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<document id="9B3F9ECB786F3368E3D4BEB9D8761A40" ID-CLB-Dataset="38764" ID-DOI="10.1080/00222933.2016.1180718" ID-GBIF-Dataset="ed0c7c1a-19b2-4497-9314-1ed20bda20dc" ID-ISSN="1464-5262" ID-Zenodo-Dep="269029" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1466575103704" checkinUser="plazi" docAuthor="Eknarin Rodcharoen, Niel L. Bruce &amp; Pornsilp Pholpunthin" docDate="2016" docId="03EE87E6E218FFC51AC9F996FC3A2104" docLanguage="en" docName="JNatHist.50.31-32.1935-1981.pdf" docOrigin="Journal of Natural History 50 (31 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2016.1180718" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Cirolana Leach 1818" docType="treatment" docVersion="17" lastPageNumber="6" masterDocId="FFD7FF9EE21DFFC31B20FFE6FF992579" masterDocTitle="Description of four new species of the Cirolana parva group (Crustacea: Isopoda: Cirolanidae) from Thailand, with supporting molecular (COI) data" masterLastPageNumber="1981" masterPageNumber="1935" pageNumber="5" updateTime="1698663086370" updateUser="ExternalLinkService">
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<mods:title id="AC5649F3983D7DDD6EB268C20EAACED9">Description of four new species of the Cirolana parva group (Crustacea: Isopoda: Cirolanidae) from Thailand, with supporting molecular (COI) data</mods:title>
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<mods:namePart id="900461C1D574C9CAFA115287E2475ED4">Eknarin Rodcharoen</mods:namePart>
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<mods:namePart id="2A2D19620A9F7DAB9815BFE9DF717768">Niel L. Bruce</mods:namePart>
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<treatment id="03EE87E6E218FFC51AC9F996FC3A2104" ID-DOI="http://doi.org/10.5281/zenodo.5195239" ID-GBIF-Taxon="121061468" ID-Zenodo-Dep="5195239" LSID="urn:lsid:plazi:treatment:03EE87E6E218FFC51AC9F996FC3A2104" httpUri="http://treatment.plazi.org/id/03EE87E6E218FFC51AC9F996FC3A2104" lastPageId="6" lastPageNumber="6" pageId="5" pageNumber="5">
<subSubSection id="C35D657BE218FFC61AC9F996FCA323F0" box="[489,826,1647,1673]" pageId="5" pageNumber="5" type="nomenclature">
<paragraph id="8BF836F0E218FFC61AC9F996FCA323F0" blockId="5.[407,908,1475,1673]" box="[489,826,1647,1673]" pageId="5" pageNumber="5">
Genus
<taxonomicName id="4C474D73E218FFC6191CF989FCA323F0" ID-CoL="7NSYL" authority="Leach, 1818" authorityName="Leach" authorityYear="1818" box="[572,826,1647,1673]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAE2E218FFC6191CF989FD3D23F0" bold="true" box="[572,676,1647,1673]" italics="true" pageId="5" pageNumber="5">Cirolana</emphasis>
<bibRefCitation id="EFD64B01E218FFC6198DF996FCA323F0" author="Leach WE." box="[685,826,1648,1673]" editor="Cuvier F" journalOrPublisher="Paris" pageId="5" pageNumber="5" pagination="338 - 354" part="12" refString="Leach WE. 1818. Cymothoades. In: Cuvier F, editor. Dictionnaire des sciences naturelles. Paris. 12: 338 - 354." title="Cymothoades" type="journal article" volumeTitle="Dictionnaire des sciences naturelles" year="1818">Leach, 1818</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="C35D657BE21BFFC51B80FF68FC3625B3" pageId="6" pageNumber="6" type="reference_group">
<paragraph id="8BF836F0E21BFFC51B80FF68FE0C25DE" blockId="6.[160,943,142,202]" box="[160,405,142,167]" pageId="6" pageNumber="6">
<heading id="D0B0819CE21BFFC51B80FF68FE0C25DE" bold="true" box="[160,405,142,167]" fontSize="11" level="2" pageId="6" pageNumber="6" reason="0">
<emphasis id="B933EAE2E21BFFC51B80FF68FE0C25DE" bold="true" box="[160,405,142,167]" italics="true" pageId="6" pageNumber="6">Restricted synonymy</emphasis>
</heading>
</paragraph>
<paragraph id="8BF836F0E21BFFC51B80FF54FC3625B3" blockId="6.[160,943,142,202]" box="[160,943,178,202]" pageId="6" pageNumber="6">
<bibRefCitation id="EFD64B01E21BFFC51B80FF54FE8625B3" author="Bruce NL." box="[160,287,178,202]" journalOrPublisher="Records Aust Museum, Suppl. 6" pageId="6" pageNumber="6" pagination="1 - 239" refString="Bruce NL. 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records Aust Museum, Suppl. 6: 1 - 239." title="Cirolanidae (Crustacea: Isopoda) of Australia" type="book chapter" year="1986">Bruce 1986</bibRefCitation>
, 139;
<bibRefCitation id="EFD64B01E21BFFC51A44FF54FDE825B3" author="Kensley B &amp; Schotte M." box="[356,625,178,202]" journalOrPublisher="Washington, DC: Smithsonian Institution Press" pageId="6" pageNumber="6" refString="Kensley B, Schotte M. 1989. Guide to the marine isopod crustaceans of the Caribbean. Washington, DC: Smithsonian Institution Press." title="Guide to the marine isopod crustaceans of the Caribbean" type="book" year="1989">Kensley &amp; Schotte 1989</bibRefCitation>
, 132;
<bibRefCitation id="EFD64B01E21BFFC51996FF54FC1825B3" author="Brusca RC &amp; Wetzer R &amp; France SC." box="[694,897,178,202]" journalOrPublisher="Proc San Diego Nat Hist Museum" pageId="6" pageNumber="6" pagination="1 - 96" part="30" refString="Brusca RC, Wetzer R, France SC. 1995. Cirolanidae (Crustacea: Isopoda: Flabellifera) of the Tropical Eastern Pacific. Proc San Diego Nat Hist Museum. 30: 1 - 96." title="Cirolanidae (Crustacea: Isopoda: Flabellifera) of the Tropical Eastern Pacific" type="journal article" year="1995">Brusca et al. 1995</bibRefCitation>
, 17.
</paragraph>
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<subSubSection id="C35D657BE21BFFC51B80FF1AFC222441" pageId="6" pageNumber="6" type="nomenclature">
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<heading id="D0B0819CE21BFFC51B80FF1AFEAD246C" bold="true" box="[160,308,252,277]" fontSize="11" level="2" pageId="6" pageNumber="6" reason="0">
<emphasis id="B933EAE2E21BFFC51B80FF1AFEAD246C" bold="true" box="[160,308,252,277]" italics="true" pageId="6" pageNumber="6">
<typeStatus id="54FC8852E21BFFC51B80FF1AFF41246C" box="[160,216,252,277]" pageId="6" pageNumber="6">Type</typeStatus>
species
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</paragraph>
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<taxonomicName id="4C474D73E21BFFC51B80FEF9FE7A2441" authority="Leach, 1818" authorityName="Leach" authorityYear="1818" box="[160,483,287,312]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="6" phylum="Arthropoda" rank="species" species="cranchi">
<emphasis id="B933EAE2E21BFFC51B80FEF9FF62244E" box="[160,251,287,311]" italics="true" pageId="6" pageNumber="6">Cirolana</emphasis>
<emphasis id="B933EAE2E21BFFC51A24FEF9FECD244E" box="[260,340,287,311]" italics="true" pageId="6" pageNumber="6">cranchi</emphasis>
<bibRefCitation id="EFD64B01E21BFFC51A7DFEC6FE7A2441" author="Leach WE." box="[349,483,288,312]" editor="Cuvier F" journalOrPublisher="Paris" pageId="6" pageNumber="6" pagination="338 - 354" part="12" refString="Leach WE. 1818. Cymothoades. In: Cuvier F, editor. Dictionnaire des sciences naturelles. Paris. 12: 338 - 354." title="Cymothoades" type="journal article" volumeTitle="Dictionnaire des sciences naturelles" year="1818">Leach, 1818</bibRefCitation>
</taxonomicName>
, by monotypy (see
<bibRefCitation id="EFD64B01E21BFFC519E8FEC6FC372441" author="Downloaded Bruce NL &amp; Ellis J." box="[712,942,288,312]" journalOrPublisher="Bull Br Museum Nat Hist (Zoology)" pageId="6" pageNumber="6" pagination="75 - 84" part="44" refString="2016 June 21 02: 00 at] Cambridge of University [by Downloaded Bruce NL, Ellis J. 1983. Cirolana cranchi Leach, 1818 (Crustacea: Isopoda: Cirolanidae) redescribed, with notes on its distribution. Bull Br Museum Nat Hist (Zoology). 44: 75 - 84." title="Cirolana cranchi Leach, 1818 (Crustacea: Isopoda: Cirolanidae) redescribed, with notes on its distribution" type="journal article" year="1983">Bruce and Ellis 1983</bibRefCitation>
).
</paragraph>
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<subSubSection id="C35D657BE21BFFC51B80FE8CFC3A2104" pageId="6" pageNumber="6" type="discussion">
<paragraph id="8BF836F0E21BFFC51B80FE8CFE9F24FA" blockId="6.[160,1156,362,1149]" box="[160,262,362,387]" pageId="6" pageNumber="6">
<heading id="D0B0819CE21BFFC51B80FE8CFE9F24FA" bold="true" box="[160,262,362,387]" fontSize="11" level="2" pageId="6" pageNumber="6" reason="0">
<emphasis id="B933EAE2E21BFFC51B80FE8CFE9F24FA" bold="true" box="[160,262,362,387]" italics="true" pageId="6" pageNumber="6">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BF836F0E21BFFC51B80FE6BFDE02749" blockId="6.[160,1156,362,1149]" pageId="6" pageNumber="6">
Species of
<taxonomicName id="4C474D73E21BFFC51A03FE6BFEE724DC" box="[291,382,397,421]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAE2E21BFFC51A03FE6BFEE724DC" box="[291,382,397,421]" italics="true" pageId="6" pageNumber="6">Cirolana</emphasis>
</taxonomicName>
are found principally in marine environments, occupying various substrates such as rocky shore, coral reef, dead wood and sediments from the continental slope, to intertidal and shallow subtidal marine zones. They are also occasionally found in low-salinity or variable-salinity habitats such as mangroves and the estuarine reaches of rivers and creeks (
<bibRefCitation id="EFD64B01E21BFFC51ACDFDFEFDF52749" author="Bruce NL." box="[493,620,536,560]" journalOrPublisher="Records Aust Museum, Suppl. 6" pageId="6" pageNumber="6" pagination="1 - 239" refString="Bruce NL. 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records Aust Museum, Suppl. 6: 1 - 239." title="Cirolanidae (Crustacea: Isopoda) of Australia" type="book chapter" year="1986">Bruce 1986</bibRefCitation>
).
</paragraph>
<paragraph id="8BF836F0E21BFFC51BE0FDDDFB1D27A4" blockId="6.[160,1156,362,1149]" pageId="6" pageNumber="6">
Despite recent revisions (e.g.
<bibRefCitation id="EFD64B01E21BFFC5192EFDDDFD14272A" author="Bruce NL." box="[526,653,571,595]" journalOrPublisher="Aust J Marine Freshwater Res" pageId="6" pageNumber="6" pagination="945 - 966" part="32" refString="Bruce NL. 1981. Cirolanidae (Crustacea: Isopoda) of Australia: diagnoses of Cirolana Leach, Metacirolana Nierstrasz, Neocirolana Hale, Anopsilana Paulian &amp; Debouteville, and three new genera - Natatolana, Politolana and Cartetolana. Aust J Marine Freshwater Res. 32: 945 - 966." title="Cirolanidae (Crustacea: Isopoda) of Australia: diagnoses of Cirolana Leach, Metacirolana Nierstrasz, Neocirolana Hale, Anopsilana Paulian &amp; Debouteville, and three new genera - Natatolana, Politolana and Cartetolana" type="journal article" year="1981">Bruce 1981</bibRefCitation>
,
<bibRefCitation id="EFD64B01E21BFFC519BAFDDDFD48272A" author="Bruce NL." box="[666,721,571,595]" journalOrPublisher="Records Aust Museum, Suppl. 6" pageId="6" pageNumber="6" pagination="1 - 239" refString="Bruce NL. 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records Aust Museum, Suppl. 6: 1 - 239." title="Cirolanidae (Crustacea: Isopoda) of Australia" type="book chapter" year="1986">1986</bibRefCitation>
;
<bibRefCitation id="EFD64B01E21BFFC519FFFDDDFC3F272A" author="Brusca RC &amp; Wetzer R &amp; France SC." box="[735,934,571,595]" journalOrPublisher="Proc San Diego Nat Hist Museum" pageId="6" pageNumber="6" pagination="1 - 96" part="30" refString="Brusca RC, Wetzer R, France SC. 1995. Cirolanidae (Crustacea: Isopoda: Flabellifera) of the Tropical Eastern Pacific. Proc San Diego Nat Hist Museum. 30: 1 - 96." title="Cirolanidae (Crustacea: Isopoda: Flabellifera) of the Tropical Eastern Pacific" type="journal article" year="1995">Brusca et al. 1995</bibRefCitation>
), there is still some lack of resolution between genera close to
<taxonomicName id="4C474D73E21BFFC519A7FDBBFD7B270C" box="[647,738,605,629]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAE2E21BFFC519A7FDBBFD7B270C" box="[647,738,605,629]" italics="true" pageId="6" pageNumber="6">Cirolana</emphasis>
</taxonomicName>
. For example,
<bibRefCitation id="EFD64B01E21BFFC518A7FDBBFF7F27E1" author="Botosaneanu L &amp; Iliffe TM." journalOrPublisher="Bull Inst R Sci Nat Belg Biol" pageId="6" pageNumber="6" pagination="77 - 94" part="67" refString="Botosaneanu L, Iliffe TM. 1997. Four new stygobitic cirolanids (Crustacea: Isopoda) from the Caribbean - with remarks on intergeneric limits in some cirolanids. Bull Inst R Sci Nat Belg Biol. 67: 77 - 94." title="Four new stygobitic cirolanids (Crustacea: Isopoda) from the Caribbean - with remarks on intergeneric limits in some cirolanids" type="journal article" year="1997">
Botosaneanu and Iliffe (
<date id="FFF91030E21BFFC51B88FD66FF4627E1" box="[168,223,640,664]" pageId="6" pageNumber="6" value="1997">1997</date>
)
</bibRefCitation>
placed
<taxonomicName id="4C474D73E21BFFC51A68FD66FCB127E1" authority="Paulian &amp; Deboutteville, 1956" authorityName="Paulian &amp; Deboutteville" authorityYear="1956" box="[328,808,640,664]" class="Malacostraca" family="Cirolanidae" genus="Anopsilana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAE2E21BFFC51A68FD66FE5B27E1" box="[328,450,640,664]" italics="true" pageId="6" pageNumber="6">Anopsilana</emphasis>
Paulian &amp; Deboutteville, 1956
</taxonomicName>
into synonymy with
<taxonomicName id="4C474D73E21BFFC51F08FD66FB1A27E1" box="[1064,1155,640,664]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAE2E21BFFC51F08FD66FB1A27E1" box="[1064,1155,640,664]" italics="true" pageId="6" pageNumber="6">Cirolana</emphasis>
</taxonomicName>
(subgenus
<taxonomicName id="4C474D73E21BFFC51A00FD44FE0327C3" box="[288,410,674,698]" class="Malacostraca" family="Cirolanidae" genus="Anopsilana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAE2E21BFFC51A00FD44FE0327C3" box="[288,410,674,698]" italics="true" pageId="6" pageNumber="6">Anopsilana</emphasis>
</taxonomicName>
), as the principal and only differentiating character, the lack of plumose setae endopod of pleopods 3 and 4, was regarded as manifestly homoplasious.
</paragraph>
<paragraph id="8BF836F0E21BFFC51BE0FD0EFC3A2104" blockId="6.[160,1156,362,1149]" pageId="6" pageNumber="6">
The
<emphasis id="B933EAE2E21BFFC51BD9FD0EFE312679" box="[249,424,744,768]" italics="true" pageId="6" pageNumber="6">
<taxonomicName id="4C474D73E21BFFC51BD9FD0EFECD2679" box="[249,340,744,768]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="6" phylum="Arthropoda" rank="genus">Cirolana</taxonomicName>
<emphasis id="B933EAE2E21BFFC51A45FD0EFEF32679" bold="true" box="[357,362,744,768]" italics="true" pageId="6" pageNumber="6"></emphasis>
<taxonomicName id="4C474D73E21BFFC51A4AFD0FFE312679" box="[362,424,745,768]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="6" phylum="Arthropoda" rank="species" species="parva">parva</taxonomicName>
</emphasis>
group
<emphasis id="B933EAE2E21BFFC51ADCFD0EFD9B2679" bold="true" box="[508,514,744,768]" italics="true" pageId="6" pageNumber="6"></emphasis>
species complex was first defined by
<bibRefCitation id="EFD64B01E21BFFC518C9FD0EFBE62679" author="Bruce NL." box="[1001,1151,744,768]" journalOrPublisher="Records Aust Museum, Suppl. 6" pageId="6" pageNumber="6" pagination="1 - 239" refString="Bruce NL. 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records Aust Museum, Suppl. 6: 1 - 239." title="Cirolanidae (Crustacea: Isopoda) of Australia" type="book chapter" year="1986">
Bruce (
<date id="FFF91030E21BFFC51F1FFD0EFBEF2679" box="[1087,1142,744,768]" pageId="6" pageNumber="6" value="1986">1986</date>
)
</bibRefCitation>
. Subsequently,
<bibRefCitation id="EFD64B01E21BFFC51A69FCEDFE43265A" author="Bruce NL." box="[329,474,779,803]" journalOrPublisher="Cah Biol Mar" pageId="6" pageNumber="6" pagination="375 - 413" part="35" refString="Bruce NL. 1994. Cirolana and related marine isopod crustacean genera (family Cirolanidae) from the coral reefs of Madang, Papua New Guinea. Cah Biol Mar. 35: 375 - 413." title="Cirolana and related marine isopod crustacean genera (family Cirolanidae) from the coral reefs of Madang, Papua New Guinea" type="journal article" year="1994">
Bruce (
<date id="FFF91030E21BFFC51ABBFCEDFE4B265A" box="[411,466,779,803]" pageId="6" pageNumber="6" value="1994">1994</date>
)
</bibRefCitation>
listed 23 morphologically similar species in the group.
<bibRefCitation id="EFD64B01E21BFFC51F48FCEDFF4E263C" author="Khalaji-Pirbalouty V &amp; Wagele JW." journalOrPublisher="Zootaxa" pageId="6" pageNumber="6" pagination="33 - 46" part="2930" refString="Khalaji-Pirbalouty V, Wagele JW. 2011. Two new species of cirolanid isopods (Crustacea: Isopoda: Cirolanidae) from Qeshm and Kish Islands in the Persian Gulf. Zootaxa. 2930: 33 - 46." title="Two new species of cirolanid isopods (Crustacea: Isopoda: Cirolanidae) from Qeshm and Kish Islands in the Persian Gulf" type="journal article" year="2011">
By
<date id="FFF91030E21BFFC51B80FCCBFF4E263C" box="[160,215,813,837]" pageId="6" pageNumber="6" value="2011">2011</date>
</bibRefCitation>
the number of species in this group had grown to 28 (see Appendix 1) (
<bibRefCitation id="EFD64B01E21BFFC51F19FCCBFE8E2611" author="Brusca RC &amp; Wetzer R &amp; France SC." journalOrPublisher="Proc San Diego Nat Hist Museum" pageId="6" pageNumber="6" pagination="1 - 96" part="30" refString="Brusca RC, Wetzer R, France SC. 1995. Cirolanidae (Crustacea: Isopoda: Flabellifera) of the Tropical Eastern Pacific. Proc San Diego Nat Hist Museum. 30: 1 - 96." title="Cirolanidae (Crustacea: Isopoda: Flabellifera) of the Tropical Eastern Pacific" type="journal article" year="1995">
Brusca et al.
<date id="FFF91030E21BFFC51BC0FCB6FE8E2611" box="[224,279,848,872]" pageId="6" pageNumber="6" value="1995">1995</date>
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;
<bibRefCitation id="EFD64B01E21BFFC51A04FCB6FE3A2611" author="Bruce NL." box="[292,419,848,872]" journalOrPublisher="Species Divers" pageId="6" pageNumber="6" pagination="47 - 66" part="9" refString="Bruce NL. 2004. New species of the Cirolana &quot; parva - group &quot; (Crustacea: isopoda: Cirolanidae) from coastal habitats around New Zealand. Species Divers. 9: 47 - 66." title="New species of the Cirolana &quot; parva - group &quot; (Crustacea: isopoda: Cirolanidae) from coastal habitats around New Zealand" type="journal article" year="2004">
Bruce
<date id="FFF91030E21BFFC51A4CFCB6FE3A2611" box="[364,419,848,872]" pageId="6" pageNumber="6" value="2004">2004</date>
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;
<bibRefCitation id="EFD64B01E21BFFC51A90FCB6FD4F2611" author="Schotte M &amp; Kensley B." box="[432,726,848,872]" journalOrPublisher="J Nat Hist" pageId="6" pageNumber="6" pagination="1211 - 1282" part="39" refString="Schotte M, Kensley B. 2005. New species and records of Flabellifera from the Indian Ocean (Crustacea: Peracarida: Isopoda). J Nat Hist. 39: 1211 - 1282." title="New species and records of Flabellifera from the Indian Ocean (Crustacea: Peracarida: Isopoda)" type="journal article" year="2005">
Schotte and Kensley
<date id="FFF91030E21BFFC519BFFCB6FD4F2611" box="[671,726,848,872]" pageId="6" pageNumber="6" value="2005">2005</date>
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;
<bibRefCitation id="EFD64B01E21BFFC519C4FCB6FBEF2611" author="Khalaji-Pirbalouty V &amp; Wagele JW." box="[740,1142,848,872]" journalOrPublisher="Zootaxa" pageId="6" pageNumber="6" pagination="33 - 46" part="2930" refString="Khalaji-Pirbalouty V, Wagele JW. 2011. Two new species of cirolanid isopods (Crustacea: Isopoda: Cirolanidae) from Qeshm and Kish Islands in the Persian Gulf. Zootaxa. 2930: 33 - 46." title="Two new species of cirolanid isopods (Crustacea: Isopoda: Cirolanidae) from Qeshm and Kish Islands in the Persian Gulf" type="journal article" year="2011">
Khalaji-Pirbalouty and Wägele
<date id="FFF91030E21BFFC51F1FFCB6FBEF2611" box="[1087,1142,848,872]" pageId="6" pageNumber="6" value="2011">2011</date>
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).
<bibRefCitation id="EFD64B01E21BFFC51B80FC95FEEB26F2" author="Bruce NL." box="[160,370,883,907]" journalOrPublisher="Species Divers" pageId="6" pageNumber="6" pagination="47 - 66" part="9" refString="Bruce NL. 2004. New species of the Cirolana &quot; parva - group &quot; (Crustacea: isopoda: Cirolanidae) from coastal habitats around New Zealand. Species Divers. 9: 47 - 66." title="New species of the Cirolana &quot; parva - group &quot; (Crustacea: isopoda: Cirolanidae) from coastal habitats around New Zealand" type="journal article" year="2004" yearSuffix="p">
Bruce (
<date id="FFF91030E21BFFC51BCFFC95FEBF26F2" box="[239,294,883,907]" pageId="6" pageNumber="6" value="2004">2004</date>
, p. 49)
</bibRefCitation>
defined the
<emphasis id="B933EAE2E21BFFC51925FC94FD3226F3" box="[517,683,882,906]" italics="true" pageId="6" pageNumber="6">
<taxonomicName id="4C474D73E21BFFC51925FC94FDF926F3" box="[517,608,882,906]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="6" phylum="Arthropoda" rank="genus">Cirolana</taxonomicName>
<emphasis id="B933EAE2E21BFFC51948FC94FDF426F3" bold="true" box="[616,621,882,906]" italics="true" pageId="6" pageNumber="6"></emphasis>
<taxonomicName id="4C474D73E21BFFC5194DFC95FD3226F3" box="[621,683,883,906]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="6" phylum="Arthropoda" rank="species" species="parva">parva</taxonomicName>
</emphasis>
group
<emphasis id="B933EAE2E21BFFC519D8FC94FD6726F3" bold="true" box="[760,766,882,906]" italics="true" pageId="6" pageNumber="6"></emphasis>
as being characterised by smooth body dorsal surfaces with brown or black chromatophores; rostral point present, folded ventrally and posteriorly, in contact with frontal lamina; frontal lamina always pentagonal in shape; pleonites 3 and 4 with posterolateral margins produced, acute or round, pleotelson linguiform, with small distinct median point; the uropodal exopod with a continuous row of robust setae and plumose marginal setae along the lateral margin; uropodal rami apically acute, with bifid or sub bifid apices; penial process are absent or present as small papillae (occasionally mesially in contact or fused).
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>