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<document id="8E577E241097AF1856280EF4BAB6DDEE" ID-DOI="10.5281/zenodo.206017" ID-GBIF-Dataset="1d6d18c3-7ab0-4afc-b111-4a14a6811442" ID-ISSN="1175-5326" ID-Zenodo-Dep="206017" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460249045320" checkinUser="plazi" docAuthor="Deltshev, Christo, Ć, Božidar P. M. &amp; Ć, Ur Č I" docDate="2011" docId="0398A3050735D07559C6FA2AFE7EFA7C" docLanguage="en" docName="zt02782p038.pdf" docOrigin="Zootaxa 2782" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Harpactea complicata Deltshev, sp. nov." docType="treatment" docVersion="9" lastPageNumber="38" masterDocId="FFA1DB7D0735D0715951FF7CFF8BFFEE" masterDocTitle="A new spider species Harpactea complicata Deltshev sp. nov. from caves of Serbia (Araneae: Dysderidae)" masterLastPageNumber="38" masterPageNumber="34" pageNumber="34" updateTime="1698242381266" updateUser="plazi">
<mods:mods id="0650D1B75E43B91C2509AB82B684C809" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="837752FCCE1767D6B29BAEFDA84BBE5D">
<mods:title id="54A5AF08942F1901AAD34725DC0B5148">A new spider species Harpactea complicata Deltshev sp. nov. from caves of Serbia (Araneae: Dysderidae)</mods:title>
</mods:titleInfo>
<mods:name id="D5824CCE0F29550DA7F240815F7ECD0E" type="personal">
<mods:role id="4474EF788C35C5805632E7209C64C6F6">
<mods:roleTerm id="23BC10B47DC7B486AF715AC521612592">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08C700EE4E8B8F2209A7F3B673A8119F">Deltshev, Christo</mods:namePart>
</mods:name>
<mods:name id="4620CBE7C58DED96901209930302AB4C" type="personal">
<mods:role id="4D8C9820E3B5B7EF045969ED410255EC">
<mods:roleTerm id="0131B7FB3E0D16C7522710956B5FEA4C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6BEAEB9A1B6E02B094E33FC5D0D72637">Ć, Božidar P. M.</mods:namePart>
</mods:name>
<mods:name id="821362DC078C6EDFAC9AEAAA370B3DE2" type="personal">
<mods:role id="9D6EE6F620823E00643D80EF8F2F8A4C">
<mods:roleTerm id="600233921F416A4E78BC8E425691EDC8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8B02EF67FEA50C5485B34D0BD745AE43">Ć, Ur Č I</mods:namePart>
</mods:name>
<mods:typeOfResource id="2019E4C62FFB35FFF9A63FBBFC7D32F3">text</mods:typeOfResource>
<mods:relatedItem id="D4C0127D7F36E9D8B8F9206C95554989" type="host">
<mods:titleInfo id="7867EE0F243FD08D0435FAC9B025550D">
<mods:title id="6F98B83301F6B1358B43105D2F980F06">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CF6614FDBEE31ADB54943BDADA20518C">
<mods:date id="F5C8314036B720454484E3E7EB7BB769">2011</mods:date>
<mods:detail id="B4B89EF0BF0C81129E5A32849F71BD6F" type="volume">
<mods:number id="4729BAACDD001D96CDB3322243FD7698">2782</mods:number>
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<mods:extent id="503FDCAA54B01D042441A5F2206CD2A5" unit="page">
<mods:start id="164FCFA8EE9DBFE6FD29EF08E652ED46">34</mods:start>
<mods:end id="048B08ACEA8172EBE46B145B7043DA0E">38</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3F7DE2AADA2C5878F6150FC449ADE7A8">journal article</mods:classification>
<mods:identifier id="C194B1578A56299BA63FD674DA546326" type="DOI">10.5281/zenodo.206017</mods:identifier>
<mods:identifier id="0672D6CEB29A014B88002F9317427A53" type="GBIF-Dataset">1d6d18c3-7ab0-4afc-b111-4a14a6811442</mods:identifier>
<mods:identifier id="C4CE27C760FA5F0776DC4FF44A24FA58" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9801D27B78947040BB675E1ED77C4E3A" type="Zenodo-Dep">206017</mods:identifier>
</mods:mods>
<treatment id="0398A3050735D07559C6FA2AFE7EFA7C" ID-DOI="http://doi.org/10.5281/zenodo.6184902" ID-GBIF-Taxon="119391106" ID-Zenodo-Dep="6184902" LSID="urn:lsid:plazi:treatment:0398A3050735D07559C6FA2AFE7EFA7C" httpUri="http://treatment.plazi.org/id/0398A3050735D07559C6FA2AFE7EFA7C" lastPageId="4" lastPageNumber="38" pageId="0" pageNumber="34">
<subSubSection id="C32B41980735D07159C6FA2AFDD9FA3B" pageId="0" pageNumber="34" type="nomenclature">
<paragraph id="8B8E12130735D07159C6FA2AFDC6FA83" blockId="0.[151,589,1365,1423]" box="[151,589,1365,1390]" pageId="0" pageNumber="34">
<heading id="D0C6A57F0735D07159C6FA2AFDC6FA83" bold="true" box="[151,589,1365,1390]" fontSize="10" level="2" pageId="0" pageNumber="34" reason="2">
<emphasis id="B945CE010735D07159C6FA2AFDC6FA83" bold="true" box="[151,589,1365,1390]" pageId="0" pageNumber="34">
<taxonomicName id="4C3169900735D07159C6FA2AFE7AFA83" ID-CoL="3JRS8" authority="Deltshev" authorityName="Deltshev" box="[151,497,1365,1390]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="34" phylum="Arthropoda" rank="species" species="complicata" status="sp. nov.">
<emphasis id="B945CE010735D07159C6FA2AFE03FA80" bold="true" box="[151,392,1366,1390]" italics="true" pageId="0" pageNumber="34">Harpactea complicata</emphasis>
Deltshev
</taxonomicName>
<taxonomicNameLabel id="A276737A0735D07158A6FA29FDC6FA83" box="[503,589,1365,1389]" pageId="0" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E12130735D07159C6FA04FE8CFA61" blockId="0.[151,589,1365,1423]" box="[151,263,1400,1423]" pageId="0" pageNumber="34">
<figureCitation id="130A0E960735D07159C6FA04FE8CFA61" box="[151,263,1400,1423]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6, 7" captionStart-2="FIGURES 8 – 13" captionStart-3="FIGURES 14, 15" captionStartId-0="1.[151,269,1486,1510]" captionStartId-1="2.[151,269,1308,1332]" captionStartId-2="3.[151,270,1979,2003]" captionStartId-3="4.[151,269,704,728]" captionTargetBox-0="[216,1400,204,1450]" captionTargetBox-1="[152,1405,217,1266]" captionTargetBox-2="[271,1311,206,1943]" captionTargetBox-3="[151,1436,182,680]" captionTargetId-0="figure@1.[181,1405,166,1464]" captionTargetId-1="figure@2.[151,1436,194,1284]" captionTargetId-2="figure@3.[268,1319,194,1955]" captionTargetId-3="figure@4.[151,1436,182,680]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURES 1 – 5. Harpactea complicata sp. nov., male palp. 1: prolateral view; 2: retrolateral view; 3: detail of tip, anterior view; 4: detail of tip, posterior view; 5: detail of tip, ventral view. Scale lines: 0.3 mm." captionText-1="FIGURES 6, 7. Harpactea complicata sp. nov., epigyne and vulva. 6: ventral view; 7: vulva, dorsal view. Scale lines: 0.3 mm." captionText-2="FIGURES 8 – 13. Harpactea complicata sp. nov. 8 – 9: male palp, prolateral view; 10: male palp, retrolateral view; 11: detail of tip of bulb, anterior view; 12: detail of tip of bulb, ventral view; 13: eyes. Scale lines: 0.2 mm." captionText-3="FIGURES 14, 15. Harpactea complicata sp. nov., epigyne and vulva. 14: ventral view; 15: vulva, dorsal view. Scale lines: 0.2 mm." httpUri-0="https://zenodo.org/record/206018/files/figure.png" httpUri-1="https://zenodo.org/record/206019/files/figure.png" httpUri-2="https://zenodo.org/record/206020/files/figure.png" httpUri-3="https://zenodo.org/record/206021/files/figure.png" pageId="0" pageNumber="34">Figs. 1–15</figureCitation>
</paragraph>
<paragraph id="8B8E12130735D07159C6FAC2FDD9FA3B" blockId="0.[151,594,1464,1493]" box="[151,594,1464,1493]" pageId="0" pageNumber="34">
<taxonomicName id="4C3169900735D07159C6FAC2FE82FA3B" box="[151,265,1470,1493]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="34" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B945CE010735D07159C6FAC2FE82FA3B" box="[151,265,1470,1493]" italics="true" pageId="0" pageNumber="34">Harpactea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276737A0735D071585EFAC2FEC7FA3B" box="[271,332,1470,1493]" pageId="0" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
: Ć urċiċ
<emphasis id="B945CE010735D07158F5FAC2FE51FA3B" box="[420,474,1470,1493]" italics="true" pageId="0" pageNumber="34">et al.</emphasis>
2007:
<specimenCount id="9D37D99A0735D0715B72FAC2FDD9FA3B" box="[547,594,1470,1493]" pageId="0" pageNumber="34" type="generic">19P.</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C32B41980735D07159C6F978FA94F918" pageId="0" pageNumber="34" type="materials_examined">
<paragraph id="8B8E12130735D07159C6F978FE86F98B" blockId="0.[151,1436,1539,2026]" pageId="0" pageNumber="34">
<emphasis id="B945CE010735D07159C6F978FEB6F9F2" bold="true" box="[151,317,1540,1564]" pageId="0" pageNumber="34">
<typeStatus id="548AACB10735D07159C6F978FF44F9F2" box="[151,207,1540,1564]" pageId="0" pageNumber="34">Type</typeStatus>
material.
</emphasis>
SW
<collectingCountry id="F32652830735D0715822F979FE31F9F2" box="[371,442,1541,1564]" name="Serbia" pageId="0" pageNumber="34">Serbia</collectingCountry>
, Đurđeva peċina Cave, Erċege, Ivanjica, 1 ɗ
<typeStatus id="548AACB10735D0715AF5F979FB8AF9F2" box="[932,1025,1541,1564]" pageId="0" pageNumber="34" type="holotype">holotype</typeStatus>
and 1 Ψ
<typeStatus id="548AACB10735D0715D35F979FB49F9F2" box="[1124,1218,1541,1564]" pageId="0" pageNumber="34" type="paratype">paratype</typeStatus>
,
<date id="FF8F34D30735D0715D9FF979FAC2F9F2" box="[1230,1353,1541,1564]" pageId="0" pageNumber="34" value="2004-04-30">30.04.2004</date>
; Peċina Potkapilijama Cave, Trudovo, Ivanjica, 1.05.2003, 1 ɗ and 3 Ψ
<typeStatus id="548AACB10735D0715A15F956FC21F9AF" box="[836,938,1578,1601]" pageId="0" pageNumber="34" type="paratype">paratypes</typeStatus>
<date id="FF8F34D30735D0715AE1F956FBA3F9AF" box="[944,1064,1578,1601]" pageId="0" pageNumber="34" value="1988-06-14">14.06.1988</date>
(leg. B. P. M. Ć urċiċ, V. Tomiċ, S. B. Ć urċiċ).
</paragraph>
<paragraph id="8B8E12130735D0715996F90EFA94F918" blockId="0.[151,1436,1539,2026]" pageId="0" pageNumber="34">
Material used for comparison.
<taxonomicName id="4C3169900735D0715B46F90EFCE4F967" authority="C. L. Koch" authorityName="C. L. Koch" box="[535,879,1650,1673]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="34" phylum="Arthropoda" rank="species" species="lepida">
<emphasis id="B945CE010735D0715B46F90EFD5FF967" box="[535,724,1650,1673]" italics="true" pageId="0" pageNumber="34">Harpactea lepida</emphasis>
(C. L. Koch)
</taxonomicName>
: SW
<collectingCountry id="F32652830735D0715AFAF90EFB84F967" box="[939,1039,1650,1673]" name="Germany" pageId="0" pageNumber="34">Germany</collectingCountry>
, Schwarzwald, Posthalde, 1 ɗ, 1 Ψ,
<date id="FF8F34D30735D07159C6F9EBFE84F940" box="[151,271,1687,1710]" pageId="0" pageNumber="34" value="1904-08-29">29.08.1904</date>
(leg. F. Dahl) (N 928
<collectionCode id="ED208AD60735D07158AEF9EBFDB6F940" box="[511,573,1687,1710]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="0" pageNumber="34">ZMB</collectionCode>
);
<collectingCountry id="F32652830735D0715B01F9EBFD1CF940" box="[592,663,1687,1710]" name="Serbia" pageId="0" pageNumber="34">Serbia</collectingCountry>
, Lovaċka ċesma, Mt. Javor, 1 ɗ, 2 Ψ,
<date id="FF8F34D30735D0715D67F9EBFB2FF940" box="[1078,1188,1687,1710]" pageId="0" pageNumber="34" value="2003-05-29">29.5.2003</date>
; Near Ršumova peċina Cave (leaf litter), Ljubiš, Mt. Zlatibor,
<date id="FF8F34D30735D0715B6CF9C7FD16F93C" box="[573,669,1723,1746]" pageId="0" pageNumber="34" value="2004-05-02">2.5.2004</date>
, (leg. B. P. M. Ć urċiċ, V. Tomiċ, S. B. Ć urċiċ);
<taxonomicName id="4C3169900735D0715DF9F9C7FA17F93C" box="[1192,1436,1723,1746]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="34" phylum="Arthropoda" rank="species" species="srednagora">
<emphasis id="B945CE010735D0715DF9F9C7FA17F93C" box="[1192,1436,1723,1746]" italics="true" pageId="0" pageNumber="34">Harpactea srednagora</emphasis>
</taxonomicName>
Dimitrov &amp; Lazarov:
<collectingCountry id="F32652830735D07158D1F9A3FE54F918" box="[384,479,1759,1782]" name="Bulgaria" pageId="0" pageNumber="34">Bulgaria</collectingCountry>
, Sredna Gora Mountain, Panagjuriste, 1 ɗ, 1 Ψ,
<date id="FF8F34D30735D0715AB4F9A3FBD6F918" box="[997,1117,1759,1782]" pageId="0" pageNumber="34" value="2006-05-29">29.05.2006</date>
(leg. S. Lazarov).
</paragraph>
</subSubSection>
<subSubSection id="C32B41980735D0715996F87FFA10F8F4" box="[199,1435,1795,1819]" pageId="0" pageNumber="34" type="etymology">
<paragraph id="8B8E12130735D0715996F87FFA10F8F4" blockId="0.[151,1436,1539,2026]" box="[199,1435,1795,1819]" pageId="0" pageNumber="34">
<emphasis id="B945CE010735D0715996F87FFECCF8F5" bold="true" box="[199,327,1795,1819]" pageId="0" pageNumber="34">Etymology.</emphasis>
The specific name refers to the complicated structure of the bulb (latin
<emphasis id="B945CE010735D0715D15F87FFB4EF8F4" box="[1092,1221,1795,1818]" italics="true" pageId="0" pageNumber="34">complicatus</emphasis>
means
<emphasis id="B945CE010735D0715C47F87FFA00F8F4" box="[1302,1419,1795,1818]" italics="true" pageId="0" pageNumber="34">complicate</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B41980735D0715996F859FC55F804" pageId="0" pageNumber="34" type="diagnosis">
<paragraph id="8B8E12130735D0715996F859FC55F804" blockId="0.[151,1436,1539,2026]" pageId="0" pageNumber="34">
<emphasis id="B945CE010735D0715996F859FEB7F8D3" bold="true" box="[199,316,1829,1853]" pageId="0" pageNumber="34">Diagnosis.</emphasis>
Somatic characters of the new species corresponding to those of the genus. The new species fits well to the
<taxonomicName id="4C3169900735D07159EFF835FE8AF88E" box="[190,257,1865,1888]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="34" phylum="Arthropoda" rank="species" species="lepida">
<emphasis id="B945CE010735D07159EFF835FE8AF88E" box="[190,257,1865,1888]" italics="true" pageId="0" pageNumber="34">lepida</emphasis>
</taxonomicName>
species group, but the genitalia differ markedly from all other known species of the genus. Morphologically it resembles most to the species
<taxonomicName id="4C3169900735D07158BFF817FD57F86C" authority="Brignoli" authorityName="Brignoli" box="[494,732,1899,1922]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="34" phylum="Arthropoda" rank="species" species="apollinea">
<emphasis id="B945CE010735D07158BFF817FDF3F86C" box="[494,632,1899,1922]" italics="true" pageId="0" pageNumber="34">H. apollinea</emphasis>
Brignoli
</taxonomicName>
,
<taxonomicName id="4C3169900735D0715BBCF817FC6AF86C" authority="Canestrini" authorityName="Canestrini" box="[749,993,1899,1922]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="34" phylum="Arthropoda" rank="species" species="grisea">
<emphasis id="B945CE010735D0715BBCF817FCDEF86C" box="[749,853,1899,1922]" italics="true" pageId="0" pageNumber="34">H. grisea</emphasis>
(Canestrini)
</taxonomicName>
,
<taxonomicName id="4C3169900735D0715AA1F817FB78F86C" authority="C. L. Koch" authorityName="C. L. Koch" box="[1008,1267,1899,1922]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="34" phylum="Arthropoda" rank="species" species="lepida">
<emphasis id="B945CE010735D0715AA1F817FBD3F86C" box="[1008,1112,1899,1922]" italics="true" pageId="0" pageNumber="34">H. lepida</emphasis>
(C. L. Koch)
</taxonomicName>
and
<taxonomicName id="4C3169900735D0715C61F817FF6CF84B" authority="Alicata" authorityName="Alicata" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="34" phylum="Arthropoda" rank="species" species="thaleri">
<emphasis id="B945CE010735D0715C61F817FA17F86C" box="[1328,1436,1899,1922]" italics="true" pageId="0" pageNumber="34">H. thaleri</emphasis>
Alicata
</taxonomicName>
, but it is easily distinguished from them by the long and slender embolus, the broader lamellar part of the conductor with a lanceolate projection, and an accessory apophysis with a characteristic triangular shape. The female differs mainly in the shape of the spermatheca that is globular at its upper part.
</paragraph>
</subSubSection>
<caption id="DF4E429B0734D07059C6FAB2FB00F9E7" httpUri="https://zenodo.org/record/206018/files/figure.png" pageId="1" pageNumber="35" targetBox="[216,1400,204,1450]" targetPageId="1">
<paragraph id="8B8E12130734D07059C6FAB2FB00F9E7" blockId="1.[151,1436,1486,1545]" pageId="1" pageNumber="35">
<emphasis id="B945CE010734D07059C6FAB2FECDFA08" bold="true" box="[151,326,1486,1510]" pageId="1" pageNumber="35">FIGURES 1–5.</emphasis>
<taxonomicName id="4C3169900734D0705800FAB3FDCFFA08" box="[337,580,1487,1510]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="35" phylum="Arthropoda" rank="species" species="complicata" status="sp. nov.">
<emphasis id="B945CE010734D0705800FAB3FDCFFA08" box="[337,580,1487,1510]" italics="true" pageId="1" pageNumber="35">Harpactea complicata</emphasis>
</taxonomicName>
<emphasis id="B945CE010734D0705B1EFAB2FD22FA08" bold="true" box="[591,681,1486,1510]" pageId="1" pageNumber="35">
<taxonomicNameLabel id="A276737A0734D0705B1EFAB2FD22FA08" box="[591,681,1486,1510]" pageId="1" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male palp. 1: prolateral view; 2: retrolateral view; 3: detail of tip, anterior view; 4: detail of tip, posterior view; 5: detail of tip, ventral view. Scale lines: 0.3 mm.
</paragraph>
</caption>
<subSubSection id="C32B41980734D0755996F944FB48FC49" lastPageId="4" lastPageNumber="38" pageId="1" pageNumber="35" type="description">
<paragraph id="8B8E12130734D0705996F944FE21F99D" blockId="1.[151,1437,1590,2033]" pageId="1" pageNumber="35">
<emphasis id="B945CE010734D0705996F944FED9F9BE" bold="true" box="[199,338,1592,1616]" pageId="1" pageNumber="35">Description.</emphasis>
ɗ
<typeStatus id="548AACB10734D0705829F945FE5EF9BE" box="[376,469,1593,1616]" pageId="1" pageNumber="35" type="holotype">holotype</typeStatus>
: Total length 5.25; prosoma, length 3.00, width 2.25; sternum length 2.02, width 1.35; opisthosoma, length 3.37.
</paragraph>
<paragraph id="8B8E12130734D0705996F903FD1AF935" blockId="1.[151,1437,1590,2033]" pageId="1" pageNumber="35">
Carapace with smooth surface, yellow to yellow brown on the cephalic part; hairs sparse. AME larger than the other eyes, distance between AME equal to their diameter (
<figureCitation id="130A0E960734D0705BB7F9DEFCB9F957" box="[742,818,1698,1721]" captionStart="FIGURES 8 – 13" captionStartId="3.[151,270,1979,2003]" captionTargetBox="[271,1311,206,1943]" captionTargetId="figure@3.[268,1319,194,1955]" captionTargetPageId="3" captionText="FIGURES 8 – 13. Harpactea complicata sp. nov. 8 – 9: male palp, prolateral view; 10: male palp, retrolateral view; 11: detail of tip of bulb, anterior view; 12: detail of tip of bulb, ventral view; 13: eyes. Scale lines: 0.2 mm." httpUri="https://zenodo.org/record/206020/files/figure.png" pageId="1" pageNumber="35">Fig 13</figureCitation>
). Labium length two times its width, with hairy whitish upper part. Sternum with hairs in the periphery.
</paragraph>
<paragraph id="8B8E12130734D0705996F99BFB67F8CF" blockId="1.[151,1437,1590,2033]" pageId="1" pageNumber="35">Chelicerae light brown. Retromargin with one tiny tooth at the base of the groove and a larger one on its middle part. Promargin with two teeth of equal size (larger than those on retromargin), which are close to each other.</paragraph>
<paragraph id="8B8E12130734D0705996F850FE20F888" blockId="1.[151,1437,1590,2033]" pageId="1" pageNumber="35">Abdomen white to light grey, elongate and slender. Legs yellow to yellow-brown. Leg spination and measurements are given in table 1 and 2.</paragraph>
<paragraph id="8B8E12130734D0705996F80EFF43F81F" blockId="1.[151,1437,1590,2033]" pageId="1" pageNumber="35">
Male palp (
<figureCitation id="130A0E960734D0705810F80EFE2AF867" box="[321,417,1906,1929]" captionStart="FIGURES 1 – 5" captionStartId="1.[151,269,1486,1510]" captionTargetBox="[216,1400,204,1450]" captionTargetId="figure@1.[181,1405,166,1464]" captionTargetPageId="1" captionText="FIGURES 1 – 5. Harpactea complicata sp. nov., male palp. 1: prolateral view; 2: retrolateral view; 3: detail of tip, anterior view; 4: detail of tip, posterior view; 5: detail of tip, ventral view. Scale lines: 0.3 mm." httpUri="https://zenodo.org/record/206018/files/figure.png" pageId="1" pageNumber="35">Figs 1–5</figureCitation>
,
<figureCitation id="130A0E960734D07058FCF80EFE6DF867" box="[429,486,1906,1929]" captionStart="FIGURES 8 – 13" captionStartId="3.[151,270,1979,2003]" captionTargetBox="[271,1311,206,1943]" captionTargetId="figure@3.[268,1319,194,1955]" captionTargetPageId="3" captionText="FIGURES 8 – 13. Harpactea complicata sp. nov. 8 – 9: male palp, prolateral view; 10: male palp, retrolateral view; 11: detail of tip of bulb, anterior view; 12: detail of tip of bulb, ventral view; 13: eyes. Scale lines: 0.2 mm." httpUri="https://zenodo.org/record/206020/files/figure.png" pageId="1" pageNumber="35">8–12</figureCitation>
): Tibia longer than tarsus, straight. Tarsus triangular in lateral view. All segments setose. Tegulum pear-shaped with complex embolar division. Embolus transverse, long and slender. Conductor bifurcated with lamella shaped like large plate and delicate lanceolate projection. Triangular accessory apophysis curved in its apical part.
</paragraph>
<caption id="DF4E429B0737D07359C6FA60FF7BFAB8" httpUri="https://zenodo.org/record/206019/files/figure.png" pageId="2" pageNumber="36" targetBox="[152,1405,217,1266]" targetPageId="2">
<paragraph id="8B8E12130737D07359C6FA60FF7BFAB8" blockId="2.[151,1436,1308,1366]" pageId="2" pageNumber="36">
<emphasis id="B945CE010737D07359C6FA60FECEFADA" bold="true" box="[151,325,1308,1332]" pageId="2" pageNumber="36">FIGURES 6, 7.</emphasis>
<taxonomicName id="4C3169900737D073581CFA60FDB5FADD" box="[333,574,1308,1331]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="36" phylum="Arthropoda" rank="species" species="complicata" status="sp. nov.">
<emphasis id="B945CE010737D073581CFA60FDB5FADD" box="[333,574,1308,1331]" italics="true" pageId="2" pageNumber="36">Harpactea complicata</emphasis>
</taxonomicName>
<emphasis id="B945CE010737D0735B14FA60FD17FADA" bold="true" box="[581,668,1308,1332]" pageId="2" pageNumber="36">
<taxonomicNameLabel id="A276737A0737D0735B14FA60FD17FADA" box="[581,668,1308,1332]" pageId="2" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, epigyne and vulva. 6: ventral view; 7: vulva, dorsal view. Scale lines: 0.3 mm.
</paragraph>
</caption>
<paragraph id="8B8E12130737D07359C6FAFAFCE9FA73" blockId="2.[151,1209,1412,1702]" box="[151,866,1412,1438]" pageId="2" pageNumber="36">
<emphasis id="B945CE010737D07359C6FAFAFE83FA70" bold="true" box="[151,264,1414,1438]" pageId="2" pageNumber="36">TABLE 1.</emphasis>
<taxonomicName id="4C3169900737D0735841FAFAFE74FA73" box="[272,511,1414,1437]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="36" phylum="Arthropoda" rank="species" species="complicata" status="sp. nov.">
<emphasis id="B945CE010737D0735841FAFAFE74FA73" box="[272,511,1414,1437]" italics="true" pageId="2" pageNumber="36">Harpactea complicata</emphasis>
</taxonomicName>
<emphasis id="B945CE010737D0735B57FAFAFDE9FA70" bold="true" box="[518,610,1414,1438]" pageId="2" pageNumber="36">
<taxonomicNameLabel id="A276737A0737D0735B57FAFAFDD6FA70" box="[518,605,1414,1438]" pageId="2" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
spination (ɗ
<typeStatus id="548AACB10737D0735BA5FAFAFCDDFA73" box="[756,854,1414,1437]" pageId="2" pageNumber="36" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8B8E12130737D07359CEFAC6FC4BF8EB" blockId="2.[151,1209,1412,1702]" lastBlockId="2.[151,1327,1772,2019]" pageId="2" pageNumber="36">
Legs I II III IV Fe 5 p 4 p 8 d 12 d Pa 0 0 0 0 Ti 0 0 6 p,
<date id="FF8F34D30737D0735A72F946FCCEF9BF" box="[803,837,1594,1617]" pageId="2" pageNumber="36">3 v</date>
, 5 r 6p,
<date id="FF8F34D30737D0735D27F946FB19F9BF" box="[1142,1170,1594,1617]" pageId="2" pageNumber="36">3v</date>
, 5r Me 0 0 6 p,
<date id="FF8F34D30737D0735A72F918FCCEF995" box="[803,837,1636,1659]" pageId="2" pageNumber="36">2 v</date>
, 3 r 7p,
<date id="FF8F34D30737D0735D27F918FB19F995" box="[1142,1170,1636,1659]" pageId="2" pageNumber="36">3v</date>
, 3 r Ta 0 0 0 0
<emphasis id="B945CE010737D07359C6F992FE83F8E8" bold="true" box="[151,264,1774,1798]" pageId="2" pageNumber="36">TABLE 2.</emphasis>
<taxonomicName id="4C3169900737D0735841F992FE74F8EB" box="[272,511,1774,1797]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="36" phylum="Arthropoda" rank="species" species="complicata" status="sp. nov.">
<emphasis id="B945CE010737D0735841F992FE74F8EB" box="[272,511,1774,1797]" italics="true" pageId="2" pageNumber="36">Harpactea complicata</emphasis>
</taxonomicName>
<emphasis id="B945CE010737D0735B57F992FDD0F8E8" bold="true" box="[518,603,1774,1798]" pageId="2" pageNumber="36">
<taxonomicNameLabel id="A276737A0737D0735B57F992FDD0F8E8" box="[518,603,1774,1798]" pageId="2" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, leg measurements (ɗ
<typeStatus id="548AACB10737D0735A03F992FC3FF8EB" box="[850,948,1774,1797]" pageId="2" pageNumber="36" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8B8E12130737D07559CEF85EFB8CFC8C" blockId="2.[151,1327,1772,2019]" lastBlockId="4.[151,1436,808,971]" lastPageId="4" lastPageNumber="38" pageId="2" pageNumber="36">Legs Femur Patella Tibia Metatarsus Tarsus Total I 2.85 1.87 2.40 2.40 0.75 10.27 II 2.77 1.35 2.02 2.10 0.67 8.91 III 2.25 1.12 1.57 1.95 0.60 7.49 IV 2.85 1.35 2.40 2.55 0.75 9.90 Female: Total length 3.96; prosoma, length 1.62, width 1.01; sternum length 0.86, width 0.72; opisthosoma, length 2.70. All characters as described for male. Leg measurements are given in Table 3.</paragraph>
<caption id="DF4E429B0736D07259C6F8C7FB68F81B" httpUri="https://zenodo.org/record/206020/files/figure.png" pageId="3" pageNumber="37" targetBox="[271,1311,206,1943]" targetPageId="3">
<paragraph id="8B8E12130736D07259C6F8C7FB68F81B" blockId="3.[151,1436,1979,2037]" pageId="3" pageNumber="37">
<emphasis id="B945CE010736D07259C6F8C7FED9F83D" bold="true" box="[151,338,1979,2003]" pageId="3" pageNumber="37">FIGURES 8–13.</emphasis>
<taxonomicName id="4C3169900736D072580BF8C0FDC1F83D" box="[346,586,1980,2003]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="37" phylum="Arthropoda" rank="species" species="complicata" status="sp. nov.">
<emphasis id="B945CE010736D072580BF8C0FDC1F83D" box="[346,586,1980,2003]" italics="true" pageId="3" pageNumber="37">Harpactea complicata</emphasis>
</taxonomicName>
<emphasis id="B945CE010736D0725B02F8C7FD21F83D" bold="true" box="[595,682,1979,2003]" pageId="3" pageNumber="37">
<taxonomicNameLabel id="A276737A0736D0725B02F8C7FD21F83D" box="[595,682,1979,2003]" pageId="3" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
8–9: male palp, prolateral view; 10: male palp, retrolateral view; 11: detail of tip of bulb, anterior view; 12: detail of tip of bulb, ventral view; 13: eyes. Scale lines: 0.2 mm.
</paragraph>
</caption>
<caption id="DF4E429B0731D07559C6FDBCFEBBFD14" httpUri="https://zenodo.org/record/206021/files/figure.png" pageId="4" pageNumber="38" targetBox="[151,1436,182,680]" targetPageId="4">
<paragraph id="8B8E12130731D07559C6FDBCFEBBFD14" blockId="4.[151,1436,704,762]" pageId="4" pageNumber="38">
<emphasis id="B945CE010731D07559C6FDBCFEEAFD36" bold="true" box="[151,353,704,728]" pageId="4" pageNumber="38">FIGURES 14, 15.</emphasis>
<taxonomicName id="4C3169900731D075583BFDBCFDD1FD39" box="[362,602,704,727]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="complicata" status="sp. nov.">
<emphasis id="B945CE010731D075583BFDBCFDD1FD39" box="[362,602,704,727]" italics="true" pageId="4" pageNumber="38">Harpactea complicata</emphasis>
</taxonomicName>
<emphasis id="B945CE010731D0755B35FDBCFD4AFD36" bold="true" box="[612,705,704,728]" pageId="4" pageNumber="38">
<taxonomicNameLabel id="A276737A0731D0755B35FDBCFD37FD36" box="[612,700,704,728]" pageId="4" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
epigyne and vulva. 14: ventral view; 15: vulva, dorsal view. Scale lines: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8E12130731D0755996FC11FB48FC49" blockId="4.[151,1436,808,971]" pageId="4" pageNumber="38">
Epigyne and vulva (
<figureCitation id="130A0E960731D07558F5FC11FD86FC6A" box="[420,525,877,900]" captionStart="FIGURES 6, 7" captionStartId="2.[151,269,1308,1332]" captionTargetBox="[152,1405,217,1266]" captionTargetId="figure@2.[151,1436,194,1284]" captionTargetPageId="2" captionText="FIGURES 6, 7. Harpactea complicata sp. nov., epigyne and vulva. 6: ventral view; 7: vulva, dorsal view. Scale lines: 0.3 mm." httpUri="https://zenodo.org/record/206019/files/figure.png" pageId="4" pageNumber="38">Figs. 6–7</figureCitation>
,
<figureCitation id="130A0E960731D0755B4AFC11FDE8FC6A" box="[539,611,877,900]" captionStart="FIGURES 14, 15" captionStartId="4.[151,269,704,728]" captionTargetBox="[151,1436,182,680]" captionTargetId="figure@4.[151,1436,182,680]" captionTargetPageId="4" captionText="FIGURES 14, 15. Harpactea complicata sp. nov., epigyne and vulva. 14: ventral view; 15: vulva, dorsal view. Scale lines: 0.2 mm." httpUri="https://zenodo.org/record/206021/files/figure.png" pageId="4" pageNumber="38">14, 15</figureCitation>
): short and sclerotized spermatheca, globular at its upper part. Anterior arc with thin lateral edges. Transversal bar indistinguishable; posterior diverticulum small, oval-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C32B41980731D0755996FCCFFE7EFA7C" pageId="4" pageNumber="38" type="distribution">
<paragraph id="8B8E12130731D0755996FCCFFBA0FC24" blockId="4.[151,1436,808,971]" box="[199,1067,947,971]" pageId="4" pageNumber="38">
<emphasis id="B945CE010731D0755996FCCFFED2FC25" bold="true" box="[199,345,947,971]" pageId="4" pageNumber="38">Distribution.</emphasis>
Only known from the localities where the
<typeStatus id="548AACB10731D0755A73FCCFFCDBFC24" box="[802,848,947,970]" pageId="4" pageNumber="38">type</typeStatus>
material was found.
</paragraph>
<paragraph id="8B8E12130731D07559C6FC86FC35FBFF" blockId="4.[151,1278,1016,1263]" box="[151,958,1016,1042]" pageId="4" pageNumber="38">
<emphasis id="B945CE010731D07559C6FC86FE83FBFC" bold="true" box="[151,264,1018,1042]" pageId="4" pageNumber="38">TABLE 3.</emphasis>
<taxonomicName id="4C3169900731D0755841FC86FE74FBFF" box="[272,511,1018,1041]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="complicata" status="sp. nov.">
<emphasis id="B945CE010731D0755841FC86FE74FBFF" box="[272,511,1018,1041]" italics="true" pageId="4" pageNumber="38">Harpactea complicata</emphasis>
</taxonomicName>
<emphasis id="B945CE010731D0755B57FC86FDD0FBFC" bold="true" box="[518,603,1018,1042]" pageId="4" pageNumber="38">
<taxonomicNameLabel id="A276737A0731D0755B57FC86FDD0FBFC" box="[518,603,1018,1042]" pageId="4" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, leg measurements (Ψ
<typeStatus id="548AACB10731D0755A03FC86FC38FBFF" box="[850,947,1018,1041]" pageId="4" pageNumber="38" type="paratype">paratype</typeStatus>
).
</paragraph>
<paragraph id="8B8E12130731D07559CEFB51FE7EFA7C" blockId="4.[151,1278,1016,1263]" lastBlockId="4.[151,1436,1328,1426]" pageId="4" pageNumber="38">Legs Femur Patella Tibia Metatarsus Tarsus Total I 3.00 1.50 2.32 2.25 1.42 10.42 II 2.40 1.42 2.25 2.10 0.75 8.92 III 2.17 0.90 1.50 1.87 0.82 7.26 IV 2.74 1.12 2.40 2.85 0.97 10.08 We are especially indebted to our colleagues V. Tomiċ and S. B. Ć urċiċ (Belgrade) for the collected material, to J. Dunlop for access to the comparative material. This research was supported by a grant of the Deutsche Forschungsgemeinschaft (DFG).</paragraph>
</subSubSection>
</treatment>
</document>
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