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<mods:title id="34F72F6073EEB1750FDDF9260CA81EBD">Revision of the Hawaiian endemic leaf-mining moth genus Philodoria Walsingham (Lepidoptera: Gracillariidae): its conservation status, host plants and descriptions of thirteen new species</mods:title>
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<taxonomicName id="4C294D5EFFC80729FF75932EFD09A674" ID-CoL="4G8NY" authority="Walsingham, 1907" authorityName="Walsingham" authorityYear="1907" box="[151,640,1831,1858]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="splendida">
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<emphasis id="B95DEACFFFC80729FF75932EFE1AA674" bold="true" box="[151,403,1831,1858]" italics="true" pageId="38" pageNumber="39">Philodoria splendida</emphasis>
<bibRefCitation id="EFB84B2CFFC80729FE789321FD09A674" author="Walsingham, Thomas" box="[410,640,1832,1858]" pageId="38" pageNumber="39" pagination="469 - 759" refId="ref102986" refString="Walsingham, Thomas, de Grey (Lord) (1907) Microlepidoptera. In: Sharp, D. (Ed.), Fauna Hawaiiensis. 1 (5). Cambridge University Press, Cambridge, pp. 469 - 759." type="book chapter" year="1907">Walsingham, 1907</bibRefCitation>
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<figureCitation id="13122A58FFC80729FF759345FE92A650" box="[151,283,1868,1894]" captionStart="FIGURE 9" captionStartId="24.[151,250,1194,1218]" captionTargetBox="[155,1433,204,1166]" captionTargetId="figure-23@24.[151,1437,200,1170]" captionTargetPageId="24" captionText="FIGURE 9. Adults of Philodoria species. A P. splendida Walsingham, 1907 lectotype male, B Female, Oahu, C P. lama sp. n. holotype male, D P. napaliensis sp. n. holotype (sex unknown). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4681831" httpUri="https://zenodo.org/record/4681831/files/figure.png" pageId="38" pageNumber="39">Figs. 9A, B</figureCitation>
, 
<figureCitation id="13122A58FFC80729FEC49345FEF1A650" box="[294,376,1868,1894]" captionStart="FIGURE 26" captionStartId="49.[151,250,1720,1744]" captionTargetBox="[195,1392,185,1692]" captionTargetId="figure-20@49.[191,1395,181,1696]" captionTargetPageId="49" captionText="FIGURE 26. Forewing pattern variation in Philodoria. Specimens from recent collection efforts that were preserved in ethanol or RNA stabilization reagent, originally for molecular studies. Wings were placed without mountant under a coverslip. A P. succedanea (sex unknown, CJ419), B P. auromagnifica, male (CJ148/SK689), C P. basalis, female (CJ410), D P. splendida, female (CJ488a), host: Syzygium sp., E P. splendida (sex unknown, CJ488b), host: Syzygium sp., F P. molokaiensis, male (CJ509b/ SK667), G P. sp. A nr. P. lysimachiella, female (CJ346a/SK753), H P. lysimachiella, female (CJ530), lateral view, I P. pipturiella, female (CJ529), J Same species, male (CJ529), K P. haelaauensis, female (CJ209), L Same species, female (CJ216), M P. floscula, female (CJ426/SK845), N P. wilkesiella, male (CJ009/SK649), O P. epibathra, male (CJ506a/SK678), P Same species, female (CJ506b/SK681). Arrows point to diagnostic characters of species: black patch at base (A), basal fascia (D), transverse fasciae (tf) (F, G), white spot (ws) (I, J), white patch (K–M). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4681867" httpUri="https://zenodo.org/record/4681867/files/figure.png" pageId="38" pageNumber="39">26D, E</figureCitation>
, 
<figureCitation id="13122A58FFC80729FE669345FE59A650" box="[388,464,1868,1894]" captionStart="FIGURE 41" captionStartId="76.[151,250,1685,1709]" captionTargetBox="[161,1426,183,1654]" captionTargetId="figure-23@76.[151,1436,181,1662]" captionTargetPageId="76" captionText="FIGURE 41. Male valva of Philodoria, inner view. A–F, H–J, L, M: Left valva, G, K, N: Right valva. A P. succedanea, Maui, SK641, B P. auromagnifica, Hawaii (Big Island), SK800, C P. auromagnifica, Kauai, SK689, D P. kolea, holotype, Hawaii (Big Island), SK851, E P. lysimachiella, Oahu, SK656, F, G P. sp. 7, Kauai, SK754, H P. molokaiensis, Molokai, SK666, I P. basalis, Hawaii (Big Island), SK614, J P. splendida, Maui, SK696, K Same species, Oahu, SK747, host Syzygium sp., L P. lama, holotype, Maui SK744, M P. napaliensis, paratype, Kauai, SK767, N Same as L, paratype, Kauai, SK745." figureDoi="http://doi.org/10.5281/zenodo.4681901" httpUri="https://zenodo.org/record/4681901/files/figure.png" pageId="38" pageNumber="39">41J, K</figureCitation>
, 
<figureCitation id="13122A58FFC80729FE399345FD92A650" box="[475,539,1868,1894]" captionStart="FIGURE 42" captionStartId="77.[151,250,1773,1797]" captionTargetBox="[168,1421,187,1741]" captionTargetId="figure-20@77.[158,1430,181,1748]" captionTargetPageId="77" captionText="FIGURE 42. Genital capsule of Philodoria, excluding right portion of male genitalia, ventral view. A P. succedanea, Maui, SK641, B P. auromagnifica, Hawaii (Big Island), SK800, C P. auromagnifica, Kauai, SK689, D P. kolea, holotype, Hawaii (Big Island), SK851, E P. lysimachiella, Oahu, SK656, F P. sp. 7, Kauai, SK754, G P. molokaiensis, Molokai, SK666, H P. basalis, Hawaii (Big Island), SK614, I P. splendida, Maui, SK696, J Same species, Oahu, SK747, host Syzygium sp., K P. lama, holotype, Maui, SK744, L P. napaliensis, paratype, Kauai, SK745. Abbreviation: ta: tuba analis." figureDoi="http://doi.org/10.5281/zenodo.4681903" httpUri="https://zenodo.org/record/4681903/files/figure.png" pageId="38" pageNumber="39">42I, J</figureCitation>
, 
<figureCitation id="13122A58FFC80729FDC59345FDEEA650" box="[551,615,1868,1894]" captionStart="FIGURE 43" captionStartId="78.[151,250,1790,1814]" captionTargetBox="[159,1418,191,1759]" captionTargetId="figure-23@78.[151,1436,181,1766]" captionTargetPageId="78" captionText="FIGURE 43. Phallus of Philodoria, lateral view. A P. succedanea, Maui, SK641, B P. auromagnifica, Hawaii (Big Island), SK800, C P. auromagnifica, Kauai, SK689, D P. kolea, holotype, Hawaii (Big Island), SK851, E P. lysimachiella, Oahu, SK656, F P. sp. 7, Kauai, SK755, G P. molokaiensis, Molokai, SK666, H P. basalis, Hawaii (Big Island), SK614, I P. splendida, Maui, SK696, J Same species, Oahu, SK747, host Syzygium sp., K P. lama, holotype, Maui, SK744, L P. napaliensis, paratype, Kauai, SK767." figureDoi="http://doi.org/10.5281/zenodo.4681905" httpUri="https://zenodo.org/record/4681905/files/figure.png" pageId="38" pageNumber="39">43I, J</figureCitation>
, 
<figureCitation id="13122A58FFC80729FD919345FD2FA651" box="[627,678,1868,1895]" captionStart="FIGURE 56" captionStartId="106.[151,250,1881,1905]" captionTargetBox="[170,1411,192,1821]" captionTargetId="figure-23@106.[165,1422,181,1857]" captionTargetPageId="106" captionText="FIGURE 56. Female genitalia of Philodoria. A P. succedanea, paralectotype, Maui, SK714, B Same species, Hawaii (Big Island), SK624, C P. auromagnifica, Hawaii (Big Island), SK623, D P. kauaulaensis, holotype, SK690, E Same species, lateral view, F P. kolea, paratype, Hawaii (Big Island), SK634, G P. lysimachiella, SK654, H P. sp. A, SK753, I P. molokaiensis, SK668, lateral view, J Same ventral view, K P. splendida, SK621, L P. lama, paratype, SK813." figureDoi="http://doi.org/10.5281/zenodo.4681931" httpUri="https://zenodo.org/record/4681931/files/figure.png" pageId="38" pageNumber="39">56K</figureCitation>
, 
<figureCitation id="13122A58FFC80729FD539345FD75A650" box="[689,764,1868,1894]" captionStart="FIGURE 64" captionStartId="115.[151,250,1375,1399]" captionTargetBox="[167,1422,200,1348]" captionTargetId="figure-20@115.[162,1425,198,1351]" captionTargetPageId="115" captionText="FIGURE 64. BPBM genitalia slides prepared by E. C. Zimmerman. A–D: P. haelaauensis, paratype male, Maui, Z-XII-19-62- 23, E, F: P. pipturiella paratype female, Oahu, Z-XII-1962-22, G–I: P. splendida, male, Oahu, Z-XII-19-62-20. A Vinculum and valvae, tegumen broken, B, H Phallus, C Tegumen? D, I Abdominal segments, E Genitalia opened from left lateral side, F Abdominal segments II–VI, G Genitalia excluding phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4681949" httpUri="https://zenodo.org/record/4681949/files/figure.png" pageId="38" pageNumber="39">64G–I</figureCitation>
, 
<figureCitation id="13122A58FFC80729FCE59345FCD2A650" box="[775,859,1868,1894]" captionStart="FIGURE 66" captionStartId="118.[151,250,1829,1853]" captionTargetBox="[158,1430,528,1803]" captionTargetId="figure-197@118.[154,1433,524,1805]" captionTargetPageId="118" captionText="FIGURE 66. Habitats and host plants of Philodoria on Kauai. A–D: Upper Limahuli, E, F: Waimea Canyon. A Mountains of Limahuli, B Habitat with senior (last) author collecting specimens, C Same location as B, leaves with mines, D Metrosideros polymorpha leaves with mine on the ground with active larva, E Hibiscus waimeae subsp. hannerae flowers and leaf mines, F Leaf mine on same host plant as E. Arrows point to mines or larvae." figureDoi="http://doi.org/10.5281/zenodo.4681953" httpUri="https://zenodo.org/record/4681953/files/figure.png" pageId="38" pageNumber="39">66C, D</figureCitation>
, 
<figureCitation id="13122A58FFC80729FC849345FC31A650" box="[870,952,1868,1894]" captionStart="FIGURE 73" captionStartId="125.[151,250,1698,1722]" captionTargetBox="[159,1429,241,1669]" captionTargetId="figure-57@125.[151,1436,237,1675]" captionTargetPageId="125" captionText="FIGURE 73. Resting posture of adult Philodoria and Caloptilia. A P. molokaiensis, Molokai, CJ229, host: Lysimachia hillebrandii, B P. splendida, Oahu, CJ319, host: Metrosideros polymorpha, C Same species, Oahu, D Caloptilia mabaella, Lanai, CJ333, host: Diospyros sandwicensis, E P. lama, Maui, CJ190, F Same species, Maui, CJ194." figureDoi="http://doi.org/10.5281/zenodo.4681967" httpUri="https://zenodo.org/record/4681967/files/figure.png" pageId="38" pageNumber="39">73B, C</figureCitation>
, 
<figureCitation id="13122A58FFC80729FC219345FC6BA650" box="[963,994,1868,1894]" captionStart="FIGURE 82" captionStartId="144.[151,250,1255,1279]" captionTargetBox="[159,1425,187,1227]" captionTargetId="figure-23@144.[155,1432,181,1230]" captionTargetPageId="144" captionText="FIGURE 82. Philodoria splendida and its host plants. A, D, E: Metrosideros polymorpha, B, C, F: Syzigium sandwicensis. A: Maui, B–F: Oahu. A, B Leaf mine, C Cocoon and old mine, D, E Host plant with leaf mines, F Mine of late instar larva." figureDoi="http://doi.org/10.5281/zenodo.4681986" httpUri="https://zenodo.org/record/4681986/files/figure.png" pageId="38" pageNumber="39">82</figureCitation>
.
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<treatmentCitation id="0A8810CCFFC80729FF759399FD11A69E" author="Walsingham, Thomas" box="[151,664,1936,1960]" page="720" pageId="38" pageNumber="39" year="1907">
<taxonomicName id="4C294D5EFFC80729FF759399FD11A69E" ID-CoL="4G8NY" authority="Walsingham, 1907: 719 - 720" authorityName="Walsingham" authorityPageNumber="719 - 720" authorityYear="1907" box="[151,664,1936,1960]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="splendida">
<emphasis id="B95DEACFFFC80729FF759399FEE7A69E" box="[151,366,1936,1960]" italics="true" pageId="38" pageNumber="39">Philodoria splendida</emphasis>
<bibRefCitation id="EFB84B2CFFC80729FE919399FD11A69E" author="Walsingham, Thomas" box="[371,664,1936,1960]" pageId="38" pageNumber="39" pagination="469 - 759" refId="ref102986" refString="Walsingham, Thomas, de Grey (Lord) (1907) Microlepidoptera. In: Sharp, D. (Ed.), Fauna Hawaiiensis. 1 (5). Cambridge University Press, Cambridge, pp. 469 - 759." type="book chapter" year="1907">Walsingham, 1907: 719–720</bibRefCitation>
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, pl. 25, fig. 23; 
<treatmentCitation id="0A8810CCFFC80729FCD59399FB88A69E" author="Swezey, O. H." box="[823,1025,1936,1960]" page="223" pageId="38" pageNumber="39" year="1913">
<bibRefCitation id="EFB84B2CFFC80729FCD59399FB88A69E" author="Swezey, O. H." box="[823,1025,1936,1960]" pageId="38" pageNumber="39" pagination="221 - 227" refId="ref102406" refString="Swezey, O. H. (1913 b) Leaf-miners of the Hawaiian Islands. Proceedings of the Hawaiian Entomological Society, 2 (5), 221 - 227." type="journal article" year="1913">Swezey 1913b: 223</bibRefCitation>
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; 
<treatmentCitation id="0A8810CCFFC80729FBE99399FA88A69E" author="Zimmerman, E. C." box="[1035,1281,1936,1960]" page="718" pageId="38" pageNumber="39" year="1978">
<bibRefCitation id="EFB84B2CFFC80729FBE99399FA88A69E" author="Zimmerman, E. C." box="[1035,1281,1936,1960]" pageId="38" pageNumber="39" refId="ref103182" refString="Zimmerman, E. C. (1978 a) Gracillarioidea. In: Microlepidoptera. Part 1. Insects of Hawaii, Vol. 9. University of Hawaii Press, Honolulu, pp. 644 - 718." type="book" year="1978">Zimmerman 1978a: 718</bibRefCitation>
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, figs. 466, 471, 477, 481.
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<emphasis id="B95DEACFFFC90728FF75949FFEBFA186" bold="true" box="[151,310,150,176]" pageId="39" pageNumber="40">
<typeStatus id="5492887FFFC90728FF75949FFF5BA186" box="[151,210,150,176]" pageId="39" pageNumber="40">Type</typeStatus>
locality.
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Lanai, 
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<paragraph id="8B9636DDFFC90728FF2594B3FC0AA222" blockId="39.[151,1437,150,2042]" pageId="39" pageNumber="40">
<emphasis id="B95DEACFFFC90728FF2594B3FEFDA1E2" bold="true" box="[199,372,186,212]" pageId="39" pageNumber="40">Type material.</emphasis>
<typeStatus id="5492887FFFC90728FE9994B2FE62A1E3" box="[379,491,187,213]" pageId="39" pageNumber="40" type="lectotype">Lectotype</typeStatus>
♂, Lanai, 
<quantity id="4CD19B38FFC90728FD8394B3FD24A1E2" box="[609,685,186,212]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="39" pageNumber="40" unit="ft" value="3000.0">3000ft</quantity>
, 
<date id="FF97101DFFC90728FD5594B3FC80A1E2" box="[695,777,186,212]" pageId="39" pageNumber="40" value="1894-02">ii.1894</date>
, Perkins 26452 |♂ genitalia on slide 101, Oct.17.1940| Philodoria 
<taxonomicName id="4C294D5EFFC90728FF0E94D7FE32A1CE" authority="D Wlsm" authorityName="D Wlsm" box="[236,443,222,249]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="splendida">splendida ♂ Wlsm</taxonomicName>
<typeStatus id="5492887FFFC90728FE2394D7FDC5A1CE" box="[449,588,222,248]" pageId="39" pageNumber="40" type="paratype">PARATYPE</typeStatus>
2/9| Fauna Hawaiiensis Collection| No. 34284 Hawaiian Coll. BISHOP Museum in 
<collectionCode id="ED38AE18FFC90728FF14950AFECDA02B" box="[246,324,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
(here designated). 
<typeStatus id="5492887FFFC90728FDF8950BFD3BA02A" box="[538,690,258,284]" pageId="39" pageNumber="40" type="paralectotype">Paralectotype</typeStatus>
2 (1♂, 1 sex unknown). 1 (sex unknown), Lanai, 
<quantity id="4CD19B38FFC90728FB00950BFAA7A02A" box="[1250,1326,258,284]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="39" pageNumber="40" unit="ft" value="2000.0">2000ft</quantity>
, 
<date id="FF97101DFFC90728FADB950BFA11A02A" box="[1337,1432,258,284]" pageId="39" pageNumber="40" value="1894-07">vii.1894</date>
, Perkins 26493 | 
<taxonomicName id="4C294D5EFFC90728FEB7952FFD02A076" authority="Wlsm" authorityName="Walsingham" authorityYear="1907" box="[341,651,294,320]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="splendida">Philodoria splendida Wlsm</taxonomicName>
<typeStatus id="5492887FFFC90728FD77952FFCA9A076" box="[661,800,294,320]" pageId="39" pageNumber="40" type="paratype">PARATYPE</typeStatus>
3/9| in 
<collectionCode id="ED38AE18FFC90728FC9F952EFC47A077" box="[893,974,295,321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
; 1♂, Kauai Mts., 
<quantity id="4CD19B38FFC90728FB42952FFABAA076" box="[1184,1331,294,320]" metricMagnitude="3" metricUnit="m" metricValue="1.0668" metricValueMax="1.2192" metricValueMin="0.9144" pageId="39" pageNumber="40" unit="ft" value="3500.0" valueMax="4000.0" valueMin="3000.0">3000–4000ft</quantity>
, 
<date id="FF97101DFFC90728FAA2952FFA1EA076" box="[1344,1431,294,320]" pageId="39" pageNumber="40" value="1894-06">vi.1894</date>
, Perkins 27188, | 
<taxonomicName id="4C294D5EFFC90728FEB19543FD29A052" authority="D Wlsm" authorityName="D Wlsm" box="[339,672,330,357]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="splendida">Philodoria splendida ♂ Wlsm</taxonomicName>
<typeStatus id="5492887FFFC90728FD459543FCBBA052" box="[679,818,330,356]" pageId="39" pageNumber="40" type="paratype">PARATYPE</typeStatus>
9/9| in 
<collectionCode id="ED38AE18FFC90728FC659542FC51A053" box="[903,984,331,357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
. Described from ten specimens: ‘type ♂ (26430) and nine ‘paratypes’ from Kauai, Oahu, Molokai and Lanai. Given the manner in which the descriptions were written, Walsingham likely considered them as the 
<typeStatus id="5492887FFFC90728FCC4959BFC01A09A" box="[806,904,402,428]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
and 
<typeStatus id="5492887FFFC90728FC21959AFBB8A09B" box="[963,1073,403,429]" pageId="39" pageNumber="40" type="paratype">paratypes</typeStatus>
, as indicated on their specimen labels. However, because a 
<typeStatus id="5492887FFFC90728FE2D95BFFDB8A0E6" box="[463,561,438,464]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
was not specified in the description, the so-labeled types and 
<typeStatus id="5492887FFFC90728FB0895BEFADCA0E7" box="[1258,1365,439,465]" pageId="39" pageNumber="40" type="paratype">paratypes</typeStatus>
are to be considered 
<typeStatus id="5492887FFFC90728FED995D2FE14A0C3" box="[315,413,475,501]" pageId="39" pageNumber="40" type="syntype">syntypes</typeStatus>
under Article 73.2 of the Code (
<bibRefCitation id="EFB84B2CFFC90728FCEF95D3FC1DA0C2" author="ICZN" box="[781,916,474,500]" pageId="39" pageNumber="40" refId="ref101112" refString="ICZN (1999) International code of zoological nomenclature. Fourth Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">
<collectionCode id="ED38AE18FFC90728FCEF95D3FCD9A0C2" box="[781,848,474,500]" pageId="39" pageNumber="40">ICZN</collectionCode>
1999
</bibRefCitation>
), and any one of them is thus eligible for designation as 
<typeStatus id="5492887FFFC90728FEFA95F7FEF6A32E" box="[280,383,510,536]" pageId="39" pageNumber="40" type="lectotype">lectotype</typeStatus>
under Article 74 of the Code (
<bibRefCitation id="EFB84B2CFFC90728FD3695F7FCD3A32E" author="ICZN" box="[724,858,510,536]" pageId="39" pageNumber="40" refId="ref101112" refString="ICZN (1999) International code of zoological nomenclature. Fourth Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">
<collectionCode id="ED38AE18FFC90728FD3695F7FC9EA32E" box="[724,791,510,536]" pageId="39" pageNumber="40">ICZN</collectionCode>
1999
</bibRefCitation>
). We could not find ‘type ♂ (26430)’ from 
<quantity id="4CD19B38FFC90728FADC95F7FA11A32E" box="[1342,1432,510,536]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="39" pageNumber="40" unit="ft" value="2000.0">2000 ft.</quantity>
, Lanai, 
<date id="FF97101DFFC90728FF04962BFECBA30A" box="[230,322,546,572]" pageId="39" pageNumber="40" value="1894-12">xii.1894</date>
and six 
<typeStatus id="5492887FFFC90728FE41962AFD80A30B" box="[419,521,547,573]" pageId="39" pageNumber="40" type="syntype">syntypes</typeStatus>
: 1 (sex unknown), Halemanu, 
<quantity id="4CD19B38FFC90728FC86962BFC31A30A" box="[868,952,546,572]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="39" pageNumber="40" unit="ft" value="4000.0">4000 ft</quantity>
, Kauai, 
<date id="FF97101DFFC90728FBF7962BFBEDA30B" box="[1045,1124,546,573]" pageId="39" pageNumber="40" value="1895-05">v.1895</date>
; 1 (sex unknown), Waianae Mts., 
<quantity id="4CD19B38FFC90728FF3B964FFEBAA356" box="[217,307,582,608]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="39" pageNumber="40" unit="ft" value="2000.0">2000 ft.</quantity>
, 
<date id="FF97101DFFC90728FEA2964FFE1CA356" box="[320,405,582,608]" pageId="39" pageNumber="40" value="1892-04">iv.1892</date>
, beaten from “Lefua” (
<taxonomicName id="4C294D5EFFC90728FD7F964FFC36A356" authorityName="Gaudich. (Walsingham" authorityYear="1907" box="[669,959,582,608]" class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="polymorpha">
<emphasis id="B95DEACFFFC90728FD7F964FFC36A356" box="[669,959,582,608]" italics="true" pageId="39" pageNumber="40">Metrosideros polymorpha</emphasis>
</taxonomicName>
?); 1 (sex unknown), Mts., near Honolulu, 
<quantity id="4CD19B38FFC90728FF759663FF78A3B2" box="[151,241,618,644]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="39" pageNumber="40" unit="ft" value="2000.0">2000 ft.</quantity>
, 
<date id="FF97101DFFC90728FF1C9663FEF8A3B3" box="[254,369,618,645]" pageId="39" pageNumber="40" value="1892-10-25">25.x.1892</date>
; 1 (sex unknown), Kalae, 
<quantity id="4CD19B38FFC90728FD789663FD7DA3B2" box="[666,756,618,645]" metricMagnitude="2" metricUnit="m" metricValue="4.572" pageId="39" pageNumber="40" unit="ft" value="1500.0">1500 ft.</quantity>
, 
<date id="FF97101DFFC90728FCE29663FCC6A3B2" box="[768,847,618,644]" pageId="39" pageNumber="40" value="1893-05">v.1893</date>
; 1 (sex unknown), Mts., about 
<quantity id="4CD19B38FFC90728FB4C9663FA80A3B2" box="[1198,1289,618,644]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="39" pageNumber="40" unit="ft" value="4000.0">4000 ft.</quantity>
, 
<date id="FF97101DFFC90728FAF79663FA0BA3B2" box="[1301,1410,618,644]" pageId="39" pageNumber="40" value="1893-06-09">9.vi.1893</date>
; 1 (sex unknown), 
<quantity id="4CD19B38FFC90728FEB39687FE25A39E" box="[337,428,654,680]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="39" pageNumber="40" unit="ft" value="3000.0">3000 ft.</quantity>
, Lanai, 
<date id="FF97101DFFC90728FDE89687FDD2A39E" box="[522,603,654,680]" pageId="39" pageNumber="40" value="1894-02">ii.1894</date>
. 
<bibRefCitation id="EFB84B2CFFC90728FD8A9687FC0FA39E" author="Zimmerman, E. C." box="[616,902,654,680]" pageId="39" pageNumber="40" refId="ref103182" refString="Zimmerman, E. C. (1978 a) Gracillarioidea. In: Microlepidoptera. Part 1. Insects of Hawaii, Vol. 9. University of Hawaii Press, Honolulu, pp. 644 - 718." type="book" year="1978">Zimmerman (1978a: 701</bibRefCitation>
, fig. 466) provided the last photograph for the ‘holotype’ (=type ♂ (26430)) in the 
<collectionCode id="ED38AE18FFC90728FDD996BAFD20A3FB" box="[571,681,691,717]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="39" pageNumber="40" type="Museum">NHMUK</collectionCode>
. The current curator of 
<collectionCode id="ED38AE18FFC90728FC5F96BAFBA3A3FB" box="[957,1066,691,717]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="39" pageNumber="40" type="Museum">NHMUK</collectionCode>
, D.C. Lees informed us that the ‘holotype’ of 
<taxonomicName id="4C294D5EFFC90728FECE96DEFE3AA3C6" box="[300,435,726,752]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="splendida">
<emphasis id="B95DEACFFFC90728FECE96DEFE3AA3C6" box="[300,435,726,752]" italics="true" pageId="39" pageNumber="40">P. splendida</emphasis>
</taxonomicName>
is not stored in the designated location of the 
<collectionCode id="ED38AE18FFC90728FC4896DEFB91A3C7" box="[938,1048,727,753]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="39" pageNumber="40" type="Museum">NHMUK</collectionCode>
. We selected a 
<typeStatus id="5492887FFFC90728FB5E96DFFAAAA3C6" box="[1212,1315,726,752]" pageId="39" pageNumber="40" type="lectotype">lectotype</typeStatus>
from three 
<typeStatus id="5492887FFFC90728FF7596F2FF70A223" box="[151,249,763,789]" pageId="39" pageNumber="40" type="syntype">syntypes</typeStatus>
(
<figureCitation id="13122A58FFC90728FEEB96F3FEEBA222" box="[265,354,762,788]" captionStart="FIGURE 9" captionStartId="24.[151,250,1194,1218]" captionTargetBox="[155,1433,204,1166]" captionTargetId="figure-23@24.[151,1437,200,1170]" captionTargetPageId="24" captionText="FIGURE 9. Adults of Philodoria species. A P. splendida Walsingham, 1907 lectotype male, B Female, Oahu, C P. lama sp. n. holotype male, D P. napaliensis sp. n. holotype (sex unknown). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4681831" httpUri="https://zenodo.org/record/4681831/files/figure.png" pageId="39" pageNumber="40">Fig. 9A</figureCitation>
), and the remaining 
<typeStatus id="5492887FFFC90728FDAA96F2FD23A223" box="[584,682,763,789]" pageId="39" pageNumber="40" type="syntype">syntypes</typeStatus>
are 
<typeStatus id="5492887FFFC90728FD3896F3FCF6A222" box="[730,895,762,788]" pageId="39" pageNumber="40" type="paralectotype">paralectotypes</typeStatus>
.
</paragraph>
<paragraph id="8B9636DDFFC90728FF259717FB72A71A" blockId="39.[151,1437,150,2042]" pageId="39" pageNumber="40">
<emphasis id="B95DEACFFFC90728FF259717FE37A20E" bold="true" box="[199,446,798,824]" pageId="39" pageNumber="40">Additional material.</emphasis>
28 (14 ♂, 
<specimenCount id="9D2FFD54FFC90728FDA79717FD09A20F" box="[581,640,798,825]" pageId="39" pageNumber="40" type="female">11 ♀</specimenCount>
, 3 sex unknown). 
<emphasis id="B95DEACFFFC90728FCBF9717FC2FA20E" bold="true" box="[861,934,798,824]" pageId="39" pageNumber="40">Kauai</emphasis>
: 1♂, Kauai Na Pali, Kona Forest Reserve, Alakai Swamp trail, 3800’, 
<date id="FF97101DFFC90728FE29974BFDB7A26A" box="[459,574,834,860]" pageId="39" pageNumber="40" value="1973-09-29">29ix.1973</date>
, K. &amp; E. Sattler, B.M.1973-498, 
<taxonomicName id="4C294D5EFFC90728FC4E974BFBACA26A" box="[940,1061,834,860]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEACFFFC90728FC4E974BFBACA26A" box="[940,1061,834,860]" italics="true" pageId="39" pageNumber="40">Philodoria</emphasis>
</taxonomicName>
sp. 12 (Kauai) Sattler Coll., color slide 140, D. C. Lees 
<date id="FF97101DFFC90728FE6F976FFD88A2B6" box="[397,513,870,896]" pageId="39" pageNumber="40" value="2016-09">Sep. 2016</date>
, BMNH(E)1621068. 
<emphasis id="B95DEACFFFC90728FD17976FFCB1A2B6" bold="true" box="[757,824,870,896]" pageId="39" pageNumber="40">Oahu</emphasis>
: 1♂, Waianae, 
<date id="FF97101DFFC90728FC05976FFBCCA2B7" box="[999,1093,870,897]" pageId="39" pageNumber="40" value="1913-01-05">5.i.1913</date>
, ex 
<taxonomicName id="4C294D5EFFC90728FB91976FFF37A292" authority=", SK" authorityName="SK" class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="polymorpha">
<emphasis id="B95DEACFFFC90728FB91976FFA1CA2B6" box="[1139,1429,870,896]" italics="true" pageId="39" pageNumber="40">Metrosideros polymorpha</emphasis>
, SK
</taxonomicName>
697♂, 
<collectionCode id="ED38AE18FFC90728FEE59782FEDCA293" box="[263,341,907,933]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
34316; 1♂, Waianae, 
<date id="FF97101DFFC90728FDA99783FD20A293" box="[587,681,906,933]" pageId="39" pageNumber="40" value="1913-01-05">5.i.1913</date>
, ex Ohia (= 
<taxonomicName id="4C294D5EFFC90728FCD29782FC52A292" box="[816,987,906,932]" class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="polymorpha">
<emphasis id="B95DEACFFFC90728FCD29782FC52A292" box="[816,987,906,932]" italics="true" pageId="39" pageNumber="40">M. polymorpha</emphasis>
</taxonomicName>
) in 
<collectionCode id="ED38AE18FFC90728FBE59783FBECA292" box="[1031,1125,906,932]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40" type="Museum">USNM</collectionCode>
;1♂, 
<specimenCount id="9D2FFD54FFC90728FB6D9783FB3BA292" box="[1167,1202,906,932]" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
, Lauiluli, 
<date id="FF97101DFFC90728FAC79783FA11A293" box="[1317,1432,906,933]" pageId="39" pageNumber="40" value="1914-10-25">25.x.1914</date>
, “Ohia Lehua” (= 
<taxonomicName id="4C294D5EFFC90728FE8297A6FD87A2FE" box="[352,526,942,968]" class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="polymorpha">
<emphasis id="B95DEACFFFC90728FE8297A6FD87A2FE" box="[352,526,942,968]" italics="true" pageId="39" pageNumber="40">M. polymorpha</emphasis>
</taxonomicName>
), 
<collectionCode id="ED38AE18FFC90728FDC597A6FDFCA2FF" box="[551,629,943,969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
34312; 3♂, 
<specimenCount id="9D2FFD54FFC90728FCE597A7FCA3A2FE" box="[775,810,942,968]" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
, Malamalama, 
<date id="FF97101DFFC90728FC3E97A7FBC9A2FE" box="[988,1088,942,968]" pageId="39" pageNumber="40" value="1916-10-08">8.x.1916</date>
, “Ohia Lehua”, 34293–34295, ♂ 
<specimenCount id="9D2FFD54FFC90728FF5597DBFF4CA2DA" box="[183,197,978,1004]" pageId="39" pageNumber="40" type="female">♀</specimenCount>
slide aB 
<date id="FF97101DFFC90728FED197DBFE4BA2DA" box="[307,450,978,1004]" pageId="39" pageNumber="40" value="1941-08-26">
Aug26 
<quantity id="4CD19B38FFC90728FE6897DBFE4BA2DA" box="[394,450,978,1004]" metricMagnitude="1" metricUnit="m" metricValue="4.93014" pageId="39" pageNumber="40" unit="in" value="1941.0">1941</quantity>
</date>
in 
<collectionCode id="ED38AE18FFC90728FE0F97DBFDCBA2DA" box="[493,578,978,1004]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40" type="Museum">USNM</collectionCode>
; 
<specimenCount id="9D2FFD54FFC90728FDB297DBFDFAA2DA" box="[592,627,978,1004]" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
, Konahuanui, 
<quantity id="4CD19B38FFC90728FCFD97DBFCE2A2DA" box="[799,875,978,1004]" metricMagnitude="2" metricUnit="m" metricValue="7.0104000000000015" pageId="39" pageNumber="40" unit="ft" value="2300.0">2300ft</quantity>
, 
<date id="FF97101DFFC90728FC9B97DBFC55A2DA" box="[889,988,978,1004]" pageId="39" pageNumber="40" value="1943-05-09">9.v.1943</date>
, Resting on 
<taxonomicName id="4C294D5EFFC90728FB9097DBFED2A527" authority=", EC Zimmerman Collector, BPBM" authorityName="EC Zimmerman Collector, BPBM" class="Liliopsida" family="Cyperaceae" genus="Cladium" kingdom="Plantae" order="Poales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="genus">Cladium, EC Zimmerman Collector, BPBM</taxonomicName>
34319; 
<specimenCount id="9D2FFD54FFC90728FE5B97FFFE55A526" box="[441,476,1014,1040]" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
, Kalihi, Coll. O.H.S, “Ohia Lehua”, 
<collectionCode id="ED38AE18FFC90728FC6297FFFC59A526" box="[896,976,1014,1040]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
34311; 1 (sex unknown), Olympus, Coll. O.H.S, no. “35” in 
<collectionCode id="ED38AE18FFC90728FE9F9013FE52A502" box="[381,475,1050,1076]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40" type="Museum">USNM</collectionCode>
.1♂, Waialae, Nui, 
<date id="FF97101DFFC90728FD539013FCADA502" box="[689,804,1050,1076]" pageId="39" pageNumber="40" value="1919-02-16">16.ii.1919</date>
, Z-XII-19-62-20♂ in 
<collectionCode id="ED38AE18FFC90728FBC49012FBFEA503" box="[1062,1143,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
. 1 (sex unknown), Kaala Road, 
<date id="FF97101DFFC90728FF029037FEDFA56E" box="[224,342,1086,1112]" pageId="39" pageNumber="40" value="2013-04-23">23.iv.2013</date>
(stored), C.A. Johns leg., Adult on 
<taxonomicName id="4C294D5EFFC90728FD069037FC4AA56E" box="[740,963,1086,1112]" class="Magnoliopsida" family="Rutaceae" genus="Melicope" kingdom="Plantae" order="Sapindales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="clusiifolia">
<emphasis id="B95DEACFFFC90728FD069037FC4AA56E" box="[740,963,1086,1112]" italics="true" pageId="39" pageNumber="40">Melicope clusiifolia</emphasis>
</taxonomicName>
, 
<date id="FF97101DFFC90728FC329037FBC0A56E" box="[976,1097,1086,1112]" pageId="39" pageNumber="40" value="2013-04-21">21.iv.2013</date>
, CJ049, 
<accessionNumber id="947AAB3EFFC90728FB489037FAAAA56E" box="[1194,1315,1086,1112]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KT982402" pageId="39" pageNumber="40">
KT 
<quantity id="4CD19B38FFC90728FB2D9037FAAAA56E" box="[1231,1315,1086,1112]" metricMagnitude="4" metricUnit="m" metricValue="2.49530108" pageId="39" pageNumber="40" unit="in" value="982402.0">982402</quantity>
</accessionNumber>
in 
<collectionCode id="ED38AE18FFC90728FAA59036FA11A56F" box="[1351,1432,1087,1113]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
; 1♂, 
<specimenCount id="9D2FFD54FFC90728FF28906BFF66A54A" box="[202,239,1122,1148]" pageId="39" pageNumber="40" type="female">2♀</specimenCount>
, Kanahuanui, 30.i.&amp; 
<date id="FF97101DFFC90728FE38906BFDC9A54A" box="[474,576,1122,1148]" pageId="39" pageNumber="40" value="2014-02-04">4.ii.2014</date>
(stored), C.A. Johns leg., host: 
<taxonomicName id="4C294D5EFFC90728FC53906AFBD6A54A" box="[945,1119,1122,1148]" class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="polymorpha">
<emphasis id="B95DEACFFFC90728FC53906AFBD6A54A" box="[945,1119,1122,1148]" italics="true" pageId="39" pageNumber="40">M. polymorpha</emphasis>
</taxonomicName>
, 
<date id="FF97101DFFC90728FB92906BFB44A54A" box="[1136,1229,1122,1148]" pageId="39" pageNumber="40" value="2014-01-03">3.i.2014</date>
, CJ246ab, CJ247, SK780♂, 
<specimenCount id="9D2FFD54FFC90728FEE8908FFEC5A596" box="[266,332,1158,1184]" pageId="39" pageNumber="40" type="female">781♀</specimenCount>
, 
<specimenCount id="9D2FFD54FFC90728FEB8908FFE10A596" box="[346,409,1158,1184]" pageId="39" pageNumber="40" type="female">773♀</specimenCount>
in 
<collectionCode id="ED38AE18FFC90728FE23908EFD9BA597" box="[449,530,1159,1185]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
. In 
<collectionCode id="ED38AE18FFC90728FDA2908EFD27A597" box="[576,686,1159,1185]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="39" pageNumber="40" type="Museum">NHMUK</collectionCode>
: 1♂, Oahu 4000’, Waianae Range, Ma Kaala, 
<date id="FF97101DFFC90728FB24908FFABBA596" box="[1222,1330,1158,1184]" pageId="39" pageNumber="40" value="1976-09-04">4.ix.1976</date>
, K. &amp; E. Sattler, B.M.1976-605, 
<taxonomicName id="4C294D5EFFC90728FE4690A3FD94A5F2" box="[420,541,1194,1220]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEACFFFC90728FE4690A3FD94A5F2" box="[420,541,1194,1220]" italics="true" pageId="39" pageNumber="40">Philodoria</emphasis>
</taxonomicName>
sp. 10D (Lanai, Mauai, Oahu) 
<taxonomicName id="4C294D5EFFC90728FC6490A3FC7AA5F2" authority="Sattler Coll., D.C. Lees, Sep. 2016" authorityName="Sattler Coll., D. C. Lees, Sep." authorityYear="2016" box="[902,1011,1194,1220]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="splendida">
<emphasis id="B95DEACFFFC90728FC6490A3FC7AA5F2" box="[902,1011,1194,1220]" italics="true" pageId="39" pageNumber="40">splendida</emphasis>
</taxonomicName>
? Sattler Coll., D.C. Lees, 
<date id="FF97101DFFC90728FAC190A3FA11A5F2" box="[1315,1432,1194,1220]" pageId="39" pageNumber="40" value="2016-09">Sep. 2016</date>
, BMNH(E)1621292; 
<specimenCount id="9D2FFD54FFC90728FE9D90C7FE2BA5DE" box="[383,418,1230,1256]" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
, Oahu 2200’, Waianae Range, above Makua Valley, 
<date id="FF97101DFFC90728FC0E90C7FBECA5DE" box="[1004,1125,1230,1256]" pageId="39" pageNumber="40" value="1976-09-11">11.ix.1976</date>
, K. &amp; E. Sattler, B.M.1976- 605, 
<taxonomicName id="4C294D5EFFC90728FF3290FBFEC0A43A" box="[208,329,1266,1292]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEACFFFC90728FF3290FBFEC0A43A" box="[208,329,1266,1292]" italics="true" pageId="39" pageNumber="40">Philodoria</emphasis>
</taxonomicName>
sp. 10C cf 
<taxonomicName id="4C294D5EFFC90728FE2D90FBFBD0A43A" authority="(Oahu) Sattler Coll., D. C. Lees. Sep. 2016" authorityName="Sattler Coll., D. C. Lees. Sep." authorityYear="2016" baseAuthorityName="Oahu" box="[463,1113,1266,1293]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="lysimachiella">
<emphasis id="B95DEACFFFC90728FE2D90FBFDEDA43A" box="[463,612,1266,1292]" italics="true" pageId="39" pageNumber="40">lysimachiella</emphasis>
(Oahu) Sattler Coll., D. C. Lees. Sep. 2016
</taxonomicName>
, BMNH(E)1621295. 
<emphasis id="B95DEACFFFC90728FAAC90FBFA1DA43A" bold="true" box="[1358,1428,1266,1292]" pageId="39" pageNumber="40">Lanai</emphasis>
: 1♂, 2750’, Munro Trail, 
<date id="FF97101DFFC90728FE4A911FFD85A406" box="[424,524,1302,1328]" pageId="39" pageNumber="40" value="1976-10-02">2.x.1976</date>
, 
<taxonomicName id="4C294D5EFFC90728FDF4911FFD02A406" box="[534,651,1302,1328]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Philodoria</taxonomicName>
sp. 10D (Lanai, Mauai, Oahu) splendida? Sattler Coll., D. C. Lees 
<date id="FF97101DFFC90728FA89911FFF5BA462" pageId="39" pageNumber="40" value="2016-09">Sep. 2016</date>
, BMNH(E) 
<quantity id="4CD19B38FFC90728FEB69133FE5AA462" box="[340,467,1338,1365]" metricMagnitude="4" metricUnit="m" metricValue="4.11906212" pageId="39" pageNumber="40" unit="in" value="1621678.0">1621678 in</quantity>
<collectionCode id="ED38AE18FFC90728FE389132FDCEA463" box="[474,583,1339,1365]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="39" pageNumber="40" type="Museum">NHMUK</collectionCode>
; 
<emphasis id="B95DEACFFFC90728FDB09133FD1BA462" bold="true" box="[594,658,1338,1364]" pageId="39" pageNumber="40">Maui</emphasis>
: 1♂, Haelaau, 
<date id="FF97101DFFC90728FCD89133FC35A462" box="[826,956,1338,1364]" pageId="39" pageNumber="40" value="1928-12-17">17.xii.1928</date>
, O.H. Swezey, 
<quantity id="4CD19B38FFC90728FB859133FB43A462" box="[1127,1226,1338,1364]" metricMagnitude="2" metricUnit="m" metricValue="8.709405999999998" pageId="39" pageNumber="40" unit="in" value="34289.0">34289 in</quantity>
<collectionCode id="ED38AE18FFC90728FB339132FAABA463" box="[1233,1314,1339,1365]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
; 1♂, 
<specimenCount id="9D2FFD54FFC90728FABC9133FA09A462" box="[1374,1408,1338,1364]" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
, 1 (sex unknown), Flemming Arboretum Trail, [
<date id="FF97101DFFC90728FD709157FD7CA44E" box="[658,757,1374,1400]" pageId="39" pageNumber="40" value="2013-05-03">3.v.2013</date>
(stored), C.A. Johns leg., host: 
<taxonomicName id="4C294D5EFFC90728FBB99156FA8EA44E" box="[1115,1287,1374,1400]" class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="polymorpha">
<emphasis id="B95DEACFFFC90728FBB99156FA8EA44E" box="[1115,1287,1374,1400]" italics="true" pageId="39" pageNumber="40">M. polymorpha</emphasis>
</taxonomicName>
, 12.iv.]2013, SK 
<specimenCount id="9D2FFD54FFC90728FF22918BFF74A4AA" box="[192,253,1410,1436]" pageId="39" pageNumber="40" type="female">621♀</specimenCount>
, SK696♂, CJ105 (sex unknown) / 
<accessionNumber id="947AAB3EFFC90728FD7B918BFC9BA4AA" box="[665,786,1410,1436]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KT982410" pageId="39" pageNumber="40">
KT 
<quantity id="4CD19B38FFC90728FD5C918BFC9BA4AA" box="[702,786,1410,1436]" metricMagnitude="4" metricUnit="m" metricValue="2.4953213999999995" pageId="39" pageNumber="40" unit="in" value="982410.0">982410</quantity>
</accessionNumber>
in 
<collectionCode id="ED38AE18FFC90728FCDF918AFC07A4AB" box="[829,910,1411,1437]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
. 
<specimenCount id="9D2FFD54FFC90728FC7E918BFC36A4AA" box="[924,959,1410,1436]" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
, W. Mauai, 3000’, W. Maui Forest Res., Kaulalewelewe (Puu Kukui Trail), 
<date id="FF97101DFFC90728FDC291AFFD23A4F6" box="[544,682,1446,1472]" pageId="39" pageNumber="40" value="1976-08-26">26.viii.1976</date>
, K. &amp; E. Sattler, B.M.1976-605, 
<taxonomicName id="4C294D5EFFC90728FBFF91AFFB1BA4F6" box="[1053,1170,1446,1472]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Philodoria</taxonomicName>
sp. 10D (Lanai, Mauai, Oahu) splendida? Sattler Coll., D. C. Lees 
<date id="FF97101DFFC90728FD9D91C3FD7AA4D2" box="[639,755,1482,1508]" pageId="39" pageNumber="40" value="2016-09">Sep. 2016</date>
, BMNH(E) 
<quantity id="4CD19B38FFC90728FC9491C3FC7EA4D2" box="[886,1015,1482,1509]" metricMagnitude="4" metricUnit="m" metricValue="4.1178683199999995" pageId="39" pageNumber="40" unit="in" value="1621208.0">1621208 in</quantity>
<collectionCode id="ED38AE18FFC90728FC1D91C3FBE5A4D2" box="[1023,1132,1482,1508]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="39" pageNumber="40" type="Museum">NHMUK</collectionCode>
; 
<emphasis id="B95DEACFFFC90728FB9A91C3FA99A4D2" bold="true" box="[1144,1296,1482,1508]" pageId="39" pageNumber="40">
<taxonomicName id="4C294D5EFFC90728FB9A91C3FB6FA4D2" box="[1144,1254,1482,1508]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Myrtales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="B95DEACFFFC90728FB9A91C3FB6FA4D2" bold="true" box="[1144,1254,1482,1508]" italics="true" pageId="39" pageNumber="40">Syzygium</emphasis>
</taxonomicName>
sp.
</emphasis>
, C.A. Johns leg., in 
<collectionCode id="ED38AE18FFC90728FF1291E7FECBA73E" box="[240,322,1518,1544]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="39" pageNumber="40" type="Museum">BPBM</collectionCode>
: 
<specimenCount id="9D2FFD54FFC90728FEAC91E4FEFAA731" box="[334,371,1517,1543]" pageId="39" pageNumber="40" type="female">2♀</specimenCount>
, Limahuli, Kauai, 
<date id="FF97101DFFC90728FDA991E4FD42A73E" box="[587,715,1517,1544]" pageId="39" pageNumber="40" value="2015-07-21">21.vii.2015</date>
(stored), 
<date id="FF97101DFFC90728FCD991E4FC3FA73E" box="[827,950,1517,1544]" pageId="39" pageNumber="40" value="2015-06-20">20.vi.2015</date>
, CJ488ab, SK 
<specimenCount id="9D2FFD54FFC90728FBBC91E4FB12A731" box="[1118,1179,1517,1543]" pageId="39" pageNumber="40" type="female">748♀</specimenCount>
, SK 
<specimenCount id="9D2FFD54FFC90728FB3291E4FA87A73E" box="[1232,1294,1517,1544]" pageId="39" pageNumber="40" type="female">752♀</specimenCount>
; 1♂, Waiaai Trail, Oahu, 
<date id="FF97101DFFC90728FEC79218FE2CA71D" box="[293,421,1553,1579]" pageId="39" pageNumber="40" value="2014-07-30">30.vii.2014</date>
(stored in RNAlater solution), 
<date id="FF97101DFFC90728FCE19219FCFEA71D" box="[771,887,1552,1579]" pageId="39" pageNumber="40" value="2014-07-05">5.vii.2014</date>
, CJ312, SK747♂ (only abdomen).
</paragraph>
</subSubSection>
<subSubSection id="C3336556FFC90728FF25923DFA94A7EA" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="8B9636DDFFC90728FF25923DFA94A7EA" blockId="39.[151,1437,150,2042]" pageId="39" pageNumber="40">
<emphasis id="B95DEACFFFC90728FF25923DFECBA778" bold="true" box="[199,322,1588,1614]" pageId="39" pageNumber="40">Diagnosis.</emphasis>
This species and 
<taxonomicName id="4C294D5EFFC90728FDEF923CFDEBA778" box="[525,610,1588,1614]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="lama" status="sp. nov.">
<emphasis id="B95DEACFFFC90728FDEF923CFDEBA778" box="[525,610,1588,1614]" italics="true" pageId="39" pageNumber="40">P. lama</emphasis>
</taxonomicName>
<emphasis id="B95DEACFFFC90728FD88923DFD22A778" bold="true" box="[618,683,1588,1614]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="A26E57B4FFC90728FD88923DFD22A778" box="[618,683,1588,1614]" pageId="39" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
have a brownish ocherous basal oblique fascia on forewing (
<figureCitation id="13122A58FFC90728FA86923DFF55A744" captionStart="FIGURE 9" captionStartId="24.[151,250,1194,1218]" captionTargetBox="[155,1433,204,1166]" captionTargetId="figure-23@24.[151,1437,200,1170]" captionTargetPageId="24" captionText="FIGURE 9. Adults of Philodoria species. A P. splendida Walsingham, 1907 lectotype male, B Female, Oahu, C P. lama sp. n. holotype male, D P. napaliensis sp. n. holotype (sex unknown). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4681831" httpUri="https://zenodo.org/record/4681831/files/figure.png" pageId="39" pageNumber="40">Figs. 9A, B</figureCitation>
, 
<figureCitation id="13122A58FFC90728FF059251FEB1A744" box="[231,312,1624,1650]" captionStart="FIGURE 73" captionStartId="125.[151,250,1698,1722]" captionTargetBox="[159,1429,241,1669]" captionTargetId="figure-57@125.[151,1436,237,1675]" captionTargetPageId="125" captionText="FIGURE 73. Resting posture of adult Philodoria and Caloptilia. A P. molokaiensis, Molokai, CJ229, host: Lysimachia hillebrandii, B P. splendida, Oahu, CJ319, host: Metrosideros polymorpha, C Same species, Oahu, D Caloptilia mabaella, Lanai, CJ333, host: Diospyros sandwicensis, E P. lama, Maui, CJ190, F Same species, Maui, CJ194." figureDoi="http://doi.org/10.5281/zenodo.4681967" httpUri="https://zenodo.org/record/4681967/files/figure.png" pageId="39" pageNumber="40">73B, C</figureCitation>
). Genital features very similar to those of 
<taxonomicName id="4C294D5EFFC90728FCE89251FCFDA747" box="[778,884,1623,1649]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="B95DEACFFFC90728FCE89251FCFDA747" box="[778,884,1623,1649]" italics="true" pageId="39" pageNumber="40">P. basalis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C294D5EFFC90728FC4B9251FC75A747" box="[937,1020,1623,1649]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="lama">
<emphasis id="B95DEACFFFC90728FC4B9251FC75A747" box="[937,1020,1623,1649]" italics="true" pageId="39" pageNumber="40">P. lama</emphasis>
</taxonomicName>
, but distinguished by a rather slender dorsal process of valva in male. Kauai specimens (CJ488, male unknown) on 
<taxonomicName id="4C294D5EFFC90728FBE99272FBFCA7A3" box="[1035,1141,1659,1685]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Myrtales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="B95DEACFFFC90728FBE99272FBFCA7A3" box="[1035,1141,1659,1685]" italics="true" pageId="39" pageNumber="40">Syzygium</emphasis>
</taxonomicName>
sp. have similar forewing pattern (
<figureCitation id="13122A58FFC90728FF169297FEFEA78F" box="[244,375,1694,1721]" captionStart="FIGURE 26" captionStartId="49.[151,250,1720,1744]" captionTargetBox="[195,1392,185,1692]" captionTargetId="figure-20@49.[191,1395,181,1696]" captionTargetPageId="49" captionText="FIGURE 26. Forewing pattern variation in Philodoria. Specimens from recent collection efforts that were preserved in ethanol or RNA stabilization reagent, originally for molecular studies. Wings were placed without mountant under a coverslip. A P. succedanea (sex unknown, CJ419), B P. auromagnifica, male (CJ148/SK689), C P. basalis, female (CJ410), D P. splendida, female (CJ488a), host: Syzygium sp., E P. splendida (sex unknown, CJ488b), host: Syzygium sp., F P. molokaiensis, male (CJ509b/ SK667), G P. sp. A nr. P. lysimachiella, female (CJ346a/SK753), H P. lysimachiella, female (CJ530), lateral view, I P. pipturiella, female (CJ529), J Same species, male (CJ529), K P. haelaauensis, female (CJ209), L Same species, female (CJ216), M P. floscula, female (CJ426/SK845), N P. wilkesiella, male (CJ009/SK649), O P. epibathra, male (CJ506a/SK678), P Same species, female (CJ506b/SK681). Arrows point to diagnostic characters of species: black patch at base (A), basal fascia (D), transverse fasciae (tf) (F, G), white spot (ws) (I, J), white patch (K–M). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4681867" httpUri="https://zenodo.org/record/4681867/files/figure.png" pageId="39" pageNumber="40">Fig. 26D, E</figureCitation>
) and female genitalia, but the Oahu specimen (CJ312, thorax sacrificed for molecular analysis) have thickened dorsal process with slightly curved dorsal side (
<figureCitation id="13122A58FFC90728FCBE92CBFC4BA7EA" box="[860,962,1730,1756]" captionStart="FIGURE 41" captionStartId="76.[151,250,1685,1709]" captionTargetBox="[161,1426,183,1654]" captionTargetId="figure-23@76.[151,1436,181,1662]" captionTargetPageId="76" captionText="FIGURE 41. Male valva of Philodoria, inner view. A–F, H–J, L, M: Left valva, G, K, N: Right valva. A P. succedanea, Maui, SK641, B P. auromagnifica, Hawaii (Big Island), SK800, C P. auromagnifica, Kauai, SK689, D P. kolea, holotype, Hawaii (Big Island), SK851, E P. lysimachiella, Oahu, SK656, F, G P. sp. 7, Kauai, SK754, H P. molokaiensis, Molokai, SK666, I P. basalis, Hawaii (Big Island), SK614, J P. splendida, Maui, SK696, K Same species, Oahu, SK747, host Syzygium sp., L P. lama, holotype, Maui SK744, M P. napaliensis, paratype, Kauai, SK767, N Same as L, paratype, Kauai, SK745." figureDoi="http://doi.org/10.5281/zenodo.4681901" httpUri="https://zenodo.org/record/4681901/files/figure.png" pageId="39" pageNumber="40">Fig. 41K</figureCitation>
) and slender saccus (
<figureCitation id="13122A58FFC90728FB5392CBFA98A7EA" box="[1201,1297,1730,1756]" captionStart="FIGURE 42" captionStartId="77.[151,250,1773,1797]" captionTargetBox="[168,1421,187,1741]" captionTargetId="figure-20@77.[158,1430,181,1748]" captionTargetPageId="77" captionText="FIGURE 42. Genital capsule of Philodoria, excluding right portion of male genitalia, ventral view. A P. succedanea, Maui, SK641, B P. auromagnifica, Hawaii (Big Island), SK800, C P. auromagnifica, Kauai, SK689, D P. kolea, holotype, Hawaii (Big Island), SK851, E P. lysimachiella, Oahu, SK656, F P. sp. 7, Kauai, SK754, G P. molokaiensis, Molokai, SK666, H P. basalis, Hawaii (Big Island), SK614, I P. splendida, Maui, SK696, J Same species, Oahu, SK747, host Syzygium sp., K P. lama, holotype, Maui, SK744, L P. napaliensis, paratype, Kauai, SK745. Abbreviation: ta: tuba analis." figureDoi="http://doi.org/10.5281/zenodo.4681903" httpUri="https://zenodo.org/record/4681903/files/figure.png" pageId="39" pageNumber="40">Fig. 42J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3336556FFC90727FF2592ECFD3EA6C3" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
<paragraph id="8B9636DDFFC90727FF2592ECFACAA62B" blockId="39.[151,1437,150,2042]" lastBlockId="40.[151,1437,1759,2037]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<emphasis id="B95DEACFFFC90728FF2592ECFE49A7C9" bold="true" box="[199,448,1765,1791]" pageId="39" pageNumber="40">Redescription: Adult</emphasis>
(
<figureCitation id="13122A58FFC90728FE3292ECFDDCA636" box="[464,597,1765,1792]" captionStart="FIGURE 9" captionStartId="24.[151,250,1194,1218]" captionTargetBox="[155,1433,204,1166]" captionTargetId="figure-23@24.[151,1437,200,1170]" captionTargetPageId="24" captionText="FIGURE 9. Adults of Philodoria species. A P. splendida Walsingham, 1907 lectotype male, B Female, Oahu, C P. lama sp. n. holotype male, D P. napaliensis sp. n. holotype (sex unknown). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4681831" httpUri="https://zenodo.org/record/4681831/files/figure.png" pageId="39" pageNumber="40">Figs. 9A, B</figureCitation>
, 
<figureCitation id="13122A58FFC90728FD8392ECFD3BA636" box="[609,690,1765,1792]" captionStart="FIGURE 26" captionStartId="49.[151,250,1720,1744]" captionTargetBox="[195,1392,185,1692]" captionTargetId="figure-20@49.[191,1395,181,1696]" captionTargetPageId="49" captionText="FIGURE 26. Forewing pattern variation in Philodoria. Specimens from recent collection efforts that were preserved in ethanol or RNA stabilization reagent, originally for molecular studies. Wings were placed without mountant under a coverslip. A P. succedanea (sex unknown, CJ419), B P. auromagnifica, male (CJ148/SK689), C P. basalis, female (CJ410), D P. splendida, female (CJ488a), host: Syzygium sp., E P. splendida (sex unknown, CJ488b), host: Syzygium sp., F P. molokaiensis, male (CJ509b/ SK667), G P. sp. A nr. P. lysimachiella, female (CJ346a/SK753), H P. lysimachiella, female (CJ530), lateral view, I P. pipturiella, female (CJ529), J Same species, male (CJ529), K P. haelaauensis, female (CJ209), L Same species, female (CJ216), M P. floscula, female (CJ426/SK845), N P. wilkesiella, male (CJ009/SK649), O P. epibathra, male (CJ506a/SK678), P Same species, female (CJ506b/SK681). Arrows point to diagnostic characters of species: black patch at base (A), basal fascia (D), transverse fasciae (tf) (F, G), white spot (ws) (I, J), white patch (K–M). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4681867" httpUri="https://zenodo.org/record/4681867/files/figure.png" pageId="39" pageNumber="40">26D, E</figureCitation>
, 
<figureCitation id="13122A58FFC90728FD5C92EFFC98A636" box="[702,785,1766,1792]" captionStart="FIGURE 73" captionStartId="125.[151,250,1698,1722]" captionTargetBox="[159,1429,241,1669]" captionTargetId="figure-57@125.[151,1436,237,1675]" captionTargetPageId="125" captionText="FIGURE 73. Resting posture of adult Philodoria and Caloptilia. A P. molokaiensis, Molokai, CJ229, host: Lysimachia hillebrandii, B P. splendida, Oahu, CJ319, host: Metrosideros polymorpha, C Same species, Oahu, D Caloptilia mabaella, Lanai, CJ333, host: Diospyros sandwicensis, E P. lama, Maui, CJ190, F Same species, Maui, CJ194." figureDoi="http://doi.org/10.5281/zenodo.4681967" httpUri="https://zenodo.org/record/4681967/files/figure.png" pageId="39" pageNumber="40">73B, C</figureCitation>
). Wingspan 
<quantity id="4CD19B38FFC90728FC7E92EFFC70A636" box="[924,1017,1766,1792]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="39" pageNumber="40" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
in type series; forewing length 
<quantity id="4CD19B38FFC90728FABE92EFFA14A636" box="[1372,1437,1766,1792]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="39" pageNumber="40" unit="mm" value="4.0">4 mm</quantity>
in 
<typeStatus id="5492887FFFC90728FF569300FE90A615" box="[180,281,1801,1827]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
, 
<quantity id="4CD19B38FFC90728FEC69300FEF3A615" box="[292,378,1801,1827]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="39" pageNumber="40" unit="mm" value="3.9">3.9 mm</quantity>
in 
<typeStatus id="5492887FFFC90728FE7C9300FD8BA615" box="[414,514,1801,1827]" pageId="39" pageNumber="40" type="paratype">paratype</typeStatus>
. Head dark brown; frons white; maxillary palpus reduced; labial palpus white, externally fuscous at apical portion. Antenna grayish fuscous. Thorax leaden gray. Forewing shining, pale leaden gray with dark brownish ocherous patches enclosing gray fascia: 
<emphasis id="B95DEACFFFC90728FCDC9359FCD4A65C" bold="true" box="[830,861,1872,1898]" pageId="39" pageNumber="40">bp</emphasis>
from near base of costal fold to dorsal 1/4; 
<emphasis id="B95DEACFFFC90728FAAE9346FAD6A65F" bold="true" box="[1356,1375,1871,1897]" pageId="39" pageNumber="40">tf</emphasis>
from costal 1/3 to dorsal 1/2, 
<emphasis id="B95DEACFFFC90728FE4C937DFE4EA6B8" bold="true" box="[430,455,1908,1934]" pageId="39" pageNumber="40">tp</emphasis>
after the middle to apical portion, distinctly narrowing in the dorsum, containing two short, shining, steel gray costal spot; apical portion with large jet-black terminal patch from tornus to apex, containing two upright steel-gray streaks near its opposite extremities; a single bright silvery dot in its middle; a very narrow ocherous line along the base of tornus; cilia shiny, leaden gray with a black fringe line; two white costal lines, long one extend apex; terminal cilia often black. Hindwing coppery brown; cilia tawny. Abdomen tawny brown, silver below. Legs grayish fuscous, with white spots on tarsal joints and at distal end of tibiae; spurs white.
</paragraph>
<caption id="DF566655FFC60727FF75925FFE07A783" ID-DOI="http://doi.org/10.5281/zenodo.4681855" ID-Zenodo-Dep="4681855" httpUri="https://zenodo.org/record/4681855/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,250,1622,1646]" targetBox="[169,1418,184,1593]" targetPageId="40">
<paragraph id="8B9636DDFFC60727FF75925FFE07A783" blockId="40.[151,1437,1622,1718]" pageId="40" pageNumber="41">
<emphasis id="B95DEACFFFC60727FF75925FFE96A758" bold="true" box="[151,287,1622,1646]" pageId="40" pageNumber="41">FIGURE 21.</emphasis>
Adults of 
<taxonomicName id="4C294D5EFFC60727FE6A925FFE7FA758" box="[392,502,1622,1646]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEACFFFC60727FE6A925FFE7FA758" box="[392,502,1622,1646]" italics="true" pageId="40" pageNumber="41">Philodoria</emphasis>
</taxonomicName>
species. 
<emphasis id="B95DEACFFFC60727FDB2925FFDEBA758" bold="true" box="[592,610,1622,1646]" pageId="40" pageNumber="41">A</emphasis>
Wings of 
<taxonomicName id="4C294D5EFFC60727FD2B925EFCACA758" box="[713,805,1622,1646]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="funkae">
<emphasis id="B95DEACFFFC60727FD2B925EFCACA758" box="[713,805,1622,1646]" italics="true" pageId="40" pageNumber="41">P. funkae</emphasis>
</taxonomicName>
sp n., holotype female, 
<emphasis id="B95DEACFFFC60727FBF1925FFBADA758" bold="true" box="[1043,1060,1622,1646]" pageId="40" pageNumber="41">B</emphasis>
Right forewing of holotype shown in A, 
<emphasis id="B95DEACFFFC60727FF549273FF41A7A4" bold="true" box="[182,200,1658,1682]" pageId="40" pageNumber="41">C</emphasis>
Wings of paratype female, 
<emphasis id="B95DEACFFFC60727FE009273FE7DA7A4" bold="true" box="[482,500,1658,1682]" pageId="40" pageNumber="41">D</emphasis>
<taxonomicName id="4C294D5EFFC60727FE199272FCB1A7A4" authority="(Swezey, 1928)" baseAuthorityName="Swezey" baseAuthorityYear="1928" box="[507,824,1658,1682]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pittosporella">
<emphasis id="B95DEACFFFC60727FE199272FD1CA7A4" box="[507,661,1658,1682]" italics="true" pageId="40" pageNumber="41">P. pittosporella</emphasis>
(
<bibRefCitation id="EFB84B2CFFC60727FD409273FCB8A7A4" author="Swezey, O. H." box="[674,817,1658,1682]" pageId="40" pageNumber="41" pagination="187 - 191" refId="ref102595" refString="Swezey, O. H. (1928) Some new species of lepidopterous leaf-miners in Hawaii. Proceedings of the Hawaiian Entomological Society, 7 (1), 187 - 191." type="journal article" year="1928">Swezey, 1928</bibRefCitation>
)
</taxonomicName>
holotype male, 
<emphasis id="B95DEACFFFC60727FC3E9273FC64A7A4" bold="true" box="[988,1005,1658,1682]" pageId="40" pageNumber="41">E</emphasis>
<taxonomicName id="4C294D5EFFC60727FC169272FAB0A7A4" authority="(Swezey, 1928)" baseAuthorityName="Swezey" baseAuthorityYear="1928" box="[1012,1337,1658,1682]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="touchardiella">
<emphasis id="B95DEACFFFC60727FC169272FB1CA7A4" box="[1012,1173,1658,1682]" italics="true" pageId="40" pageNumber="41">P. touchardiella</emphasis>
(
<bibRefCitation id="EFB84B2CFFC60727FB409273FABBA7A4" author="Swezey, O. H." box="[1186,1330,1658,1682]" pageId="40" pageNumber="41" pagination="187 - 191" refId="ref102595" refString="Swezey, O. H. (1928) Some new species of lepidopterous leaf-miners in Hawaii. Proceedings of the Hawaiian Entomological Society, 7 (1), 187 - 191." type="journal article" year="1928">Swezey, 1928</bibRefCitation>
)
</taxonomicName>
lectotype female. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="8B9636DDFFC60727FF25932EFF44A69B" blockId="40.[151,1437,1759,2037]" pageId="40" pageNumber="41">
<emphasis id="B95DEACFFFC60727FF25932EFEFBA677" bold="true" box="[199,370,1831,1857]" pageId="40" pageNumber="41">Male genitalia</emphasis>
(
<figureCitation id="13122A58FFC60727FE62932EFD82A677" box="[384,523,1831,1857]" captionStart="FIGURE 41" captionStartId="76.[151,250,1685,1709]" captionTargetBox="[161,1426,183,1654]" captionTargetId="figure-23@76.[151,1436,181,1662]" captionTargetPageId="76" captionText="FIGURE 41. Male valva of Philodoria, inner view. A–F, H–J, L, M: Left valva, G, K, N: Right valva. A P. succedanea, Maui, SK641, B P. auromagnifica, Hawaii (Big Island), SK800, C P. auromagnifica, Kauai, SK689, D P. kolea, holotype, Hawaii (Big Island), SK851, E P. lysimachiella, Oahu, SK656, F, G P. sp. 7, Kauai, SK754, H P. molokaiensis, Molokai, SK666, I P. basalis, Hawaii (Big Island), SK614, J P. splendida, Maui, SK696, K Same species, Oahu, SK747, host Syzygium sp., L P. lama, holotype, Maui SK744, M P. napaliensis, paratype, Kauai, SK767, N Same as L, paratype, Kauai, SK745." figureDoi="http://doi.org/10.5281/zenodo.4681901" httpUri="https://zenodo.org/record/4681901/files/figure.png" pageId="40" pageNumber="41">Figs. 41J, K</figureCitation>
, 
<figureCitation id="13122A58FFC60727FDF4932EFDDFA677" box="[534,598,1831,1857]" captionStart="FIGURE 42" captionStartId="77.[151,250,1773,1797]" captionTargetBox="[168,1421,187,1741]" captionTargetId="figure-20@77.[158,1430,181,1748]" captionTargetPageId="77" captionText="FIGURE 42. Genital capsule of Philodoria, excluding right portion of male genitalia, ventral view. A P. succedanea, Maui, SK641, B P. auromagnifica, Hawaii (Big Island), SK800, C P. auromagnifica, Kauai, SK689, D P. kolea, holotype, Hawaii (Big Island), SK851, E P. lysimachiella, Oahu, SK656, F P. sp. 7, Kauai, SK754, G P. molokaiensis, Molokai, SK666, H P. basalis, Hawaii (Big Island), SK614, I P. splendida, Maui, SK696, J Same species, Oahu, SK747, host Syzygium sp., K P. lama, holotype, Maui, SK744, L P. napaliensis, paratype, Kauai, SK745. Abbreviation: ta: tuba analis." figureDoi="http://doi.org/10.5281/zenodo.4681903" httpUri="https://zenodo.org/record/4681903/files/figure.png" pageId="40" pageNumber="41">42I, J</figureCitation>
, 
<figureCitation id="13122A58FFC60727FD80932EFD2BA677" box="[610,674,1831,1857]" captionStart="FIGURE 43" captionStartId="78.[151,250,1790,1814]" captionTargetBox="[159,1418,191,1759]" captionTargetId="figure-23@78.[151,1436,181,1766]" captionTargetPageId="78" captionText="FIGURE 43. Phallus of Philodoria, lateral view. A P. succedanea, Maui, SK641, B P. auromagnifica, Hawaii (Big Island), SK800, C P. auromagnifica, Kauai, SK689, D P. kolea, holotype, Hawaii (Big Island), SK851, E P. lysimachiella, Oahu, SK656, F P. sp. 7, Kauai, SK755, G P. molokaiensis, Molokai, SK666, H P. basalis, Hawaii (Big Island), SK614, I P. splendida, Maui, SK696, J Same species, Oahu, SK747, host Syzygium sp., K P. lama, holotype, Maui, SK744, L P. napaliensis, paratype, Kauai, SK767." figureDoi="http://doi.org/10.5281/zenodo.4681905" httpUri="https://zenodo.org/record/4681905/files/figure.png" pageId="40" pageNumber="41">43I, J</figureCitation>
, 
<figureCitation id="13122A58FFC60727FD4C932EFD7FA677" box="[686,758,1831,1857]" captionStart="FIGURE 64" captionStartId="115.[151,250,1375,1399]" captionTargetBox="[167,1422,200,1348]" captionTargetId="figure-20@115.[162,1425,198,1351]" captionTargetPageId="115" captionText="FIGURE 64. BPBM genitalia slides prepared by E. C. Zimmerman. A–D: P. haelaauensis, paratype male, Maui, Z-XII-19-62- 23, E, F: P. pipturiella paratype female, Oahu, Z-XII-1962-22, G–I: P. splendida, male, Oahu, Z-XII-19-62-20. A Vinculum and valvae, tegumen broken, B, H Phallus, C Tegumen? D, I Abdominal segments, E Genitalia opened from left lateral side, F Abdominal segments II–VI, G Genitalia excluding phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4681949" httpUri="https://zenodo.org/record/4681949/files/figure.png" pageId="40" pageNumber="41">64G, I</figureCitation>
) (n=3). Capsule 
<quantity id="4CD19B38FFC60727FC51932EFB85A677" box="[947,1036,1831,1857]" metricMagnitude="-1" metricUnit="m" metricValue="8.3" pageId="40" pageNumber="41" unit="mm" value="830.0">830 µm</quantity>
. Tegumen same length as valva and rather wide; valva 
<quantity id="4CD19B38FFC60727FE889342FE4BA653" box="[362,450,1867,1893]" metricMagnitude="-1" metricUnit="m" metricValue="4.9" pageId="40" pageNumber="41" unit="mm" value="490.0">490 µm</quantity>
long, similar to 
<taxonomicName id="4C294D5EFFC60727FD9F9345FD61A653" box="[637,744,1867,1893]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="B95DEACFFFC60727FD9F9345FD61A653" box="[637,744,1867,1893]" italics="true" pageId="40" pageNumber="41">P. basalis</emphasis>
</taxonomicName>
except dorsal process rather slender (
<figureCitation id="13122A58FFC60727FB719342FB7BA653" box="[1171,1266,1867,1893]" captionStart="FIGURE 41" captionStartId="76.[151,250,1685,1709]" captionTargetBox="[161,1426,183,1654]" captionTargetId="figure-23@76.[151,1436,181,1662]" captionTargetPageId="76" captionText="FIGURE 41. Male valva of Philodoria, inner view. A–F, H–J, L, M: Left valva, G, K, N: Right valva. A P. succedanea, Maui, SK641, B P. auromagnifica, Hawaii (Big Island), SK800, C P. auromagnifica, Kauai, SK689, D P. kolea, holotype, Hawaii (Big Island), SK851, E P. lysimachiella, Oahu, SK656, F, G P. sp. 7, Kauai, SK754, H P. molokaiensis, Molokai, SK666, I P. basalis, Hawaii (Big Island), SK614, J P. splendida, Maui, SK696, K Same species, Oahu, SK747, host Syzygium sp., L P. lama, holotype, Maui SK744, M P. napaliensis, paratype, Kauai, SK767, N Same as L, paratype, Kauai, SK745." figureDoi="http://doi.org/10.5281/zenodo.4681901" httpUri="https://zenodo.org/record/4681901/files/figure.png" pageId="40" pageNumber="41">Fig. 41J</figureCitation>
). Saccus broad and straight (
<figureCitation id="13122A58FFC60727FECB9366FE0FA6BF" box="[297,390,1903,1929]" captionStart="FIGURE 42" captionStartId="77.[151,250,1773,1797]" captionTargetBox="[168,1421,187,1741]" captionTargetId="figure-20@77.[158,1430,181,1748]" captionTargetPageId="77" captionText="FIGURE 42. Genital capsule of Philodoria, excluding right portion of male genitalia, ventral view. A P. succedanea, Maui, SK641, B P. auromagnifica, Hawaii (Big Island), SK800, C P. auromagnifica, Kauai, SK689, D P. kolea, holotype, Hawaii (Big Island), SK851, E P. lysimachiella, Oahu, SK656, F P. sp. 7, Kauai, SK754, G P. molokaiensis, Molokai, SK666, H P. basalis, Hawaii (Big Island), SK614, I P. splendida, Maui, SK696, J Same species, Oahu, SK747, host Syzygium sp., K P. lama, holotype, Maui, SK744, L P. napaliensis, paratype, Kauai, SK745. Abbreviation: ta: tuba analis." figureDoi="http://doi.org/10.5281/zenodo.4681903" httpUri="https://zenodo.org/record/4681903/files/figure.png" pageId="40" pageNumber="41">Fig. 42I</figureCitation>
). Phallus 
<quantity id="4CD19B38FFC60727FE139366FDCEA6BF" box="[497,583,1903,1929]" metricMagnitude="-1" metricUnit="m" metricValue="6.6" pageId="40" pageNumber="41" unit="mm" value="660.0">660 µm</quantity>
long, sinuous in lateral view; cornuti large and well developed apically (
<figureCitation id="13122A58FFC60727FA8D9366FF49A69B" captionStart="FIGURE 43" captionStartId="78.[151,250,1790,1814]" captionTargetBox="[159,1418,191,1759]" captionTargetId="figure-23@78.[151,1436,181,1766]" captionTargetPageId="78" captionText="FIGURE 43. Phallus of Philodoria, lateral view. A P. succedanea, Maui, SK641, B P. auromagnifica, Hawaii (Big Island), SK800, C P. auromagnifica, Kauai, SK689, D P. kolea, holotype, Hawaii (Big Island), SK851, E P. lysimachiella, Oahu, SK656, F P. sp. 7, Kauai, SK755, G P. molokaiensis, Molokai, SK666, H P. basalis, Hawaii (Big Island), SK614, I P. splendida, Maui, SK696, J Same species, Oahu, SK747, host Syzygium sp., K P. lama, holotype, Maui, SK744, L P. napaliensis, paratype, Kauai, SK767." figureDoi="http://doi.org/10.5281/zenodo.4681905" httpUri="https://zenodo.org/record/4681905/files/figure.png" pageId="40" pageNumber="41">Fig. 43I</figureCitation>
).
</paragraph>
<paragraph id="8B9636DDFFC60727FF2593BEFD3EA6C3" blockId="40.[151,1437,1759,2037]" pageId="40" pageNumber="41">
<emphasis id="B95DEACFFFC60727FF2593BEFE05A6E7" bold="true" box="[199,396,1975,2001]" pageId="40" pageNumber="41">Female genitalia</emphasis>
(
<figureCitation id="13122A58FFC60727FE7893BEFE76A6E4" box="[410,511,1975,2002]" captionStart="FIGURE 56" captionStartId="106.[151,250,1881,1905]" captionTargetBox="[170,1411,192,1821]" captionTargetId="figure-23@106.[165,1422,181,1857]" captionTargetPageId="106" captionText="FIGURE 56. Female genitalia of Philodoria. A P. succedanea, paralectotype, Maui, SK714, B Same species, Hawaii (Big Island), SK624, C P. auromagnifica, Hawaii (Big Island), SK623, D P. kauaulaensis, holotype, SK690, E Same species, lateral view, F P. kolea, paratype, Hawaii (Big Island), SK634, G P. lysimachiella, SK654, H P. sp. A, SK753, I P. molokaiensis, SK668, lateral view, J Same ventral view, K P. splendida, SK621, L P. lama, paratype, SK813." figureDoi="http://doi.org/10.5281/zenodo.4681931" httpUri="https://zenodo.org/record/4681931/files/figure.png" pageId="40" pageNumber="41">Fig. 56K</figureCitation>
) (n=4). 
<quantity id="4CD19B38FFC60727FDBB93BEFD37A6E7" box="[601,702,1975,2001]" metricMagnitude="0" metricUnit="m" metricValue="1.4" pageId="40" pageNumber="41" unit="mm" value="1400.0">1400 µm</quantity>
long. Lamella antevaginalis 
<quantity id="4CD19B38FFC60727FC1D93BEFBDFA6E7" box="[1023,1110,1975,2001]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="41" unit="mm" value="200.0">200 µm</quantity>
in length. Corpus bursae 
<quantity id="4CD19B38FFC60727FA9093BEFF34A6C3" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="40" pageNumber="41" unit="mm" value="800.0">800 µm</quantity>
long. Similar in the appearance to 
<taxonomicName id="4C294D5EFFC60727FDA793D5FD26A6C3" box="[581,687,2011,2037]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="B95DEACFFFC60727FDA793D5FD26A6C3" box="[581,687,2011,2037]" italics="true" pageId="40" pageNumber="41">P. basalis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF566655FFC70726FF7590A6FACAA5DD" ID-DOI="http://doi.org/10.5281/zenodo.4681857" ID-Zenodo-Dep="4681857" httpUri="https://zenodo.org/record/4681857/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,250,1199,1223]" targetBox="[155,1432,186,1146]" targetPageId="41">
<paragraph id="8B9636DDFFC70726FF7590A6FACAA5DD" blockId="41.[151,1437,1199,1259]" pageId="41" pageNumber="42">
<emphasis id="B95DEACFFFC70726FF7590A6FEA8A5F1" bold="true" box="[151,289,1199,1223]" pageId="41" pageNumber="42">FIGURE 22.</emphasis>
Adults of 
<taxonomicName id="4C294D5EFFC70726FE6D90A6FE74A5F1" box="[399,509,1199,1223]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEACFFFC70726FE6D90A6FE74A5F1" box="[399,509,1199,1223]" italics="true" pageId="41" pageNumber="42">Philodoria</emphasis>
</taxonomicName>
species. 
<emphasis id="B95DEACFFFC70726FDB990A6FDE4A5F1" bold="true" box="[603,621,1199,1223]" pageId="41" pageNumber="42">A</emphasis>
<taxonomicName id="4C294D5EFFC70726FD9790B9FD6DA5F1" box="[629,740,1199,1223]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="keaensis" status="sp. nov.">
<emphasis id="B95DEACFFFC70726FD9790B9FD6DA5F1" box="[629,740,1199,1223]" italics="true" pageId="41" pageNumber="42">P. keaensis</emphasis>
</taxonomicName>
<emphasis id="B95DEACFFFC70726FD0E90A6FCACA5F1" bold="true" box="[748,805,1199,1223]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A26E57B4FFC70726FD0E90A6FCACA5F1" box="[748,805,1199,1223]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
holotype female, lateral view, 
<emphasis id="B95DEACFFFC70726FB8190A6FBFDA5F1" bold="true" box="[1123,1140,1199,1223]" pageId="41" pageNumber="42">B</emphasis>
Forewings of holotype, 
<emphasis id="B95DEACFFFC70726FA9290A6FA0BA5F1" bold="true" box="[1392,1410,1199,1223]" pageId="41" pageNumber="42">C</emphasis>
<taxonomicName id="4C294D5EFFC70726FA6890B9FE3AA5DD" authority="(Swezey, 1946)" baseAuthorityName="Swezey" baseAuthorityYear="1946" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="nigrelloides">
<emphasis id="B95DEACFFFC70726FA6890B9FE86A5DD" italics="true" pageId="41" pageNumber="42">P. nigrelloides</emphasis>
(
<bibRefCitation id="EFB84B2CFFC70726FEFF90DAFE25A5DD" author="Swezey, O. H." box="[285,428,1235,1259]" pageId="41" pageNumber="42" pagination="625 - 628" refId="ref102759" refString="Swezey, O. H. (1946) New species of Hawaiian Lepidoptera. Proceedings of the Hawaiian Entomological Society, 12 (3), 625 - 628." type="journal article" year="1946">Swezey, 1946</bibRefCitation>
)
</taxonomicName>
holotype female, 
<emphasis id="B95DEACFFFC70726FD8890DAFDF5A5DD" bold="true" box="[618,636,1235,1259]" pageId="41" pageNumber="42">D</emphasis>
Forewings of 
<taxonomicName id="4C294D5EFFC70726FCED90DDFC33A5DD" box="[783,954,1235,1259]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="platyphylliella" status="sp. nov.">
<emphasis id="B95DEACFFFC70726FCED90DDFC33A5DD" box="[783,954,1235,1259]" italics="true" pageId="41" pageNumber="42">P. platyphylliella</emphasis>
</taxonomicName>
<emphasis id="B95DEACFFFC70726FC2390DAFC73A5DD" bold="true" box="[961,1018,1235,1259]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A26E57B4FFC70726FC2390DAFC73A5DD" box="[961,1018,1235,1259]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
holotype male. Scale bar 1 mm.
</paragraph>
</caption>
<subSubSection id="C3336556FFC70726FF259110FA14A402" box="[199,1437,1305,1332]" pageId="41" pageNumber="42" type="distribution">
<paragraph id="8B9636DDFFC70726FF259110FA14A402" blockId="41.[151,1437,1305,1728]" box="[199,1437,1305,1332]" pageId="41" pageNumber="42">
<emphasis id="B95DEACFFFC70726FF259110FEE8A405" bold="true" box="[199,353,1305,1331]" pageId="41" pageNumber="42">Distribution.</emphasis>
Kauai, Oahu, Molokai, Lanai (
<bibRefCitation id="EFB84B2CFFC70726FD589110FC00A402" author="Walsingham, Thomas" box="[698,905,1305,1332]" pageId="41" pageNumber="42" pagination="469 - 759" refId="ref102986" refString="Walsingham, Thomas, de Grey (Lord) (1907) Microlepidoptera. In: Sharp, D. (Ed.), Fauna Hawaiiensis. 1 (5). Cambridge University Press, Cambridge, pp. 469 - 759." type="book chapter" year="1907">Walsingham 1907</bibRefCitation>
), and 
<collectingRegion id="49EDF83FFFC70726FC299110FB95A405" box="[971,1052,1305,1331]" country="United States of America" name="Hawaii" pageId="41" pageNumber="42">Hawaii</collectingRegion>
(Big Island) (
<bibRefCitation id="EFB84B2CFFC70726FB579110FA19A402" author="Zimmerman, E. C." box="[1205,1424,1305,1332]" pageId="41" pageNumber="42" refId="ref103182" refString="Zimmerman, E. C. (1978 a) Gracillarioidea. In: Microlepidoptera. Part 1. Insects of Hawaii, Vol. 9. University of Hawaii Press, Honolulu, pp. 644 - 718." type="book" year="1978">Zimmerman 1978a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3336556FFC70726FF259137FDC7A762" pageId="41" pageNumber="42" type="biology_ecology">
<paragraph id="8B9636DDFFC70726FF259137FA07A461" blockId="41.[151,1437,1305,1728]" box="[199,1422,1341,1368]" pageId="41" pageNumber="42">
<emphasis id="B95DEACFFFC70726FF259137FEDFA46E" bold="true" box="[199,342,1342,1368]" pageId="41" pageNumber="42">Host plants.</emphasis>
<taxonomicName id="4C294D5EFFC70726FEBF9137FE51A46E" box="[349,472,1342,1368]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
: 
<taxonomicName id="4C294D5EFFC70726FE009134FBD7A46E" authority="Gaudich. (Walsingham, 1907)" authorityName="Gaudich. (Walsingham" authorityYear="1907" box="[482,1118,1341,1368]" class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="polymorpha">
<emphasis id="B95DEACFFFC70726FE009134FC8AA461" box="[482,771,1341,1367]" italics="true" pageId="41" pageNumber="42">Metrosideros polymorpha</emphasis>
Gaudich. (
<bibRefCitation id="EFB84B2CFFC70726FC9D9134FBDFA46E" author="Walsingham, Thomas" box="[895,1110,1341,1368]" pageId="41" pageNumber="42" pagination="469 - 759" refId="ref102986" refString="Walsingham, Thomas, de Grey (Lord) (1907) Microlepidoptera. In: Sharp, D. (Ed.), Fauna Hawaiiensis. 1 (5). Cambridge University Press, Cambridge, pp. 469 - 759." type="book chapter" year="1907">Walsingham, 1907</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C294D5EFFC70726FB889137FB5DA46E" box="[1130,1236,1342,1368]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Myrtales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="B95DEACFFFC70726FB889137FB5DA46E" box="[1130,1236,1342,1368]" italics="true" pageId="41" pageNumber="42">Syzygium</emphasis>
</taxonomicName>
sp.: new record.
</paragraph>
<paragraph id="8B9636DDFFC70726FF259168FB55A73D" blockId="41.[151,1437,1305,1728]" pageId="41" pageNumber="42">
<emphasis id="B95DEACFFFC70726FF259168FEAEA44D" bold="true" box="[199,295,1377,1403]" pageId="41" pageNumber="42">Biology.</emphasis>
(
<figureCitation id="13122A58FFC70726FEDB9168FE5BA44A" box="[313,466,1377,1404]" captionStart="FIGURE 66" captionStartId="118.[151,250,1829,1853]" captionTargetBox="[158,1430,528,1803]" captionTargetId="figure-197@118.[154,1433,524,1805]" captionTargetPageId="118" captionText="FIGURE 66. Habitats and host plants of Philodoria on Kauai. A–D: Upper Limahuli, E, F: Waimea Canyon. A Mountains of Limahuli, B Habitat with senior (last) author collecting specimens, C Same location as B, leaves with mines, D Metrosideros polymorpha leaves with mine on the ground with active larva, E Hibiscus waimeae subsp. hannerae flowers and leaf mines, F Leaf mine on same host plant as E. Arrows point to mines or larvae." figureDoi="http://doi.org/10.5281/zenodo.4681953" httpUri="https://zenodo.org/record/4681953/files/figure.png" pageId="41" pageNumber="42">Figs. 66C, D</figureCitation>
, 
<figureCitation id="13122A58FFC70726FE02916BFD89A44A" box="[480,512,1378,1404]" captionStart="FIGURE 82" captionStartId="144.[151,250,1255,1279]" captionTargetBox="[159,1425,187,1227]" captionTargetId="figure-23@144.[155,1432,181,1230]" captionTargetPageId="144" captionText="FIGURE 82. Philodoria splendida and its host plants. A, D, E: Metrosideros polymorpha, B, C, F: Syzigium sandwicensis. A: Maui, B–F: Oahu. A, B Leaf mine, C Cocoon and old mine, D, E Host plant with leaf mines, F Mine of late instar larva." figureDoi="http://doi.org/10.5281/zenodo.4681986" httpUri="https://zenodo.org/record/4681986/files/figure.png" pageId="41" pageNumber="42">82</figureCitation>
). 
<bibRefCitation id="EFB84B2CFFC70726FDF4916BFC85A44A" author="Swezey, O. H." box="[534,780,1377,1404]" pageId="41" pageNumber="42" pagination="221 - 227" refId="ref102406" refString="Swezey, O. H. (1913 b) Leaf-miners of the Hawaiian Islands. Proceedings of the Hawaiian Entomological Society, 2 (5), 221 - 227." type="journal article" year="1913">Swezey (1913b: 223)</bibRefCitation>
reported the “taco” like cocoon and mature larval habit, which are similar to that of 
<taxonomicName id="4C294D5EFFC70726FE24918FFDB9A4A9" box="[454,560,1413,1439]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="B95DEACFFFC70726FE24918FFDB9A4A9" box="[454,560,1413,1439]" italics="true" pageId="41" pageNumber="42">P. basalis</emphasis>
</taxonomicName>
. The biology is poorly studied, but it is presumably similar to that of 
<taxonomicName id="4C294D5EFFC70726FAC9918FFA1CA4A9" box="[1323,1429,1413,1439]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="B95DEACFFFC70726FAC9918FFA1CA4A9" box="[1323,1429,1413,1439]" italics="true" pageId="41" pageNumber="42">P. basalis</emphasis>
</taxonomicName>
. In general, larvae 
<taxonomicName id="4C294D5EFFC70726FE8291A0FE67A4F5" box="[352,494,1449,1475]" form="slender" pageId="41" pageNumber="42" rank="form">form slender</taxonomicName>
linear mine on the adaxial leaf surface (
<figureCitation id="13122A58FFC70726FC4F91A0FB9BA4F2" box="[941,1042,1449,1476]" captionStart="FIGURE 82" captionStartId="144.[151,250,1255,1279]" captionTargetBox="[159,1425,187,1227]" captionTargetId="figure-23@144.[155,1432,181,1230]" captionTargetPageId="144" captionText="FIGURE 82. Philodoria splendida and its host plants. A, D, E: Metrosideros polymorpha, B, C, F: Syzigium sandwicensis. A: Maui, B–F: Oahu. A, B Leaf mine, C Cocoon and old mine, D, E Host plant with leaf mines, F Mine of late instar larva." figureDoi="http://doi.org/10.5281/zenodo.4681986" httpUri="https://zenodo.org/record/4681986/files/figure.png" pageId="41" pageNumber="42">Fig. 82E</figureCitation>
), which gradually expands as lavae feed and grow (
<figureCitation id="13122A58FFC70726FEA191C4FE25A4DE" box="[323,428,1485,1512]" captionStart="FIGURE 82" captionStartId="144.[151,250,1255,1279]" captionTargetBox="[159,1425,187,1227]" captionTargetId="figure-23@144.[155,1432,181,1230]" captionTargetPageId="144" captionText="FIGURE 82. Philodoria splendida and its host plants. A, D, E: Metrosideros polymorpha, B, C, F: Syzigium sandwicensis. A: Maui, B–F: Oahu. A, B Leaf mine, C Cocoon and old mine, D, E Host plant with leaf mines, F Mine of late instar larva." figureDoi="http://doi.org/10.5281/zenodo.4681986" httpUri="https://zenodo.org/record/4681986/files/figure.png" pageId="41" pageNumber="42">Fig. 82A</figureCitation>
). One or two mines per leaf. We often observed the fallen mined leaves under the host plant (
<figureCitation id="13122A58FFC70726FF4291F8FEAFA73A" box="[160,294,1521,1548]" captionStart="FIGURE 66" captionStartId="118.[151,250,1829,1853]" captionTargetBox="[158,1430,528,1803]" captionTargetId="figure-197@118.[154,1433,524,1805]" captionTargetPageId="118" captionText="FIGURE 66. Habitats and host plants of Philodoria on Kauai. A–D: Upper Limahuli, E, F: Waimea Canyon. A Mountains of Limahuli, B Habitat with senior (last) author collecting specimens, C Same location as B, leaves with mines, D Metrosideros polymorpha leaves with mine on the ground with active larva, E Hibiscus waimeae subsp. hannerae flowers and leaf mines, F Leaf mine on same host plant as E. Arrows point to mines or larvae." figureDoi="http://doi.org/10.5281/zenodo.4681953" httpUri="https://zenodo.org/record/4681953/files/figure.png" pageId="41" pageNumber="42">Fig. 66C, D</figureCitation>
) with late instar larvae in the mine and/or pupae in a taco cocoon on or near the leaf.
</paragraph>
<paragraph id="8B9636DDFFC70726FF25921CFDC7A762" blockId="41.[151,1437,1305,1728]" pageId="41" pageNumber="42">
<emphasis id="B95DEACFFFC70726FF25921CFEDDA719" bold="true" box="[199,340,1557,1583]" pageId="41" pageNumber="42">Parasitoids.</emphasis>
(
<figureCitation id="13122A58FFC70726FE80921CFE71A706" box="[354,504,1557,1584]" captionStart="FIGURE 101" captionStartId="163.[151,250,1042,1066]" captionTargetBox="[157,1429,184,1016]" captionTargetId="figure-20@163.[151,1437,181,1018]" captionTargetPageId="163" captionText="FIGURE 101. Parasitoid wasps (Chalcidoidea) that emerged from Philodoria basalis or P. splendida. A–F: from P. basalis or P. splendida on Metrosideros polymorpha. A, B CJ123-1, Maui, C, D CJ123-2, E, F CJ417, Hawaii (Big Island), G, H CJ515 from P. splendida on Syzygium sp., Kauai." figureDoi="http://doi.org/10.5281/zenodo.4682033" httpUri="https://zenodo.org/record/4682033/files/figure.png" pageId="41" pageNumber="42">Fig. 101G, H</figureCitation>
). 
<taxonomicName id="4C294D5EFFC70726FDE8921CFD01A719" box="[522,648,1557,1583]" class="Insecta" family="Bethylidae" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="family">Bethylidae</taxonomicName>
: 
<taxonomicName id="4C294D5EFFC70726FD73921CFB95A706" authority="Fullaway, 1920" authorityName="Fullaway" authorityYear="1920" box="[657,1052,1557,1584]" class="Insecta" family="Bethylidae" genus="Sierola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="philodoriae">
<emphasis id="B95DEACFFFC70726FD73921CFCE0A719" box="[657,873,1557,1583]" italics="true" pageId="41" pageNumber="42">Sierola philodoriae</emphasis>
Fullaway, 1920
</taxonomicName>
; 
<taxonomicName id="4C294D5EFFC70726FBC5921CFB22A719" baseAuthorityName="Haliday" baseAuthorityYear="1833" box="[1063,1195,1557,1583]" class="Insecta" family="Eulophidae" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="family">Eulophidae</taxonomicName>
: 
<taxonomicName id="4C294D5EFFC70726FB56921CFDC0A762" authority="Yoshimoto, 1965" authorityName="Yoshimoto" authorityYear="1965" class="Insecta" family="Eulophidae" genus="Pauahiana" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="B95DEACFFFC70726FB56921CFA15A719" box="[1204,1436,1557,1583]" italics="true" pageId="41" pageNumber="42">Pauahiana metallica</emphasis>
<bibRefCitation id="EFB84B2CFFC70726FF759230FEDEA762" author="Yoshimoto, C. M." box="[151,343,1593,1620]" pageId="41" pageNumber="42" pagination="665 - 699" refId="ref103100" refString="Yoshimoto, C. M. (1965) Synopsis of Hawaiian Eulophidae including Aphelininae (Hym.: Chalcidoidea). Pacific Insects, 7 (4), 665 - 699." type="journal article" year="1965">Yoshimoto, 1965</bibRefCitation>
(
<bibRefCitation id="EFB84B2CFFC70726FE879230FDC8A762" author="Zimmerman, E. C." box="[357,577,1593,1620]" pageId="41" pageNumber="42" refId="ref103182" refString="Zimmerman, E. C. (1978 a) Gracillarioidea. In: Microlepidoptera. Part 1. Insects of Hawaii, Vol. 9. University of Hawaii Press, Honolulu, pp. 644 - 718." type="book" year="1978">Zimmerman 1978a</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3336556FFC70726FF259257FB8EA7F6" pageId="41" pageNumber="42" type="discussion">
<paragraph id="8B9636DDFFC70726FF259257FB8EA7F6" blockId="41.[151,1437,1305,1728]" pageId="41" pageNumber="42">
<emphasis id="B95DEACFFFC70726FF259257FEB2A74E" bold="true" box="[199,315,1630,1656]" pageId="41" pageNumber="42">Remarks.</emphasis>
We identified two adult moths (Coll ID CJ-049 / GenBank accession no. ID 
<accessionNumber id="947AAB3EFFC70726FB6D9257FA81A74E" box="[1167,1288,1630,1656]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KT982402" pageId="41" pageNumber="42">KT982402</accessionNumber>
and CJ-105 / 
<accessionNumber id="947AAB3EFFC70726FF75928BFE9AA7AA" box="[151,275,1666,1692]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KT982410" pageId="41" pageNumber="42">KT982410</accessionNumber>
) as 
<taxonomicName id="4C294D5EFFC70726FEDC928BFE40A7AD" box="[318,457,1665,1692]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="splendida">
<emphasis id="B95DEACFFFC70726FEDC928BFEC6A7AD" box="[318,335,1666,1691]" italics="true" pageId="41" pageNumber="42">P</emphasis>
. 
<emphasis id="B95DEACFFFC70726FEBE9288FE40A7AD" box="[348,457,1665,1691]" italics="true" pageId="41" pageNumber="42">splendida</emphasis>
</taxonomicName>
, based on the presence of a brown basal oblique fascia on forewing and host plant, from which whole bodies were sacrificed for molecular analysis (
<bibRefCitation id="EFB84B2CFFC70726FCD592ACFC72A7F6" author="Johns, C. A. &amp; Moore, M. R. &amp; Kawahara, A. Y." box="[823,1019,1701,1728]" pageId="41" pageNumber="42" pagination="361 - 372" refId="ref101228" refString="Johns, C. A., Moore, M. R. &amp; Kawahara, A. Y. (2016) Molecular phylogeny, revised higher classification, and implications for conservation of endangered Hawaiian leaf-mining moths (Lepidoptera: Gracillariidae: Philodoria). Pacific Science, 70 (3), 361 - 372. https: // doi. org / 10.2984 / 70.3.7" type="journal article" year="2016">
Johns 
<emphasis id="B95DEACFFFC70726FC9D92AFFC39A789" box="[895,944,1701,1727]" italics="true" pageId="41" pageNumber="42">et al</emphasis>
. 2016
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>