236 lines
30 KiB
XML
236 lines
30 KiB
XML
<document id="15CC14B6670BD3A2AE8AED324F384FD5" ID-DOI="10.11646/zootaxa.5369.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10146170" ID-ZooBank="42563452-CF2B-47A6-B0B0-485AC4F320BD" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1700185584045" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="038DBF6E540BCC6FFF06FBBAFE0BFD73" docLanguage="en" docName="zootaxa.5369.2.1.pdf" docOrigin="Zootaxa 5369 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5369.2.1/52237" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omalium latocavum Shavrin 2023, sp. n." docType="treatment" docVersion="1" lastPageNumber="199" masterDocId="FFB4C7165425CC5FFF91FFD8FFD8FF82" masterDocTitle="A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region" masterLastPageNumber="206" masterPageNumber="151" pageNumber="197" updateTime="1700578467175" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="FD2AD0AA44B3746D5AED5FACECDB42E5">A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region</mods:title>
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<mods:namePart id="A51F50E36F4632308771953FF2AA0363">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="6B401DA53836551D52B338C30B6E194E">Zootaxa</mods:title>
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<mods:part id="2E882801F637D0C1BDC7178F86849335">
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<mods:date id="EBC3FF0C4E6D5CDD65589DDD58B2F6DF">2023</mods:date>
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<mods:number id="7D2168EE531CF06B49DA6B12171FAD2F">2023-11-09</mods:number>
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<mods:number id="FF669E490384F397319093D2EC6F53CA">5369</mods:number>
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<mods:start id="B3EF654884FFE3114CE6E5277231AF4F">151</mods:start>
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<mods:url id="4ADB6C328AD2F0FD24A8D31DE06C2F94">https://www.mapress.com/zt/article/download/zootaxa.5369.2.1/52237</mods:url>
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<mods:classification id="9149B9B3C6E9EF4AF9E3CF3BC196DF0D">journal article</mods:classification>
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<mods:identifier id="FFB662CBAEF37148C0C301118F363486" type="DOI">10.11646/zootaxa.5369.2.1</mods:identifier>
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<mods:identifier id="E859E8A07435725CF1A3A02954E8030C" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038DBF6E540BCC6FFF06FBBAFE0BFD73" LSID="urn:lsid:plazi:treatment:038DBF6E540BCC6FFF06FBBAFE0BFD73" httpUri="http://treatment.plazi.org/id/038DBF6E540BCC6FFF06FBBAFE0BFD73" lastPageId="48" lastPageNumber="199" pageId="46" pageNumber="197">
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<subSubSection id="C33E5DF3540BCC71FF06FBBAFD95FBFF" box="[151,589,1122,1149]" pageId="46" pageNumber="197" type="nomenclature">
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<paragraph id="8B9B0E78540BCC71FF06FBBAFD95FBFF" blockId="46.[151,589,1122,1185]" box="[151,589,1122,1149]" pageId="46" pageNumber="197">
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<heading id="D0D3B914540BCC71FF06FBBAFD95FBFF" bold="true" box="[151,589,1122,1149]" fontSize="11" level="1" pageId="46" pageNumber="197" reason="1">
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<emphasis id="B950D26A540BCC71FF06FBBAFD95FBFF" bold="true" box="[151,589,1122,1149]" pageId="46" pageNumber="197">
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3.2.5.2.5.
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<taxonomicName id="4C2475FB540BCC71FE9CFBBAFDDBFBFF" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[269,515,1122,1149]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="197" phylum="Arthropoda" rank="species" species="latocavum" status="sp. nov.">
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<emphasis id="B950D26A540BCC71FE9CFBBAFDDBFBFF" bold="true" box="[269,515,1122,1149]" italics="true" pageId="46" pageNumber="197">Omalium latocavum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2636F11540BCC71FD9AFBBBFD95FBFF" box="[523,589,1123,1149]" pageId="46" pageNumber="197" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E5DF3540BCC71FF06FB5FFE7DFB23" box="[151,421,1158,1185]" pageId="46" pageNumber="197" type="description">
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<paragraph id="8B9B0E78540BCC71FF06FB5FFE7DFB23" blockId="46.[151,589,1122,1185]" box="[151,421,1158,1185]" pageId="46" pageNumber="197">
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(
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<figureCitation id="131F12FD540BCC71FF0EFB5FFF2EFB23" box="[159,246,1159,1185]" captionStart="FIGURE 69" captionStartId="38.[151,250,1110,1134]" captionTargetBox="[151,1435,273,1086]" captionTargetId="figure-65@38.[151,1435,273,1086]" captionTargetPageId="38" captionText="FIGURE 69. Distribution of Omalium in the Himalayan region: O. angustissimum sp. n. (circle), O. kumari sp. n. (triangles), O. latocavum sp. n. (diamond)." figureDoi="http://doi.org/10.5281/zenodo.10146218" httpUri="https://zenodo.org/record/10146218/files/figure.png" pageId="46" pageNumber="197">Figs 69</figureCitation>
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,
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<figureCitation id="131F12FD540BCC71FE93FB5FFEF8FB23" box="[258,288,1159,1185]" captionStart="FIGURES 76–83" captionStartId="41.[151,264,1651,1675]" captionTargetBox="[154,1432,816,1625]" captionTargetId="figure-275@41.[151,1435,813,1628]" captionTargetPageId="41" captionText="FIGURES 76–83. Female accessory sclerite (Figs 76–77, 79, 81–82) and spermatheca (Figs 78, 80, 83) of Omalium: 76— O.cariosum, 77–78—O. flavotestaceum, 79–80—O. kumari, 81—O. latocavum, 82–83—O. secretum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146224" httpUri="https://zenodo.org/record/10146224/files/figure.png" pageId="46" pageNumber="197">81</figureCitation>
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,
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<figureCitation id="131F12FD540BCC71FEBDFB5FFE93FB23" box="[300,331,1159,1185]" captionStart="FIGURES 90–92" captionStartId="45.[151,264,1333,1357]" captionTargetBox="[164,1418,397,1298]" captionTargetId="figure-104@45.[151,1435,381,1310]" captionTargetPageId="45" captionText="FIGURES 90–92. Habitus of Omalium: 90—O. latocavum (paratype), 91—O. nigrum (holotype), 92—O. secretum (Nepal: Parbat). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146230" httpUri="https://zenodo.org/record/10146230/files/figure.png" pageId="46" pageNumber="197">90</figureCitation>
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,
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<figureCitation id="131F12FD540BCC71FEC7FB5EFE45FB23" box="[342,413,1158,1185]" captionStart="FIGURES 93–95" captionStartId="47.[151,264,1650,1674]" captionTargetBox="[154,1433,420,1624]" captionTargetId="figure-123@47.[151,1436,417,1627]" captionTargetPageId="47" captionText="FIGURES 93–95. Aedeagus of Omalium latocavum (holotype): 93—aedeagus, parameral view, 94—aedeagus, lateral view, 95—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146232" httpUri="https://zenodo.org/record/10146232/files/figure.png" pageId="46" pageNumber="197">93–95</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E5DF3540BCC71FF06FB17FB43FAFB" pageId="46" pageNumber="197" type="materials_examined">
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<paragraph id="8B9B0E78540BCC71FF06FB17FCADFAB3" blockId="46.[151,1437,1230,2013]" pageId="46" pageNumber="197">
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<emphasis id="B950D26A540BCC71FF06FB17FE9EFB6A" bold="true" box="[151,326,1230,1257]" pageId="46" pageNumber="197">Type material.</emphasis>
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<materialsCitation id="3B4C0425540BCC71FEDEFB17FC00FA8F" collectorName="Ahrens & Kulbe & Rulik'" country="Nepal" location="Abuthum Lekh" pageId="46" pageNumber="197" specimenCount="1" stateProvince="SW Ganesh-H." typeStatus="holotype">
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<typeStatus id="549FB0DA540BCC71FEDEFB17FE6FFB6B" box="[335,439,1231,1257]" pageId="46" pageNumber="197" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D22C5F1540BCC71FE51FB08FE0DFB69" box="[448,469,1232,1259]" pageId="46" pageNumber="197">♁</specimenCount>
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(dissected): ‘N Mongeythanti | Bhanjyang 3600- |
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<quantity id="4CDCA39D540BCC71FB8BFB17FBB0FB6B" box="[1050,1128,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="46" pageNumber="197" unit="m" value="4000.0">4000m</quantity>
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19/20.5.[19]96’ <printed>, ‘
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<collectingCountry id="F3334EE8540BCC71FF0AFB2BFF2FFA8F" box="[155,247,1267,1293]" name="Nepal" pageId="46" pageNumber="197">NEPAL</collectingCountry>
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<collectingRegion id="49E0C09A540BCC71FE92FB2BFE6AFA8F" box="[259,434,1267,1293]" pageId="46" pageNumber="197">SW Ganesh-H.</collectingRegion>
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<location id="8EFB58A3540BCC71FE5FFB2BFD58FA8F" LSID="urn:lsid:plazi:treatment:038DBF6E540BCC6FFF06FBBAFE0BFD73:8EFB58A3540BCC71FE5FFB2BFD58FA8F" box="[462,640,1267,1293]" country="Nepal" name="Abuthum Lekh" pageId="46" pageNumber="197" stateProvince="SW Ganesh-H.">Abuthum Lekh</location>
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, leg. |
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<collectorName id="26D16BAE540BCC71FD46FB2BFCF3FA8F" box="[727,811,1267,1293]" pageId="46" pageNumber="197">Ahrens</collectorName>
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,
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<collectorName id="26D16BAE540BCC71FCAAFB2BFC5AFA8F" box="[827,898,1267,1293]" pageId="46" pageNumber="197">Kulbe</collectorName>
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,
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<collectorName id="26D16BAE540BCC71FC02FB2BFC00FA8F" box="[915,984,1267,1293]" pageId="46" pageNumber="197">Rulik’</collectorName>
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</materialsCitation>
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<printed>, ‘
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<typeStatus id="549FB0DA540BCC71FBFAFB2BFADCFA8F" box="[1131,1284,1267,1293]" pageId="46" pageNumber="197" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="4C2475FB540BCC71FAB3FB2BFA52FA8F" authorityName="Gravenhorst" authorityYear="1802" box="[1314,1418,1267,1293]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="197" phylum="Arthropoda" rank="genus">
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<emphasis id="B950D26A540BCC71FAB3FB2BFA52FA8F" box="[1314,1418,1267,1293]" italics="true" pageId="46" pageNumber="197">Omalium</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="4C2475FB540BCC71FF06FACFFED4FAB3" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[151,268,1303,1329]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="197" phylum="Arthropoda" rank="species" species="latocavum" status="sp. nov.">
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<emphasis id="B950D26A540BCC71FF06FACFFED4FAB3" box="[151,268,1303,1329]" italics="true" pageId="46" pageNumber="197">latocavum</emphasis>
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</taxonomicName>
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<emphasis id="B950D26A540BCC71FE82FACFFE8AFAB3" bold="true" box="[275,338,1303,1329]" pageId="46" pageNumber="197">
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<taxonomicNameLabel id="A2636F11540BCC71FE82FACFFE8AFAB3" box="[275,338,1303,1329]" pageId="46" pageNumber="197" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2023’ <red, printed> (NME).
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</paragraph>
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<paragraph id="8B9B0E78540BCC71FF56FAE3FB43FAFB" blockId="46.[151,1437,1230,2013]" pageId="46" pageNumber="197">
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<materialsCitation id="3B4C0425540BCC71FF56FAE3FA41FAD7" box="[199,1433,1339,1367]" collectorName="N Mongeythanti & Bhanjyang & Abuthum Lekh" country="Nepal" county="Rulik'" elevation="4000" location="Ahrens" municipality="Kulbe" pageId="46" pageNumber="197" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<typeStatus id="549FB0DA540BCC71FF56FAE3FEE0FAD7" box="[199,312,1339,1365]" pageId="46" pageNumber="197" type="paratype">Paratypes</typeStatus>
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: 1 ♁ (dissected),
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<specimenCount id="9D22C5F1540BCC71FD9FFAE3FDE5FAD7" box="[526,573,1339,1365]" count="1" pageId="46" pageNumber="197" type="female">1 ♀</specimenCount>
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(dissected): same data as the holotype, with additional red printed label
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</materialsCitation>
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: ‘
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<typeStatus id="549FB0DA540BCC71FF0AFA87FEF3FAFB" box="[155,299,1375,1401]" pageId="46" pageNumber="197" type="paratype">PARATYPE</typeStatus>
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<taxonomicName id="4C2475FB540BCC71FEAEFA87FE7FFAFB" authorityName="Gravenhorst" authorityYear="1802" box="[319,423,1375,1401]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="197" phylum="Arthropoda" rank="genus">
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<emphasis id="B950D26A540BCC71FEAEFA87FE7FFAFB" box="[319,423,1375,1401]" italics="true" pageId="46" pageNumber="197">Omalium</emphasis>
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</taxonomicName>
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<taxonomicName id="4C2475FB540BCC71FE2AFA87FDE8FAFB" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[443,560,1375,1401]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="197" phylum="Arthropoda" rank="species" species="latocavum" status="sp. nov.">
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<emphasis id="B950D26A540BCC71FE2AFA87FDE8FAFB" box="[443,560,1375,1401]" italics="true" pageId="46" pageNumber="197">latocavum</emphasis>
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</taxonomicName>
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<emphasis id="B950D26A540BCC71FDA6FA87FDAEFAFB" bold="true" box="[567,630,1375,1401]" pageId="46" pageNumber="197">
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<taxonomicNameLabel id="A2636F11540BCC71FDA6FA87FDAEFAFB" box="[567,630,1375,1401]" pageId="46" pageNumber="197" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2023’ (1 ♁: cSh,
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<specimenCount id="9D22C5F1540BCC71FB82FA87FBE7FAFB" box="[1043,1087,1375,1401]" count="1" pageId="46" pageNumber="197" type="female">1 ♀</specimenCount>
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: NME).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E5DF3540BCC6FFF56FA5AFE04FE2F" lastPageId="48" lastPageNumber="199" pageId="46" pageNumber="197" type="description">
|
||
<paragraph id="8B9B0E78540BCC71FF56FA5AFF08F98B" blockId="46.[151,1437,1230,2013]" pageId="46" pageNumber="197">
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||
<emphasis id="B950D26A540BCC71FF56FA5AFE81FA1E" bold="true" box="[199,345,1410,1436]" pageId="46" pageNumber="197">Description.</emphasis>
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Measurements (n=3): HW: 0.60–0.62; HL: 0.43–0.49; OL: 0.15; TL: 0.10–0.12; AL (
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<typeStatus id="549FB0DA540BCC71FAB9FA5BFA49FA1F" box="[1320,1425,1411,1437]" pageId="46" pageNumber="197" type="holotype">holotype</typeStatus>
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): 1.15; PL: 0.47–0.50; PWmax: 0.71–0.72; PWmin: 0.67; ESL: 0.90–0.96; EW: 0.92–0.97; MTbL (
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<typeStatus id="549FB0DA540BCC71FB76FA7FFA88FA43" box="[1255,1360,1447,1473]" pageId="46" pageNumber="197" type="holotype">holotype</typeStatus>
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): 0.55; MTrL (
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<typeStatus id="549FB0DA540BCC71FF7BFA13FE8BFA67" box="[234,339,1483,1509]" pageId="46" pageNumber="197" type="holotype">holotype</typeStatus>
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): 0.22 (MTrL 1–4: 0.09; MTrL 5: 0.13); AW: 0.90–0.95; AedL: 0.70–0.72; BL: 3.06 (
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<typeStatus id="549FB0DA540BCC71FAB0FA13FA5EFA67" box="[1313,1414,1483,1509]" pageId="46" pageNumber="197" type="holotype">holotype</typeStatus>
|
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)– 3.28.
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</paragraph>
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<paragraph id="8B9B0E78540BCC71FF56F9CBFDBAF887" blockId="46.[151,1437,1230,2013]" pageId="46" pageNumber="197">
|
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Habitus as in
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<figureCitation id="131F12FD540BCC71FECCF9CBFE77F9AF" box="[349,431,1555,1581]" captionStart="FIGURES 90–92" captionStartId="45.[151,264,1333,1357]" captionTargetBox="[164,1418,397,1298]" captionTargetId="figure-104@45.[151,1435,381,1310]" captionTargetPageId="45" captionText="FIGURES 90–92. Habitus of Omalium: 90—O. latocavum (paratype), 91—O. nigrum (holotype), 92—O. secretum (Nepal: Parbat). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146230" httpUri="https://zenodo.org/record/10146230/files/figure.png" pageId="46" pageNumber="197">Fig. 90</figureCitation>
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. Body reddish-brown, with slightly paler lateral and basal portions of pronotum, elytra and abdomen; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellow. Punctation of head moderately small and sparse, denser in middle and coarser on infraorbital portions; punctation of pronotum moderately dense, larger and deeper than that of middle portion of head, slightly finer in middle and sparser in mediobasal third; elytra with dense and coarse punctation, slightly larger and deeper than that on pronotum, finer and sparser along suture. Anterior portion of clypeus with fine transverse microreticulation; pronotum and elytra without microsculpture; scutellum with fine isodiametric meshes.
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</paragraph>
|
||
<paragraph id="8B9B0E78540BCC70FF56F8D7FAA8FF57" blockId="46.[151,1437,1230,2013]" lastBlockId="47.[151,1437,150,393]" lastPageId="47" lastPageNumber="198" pageId="46" pageNumber="197">Head 1.2–1.3 times as broad as long, with broad clypeus and slightly convex supra-antennal elevations, lateral portions of clypeus with wide shallow anteriomedian depressions, reaching level of anterior third of eyes. Surface with elongate elevations in middle and slightly stronger longitudinal wrinkles between punctures on infraorbital portions. Anteocellar foveae very deep and wide, oval, short, reaching level of posterior third of eyes. Temples 1.2–1.5 times as long as longitudinal length of eyes. Distance between ocelli distinctly more than twice as long as distance between ocellus and posterior margin of eye. Antennomeres 4–5 distinctly shorter than 3, about as long as broad, 6 longer and broader than 5, 7–8 distinctly broader than 6, 9–10 longer and broader than 8, apical antennomere 1.3 times as long as 10, from about middle gradually narrowed toward rounded or subacute apex.</paragraph>
|
||
<paragraph id="8B9B0E78540ACC70FF56FF07FCA2FE0B" blockId="47.[151,1437,150,393]" pageId="47" pageNumber="198">
|
||
Pronotum moderately narrow, 1.4–1.5 times as broad as long, from widest apical middle gradually narrowed both posteriad and anteriad. Surface of disc with two very deep and wide longitudinal depressions, slightly broadened posteriad toward mediobasal third, and with additional oval shallow or very deep depression in medioapical third. Median surface between outer margins of longitudinal depressions and lateral portions with irregular longidudinal elevations between punctures (
|
||
<typeStatus id="549FB0DA540ACC70FE7EFEB7FD8BFE0B" box="[495,595,367,393]" pageId="47" pageNumber="198" type="holotype">holotype</typeStatus>
|
||
without these elevations).
|
||
</paragraph>
|
||
<caption id="DF5B5EF0540ACC70FF06F9AAFD4DF92C" ID-DOI="http://doi.org/10.5281/zenodo.10146232" ID-Zenodo-Dep="10146232" httpUri="https://zenodo.org/record/10146232/files/figure.png" pageId="47" pageNumber="198" startId="47.[151,264,1650,1674]" targetBox="[154,1433,420,1624]" targetPageId="47" targetType="figure">
|
||
<paragraph id="8B9B0E78540ACC70FF06F9AAFD4DF92C" blockId="47.[151,1436,1650,1711]" pageId="47" pageNumber="198">
|
||
<emphasis id="B950D26A540ACC70FF06F9AAFE8EF908" bold="true" box="[151,342,1650,1674]" pageId="47" pageNumber="198">FIGURES 93–95.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="4C2475FB540ACC70FE76F9ABFD6FF909" authorityName="Shavrin" authorityYear="2023" box="[487,695,1651,1675]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="198" phylum="Arthropoda" rank="species" species="latocavum">
|
||
<emphasis id="B950D26A540ACC70FE76F9ABFD6FF909" box="[487,695,1651,1675]" italics="true" pageId="47" pageNumber="198">Omalium latocavum</emphasis>
|
||
</taxonomicName>
|
||
(holotype): 93—aedeagus, parameral view, 94—aedeagus, lateral view, 95—median lobe, lateral view. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B0E78540ACC70FF56F8D9FC46F8BD" blockId="47.[151,1437,1793,2035]" pageId="47" pageNumber="198">Elytra slightly broader than long, slightly less than twice as long as pronotum, subparallel in middle or slightly broadened posteriad, with slightly concave lateral margins in middle.</paragraph>
|
||
<paragraph id="8B9B0E78540ACC6FFF56F891FAC2FEC3" blockId="47.[151,1437,1793,2035]" lastBlockId="48.[151,1437,151,753]" lastPageId="48" lastPageNumber="199" pageId="47" pageNumber="198">
|
||
Male. Antennomeres 9–10 sligthly transverse. Posterior margin of abdominal tergite VIII straight or rounded. Posterior margin of abdominal sternite VIII rounded. Aedeagus with wide basal portion, gradually narrowed toward middle, with long moderately narrow spear-shaped median lobe, slightly broadened in apical portion, with irregularly crenulate lateral margins and rounded apex, each preapical margins forming tooth indistinctly protruded laterobasad; mediolateral portions with narrow elongate sclerotized accessory plates connected with median and preapical portions of parameres; parameres significantly shorter than apex of median lobe, strongly broadened apically, with slightly asymmetric apical portions and broad apices, each with four moderately long apical setae; internal sac moderately wide and long, with four sclerotized transverse structures in basal portion (
|
||
<figureCitation id="131F12FD5415CC6FFB77FF07FAE2FF7B" box="[1254,1338,223,249]" captionStart="FIGURES 93–95" captionStartId="47.[151,264,1650,1674]" captionTargetBox="[154,1433,420,1624]" captionTargetId="figure-123@47.[151,1436,417,1627]" captionTargetPageId="47" captionText="FIGURES 93–95. Aedeagus of Omalium latocavum (holotype): 93—aedeagus, parameral view, 94—aedeagus, lateral view, 95—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146232" httpUri="https://zenodo.org/record/10146232/files/figure.png" pageId="48" pageNumber="199">Fig. 93</figureCitation>
|
||
). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="131F12FD5415CC6FFE4EFEDBFDEBFE9F" box="[479,563,259,285]" captionStart="FIGURES 93–95" captionStartId="47.[151,264,1650,1674]" captionTargetBox="[154,1433,420,1624]" captionTargetId="figure-123@47.[151,1436,417,1627]" captionTargetPageId="47" captionText="FIGURES 93–95. Aedeagus of Omalium latocavum (holotype): 93—aedeagus, parameral view, 94—aedeagus, lateral view, 95—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146232" httpUri="https://zenodo.org/record/10146232/files/figure.png" pageId="48" pageNumber="199">Fig. 94</figureCitation>
|
||
; apical part of median lobe narrow, inner margin forming distinct hook with subacute hind angle, preapical inner margin crenulate, with rounded and slightly curved apex (
|
||
<figureCitation id="131F12FD5415CC6FFB28FEFFFAD6FEC3" box="[1209,1294,294,321]" captionStart="FIGURES 93–95" captionStartId="47.[151,264,1650,1674]" captionTargetBox="[154,1433,420,1624]" captionTargetId="figure-123@47.[151,1436,417,1627]" captionTargetPageId="47" captionText="FIGURES 93–95. Aedeagus of Omalium latocavum (holotype): 93—aedeagus, parameral view, 94—aedeagus, lateral view, 95—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146232" httpUri="https://zenodo.org/record/10146232/files/figure.png" pageId="48" pageNumber="199">Fig. 95</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9B0E785415CC6FFF56FE93FE04FE2F" blockId="48.[151,1437,151,753]" pageId="48" pageNumber="199">Female. Antennomeres 9–10 less transverse. Posterior margin of abdominal tergite VIII and sternite VIII rounded. Accessory sclerite leaf-shaped, from widest basal portion gradually narrowed toward small rounded apex. Spermatheca not recognized.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF35415CC6FFF56FE6EFC8CFDBF" pageId="48" pageNumber="199" type="diagnosis">
|
||
<paragraph id="8B9B0E785415CC6FFF56FE6EFC8CFDBF" blockId="48.[151,1437,151,753]" pageId="48" pageNumber="199">
|
||
<emphasis id="B950D26A5415CC6FFF56FE6EFE76FE53" bold="true" box="[199,430,438,465]" pageId="48" pageNumber="199">Comparative notes.</emphasis>
|
||
Regarding the general shape of the pronotum gradually narrowed both apicad and posteriad,
|
||
<taxonomicName id="4C2475FB5415CC6FFF06FE03FEE8FE77" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[151,304,475,501]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="199" phylum="Arthropoda" rank="species" species="latocavum" status="sp. n.">
|
||
<emphasis id="B950D26A5415CC6FFF06FE03FF73FE77" box="[151,171,475,501]" italics="true" pageId="48" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="B950D26A5415CC6FFF2AFE03FEE8FE77" box="[187,304,475,501]" italics="true" pageId="48" pageNumber="199">latocavum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A5415CC6FFEA9FE03FEA2FE77" bold="true" box="[312,378,475,501]" pageId="48" pageNumber="199">
|
||
<taxonomicNameLabel id="A2636F115415CC6FFEA9FE03FEA2FE77" box="[312,378,475,501]" pageId="48" pageNumber="199" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C2475FB5415CC6FFD83FE03FD42FE76" authorityName="Cameron" authorityYear="1924" box="[530,666,475,501]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="199" phylum="Arthropoda" rank="species" species="secretum">
|
||
<emphasis id="B950D26A5415CC6FFD83FE03FDFEFE77" box="[530,550,475,501]" italics="true" pageId="48" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="B950D26A5415CC6FFDA7FE03FD42FE76" box="[566,666,475,500]" italics="true" pageId="48" pageNumber="199">secretum</emphasis>
|
||
</taxonomicName>
|
||
, from which it can be distinguished by the slightly larger body, the elytra slightly less than twice as long as pronotum, denser and coarser sculpture of the pronotum and the elytra, and details of the morphology of the aedeagus (see the key above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF35415CC6FFF56FD9EFA82FDE3" box="[199,1370,582,609]" pageId="48" pageNumber="199" type="distribution">
|
||
<paragraph id="8B9B0E785415CC6FFF56FD9EFA82FDE3" blockId="48.[151,1437,151,753]" box="[199,1370,582,609]" pageId="48" pageNumber="199">
|
||
<emphasis id="B950D26A5415CC6FFF56FD9EFEB9FDE2" bold="true" box="[199,353,582,608]" pageId="48" pageNumber="199">Distribution.</emphasis>
|
||
<taxonomicName id="4C2475FB5415CC6FFEF6FD9FFD94FDE3" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[359,588,583,609]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="199" phylum="Arthropoda" rank="species" species="latocavum" status="sp. nov.">
|
||
<emphasis id="B950D26A5415CC6FFEF6FD9FFD94FDE3" box="[359,588,583,609]" italics="true" pageId="48" pageNumber="199">Omalium latocavum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A5415CC6FFDC3FD9FFD4AFDE3" bold="true" box="[594,658,583,609]" pageId="48" pageNumber="199">
|
||
<taxonomicNameLabel id="A2636F115415CC6FFDC3FD9FFD4AFDE3" box="[594,658,583,609]" pageId="48" pageNumber="199" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from the
|
||
<typeStatus id="549FB0DA5415CC6FFC34FD9FFC0DFDE3" box="[933,981,583,609]" pageId="48" pageNumber="199">type</typeStatus>
|
||
locality in central
|
||
<collectingCountry id="F3334EE85415CC6FFB36FD9FFB32FDE3" box="[1191,1258,583,609]" name="Nepal" pageId="48" pageNumber="199">Nepal</collectingCountry>
|
||
(
|
||
<figureCitation id="131F12FD5415CC6FFB68FD9FFA95FDE3" box="[1273,1357,583,609]" captionStart="FIGURE 69" captionStartId="38.[151,250,1110,1134]" captionTargetBox="[151,1435,273,1086]" captionTargetId="figure-65@38.[151,1435,273,1086]" captionTargetPageId="38" captionText="FIGURE 69. Distribution of Omalium in the Himalayan region: O. angustissimum sp. n. (circle), O. kumari sp. n. (triangles), O. latocavum sp. n. (diamond)." figureDoi="http://doi.org/10.5281/zenodo.10146218" httpUri="https://zenodo.org/record/10146218/files/figure.png" pageId="48" pageNumber="199">Fig. 69</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF35415CC6FFF56FDB3FEFEFD2B" pageId="48" pageNumber="199" type="etymology">
|
||
<paragraph id="8B9B0E785415CC6FFF56FDB3FEFEFD2B" blockId="48.[151,1437,151,753]" pageId="48" pageNumber="199">
|
||
<emphasis id="B950D26A5415CC6FFF56FDB3FE96FD07" bold="true" box="[199,334,619,645]" pageId="48" pageNumber="199">Etymology.</emphasis>
|
||
The specific epithet is the Latin
|
||
<emphasis id="B950D26A5415CC6FFD51FDB3FCF3FD07" box="[704,811,619,645]" italics="true" pageId="48" pageNumber="199">latocavus</emphasis>
|
||
, -
|
||
<emphasis id="B950D26A5415CC6FFCD2FDB3FC89FD06" box="[835,849,619,644]" italics="true" pageId="48" pageNumber="199">a</emphasis>
|
||
, -
|
||
<emphasis id="B950D26A5415CC6FFCF9FDB3FC52FD06" box="[872,906,619,644]" italics="true" pageId="48" pageNumber="199">um</emphasis>
|
||
(wide hole). It alludes to the coarse punctation of the elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF35415CC6FFF56FD6AFE0BFD73" pageId="48" pageNumber="199" type="biology_ecology">
|
||
<paragraph id="8B9B0E785415CC6FFF56FD6AFE0BFD73" blockId="48.[151,1437,151,753]" pageId="48" pageNumber="199">
|
||
<emphasis id="B950D26A5415CC6FFF56FD6AFE93FD4E" bold="true" box="[199,331,690,716]" pageId="48" pageNumber="199">Bionomics.</emphasis>
|
||
Specimens were collected at elevations from
|
||
<quantity id="4CDCA39D5415CC6FFCD6FD6BFC2DFD4F" box="[839,1013,691,717]" metricMagnitude="3" metricUnit="m" metricValue="3.8" metricValueMax="4.0" metricValueMin="3.6" pageId="48" pageNumber="199" unit="m" value="3800.0" valueMax="4000.0" valueMin="3600.0">3600 to 4000 m</quantity>
|
||
a.s.l. Specimens were collected under stones along the main ridge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |