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<document id="366A1DA6255C4746FBCA7AC6FF78646A" ID-DOI="10.11646/zootaxa.3949.1.4" ID-GBIF-Dataset="60b71b57-6ebd-42e4-9d9a-3689579bab1b" ID-ISSN="1175-5326" ID-Zenodo-Dep="241799" ID-ZooBank="22266A34-5663-4180-91F3-814AA12D8ABE" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461045206382" checkinUser="plazi" docAuthor="Ariyama, Hiroyuki" docDate="2015" docId="038D5362FF9BFFCCFF7DFC2FFB0BFF1E" docLanguage="en" docName="zt03949p110.pdf" docOrigin="Zootaxa 3949 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraflagitopisa" docType="treatment" docVersion="10" lastPageNumber="92" masterDocId="FFB42B1AFF9AFFCEFFEAFF86FF81FFCB" masterDocTitle="Three new species of the Eriopisa group (Crustacea: Amphipoda: Eriopisidae) from Japan, with the description of a new genus" masterLastPageNumber="110" masterPageNumber="91" pageNumber="92" updateTime="1698576547859" updateUser="plazi">
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<mods:title id="82363F6A09F1D3F00037A2D7064333DE">Three new species of the Eriopisa group (Crustacea: Amphipoda: Eriopisidae) from Japan, with the description of a new genus</mods:title>
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<mods:name id="7B9B9314448D0E5A56256CAFB5D4857C" type="personal">
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<mods:namePart id="973C5B6D0D9458B690515798EFDA374E">Ariyama, Hiroyuki</mods:namePart>
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<mods:typeOfResource id="0DEFE5EBDD35CE680C06F961661E7EC7">text</mods:typeOfResource>
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<mods:title id="1F34669104455D7CA1B022487E280E79">Zootaxa</mods:title>
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<mods:date id="11BBAAE88D46AB23F2522207E5371812">2015</mods:date>
<mods:detail id="65DBD4E79D31181B0F91D4C8E09CC17E" type="volume">
<mods:number id="DD52D6EAE20254ADFD3475DA9953BBED">3949</mods:number>
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<mods:identifier id="64583CE49C10C3C8D49BE77F77934C6E" type="DOI">10.11646/zootaxa.3949.1.4</mods:identifier>
<mods:identifier id="0B01DB040FE5D2FC34C70DE19A601227" type="GBIF-Dataset">60b71b57-6ebd-42e4-9d9a-3689579bab1b</mods:identifier>
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<mods:identifier id="A78260D0A0ABF1BF2F738D18A81C2E6C" type="Zenodo-Dep">241799</mods:identifier>
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<treatment id="038D5362FF9BFFCCFF7DFC2FFB0BFF1E" ID-DOI="http://doi.org/10.5281/zenodo.6108623" ID-GBIF-Taxon="119610329" ID-Zenodo-Dep="6108623" LSID="urn:lsid:plazi:treatment:038D5362FF9BFFCCFF7DFC2FFB0BFF1E" httpUri="http://treatment.plazi.org/id/038D5362FF9BFFCCFF7DFC2FFB0BFF1E" lastPageId="2" lastPageNumber="92" pageId="1" pageNumber="92">
<subSubSection id="C33EB1FFFF9BFFCFFF7DFC2FFE68FC08" box="[151,489,937,963]" pageId="1" pageNumber="92" type="nomenclature">
<paragraph id="8B9BE274FF9BFFCFFF7DFC2FFE68FC08" blockId="1.[151,489,937,963]" box="[151,489,937,963]" pageId="1" pageNumber="92">
<heading id="D0D35518FF9BFFCFFF7DFC2FFE68FC08" bold="true" box="[151,489,937,963]" fontSize="11" level="1" pageId="1" pageNumber="92" reason="1">
<emphasis id="B9503E66FF9BFFCFFF7DFC2FFE68FC08" bold="true" box="[151,489,937,963]" pageId="1" pageNumber="92">
Genus
<taxonomicName id="4C2499F7FF9BFFCFFF04FC2FFE29FC08" ID-CoL="7PRTL" box="[238,424,937,963]" class="Malacostraca" family="Eriopisidae" genus="Paraflagitopisa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9503E66FF9BFFCFFF04FC2FFE29FC08" bold="true" box="[238,424,937,963]" italics="true" pageId="1" pageNumber="92">Paraflagitopisa</emphasis>
</taxonomicName>
, new
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33EB1FFFF9BFFCFFF7DFC69FCD4FBC3" box="[151,853,1007,1032]" pageId="1" pageNumber="92" type="materials_examined">
<paragraph id="8B9BE274FF9BFFCFFF7DFC69FCD4FBC3" blockId="1.[151,1437,1007,2005]" box="[151,853,1007,1032]" pageId="1" pageNumber="92">
<emphasis id="B9503E66FF9BFFCFFF7DFC69FEB3FBC3" bold="true" box="[151,306,1007,1032]" pageId="1" pageNumber="92">
<typeStatus id="549F5CD6FF9BFFCFFF7DFC69FF53FBC3" box="[151,210,1007,1032]" pageId="1" pageNumber="92">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C2499F7FF9BFFCFFED3FC69FDD5FBC3" box="[313,596,1007,1032]" class="Malacostraca" family="Eriopisidae" genus="Paraflagitopisa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="92" phylum="Arthropoda" rank="species" species="excavata" status="sp. nov.">
<emphasis id="B9503E66FF9BFFCFFED3FC69FDD5FBC3" box="[313,596,1007,1032]" italics="true" pageId="1" pageNumber="92">Paraflagitopisa excavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A263831DFF9BFFCFFDB0FC69FD32FBC3" box="[602,691,1007,1032]" pageId="1" pageNumber="92" rank="species">
<emphasis id="B9503E66FF9BFFCFFDB0FC69FD32FBC3" bold="true" box="[602,691,1007,1032]" pageId="1" pageNumber="92">sp. nov.</emphasis>
</taxonomicNameLabel>
by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C33EB1FFFF9BFFCFFF2DFB92FD9DF91C" pageId="1" pageNumber="92" type="diagnosis">
<paragraph id="8B9BE274FF9BFFCFFF2DFB92FD9DF91C" blockId="1.[151,1437,1007,2005]" pageId="1" pageNumber="92">
<emphasis id="B9503E66FF9BFFCFFF2DFB92FEC0FBE6" bold="true" box="[199,321,1044,1069]" pageId="1" pageNumber="92">Diagnosis.</emphasis>
<emphasis id="B9503E66FF9BFFCFFEBBFB92FE0AFBE6" box="[337,395,1044,1069]" italics="true" pageId="1" pageNumber="92">Body</emphasis>
depressed laterally, smooth; urosomites free.
<emphasis id="B9503E66FF9BFFCFFC28FB92FC7FFBE6" box="[962,1022,1044,1069]" italics="true" pageId="1" pageNumber="92">Head</emphasis>
longer than pereonite 1, rostrum obsolescent, eyes absent; lateral cephalic lobe rounded without incision, anteroventral sinus indistinct.
<emphasis id="B9503E66FF9BFFCFFAC2FBBEFA1DFB9B" box="[1320,1436,1080,1104]" italics="true" pageId="1" pageNumber="92">Antenna 1</emphasis>
long; peduncular articles 12 nearly subequal long, article 3 short; accessory flagellum short, 2-articulated; primary flagellum with many articles.
<emphasis id="B9503E66FF9BFFCFFE1AFB06FDE5FB53" box="[496,612,1152,1176]" italics="true" pageId="1" pageNumber="92">Antenna 2</emphasis>
short, slender; flagellum not fused, slightly longer than last peduncular article.
<emphasis id="B9503E66FF9BFFCFFF07FB23FEDBFB76" box="[237,346,1188,1213]" italics="true" pageId="1" pageNumber="92">Upper lip</emphasis>
entire.
<emphasis id="B9503E66FF9BFFCFFE5EFB22FD9EFB76" box="[436,543,1188,1213]" italics="true" pageId="1" pageNumber="92">Mandible</emphasis>
with toothed incisor, lacinia mobilis, accessory blades and strong molar; palp with 3 articles, article 1 short, article 2 longer than article 3.
<emphasis id="B9503E66FF9BFFCFFCDCFB4FFC23FB2B" box="[822,930,1223,1248]" italics="true" pageId="1" pageNumber="92">Lower lip</emphasis>
with small inner lobes.
<emphasis id="B9503E66FF9BFFCFFB46FB41FA99FB2B" box="[1196,1304,1223,1248]" italics="true" pageId="1" pageNumber="92">Maxilla 1</emphasis>
, inner plate triangular, with many distolateral setae; outer plate with 9 robust setae apically; palp 2-articulated, with distal robust setae.
<emphasis id="B9503E66FF9BFFCFFED2FA89FE2AFAE3" box="[312,427,1295,1320]" italics="true" pageId="1" pageNumber="92">Maxilla 2</emphasis>
, inner plate with dorsal oblique row of setae, terminal ends of both plates setose.
<emphasis id="B9503E66FF9BFFCFFF7DFAB2FE91FA86" box="[151,272,1332,1357]" italics="true" pageId="1" pageNumber="92">Maxilliped</emphasis>
, inner plate truncate, with a few robust setae distally; outer plate bearing row of robust setae distomedially; palp 4-articulated, article 4 with short nail.
<emphasis id="B9503E66FF9BFFCFFCD1FADEFC02FAA4" box="[827,899,1368,1391]" italics="true" pageId="1" pageNumber="92">Coxae</emphasis>
short, weakly overlapping; coxae 12 slightly produced forward, coxae 56 lobed, coxa 7 smallest; gills large, present on coxae 26.
<emphasis id="B9503E66FF9BFFCFFB87FAFAFA80FA5F" box="[1133,1281,1404,1429]" italics="true" pageId="1" pageNumber="92">
Gnathopod
<quantity id="4CDC4F91FF9BFFCFFB19FAFAFA80FA5F" box="[1267,1281,1404,1428]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="92" unit="in" value="1.0">1</quantity>
</emphasis>
in both sexes similar, small, subchelate; carpus longer than propodus, palm transverse.
<emphasis id="B9503E66FF9BFFCFFBE2FA19FB23FA73" box="[1032,1186,1439,1464]" italics="true" pageId="1" pageNumber="92">
Gnathopod
<quantity id="4CDC4F91FF9BFFCFFB7EFA26FB23FA73" box="[1172,1186,1440,1464]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="92" unit="in" value="2.0">2</quantity>
</emphasis>
in both sexes large, subchelate; propodus dilated, palm oblique; female gnathopod 2 smaller than that of male.
<emphasis id="B9503E66FF9BFFCFFB72FA42FAC0FA17" box="[1176,1345,1476,1501]" italics="true" pageId="1" pageNumber="92">Pereopods 34</emphasis>
similar, slender; bases linear.
<emphasis id="B9503E66FF9BFFCFFE61FA61FDB2FA34" box="[395,563,1511,1536]" italics="true" pageId="1" pageNumber="92">Pereopods 57</emphasis>
, bases similarly dilated, meri not expanded.
<emphasis id="B9503E66FF9BFFCFFBD9FA6EFB28FA34" box="[1075,1193,1512,1535]" italics="true" pageId="1" pageNumber="92">Oostegites</emphasis>
of female present on coxae 25.
<emphasis id="B9503E66FF9BFFCFFEF4F98AFD8EF9EF" box="[286,527,1548,1573]" italics="true" pageId="1" pageNumber="92">Pleonal epimera 13</emphasis>
, posteroventral corners quadrate, ventral margins scarcely setose; first pleonal segment without gills.
<emphasis id="B9503E66FF9BFFCFFE77F9A9FDB9F983" box="[413,568,1583,1608]" italics="true" pageId="1" pageNumber="92">Pleopods 13</emphasis>
, peduncles long, each with 2 coupling hooks; inner rami longer than outer.
<emphasis id="B9503E66FF9BFFCFFF7DF9D2FEA8F9A7" box="[151,297,1620,1645]" italics="true" pageId="1" pageNumber="92">Uropods 12</emphasis>
biramous, peduncles each with dorsal and distal robust setae, peduncle of uropod 1 with ventrofacial robust setae; both rami bearing marginal and distal robust setae.
<emphasis id="B9503E66FF9BFFCFFC86F9F1FC59F95B" box="[876,984,1655,1680]" italics="true" pageId="1" pageNumber="92">Uropod 3</emphasis>
very long, biramous; inner ramus short, slender; outer ramus long, slender, 2-articulated, length of article 2 subequal to article 1.
<emphasis id="B9503E66FF9BFFCFFB75F91AFB64F97E" box="[1183,1253,1692,1717]" italics="true" pageId="1" pageNumber="92">Telson</emphasis>
deeply incised, with lateral and distal robust setae.
</paragraph>
</subSubSection>
<subSubSection id="C33EB1FFFF9BFFCFFF2DF962FBEBF937" box="[199,1130,1764,1789]" pageId="1" pageNumber="92" type="etymology">
<paragraph id="8B9BE274FF9BFFCFFF2DF962FBEBF937" blockId="1.[151,1437,1007,2005]" box="[199,1130,1764,1789]" pageId="1" pageNumber="92">
<emphasis id="B9503E66FF9BFFCFFF2DF962FECFF936" bold="true" box="[199,334,1764,1789]" pageId="1" pageNumber="92">Etymology.</emphasis>
From the Latin
<emphasis id="B9503E66FF9BFFCFFDE9F963FDB9F937" box="[515,568,1765,1788]" italics="true" pageId="1" pageNumber="92">para</emphasis>
(= beside) and
<taxonomicName id="4C2499F7FF9BFFCFFD0CF962FCDFF936" box="[742,862,1764,1789]" class="Malacostraca" family="Melitidae" genus="Flagitopisa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B9503E66FF9BFFCFFD0CF962FCDFF936" box="[742,862,1764,1789]" italics="true" pageId="1" pageNumber="92">flagitopisa</emphasis>
</taxonomicName>
(closely related genus).
</paragraph>
</subSubSection>
<subSubSection id="C33EB1FFFF9BFFCCFF2DF881FB0BFF1E" lastPageId="2" lastPageNumber="93" pageId="1" pageNumber="92" type="discussion">
<paragraph id="8B9BE274FF9BFFCCFF2DF881FB0BFF1E" blockId="1.[151,1437,1007,2005]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="93" pageId="1" pageNumber="92">
<emphasis id="B9503E66FF9BFFCFFF2DF881FEBAF8EB" bold="true" box="[199,315,1799,1824]" pageId="1" pageNumber="92">Remarks.</emphasis>
<taxonomicName id="4C2499F7FF9BFFCFFEA8F881FE73F8EB" box="[322,498,1799,1824]" class="Malacostraca" family="Eriopisidae" genus="Paraflagitopisa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9503E66FF9BFFCFFEA8F881FE73F8EB" box="[322,498,1799,1824]" italics="true" pageId="1" pageNumber="92">Paraflagitopisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A263831DFF9BFFCFFE10F881FDE5F8EB" box="[506,612,1799,1824]" pageId="1" pageNumber="92" rank="genus">
<emphasis id="B9503E66FF9BFFCFFE10F881FDE5F8EB" bold="true" box="[506,612,1799,1824]" pageId="1" pageNumber="92">gen. nov.</emphasis>
</taxonomicNameLabel>
is characterized by (1) entire lateral cephalic lobe, (2) unfused flagellum of antenna 2, (3) 3-articulated mandibular palp, (4) carpus of gnathopod 1 longer than propodus, (5) transverse palm of gnathopod 1, and (6) slender outer ramus of uropod 3 with long second article. This new genus share these characters with
<taxonomicName id="4C2499F7FF9BFFCFFE82F8F2FE68F846" box="[360,489,1908,1933]" class="Malacostraca" family="Melitidae" genus="Flagitopisa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B9503E66FF9BFFCFFE82F8F2FE68F846" box="[360,489,1908,1933]" italics="true" pageId="1" pageNumber="92">Flagitopisa</emphasis>
</taxonomicName>
; however,
<taxonomicName id="4C2499F7FF9BFFCFFD6CF8F2FC86F846" box="[646,775,1908,1933]" class="Malacostraca" family="Melitidae" genus="Flagitopisa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B9503E66FF9BFFCFFD6CF8F2FC86F846" box="[646,775,1908,1933]" italics="true" pageId="1" pageNumber="92">Flagitopisa</emphasis>
</taxonomicName>
can be distinguished by the following characters (
<taxonomicName id="4C2499F7FF9BFFCFFF4AF811FED1F87B" box="[160,336,1943,1968]" class="Malacostraca" family="Eriopisidae" genus="Paraflagitopisa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9503E66FF9BFFCFFF4AF811FED1F87B" box="[160,336,1943,1968]" italics="true" pageId="1" pageNumber="92">Paraflagitopisa</emphasis>
</taxonomicName>
in parentheses): (1) marked anteroventral sinus of head (indistinct), (2) article 2 of mandibular palp nearly as long as article 3 (longer than article 3), (3) dilated bases of pereopods 34 (undilated), (4) first pleonal segment with pair of gills (without gills), and (5) scale-like inner ramus of uropod 3 (slender) [see
<bibRefCitation id="EFB59F85FF98FFCCFA84FF1EFECEFF1F" author="Karaman" pageId="2" pageNumber="93" refString="Karaman, G. S. (1984) Revision of Eriopisa - complex of genera (Gammaridea) (Contribution to the knowledge of the Amphipoda 139). Poljoprivreda i Sumarstvo, Titograd, 30, 39 - 72." type="journal article" year="1984">G.S. Karaman (1984)</bibRefCitation>
and
<bibRefCitation id="EFB59F85FF98FFCCFE6FFF3AFDF4FF1E" box="[389,629,188,213]" pageId="2" pageNumber="93" refString="Sawicki, T. R., Holsinger, J. R. &amp; Sket, B. (2005) Redescription of the subterranean amphipod crustacean Flagitopisa philippensis (Hadzioidea: Melitidae), with notes on its unique morphology and clarification of the taxonomic status of Psammogammarus fluviatilis. Raffles Bulletin of Zoology, 53, 59 - 68." type="journal article">
Sawicki
<emphasis id="B9503E66FF98FFCCFE0DFF3BFD9EFF1E" box="[487,543,188,213]" italics="true" pageId="2" pageNumber="93">et al.</emphasis>
(2005)
</bibRefCitation>
]. In addition,
<taxonomicName id="4C2499F7FF98FFCCFCFAFF3AFC10FF1E" box="[784,913,188,213]" class="Malacostraca" family="Melitidae" genus="Flagitopisa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B9503E66FF98FFCCFCFAFF3AFC10FF1E" box="[784,913,188,213]" italics="true" pageId="2" pageNumber="93">Flagitopisa</emphasis>
</taxonomicName>
is a freshwater genus.
</paragraph>
</subSubSection>
</treatment>
</document>