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<document id="ED6550AD21EF9C39D134813CE6706D56" ID-DOI="10.5852/ejt.2024.922.2443" ID-ISSN="2118-9773" ID-Zenodo-Dep="10715279" ID-ZooBank="CEE6866E-B1DD-4BF9-8461-FA99D0A9FE9A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709025066059" checkinUser="plazi" docAuthor="Orsetti, Artur &amp; Lopes-Andrade, Cristiano" docDate="2024" docId="038C9540AB2BFFB19359FB952800A1A3" docLanguage="en" docName="EJT.2024.922.1-61.pdf" docOrigin="European Journal of Taxonomy 922" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2443/10823" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pelecium helenae Straneo &amp; Ball 1989" docType="treatment" docVersion="2" lastPageNumber="36" masterDocId="FFB5ED38AB0AFF929177FFFD2D0AA24C" masterDocTitle="A contribution to the taxonomy of the genus Pelecium Kirby (Coleoptera: Carabidae: Peleciini)" masterLastPageNumber="61" masterPageNumber="1" pageNumber="34" updateTime="1709025674441" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DF8E939856518A1192AAE874C589E8E8">A contribution to the taxonomy of the genus Pelecium Kirby (Coleoptera: Carabidae: Peleciini)</mods:title>
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<mods:namePart id="21F2776DAE554AD111E7C8E412D39DEA">Orsetti, Artur</mods:namePart>
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<mods:affiliation id="0AF15DBA6757786CBC38B5AEFE393674">Pós-Graduação em Ecologia, Departamento de Biologia Geral, Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil. &amp; Laboratório de Sistemática e Biologia de Coleoptera, Departamento de Biologia Animal, Universidade Federal de Viçosa, Viçosa, Minas Gerais, Brazil.</mods:affiliation>
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<mods:namePart id="9FDB954F64FF28420246B34742010932">Lopes-Andrade, Cristiano</mods:namePart>
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<mods:affiliation id="2F213F915C9B91EFA7B60C1BE4164823">Laboratório de Sistemática e Biologia de Coleoptera, Departamento de Biologia Animal, Universidade Federal de Viçosa, Viçosa, Minas Gerais, Brazil.</mods:affiliation>
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<treatment id="038C9540AB2BFFB19359FB952800A1A3" ID-DOI="http://doi.org/10.5281/zenodo.10715338" ID-Zenodo-Dep="10715338" LSID="urn:lsid:plazi:treatment:038C9540AB2BFFB19359FB952800A1A3" httpUri="http://treatment.plazi.org/id/038C9540AB2BFFB19359FB952800A1A3" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
<subSubSection id="C33F77DDAB2BFFB39359FB95290CA6CF" box="[558,1030,1128,1155]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8B9A2456AB2BFFB39359FB95290CA6CF" blockId="33.[558,1030,1128,1193]" box="[558,1030,1128,1155]" pageId="33" pageNumber="34">
<heading id="D0D2933AAB2BFFB39359FB95290CA6CF" box="[558,1030,1128,1155]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="34" reason="2">
<taxonomicName id="4C255FD5AB2BFFB39359FB95290CA6CF" ID-CoL="6TZ92" authority="Straneo &amp; Ball, 1989" authorityName="Straneo &amp; Ball" authorityYear="1989" box="[558,1030,1128,1155]" class="Insecta" family="Carabidae" genus="Pelecium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="helenae">
<emphasis id="B951F844AB2BFFB39359FB952E0FA6CF" bold="true" box="[558,773,1128,1155]" italics="true" pageId="33" pageNumber="34">Pelecium helenae</emphasis>
<bibRefCitation id="EFB459A7AB2BFFB3927BFB94290CA6CF" author="Straneo S. L. &amp; Ball G. B." box="[780,1030,1129,1155]" pageId="33" pageNumber="34" pagination="73 - 178" refId="ref27763" refString="Straneo S. L. &amp; Ball G. B. 1989. Synopsis of the genera and subgenera of the tribe Peleciini, and revision of the Neotropical and Oriental species (Coleoptera: Carabidae). Insecta Mundi 3 (2): 73 - 178." type="journal article" year="1989">Straneo &amp; Ball, 1989</bibRefCitation>
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<subSubSection id="C33F77DDAB2BFFB39317FB732CCBA6A7" pageId="33" pageNumber="34" type="description">
<paragraph id="8B9A2456AB2BFFB39317FB732ED9A6E4" blockId="33.[558,1030,1128,1193]" box="[608,979,1166,1193]" pageId="33" pageNumber="34">
<figureCitation id="131E38D3AB2BFFB39317FB732FEFA6E4" box="[608,741,1166,1192]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[302,1289,266,1760]" captionTargetId="figure-126@34.[296,1295,259,1764]" captionTargetPageId="34" captionText="Fig. 9. A–H. Pelecium helenae Straneo &amp; Ball, 1989, male from Piraquara, Paraná, Brazil (A–G) (CERPE) and female from Mogi das Cruzes, São Paulo, Brazil (H) (CEMT). A. Dorsal view. B. Detail of head. C–D. Median lobe. E. Tergite IX. F. Left paramere. G. Right paramere. H. Genitalia. – I–P. Pelecium laeve Chaudoir, 1854, male from Porto Alegre, Rio Grande do Sul, Brazil (I–O) (FZRBS) and female from Porto Alegre, Rio Grande do Sul, Brazil (P) (FZRBS). I. Dorsal view. J. Detail of head. K–L. Median lobe. M. Tergite IX. N. Left paramere. O. Right paramere. P. Genitalia. Scale bars: A–B, I–J = 1 mm; C–H, K–P = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10715310" httpUri="https://zenodo.org/record/10715310/files/figure.png" pageId="33" pageNumber="34">Figs 9A–H</figureCitation>
,
<figureCitation id="131E38D3AB2BFFB39386FB732E42A6E5" box="[753,840,1166,1193]" captionStart="Fig" captionStartId="45.[189,232,1891,1917]" captionTargetBox="[304,1287,268,1853]" captionTargetId="figure-61@45.[298,1290,265,1854]" captionTargetPageId="45" captionText="Fig. 15. Types of Pelecium Kirby, 1817, dorsal view. A. Pelecium fulgidum Straneo, 1962, allotype (MCSN), photograph by Michele Zilioli. B. Pelecium helenae Straneo &amp; Ball, 1989, holotype (MZSP), photograph by Guilherme Ide. C. Pelecium helenae, paratype (MCSN), photograph by Michele Zilioli. D. Pelecium helenae, allotype (MCSN), photograph by Michele Zilioli. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10715326" httpUri="https://zenodo.org/record/10715326/files/figure.png" pageId="33" pageNumber="34">15B–D</figureCitation>
,
<figureCitation id="131E38D3AB2BFFB39223FB732EA0A6E4" box="[852,938,1166,1192]" captionStart="Fig" captionStartId="51.[189,232,1711,1737]" captionTargetBox="[330,1265,266,1656]" captionTargetId="figure-132@51.[320,1268,249,1671]" captionTargetPageId="51" captionText="Fig. 21.Labels of type specimens of some species of Pelecium Kirby, 1817. A. Pelecium cyanipes Kirby, 1817, holotype (BMNH), photograph by Keita Matsumoto. B. Pelecium drakei Quedenfeldt, 1890, lectotype (MUB), photograph by Bernd Jaeger. C. Pelecium fulgidum Straneo, 1962, holotype (MCSN), photograph by Michele Zilioli. D. Pelecium fulgidum, allotype (MCSN), photograph by Michele Zilioli. E. Pelecium helenae Straneo &amp; Ball, 1989, holotype (MZSP), photograph by Guilherme Ide.F. Pelecium helenae, paratype (MCSN), photograph by Michele Zilioli. G. Pelecium helenae, allotype (MCSN), photograph by Michele Zilioli. H. Pelecium laeve Chaudoir, 1854, holotype (MNHN), photograph by Azadeh Taghavian. I. Pelecium longicolle Straneo, 1953, allotype (MCSN), photograph by Michele Zilioli. J. Pelecium negrei Straneo, 1962, holotype (MCSN), photograph by Michele Zilioli." figureDoi="http://doi.org/10.5281/zenodo.10715347" httpUri="https://zenodo.org/record/10715347/files/figure.png" pageId="33" pageNumber="34">21E–G</figureCitation>
,
<figureCitation id="131E38D3AB2BFFB392C1FB732ED9A6E4" box="[950,979,1166,1192]" captionStart="Fig" captionStartId="56.[189,232,1890,1916]" captionTargetBox="[189,1399,771,1851]" captionTargetId="figure-209@56.[189,1399,771,1851]" captionTargetPageId="56" captionText="Fig. 26. Map highlighting part of South America with the known occurrences, including new records, of Pelecium atroviolaceum Straneo &amp; Ball, 1989, P. bolivianum Straneo &amp; Ball, 1989, P. cyanipes Kirby, 1817, P. drakei Quedenfeldt, 1890, P. fulgidum Straneo, 1962, P. helenae Straneo &amp; Ball, 1989 and P. nicki Straneo, 1955." figureDoi="http://doi.org/10.5281/zenodo.10715361" httpUri="https://zenodo.org/record/10715361/files/figure.png" pageId="33" pageNumber="34">26</figureCitation>
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<heading id="D0D2933AAB2BFFB391CAFB2C2CCBA6A7" bold="true" box="[189,449,1233,1259]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B951F844AB2BFFB391CAFB2C2CCBA6A7" bold="true" box="[189,449,1233,1259]" pageId="33" pageNumber="34">Specimens examined</emphasis>
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</paragraph>
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•
<specimenCount id="9D23EFDFAB2BFFB39030FAFF2C9BA751" box="[327,401,1282,1309]" count="2" pageId="33" pageNumber="34" type="male">2 ♂♂</specimenCount>
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<collectingMunicipality id="6BFEBE2CAB2BFFB39373FAFF2F72A750" box="[516,632,1282,1308]" pageId="33" pageNumber="34">Piraquara</collectingMunicipality>
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;
<date id="FF9B0296AB2BFFB392FFFAFF2921A750" box="[904,1067,1282,1308]" pageId="33" pageNumber="34" value="2011-10-19">
<collectingDate id="EFDFFB7EAB2BFFB392FFFAFF2921A750" box="[904,1067,1282,1308]" pageId="33" pageNumber="34" value="2011-10-19">19 Oct. 2011</collectingDate>
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leg.;
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•
<materialsCitation id="3B4D2E0BAB2BFFB3904CFADA2C12A72B" collectingDate="2006-04-09" collectionCode="CERPE" collectorName="Grossi &amp; Parizotto" country="Brazil" location="Estr. Castro-Tibagi" municipality="Castro" pageId="33" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Parana">
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<collectingMunicipality id="6BFEBE2CAB2BFFB39094FADA2F39A70D" box="[483,563,1319,1345]" pageId="33" pageNumber="34">Castro</collectingMunicipality>
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km 15;
<date id="FF9B0296AB2BFFB392E1FADA2927A70D" box="[918,1069,1319,1345]" pageId="33" pageNumber="34" value="2006-04-09">
<collectingDate id="EFDFFB7EAB2BFFB392E1FADA2927A70D" box="[918,1069,1319,1345]" pageId="33" pageNumber="34" value="2006-04-09">9 Apr. 2006</collectingDate>
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leg.;
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<materialsCitation id="3B4D2E0BAB2BFFB39047FAB029FDA72B" box="[304,1271,1356,1383]" collectingDate="2005-11-30" collectionCode="CERPE" collectorName="P. Grossi" country="Brazil" location="Mananciais da Serra" municipality="Piraquara" pageId="33" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Parana">
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<collectingMunicipality id="6BFEBE2CAB2BFFB390BEFAB12F37A72A" box="[457,573,1356,1382]" pageId="33" pageNumber="34">Piraquara</collectingMunicipality>
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<date id="FF9B0296AB2BFFB39231FAB02EEDA72B" box="[838,999,1356,1383]" pageId="33" pageNumber="34" value="2005-11-30">
<collectingDate id="EFDFFB7EAB2BFFB39231FAB02EEDA72B" box="[838,999,1356,1383]" pageId="33" pageNumber="34" value="2005-11-30">30 Nov. 2005</collectingDate>
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leg.;
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•
<materialsCitation id="3B4D2E0BAB2BFFB39467FAB029E7A7C0" collectingDate="2015-02-24" collectingDateMax="2015-02-28" collectingDateMin="2015-02-24" collectionCode="R, V, CEMT" collectorName="Mogi &amp; Nunes" country="Brazil" county="Cruzes" location="Neblinas" municipality="Parque" pageId="33" pageNumber="34" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo">
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das
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<location id="8EFA728DAB2BFFB3930FFA8F2FEEA7C0" LSID="urn:lsid:plazi:treatment:038C9540AB2BFFB19359FB952800A1A3:8EFA728DAB2BFFB3930FFA8F2FEEA7C0" box="[632,740,1394,1420]" country="Brazil" county="Cruzes" municipality="Parque" name="Neblinas" pageId="33" pageNumber="34" stateProvince="Sao Paulo">Neblinas</location>
;
<date id="FF9B0296AB2BFFB39385FA8F2ECAA7C1" box="[754,960,1394,1421]" pageId="33" pageNumber="34" value="2015-02-24" valueMax="2015-02-28" valueMin="2015-02-24">
<collectingDate id="EFDFFB7EAB2BFFB39385FA8F2ECAA7C1" box="[754,960,1394,1421]" pageId="33" pageNumber="34" value="2015-02-24" valueMax="2015-02-28" valueMin="2015-02-24">24–28 Feb. 2015</collectingDate>
</date>
;
<collectionCode id="ED34BC93AB2BFFB392B9FA8F2EE9A7C0" box="[974,995,1394,1420]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="33" pageNumber="34">R</collectionCode>
.
<collectionCode id="ED34BC93AB2BFFB39290FA8F2EF4A7C0" box="[999,1022,1394,1420]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="33" pageNumber="34" type="Museum">V</collectionCode>
.
<collectorName id="26D04180AB2BFFB3957CFA8F295CA7C0" box="[1035,1110,1394,1420]" pageId="33" pageNumber="34">Nunes</collectorName>
leg.;
<collectionCode id="ED34BC93AB2BFFB395EBFA8F29E7A7C0" box="[1180,1261,1394,1420]" pageId="33" pageNumber="34">CEMT</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B4D2E0BAB2BFFB3947CFA8F2868A7FD" collectingDate="2007-01" collectionCode="BELLO, CAMB" collectorName="M. Huhara" country="Brazil" location="Est. Biologica Paranapiacaba" pageId="33" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo">
<specimenCount id="9D23EFDFAB2BFFB3947CFA8F2836A7C0" box="[1291,1340,1394,1420]" count="1" pageId="33" pageNumber="34" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49E1EAB4AB2BFFB3943DFA8F2C09A7FD" country="Brazil" name="Sao Paulo" pageId="33" pageNumber="34">São Paulo</collectingRegion>
,
<location id="8EFA728DAB2BFFB39079FA652F63A7FD" LSID="urn:lsid:plazi:treatment:038C9540AB2BFFB19359FB952800A1A3:8EFA728DAB2BFFB39079FA652F63A7FD" box="[270,617,1431,1458]" country="Brazil" name="Est. Biologica Paranapiacaba" pageId="33" pageNumber="34" stateProvince="Sao Paulo">Est. Biologica Paranapiacaba</location>
;
<date id="FF9B0296AB2BFFB39302FA652FE3A7FD" box="[629,745,1431,1458]" pageId="33" pageNumber="34" value="2007-01">
<collectingDate id="EFDFFB7EAB2BFFB39302FA652FE3A7FD" box="[629,745,1431,1458]" pageId="33" pageNumber="34" value="2007-01">Jan. 2007</collectingDate>
</date>
;
<collectorName id="26D04180AB2BFFB39382FA652E7EA7FD" box="[757,884,1431,1458]" pageId="33" pageNumber="34">M. Huhara</collectorName>
leg.; “\\ Coleção A. M.
<collectionCode id="ED34BC93AB2BFFB395FAFA6A29F8A7FD" box="[1165,1266,1431,1457]" pageId="33" pageNumber="34">BELLO</collectionCode>
”;
<collectionCode id="ED34BC93AB2BFFB3947FFA6A2868A7FD" box="[1288,1378,1431,1457]" pageId="33" pageNumber="34">CAMB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A2456AB2BFFB391CAFA262C09A7B9" blockId="33.[189,259,1499,1525]" box="[189,259,1499,1525]" pageId="33" pageNumber="34">
<heading id="D0D2933AAB2BFFB391CAFA262C09A7B9" bold="true" box="[189,259,1499,1525]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B951F844AB2BFFB391CAFA262C09A7B9" bold="true" box="[189,259,1499,1525]" pageId="33" pageNumber="34">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A2456AB2BFFB391CAF9F1283AA566" blockId="33.[188,1399,1547,1834]" pageId="33" pageNumber="34">
Most known specimens of
<taxonomicName id="4C255FD5AB2BFFB39081F9F12F64A469" authorityName="Straneo &amp; Ball" authorityYear="1989" box="[502,622,1547,1573]" class="Insecta" family="Carabidae" genus="Pelecium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="helenae">
<emphasis id="B951F844AB2BFFB39081F9F12F64A469" box="[502,622,1547,1573]" italics="true" pageId="33" pageNumber="34">P. helenae</emphasis>
</taxonomicName>
are from Southeast
<collectingCountry id="F33264C6AB2BFFB3922EF9F62EAEA469" box="[857,932,1547,1573]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
, including the
<typeStatus id="549E9AF4AB2BFFB39538F9F629BCA469" box="[1103,1206,1547,1573]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
and the
<typeStatus id="549E9AF4AB2BFFB3946FF9F6287DA469" box="[1304,1399,1547,1573]" pageId="33" pageNumber="34" type="allotype">allotype</typeStatus>
(both from the state of
<collectingRegion id="49E1EAB4AB2BFFB390B2F9CD2F35A406" box="[453,575,1584,1610]" country="Brazil" name="Sao Paulo" pageId="33" pageNumber="34">São Paulo</collectingRegion>
), and
<specimenCount id="9D23EFDFAB2BFFB393F4F9CD2E3EA407" box="[643,820,1584,1611]" count="8" pageId="33" pageNumber="34" type="generic" typeStatus="paratype">eight paratypes</specimenCount>
(states of
<collectingRegion id="49E1EAB4AB2BFFB392DCF9CD294DA406" box="[939,1095,1584,1610]" country="Brazil" name="Minas Gerais" pageId="33" pageNumber="34">Minas Gerais</collectingRegion>
and
<collectingRegion id="49E1EAB4AB2BFFB39509F9CD29F2A406" box="[1150,1272,1584,1610]" country="Brazil" name="Sao Paulo" pageId="33" pageNumber="34">São Paulo</collectingRegion>
). Only
<specimenCount id="9D23EFDFAB2BFFB3943BF9CC2C2BA43C" count="1" pageId="33" pageNumber="34" type="generic" typeStatus="paratype">one paratype</specimenCount>
was collected in the state of
<collectingRegion id="49E1EAB4AB2BFFB39305F9AB2FA3A43C" box="[626,681,1622,1648]" country="Brazil" name="Para" pageId="33" pageNumber="34">Pará</collectingRegion>
, North
<collectingCountry id="F33264C6AB2BFFB39388F9A82E40A423" box="[767,842,1621,1647]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
. The new records are from the Araucaria Forest province, while the
<typeStatus id="549E9AF4AB2BFFB390D9F9862F07A4D9" box="[430,525,1659,1685]" pageId="33" pageNumber="34" type="allotype">allotype</typeStatus>
is from the Atlantic province and the
<typeStatus id="549E9AF4AB2BFFB392A8F9862945A4D9" box="[991,1103,1659,1685]" pageId="33" pageNumber="34" type="paratype">paratypes</typeStatus>
from the state of
<collectingRegion id="49E1EAB4AB2BFFB3945AF9862C00A4F6" country="Brazil" name="Minas Gerais" pageId="33" pageNumber="34">Minas Gerais</collectingRegion>
are from the
<collectingRegion id="49E1EAB4AB2BFFB390DDF95D2CF0A4F6" box="[426,506,1696,1722]" country="Brazil" name="Parana" pageId="33" pageNumber="34">Paraná</collectingRegion>
Forest province, but all the three provinces belong to the
<collectingRegion id="49E1EAB4AB2BFFB395D0F95D29FDA4F6" box="[1191,1271,1696,1722]" country="Brazil" name="Parana" pageId="33" pageNumber="34">Paraná</collectingRegion>
dominion.
<typeStatus id="549E9AF4AB2BFFB391C9F93B2C3AA4AC" box="[190,304,1734,1760]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
from the state of
<collectingRegion id="49E1EAB4AB2BFFB39372F9382F33A493" box="[517,569,1733,1759]" country="Brazil" name="Para" pageId="33" pageNumber="34">Pará</collectingRegion>
are from the
<collectingRegion id="49E1EAB4AB2BFFB393AAF9382E8AA493" box="[733,896,1733,1759]" country="Brazil" name="Para" pageId="33" pageNumber="34">Pará province</collectingRegion>
of the Brazilian boreal dominion. The red label (
<figureCitation id="131E38D3AB2BFFB39071F9172C67A549" box="[262,365,1770,1797]" captionStart="Fig" captionStartId="51.[189,232,1711,1737]" captionTargetBox="[330,1265,266,1656]" captionTargetId="figure-132@51.[320,1268,249,1671]" captionTargetPageId="51" captionText="Fig. 21.Labels of type specimens of some species of Pelecium Kirby, 1817. A. Pelecium cyanipes Kirby, 1817, holotype (BMNH), photograph by Keita Matsumoto. B. Pelecium drakei Quedenfeldt, 1890, lectotype (MUB), photograph by Bernd Jaeger. C. Pelecium fulgidum Straneo, 1962, holotype (MCSN), photograph by Michele Zilioli. D. Pelecium fulgidum, allotype (MCSN), photograph by Michele Zilioli. E. Pelecium helenae Straneo &amp; Ball, 1989, holotype (MZSP), photograph by Guilherme Ide.F. Pelecium helenae, paratype (MCSN), photograph by Michele Zilioli. G. Pelecium helenae, allotype (MCSN), photograph by Michele Zilioli. H. Pelecium laeve Chaudoir, 1854, holotype (MNHN), photograph by Azadeh Taghavian. I. Pelecium longicolle Straneo, 1953, allotype (MCSN), photograph by Michele Zilioli. J. Pelecium negrei Straneo, 1962, holotype (MCSN), photograph by Michele Zilioli." figureDoi="http://doi.org/10.5281/zenodo.10715347" httpUri="https://zenodo.org/record/10715347/files/figure.png" pageId="33" pageNumber="34">Fig. 21E</figureCitation>
) of the
<typeStatus id="549E9AF4AB2BFFB390B2F9172F26A548" box="[453,556,1770,1796]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C255FD5AB2BFFB39338F9162FCDA548" authorityName="Straneo &amp; Ball" authorityYear="1989" box="[591,711,1770,1796]" class="Insecta" family="Carabidae" genus="Pelecium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="helenae">
<emphasis id="B951F844AB2BFFB39338F9162FCDA548" box="[591,711,1770,1796]" italics="true" pageId="33" pageNumber="34">P. helenae</emphasis>
</taxonomicName>
is wrongly labelled as
<taxonomicName id="4C255FD5AB2BFFB392A5F9162934A548" authorityName="Straneo &amp; Ball" authorityYear="1989" box="[978,1086,1770,1796]" class="Insecta" family="Carabidae" genus="Pelecium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="paulae">
<emphasis id="B951F844AB2BFFB392A5F9162934A548" box="[978,1086,1770,1796]" italics="true" pageId="33" pageNumber="34">P. paulae</emphasis>
</taxonomicName>
, but following the original description and comparing the locality label we can conclude that it is the
<typeStatus id="549E9AF4AB2BFFB39554F8ED2980A566" box="[1059,1162,1808,1834]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C255FD5AB2BFFB395C6F8EC2823A566" authorityName="Straneo &amp; Ball" authorityYear="1989" box="[1201,1321,1808,1834]" class="Insecta" family="Carabidae" genus="Pelecium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="helenae">
<emphasis id="B951F844AB2BFFB395C6F8EC2823A566" box="[1201,1321,1808,1834]" italics="true" pageId="33" pageNumber="34">P. helenae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9A2456AB2BFFB391CAF8AE2F16A521" blockId="33.[189,540,1875,1901]" box="[189,540,1875,1901]" pageId="33" pageNumber="34">
<heading id="D0D2933AAB2BFFB391CAF8AE2F16A521" bold="true" box="[189,540,1875,1901]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B951F844AB2BFFB391CAF8AE2F16A521" bold="true" box="[189,540,1875,1901]" pageId="33" pageNumber="34">Complementary description</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F77DDAB2BFFB191CAF8792800A1A3" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34" type="description">
<paragraph id="8B9A2456AB2BFFB391CAF8792C2CA5A4" blockId="33.[189,1399,1924,2024]" pageId="33" pageNumber="34">
<smallCapsWord id="8D7CB28AAB2BFFB391CAF8792C4BA5D0" baselines="1943,1943" box="[189,321,1924,1950]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdominal" pageId="33" pageNumber="34">ABDOMINAL</smallCapsWord>
<smallCapsWord id="8D7CB28AAB2BFFB3903CF8742CABA5D0" baselines="1943" box="[331,417,1929,1948]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="tergite" pageId="33" pageNumber="34">TERGITE</smallCapsWord>
IX (
<figureCitation id="131E38D3AB2BFFB390AAF8792F30A5D2" box="[477,570,1924,1950]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[302,1289,266,1760]" captionTargetId="figure-126@34.[296,1295,259,1764]" captionTargetPageId="34" captionText="Fig. 9. A–H. Pelecium helenae Straneo &amp; Ball, 1989, male from Piraquara, Paraná, Brazil (A–G) (CERPE) and female from Mogi das Cruzes, São Paulo, Brazil (H) (CEMT). A. Dorsal view. B. Detail of head. C–D. Median lobe. E. Tergite IX. F. Left paramere. G. Right paramere. H. Genitalia. – I–P. Pelecium laeve Chaudoir, 1854, male from Porto Alegre, Rio Grande do Sul, Brazil (I–O) (FZRBS) and female from Porto Alegre, Rio Grande do Sul, Brazil (P) (FZRBS). I. Dorsal view. J. Detail of head. K–L. Median lobe. M. Tergite IX. N. Left paramere. O. Right paramere. P. Genitalia. Scale bars: A–B, I–J = 1 mm; C–H, K–P = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10715310" httpUri="https://zenodo.org/record/10715310/files/figure.png" pageId="33" pageNumber="34">Fig. 9E</figureCitation>
). Oval, asymmetrical, sides converging to anterior portion forming a rounded angle; left side more angulate them right side; inner anterior angle rounded; posterior portion rounded.
</paragraph>
<caption id="DF5A74DEAB28FFB091CAF9062F47A5A6" ID-DOI="http://doi.org/10.5281/zenodo.10715310" ID-Zenodo-Dep="10715310" httpUri="https://zenodo.org/record/10715310/files/figure.png" pageId="34" pageNumber="35" startId="34.[189,232,1787,1813]" targetBox="[302,1289,266,1760]" targetPageId="34">
<paragraph id="8B9A2456AB28FFB091CAF9062F47A5A6" blockId="34.[188,1399,1787,2026]" pageId="34" pageNumber="35">
<emphasis id="B951F844AB28FFB091CAF9062C5FA559" bold="true" box="[189,341,1787,1814]" pageId="34" pageNumber="35">Fig. 9. A–H</emphasis>
.
<taxonomicName id="4C255FD5AB28FFB0901FF9012E59A55A" authority="Straneo &amp; Ball, 1989" authorityName="Straneo &amp; Ball" authorityYear="1989" box="[360,851,1788,1814]" class="Insecta" family="Carabidae" genus="Pelecium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="helenae">
<emphasis id="B951F844AB28FFB0901FF9012F36A55A" box="[360,572,1788,1814]" italics="true" pageId="34" pageNumber="35">Pelecium helenae</emphasis>
<bibRefCitation id="EFB459A7AB28FFB0933EF9012E59A55A" author="Straneo S. L. &amp; Ball G. B." box="[585,851,1788,1814]" pageId="34" pageNumber="35" pagination="73 - 178" refId="ref27763" refString="Straneo S. L. &amp; Ball G. B. 1989. Synopsis of the genera and subgenera of the tribe Peleciini, and revision of the Neotropical and Oriental species (Coleoptera: Carabidae). Insecta Mundi 3 (2): 73 - 178." type="journal article" year="1989">Straneo &amp; Ball, 1989</bibRefCitation>
</taxonomicName>
, male from Piraquara, Paraná, Brazil (A–G) (CERPE) and female from Mogi das Cruzes, São Paulo, Brazil (H) (CEMT).
<emphasis id="B951F844AB28FFB0953BF8E2296BA575" bold="true" box="[1100,1121,1823,1849]" pageId="34" pageNumber="35">A</emphasis>
. Dorsal view.
<emphasis id="B951F844AB28FFB0947AF8E2282AA575" bold="true" box="[1293,1312,1823,1849]" pageId="34" pageNumber="35">B</emphasis>
. Detail of head.
<emphasis id="B951F844AB28FFB0905AF8BF2C6CA510" bold="true" box="[301,358,1858,1884]" pageId="34" pageNumber="35">C–D</emphasis>
. Median lobe.
<emphasis id="B951F844AB28FFB09350F8BF2F31A510" bold="true" box="[551,571,1858,1884]" pageId="34" pageNumber="35">E</emphasis>
. Tergite IX.
<emphasis id="B951F844AB28FFB09397F8BF2FF8A510" bold="true" box="[736,754,1858,1884]" pageId="34" pageNumber="35">F</emphasis>
. Left paramere.
<emphasis id="B951F844AB28FFB092B3F8BF2ED1A510" bold="true" box="[964,987,1858,1884]" pageId="34" pageNumber="35">G</emphasis>
. Right paramere.
<emphasis id="B951F844AB28FFB095C9F8BF29DFA510" bold="true" box="[1214,1237,1858,1884]" pageId="34" pageNumber="35">H</emphasis>
. Genitalia.
<emphasis id="B951F844AB28FFB0941FF8BF2DE3A533" bold="true" pageId="34" pageNumber="35">– I–P</emphasis>
.
<taxonomicName id="4C255FD5AB28FFB09181F8982F6EA5CC" authority="Chaudoir, 1854" authorityName="Chaudoir" authorityYear="1854" box="[246,612,1893,1920]" class="Insecta" family="Carabidae" genus="Pelecium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="laeve">
<emphasis id="B951F844AB28FFB09181F8982CA2A533" box="[246,424,1893,1919]" italics="true" pageId="34" pageNumber="35">Pelecium laeve</emphasis>
<bibRefCitation id="EFB459A7AB28FFB090D9F8982F6EA5CC" author="Chaudoir M." box="[430,612,1893,1920]" pageId="34" pageNumber="35" pagination="279 - 352" refId="ref26834" refString="Chaudoir M. 1854. Memoire sur la famille des carabiques. Bulletin de la Societe imperiale des Naturalistes de Moscou 27: 112 - 144, 279 - 352." type="journal article" year="1854">Chaudoir, 1854</bibRefCitation>
</taxonomicName>
, male from Porto Alegre, Rio Grande do Sul, Brazil (I–O) (FZRBS) and female from Porto Alegre, Rio Grande do Sul, Brazil (P) (FZRBS).
<emphasis id="B951F844AB28FFB0928BF874290DA5EF" bold="true" box="[1020,1031,1929,1955]" pageId="34" pageNumber="35">I</emphasis>
. Dorsal view.
<emphasis id="B951F844AB28FFB095D9F87429B7A5EF" bold="true" box="[1198,1213,1929,1955]" pageId="34" pageNumber="35">J</emphasis>
. Detail of head.
<emphasis id="B951F844AB28FFB091CBF8512DFFA58A" bold="true" box="[188,245,1964,1990]" pageId="34" pageNumber="35">K–L</emphasis>
. Median lobe.
<emphasis id="B951F844AB28FFB090D1F8512CC8A58A" bold="true" box="[422,450,1964,1990]" pageId="34" pageNumber="35">M</emphasis>
. Tergite IX.
<emphasis id="B951F844AB28FFB09321F8512F61A58A" bold="true" box="[598,619,1964,1990]" pageId="34" pageNumber="35">N</emphasis>
. Left paramere.
<emphasis id="B951F844AB28FFB09259F8562E4FA589" bold="true" box="[814,837,1963,1989]" pageId="34" pageNumber="35">O</emphasis>
. Right paramere.
<emphasis id="B951F844AB28FFB0956FF8512920A58A" bold="true" box="[1048,1066,1964,1990]" pageId="34" pageNumber="35">P</emphasis>
. Genitalia. Scale bars: A–B, I–J = 1 mm; C–H, K–P = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9A2456AB29FFB191CAFEF42C0FA3FA" blockId="35.[189,1400,265,438]" pageId="35" pageNumber="36">
<smallCapsWord id="8D7CB28AAB29FFB191CAFEF42C38A36D" baselines="284,284" box="[189,306,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Aedeagus" pageId="35" pageNumber="36">AEDEAGUS</smallCapsWord>
(
<figureCitation id="131E38D3AB29FFB19034FEF42F0CA36F" box="[323,518,265,291]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[302,1289,266,1760]" captionTargetId="figure-126@34.[296,1295,259,1764]" captionTargetPageId="34" captionText="Fig. 9. A–H. Pelecium helenae Straneo &amp; Ball, 1989, male from Piraquara, Paraná, Brazil (A–G) (CERPE) and female from Mogi das Cruzes, São Paulo, Brazil (H) (CEMT). A. Dorsal view. B. Detail of head. C–D. Median lobe. E. Tergite IX. F. Left paramere. G. Right paramere. H. Genitalia. – I–P. Pelecium laeve Chaudoir, 1854, male from Porto Alegre, Rio Grande do Sul, Brazil (I–O) (FZRBS) and female from Porto Alegre, Rio Grande do Sul, Brazil (P) (FZRBS). I. Dorsal view. J. Detail of head. K–L. Median lobe. M. Tergite IX. N. Left paramere. O. Right paramere. P. Genitalia. Scale bars: A–B, I–J = 1 mm; C–H, K–P = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10715310" httpUri="https://zenodo.org/record/10715310/files/figure.png" pageId="35" pageNumber="36">Fig. 9C–D, F–G</figureCitation>
). With
<emphasis id="B951F844AB29FFB1932EFEF42FFBA36F" bold="true" box="[601,753,265,291]" pageId="35" pageNumber="36">median lobe</emphasis>
(
<figureCitation id="131E38D3AB29FFB19276FEF42E75A36F" box="[769,895,265,291]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[302,1289,266,1760]" captionTargetId="figure-126@34.[296,1295,259,1764]" captionTargetPageId="34" captionText="Fig. 9. A–H. Pelecium helenae Straneo &amp; Ball, 1989, male from Piraquara, Paraná, Brazil (A–G) (CERPE) and female from Mogi das Cruzes, São Paulo, Brazil (H) (CEMT). A. Dorsal view. B. Detail of head. C–D. Median lobe. E. Tergite IX. F. Left paramere. G. Right paramere. H. Genitalia. – I–P. Pelecium laeve Chaudoir, 1854, male from Porto Alegre, Rio Grande do Sul, Brazil (I–O) (FZRBS) and female from Porto Alegre, Rio Grande do Sul, Brazil (P) (FZRBS). I. Dorsal view. J. Detail of head. K–L. Median lobe. M. Tergite IX. N. Left paramere. O. Right paramere. P. Genitalia. Scale bars: A–B, I–J = 1 mm; C–H, K–P = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10715310" httpUri="https://zenodo.org/record/10715310/files/figure.png" pageId="35" pageNumber="36">Fig. 9C–D</figureCitation>
) cylindrical, slightly asymmetrical, bent to left, slightly broadening apically; ostium short, confined to apical fourth; apical lamella long, not evenly narrowed, apical edges narrow; basal bulb thin.
<emphasis id="B951F844AB29FFB19274FEAE2E80A321" bold="true" box="[771,906,339,365]" pageId="35" pageNumber="36">Parameres</emphasis>
(
<figureCitation id="131E38D3AB29FFB192EEFEAF291DA321" box="[921,1047,338,365]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[302,1289,266,1760]" captionTargetId="figure-126@34.[296,1295,259,1764]" captionTargetPageId="34" captionText="Fig. 9. A–H. Pelecium helenae Straneo &amp; Ball, 1989, male from Piraquara, Paraná, Brazil (A–G) (CERPE) and female from Mogi das Cruzes, São Paulo, Brazil (H) (CEMT). A. Dorsal view. B. Detail of head. C–D. Median lobe. E. Tergite IX. F. Left paramere. G. Right paramere. H. Genitalia. – I–P. Pelecium laeve Chaudoir, 1854, male from Porto Alegre, Rio Grande do Sul, Brazil (I–O) (FZRBS) and female from Porto Alegre, Rio Grande do Sul, Brazil (P) (FZRBS). I. Dorsal view. J. Detail of head. K–L. Median lobe. M. Tergite IX. N. Left paramere. O. Right paramere. P. Genitalia. Scale bars: A–B, I–J = 1 mm; C–H, K–P = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10715310" httpUri="https://zenodo.org/record/10715310/files/figure.png" pageId="35" pageNumber="36">Fig. 9F–G</figureCitation>
) asymmetrical; left paramere elongate with posterior angle rounded; right paramere square-shaped with protuberant, rounded lateral angle.
</paragraph>
<paragraph id="8B9A2456AB29FFB191CAFE222F1BA052" blockId="35.[189,1398,479,542]" pageId="35" pageNumber="36">
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1 (
<figureCitation id="131E38D3AB29FFB19006FE222CC4A3B5" box="[369,462,479,506]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[302,1289,266,1760]" captionTargetId="figure-126@34.[296,1295,259,1764]" captionTargetPageId="34" captionText="Fig. 9. A–H. Pelecium helenae Straneo &amp; Ball, 1989, male from Piraquara, Paraná, Brazil (A–G) (CERPE) and female from Mogi das Cruzes, São Paulo, Brazil (H) (CEMT). A. Dorsal view. B. Detail of head. C–D. Median lobe. E. Tergite IX. F. Left paramere. G. Right paramere. H. Genitalia. – I–P. Pelecium laeve Chaudoir, 1854, male from Porto Alegre, Rio Grande do Sul, Brazil (I–O) (FZRBS) and female from Porto Alegre, Rio Grande do Sul, Brazil (P) (FZRBS). I. Dorsal view. J. Detail of head. K–L. Median lobe. M. Tergite IX. N. Left paramere. O. Right paramere. P. Genitalia. Scale bars: A–B, I–J = 1 mm; C–H, K–P = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10715310" httpUri="https://zenodo.org/record/10715310/files/figure.png" pageId="35" pageNumber="36">Fig. 9H</figureCitation>
). Outer and inner sides arcuate; inner side forming acute projection on posterior margin; anterior margin dull.
</paragraph>
<paragraph id="8B9A2456AB29FFB191CAFDBA2E35A0CA" blockId="35.[189,1399,583,646]" pageId="35" pageNumber="36">
<smallCapsWord id="8D7CB28AAB29FFB191CAFDBA2C40A013" baselines="602,602" box="[189,330,583,609]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Gonocoxite" pageId="35" pageNumber="36">GONOCOXITE</smallCapsWord>
2 (
<figureCitation id="131E38D3AB29FFB19006FDBA2CC4A02D" box="[369,462,583,609]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[302,1289,266,1760]" captionTargetId="figure-126@34.[296,1295,259,1764]" captionTargetPageId="34" captionText="Fig. 9. A–H. Pelecium helenae Straneo &amp; Ball, 1989, male from Piraquara, Paraná, Brazil (A–G) (CERPE) and female from Mogi das Cruzes, São Paulo, Brazil (H) (CEMT). A. Dorsal view. B. Detail of head. C–D. Median lobe. E. Tergite IX. F. Left paramere. G. Right paramere. H. Genitalia. – I–P. Pelecium laeve Chaudoir, 1854, male from Porto Alegre, Rio Grande do Sul, Brazil (I–O) (FZRBS) and female from Porto Alegre, Rio Grande do Sul, Brazil (P) (FZRBS). I. Dorsal view. J. Detail of head. K–L. Median lobe. M. Tergite IX. N. Left paramere. O. Right paramere. P. Genitalia. Scale bars: A–B, I–J = 1 mm; C–H, K–P = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10715310" httpUri="https://zenodo.org/record/10715310/files/figure.png" pageId="35" pageNumber="36">Fig. 9H</figureCitation>
). Elongate; basal lobe bearing ensiform setae on each side; ensiform setae long, with half length of gonocoxite 2; slender setae at apex.
</paragraph>
<paragraph id="8B9A2456AB29FFB191CAFD522800A1A3" blockId="35.[189,1399,687,1008]" pageId="35" pageNumber="36">
<smallCapsWord id="8D7CB28AAB29FFB191CAFD522C66A08B" baselines="706,706" box="[189,364,687,713]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="35" pageNumber="36">MEASUREMENTS</smallCapsWord>
(in mm). Males (n = 4): AL1 0.8–0.9 (0.83 ± 0.05), AL2 0.3–0.5 (0.43 ± 0.1), EL 4.7–6.7 (5.78 ± 0.85), EW 3.2–4.3 (3.98± 0.52), PL 2.7–3.6 (3.28 ± 0.39), PW 2.6–3.6 (2.25 ±0.39), HL 1.3–1.6 (1.53± 0.15), HW 1.7–2.4 (2.15± 0.31), TL 8.7–11.9 (10.58 ± 1.36), TL/EW 2.5–2.8 (2.66 ± 0.11), lp2L 0.5–0.7 (0.6 ±0.08), lp3L 0.6–0.7 (0.68 ± 0.1), lp3
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–0.4 (0.35± 0.06), mp3L 0.2–03 (0.28± 0.05), mp4L 0.5–0.7 (0.63 ±0.1), mp4
<geoCoordinate id="EE114291AB29FFB1934DFCBF2F8FA110" box="[570,645,834,860]" degrees="0.3" direction="west" orientation="longitude" pageId="35" pageNumber="36" precision="5555" value="-0.3">W 0.3</geoCoordinate>
–0.4 (0.33 ± 0.05). Females (n = 2): AL1 1.1–1.2 (1.15± 0.07), AL2 0.6–0.6 (0.6 ± 0.0), EL 8.95–0.35 (8.95 ± 0.35), EW 5.7–5.9 (5.8 ±0.14), PL 4.8–4.9 (4.85± 0.07), PW 4.3–4.7 (4.5 ± 0.28), HL 2.8–3.0 (2.1 ±0.28), HW 2.8–3.0 (2.9± 0.14), TL 15.5–16.3 (15.9± 0.57), TL/EW 2.7–2.8 (2.74 ± 0.03), lp2L 0.8–0.9 (0.85± 0.07), lp3L 0.8–0.9 (0.85± 0.07), lp3
<geoCoordinate id="EE114291AB29FFB19584FC4D2848A186" box="[1267,1346,944,970]" degrees="0.3" direction="west" orientation="longitude" pageId="35" pageNumber="36" precision="5555" value="-0.3">W 0.3</geoCoordinate>
–0.3 (0.3 ± 0.0), mp3L 0.4–0.5 (0.45± 0.07), mp4L 0.8–0.8 (0.8 ±0.0), mp4
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–0.4 (0.35± 0.07).
</paragraph>
</subSubSection>
</treatment>
</document>
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