110 lines
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110 lines
12 KiB
XML
<document ID-DOI="10.5281/zenodo.3245327" ID-GBIF-Dataset="9bf3a9f5-cd36-4be6-bba9-49607815bcd0" ID-Zenodo-Dep="3245327" approvalRequired="3" approvalRequired_for_bibRefs="3" checkinTime="1559900965784" checkinUser="jeremy" docAuthor="Holtz, T. R." docDate="2001" docId="03EE87AEFFA6FF8AFEEDFBAC8E9C0DDE" docLanguage="en" docName="Holtz2001TyrannosauridaeGS.pdf.imf" docOrigin="Mesozoic Vertebrate Life, Bloomington: Indiana University Press" docStyle="DocumentStyle{}" docTitle="Aublysodontinae Leidy 1868" docType="treatment" docVersion="18" lastPageNumber="67" masterDocId="FFD7FFD6FFA5FF89FFE1FFFB8A660428" masterDocTitle="The phylogeny and taxonomy of the Tyrannosauridae" masterLastPageNumber="83" masterPageNumber="64" pageNumber="67" updateTime="1645527105248" updateUser="jeremy">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The phylogeny and taxonomy of the Tyrannosauridae</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Holtz, T. R.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:originInfo>
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<mods:dateIssued>2001</mods:dateIssued>
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<mods:dateOther type="pubDate">2001-12-31</mods:dateOther>
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<mods:publisher>Indiana University Press</mods:publisher>
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<mods:place>
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<mods:placeTerm>Bloomington</mods:placeTerm>
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</mods:place>
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</mods:originInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Editor</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tanke D. H.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Editor</mods:roleTerm>
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</mods:role>
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<mods:namePart>Carpenter K.</mods:namePart>
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</mods:name>
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<mods:titleInfo>
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<mods:title>Mesozoic Vertebrate Life</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:extent unit="page">
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<mods:start>64</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>book chapter</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.3245327</mods:identifier>
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<mods:identifier type="GBIF-Dataset">9bf3a9f5-cd36-4be6-bba9-49607815bcd0</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3245327</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5204480" ID-GBIF-Taxon="157436187" ID-Zenodo-Dep="5204480" LSID="urn:lsid:plazi:treatment:03EE87AEFFA6FF8AFEEDFBAC8E9C0DDE" httpUri="http://treatment.plazi.org/id/03EE87AEFFA6FF8AFEEDFBAC8E9C0DDE" lastPageNumber="67" pageId="3" pageNumber="67">
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<subSubSection box="[268,561,1111,1148]" pageId="3" pageNumber="67" type="nomenclature">
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<paragraph blockId="3.[191,1394,157,2652]" box="[268,561,1111,1148]" pageId="3" pageNumber="67">
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<taxonomicName authorityName="Leidy" authorityYear="1868" box="[268,561,1111,1148]" class="Reptilia" family="Tyrannosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="67" phylum="Chordata" rank="subFamily" subFamily="Aublysodontinae">Aublysodontinae</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="67" type="discussion">
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<paragraph blockId="3.[191,1394,157,2652]" pageId="3" pageNumber="67">
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is comprised at present of only incompletely known taxa. These forms are united by two dental synapomorphies: 76.I and 77.1 (unserrated premaxillary teeth with prominent vertical ridges on the caudal surface). These derived features are not present on other tyrannosaurids, including the oldest confirmed tyrannosaurid premaxillary tooth (from the Early Cretaceous Jobu Formation of Japan:
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<bibRefCitation author="Manabe" box="[316,564,1411,1448]" firstAuthor="Manabe" journalOrPublisher="Journal of Paleontology" pageId="3" pageNumber="67" pagination="1176 - 1178" part="73" refId="ref5697" refString="Manabe, M. 1999. The early evolution of the Tyrannosauridae in Asia. Journal of Paleontology 73: 1176 - 1178." title="The early evolution of the Tyrannosauridae in Asia" type="journal volume" year="1999">Manabe 1999</bibRefCitation>
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). Although
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<bibRefCitation author="Molnar and Carpenter" box="[771,1293,1408,1448]" firstAuthor="Molnar" journalOrPublisher="Geobios" pageId="3" pageNumber="67" pagination="445 - 454" part="22" refId="ref5793" refString="Molnar, R. E., and K. Carpenter. 1989. The Jordan theropod (Maastrichtian, Montana, U. S. A.) referred to the genus Aublysodon. Geobios 22: 445 - 454." title="The Jordan theropod (Maastrichtian, Montana, U. S. A.) referred to the genus Aublysodon" type="journal article" year="1989">Molnar and Carpenter (1989)</bibRefCitation>
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separated
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<taxonomicName authorityName="Leidy" authorityYear="1868" box="[293,500,1461,1498]" class="Reptilia" family="Tyrannosauridae" genus="Aublysodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="67" phylum="Chordata" rank="genus">
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<emphasis box="[293,500,1461,1498]" italics="true" pageId="3" pageNumber="67">Aublysodon</emphasis>
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</taxonomicName>
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from
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<taxonomicName box="[610,891,1462,1497]" class="Reptilia" family="Tyrannosauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="67" phylum="Chordata" rank="family">Tyrannosauridae</taxonomicName>
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as Family
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<taxonomicName authorityName="Nopcsa" authorityYear="1928" box="[1089,1371,1462,1497]" class="Reptilia" family="Aublysodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="67" phylum="Chordata" rank="family">Aublysodontidae</taxonomicName>
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, the presence of Aublysodon-llke teeth in specimens of
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<taxonomicName class="Reptilia" family="Not" genus="Alectrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="67" phylum="Chordata" rank="species" species="olseni">
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<emphasis box="[1155,1384,1512,1547]" italics="true" pageId="3" pageNumber="67">Alectrosaurus</emphasis>
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<emphasis box="[194,296,1562,1597]" italics="true" pageId="3" pageNumber="67">olseni</emphasis>
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</taxonomicName>
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, a taxon with tyrannosaurid synapomorphies (
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<bibRefCitation author="Mader and Bradley" firstAuthor="Mader" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="3" pageNumber="67" pagination="41, - 55" part="9" refId="ref5532" refString="Mader, B. J., and R. L. Bradley. 1989. A redescription and revised diagnosis of the syntypes of the Mongolian tyrannosaur Alectrosaurus olseni. Journal of Vertebrate Paleontology 9: 41, - 55." title="A redescription and revised diagnosis of the syntypes of the Mongolian tyrannosaur Alectrosaurus olseni" type="journal article" year="1989">Mader and Bradley 1989</bibRefCitation>
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) led
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<bibRefCitation author="Currie" box="[431,763,1611,1648]" editor="K. Carpenter & P. J. Currie" etAl="et al." firstAuthor="Currie" journalOrPublisher="Cambridge: Cambridge University Press" pageId="3" pageNumber="67" pagination="107 - 135" refId="ref4897" refString="Currie, P. J., J. K. Rigby Jr., and R. E. Sloan. 1990. Theropod teeth from the Judith River Formation of southern Alberta, Canada. In K. Carpenter and P. J. Currie (eds.), Dinosaur Systematics: Approaches and Perspectives, pp. 107 - 135. Cambridge: Cambridge University Press." title="Theropod teeth from the Judith River Formation of southern Alberta, Canada" type="book chapter" volumeTitle="Dinosaur Systematics: Approaches and Perspectives" year="1990">
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Currie et
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<emphasis box="[600,634,1614,1647]" italics="true" pageId="3" pageNumber="67">al</emphasis>
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. (1990)
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</bibRefCitation>
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to include this group as a subfamily within
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<taxonomicName box="[311,597,1662,1697]" class="Reptilia" family="Tyrannosauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="67" phylum="Chordata" rank="family">Tyrannosauridae</taxonomicName>
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, a nomenclatural practice followed here. Other than the specialized premaxillary teeth, aublysodontines generally retain more primitive coelurosaurian features lost in other tyrannosaurids: for example, the skulls of aublysodontines are longer and lower, the tooth count higher, and the lateral teeth less labiolingually expanded than in derived tyrannosaurines (
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<bibRefCitation author="Currie, P. J." box="[960,1225,1909,1949]" editor="M. Benton & E. Kurochkin & M. Shiskin & D. Unwin" journalOrPublisher="Cambridge Univer- sity Press" pageId="3" pageNumber="67" refId="ref4795" refString="Currie, P. J. in press. Theropods from the Cretaceous of Mongolia. In M. Benton, E. Kurochkin, M. Shiskin, and D. Unwin (eds.), The Age of Dinosaurs in Russia and Mongolia. Cambridge: Cambridge Univer- sity Press." title="Theropods from the Cretaceous of Mongolia" type="book chapter" volumeTitle="The Age of Dinosaurs in Russia and Mongolia" year="in press">Currie in press</bibRefCitation>
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).
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<taxonomicName authorityName="Leidy" authorityYear="1868" class="Reptilia" family="Tyrannosauridae" genus="Aublysodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="67" phylum="Chordata" rank="genus">
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<emphasis box="[1260,1385,1913,1948]" italics="true" pageId="3" pageNumber="67">Aublysodon</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Gilmore" authorityYear="1933" box="[365,594,1962,1997]" class="Reptilia" family="Tyrannosauridae" genus="Alectrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="67" phylum="Chordata" rank="genus">
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<emphasis box="[365,594,1962,1997]" italics="true" pageId="3" pageNumber="67">Alectrosaurus</emphasis>
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</taxonomicName>
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(but not the Kirtland Shale taxon) are smaller than typical tyrannosaurines: however the ontogenetic status of known aublysodontine specimens is hindered by the incompleteness of their fossils, and these might represent juvenile individuals of taxa which reached larger sizes. Aithough
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<taxonomicName authorityName="Leidy" authorityYear="1868" box="[717,924,2162,2199]" class="Reptilia" family="Tyrannosauridae" genus="Aublysodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="67" phylum="Chordata" rank="genus">
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<emphasis box="[717,924,2162,2199]" italics="true" pageId="3" pageNumber="67">Aublysodon</emphasis>
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</taxonomicName>
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is provisionally recognized as a valid name here,
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<bibRefCitation author="Currie" box="[566,893,2211,2249]" editor="K. Carpenter & P. J. Currie" etAl="et al." firstAuthor="Currie" journalOrPublisher="Cambridge: Cambridge University Press" pageId="3" pageNumber="67" pagination="107 - 135" refId="ref4897" refString="Currie, P. J., J. K. Rigby Jr., and R. E. Sloan. 1990. Theropod teeth from the Judith River Formation of southern Alberta, Canada. In K. Carpenter and P. J. Currie (eds.), Dinosaur Systematics: Approaches and Perspectives, pp. 107 - 135. Cambridge: Cambridge University Press." title="Theropod teeth from the Judith River Formation of southern Alberta, Canada" type="book chapter" volumeTitle="Dinosaur Systematics: Approaches and Perspectives" year="1990">
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Currie et
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<emphasis box="[730,766,2212,2249]" italics="true" pageId="3" pageNumber="67">al</emphasis>
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. (1990)
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</bibRefCitation>
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caution that this may not be a taxon, but instead an ontogenetic stage or a sexual dimorph. It might be further added that tooth wear in life, postmortem abrasion, and digestion might also conceivably render a typical tyrannosaurid premaxilla tooth into a nonserrated one. In such a case, the synapomorphies uniting "
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<taxonomicName authorityName="Leidy" authorityYear="1868" box="[508,803,2465,2502]" class="Reptilia" family="Tyrannosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="67" phylum="Chordata" rank="subFamily" subFamily="Aublysodontinae">Aublysodontinae</taxonomicName>
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" would be lost, and the remaining resemblances between these taxa would be symplesiomorphies.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |