageId="7" pageNumber="308">0 1101</td>
<td id="76A65DD6FFF4000B9574FB04FB84A4F1" box="[1093,1148,1209,1231]" gridcol="4" gridrow="19" pageId="7" pageNumber="308">0 111</td>
</tr>
<tr id="357734AAFFF4000B9191FB5FFB84A4C6" box="[160,1148,1250,1272]" gridrow="20" pageId="7" pageNumber="308">
<th id="76A65DD6FFF4000B9191FB5FFE6FA4C6" box="[160,407,1250,1272]" gridcol="0" gridrow="20" pageId="7" pageNumber="308">
<taxonomicName id="4C474D6BFFF4FFF39191FB5EFE69A4C6" box="[160,401,1251,1272]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="B933EAFAFFF4FFF39191FB5EFE69A4C6" box="[160,401,1251,1272]" italics="true" pageId="7" pageNumber="308">Laccomimus variegatus</emphasis>
</taxonomicName>
</th>
<td id="76A65DD6FFF4000B9254FB5FFC5DA4C6" box="[869,933,1250,1272]" gridcol="1" gridrow="20" pageId="7" pageNumber="308">20020</td>
<td id="76A65DD6FFF4000B929FFB5FFC09A4C6" box="[942,1009,1250,1272]" gridcol="2" gridrow="20" pageId="7" pageNumber="308">20010</td>
<td id="76A65DD6FFF4000B92CBFB5FFBC5A4C6" box="[1018,1085,1250,1272]" gridcol="3" gridrow="20" pageId="7" pageNumber="308">0 1101</td>
<td id="76A65DD6FFF4000B9574FB5FFB84A4C6" box="[1093,1148,1250,1272]" gridcol="4" gridrow="20" pageId="7" pageNumber="308">0 111</td>
</tr>
<tr id="357734AAFFF4000B9191FAB1FB84A51C" box="[160,1148,1292,1314]" gridrow="21" pageId="7" pageNumber="308">
<th id="76A65DD6FFF4000B9191FAB1FE6FA51C" box="[160,407,1292,1314]" gridcol="0" gridrow="21" pageId="7" pageNumber="308">
<taxonomicName id="4C474D6BFFF4FFF39191FAB0FE93A51C" box="[160,363,1293,1314]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="youngi">
<emphasis id="B933EAFAFFF4FFF39191FAB0FE93A51C" box="[160,363,1293,1314]" italics="true" pageId="7" pageNumber="308">Laccomimus youngi</emphasis>
</taxonomicName>
</th>
<td id="76A65DD6FFF4000B9254FAB1FC5DA51C" box="[869,933,1292,1314]" gridcol="1" gridrow="21" pageId="7" pageNumber="308">10020</td>
<td id="76A65DD6FFF4000B929FFAB1FC09A51C" box="[942,1009,1292,1314]" gridcol="2" gridrow="21" pageId="7" pageNumber="308">0 0 0 10</td>
<td id="76A65DD6FFF4000B92CBFAB1FBC5A51C" box="[1018,1085,1292,1314]" gridcol="3" gridrow="21" pageId="7" pageNumber="308">0 0 101</td>
<td id="76A65DD6FFF4000B9574FAB1FB84A51C" box="[1093,1148,1292,1314]" gridcol="4" gridrow="21" pageId="7" pageNumber="308">0 111</td>
</tr>
</table>
</paragraph>
<subSubSection id="C35D6563FFF4FFFC91F6F9DEFF18A1CA" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308" type="reference_group">
<paragraph id="8BF836E8FFF4FFFC91F6F9DEFF18A1CA" blockId="7.[151,1436,1382,2019]" lastBlockId="8.[151,1436,151,500]" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308">
<taxonomicName id="4C474D6BFFF4FFF391F6F9DEFEBDA645" ID-CoL="59RJ" box="[199,325,1635,1659]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF4FFF391F6F9DEFEBDA645" box="[199,325,1635,1659]" italics="true" pageId="7" pageNumber="308">Laccodytes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFF4FFF390A0F9DEFDFFA645" ID-CoL="5XL2" box="[401,519,1635,1659]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF4FFF390A0F9DEFDFFA645" box="[401,519,1635,1659]" italics="true" pageId="7" pageNumber="308">Napodytes</emphasis>
</taxonomicName>
are grouped together with the Oriental and Afrotropical reophilic genus 
<taxonomicName id="4C474D6BFFF4FFF391A6F93AFED3A6A0" ID-CoL="639Q" box="[151,299,1671,1694]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF4FFF391A6F93AFED3A6A0" box="[151,299,1671,1694]" italics="true" pageId="7" pageNumber="308">Neptosternus</emphasis>
</taxonomicName>
based on a clearly visible suture between the elytron and epipleuron in dorsal view (= “lateral side of elytron bordered”) (
<bibRefCitation id="EFD64B19FFF4FFF3904CF917FDAEA6FD" author="Ribera" box="[381,598,1706,1731]" pageId="7" pageNumber="308" refString="Ribera, I., Vogler, A. P. &amp; Balke, M. (2008) Phylogeny and diversification of diving beetles (Coleoptera: Dytiscidae). Cladistics, 24, 563 - 590. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2007.00192. x" type="journal article" year="2008">
Ribera 
<emphasis id="B933EAFAFFF4FFF390E5F916FDE8A6FD" box="[468,528,1707,1731]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2008
</bibRefCitation>
; 
<bibRefCitation id="EFD64B19FFF4FFF39355F917FCB9A6FD" author="Toledo" box="[612,833,1706,1731]" pageId="7" pageNumber="308" refString="Toledo, M., Spangler, P. J. &amp; Balke, M. (2010) Taxonomic revision of the Neotropical diving beetles genus Laccodytes Regimbart, 1895 (Coleoptera: Dytiscidae). Zootaxa, 2347, 37 - 58." type="journal article" year="2010">
Toledo 
<emphasis id="B933EAFAFFF4FFF3938CF916FD01A6FD" box="[701,761,1707,1731]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2010
</bibRefCitation>
). 
<taxonomicName id="4C474D6BFFF4FFF39266F916FC1DA6FC" ID-CoL="8LY8Y" box="[855,997,1707,1730]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF4FFF39266F916FC1DA6FC" box="[855,997,1707,1730]" italics="true" pageId="7" pageNumber="308">Laccomimus</emphasis>
</taxonomicName>
, on the other hand, is closer to other 
<taxonomicName id="4C474D6BFFF4FFF391A6F973FECEA6D9" ID-CoL="JS5" box="[151,310,1742,1767]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="subFamily" subFamily="Laccophilinae">Laccophilinae</taxonomicName>
genera such as the poorly known, Oriental genus 
<taxonomicName id="4C474D6BFFF4FFF39243F972FBFFA6D8" ID-CoL="59S8" box="[882,1031,1743,1766]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF4FFF39243F972FBFFA6D8" box="[882,1031,1743,1766]" italics="true" pageId="7" pageNumber="308">Laccosternus</emphasis>
</taxonomicName>
, with which it shares the following features: the prosternal process more or less cordiform (depending on the species in 
<taxonomicName id="4C474D6BFFF4FFF3954DF94EFAF2A734" ID-CoL="8LY8Y" box="[1148,1290,1779,1802]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFF4FFF3954DF94EFAF2A734" box="[1148,1290,1779,1802]" italics="true" pageId="7" pageNumber="308">Laccomimus</emphasis>
</taxonomicName>
), previously regarded as an apomorphy of 
<taxonomicName id="4C474D6BFFF4FFF390C8F8AAFD76A710" ID-CoL="59S8" box="[505,654,1815,1838]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF4FFF390C8F8AAFD76A710" box="[505,654,1815,1838]" italics="true" pageId="7" pageNumber="308">Laccosternus</emphasis>
</taxonomicName>
(character 3); the metacoxal lines sinuate and strongly converging anteriorly (character 9), with the lobes of the metacoxal processes rounded and divided by an evident notch (character 10); similarly small size and elongate body, attenuate caudally; females with the gonocoxae with one tooth (character 17) and with a short and rudimentary lateroproximal process of the ramus (character 18). The mesotibial spurs, however, are short in 
<taxonomicName id="4C474D6BFFF4FFF39368F81AFD16A780" ID-CoL="59S8" box="[601,750,1959,1982]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF4FFF39368F81AFD16A780" box="[601,750,1959,1982]" italics="true" pageId="7" pageNumber="308">Laccosternus</emphasis>
</taxonomicName>
, similar to those found in 
<taxonomicName id="4C474D6BFFF4FFF39510F81BFB52A780" ID-CoL="59RR" box="[1057,1194,1958,1982]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFF4FFF39510F81BFB52A780" box="[1057,1194,1958,1982]" italics="true" pageId="7" pageNumber="308">Laccophilus</emphasis>
</taxonomicName>
(character 12). After the recent discovery of 
<taxonomicName id="4C474D6BFFF4FFF390F3F876FBDAA7DD" ID-CoL="8LZ8G" authority="Brancucci &amp; Vongsana, 2013" authorityName="Brancucci &amp; Vongsana" authorityYear="2013" box="[450,1058,1994,2019]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="krausi">
<emphasis id="B933EAFAFFF4FFF390F3F876FD55A7DD" box="[450,685,1995,2019]" italics="true" pageId="7" pageNumber="308">Laccosternus krausi</emphasis>
<bibRefCitation id="EFD64B19FFF4FFF3938CF877FBDAA7DD" author="Brancucci" box="[701,1058,1994,2019]" pageId="7" pageNumber="308" refString="Brancucci, M. &amp; Vongsana, K. (2013) A revision of the genus Laccosternus Brancucci, 1983 (Coleoptera, Dytiscidae). Entomologica Basiliensia et Collectionis Frey, 34, 129 - 135." type="journal article" year="2013">Brancucci &amp; Vongsana, 2013</bibRefCitation>
</taxonomicName>
, the second known species of 
<taxonomicName id="4C474D6BFFFBFFFC91A6FF24FED4A08E" ID-CoL="59S8" box="[151,300,153,176]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFFBFFFC91A6FF24FED4A08E" box="[151,300,153,176]" italics="true" pageId="8" pageNumber="309">Laccosternus</emphasis>
</taxonomicName>
and the first description of a male of this genus, it is clear that male characters differ from those of 
<taxonomicName id="4C474D6BFFFBFFFC91A6FF00FEDCA0EA" ID-CoL="8LY8Y" box="[151,292,189,212]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFFBFFFC91A6FF00FEDCA0EA" box="[151,292,189,212]" italics="true" pageId="8" pageNumber="309">Laccomimus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C474D6BFFFBFFFC9002FF00FE30A0EA" ID-CoL="59S8" box="[307,456,189,212]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFFBFFFC9002FF00FE30A0EA" box="[307,456,189,212]" italics="true" pageId="8" pageNumber="309">Laccosternus</emphasis>
</taxonomicName>
have parameres that are more similar to 
<taxonomicName id="4C474D6BFFFBFFFC92A2FF01FBE4A0EA" ID-CoL="59RR" box="[915,1052,188,212]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFFBFFFC92A2FF01FBE4A0EA" box="[915,1052,188,212]" italics="true" pageId="8" pageNumber="309">Laccophilus</emphasis>
</taxonomicName>
(character 16). This confirms that 
<taxonomicName id="4C474D6BFFFBFFFC91A6FF5CFED4A0C6" ID-CoL="59S8" box="[151,300,225,248]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFFBFFFC91A6FF5CFED4A0C6" box="[151,300,225,248]" italics="true" pageId="8" pageNumber="309">Laccosternus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D6BFFFBFFFC9041FF5DFE06A0C9" ID-CoL="8LY8Y" box="[368,510,224,247]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFFBFFFC9041FF5DFE06A0C9" box="[368,510,224,247]" italics="true" pageId="8" pageNumber="309">Laccomimus</emphasis>
</taxonomicName>
are two separate genera, although closely related, and may be sister groups depending on where the phylogeny presented here is rooted (
<figureCitation id="137C2A6DFFFBFFFC920AFEB9FC65A122" box="[827,925,260,285]" captionStart="FIGURES 116 – 121" captionStartId="47.[151,264,1829,1851]" captionTargetBox="[229,1344,199,1783]" captionTargetId="figure@47.[198,1370,193,1789]" captionTargetPageId="47" captionText="FIGURES 116 – 121. Female genitalia and cladogram. 116 – 120) Female genitalia in Laccomimus species, left lateral view: 116) L. distinctus (Paraguay), a) left gonocoxa, b) ramus; 117) L. malkini (Perú), a) left gonocoxa, b) ramus; 118) L. alvarengi (Brazil), a) left gonocoxa, b) ramus; 119) L. variegatus (Perú), a) left gonocoxa, b) ramus; 120) L. bordoni (Bolivia), ramus (arrow indicates lateral proximal process of ramus). 121) Single most parsimonious cladogram rooted between ingroup and outgroup, with character changes mapped for each clade. Solid rectangles indicate unique character state transformations; open rectangles indicate homoplasious character state transformations. Support values are indicated on each branch as Bremer / Bootstrap." httpUri="https://zenodo.org/record/244320/files/figure.png" pageId="8" pageNumber="309">Fig. 121</figureCitation>
). With 
<taxonomicName id="4C474D6BFFFBFFFC92DCFEB8FB79A122" ID-CoL="639Q" box="[1005,1153,261,284]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFFBFFFC92DCFEB8FB79A122" box="[1005,1153,261,284]" italics="true" pageId="8" pageNumber="309">Neptosternus</emphasis>
</taxonomicName>
as outgroup (a procedure supported by the distant relationship of this genus with 
<taxonomicName id="4C474D6BFFFBFFFC9221FE95FC65A101" ID-CoL="8LY8Y" box="[784,925,296,319]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFFBFFFC9221FE95FC65A101" box="[784,925,296,319]" italics="true" pageId="8" pageNumber="309">Laccomimus</emphasis>
</taxonomicName>
, see 
<bibRefCitation id="EFD64B19FFFBFFFC92E8FE9AFB5FA17E" author="Ribera" box="[985,1191,295,320]" pageId="8" pageNumber="309" refString="Ribera, I., Vogler, A. P. &amp; Balke, M. (2008) Phylogeny and diversification of diving beetles (Coleoptera: Dytiscidae). Cladistics, 24, 563 - 590. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2007.00192. x" type="journal article" year="2008">
Ribera 
<emphasis id="B933EAFAFFFBFFFC951DFE94FB9DA17E" box="[1068,1125,296,320]" italics="true" pageId="8" pageNumber="309">et al.</emphasis>
2008
</bibRefCitation>
) the result supports a sister relationship of these two genera. The strongly scattered punctation on the elytra described for 
<taxonomicName id="4C474D6BFFFBFFFC9437FEF0FEFFA1B6" ID-CoL="3RMGX" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="grouvellei">
<emphasis id="B933EAFAFFFBFFFC9437FEF0FEFFA1B6" italics="true" pageId="8" pageNumber="309">Laccosternus grouvellei</emphasis>
</taxonomicName>
is also visible (although less marked) in 
<taxonomicName id="4C474D6BFFFBFFFC93E5FECCFCC0A1B6" ID-CoL="8LZ8G" box="[724,824,368,392]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="krausi">
<emphasis id="B933EAFAFFFBFFFC93E5FECCFCC0A1B6" box="[724,824,368,392]" italics="true" pageId="8" pageNumber="309">L. krausi</emphasis>
</taxonomicName>
, suggesting that a diffuse elytral punctation (character 6) might be another character state defining 
<taxonomicName id="4C474D6BFFFBFFFC93A3FE28FCDFA192" ID-CoL="59S8" box="[658,807,405,428]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAFAFFFBFFFC93A3FE28FCDFA192" box="[658,807,405,428]" italics="true" pageId="8" pageNumber="309">Laccosternus</emphasis>
</taxonomicName>
. Male features of 
<taxonomicName id="4C474D6BFFFBFFFC92CAFE28FB73A192" ID-CoL="3RMGX" box="[1019,1163,404,428]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="grouvellei">
<emphasis id="B933EAFAFFFBFFFC92CAFE28FB73A192" box="[1019,1163,404,428]" italics="true" pageId="8" pageNumber="309">L. grouvellei</emphasis>
</taxonomicName>
are still unknown since this species is at present only known from three females collected in Sumatra, 
<collectingCountry id="F3507678FFFBFFFC952DFE05FBADA1F1" box="[1052,1109,440,463]" name="Laos" pageId="8" pageNumber="309">Laos</collectingCountry>
, and 
<collectingCountry id="F3507678FFFBFFFC95A2FE0AFB02A1EE" box="[1171,1274,439,464]" name="Malaysia" pageId="8" pageNumber="309">Malaysia</collectingCountry>
(
<bibRefCitation id="EFD64B19FFFBFFFC943AFE0AFF2CA1CA" author="Toledo" pageId="8" pageNumber="309" refString="Toledo, M., Hendrich, L. &amp; Stastny, J. (2002) Two new species of Laccophilus from Sulawesi with notes on other Laccophilinae in Southeast Asia (Coleoptera: Dytiscidae). Linzer Biologische Beitrage, 35, 189 - 200." type="journal article" year="2002">
Toledo 
<emphasis id="B933EAFAFFFBFFFC9453FE04FA64A1EE" box="[1378,1436,440,464]" italics="true" pageId="8" pageNumber="309">et al.</emphasis>
2002
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>