treatments-xml/data/03/EE/87/03EE878B3970B020FF7CFDFDFD96FF17.xml

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<mods:title id="7F746CDC3A7F799112427287730804F9">A new species of king cricket Glaphyrosoma Brunner von Wattenwyl, 1888 (Orthoptera: Anostostomatidae: Glaphyrosomatini) from Costa Rica with behavioral observations</mods:title>
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<mods:namePart id="8CC1CAE5EA93F17C034F690BBF0093E8">Koehl, Jonathan</mods:namePart>
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<paragraph id="8BF8369D3970B02AFF7CFDFDFE8CFDD3" blockId="2.[151,1181,513,576]" pageId="2" pageNumber="95">
<heading id="D0B081F13970B02AFF7CFDFDFE8CFDD3" bold="true" fontSize="11" level="1" pageId="2" pageNumber="95" reason="1">
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<taxonomicName id="4C474D1E3970B02AFF7CFDFDFC4CFDF0" authority="Richardson, Trimm, Paderes, Koehl" authorityName="Richardson, Trimm, Paderes, Koehl" box="[151,966,513,540]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="stephanosoltis">
<emphasis id="B933EA8F3970B02AFF7CFDFDFE71FDF0" bold="true" box="[151,507,513,540]" italics="true" pageId="2" pageNumber="95">Glaphyrosoma stephanosoltis</emphasis>
Richardson, Trimm, Paderes, Koehl
</taxonomicName>
, &amp; Song, sp. nov.
</emphasis>
<figureCitation id="137C2A183970B02AFF7CFDD9FE8CFDD3" box="[151,262,549,576]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="2.[151,250,1967,1992]" captionStartId-1="3.[151,250,1814,1839]" captionStartId-10="10.[151,250,1847,1872]" captionStartId-2="4.[151,250,1199,1224]" captionStartId-3="4.[151,250,1978,2003]" captionStartId-4="5.[151,250,682,707]" captionStartId-5="5.[151,250,1928,1953]" captionStartId-6="6.[151,250,1401,1426]" captionStartId-7="7.[151,250,917,942]" captionStartId-8="8.[151,250,1415,1440]" captionStartId-9="9.[151,250,1225,1250]" captionTargetBox-0="[182,1410,1369,1918]" captionTargetBox-1="[183,1403,1306,1763]" captionTargetBox-10="[203,1385,376,1778]" captionTargetBox-2="[175,1414,205,1154]" captionTargetBox-3="[279,1281,1264,1949]" captionTargetBox-4="[170,1418,201,637]" captionTargetBox-5="[270,1318,787,1886]" captionTargetBox-6="[186,1407,211,1350]" captionTargetBox-7="[201,1386,220,856]" captionTargetBox-8="[239,1394,334,1360]" captionTargetBox-9="[184,1406,205,1179]" captionTargetId-0="figure@2.[158,1429,1353,1941]" captionTargetId-1="figure@3.[159,1429,1281,1788]" captionTargetId-10="figure@10.[162,1425,345,1822]" captionTargetId-2="figure@4.[151,1437,181,1174]" captionTargetId-3="figure@4.[279,1308,1246,1953]" captionTargetId-4="figure@5.[154,1434,181,657]" captionTargetId-5="figure@5.[248,1339,765,1903]" captionTargetId-6="figure@6.[156,1431,181,1376]" captionTargetId-7="figure@7.[166,1421,181,891]" captionTargetId-8="figure@8.[170,1418,273,1390]" captionTargetId-9="figure@9.[159,1429,181,1199]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-10="10" captionTargetPageId-2="4" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="5" captionTargetPageId-6="6" captionTargetPageId-7="7" captionTargetPageId-8="8" captionTargetPageId-9="9" captionText-0="FIGURE 1. Habitus of live specimens of Glaphyrosoma stephanosoltis sp. nov. taken in the field. (A) dorso-lateral view; (B) frontal view." captionText-1="FIGURE 2. (A) frontal view of head, showing the long maxillary palpi; (B) lateral view showing head and thorax." captionText-10="FIGURE 11. Female ovipositor of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of ovipositor; (B) dorsal view of female abdominal terminalia showing cerci and ovipositor." captionText-2="FIGURE 3. Holotype of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view; (B) dorsal view." captionText-3="FIGURE 4. Legs of Glaphyrosoma stephanosoltis sp. nov. (A) fore tibia to tarsi; (B) mid femur to tarsi; (C) Hind tibia to tarsi." captionText-4="FIGURE 5. Tibial tympana of Glaphyrosoma stephanosoltis sp. nov. (A) male, showing outer tympanum in the right fore tibia and inner tympanum in the left fore tibia; (B) female outer tympanum; (C) female inner tympanum." captionText-5="FIGURE 6. Stridulatory apparatus of Glaphyrosoma stephanosoltis sp. nov. (A) male and (B) female, arrows pointing to patches of granular stridulatory pegs in the first and second abdominal tergites; (C) male and (D) female, arrow pointing to diagonal rows of stridulatory pegs in the inner side of hind femur." captionText-6="FIGURE 7. Male external genitalia of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of abdominal segments 4–10; (B) posterior view of abdomen, showing cerci, paraproctal processes, and tenth tergites; (C) dorsal view of ninth and tenth abdominal tergites; (D) ventral view showing male subgenital plate." captionText-7="FIGURE 8. Male internal genitalia of Glaphyrosoma stephanosoltis sp. nov. (A) male internal genitalia in their natural position inside abdomen, showing their relative position to the subgenital plate; (B) dissected and magnified view of male internal genitalia." captionText-8="FIGURE 9. (A) posterior view of male external genitalia showing paraproctal processes; (B) ventral view of male subgenital plate; (C) lateral view of male ninth and tenth abdominal tergites; (D) lateral view of male paraproctal process; (E) ventral view of female subgenital plate." captionText-9="FIGURE 10. Female of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view; (B) dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.3450270" figureDoi-1="http://doi.org/10.5281/zenodo.3450274" figureDoi-10="http://doi.org/10.5281/zenodo.3450304" figureDoi-2="http://doi.org/10.5281/zenodo.3450280" figureDoi-3="http://doi.org/10.5281/zenodo.3450284" figureDoi-4="http://doi.org/10.5281/zenodo.3450286" figureDoi-5="http://doi.org/10.5281/zenodo.3450290" figureDoi-6="http://doi.org/10.5281/zenodo.3450296" figureDoi-7="http://doi.org/10.5281/zenodo.3450298" figureDoi-8="http://doi.org/10.5281/zenodo.3450300" figureDoi-9="http://doi.org/10.5281/zenodo.3450302" httpUri-0="https://zenodo.org/record/3450270/files/figure.png" httpUri-1="https://zenodo.org/record/3450274/files/figure.png" httpUri-10="https://zenodo.org/record/3450304/files/figure.png" httpUri-2="https://zenodo.org/record/3450280/files/figure.png" httpUri-3="https://zenodo.org/record/3450284/files/figure.png" httpUri-4="https://zenodo.org/record/3450286/files/figure.png" httpUri-5="https://zenodo.org/record/3450290/files/figure.png" httpUri-6="https://zenodo.org/record/3450296/files/figure.png" httpUri-7="https://zenodo.org/record/3450298/files/figure.png" httpUri-8="https://zenodo.org/record/3450300/files/figure.png" httpUri-9="https://zenodo.org/record/3450302/files/figure.png" pageId="2" pageNumber="95">
<emphasis id="B933EA8F3970B02AFF7CFDD9FE8CFDD3" bold="true" box="[151,262,549,576]" pageId="2" pageNumber="95">Figs 1–11</emphasis>
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urn:lsid: 
<taxonomicName id="4C474D1E3970B02AFF05FD91FEE4FD64" authorityName=", Barranco" authorityYear="2010" box="[238,366,621,648]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
.speciesfile.org:TaxonName:506875
</paragraph>
</subSubSection>
<subSubSection id="C35D65163970B02BFF7CFD49FAE0F853" lastPageId="3" lastPageNumber="96" pageId="2" pageNumber="95" type="diagnosis">
<paragraph id="8BF8369D3970B02AFF7CFD49FA82FBFF" blockId="2.[151,1437,693,1332]" pageId="2" pageNumber="95">
<emphasis id="B933EA8F3970B02AFF7CFD49FDE3FD23" box="[151,617,693,719]" italics="true" pageId="2" pageNumber="95">
<emphasis id="B933EA8F3970B02AFF7CFD49FE9BFD23" bold="true" box="[151,273,693,719]" italics="true" pageId="2" pageNumber="95">Diagnosis.</emphasis>
<taxonomicName id="4C474D1E3970B02AFEF1FD49FDE3FD23" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[282,617,693,719]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">Glaphyrosoma stephanosoltis</taxonomicName>
</emphasis>
<emphasis id="B933EA8F3970B02AFD99FD4AFD44FD3C" bold="true" box="[626,718,694,720]" pageId="2" pageNumber="95">
<taxonomicNameLabel id="A20057F43970B02AFD99FD4AFD44FD3C" box="[626,718,694,720]" pageId="2" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is dorsally colored in solid dark brown (
<figureCitation id="137C2A183970B02AFB4EFD49FB7EFD23" box="[1189,1268,693,719]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1992]" captionTargetBox="[182,1410,1369,1918]" captionTargetId="figure@2.[158,1429,1353,1941]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of live specimens of Glaphyrosoma stephanosoltis sp. nov. taken in the field. (A) dorso-lateral view; (B) frontal view." figureDoi="http://doi.org/10.5281/zenodo.3450270" httpUri="https://zenodo.org/record/3450270/files/figure.png" pageId="2" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="137C2A183970B02AFAEEFD49FA9EFD23" box="[1285,1300,693,719]" captionStart="FIGURE 3" captionStartId="4.[151,250,1199,1224]" captionTargetBox="[175,1414,205,1154]" captionTargetId="figure@4.[151,1437,181,1174]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3450280" httpUri="https://zenodo.org/record/3450280/files/figure.png" pageId="2" pageNumber="95">3</figureCitation>
, 
<figureCitation id="137C2A183970B02AFAC8FD49FACBFD23" box="[1315,1345,693,719]" captionStart="FIGURE 10" captionStartId="9.[151,250,1225,1250]" captionTargetBox="[184,1406,205,1179]" captionTargetId="figure@9.[159,1429,181,1199]" captionTargetPageId="9" captionText="FIGURE 10. Female of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3450302" httpUri="https://zenodo.org/record/3450302/files/figure.png" pageId="2" pageNumber="95">10</figureCitation>
), and is similar in overall coloration to 
<emphasis id="B933EA8F3970B02AFE06FD26FEEDFCFB" italics="true" pageId="2" pageNumber="95">
<taxonomicName id="4C474D1E3970B02AFE06FD26FDEBFD1F" authorityName="Brunner von Wattenwyl" authorityYear="1888" box="[493,609,729,756]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="gracile">G. gracile</taxonomicName>
, 
<taxonomicName id="4C474D1E3970B02AFD80FD26FD52FD1F" authorityName="Cadena-Castaneda &amp; Gorochov" authorityYear="2016" box="[619,728,729,756]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="anderi">G. anderi</taxonomicName>
, 
<taxonomicName id="4C474D1E3970B02AFD09FD26FCD3FD1F" authorityName="Gorochov" authorityYear="2016" box="[738,857,729,756]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="beretka">G. beretka</taxonomicName>
, 
<taxonomicName id="4C474D1E3970B02AFC88FD26FC7EFD1F" authorityName="Gorochov &amp; Cadena-Castaneda" authorityYear="2016" box="[867,1012,729,756]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="bulbosum">G. bulbosum</taxonomicName>
, 
<taxonomicName id="4C474D1E3970B02AFC15FD26FB2AFD1F" authorityName="Gorochov" authorityYear="2016" box="[1022,1184,729,756]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="pushenkovi">G. pushenkovi</taxonomicName>
, 
<taxonomicName id="4C474D1E3970B02AFB41FD26FA12FD18" authorityName="Cadena-Castaneda &amp; Monzon-Sierra" authorityYear="2017" box="[1194,1432,730,756]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="franciscoasturiasi">G. franciscoasturiasi</taxonomicName>
, 
<taxonomicName id="4C474D1E3970B02AFF7CFD02FEE9FCFB" authorityName="Cadena-Castaneda &amp; Monzon-Sierra" authorityYear="2017" box="[151,355,765,792]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="hectorcentenoi">G. hectorcentenoi</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C474D1E3970B02AFE4AFD02FDFEFCFB" authorityName="Cadena-Castaneda &amp; Monzon-Sierra" authorityYear="2017" box="[417,628,765,792]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="magnaproctalis">
<emphasis id="B933EA8F3970B02AFE4AFD02FDFEFCFB" box="[417,628,765,792]" italics="true" pageId="2" pageNumber="95">G. magnaproctalis</emphasis>
</taxonomicName>
. However, it can be separated from all other members of the genus by having quadrangular male paraproctal process with a spine protruding medially facing upwards (
<figureCitation id="137C2A183970B02AFB3EFCDDFAECFCD7" box="[1237,1382,801,828]" captionStart="FIGURE 7" captionStartId="6.[151,250,1401,1426]" captionTargetBox="[186,1407,211,1350]" captionTargetId="figure@6.[156,1431,181,1376]" captionTargetPageId="6" captionText="FIGURE 7. Male external genitalia of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of abdominal segments 4–10; (B) posterior view of abdomen, showing cerci, paraproctal processes, and tenth tergites; (C) dorsal view of ninth and tenth abdominal tergites; (D) ventral view showing male subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450296" httpUri="https://zenodo.org/record/3450296/files/figure.png" pageId="2" pageNumber="95">Figs. 7B, 7C</figureCitation>
, 
<figureCitation id="137C2A183970B02AFA98FCDDFF37FCB3" captionStart="FIGURE 9" captionStartId="8.[151,250,1415,1440]" captionTargetBox="[239,1394,334,1360]" captionTargetId="figure@8.[170,1418,273,1390]" captionTargetPageId="8" captionText="FIGURE 9. (A) posterior view of male external genitalia showing paraproctal processes; (B) ventral view of male subgenital plate; (C) lateral view of male ninth and tenth abdominal tergites; (D) lateral view of male paraproctal process; (E) ventral view of female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450300" httpUri="https://zenodo.org/record/3450300/files/figure.png" pageId="2" pageNumber="95">9A, 9D</figureCitation>
). This species is morphologically most similar to 
<taxonomicName id="4C474D1E3970B02AFD03FCBAFC33FCB3" authorityName="Cadena-Castaneda &amp; Monzon-Sierra" authorityYear="2017" box="[744,953,837,864]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="magnaproctalis">
<emphasis id="B933EA8F3970B02AFD03FCBAFC33FCB3" box="[744,953,837,864]" italics="true" pageId="2" pageNumber="95">G. magnaproctalis</emphasis>
</taxonomicName>
according to the key presented by Cadena- Castañeda &amp; Monzón-Sierra (2017), but differs in the position of the spine on male paraproctal process and the appearance of the male subgenital plate as well as the length of styli. The spine arises medially in 
<taxonomicName id="4C474D1E3970B02AFB58FC72FAFFFC4B" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[1203,1397,909,936]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">
<emphasis id="B933EA8F3970B02AFB58FC72FAFFFC4B" box="[1203,1397,909,936]" italics="true" pageId="2" pageNumber="95">G. stephanosoltis</emphasis>
</taxonomicName>
<emphasis id="B933EA8F3970B02AFA90FC72FF42FC20" bold="true" pageId="2" pageNumber="95">
<taxonomicNameLabel id="A20057F43970B02AFA90FC72FF42FC20" pageId="2" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
instead of being located at the top of the paraproctal process. The subgenital plate also shows more curvature between the styli forming a deep “V” shaped notch half the length of the subgenital plate (
<figureCitation id="137C2A183970B02AFB65FC29FB6DFC1C" box="[1166,1255,981,1008]" captionStart="FIGURE 7" captionStartId="6.[151,250,1401,1426]" captionTargetBox="[186,1407,211,1350]" captionTargetId="figure@6.[156,1431,181,1376]" captionTargetPageId="6" captionText="FIGURE 7. Male external genitalia of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of abdominal segments 4–10; (B) posterior view of abdomen, showing cerci, paraproctal processes, and tenth tergites; (C) dorsal view of ninth and tenth abdominal tergites; (D) ventral view showing male subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450296" httpUri="https://zenodo.org/record/3450296/files/figure.png" pageId="2" pageNumber="95">Fig. 7D</figureCitation>
, 
<figureCitation id="137C2A183970B02AFB18FC29FA9EFC03" box="[1267,1300,981,1007]" captionStart="FIGURE 9" captionStartId="8.[151,250,1415,1440]" captionTargetBox="[239,1394,334,1360]" captionTargetId="figure@8.[170,1418,273,1390]" captionTargetPageId="8" captionText="FIGURE 9. (A) posterior view of male external genitalia showing paraproctal processes; (B) ventral view of male subgenital plate; (C) lateral view of male ninth and tenth abdominal tergites; (D) lateral view of male paraproctal process; (E) ventral view of female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450300" httpUri="https://zenodo.org/record/3450300/files/figure.png" pageId="2" pageNumber="95">9B</figureCitation>
) and differs from the subgenital plate of 
<taxonomicName id="4C474D1E3970B02AFE3FFC06FD2FFBFF" authorityName="Cadena-Castaneda &amp; Monzon-Sierra" authorityYear="2017" box="[468,677,1017,1044]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="magnaproctalis">
<emphasis id="B933EA8F3970B02AFE3FFC06FD2FFBFF" box="[468,677,1017,1044]" italics="true" pageId="2" pageNumber="95">G. magnaproctalis</emphasis>
</taxonomicName>
in which there is almost no notch in between the styli.
</paragraph>
<paragraph id="8BF8369D3970B02AFF2CFBE1FC0EFADF" blockId="2.[151,1437,693,1332]" pageId="2" pageNumber="95">
<emphasis id="B933EA8F3970B02AFF2CFBE1FEC0FBD4" bold="true" box="[199,330,1053,1080]" italics="true" pageId="2" pageNumber="95">Coloration.</emphasis>
Dorsal portion of head dark brown, nearing black; gena and frons dark brown; clypeus light yellow. Median ocellus and two upper ocelli pinkish cream white (when alive) with light beige rings (
<figureCitation id="137C2A183970B02AFB48FBBDFB69FBB7" box="[1187,1251,1089,1115]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1992]" captionTargetBox="[182,1410,1369,1918]" captionTargetId="figure@2.[158,1429,1353,1941]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of live specimens of Glaphyrosoma stephanosoltis sp. nov. taken in the field. (A) dorso-lateral view; (B) frontal view." figureDoi="http://doi.org/10.5281/zenodo.3450270" httpUri="https://zenodo.org/record/3450270/files/figure.png" pageId="2" pageNumber="95">Fig. 1</figureCitation>
). Antennae dark brown. Labrum white (when alive); mandibles light brown at base and dark brown at the tip; maxillary palpi and labial palpi creamy white. Entire dorsal region from thorax to abdomen dark brown with no delineated coloration patterns. Lateral margin of thoracic and abdominal tergites lighter brown. Sternum yellow/brown. Fore femora and mid femora yellowish brown distally, and darker brown apically. Hind femora lighter brown distally, gradually becoming darker apically and dorsally, with apical end (knee) abruptly becoming creamy white (
<figureCitation id="137C2A183970B02AFB32FB09FA91FAE3" box="[1241,1307,1269,1295]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1992]" captionTargetBox="[182,1410,1369,1918]" captionTargetId="figure@2.[158,1429,1353,1941]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of live specimens of Glaphyrosoma stephanosoltis sp. nov. taken in the field. (A) dorso-lateral view; (B) frontal view." figureDoi="http://doi.org/10.5281/zenodo.3450270" httpUri="https://zenodo.org/record/3450270/files/figure.png" pageId="2" pageNumber="95">Fig. 1</figureCitation>
). All tibiae distally dark brown, apically creamy white. All tarsi creamy white.
</paragraph>
<caption id="DF3866153970B02AFF7CF853FE92F800" ID-DOI="http://doi.org/10.5281/zenodo.3450270" ID-Zenodo-Dep="3450270" httpUri="https://zenodo.org/record/3450270/files/figure.png" pageId="2" pageNumber="95" startId="2.[151,250,1967,1992]" targetBox="[182,1410,1369,1918]" targetPageId="2">
<paragraph id="8BF8369D3970B02AFF7CF853FE92F800" blockId="2.[151,1436,1967,2028]" pageId="2" pageNumber="95">
<emphasis id="B933EA8F3970B02AFF7CF853FE9EF824" bold="true" box="[151,276,1967,1992]" pageId="2" pageNumber="95">FIGURE 1.</emphasis>
Habitus of live specimens of 
<taxonomicName id="4C474D1E3970B02AFDAEF853FCF8F82B" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[581,882,1967,1991]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">
<emphasis id="B933EA8F3970B02AFDAEF853FCF8F82B" box="[581,882,1967,1991]" italics="true" pageId="2" pageNumber="95">Glaphyrosoma stephanosoltis</emphasis>
</taxonomicName>
<emphasis id="B933EA8F3970B02AFC92F853FC40F82B" bold="true" box="[889,970,1967,1991]" pageId="2" pageNumber="95">
<taxonomicNameLabel id="A20057F43970B02AFC92F853FC40F82B" box="[889,970,1967,1991]" pageId="2" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
taken in the field. (A) dorso-lateral view; (B) frontal view.
</paragraph>
</caption>
<paragraph id="8BF8369D3971B02BFF2CFF64FDE4FB2B" blockId="3.[151,1437,152,1223]" pageId="3" pageNumber="96">
<emphasis id="B933EA8F3971B02BFF2CFF64FEC4FF5F" box="[199,334,152,179]" italics="true" pageId="3" pageNumber="96">
<emphasis id="B933EA8F3971B02BFF2CFF64FE83FF5F" bold="true" box="[199,265,152,179]" italics="true" pageId="3" pageNumber="96">Male.</emphasis>
Head
</emphasis>
(
<figureCitation id="137C2A183971B02BFEB6FF65FE58FF5F" box="[349,466,153,179]" captionStart="FIGURE 2" captionStartId="3.[151,250,1814,1839]" captionTargetBox="[183,1403,1306,1763]" captionTargetId="figure@3.[159,1429,1281,1788]" captionTargetPageId="3" captionText="FIGURE 2. (A) frontal view of head, showing the long maxillary palpi; (B) lateral view showing head and thorax." figureDoi="http://doi.org/10.5281/zenodo.3450274" httpUri="https://zenodo.org/record/3450274/files/figure.png" pageId="3" pageNumber="96">Fig. 2A,B</figureCitation>
): Integument smooth. Vertex smooth without any depression, blended to frontal ridge. Gena convex. Frons, smooth, unpronounced, blended with the rest of head; two parallel depressions starting at the base of scape; a pair of small circular depressions above the mandibular condyles. Antennal scape originating from where frontal ocellus is positioned. Antenna filiform and at least three times longer than body. Ocelli round and not protruding from head. Eyes pear-shaped, narrowing dorsoventrally, half as long as the width of gena. Clypeus upside down triangular with broadly round ventrally, concave. Labrum circular. Mandibles narrowing to sharp end, articulation points pronounced below gena. Maxillary palpi very long, nearly three times the dorsoventral length of the head. Labial palpi evident, but not long, shorter than the length of mandible. Both maxillary and labial palpi with bulbous apical tips. 
<emphasis id="B933EA8F3971B02BFE58FE45FD8EFE3F" box="[435,516,441,467]" italics="true" pageId="3" pageNumber="96">Thorax</emphasis>
: Integument smooth. Completely apterous. Pronotum about as long as wide (
<figureCitation id="137C2A183971B02BFA84FE45FF37FE1B" captionStart="FIGURE 3" captionStartId="4.[151,250,1199,1224]" captionTargetBox="[175,1414,205,1154]" captionTargetId="figure@4.[151,1437,181,1174]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3450280" httpUri="https://zenodo.org/record/3450280/files/figure.png" pageId="3" pageNumber="96">Fig. 3A</figureCitation>
); sloping downward anteriorly; lateral lobe of pronotum square basally (
<figureCitation id="137C2A183971B02BFBE3FE21FBE9FE1B" box="[1032,1123,477,503]" captionStart="FIGURE 2" captionStartId="3.[151,250,1814,1839]" captionTargetBox="[183,1403,1306,1763]" captionTargetId="figure@3.[159,1429,1281,1788]" captionTargetPageId="3" captionText="FIGURE 2. (A) frontal view of head, showing the long maxillary palpi; (B) lateral view showing head and thorax." figureDoi="http://doi.org/10.5281/zenodo.3450274" httpUri="https://zenodo.org/record/3450274/files/figure.png" pageId="3" pageNumber="96">Fig. 2B</figureCitation>
). Mesonotal and metanotal lobes rounded distally (
<figureCitation id="137C2A183971B02BFE75FDFDFE7CFDF7" box="[414,502,513,539]" captionStart="FIGURE 2" captionStartId="3.[151,250,1814,1839]" captionTargetBox="[183,1403,1306,1763]" captionTargetId="figure@3.[159,1429,1281,1788]" captionTargetPageId="3" captionText="FIGURE 2. (A) frontal view of head, showing the long maxillary palpi; (B) lateral view showing head and thorax." figureDoi="http://doi.org/10.5281/zenodo.3450274" httpUri="https://zenodo.org/record/3450274/files/figure.png" pageId="3" pageNumber="96">Fig. 2B</figureCitation>
). 
<emphasis id="B933EA8F3971B02BFDE1FDFEFDB5FDF7" box="[522,575,514,539]" italics="true" pageId="3" pageNumber="96">Legs</emphasis>
: Fore tibiae ventrally with 3 pairs of symmetrically arranged spines, 6 apical spurs composed of 1 pair of short symmetrical spines and 2 pairs of apical spurs (
<figureCitation id="137C2A183971B02BFBF6FDD9FB0AFDD3" box="[1053,1152,549,575]" captionStart="FIGURE 4" captionStartId="4.[151,250,1978,2003]" captionTargetBox="[279,1281,1264,1949]" captionTargetId="figure@4.[279,1308,1246,1953]" captionTargetPageId="4" captionText="FIGURE 4. Legs of Glaphyrosoma stephanosoltis sp. nov. (A) fore tibia to tarsi; (B) mid femur to tarsi; (C) Hind tibia to tarsi." figureDoi="http://doi.org/10.5281/zenodo.3450284" httpUri="https://zenodo.org/record/3450284/files/figure.png" pageId="3" pageNumber="96">Figs. 4A</figureCitation>
, 
<figureCitation id="137C2A183971B02BFB61FDD8FB3AFDD3" box="[1162,1200,548,575]" captionStart="FIGURE 5" captionStartId="5.[151,250,682,707]" captionTargetBox="[170,1418,201,637]" captionTargetId="figure@5.[154,1434,181,657]" captionTargetPageId="5" captionText="FIGURE 5. Tibial tympana of Glaphyrosoma stephanosoltis sp. nov. (A) male, showing outer tympanum in the right fore tibia and inner tympanum in the left fore tibia; (B) female outer tympanum; (C) female inner tympanum." figureDoi="http://doi.org/10.5281/zenodo.3450286" httpUri="https://zenodo.org/record/3450286/files/figure.png" pageId="3" pageNumber="96">5A</figureCitation>
); no subapical spines along the dorsal surface; tympanum present as depression at distal portion of inner and outer surface of fore tibiae (
<figureCitation id="137C2A183971B02BFF4BFD91FF73FD6B" box="[160,249,620,647]" captionStart="FIGURE 5" captionStartId="5.[151,250,682,707]" captionTargetBox="[170,1418,201,637]" captionTargetId="figure@5.[154,1434,181,657]" captionTargetPageId="5" captionText="FIGURE 5. Tibial tympana of Glaphyrosoma stephanosoltis sp. nov. (A) male, showing outer tympanum in the right fore tibia and inner tympanum in the left fore tibia; (B) female outer tympanum; (C) female inner tympanum." figureDoi="http://doi.org/10.5281/zenodo.3450286" httpUri="https://zenodo.org/record/3450286/files/figure.png" pageId="3" pageNumber="96">Fig. 5A</figureCitation>
). Middle tibiae with 4 asymmetrical spines dorsally and 3 pairs of symmetrically arranged spines ventrally (
<figureCitation id="137C2A183971B02BFF4BFD6DFF7DFD47" box="[160,247,657,683]" captionStart="FIGURE 4" captionStartId="4.[151,250,1978,2003]" captionTargetBox="[279,1281,1264,1949]" captionTargetId="figure@4.[279,1308,1246,1953]" captionTargetPageId="4" captionText="FIGURE 4. Legs of Glaphyrosoma stephanosoltis sp. nov. (A) fore tibia to tarsi; (B) mid femur to tarsi; (C) Hind tibia to tarsi." figureDoi="http://doi.org/10.5281/zenodo.3450284" httpUri="https://zenodo.org/record/3450284/files/figure.png" pageId="3" pageNumber="96">Fig. 4B</figureCitation>
); 6 apical spurs composed of 1 pair of short symmetrical spines, 1 pair of spurs dorsally, and 1 pair of spurs ventrally (
<figureCitation id="137C2A183971B02BFEE1FD49FEE9FD23" box="[266,355,693,719]" captionStart="FIGURE 4" captionStartId="4.[151,250,1978,2003]" captionTargetBox="[279,1281,1264,1949]" captionTargetId="figure@4.[279,1308,1246,1953]" captionTargetPageId="4" captionText="FIGURE 4. Legs of Glaphyrosoma stephanosoltis sp. nov. (A) fore tibia to tarsi; (B) mid femur to tarsi; (C) Hind tibia to tarsi." figureDoi="http://doi.org/10.5281/zenodo.3450284" httpUri="https://zenodo.org/record/3450284/files/figure.png" pageId="3" pageNumber="96">Fig. 4B</figureCitation>
). Hind tibiae dorsally armed with 18–21 unarticulated spines, 8 apical spurs composed of 1 pair of short spines, 1 pair of short spurs ventrally, two pairs of long spurs (
<figureCitation id="137C2A183971B02BFC55FD25FB93FD1F" box="[958,1049,729,755]" captionStart="FIGURE 4" captionStartId="4.[151,250,1978,2003]" captionTargetBox="[279,1281,1264,1949]" captionTargetId="figure@4.[279,1308,1246,1953]" captionTargetPageId="4" captionText="FIGURE 4. Legs of Glaphyrosoma stephanosoltis sp. nov. (A) fore tibia to tarsi; (B) mid femur to tarsi; (C) Hind tibia to tarsi." figureDoi="http://doi.org/10.5281/zenodo.3450284" httpUri="https://zenodo.org/record/3450284/files/figure.png" pageId="3" pageNumber="96">Fig. 4C</figureCitation>
). Hind femora absent of stripes or patterns (
<figureCitation id="137C2A183971B02BFEEBFD01FED0FCFB" box="[256,346,765,791]" captionStart="FIGURE 3" captionStartId="4.[151,250,1199,1224]" captionTargetBox="[175,1414,205,1154]" captionTargetId="figure@4.[151,1437,181,1174]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3450280" httpUri="https://zenodo.org/record/3450280/files/figure.png" pageId="3" pageNumber="96">Fig. 3A</figureCitation>
); light muscular lining visible through cuticle (
<figureCitation id="137C2A183971B02BFC85FD01FC43FCFB" box="[878,969,765,791]" captionStart="FIGURE 3" captionStartId="4.[151,250,1199,1224]" captionTargetBox="[175,1414,205,1154]" captionTargetId="figure@4.[151,1437,181,1174]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3450280" httpUri="https://zenodo.org/record/3450280/files/figure.png" pageId="3" pageNumber="96">Fig. 3A</figureCitation>
); femoral groove present on outer side of hind femur from base to distal part (
<figureCitation id="137C2A183971B02BFDC3FCDDFD08FCD7" box="[552,642,801,827]" captionStart="FIGURE 3" captionStartId="4.[151,250,1199,1224]" captionTargetBox="[175,1414,205,1154]" captionTargetId="figure@4.[151,1437,181,1174]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3450280" httpUri="https://zenodo.org/record/3450280/files/figure.png" pageId="3" pageNumber="96">Fig. 3A</figureCitation>
). Inner side of hind femora with 3–4 diagonal rows of stridulatory pegs present (
<figureCitation id="137C2A183971B02BFF13FCB9FEDBFC8C" box="[248,337,837,864]" captionStart="FIGURE 6" captionStartId="5.[151,250,1928,1953]" captionTargetBox="[270,1318,787,1886]" captionTargetId="figure@5.[248,1339,765,1903]" captionTargetPageId="5" captionText="FIGURE 6. Stridulatory apparatus of Glaphyrosoma stephanosoltis sp. nov. (A) male and (B) female, arrows pointing to patches of granular stridulatory pegs in the first and second abdominal tergites; (C) male and (D) female, arrow pointing to diagonal rows of stridulatory pegs in the inner side of hind femur." figureDoi="http://doi.org/10.5281/zenodo.3450290" httpUri="https://zenodo.org/record/3450290/files/figure.png" pageId="3" pageNumber="96">Fig. 6C</figureCitation>
). 
<emphasis id="B933EA8F3971B02BFE8DFCB9FE5AFCB3" box="[358,464,837,863]" italics="true" pageId="3" pageNumber="96">Abdomen</emphasis>
: Integument smooth. Abdominal tergites gradually narrowing to last tergite (
<figureCitation id="137C2A183971B02BFADDFCB9FA05FCB3" box="[1334,1423,837,863]" captionStart="FIGURE 3" captionStartId="4.[151,250,1199,1224]" captionTargetBox="[175,1414,205,1154]" captionTargetId="figure@4.[151,1437,181,1174]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3450280" httpUri="https://zenodo.org/record/3450280/files/figure.png" pageId="3" pageNumber="96">Fig. 3B</figureCitation>
). Sternites rectangular and wider than its length. Lateral margin of first and second abdominal tergites with patches of granular stridulatory pegs (
<figureCitation id="137C2A183971B02BFE2EFC71FD95FC44" box="[453,543,909,936]" captionStart="FIGURE 6" captionStartId="5.[151,250,1928,1953]" captionTargetBox="[270,1318,787,1886]" captionTargetId="figure@5.[248,1339,765,1903]" captionTargetPageId="5" captionText="FIGURE 6. Stridulatory apparatus of Glaphyrosoma stephanosoltis sp. nov. (A) male and (B) female, arrows pointing to patches of granular stridulatory pegs in the first and second abdominal tergites; (C) male and (D) female, arrow pointing to diagonal rows of stridulatory pegs in the inner side of hind femur." figureDoi="http://doi.org/10.5281/zenodo.3450290" httpUri="https://zenodo.org/record/3450290/files/figure.png" pageId="3" pageNumber="96">Fig. 6A</figureCitation>
). The ninth tergite broadly bilobed and divided in the middle with a broad notch along the postero-medial margin (
<figureCitation id="137C2A183971B02BFDF9FC4DFDFEFC27" box="[530,628,945,971]" captionStart="FIGURE 7" captionStartId="6.[151,250,1401,1426]" captionTargetBox="[186,1407,211,1350]" captionTargetId="figure@6.[156,1431,181,1376]" captionTargetPageId="6" captionText="FIGURE 7. Male external genitalia of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of abdominal segments 4–10; (B) posterior view of abdomen, showing cerci, paraproctal processes, and tenth tergites; (C) dorsal view of ninth and tenth abdominal tergites; (D) ventral view showing male subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450296" httpUri="https://zenodo.org/record/3450296/files/figure.png" pageId="3" pageNumber="96">Figs. 7C</figureCitation>
, 
<figureCitation id="137C2A183971B02BFD6BFC4DFD5FFC27" box="[640,725,945,971]" captionStart="FIGURE 9" captionStartId="8.[151,250,1415,1440]" captionTargetBox="[239,1394,334,1360]" captionTargetId="figure@8.[170,1418,273,1390]" captionTargetPageId="8" captionText="FIGURE 9. (A) posterior view of male external genitalia showing paraproctal processes; (B) ventral view of male subgenital plate; (C) lateral view of male ninth and tenth abdominal tergites; (D) lateral view of male paraproctal process; (E) ventral view of female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450300" httpUri="https://zenodo.org/record/3450300/files/figure.png" pageId="3" pageNumber="96">9A, 9B</figureCitation>
). The tenth tergite forming a pair of hooks in the middle, which turn upward above the middle of the ninth tergite (
<figureCitation id="137C2A183971B02BFD26FC29FCD4FC03" box="[717,862,981,1007]" captionStart="FIGURE 7" captionStartId="6.[151,250,1401,1426]" captionTargetBox="[186,1407,211,1350]" captionTargetId="figure@6.[156,1431,181,1376]" captionTargetPageId="6" captionText="FIGURE 7. Male external genitalia of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of abdominal segments 4–10; (B) posterior view of abdomen, showing cerci, paraproctal processes, and tenth tergites; (C) dorsal view of ninth and tenth abdominal tergites; (D) ventral view showing male subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450296" httpUri="https://zenodo.org/record/3450296/files/figure.png" pageId="3" pageNumber="96">Figs. 7B, 7C</figureCitation>
, 
<figureCitation id="137C2A183971B02BFC81FC29FC35FC03" box="[874,959,981,1007]" captionStart="FIGURE 9" captionStartId="8.[151,250,1415,1440]" captionTargetBox="[239,1394,334,1360]" captionTargetId="figure@8.[170,1418,273,1390]" captionTargetPageId="8" captionText="FIGURE 9. (A) posterior view of male external genitalia showing paraproctal processes; (B) ventral view of male subgenital plate; (C) lateral view of male ninth and tenth abdominal tergites; (D) lateral view of male paraproctal process; (E) ventral view of female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450300" httpUri="https://zenodo.org/record/3450300/files/figure.png" pageId="3" pageNumber="96">9A, 9C</figureCitation>
). Paraproct projected posteriorly but distal part still remains very close to last abdominal tergite and almost flat in appearance (
<figureCitation id="137C2A183971B02BFBABFC05FB28FBFF" box="[1088,1186,1017,1043]" captionStart="FIGURE 7" captionStartId="6.[151,250,1401,1426]" captionTargetBox="[186,1407,211,1350]" captionTargetId="figure@6.[156,1431,181,1376]" captionTargetPageId="6" captionText="FIGURE 7. Male external genitalia of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of abdominal segments 4–10; (B) posterior view of abdomen, showing cerci, paraproctal processes, and tenth tergites; (C) dorsal view of ninth and tenth abdominal tergites; (D) ventral view showing male subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450296" httpUri="https://zenodo.org/record/3450296/files/figure.png" pageId="3" pageNumber="96">Figs. 7B</figureCitation>
, 
<figureCitation id="137C2A183971B02BFB45FC05FB59FBFF" box="[1198,1235,1017,1043]" captionStart="FIGURE 9" captionStartId="8.[151,250,1415,1440]" captionTargetBox="[239,1394,334,1360]" captionTargetId="figure@8.[170,1418,273,1390]" captionTargetPageId="8" captionText="FIGURE 9. (A) posterior view of male external genitalia showing paraproctal processes; (B) ventral view of male subgenital plate; (C) lateral view of male ninth and tenth abdominal tergites; (D) lateral view of male paraproctal process; (E) ventral view of female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450300" httpUri="https://zenodo.org/record/3450300/files/figure.png" pageId="3" pageNumber="96">9A</figureCitation>
). Paraproctal process rectangular with a small denticle arising from medio-posteriorly lying almost flat and directed upwards (
<figureCitation id="137C2A183971B02BFA8FFBE1FF30FBB7" captionStart="FIGURE 7" captionStartId="6.[151,250,1401,1426]" captionTargetBox="[186,1407,211,1350]" captionTargetId="figure@6.[156,1431,181,1376]" captionTargetPageId="6" captionText="FIGURE 7. Male external genitalia of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of abdominal segments 4–10; (B) posterior view of abdomen, showing cerci, paraproctal processes, and tenth tergites; (C) dorsal view of ninth and tenth abdominal tergites; (D) ventral view showing male subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450296" httpUri="https://zenodo.org/record/3450296/files/figure.png" pageId="3" pageNumber="96">Figs. 7B</figureCitation>
, 
<figureCitation id="137C2A183971B02BFF2CFBBDFE97FBB7" box="[199,285,1089,1115]" captionStart="FIGURE 9" captionStartId="8.[151,250,1415,1440]" captionTargetBox="[239,1394,334,1360]" captionTargetId="figure@8.[170,1418,273,1390]" captionTargetPageId="8" captionText="FIGURE 9. (A) posterior view of male external genitalia showing paraproctal processes; (B) ventral view of male subgenital plate; (C) lateral view of male ninth and tenth abdominal tergites; (D) lateral view of male paraproctal process; (E) ventral view of female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450300" httpUri="https://zenodo.org/record/3450300/files/figure.png" pageId="3" pageNumber="96">9A, 9D</figureCitation>
). Subgenital plate with a prominent “V” shaped posteromedian notch and relatively long styli half the length of the subgenital plate (
<figureCitation id="137C2A183971B02BFE1BFB99FDDFFB93" box="[496,597,1125,1151]" captionStart="FIGURE 7" captionStartId="6.[151,250,1401,1426]" captionTargetBox="[186,1407,211,1350]" captionTargetId="figure@6.[156,1431,181,1376]" captionTargetPageId="6" captionText="FIGURE 7. Male external genitalia of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of abdominal segments 4–10; (B) posterior view of abdomen, showing cerci, paraproctal processes, and tenth tergites; (C) dorsal view of ninth and tenth abdominal tergites; (D) ventral view showing male subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450296" httpUri="https://zenodo.org/record/3450296/files/figure.png" pageId="3" pageNumber="96">Figs. 7D</figureCitation>
, 
<figureCitation id="137C2A183971B02BFD89FB99FD0CFB93" box="[610,646,1125,1151]" captionStart="FIGURE 9" captionStartId="8.[151,250,1415,1440]" captionTargetBox="[239,1394,334,1360]" captionTargetId="figure@8.[170,1418,273,1390]" captionTargetPageId="8" captionText="FIGURE 9. (A) posterior view of male external genitalia showing paraproctal processes; (B) ventral view of male subgenital plate; (C) lateral view of male ninth and tenth abdominal tergites; (D) lateral view of male paraproctal process; (E) ventral view of female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450300" httpUri="https://zenodo.org/record/3450300/files/figure.png" pageId="3" pageNumber="96">9B</figureCitation>
). Cerci almost twice as long as the width of the tenth tergite, tapering from the base to the tip, arched inwards in dorsal view, covered in small hairs (
<figureCitation id="137C2A183971B02BFBE1FB75FB38FB4F" box="[1034,1202,1161,1187]" captionStart="FIGURE 7" captionStartId="6.[151,250,1401,1426]" captionTargetBox="[186,1407,211,1350]" captionTargetId="figure@6.[156,1431,181,1376]" captionTargetPageId="6" captionText="FIGURE 7. Male external genitalia of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of abdominal segments 4–10; (B) posterior view of abdomen, showing cerci, paraproctal processes, and tenth tergites; (C) dorsal view of ninth and tenth abdominal tergites; (D) ventral view showing male subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450296" httpUri="https://zenodo.org/record/3450296/files/figure.png" pageId="3" pageNumber="96">Figs. 7A, C, D</figureCitation>
). Male internal genitalia largely membranous and as in 
<figureCitation id="137C2A183971B02BFDCEFB51FDE2FB2B" box="[549,616,1197,1223]" captionStart="FIGURE 8" captionStartId="7.[151,250,917,942]" captionTargetBox="[201,1386,220,856]" captionTargetId="figure@7.[166,1421,181,891]" captionTargetPageId="7" captionText="FIGURE 8. Male internal genitalia of Glaphyrosoma stephanosoltis sp. nov. (A) male internal genitalia in their natural position inside abdomen, showing their relative position to the subgenital plate; (B) dissected and magnified view of male internal genitalia." figureDoi="http://doi.org/10.5281/zenodo.3450298" httpUri="https://zenodo.org/record/3450298/files/figure.png" pageId="3" pageNumber="96">Fig. 8</figureCitation>
.
</paragraph>
<caption id="DF3866153971B02BFF7CF8EAFAA9F8C3" ID-DOI="http://doi.org/10.5281/zenodo.3450274" ID-Zenodo-Dep="3450274" box="[151,1315,1814,1839]" httpUri="https://zenodo.org/record/3450274/files/figure.png" pageId="3" pageNumber="96" startId="3.[151,250,1814,1839]" targetBox="[183,1403,1306,1763]" targetPageId="3">
<paragraph id="8BF8369D3971B02BFF7CF8EAFAA9F8C3" blockId="3.[151,1315,1814,1839]" box="[151,1315,1814,1839]" pageId="3" pageNumber="96">
<emphasis id="B933EA8F3971B02BFF7CF8EAFE99F8C3" bold="true" box="[151,275,1814,1839]" pageId="3" pageNumber="96">FIGURE 2.</emphasis>
(A) frontal view of head, showing the long maxillary palpi; (B) lateral view showing head and thorax.
</paragraph>
</caption>
<paragraph id="8BF8369D3971B02BFF2CF8A0FAE0F853" blockId="3.[151,1437,1884,1983]" pageId="3" pageNumber="96">
<emphasis id="B933EA8F3971B02BFF2CF8A0FEAFF89B" bold="true" box="[199,293,1884,1911]" italics="true" pageId="3" pageNumber="96">Female.</emphasis>
Similar to male (
<figureCitation id="137C2A183971B02BFE1BF8A1FDCCF89B" box="[496,582,1885,1911]" captionStart="FIGURE 10" captionStartId="9.[151,250,1225,1250]" captionTargetBox="[184,1406,205,1179]" captionTargetId="figure@9.[159,1429,181,1199]" captionTargetPageId="9" captionText="FIGURE 10. Female of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3450302" httpUri="https://zenodo.org/record/3450302/files/figure.png" pageId="3" pageNumber="96">Fig. 10</figureCitation>
). Subgenital plate triangular with a broad base and tapering toward the tip (
<figureCitation id="137C2A183971B02BFF4BF87DFF72F877" box="[160,248,1921,1947]" captionStart="FIGURE 9" captionStartId="8.[151,250,1415,1440]" captionTargetBox="[239,1394,334,1360]" captionTargetId="figure@8.[170,1418,273,1390]" captionTargetPageId="8" captionText="FIGURE 9. (A) posterior view of male external genitalia showing paraproctal processes; (B) ventral view of male subgenital plate; (C) lateral view of male ninth and tenth abdominal tergites; (D) lateral view of male paraproctal process; (E) ventral view of female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.3450300" httpUri="https://zenodo.org/record/3450300/files/figure.png" pageId="3" pageNumber="96">Fig. 9E</figureCitation>
). Ovipositor curved medially and apically directed (
<figureCitation id="137C2A183971B02BFCA2F87DFC13F877" box="[841,921,1921,1947]" captionStart="FIGURE 11" captionStartId="10.[151,250,1847,1872]" captionTargetBox="[203,1385,376,1778]" captionTargetId="figure@10.[162,1425,345,1822]" captionTargetPageId="10" captionText="FIGURE 11. Female ovipositor of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of ovipositor; (B) dorsal view of female abdominal terminalia showing cerci and ovipositor." figureDoi="http://doi.org/10.5281/zenodo.3450304" httpUri="https://zenodo.org/record/3450304/files/figure.png" pageId="3" pageNumber="96">Fig. 11</figureCitation>
) and almost one-third length of hind femora. Paraproctal process absent. Cerci thicker than male cerci at base with prominently tapering to the tip (
<figureCitation id="137C2A183971B02BFAE0F859FAD6F853" box="[1291,1372,1957,1983]" captionStart="FIGURE 11" captionStartId="10.[151,250,1847,1872]" captionTargetBox="[203,1385,376,1778]" captionTargetId="figure@10.[162,1425,345,1822]" captionTargetPageId="10" captionText="FIGURE 11. Female ovipositor of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view of ovipositor; (B) dorsal view of female abdominal terminalia showing cerci and ovipositor." figureDoi="http://doi.org/10.5281/zenodo.3450304" httpUri="https://zenodo.org/record/3450304/files/figure.png" pageId="3" pageNumber="96">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3866153976B02CFF7CFB53FBFFFB24" ID-DOI="http://doi.org/10.5281/zenodo.3450280" ID-Zenodo-Dep="3450280" box="[151,1141,1199,1224]" httpUri="https://zenodo.org/record/3450280/files/figure.png" pageId="4" pageNumber="97" startId="4.[151,250,1199,1224]" targetBox="[175,1414,205,1154]" targetPageId="4">
<paragraph id="8BF8369D3976B02CFF7CFB53FBFFFB24" blockId="4.[151,1141,1199,1224]" box="[151,1141,1199,1224]" pageId="4" pageNumber="97">
<emphasis id="B933EA8F3976B02CFF7CFB53FE99FB24" bold="true" box="[151,275,1199,1224]" pageId="4" pageNumber="97">FIGURE 3.</emphasis>
Holotype of 
<taxonomicName id="4C474D1E3976B02CFE72FB4CFD4CFB24" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[409,710,1200,1224]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="97" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">
<emphasis id="B933EA8F3976B02CFE72FB4CFD4CFB24" box="[409,710,1200,1224]" italics="true" pageId="4" pageNumber="97">Glaphyrosoma stephanosoltis</emphasis>
</taxonomicName>
<emphasis id="B933EA8F3976B02CFD26FB4CFC97FB24" bold="true" box="[717,797,1200,1224]" pageId="4" pageNumber="97">
<taxonomicNameLabel id="A20057F43976B02CFD26FB4CFC97FB24" box="[717,797,1200,1224]" pageId="4" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) lateral view; (B) dorsal view.
</paragraph>
</caption>
<caption id="DF3866153976B02CFF7CF846FF40F81B" ID-DOI="http://doi.org/10.5281/zenodo.3450284" ID-Zenodo-Dep="3450284" httpUri="https://zenodo.org/record/3450284/files/figure.png" pageId="4" pageNumber="97" startId="4.[151,250,1978,2003]" targetBox="[279,1281,1264,1949]" targetPageId="4">
<paragraph id="8BF8369D3976B02CFF7CF846FF40F81B" blockId="4.[151,1436,1978,2039]" pageId="4" pageNumber="97">
<emphasis id="B933EA8F3976B02CFF7CF846FE9FF83F" bold="true" box="[151,277,1978,2003]" pageId="4" pageNumber="97">FIGURE 4.</emphasis>
Legs of 
<taxonomicName id="4C474D1E3976B02CFE9DF846FD2FF83E" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[374,677,1978,2002]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="97" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">
<emphasis id="B933EA8F3976B02CFE9DF846FD2FF83E" box="[374,677,1978,2002]" italics="true" pageId="4" pageNumber="97">Glaphyrosoma stephanosoltis</emphasis>
</taxonomicName>
<emphasis id="B933EA8F3976B02CFD45F846FC8BF83E" bold="true" box="[686,769,1978,2002]" pageId="4" pageNumber="97">
<taxonomicNameLabel id="A20057F43976B02CFD45F846FC8BF83E" box="[686,769,1978,2002]" pageId="4" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) fore tibia to tarsi; (B) mid femur to tarsi; (C) Hind tibia to tarsi.
</paragraph>
</caption>
<caption id="DF3866153977B02DFF7CFD56FB03FD0B" ID-DOI="http://doi.org/10.5281/zenodo.3450286" ID-Zenodo-Dep="3450286" httpUri="https://zenodo.org/record/3450286/files/figure.png" pageId="5" pageNumber="98" startId="5.[151,250,682,707]" targetBox="[170,1418,201,637]" targetPageId="5">
<paragraph id="8BF8369D3977B02DFF7CFD56FB03FD0B" blockId="5.[151,1436,682,743]" pageId="5" pageNumber="98">
<emphasis id="B933EA8F3977B02DFF7CFD56FE99FD2F" bold="true" box="[151,275,682,707]" pageId="5" pageNumber="98">FIGURE 5.</emphasis>
Tibial tympana of 
<taxonomicName id="4C474D1E3977B02DFE38FD57FC8AFD2F" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[467,768,683,707]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="98" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">
<emphasis id="B933EA8F3977B02DFE38FD57FC8AFD2F" box="[467,768,683,707]" italics="true" pageId="5" pageNumber="98">Glaphyrosoma stephanosoltis</emphasis>
</taxonomicName>
<emphasis id="B933EA8F3977B02DFCEDFD57FCDCFD2F" bold="true" box="[774,854,683,707]" pageId="5" pageNumber="98">
<taxonomicNameLabel id="A20057F43977B02DFCEDFD57FCDCFD2F" box="[774,854,683,707]" pageId="5" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) male, showing outer tympanum in the right fore tibia and inner tympanum in the left fore tibia; (B) female outer tympanum; (C) female inner tympanum.
</paragraph>
</caption>
<caption id="DF3866153977B02DFF7CF874FD5EF805" ID-DOI="http://doi.org/10.5281/zenodo.3450290" ID-Zenodo-Dep="3450290" httpUri="https://zenodo.org/record/3450290/files/figure.png" pageId="5" pageNumber="98" startId="5.[151,250,1928,1953]" targetBox="[270,1318,787,1886]" targetPageId="5">
<paragraph id="8BF8369D3977B02DFF7CF874FD5EF805" blockId="5.[151,1436,1928,2025]" pageId="5" pageNumber="98">
<emphasis id="B933EA8F3977B02DFF7CF874FE98F84D" bold="true" box="[151,274,1928,1953]" pageId="5" pageNumber="98">FIGURE 6.</emphasis>
Stridulatory apparatus of 
<taxonomicName id="4C474D1E3977B02DFDFCF874FCC9F84C" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[535,835,1928,1952]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="98" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">
<emphasis id="B933EA8F3977B02DFDFCF874FCC9F84C" box="[535,835,1928,1952]" italics="true" pageId="5" pageNumber="98">Glaphyrosoma stephanosoltis</emphasis>
</taxonomicName>
<emphasis id="B933EA8F3977B02DFCA2F875FC12F84D" bold="true" box="[841,920,1929,1953]" pageId="5" pageNumber="98">
<taxonomicNameLabel id="A20057F43977B02DFCA2F875FC12F84D" box="[841,920,1929,1953]" pageId="5" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) male and (B) female, arrows pointing to patches of granular stridulatory pegs in the first and second abdominal tergites; (C) male and (D) female, arrow pointing to diagonal rows of stridulatory pegs in the inner side of hind femur.
</paragraph>
</caption>
<caption id="DF3866153974B02EFF7CFA85FC89FA36" ID-DOI="http://doi.org/10.5281/zenodo.3450296" ID-Zenodo-Dep="3450296" httpUri="https://zenodo.org/record/3450296/files/figure.png" pageId="6" pageNumber="99" startId="6.[151,250,1401,1426]" targetBox="[186,1407,211,1350]" targetPageId="6">
<paragraph id="8BF8369D3974B02EFF7CFA85FC89FA36" blockId="6.[151,1436,1401,1498]" pageId="6" pageNumber="99">
<emphasis id="B933EA8F3974B02EFF7CFA85FE98FA7E" bold="true" box="[151,274,1401,1426]" pageId="6" pageNumber="99">FIGURE 7.</emphasis>
Male external genitalia of 
<taxonomicName id="4C474D1E3974B02EFDCBFA86FCC6FA7E" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[544,844,1402,1426]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">
<emphasis id="B933EA8F3974B02EFDCBFA86FCC6FA7E" box="[544,844,1402,1426]" italics="true" pageId="6" pageNumber="99">Glaphyrosoma stephanosoltis</emphasis>
</taxonomicName>
<emphasis id="B933EA8F3974B02EFCB9FA86FC28FA7E" bold="true" box="[850,930,1402,1426]" pageId="6" pageNumber="99">
<taxonomicNameLabel id="A20057F43974B02EFCB9FA86FC28FA7E" box="[850,930,1402,1426]" pageId="6" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) lateral view of abdominal segments 4–10; (B) posterior view of abdomen, showing cerci, paraproctal processes, and tenth tergites; (C) dorsal view of ninth and tenth abdominal tergites; (D) ventral view showing male subgenital plate.
</paragraph>
</caption>
<subSubSection id="C35D65163974B02EFF2CF9F5FB4BF95F" pageId="6" pageNumber="99" type="description">
<paragraph id="8BF8369D3974B02EFF2CF9F5FB4BF95F" blockId="6.[151,1436,1544,2039]" pageId="6" pageNumber="99">
<emphasis id="B933EA8F3974B02EFF2CF9F5FD96F9CF" box="[199,540,1545,1571]" italics="true" pageId="6" pageNumber="99">
<emphasis id="B933EA8F3974B02EFF2CF9F5FE51F9CF" bold="true" box="[199,475,1545,1571]" italics="true" pageId="6" pageNumber="99">Measurements (in mm).</emphasis>
Male
</emphasis>
(n=5): pronotum length 8.94–9.42 (9.24 ± 0.22); fore femur length 11.89–12.62 (12.23 ± 0.29); mid femur length 12.09–13.63 (12.87 ± 0.59); hind femur length 29.36–31.31 (30.02 ± 0.89); hind femur width 6.71–6.83 (6.78 ± 0.06). 
<emphasis id="B933EA8F3974B02EFDA8F9ADFD1DF987" box="[579,663,1617,1643]" italics="true" pageId="6" pageNumber="99">Female</emphasis>
(n=5): pronotum length 9.00–10.24 (9.60 ± 0.48); fore femur length 11.33–13.55 (12.33 ± 0.84); mid femur length 11.96–13.65 (12.79 ± 0.67); hind femur length 28.53–31.73 (30.15 ± 1.29); hind femur width 6.50–7.60 (7.05 ± 0.40); ovipositor length 11.74–13.95 (12.56 ± 0.82).
</paragraph>
</subSubSection>
<subSubSection id="C35D65163974B02EFF2CF940FDECF8F2" pageId="6" pageNumber="99" type="etymology">
<paragraph id="8BF8369D3974B02EFF2CF940FDECF8F2" blockId="6.[151,1436,1544,2039]" pageId="6" pageNumber="99">
<emphasis id="B933EA8F3974B02EFF2CF940FEC2F93B" bold="true" box="[199,328,1724,1751]" italics="true" pageId="6" pageNumber="99">Etymology.</emphasis>
From Latin “stephano” meaning crown and “soltis” referring to the Soltis Center for Research and Education, the 
<typeStatus id="54FC883F3974B02EFEAAF91DFEFBF917" box="[321,369,1761,1787]" pageId="6" pageNumber="99">type</typeStatus>
locality of the species. Therefore, 
<taxonomicName id="4C474D1E3974B02EFD1CF91DFC1DF917" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[759,919,1761,1787]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="stephanosoltis">
<emphasis id="B933EA8F3974B02EFD1CF91DFC1DF917" box="[759,919,1761,1787]" italics="true" pageId="6" pageNumber="99">stephanosoltis</emphasis>
</taxonomicName>
means “Crown of Soltis” referring to the first king cricket ever described at the facility.
</paragraph>
</subSubSection>
<subSubSection id="C35D65163974B02EFF2CF8D4FE09F88A" pageId="6" pageNumber="99" type="distribution">
<paragraph id="8BF8369D3974B02EFF2CF8D4FE09F88A" blockId="6.[151,1436,1544,2039]" pageId="6" pageNumber="99">
<emphasis id="B933EA8F3974B02EFF2CF8D4FED0F8AF" bold="true" box="[199,346,1832,1859]" italics="true" pageId="6" pageNumber="99">Distribution.</emphasis>
<collectingCountry id="F350760D3974B02EFE8AF8D5FE55F8AE" box="[353,479,1832,1859]" name="Costa Rica" pageId="6" pageNumber="99">Costa Rica</collectingCountry>
: 
<collectingRegion id="4983F87F3974B02EFE0CF8D4FD38F8AE" box="[487,690,1832,1858]" country="Costa Rica" name="Alajuela" pageId="6" pageNumber="99">Alajuela Province</collectingRegion>
, San Ramón. Forest floor in mid-elevation (
<quantity id="4CBF9B783974B02EFB70F8D4FB69F8AF" box="[1179,1251,1832,1859]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="99" unit="m" value="450.0">450 m</quantity>
above sea level) secondary rainforest.
</paragraph>
</subSubSection>
<subSubSection id="C35D65163974B020FF2CF88CFD96FF17" lastPageId="8" lastPageNumber="101" pageId="6" pageNumber="99" type="materials_examined">
<paragraph id="8BF8369D3974B02EFF2CF88CFCC9F81A" blockId="6.[151,1436,1544,2039]" pageId="6" pageNumber="99">
<materialsCitation id="3B2F3CC03974B02EFF2CF88CFCC9F81A" ID-GBIF-Occurrence="2416712478" collectingDate="2018-08-07" collectorName="Coll. S. J. Richardson." country="Costa Rica" county="Male" latitude="10.383459" location="San Juan de Penas Blancas. Soltis Center" longLatPrecision="1" longitude="-84.617966" municipality="San Ramon" pageId="6" pageNumber="99" specimenCount="1" stateProvince="Alajuela" typeStatus="holotype">
<emphasis id="B933EA8F3974B02EFF2CF88CFEB3F867" bold="true" box="[199,313,1904,1931]" italics="true" pageId="6" pageNumber="99">
<typeStatus id="54FC883F3974B02EFF2CF88CFEBBF867" box="[199,305,1904,1931]" pageId="6" pageNumber="99" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCounty id="62994E113974B02EFEAAF88CFEF1F866" box="[321,379,1904,1930]" pageId="6" pageNumber="99">Male</collectingCounty>
(
<figureCitation id="137C2A183974B02EFE67F88CFE5BF867" box="[396,465,1904,1931]" captionStart="FIGURE 3" captionStartId="4.[151,250,1199,1224]" captionTargetBox="[175,1414,205,1154]" captionTargetId="figure@4.[151,1437,181,1174]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Glaphyrosoma stephanosoltis sp. nov. (A) lateral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3450280" httpUri="https://zenodo.org/record/3450280/files/figure.png" pageId="6" pageNumber="99">Fig. 3</figureCitation>
). (Measurement: pronotum length 
<quantity id="4CBF9B783974B02EFCB5F88DFC4DF867" box="[862,967,1905,1931]" metricMagnitude="-3" metricUnit="m" metricValue="9.34" pageId="6" pageNumber="99" unit="mm" value="9.34">9.34 mm</quantity>
; fore femur length 
<quantity id="4CBF9B783974B02EFB48F88DFA92F867" box="[1187,1304,1905,1931]" metricMagnitude="-2" metricUnit="m" metricValue="1.199" pageId="6" pageNumber="99" unit="mm" value="11.99">11.99 mm</quantity>
; mid femur length 
<quantity id="4CBF9B783974B02EFF08F869FED2F843" box="[227,344,1941,1967]" metricMagnitude="-2" metricUnit="m" metricValue="1.209" pageId="6" pageNumber="99" unit="mm" value="12.09">12.09 mm</quantity>
; hind femur length 
<quantity id="4CBF9B783974B02EFDDBF868FD2EF843" box="[560,676,1940,1967]" metricMagnitude="-2" metricUnit="m" metricValue="2.936" pageId="6" pageNumber="99" unit="mm" value="29.36">29.36 mm</quantity>
; hind femur width 
<quantity id="4CBF9B783974B02EFC9EF868FC57F843" box="[885,989,1940,1967]" metricMagnitude="-3" metricUnit="m" metricValue="6.81" pageId="6" pageNumber="99" unit="mm" value="6.81">6.81 mm</quantity>
.) 
<collectingCountry id="F350760D3974B02EFC04F869FB1EF843" box="[1007,1172,1941,1967]" name="Costa Rica" pageId="6" pageNumber="99">COSTA RICA</collectingCountry>
: 
<collectingRegion id="4983F87F3974B02EFB77F868FAEDF842" box="[1180,1383,1940,1966]" country="Costa Rica" name="Alajuela" pageId="6" pageNumber="99">Alajuela Province</collectingRegion>
, 
<collectingMunicipality id="6B9CACE73974B02EFA99F869FF61F83F" pageId="6" pageNumber="99">San Ramón</collectingMunicipality>
, 
<location id="8E9860463974B02EFF1CF845FD4DF83F" LSID="urn:lsid:plazi:treatment:03EE878B3970B020FF7CFDFDFD96FF17:8E9860463974B02EFF1CF845FD4DF83F" box="[247,711,1976,2003]" country="Costa Rica" county="Male" latitude="10.383459" longLatPrecision="1" longitude="-84.617966" municipality="San Ramon" name="San Juan de Penas Blancas. Soltis Center" pageId="6" pageNumber="99" stateProvince="Alajuela">San Juan de Peñas Blancas. Soltis Center</location>
for 
<location id="8E9860463974B02EFD1DF844FCD7F83E" LSID="urn:lsid:plazi:treatment:03EE878B3970B020FF7CFDFDFD96FF17:8E9860463974B02EFD1DF844FCD7F83E" box="[758,861,1976,2002]" country="Costa Rica" county="Male" latitude="10.383459" longLatPrecision="1" longitude="-84.617966" municipality="San Ramon" name="Research" pageId="6" pageNumber="99" stateProvince="Alajuela">Research</location>
and 
<location id="8E9860463974B02EFC7FF844FB82F83E" LSID="urn:lsid:plazi:treatment:03EE878B3970B020FF7CFDFDFD96FF17:8E9860463974B02EFC7FF844FB82F83E" box="[916,1032,1976,2002]" country="Costa Rica" county="Male" latitude="10.383459" longLatPrecision="1" longitude="-84.617966" municipality="San Ramon" name="Education" pageId="6" pageNumber="99" stateProvince="Alajuela">Education</location>
, 
<geoCoordinate id="EE73505A3974B02EFBFFF844FB5CF83F" box="[1044,1238,1976,2003]" degrees="10" direction="north" minutes="23" orientation="latitude" pageId="6" pageNumber="99" precision="1" seconds="0.4524" value="10.383459">10°23’0.4524’’N</geoCoordinate>
, 
<geoCoordinate id="EE73505A3974B02EFB0AF844FA12F83E" box="[1249,1432,1976,2002]" degrees="84" direction="west" minutes="37" orientation="longitude" pageId="6" pageNumber="99" precision="1" seconds="4.674" value="-84.617966">84°37’4.674’’W</geoCoordinate>
, 
<date id="FFF9105D3974B02EFF7CF820FE99F81A" box="[151,275,2012,2038]" pageId="6" pageNumber="99" value="2018-08-07">
<collectingDate id="EFBDE9B53974B02EFF7CF820FE99F81A" box="[151,275,2012,2038]" pageId="6" pageNumber="99" value="2018-08-07">7.viii.2018</collectingDate>
</date>
, collected by hand at night. 
<collectorName id="26B2534B3974B02EFDA5F820FCC9F81A" box="[590,835,2012,2039]" pageId="6" pageNumber="99">Coll. S.J. Richardson.</collectorName>
</materialsCitation>
</paragraph>
<caption id="DF3866153975B02FFF7CFC69FF7CFC1A" ID-DOI="http://doi.org/10.5281/zenodo.3450298" ID-Zenodo-Dep="3450298" httpUri="https://zenodo.org/record/3450298/files/figure.png" pageId="7" pageNumber="100" startId="7.[151,250,917,942]" targetBox="[201,1386,220,856]" targetPageId="7">
<paragraph id="8BF8369D3975B02FFF7CFC69FF7CFC1A" blockId="7.[151,1437,917,1014]" pageId="7" pageNumber="100">
<emphasis id="B933EA8F3975B02FFF7CFC69FE9EFC42" bold="true" box="[151,276,917,942]" pageId="7" pageNumber="100">FIGURE 8.</emphasis>
Male internal genitalia of 
<taxonomicName id="4C474D1E3975B02FFDCDFC6AFCDEFC42" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[550,852,918,942]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">
<emphasis id="B933EA8F3975B02FFDCDFC6AFCDEFC42" box="[550,852,918,942]" italics="true" pageId="7" pageNumber="100">Glaphyrosoma stephanosoltis</emphasis>
</taxonomicName>
<emphasis id="B933EA8F3975B02FFCB7FC6AFC27FC42" bold="true" box="[860,941,918,942]" pageId="7" pageNumber="100">
<taxonomicNameLabel id="A20057F43975B02FFCB7FC6AFC27FC42" box="[860,941,918,942]" pageId="7" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) male internal genitalia in their natural position inside abdomen, showing their relative position to the subgenital plate; (B) dissected and magnified view of male internal genitalia.
</paragraph>
</caption>
<paragraph id="8BF8369D3975B02FFF2CFBDFFE79FB8E" blockId="7.[151,1437,1059,2022]" pageId="7" pageNumber="100">
<emphasis id="B933EA8F3975B02FFF2CFBDFFE7CFBD2" bold="true" box="[199,502,1059,1086]" italics="true" pageId="7" pageNumber="100">Additional Type Material.</emphasis>
<specimenCount id="9D41FD143975B02FFDEBFBD8FD1BFBD2" box="[512,657,1060,1086]" pageId="7" pageNumber="100" type="generic" typeStatus="paratypes">26 paratypes</specimenCount>
(
<specimenCount id="9D41FD143975B02FFD4EFBDFFD7FFBD2" box="[677,757,1059,1086]" pageId="7" pageNumber="100" type="adult">5 adult</specimenCount>
males, 
<specimenCount id="9D41FD143975B02FFCB9FBD8FC3AFBD2" box="[850,944,1060,1086]" pageId="7" pageNumber="100" type="adult">13 adult</specimenCount>
females, 5 nymphal males, 3 nymphal females). Same data as 
<typeStatus id="54FC883F3975B02FFE62FBB4FE65FB8E" box="[393,495,1096,1122]" pageId="7" pageNumber="100" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BF8369D3975B02FFF2CFB90FF4EFB46" blockId="7.[151,1437,1059,2022]" pageId="7" pageNumber="100">
<emphasis id="B933EA8F3975B02FFF2CFB90FE0CFB6A" bold="true" box="[199,390,1132,1158]" italics="true" pageId="7" pageNumber="100">Type Depository.</emphasis>
All 
<typeStatus id="54FC883F3975B02FFE5CFB90FE6DFB6A" box="[439,487,1132,1158]" pageId="7" pageNumber="100">type</typeStatus>
material has been deposited to the 
<collectingRegion id="4983F87F3975B02FFC80FB90FC26FB6A" box="[875,940,1132,1158]" country="United States of America" name="Texas" pageId="7" pageNumber="100">Texas</collectingRegion>
A&amp;M University Insect Collection (TAMU- IC).
</paragraph>
<paragraph id="8BF8369D3975B02FFF2CFB48FE69FA4A" blockId="7.[151,1437,1059,2022]" pageId="7" pageNumber="100">
<emphasis id="B933EA8F3975B02FFF2CFB48FEE5FB22" bold="true" box="[199,367,1203,1230]" italics="true" pageId="7" pageNumber="100">DNA Barcode.</emphasis>
We have generated a DNA barcode for 
<taxonomicName id="4C474D1E3975B02FFCC4FB48FC7BFB22" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[815,1009,1204,1230]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">
<emphasis id="B933EA8F3975B02FFCC4FB48FC7BFB22" box="[815,1009,1204,1230]" italics="true" pageId="7" pageNumber="100">G. stephanosoltis</emphasis>
</taxonomicName>
<emphasis id="B933EA8F3975B02FFC13FB48FBD8FB22" bold="true" box="[1016,1106,1204,1230]" pageId="7" pageNumber="100">
<taxonomicNameLabel id="A20057F43975B02FFC13FB48FBD8FB22" box="[1016,1106,1204,1230]" pageId="7" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which has been deposited to GenBank with accession number 
<accessionNumber id="9414AB7E3975B02FFDE6FB24FD1AFB1E" box="[525,656,1240,1266]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN128722" pageId="7" pageNumber="100">
<collectionCode id="ED56AE583975B02FFDE6FB24FDB6FB1E" box="[525,572,1240,1266]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="7" pageNumber="100">MN</collectionCode>
128722
</accessionNumber>
, and the DNA tissue voucher specimen was deposited to 
<collectionCode id="ED56AE583975B02FFAE2FB24FAFFFB1E" box="[1289,1397,1240,1266]" country="USA" httpUri="http://grbio.org/cool/wntj-faij" name="Texas A&amp;amp;M University Insect Collection" pageId="7" pageNumber="100" type="University or college">TAMUIC</collectionCode>
Insect Genomic Collection with voucher number TAMUIC-IGC-002678. Although 
<bibRefCitation id="EFD64B6C3975B02FFBCBFB00FAA0FAFA" author="Vandergast, A. G. &amp; Weissman, D. B. &amp; Wood, D. A. &amp; Rentz, D. C. F. &amp; Bazelet, C. S. &amp; Ueshima, N." box="[1056,1322,1276,1302]" pageId="7" pageNumber="100" pagination="1 - 33" refId="ref5676" refString="Vandergast, A. G., Weissman, D. B., Wood, D. A., Rentz, D. C. F., Bazelet, C. S. &amp; Ueshima, N. (2017) Tackling an intractable problem: Can greater taxon sampling help resolve relationships within the Stenopelmatoidea (Orthoptera: Ensifera)? Zootaxa, 4291 (1), 1 - 33. https: // doi. org / 10.11646 / zootaxa. 4291.1.1" type="journal article" year="2017">
Vandergast 
<emphasis id="B933EA8F3975B02FFB49FB01FB53FAFA" box="[1186,1241,1276,1302]" italics="true" pageId="7" pageNumber="100">et al.</emphasis>
(2017)
</bibRefCitation>
generated COI genes for many 
<taxonomicName id="4C474D1E3975B02FFE6EFADCFDD8FAD6" authorityName="Saussure" authorityYear="1859" box="[389,594,1312,1338]" class="Insecta" family="Anostostomatidae" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="100" phylum="Arthropoda" rank="family">Anostostomatidae</taxonomicName>
, including several unidentified specimens of 
<taxonomicName id="4C474D1E3975B02FFBBFFADCFB73FAD6" authorityName="Brunner von Wattenwyl" authorityYear="1888" box="[1108,1273,1312,1338]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="B933EA8F3975B02FFBBFFADCFB73FAD6" box="[1108,1273,1312,1338]" italics="true" pageId="7" pageNumber="100">Glaphyrosoma</emphasis>
</taxonomicName>
from 
<collectingCountry id="F350760D3975B02FFAABFADCFA12FAD6" box="[1344,1432,1312,1338]" name="Mexico" pageId="7" pageNumber="100">Mexico</collectingCountry>
, the primers that they used (C1-J-2183 and C1-N-2872) amplified the back half of the COI gene and did not overlap with the DNA barcode region, which is located in the front half of the COI gene. Therefore, our DNA barcode represents the first for the genus.
</paragraph>
<paragraph id="8BF8369D3975B020FF2CFA53FD96FF17" blockId="7.[151,1437,1059,2022]" lastBlockId="8.[151,1437,153,251]" lastPageId="8" lastPageNumber="101" pageId="7" pageNumber="100">
<emphasis id="B933EA8F3975B02FFF2CFA53FCA7FA26" box="[199,813,1455,1482]" italics="true" pageId="7" pageNumber="100">
<emphasis id="B933EA8F3975B02FFF2CFA53FE53FA26" bold="true" box="[199,473,1455,1482]" italics="true" pageId="7" pageNumber="100">Biological Information.</emphasis>
<taxonomicName id="4C474D1E3975B02FFE0AFA4CFCA7FA26" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[481,813,1456,1482]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">Glaphyrosoma stephanosoltis</taxonomicName>
</emphasis>
<emphasis id="B933EA8F3975B02FFCDEFA4CFC05FA26" bold="true" box="[821,911,1456,1482]" pageId="7" pageNumber="100">
<taxonomicNameLabel id="A20057F43975B02FFCDEFA4CFC05FA26" box="[821,911,1456,1482]" pageId="7" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was hand collected along the trails adjacent to secondary rainforest in the vicinity of the Soltis Center for Research and Education. The Soltis Center’s forests are adjacent to the Children’s Eternal Rainforest (Bosque Eterno de los Niños) and located about 
<quantity id="4CBF9B783975B02FFB4AFA0BFB6DF9FE" box="[1185,1255,1527,1554]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="100" unit="m" value="450.0">450 m</quantity>
above sea level. The insects were only found after sunset when it was sprinkling or after it had just rained, and were more abundant when temperature was above 21 
<superScript id="7C329BD53975B02FFE14F9C3FD97F9B6" attach="right" box="[511,541,1599,1626]" fontSize="6" pageId="7" pageNumber="100">oC</superScript>
. The species was never observed during the day, and when kept in a screen cage with foliage comprised of torn leaves and branches, it was observed that the insects still hid under foliage, even in a dark environment. Those that were unable to hide under the foliage started to appear dull in their cuticles, which suggested that they desiccate easily even on the rainforest floor. The species most likely lives underground in order to avoid desiccation and remain in a higher level of humidity beneath the rainforest floor and only comes out when humidity levels are higher to search for food or mates. When collecting specimens along trails, it was found they tended to hide on the overhanging dirt underside along trails and sometimes in tunnels most likely made by other burrowing animals. When baited with oatmeal, the insects were attracted within an hour of placing the bait, which suggested that these insects might be targeting a good source of carbohydrate and protein, which are rich in the oatmeal used for the trail. When presented with a choice between a plant mixture (
<taxonomicName id="4C474D1E3975B02FFBE2F878FBF6F872" box="[1033,1148,1924,1950]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="100" phylum="Tracheophyta" rank="genus">
<emphasis id="B933EA8F3975B02FFBE2F878FBF6F872" box="[1033,1148,1924,1950]" italics="true" pageId="7" pageNumber="100">Passiflora</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="4C474D1E3975B02FFB5BF878FA1FF872" box="[1200,1429,1924,1950]" class="Magnoliopsida" family="Urticaceae" genus="Cecropia" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="100" phylum="Tracheophyta" rank="species" species="obtusifolia">
<emphasis id="B933EA8F3975B02FFB5BF878FA1FF872" box="[1200,1429,1924,1950]" italics="true" pageId="7" pageNumber="100">Cecropia obtusifolia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C474D1E3975B02FFF7CF854FEE0F82E" box="[151,362,1960,1986]" class="Magnoliopsida" family="Asteraceae" genus="Neurolaena" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="100" phylum="Tracheophyta" rank="species" species="lobata">
<emphasis id="B933EA8F3975B02FFF7CF854FEE0F82E" box="[151,362,1960,1986]" italics="true" pageId="7" pageNumber="100">Neurolaena lobata</emphasis>
</taxonomicName>
, and unidentified 
<taxonomicName id="4C474D1E3975B02FFDDDF854FD3BF82F" box="[566,689,1960,1987]" class="Magnoliopsida" family="Piperaceae" kingdom="Plantae" order="Piperales" pageId="7" pageNumber="100" phylum="Tracheophyta" rank="family">Piperaceae</taxonomicName>
) and a protein mixture (land crab and katydids), all of which were commonly found in the forest, 
<taxonomicName id="4C474D1E3975B02FFE06F830FD24F80A" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[493,686,1996,2022]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">
<emphasis id="B933EA8F3975B02FFE06F830FD24F80A" box="[493,686,1996,2022]" italics="true" pageId="7" pageNumber="100">G. stephanosoltis</emphasis>
</taxonomicName>
<emphasis id="B933EA8F3975B02FFD5FF830FC86F80A" bold="true" box="[692,780,1996,2022]" pageId="7" pageNumber="100">
<taxonomicNameLabel id="A20057F43975B02FFD5FF830FC86F80A" box="[692,780,1996,2022]" pageId="7" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
showed overwhelming preference for the plants. However, when kept in a cage with other insects, they fed on dead crickets and katydids, suggesting a necrophagous behavior. Collectively, these observations suggest that 
<taxonomicName id="4C474D1E397AB020FD62FF41FCC1FF3B" authority="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song, 2019" authorityName="Richardson &amp; Trimm &amp; Paredes &amp; Koehl &amp; Song" authorityYear="2019" box="[649,843,189,215]" class="Insecta" family="Chorotypidae" genus="Glaphyrosoma" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="101" phylum="Arthropoda" rank="species" species="stephanosoltis" status="sp. nov.">
<emphasis id="B933EA8F397AB020FD62FF41FCC1FF3B" box="[649,843,189,215]" italics="true" pageId="8" pageNumber="101">G. stephanosoltis</emphasis>
</taxonomicName>
<emphasis id="B933EA8F397AB020FCBAFF41FC20FF3B" bold="true" box="[849,938,189,215]" pageId="8" pageNumber="101">
<taxonomicNameLabel id="A20057F4397AB020FCBAFF41FC20FF3B" box="[849,938,189,215]" pageId="8" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is an omnivorous insect, which is a common dietary pattern in many ensiferans.
</paragraph>
</subSubSection>
</treatment>
</document>