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<document id="69A55BAF7D5F002ED5BF09DB065D010F" ID-DOI="10.11646/zootaxa.3731.2.9" ID-GBIF-Dataset="2ed4d413-1628-4af9-8ce0-94f224479524" ID-ISSN="1175-5326" ID-Zenodo-Dep="221617" ID-ZooBank="E17CEF13-F120-4C30-B60D-BD00431B748A" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460694066611" checkinUser="plazi" docAuthor="Britz, Ralf" docDate="2013" docId="0397DB4D2E273D63FF49FF25FCF1DC2B" docLanguage="en" docName="zt03731p294.pdf" docOrigin="Zootaxa 3731 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Channa andrao Britz, 2013, new species" docType="treatment" docVersion="8" lastPageNumber="291" masterDocId="FFAEA3352E263D67FFDEFFB2FFC8DD37" masterDocTitle="Channa andrao, a new species of dwarf snakehead from West Bengal, India (Teleostei: Channidae)" masterLastPageNumber="294" masterPageNumber="287" pageNumber="288" updateTime="1698353523425" updateUser="plazi">
<mods:mods id="852F9647E0867CE75B32BE27A5BE58F9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B108BFC090289FCB706DDC9D02455CE3">
<mods:title id="D9E94DD2224EF245F734C6F3D3CC8E21">Channa andrao, a new species of dwarf snakehead from West Bengal, India (Teleostei: Channidae)</mods:title>
</mods:titleInfo>
<mods:name id="87E91552AF345232B2CAB9C8E653B2E0" type="personal">
<mods:role id="B4260854B3DE08CFFCBE628AA07FDE1E">
<mods:roleTerm id="E8994BF49F1027B423862B0A87BCFCAC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8FBE91EC85EC006FBA53781989DF41F2">Britz, Ralf</mods:namePart>
</mods:name>
<mods:typeOfResource id="970E367A1CCBD97AF2A8B0B62780C138">text</mods:typeOfResource>
<mods:relatedItem id="2C8C7F35580FD98FD19743AF156C7AB7" type="host">
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<mods:title id="4C9275F75A04EBE81C4D52355D7F0B5E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E4CD91C2689C90F53CD9A1B48EF15856">
<mods:date id="90393C59EB28117BFC9CA828EC1364D8">2013</mods:date>
<mods:detail id="F3094AD8A544486C876E3EBC87D59624" type="volume">
<mods:number id="1DA037926D29B6A595B54C19280FA59A">3731</mods:number>
</mods:detail>
<mods:detail id="76CFC0267F8F0418AFF146D7DD484C80" type="issue">
<mods:number id="000643265EE4605AA9BEADECD9F88D8F">2</mods:number>
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</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="574F47E6A1B378A14713F667D242E844">journal article</mods:classification>
<mods:identifier id="76E369B226B2023B32F262F90D38910B" type="DOI">10.11646/zootaxa.3731.2.9</mods:identifier>
<mods:identifier id="0BF2A3D74B46AE2FCBABB392A899C6E9" type="GBIF-Dataset">2ed4d413-1628-4af9-8ce0-94f224479524</mods:identifier>
<mods:identifier id="50465B36BE70D0D7E293A0F2E2913D13" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AC69FFBB14C44DB852F9C38A92640B1E" type="Zenodo-Dep">221617</mods:identifier>
<mods:identifier id="B308384D6E21D18E33BF22D3255690C8" type="ZooBank">E17CEF13-F120-4C30-B60D-BD00431B748A</mods:identifier>
</mods:mods>
<treatment id="0397DB4D2E273D63FF49FF25FCF1DC2B" ID-DOI="http://doi.org/10.5281/zenodo.5686411" ID-GBIF-Taxon="119581325" ID-Zenodo-Dep="5686411" LSID="urn:lsid:plazi:treatment:0397DB4D2E273D63FF49FF25FCF1DC2B" httpUri="http://treatment.plazi.org/id/0397DB4D2E273D63FF49FF25FCF1DC2B" lastPageId="4" lastPageNumber="291" pageId="1" pageNumber="288">
<subSubSection id="C32439D02E273D66FF49FF25FEE2DDE4" pageId="1" pageNumber="288" type="nomenclature">
<paragraph id="8B816A5B2E273D66FF49FF25FE3CDD85" blockId="1.[151,500,151,211]" box="[151,500,151,178]" pageId="1" pageNumber="288">
<heading id="D0C9DD372E273D66FF49FF25FE3CDD85" bold="true" box="[151,500,151,178]" fontSize="11" level="1" pageId="1" pageNumber="288" reason="1">
<emphasis id="B94AB6492E273D66FF49FF25FE3CDD85" bold="true" box="[151,500,151,178]" pageId="1" pageNumber="288">
<taxonomicName id="4C3E11D82E273D66FF49FF25FE9EDD86" ID-CoL="TM2K" box="[151,342,151,177]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="288" phylum="Chordata" rank="species" species="andrao" status="sp. nov.">
<emphasis id="B94AB6492E273D66FF49FF25FE9EDD86" bold="true" box="[151,342,151,177]" italics="true" pageId="1" pageNumber="288">Channa andrao</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2790B322E273D66FEBBFF2AFE3CDD85" box="[357,500,152,178]" pageId="1" pageNumber="288" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B816A5B2E273D66FF49FF08FEE2DDE4" blockId="1.[151,500,151,211]" box="[151,298,186,211]" pageId="1" pageNumber="288">
<figureCitation id="130576DE2E273D66FF49FF08FF35DDE4" box="[151,253,186,211]" captionStart="FIGURE 1" captionStartId="1.[151,250,1498,1520]" captionTargetBox="[151,1436,410,1472]" captionTargetId="figure@1.[151,1436,410,1474]" captionTargetPageId="1" captionText="FIGURE 1. Channa andrao, BMNH 2013.10. 1.2, holotype, 98.2 mm SL; India: West Bengal: Lefraguri swamp." httpUri="https://zenodo.org/record/221618/files/figure.png" pageId="1" pageNumber="288">Figures 1</figureCitation>
,
<figureCitation id="130576DE2E273D66FEDDFF08FEDDDDE4" box="[259,277,186,211]" captionStart="FIGURE 2" captionStartId="1.[151,250,1953,1975]" captionTargetBox="[294,1291,1570,1931]" captionTargetId="figure@1.[294,1292,1570,1932]" captionTargetPageId="1" captionText="FIGURE 2. Channa andrao, BMNH 2013.10.1.3 – 11, paratype, 95.5 mm SL, live coloration." httpUri="https://zenodo.org/record/221619/files/figure.png" pageId="1" pageNumber="288">2</figureCitation>
,
<figureCitation id="130576DE2E273D66FEC4FF08FEE2DDE4" box="[282,298,186,211]" captionStart="FIGURE 5" captionStartId="3.[151,250,1364,1386]" captionTargetBox="[226,1360,807,1339]" captionTargetId="figure@3.[226,1360,807,1339]" captionTargetPageId="3" captionText="FIGURE 5. Pectoral fins of A, Channa andrao, BMNH 2013.10. 1.2, holotype, 98.2 mm; B, C. orientalis ZMB 5029, 79.7 mm SL; C, C. bleheri, ZFMK 16555, holotype, 105.7 mm SL; D, C. asiatica, BMNH 1933.3. 11.900, 115.7 mm SL." httpUri="https://zenodo.org/record/221622/files/figure.png" pageId="1" pageNumber="288">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32439D02E273D66FF49FEB1FB98DC54" pageId="1" pageNumber="288" type="materials_examined">
<paragraph id="8B816A5B2E273D66FF49FEB1FDE1DC08" blockId="1.[151,1436,258,355]" pageId="1" pageNumber="288">
<emphasis id="B94AB6492E273D66FF49FEB1FEC3DC2C" bold="true" box="[151,267,259,283]" pageId="1" pageNumber="288">
<typeStatus id="5485D4F92E273D66FF49FEB1FECEDC2C" box="[151,262,259,283]" pageId="1" pageNumber="288" type="holotype">Holotype</typeStatus>
.
</emphasis>
BMNH 2013.10.1.2,
<quantity id="4CC6C7BE2E273D66FDD9FEB1FDA6DC2D" box="[519,622,259,283]" metricMagnitude="-2" metricUnit="m" metricValue="9.82" pageId="1" pageNumber="288" unit="mm" value="98.2">98.2 mm</quantity>
SL;
<collectingCountry id="F3292ACB2E273D66FD77FEB0FD2FDC2C" box="[681,743,258,283]" name="India" pageId="1" pageNumber="288">India</collectingCountry>
: West Bengal: Jalpaiguri district: Lefraguri swamp,
<geoCoordinate id="EE0A0C9C2E273D66FA9EFEB0FA54DC2C" box="[1344,1436,258,283]" direction="north" orientation="latitude" pageId="1" pageNumber="288" precision="925" value="26.516666">26°31'N</geoCoordinate>
<geoCoordinate id="EE0A0C9C2E273D66FF49FE94FF3BDC08" box="[151,243,294,319]" direction="east" orientation="longitude" pageId="1" pageNumber="288" precision="925" value="89.833336">89°50'E</geoCoordinate>
; Pradhan K.
<date id="FF804C9B2E273D66FE5AFE95FDEDDC08" box="[388,549,294,319]" pageId="1" pageNumber="288" value="2008-04-28">28 April 2008</date>
.
</paragraph>
<paragraph id="8B816A5B2E273D66FF19FEF9FB98DC54" blockId="1.[151,1436,258,355]" box="[199,1104,330,355]" pageId="1" pageNumber="288">
<emphasis id="B94AB6492E273D66FF19FEF9FE8EDC54" bold="true" box="[199,326,331,355]" pageId="1" pageNumber="288">
<typeStatus id="5485D4F92E273D66FF19FEF9FE89DC54" box="[199,321,331,355]" pageId="1" pageNumber="288" type="paratype">Paratypes</typeStatus>
.
</emphasis>
BMNH 2013.10.1.3–11, 9,
<quantity id="4CC6C7BE2E273D66FDA1FEF9FCD3DC55" box="[639,795,331,355]" metricMagnitude="-2" metricUnit="m" metricValue="9.115" metricValueMax="10.03" metricValueMin="8.2" pageId="1" pageNumber="288" unit="mm" value="91.15" valueMax="100.3" valueMin="82.0">82–100.3 mm</quantity>
SL; same data as
<typeStatus id="5485D4F92E273D66FC38FEF8FB84DC54" box="[998,1100,330,355]" pageId="1" pageNumber="288" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="DF413AD32E273D66FF49FA68FAD8D8C7" box="[151,1296,1498,1520]" httpUri="https://zenodo.org/record/221618/files/figure.png" pageId="1" pageNumber="288" targetBox="[151,1436,410,1472]" targetPageId="1">
<paragraph id="8B816A5B2E273D66FF49FA68FAD8D8C7" blockId="1.[151,1296,1498,1520]" box="[151,1296,1498,1520]" pageId="1" pageNumber="288">
<emphasis id="B94AB6492E273D66FF49FA68FEDBD8D8" bold="true" box="[151,275,1498,1520]" pageId="1" pageNumber="288">FIGURE 1.</emphasis>
<taxonomicName id="4C3E11D82E273D66FEC7FA68FE70D8C7" box="[281,440,1498,1520]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="288" phylum="Chordata" rank="species" species="andrao">
<emphasis id="B94AB6492E273D66FEC7FA68FE70D8C7" box="[281,440,1498,1520]" italics="true" pageId="1" pageNumber="288">Channa andrao</emphasis>
</taxonomicName>
, BMNH 2013.10.1.2, holotype, 98.2 mm SL; India: West Bengal: Lefraguri swamp.
</paragraph>
</caption>
<caption id="DF413AD32E273D66FF49F813FB82DA80" box="[151,1098,1953,1975]" httpUri="https://zenodo.org/record/221619/files/figure.png" pageId="1" pageNumber="288" targetBox="[294,1291,1570,1931]" targetPageId="1">
<paragraph id="8B816A5B2E273D66FF49F813FB82DA80" blockId="1.[151,1098,1953,1975]" box="[151,1098,1953,1975]" pageId="1" pageNumber="288">
<emphasis id="B94AB6492E273D66FF49F813FEDBDA81" bold="true" box="[151,275,1953,1975]" pageId="1" pageNumber="288">FIGURE 2.</emphasis>
<taxonomicName id="4C3E11D82E273D66FEC7F813FE70DA80" box="[281,440,1953,1975]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="288" phylum="Chordata" rank="species" species="andrao">
<emphasis id="B94AB6492E273D66FEC7F813FE70DA80" box="[281,440,1953,1975]" italics="true" pageId="1" pageNumber="288">Channa andrao</emphasis>
</taxonomicName>
, BMNH 2013.10.1.3–11, paratype, 95.5 mm SL, live coloration.
</paragraph>
</caption>
<caption id="DF413AD32E243D65FF49FB2DFC9BD982" box="[151,851,1183,1205]" httpUri="https://zenodo.org/record/221620/files/figure.png" pageId="2" pageNumber="289" targetBox="[158,1427,177,1161]" targetPageId="2">
<paragraph id="8B816A5B2E243D65FF49FB2DFC9BD982" blockId="2.[151,851,1183,1205]" box="[151,851,1183,1205]" pageId="2" pageNumber="289">
<emphasis id="B94AB6492E243D65FF49FB2DFEDAD983" bold="true" box="[151,274,1183,1205]" pageId="2" pageNumber="289">FIGURE 3.</emphasis>
<taxonomicName id="4C3E11D82E243D65FEC7FB2DFE7FD982" box="[281,439,1183,1205]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="bleheri">
<emphasis id="B94AB6492E243D65FEC7FB2DFE7FD982" box="[281,439,1183,1205]" italics="true" pageId="2" pageNumber="289">Channa bleheri</emphasis>
</taxonomicName>
, ZFMK 16555, holotype, 105.7 mm SL.
</paragraph>
</caption>
<caption id="DF413AD32E243D65FF49F9FBFCFBDB68" box="[151,819,1609,1631]" httpUri="https://zenodo.org/record/221621/files/figure.png" pageId="2" pageNumber="289" targetBox="[186,1415,1284,1551]" targetPageId="2">
<paragraph id="8B816A5B2E243D65FF49F9FBFCFBDB68" blockId="2.[151,819,1609,1631]" box="[151,819,1609,1631]" pageId="2" pageNumber="289">
<emphasis id="B94AB6492E243D65FF49F9FBFEDBDB69" bold="true" box="[151,275,1609,1631]" pageId="2" pageNumber="289">FIGURE 4.</emphasis>
<taxonomicName id="4C3E11D82E243D65FEC7F9FBFE77DB68" box="[281,447,1609,1631]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="asiatica">
<emphasis id="B94AB6492E243D65FEC7F9FBFE77DB68" box="[281,447,1609,1631]" italics="true" pageId="2" pageNumber="289">Channa asiatica</emphasis>
</taxonomicName>
, holotype, ZMUU 171, 170 mm SL.
</paragraph>
</caption>
<subSubSection id="C32439D02E243D64FF19F939FF35DC52" lastPageId="3" lastPageNumber="290" pageId="2" pageNumber="289" type="diagnosis">
<paragraph id="8B816A5B2E243D64FF19F939FF35DC52" blockId="2.[151,1436,1675,2025]" lastBlockId="3.[151,1436,151,752]" lastPageId="3" lastPageNumber="290" pageId="2" pageNumber="289">
<emphasis id="B94AB6492E243D65FF19F939FE89DB93" bold="true" box="[199,321,1675,1700]" pageId="2" pageNumber="289">Diagnosis.</emphasis>
<taxonomicName id="4C3E11D82E243D65FE96F93EFE32DB93" box="[328,506,1676,1700]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="andrao">
<emphasis id="B94AB6492E243D65FE96F93EFE32DB93" box="[328,506,1676,1700]" italics="true" pageId="2" pageNumber="289">Channa andrao</emphasis>
</taxonomicName>
is distinguished from all other species in the genus, except,
<taxonomicName id="4C3E11D82E243D65FB7FF93FFA54DB92" authority="Linnaeus" authorityName="Linnaeus" box="[1185,1436,1676,1701]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="asiatica">
<emphasis id="B94AB6492E243D65FB7FF93FFAD4DB93" box="[1185,1308,1677,1700]" italics="true" pageId="2" pageNumber="289">C. asiatica</emphasis>
(Linnaeus)
</taxonomicName>
(
<figureCitation id="130576DE2E243D65FF41F902FF2CDBFE" box="[159,228,1712,1737]" captionStart="FIGURE 4" captionStartId="2.[151,250,1609,1631]" captionTargetBox="[186,1415,1284,1551]" captionTargetId="figure@2.[151,1436,1255,1588]" captionTargetPageId="2" captionText="FIGURE 4. Channa asiatica, holotype, ZMUU 171, 170 mm SL." httpUri="https://zenodo.org/record/221621/files/figure.png" pageId="2" pageNumber="289">Fig. 4</figureCitation>
),
<taxonomicName id="4C3E11D82E243D65FF25F903FE76DBFE" authority="Vierke" authorityName="Vierke" box="[251,446,1712,1737]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="bleheri">
<emphasis id="B94AB6492E243D65FF25F903FEA3DBFF" box="[251,363,1712,1736]" italics="true" pageId="2" pageNumber="289">C. bleheri</emphasis>
Vierke
</taxonomicName>
(
<figureCitation id="130576DE2E243D65FE10F902FDDADBFE" box="[462,530,1712,1737]" captionStart="FIGURE 3" captionStartId="2.[151,250,1183,1205]" captionTargetBox="[158,1427,177,1161]" captionTargetId="figure@2.[158,1428,177,1162]" captionTargetPageId="2" captionText="FIGURE 3. Channa bleheri, ZFMK 16555, holotype, 105.7 mm SL." httpUri="https://zenodo.org/record/221620/files/figure.png" pageId="2" pageNumber="289">Fig. 3</figureCitation>
),
<taxonomicName id="4C3E11D82E243D65FDF7F903FC8DDBFE" authority="Chaudhuri" authorityName="Chaudhuri" box="[553,837,1712,1737]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="burmanica">
<emphasis id="B94AB6492E243D65FDF7F903FD0CDBFF" box="[553,708,1712,1736]" italics="true" pageId="2" pageNumber="289">C. burmanica</emphasis>
Chaudhuri
</taxonomicName>
, the recently described
<taxonomicName id="4C3E11D82E243D65FB8EF903FA82DBFE" authority="Nguyen" authorityName="Nguyen" box="[1104,1354,1712,1737]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="hoaluensis">
<emphasis id="B94AB6492E243D65FB8EF903FB22DBFF" box="[1104,1258,1712,1736]" italics="true" pageId="2" pageNumber="289">C. hoaluensis</emphasis>
Nguyen
</taxonomicName>
and
<taxonomicName id="4C3E11D82E243D65FA5CF903FE5EDBDA" authority="Nguyen" authorityName="Nguyen" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="ninhbinhensis">
<emphasis id="B94AB6492E243D65FA5CF903FEFBDBDB" italics="true" pageId="2" pageNumber="289">C. ninhbinhensis</emphasis>
Nguyen
</taxonomicName>
, and
<taxonomicName id="4C3E11D82E243D65FE0FF967FCF5DBDA" authority="Bloch &amp; Schneider" authorityName="Bloch &amp; Schneider" box="[465,829,1748,1773]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="B94AB6492E243D65FE0FF967FD96DBDB" box="[465,606,1748,1772]" italics="true" pageId="2" pageNumber="289">C. orientalis</emphasis>
Bloch &amp; Schneider
</taxonomicName>
by the absence of pelvic fins. It is distinguished from
<taxonomicName id="4C3E11D82E243D65FF49F94BFEDADA27" box="[151,274,1785,1808]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="asiatica">
<emphasis id="B94AB6492E243D65FF49F94BFEDADA27" box="[151,274,1785,1808]" italics="true" pageId="2" pageNumber="289">C. asiatica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3E11D82E243D65FEFEF94BFE72DA27" box="[288,442,1784,1808]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="hoaluensis">
<emphasis id="B94AB6492E243D65FEFEF94BFE72DA27" box="[288,442,1784,1808]" italics="true" pageId="2" pageNumber="289">C. hoaluensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3E11D82E243D65FE17F94BFD4FDA27" box="[457,647,1784,1808]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="ninhbinhensis">
<emphasis id="B94AB6492E243D65FE17F94BFD4FDA27" box="[457,647,1784,1808]" italics="true" pageId="2" pageNumber="289">C. ninhbinhensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3E11D82E243D65FD61F94BFC92DA27" box="[703,858,1784,1808]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="burmanica">
<emphasis id="B94AB6492E243D65FD61F94BFC92DA27" box="[703,858,1784,1808]" italics="true" pageId="2" pageNumber="289">C. burmanica</emphasis>
</taxonomicName>
by fewer vertebrae (41–43 vs. 45–57). It is further distinguished from
<emphasis id="B94AB6492E243D65FEAEF8AFFD46DA03" box="[368,654,1820,1844]" italics="true" pageId="2" pageNumber="289">
<taxonomicName id="4C3E11D82E243D65FEAEF8AFFE23DA03" box="[368,491,1821,1844]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="asiatica">C. asiatica</taxonomicName>
,
<taxonomicName id="4C3E11D82E243D65FE28F8AFFD46DA03" box="[502,654,1820,1844]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="hoaluensis">C. hoaluensis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C3E11D82E243D65FD1AF8AFFC48DA03" box="[708,896,1820,1844]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="ninhbinhensis">
<emphasis id="B94AB6492E243D65FD1AF8AFFC48DA03" box="[708,896,1820,1844]" italics="true" pageId="2" pageNumber="289">C. ninhbinhensis</emphasis>
</taxonomicName>
by fewer dorsal- (35–36 vs. 46–49) and anal-fin rays (23–26 vs. 28–32), and fewer scales in the lateral line (42–43 vs. 55–64). It differs also from
<taxonomicName id="4C3E11D82E243D65FB22F8F3FAB0DA6F" box="[1276,1400,1857,1880]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="asiatica">
<emphasis id="B94AB6492E243D65FB22F8F3FAB0DA6F" box="[1276,1400,1857,1880]" italics="true" pageId="2" pageNumber="289">C. asiatica</emphasis>
</taxonomicName>
by fewer scales above (3.5 vs 5.5) and below the lateral line (5.5 vs. 11), fewer preanal scales (21–22 vs. 39), fewer pectoral-fin rays (14–15 vs. 16), fewer principal caudal rays (12–13 vs. 16) and by its reproductive behaviour (mouthbrooder vs. floating-egg guarder). The new species differs from
<taxonomicName id="4C3E11D82E243D65FC3EF81EFB78DAF3" box="[992,1200,1964,1988]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="B94AB6492E243D65FC3EF81EFB78DAF3" box="[992,1200,1964,1988]" italics="true" pageId="2" pageNumber="289">Channa orientalis</emphasis>
</taxonomicName>
by a longer dorsal- (64.6–68.8 vs. 58.0–59.6 % SL), and anal-fin base (40.8–44.5 vs. 36.4–38.8 % SL), more lateral line scales (42–43 vs. 40–41), more vertebrae (41–43 vs. 38–40), more anal-fin rays (23–26 vs. 21–22) and the coloration of its pectoral fins (two to three basal pectoral-fin bands vs. five narrow pectoral-fin bands, see
<figureCitation id="130576DE2E253D64FBA2FF0EFB76DDE2" box="[1148,1214,188,213]" captionStart="FIGURE 5" captionStartId="3.[151,250,1364,1386]" captionTargetBox="[226,1360,807,1339]" captionTargetId="figure@3.[226,1360,807,1339]" captionTargetPageId="3" captionText="FIGURE 5. Pectoral fins of A, Channa andrao, BMNH 2013.10. 1.2, holotype, 98.2 mm; B, C. orientalis ZMB 5029, 79.7 mm SL; C, C. bleheri, ZFMK 16555, holotype, 105.7 mm SL; D, C. asiatica, BMNH 1933.3. 11.900, 115.7 mm SL." httpUri="https://zenodo.org/record/221622/files/figure.png" pageId="3" pageNumber="290">Fig. 5</figureCitation>
).
<taxonomicName id="4C3E11D82E253D64FB0DFF0EFA4BDDE3" box="[1235,1411,188,212]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="290" phylum="Chordata" rank="species" species="andrao">
<emphasis id="B94AB6492E253D64FB0DFF0EFA4BDDE3" box="[1235,1411,188,212]" italics="true" pageId="3" pageNumber="290">Channa andrao</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C3E11D82E253D64FE92FF52FE76DDCF" box="[332,446,224,248]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="290" phylum="Chordata" rank="species" species="bleheri">
<emphasis id="B94AB6492E253D64FE92FF52FE76DDCF" box="[332,446,224,248]" italics="true" pageId="3" pageNumber="290">C. bleheri</emphasis>
</taxonomicName>
(Figs. 3,5), but differs from it by a longer dorsal-fin base (64.6–68.8 vs. 59.7–63.7 % SL), absence (vs. presence) of whitish-cream (orange-red in life) blotches on the caudal fin, only two (vs. five) dark concentric bands in the pectoral fin (see
<figureCitation id="130576DE2E253D64FDBDFE95FD6EDC77" box="[611,678,295,320]" captionStart="FIGURE 4" captionStartId="2.[151,250,1609,1631]" captionTargetBox="[186,1415,1284,1551]" captionTargetId="figure@2.[151,1436,1255,1588]" captionTargetPageId="2" captionText="FIGURE 4. Channa asiatica, holotype, ZMUU 171, 170 mm SL." httpUri="https://zenodo.org/record/221621/files/figure.png" pageId="3" pageNumber="290">Fig. 4</figureCitation>
) and by its reproductive behaviour (mouthbrooder vs. floating-egg guarder).
</paragraph>
</subSubSection>
<subSubSection id="C32439D02E253D64FF19FEDDFBF9DA61" pageId="3" pageNumber="290" type="description">
<paragraph id="8B816A5B2E253D64FF19FEDDFAD9DCBF" blockId="3.[151,1436,151,752]" box="[199,1297,367,392]" pageId="3" pageNumber="290">
<emphasis id="B94AB6492E253D64FF19FEDDFE91DCBF" bold="true" box="[199,345,367,392]" pageId="3" pageNumber="290">Description.</emphasis>
For general appearance see
<figureCitation id="130576DE2E253D64FD48FEDDFCD5DCBF" box="[662,797,367,392]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,250,1498,1520]" captionStartId-1="1.[151,250,1953,1975]" captionTargetBox-0="[151,1436,410,1472]" captionTargetBox-1="[294,1291,1570,1931]" captionTargetId-0="figure@1.[151,1436,410,1474]" captionTargetId-1="figure@1.[294,1292,1570,1932]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionText-0="FIGURE 1. Channa andrao, BMNH 2013.10. 1.2, holotype, 98.2 mm SL; India: West Bengal: Lefraguri swamp." captionText-1="FIGURE 2. Channa andrao, BMNH 2013.10.1.3 – 11, paratype, 95.5 mm SL, live coloration." httpUri-0="https://zenodo.org/record/221618/files/figure.png" httpUri-1="https://zenodo.org/record/221619/files/figure.png" pageId="3" pageNumber="290">Figures 1–2</figureCitation>
. Morphometric data are provided in
<tableCitation id="C6BC5FE02E253D64FB66FEDDFAC3DCBF" box="[1208,1291,367,392]" captionStart="TABLE 1" captionStartId="4.[151,239,328,351]" captionTargetBox="[159,1413,410,1951]" captionTargetPageId="4" captionText="TABLE 1. Selected morphometric data for holotype (BMNH 2013.10.1.2) and 9 paratypes (BMNH 2013.10.1.3 – 11) of Channa andrao." httpUri="http://table.plazi.org/id/DF413AD32E223D63FF49FEFAFE8EDC48" pageId="3" pageNumber="290" tableUuid="DF413AD32E223D63FF49FEFAFE8EDC48">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8B816A5B2E253D64FF19FE26FBB4DF0A" blockId="3.[151,1436,151,752]" pageId="3" pageNumber="290">
A member of the
<taxonomicName id="4C3E11D82E253D64FE47FE26FD87DC9B" box="[409,591,404,428]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="290" phylum="Chordata" rank="species" species="gachua">
<emphasis id="B94AB6492E253D64FE47FE26FD87DC9B" box="[409,591,404,428]" italics="true" pageId="3" pageNumber="290">Channa gachua</emphasis>
</taxonomicName>
species group (sensu Britz 2008). Body elongate, round in cross section anteriorly, laterally compressed at caudal peduncle, body depth 5.7–6.6 times in SL. Head large, length 3.4–3.7 times in SL. Head widest halfway between posterior margin of eye and that of opercle. Mouth large, maxilla extending posteriorly beyond jaw angle, lips fleshy. Both jaws with multiple rows of sharp, pointed minute teeth, no canines. Vomer with small number of canine-like teeth only. Minute teeth on palatine.
</paragraph>
<paragraph id="8B816A5B2E253D64FF0FFDF5FBA9DFB2" blockId="3.[151,1436,151,752]" pageId="3" pageNumber="290">Pectoral-fin rays 13–15. Dorsal-fin rays 35–36. Anal-fin rays 23–26. Caudal-fin rays (including procurrent rays): 1+6+6+2(1), 2+6+6+2 (5), 2+6+6+3(1), 3+6+6+3(1), 2+7+6+3(1), 2+6+7+1(1).</paragraph>
<paragraph id="8B816A5B2E253D64FF19FD3DFB24DFC7" blockId="3.[151,1436,151,752]" pageId="3" pageNumber="290">Predorsal scales 13(2)–14(8). Lateral line with 42(5)–43(5) scales extending from shoulder girdle in a horizontal line, but dropping one scale row at scale 13(4) or 14 (6), then continuing horizontally to caudal peduncle. Scales above lateral line 3.5 (10) and below 5.5 (10) at anal-fin origin. Vertebrae: 41–43.</paragraph>
<caption id="DF413AD32E253D64FF49FAE6FAC8D8BF" httpUri="https://zenodo.org/record/221622/files/figure.png" pageId="3" pageNumber="290" targetBox="[226,1360,807,1339]" targetPageId="3">
<paragraph id="8B816A5B2E253D64FF49FAE6FAC8D8BF" blockId="3.[151,1435,1364,1416]" pageId="3" pageNumber="290">
<emphasis id="B94AB6492E253D64FF49FAE6FEDAD85E" bold="true" box="[151,274,1364,1386]" pageId="3" pageNumber="290">FIGURE 5.</emphasis>
Pectoral fins of A,
<taxonomicName id="4C3E11D82E253D64FE09FAE6FDBED85D" box="[471,630,1364,1386]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="290" phylum="Chordata" rank="species" species="andrao">
<emphasis id="B94AB6492E253D64FE09FAE6FDBED85D" box="[471,630,1364,1386]" italics="true" pageId="3" pageNumber="290">Channa andrao</emphasis>
</taxonomicName>
, BMNH 2013.10.1.2, holotype, 98.2 mm; B,
<taxonomicName id="4C3E11D82E253D64FB9CFAE6FB08D85D" box="[1090,1216,1364,1386]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="290" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="B94AB6492E253D64FB9CFAE6FB08D85D" box="[1090,1216,1364,1386]" italics="true" pageId="3" pageNumber="290">C. orientalis</emphasis>
</taxonomicName>
ZMB 5029, 79.7 mm SL; C,
<taxonomicName id="4C3E11D82E253D64FF01FAC1FE8CD8BF" box="[223,324,1394,1416]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="290" phylum="Chordata" rank="species" species="bleheri">
<emphasis id="B94AB6492E253D64FF01FAC1FE8CD8BF" box="[223,324,1394,1416]" italics="true" pageId="3" pageNumber="290">C. bleheri</emphasis>
</taxonomicName>
, ZFMK 16555, holotype, 105.7 mm SL; D,
<taxonomicName id="4C3E11D82E253D64FCD8FAC1FCBBD8BF" box="[774,883,1395,1416]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="290" phylum="Chordata" rank="species" species="asiatica">
<emphasis id="B94AB6492E253D64FCD8FAC1FCBBD8BF" box="[774,883,1395,1416]" italics="true" pageId="3" pageNumber="290">C. asiatica</emphasis>
</taxonomicName>
, BMNH 1933.3.11.900, 115.7 mm SL.
</paragraph>
</caption>
<paragraph id="8B816A5B2E253D64FF19FA02FF1BDB01" blockId="3.[151,1436,1456,2022]" pageId="3" pageNumber="290">
<emphasis id="B94AB6492E253D64FF19FA02FEE8D8FE" bold="true" box="[199,288,1456,1481]" pageId="3" pageNumber="290">Colour.</emphasis>
In alcohol (
<figureCitation id="130576DE2E253D64FE73FA03FE3AD8FE" box="[429,498,1457,1482]" captionStart="FIGURE 1" captionStartId="1.[151,250,1498,1520]" captionTargetBox="[151,1436,410,1472]" captionTargetId="figure@1.[151,1436,410,1474]" captionTargetPageId="1" captionText="FIGURE 1. Channa andrao, BMNH 2013.10. 1.2, holotype, 98.2 mm SL; India: West Bengal: Lefraguri swamp." httpUri="https://zenodo.org/record/221618/files/figure.png" pageId="3" pageNumber="290">Fig. 1</figureCitation>
,
<figureCitation id="130576DE2E253D64FDDDFA03FDDAD8FD" box="[515,530,1457,1482]" captionStart="FIGURE 5" captionStartId="3.[151,250,1364,1386]" captionTargetBox="[226,1360,807,1339]" captionTargetId="figure@3.[226,1360,807,1339]" captionTargetPageId="3" captionText="FIGURE 5. Pectoral fins of A, Channa andrao, BMNH 2013.10. 1.2, holotype, 98.2 mm; B, C. orientalis ZMB 5029, 79.7 mm SL; C, C. bleheri, ZFMK 16555, holotype, 105.7 mm SL; D, C. asiatica, BMNH 1933.3. 11.900, 115.7 mm SL." httpUri="https://zenodo.org/record/221622/files/figure.png" pageId="3" pageNumber="290">5</figureCitation>
), dorsum and sides of head and body uniformly dark grey. Opercular area and jaws gray, becoming lighter towards throat, with whitish cream marbling spots. Branchiostegal membrane dark grey, isthmus cream. Belly cream, ventral area adjacent to anal fin with alternating whitish cream and grey oblique lines.
</paragraph>
<paragraph id="8B816A5B2E253D64FF19F9F3FC0FDB95" blockId="3.[151,1436,1456,2022]" pageId="3" pageNumber="290">
Pectoral fin with dark grey spot at fleshy base, more distally with two alternating creamish white and dark grey semi-circular bands, distal part of pectoral fin with dark grey fin rays alternating with whitish cream fin membranes (
<figureCitation id="130576DE2E253D64FF41F93BFF2ADB96" box="[159,226,1673,1698]" captionStart="FIGURE 1" captionStartId="1.[151,250,1498,1520]" captionTargetBox="[151,1436,410,1472]" captionTargetId="figure@1.[151,1436,410,1474]" captionTargetPageId="1" captionText="FIGURE 1. Channa andrao, BMNH 2013.10. 1.2, holotype, 98.2 mm SL; India: West Bengal: Lefraguri swamp." httpUri="https://zenodo.org/record/221618/files/figure.png" pageId="3" pageNumber="290">Fig. 1</figureCitation>
,
<figureCitation id="130576DE2E253D64FF31F93BFF37DB95" box="[239,255,1673,1698]" captionStart="FIGURE 5" captionStartId="3.[151,250,1364,1386]" captionTargetBox="[226,1360,807,1339]" captionTargetId="figure@3.[226,1360,807,1339]" captionTargetPageId="3" captionText="FIGURE 5. Pectoral fins of A, Channa andrao, BMNH 2013.10. 1.2, holotype, 98.2 mm; B, C. orientalis ZMB 5029, 79.7 mm SL; C, C. bleheri, ZFMK 16555, holotype, 105.7 mm SL; D, C. asiatica, BMNH 1933.3. 11.900, 115.7 mm SL." httpUri="https://zenodo.org/record/221622/files/figure.png" pageId="3" pageNumber="290">5</figureCitation>
). Dorsal, anal and caudal fins dark grey with narrow white rim.
</paragraph>
<paragraph id="8B816A5B2E253D64FF19F91CFBF9DA61" blockId="3.[151,1436,1456,2022]" pageId="3" pageNumber="290">
In life (
<figureCitation id="130576DE2E253D64FEC0F91FFEACDBF1" box="[286,356,1709,1734]" captionStart="FIGURE 2" captionStartId="1.[151,250,1953,1975]" captionTargetBox="[294,1291,1570,1931]" captionTargetId="figure@1.[294,1292,1570,1932]" captionTargetPageId="1" captionText="FIGURE 2. Channa andrao, BMNH 2013.10.1.3 – 11, paratype, 95.5 mm SL, live coloration." httpUri="https://zenodo.org/record/221619/files/figure.png" pageId="3" pageNumber="290">Fig. 2</figureCitation>
), body light grey, series of red spots on cheek and opercle continued on body by series of red marks arranged in oblique lines, most conspicuous on ventral half of side above anal-fin base, corresponding to whitish cream marks in preserved specimens. Dorsal and anal fin with bluish, sometimes greenish reflecting membranes, distal parts of dorsal and anal-fin rays dark grey, both fins with white rim. Caudal fin with bluish membranes, but dark-grey fin rays, rimmed with red and more distally with white.
</paragraph>
</subSubSection>
<subSubSection id="C32439D02E253D64FF19F8D2FAC5DA4D" box="[199,1293,1888,1914]" pageId="3" pageNumber="290" type="distribution">
<paragraph id="8B816A5B2E253D64FF19F8D2FAC5DA4D" blockId="3.[151,1436,1456,2022]" box="[199,1293,1888,1914]" pageId="3" pageNumber="290">
<emphasis id="B94AB6492E253D64FF19F8D2FEA8DA4E" bold="true" box="[199,352,1888,1913]" pageId="3" pageNumber="290">Distribution.</emphasis>
Known only from Lefraguri swamp, Jalpaiguri district, West Bengal,
<collectingCountry id="F3292ACB2E253D64FBAFF8D3FB63DA4D" box="[1137,1195,1889,1914]" name="India" pageId="3" pageNumber="290">India</collectingCountry>
(
<figureCitation id="130576DE2E253D64FB64F8D3FB37DA4D" box="[1210,1279,1889,1914]" captionStart="FIGURE 6" captionStartId="5.[151,250,1180,1202]" captionTargetBox="[151,1436,193,1156]" captionTargetId="figure@5.[151,1436,193,1157]" captionTargetPageId="5" captionText="FIGURE 6. Lefraguri swamp, West Bengal, type locality of Channa andrao (photo: C. Kanele)." httpUri="https://zenodo.org/record/221623/files/figure.png" pageId="3" pageNumber="290">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32439D02E253D64FF19F837FC97DAF5" pageId="3" pageNumber="290" type="etymology">
<paragraph id="8B816A5B2E253D64FF19F837FC97DAF5" blockId="3.[151,1436,1456,2022]" pageId="3" pageNumber="290">
<emphasis id="B94AB6492E253D64FF19F837FE86DAAA" bold="true" box="[199,334,1925,1949]" pageId="3" pageNumber="290">Etymology.</emphasis>
The species name honours Andrew Rao, Malabar Tropicals, Calcutta, for his support of the ichthyological exploration of the freshwater fish fauna of
<collectingCountry id="F3292ACB2E253D64FCC0F81BFC93DAF5" box="[798,859,1961,1986]" name="India" pageId="3" pageNumber="290">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32439D02E253D63FF19F87FFCF1DC2B" lastPageId="4" lastPageNumber="291" pageId="3" pageNumber="290" type="discussion">
<paragraph id="8B816A5B2E253D63FF19F87FFD56DDE3" blockId="3.[151,1436,1456,2022]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="291" pageId="3" pageNumber="290">
<emphasis id="B94AB6492E253D64FF19F87FFEF3DAD2" bold="true" box="[199,315,1997,2021]" pageId="3" pageNumber="290">Remarks.</emphasis>
This species was caught in a swamp with silty bottom in a forested area (
<figureCitation id="130576DE2E253D64FBAEF87FFB7CDAD1" box="[1136,1204,1997,2022]" captionStart="FIGURE 6" captionStartId="5.[151,250,1180,1202]" captionTargetBox="[151,1436,193,1156]" captionTargetId="figure@5.[151,1436,193,1157]" captionTargetPageId="5" captionText="FIGURE 6. Lefraguri swamp, West Bengal, type locality of Channa andrao (photo: C. Kanele)." httpUri="https://zenodo.org/record/221623/files/figure.png" pageId="3" pageNumber="290">Fig. 6</figureCitation>
). The shallow turbid water had
<collectingCountry id="F3292ACB2E223D63FECBFF2AFEFFDD98" box="[277,311,152,175]" name="Philippines" pageId="4" pageNumber="291">pH</collectingCountry>
6.3, conductivity 72 µS·cm, water temperature 26.9°C, and there was an abundance of aquatic vegetation (recorded
<date id="FF804C9B2E223D63FE5BFF0EFDDCDDE2" box="[389,532,188,213]" pageId="4" pageNumber="291" value="2006-07-11">11 July 2006</date>
by A. Rao).
</paragraph>
<paragraph id="8B816A5B2E223D63FF19FF6DFCF1DC2B" blockId="4.[151,1436,151,285]" pageId="4" pageNumber="291">
This species is known in the ornamental fish trade as
<taxonomicName id="4C3E11D82E223D63FC8CFF52FC63DDCF" box="[850,939,224,248]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="291" phylum="Chordata" rank="genus">
<emphasis id="B94AB6492E223D63FC8CFF52FC63DDCF" box="[850,939,224,248]" italics="true" pageId="4" pageNumber="291">Channa</emphasis>
</taxonomicName>
“blue
<taxonomicName id="4C3E11D82E223D63FC21FF6DFB9ADDCF" box="[1023,1106,223,248]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="291" phylum="Chordata" rank="species" species="bleheri">bleheri</taxonomicName>
”,
<emphasis id="B94AB6492E223D63FBB3FF52FB4FDDC0" box="[1133,1159,224,247]" italics="true" pageId="4" pageNumber="291">C.</emphasis>
“Lal Cheng” or
<emphasis id="B94AB6492E223D63FA88FF52FAA1DDC0" box="[1366,1385,224,247]" italics="true" pageId="4" pageNumber="291">C</emphasis>
. sp. “Assam”. It is a male mouthbrooder (Brede &amp; Antler 2009).
</paragraph>
</subSubSection>
</treatment>
</document>
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