treatments-xml/data/03/80/87/038087A0FFF8FF8F5BCBB79FFB8AEE8A.xml

<document id="4C8617FD4C0DCE21D1892CF6FBB73D7D" ID-DOI="10.11646/zootaxa.4040.2.7" ID-GBIF-Dataset="8104ef08-9730-428d-a53a-5fc9e79018e0" ID-ISSN="1175-5326" ID-Zenodo-Dep="242146" ID-ZooBank="7F0C06CE-53AA-4D31-A180-191909020328" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461334719782" checkinUser="plazi" docAuthor="Sepahvand, Vahid, Momtazi, Farzaneh &amp; Tudge, Christopher" docDate="2015" docId="038087A0FFF8FF8F5BCBB79FFB8AEE8A" docLanguage="en" docName="zt04040p224.pdf" docOrigin="Zootaxa 4040 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cheramus iranicus Sepahvand, Momtazi &amp; Tudge, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="223" masterDocId="FFB9FFD8FFF9FF875B5CB100FF8FEC46" masterDocTitle="Cheramus iranicus, a new species of ghost shrimp (Decapoda: Axiidea: Callianassidae) from the Persian Gulf, Iran" masterLastPageNumber="224" masterPageNumber="215" pageNumber="216" updateTime="1698619054646" updateUser="plazi">
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<mods:title id="31242C1599657FDE68BDBA840BB9E5BE">Cheramus iranicus, a new species of ghost shrimp (Decapoda: Axiidea: Callianassidae) from the Persian Gulf, Iran</mods:title>
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<mods:namePart id="35D4C578583A0A86A8523906D2FBD993">Sepahvand, Vahid</mods:namePart>
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<mods:namePart id="9FF01781BB49728665A04C00B1B42CA7">Momtazi, Farzaneh</mods:namePart>
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<mods:namePart id="42E977FFED5566C2CFA42CFADD9C4AEE">Tudge, Christopher</mods:namePart>
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<mods:date id="FA44961BEF601CA50CA736A545A20FF5">2015</mods:date>
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<treatment id="038087A0FFF8FF8F5BCBB79FFB8AEE8A" ID-DOI="http://doi.org/10.5281/zenodo.6106635" ID-GBIF-Taxon="119644337" ID-Zenodo-Dep="6106635" LSID="urn:lsid:plazi:treatment:038087A0FFF8FF8F5BCBB79FFB8AEE8A" httpUri="http://treatment.plazi.org/id/038087A0FFF8FF8F5BCBB79FFB8AEE8A" lastPageId="8" lastPageNumber="223" pageId="1" pageNumber="216">
<subSubSection id="C333653DFFF8FF865BCBB79FFE85EA9D" pageId="1" pageNumber="216" type="nomenclature">
<paragraph id="8B9636B6FFF8FF865BCBB79FFE6BEAFC" blockId="1.[151,484,1695,1755]" box="[151,484,1695,1722]" pageId="1" pageNumber="216">
<heading id="D0DE81DAFFF8FF865BCBB79FFE6BEAFC" bold="true" box="[151,484,1695,1722]" fontSize="11" level="1" pageId="1" pageNumber="216" reason="1">
<emphasis id="B95DEAA4FFF8FF865BCBB79FFE6BEAFC" bold="true" box="[151,484,1695,1722]" pageId="1" pageNumber="216">
<taxonomicName id="4C294D35FFF8FF865BCBB79FFEF0EAFF" ID-CoL="63feac09-b778-4422-90e0-fe9e2ecd84f1" box="[151,383,1695,1721]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="iranicus" status="sp. nov.">
<emphasis id="B95DEAA4FFF8FF865BCBB79FFEF0EAFF" bold="true" box="[151,383,1695,1721]" italics="true" pageId="1" pageNumber="216">Cheramus iranicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57DFFFF8FF865ADAB7A0FE6BEAFC" box="[390,484,1696,1722]" pageId="1" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9636B6FFF8FF865BCBB7C2FE85EA9D" blockId="1.[151,484,1695,1755]" box="[151,266,1730,1755]" pageId="1" pageNumber="216">
(
<figureCitation id="13122A33FFF8FF865BC3B7C2FE8DEA9D" box="[159,258,1730,1755]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="2.[151,250,1973,1995]" captionStartId-1="3.[151,250,1960,1982]" captionStartId-2="4.[151,250,1958,1980]" captionStartId-3="5.[151,250,1477,1499]" captionTargetBox-0="[169,1418,193,1940]" captionTargetBox-1="[187,1399,194,1922]" captionTargetBox-2="[177,1407,196,1925]" captionTargetBox-3="[157,1427,193,1434]" captionTargetId-0="figure@2.[169,1419,193,1941]" captionTargetId-1="figure@3.[177,1403,193,1923]" captionTargetId-2="figure@4.[176,1408,193,1925]" captionTargetId-3="figure@5.[151,1436,193,1456]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 2. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: whole body, lateral view; B: eyestalk and rostrum; C: large cheliped (lateral); D: small cheliped (mesial); E: second pereopod (lateral). Scale bar: 1 mm." captionText-1="FIGURE 3. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm." captionText-2="FIGURE 4. Cheramus iranicus sp. nov. Allotype female, INIOC- 1 - 16 S. A: mandible; B: first maxillae; C: second maxillae; D: first maxilliped; E: second maxilliped; F and G: third maxilliped. Scale bar: 1 mm." captionText-3="FIGURE 5. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: third pereopod; B: fifth pereopod; C: first pleopod; D: third pleopod; E and F: pleopod 1 and 2 in allotype female INIOC- 1 - 16 S. Scale bar: 1 mm." httpUri-0="https://zenodo.org/record/242148/files/figure.png" httpUri-1="https://zenodo.org/record/242149/files/figure.png" httpUri-2="https://zenodo.org/record/242150/files/figure.png" httpUri-3="https://zenodo.org/record/242151/files/figure.png" pageId="1" pageNumber="216">Figs 2–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C333653DFFF8FF865BCBB60AFAFCEB90" pageId="1" pageNumber="216" type="materials_examined">
<paragraph id="8B9636B6FFF8FF865BCBB60AFB8BEB65" blockId="1.[151,1436,1802,2007]" box="[151,1028,1802,1827]" pageId="1" pageNumber="216">
<emphasis id="B95DEAA4FFF8FF865BCBB60AFEF6EB65" bold="true" box="[151,377,1802,1827]" pageId="1" pageNumber="216">Material examined</emphasis>
. 
<typeStatus id="54928814FFF8FF865ADBB60AFE7BEB65" box="[391,500,1802,1827]" pageId="1" pageNumber="216" type="holotype">Holotype</typeStatus>
: male (tl 
<quantity id="4CD19B53FFF8FF865907B60BFD4EEB64" box="[603,705,1803,1827]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="1" pageNumber="216" unit="mm" value="20.7">20.7 mm</quantity>
, cl 
<quantity id="4CD19B53FFF8FF8659B4B60AFCDFEB65" box="[744,848,1802,1827]" metricMagnitude="-3" metricUnit="m" metricValue="5.18" pageId="1" pageNumber="216" unit="mm" value="5.18">5.18 mm</quantity>
), INIOC-1-15S.
</paragraph>
<paragraph id="8B9636B6FFF8FF865B9BB62EFE73EB2D" blockId="1.[151,1436,1802,2007]" pageId="1" pageNumber="216">
Allotype: female (tl 
<quantity id="4CD19B53FFF8FF865A98B62EFDCFEB00" box="[452,576,1838,1863]" metricMagnitude="-2" metricUnit="m" metricValue="1.526" pageId="1" pageNumber="216" unit="mm" value="15.26">15.26 mm</quantity>
, cl 
<quantity id="4CD19B53FFF8FF86592BB62EFD67EB01" box="[631,744,1838,1863]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" pageId="1" pageNumber="216" unit="mm" value="3.65">3.65 mm</quantity>
), INIOC-1-16S, the Persian Gulf, west of 
<collectingCountry id="F33E7626FFF8FF865FA5B62EFAC7EB01" box="[1273,1352,1838,1863]" name="Iran" pageId="1" pageNumber="216">Qeshm</collectingCountry>
Island (
<geoCoordinate id="EE1D5071FFF8FF865BC3B652FEC6EB2D" box="[159,329,1874,1899]" direction="north" orientation="latitude" pageId="1" pageNumber="216" precision="1" value="26.586908">26°35'12.87&quot;N</geoCoordinate>
, 
<geoCoordinate id="EE1D5071FFF8FF865A09B652FE60EB2D" box="[341,495,1874,1899]" direction="east" orientation="longitude" pageId="1" pageNumber="216" precision="1" value="55.133522">55°08'0.67&quot;E</geoCoordinate>
).
</paragraph>
<paragraph id="8B9636B6FFF8FF865B9BB677FC72EBF5" blockId="1.[151,1436,1802,2007]" pageId="1" pageNumber="216">
<typeStatus id="54928814FFF8FF865B9BB677FEB6EBC9" box="[199,313,1911,1935]" pageId="1" pageNumber="216" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D2FFD3FFFF8FF865A1FB677FE00EBC9" box="[323,399,1910,1935]" pageId="1" pageNumber="216" type="male">1 male</specimenCount>
(tl 
<quantity id="4CD19B53FFF8FF865AE4B677FDA2EBC8" box="[440,557,1911,1935]" metricMagnitude="-2" metricUnit="m" metricValue="2.118" pageId="1" pageNumber="216" unit="mm" value="21.18">21.18 mm</quantity>
, cl 
<quantity id="4CD19B53FFF8FF86590AB676FD30EBC9" box="[598,703,1910,1935]" metricMagnitude="-3" metricUnit="m" metricValue="5.51" pageId="1" pageNumber="216" unit="mm" value="5.51">5.51 mm</quantity>
), 
<specimenCount id="9D2FFD3FFFF8FF86598EB677FCBBEBC9" box="[722,820,1910,1935]" pageId="1" pageNumber="216" type="female">1 female</specimenCount>
(tl 
<quantity id="4CD19B53FFF8FF865801B677FC5DEBC8" box="[861,978,1911,1935]" metricMagnitude="-2" metricUnit="m" metricValue="2.227" pageId="1" pageNumber="216" unit="mm" value="22.27">22.27 mm</quantity>
, cl 
<quantity id="4CD19B53FFF8FF8658A6B676FBECEBC9" box="[1018,1123,1910,1935]" metricMagnitude="-3" metricUnit="m" metricValue="5.58" pageId="1" pageNumber="216" unit="mm" value="5.58">5.58 mm</quantity>
), INIOC-1-17S, the Persian Gulf, west of 
<collectingCountry id="F33E7626FFF8FF865A6DB69AFE0FEBF5" box="[305,384,1946,1971]" name="Iran" pageId="1" pageNumber="216">Qeshm</collectingCountry>
Island (
<geoCoordinate id="EE1D5071FFF8FF865A87B69AFD0CEBF5" box="[475,643,1946,1971]" direction="north" orientation="latitude" pageId="1" pageNumber="216" precision="1" value="26.586908">26°35'12.87&quot;N</geoCoordinate>
<geoCoordinate id="EE1D5071FFF8FF8659D6B69AFCACEBF5" box="[650,803,1946,1971]" direction="east" orientation="longitude" pageId="1" pageNumber="216" precision="1" value="55.133522">55°08'0.67&quot;E</geoCoordinate>
), 
<date id="FF971076FFF8FF86586BB69AFC3DEBF5" box="[823,946,1946,1971]" pageId="1" pageNumber="216">5 February</date>
, 2014.
</paragraph>
<paragraph id="8B9636B6FFF8FF865B9BB6BEFAFCEB90" blockId="1.[151,1436,1802,2007]" box="[199,1395,1982,2007]" pageId="1" pageNumber="216">
<emphasis id="B95DEAA4FFF8FF865B9BB6BEFED2EB91" bold="true" box="[199,349,1982,2007]" pageId="1" pageNumber="216">
<typeStatus id="54928814FFF8FF865B9BB6BEFE8DEB91" box="[199,258,1982,2007]" pageId="1" pageNumber="216">Type</typeStatus>
locality
</emphasis>
. Persian Gulf, west of the 
<collectingCountry id="F33E7626FFF8FF8659D4B6BEFD58EB91" box="[648,727,1982,2007]" name="Iran" pageId="1" pageNumber="216">Qeshm</collectingCountry>
Island, 
<geoCoordinate id="EE1D5071FFF8FF86586DB6BEFC55EB91" box="[817,986,1982,2007]" direction="north" orientation="latitude" pageId="1" pageNumber="216" precision="1" value="26.586908">26°35'12.87&quot;N</geoCoordinate>
, 
<geoCoordinate id="EE1D5071FFF8FF8658BAB6BEFBF0EB91" box="[998,1151,1982,2007]" direction="east" orientation="longitude" pageId="1" pageNumber="216" precision="1" value="55.133522">55°08'0.67&quot;E</geoCoordinate>
; at depth of 
<quantity id="4CD19B53FFF8FF865E55B6BEFAE0EB90" box="[1289,1391,1982,2007]" metricMagnitude="1" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="216" unit="m" value="45.0" valueMax="50.0" valueMin="40.0">
40– 
<specimenCount id="9D2FFD3FFFF8FF865E68B6BEFAE0EB90" box="[1332,1391,1982,2007]" pageId="1" pageNumber="216" type="generic">50 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF56663EFFFBFF855BCBB6B5FB18EBAC" httpUri="https://zenodo.org/record/242148/files/figure.png" pageId="2" pageNumber="217" targetBox="[169,1418,193,1940]" targetPageId="2">
<paragraph id="8B9636B6FFFBFF855BCBB6B5FB18EBAC" blockId="2.[151,1436,1973,2026]" pageId="2" pageNumber="217">
<emphasis id="B95DEAA4FFFBFF855BCBB6B5FE81EB8D" bold="true" box="[151,270,1973,1995]" pageId="2" pageNumber="217">FIGURE 2</emphasis>
. 
<taxonomicName id="4C294D35FFFBFF855A48B6B5FE59EB8A" box="[276,470,1973,1996]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="iranicus" status="sp. nov.">
<emphasis id="B95DEAA4FFFBFF855A48B6B5FE59EB8A" box="[276,470,1973,1996]" italics="true" pageId="2" pageNumber="217">Cheramus iranicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57DFFFFBFF855A82B6B6FDA0EB8D" box="[478,559,1974,1995]" pageId="2" pageNumber="217" rank="species">
<emphasis id="B95DEAA4FFFBFF855A82B6B6FDA0EB8D" bold="true" box="[478,559,1974,1995]" pageId="2" pageNumber="217">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype male, INIOC-1-15S. A: whole body, lateral view; B: eyestalk and rostrum; C: large cheliped (lateral); D: small cheliped (mesial); E: second pereopod (lateral). Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF56663EFFFAFF845BCBB6A8FA9FEB9A" httpUri="https://zenodo.org/record/242149/files/figure.png" pageId="3" pageNumber="218" targetBox="[187,1399,194,1922]" targetPageId="3">
<paragraph id="8B9636B6FFFAFF845BCBB6A8FA9FEB9A" blockId="3.[151,1436,1960,2012]" pageId="3" pageNumber="218">
<emphasis id="B95DEAA4FFFAFF845BCBB6A8FE89EBF8" bold="true" box="[151,262,1960,1982]" pageId="3" pageNumber="218">FIGURE3</emphasis>
. 
<taxonomicName id="4C294D35FFFAFF845A4EB6A8FE5CEBF8" box="[274,467,1960,1982]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="218" phylum="Arthropoda" rank="species" species="iranicus" status="sp. nov.">
<emphasis id="B95DEAA4FFFAFF845A4EB6A8FE5CEBF8" box="[274,467,1960,1982]" italics="true" pageId="3" pageNumber="218">Cheramus iranicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57DFFFFAFF845A85B6A8FDA6EBFB" box="[473,553,1960,1981]" pageId="3" pageNumber="218" rank="species">
<emphasis id="B95DEAA4FFFAFF845A85B6A8FDA6EBFB" bold="true" box="[473,553,1960,1981]" pageId="3" pageNumber="218">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype male, INIOC-1-15S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF56663EFFFDFF835BCBB6A6FC52EB9C" httpUri="https://zenodo.org/record/242150/files/figure.png" pageId="4" pageNumber="219" targetBox="[177,1407,196,1925]" targetPageId="4">
<paragraph id="8B9636B6FFFDFF835BCBB6A6FC52EB9C" blockId="4.[151,1436,1958,2010]" pageId="4" pageNumber="219">
<emphasis id="B95DEAA4FFFDFF835BCBB6A6FE83EBFD" bold="true" box="[151,268,1958,1980]" pageId="4" pageNumber="219">FIGURE 4</emphasis>
. 
<taxonomicName id="4C294D35FFFDFF835A44B6A6FE57EBFA" box="[280,472,1958,1980]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="iranicus" status="sp. nov.">
<emphasis id="B95DEAA4FFFDFF835A44B6A6FE57EBFA" box="[280,472,1958,1980]" italics="true" pageId="4" pageNumber="219">Cheramus iranicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57DFFFFDFF835A82B6A6FDA1EBFD" box="[478,558,1958,1979]" pageId="4" pageNumber="219" rank="species">
<emphasis id="B95DEAA4FFFDFF835A82B6A6FDA1EBFD" bold="true" box="[478,558,1958,1979]" pageId="4" pageNumber="219">sp. nov.</emphasis>
</taxonomicNameLabel>
Allotype female, INIOC-1-16S. A: mandible; B: first maxillae; C: second maxillae; D: first maxilliped; E: second maxilliped; F and G: third maxilliped. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF56663EFFFCFF825BCBB4C5FBDDE9BC" httpUri="https://zenodo.org/record/242151/files/figure.png" pageId="5" pageNumber="220" targetBox="[157,1427,193,1434]" targetPageId="5">
<paragraph id="8B9636B6FFFCFF825BCBB4C5FBDDE9BC" blockId="5.[151,1436,1477,1530]" pageId="5" pageNumber="220">
<emphasis id="B95DEAA4FFFCFF825BCBB4C5FE82E99D" bold="true" box="[151,269,1477,1499]" pageId="5" pageNumber="220">FIGURE 5</emphasis>
. 
<taxonomicName id="4C294D35FFFCFF825A46B4C5FE54E99D" box="[282,475,1477,1499]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="iranicus" status="sp. nov.">
<emphasis id="B95DEAA4FFFCFF825A46B4C5FE54E99D" box="[282,475,1477,1499]" italics="true" pageId="5" pageNumber="220">Cheramus iranicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57DFFFFCFF825ABEB4C6FDBCE99D" box="[482,563,1478,1499]" pageId="5" pageNumber="220" rank="species">
<emphasis id="B95DEAA4FFFCFF825ABEB4C6FDBCE99D" bold="true" box="[482,563,1478,1499]" pageId="5" pageNumber="220">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype male, INIOC-1-15S. A: third pereopod; B: fifth pereopod; C: first pleopod; D: third pleopod; E and F: pleopod 1 and 2 in allotype female INIOC-1-16S. Scale bar: 1 mm.
</paragraph>
</caption>
<subSubSection id="C333653DFFFCFF825B9BB721FE7DEB54" pageId="5" pageNumber="220" type="diagnosis">
<paragraph id="8B9636B6FFFCFF825B9BB721FE7DEB54" blockId="5.[151,1437,1569,2026]" pageId="5" pageNumber="220">
<emphasis id="B95DEAA4FFFCFF825B9BB721FEB5EA7C" bold="true" box="[199,314,1569,1594]" pageId="5" pageNumber="220">Diagnosis</emphasis>
. Rostrum reaching to one third length of eyestalks; eyestalks with prominent slender distal spine; thoracic shield (plate on thoracic sternite 7) (
<figureCitation id="13122A33FFFCFF8259FCB745FD6BEA18" box="[672,740,1605,1630]" captionStart="FIGURE 3" captionStartId="3.[151,250,1960,1982]" captionTargetBox="[187,1399,194,1922]" captionTargetId="figure@3.[177,1403,193,1923]" captionTargetPageId="3" captionText="FIGURE 3. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242149/files/figure.png" pageId="5" pageNumber="220">Fig. 3</figureCitation>
D) pentagonal with strongly produced anteromedian margin; ventral surface with shallow groove posteriorly; third maxilliped with broad (suboperculiform) merus and ischium; larger cheliped with 7 denticles on ventral margin of ischium, merus oval with proximal ventral spine; telson as broad as long, broadest proximally, proximolateral margins convex, posterior margin strongly concave medially with median tooth or fixed spine and with 2 movable spines at each angle; uropodal exopod not particularly broad with truncate posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="C333653DFFFCFF805B9BB61DFC9CE976" lastPageId="7" lastPageNumber="222" pageId="5" pageNumber="220" type="description">
<paragraph id="8B9636B6FFFCFF825B9BB61DFBE2EBAC" blockId="5.[151,1437,1569,2026]" pageId="5" pageNumber="220">
<emphasis id="B95DEAA4FFFCFF825B9BB61DFED6EB70" bold="true" box="[199,345,1821,1846]" pageId="5" pageNumber="220">Description.</emphasis>
Carapace with frontal margin (
<figureCitation id="13122A33FFFCFF8259E7B61DFD72EB70" box="[699,765,1821,1846]" captionStart="FIGURE 3" captionStartId="3.[151,250,1960,1982]" captionTargetBox="[187,1399,194,1922]" captionTargetId="figure@3.[177,1403,193,1923]" captionTargetPageId="3" captionText="FIGURE 3. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242149/files/figure.png" pageId="5" pageNumber="220">Fig. 3</figureCitation>
A) bearing short, spiniform rostrum not reaching to cornea, reaching approximately one third length of eyestalks and obtuse rounded projection lateral to rostrum; carapace length about one quarter of total length; orbital margins evenly concave; linea thalassinica distinct, extending to posterolateral margin of carapace; dorsal oval clearly demarcated with deep cervical groove and shallow anterior and transverse grooves behind rostral base (
<figureCitation id="13122A33FFFCFF8259D9B6ADFD5CEB80" box="[645,723,1965,1990]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[169,1418,193,1940]" captionTargetId="figure@2.[169,1419,193,1941]" captionTargetPageId="2" captionText="FIGURE 2. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: whole body, lateral view; B: eyestalk and rostrum; C: large cheliped (lateral); D: small cheliped (mesial); E: second pereopod (lateral). Scale bar: 1 mm." httpUri="https://zenodo.org/record/242148/files/figure.png" pageId="5" pageNumber="220">Figs. 2</figureCitation>
A, 3A, D). Anterior branchiostegal lobe (pterygostomial angle) triangular, extending to anterior margin of first segment of antennal peduncle (
<figureCitation id="13122A33FFFCFF825F5AB6D1FBC7EBAC" box="[1030,1096,2001,2026]" captionStart="FIGURE 3" captionStartId="3.[151,250,1960,1982]" captionTargetBox="[187,1399,194,1922]" captionTargetId="figure@3.[177,1403,193,1923]" captionTargetPageId="3" captionText="FIGURE 3. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242149/files/figure.png" pageId="5" pageNumber="220">Fig. 3</figureCitation>
D).
</paragraph>
<paragraph id="8B9636B6FFFFFF815B9BB197FF77EDCE" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="221">
Length ratio of pleomeres to telson measured along midline 1.0: 1.6: 1.2: 1.0: 1.2: 1.6 (anterior to posterior) (
<figureCitation id="13122A33FFFFFF815BC3B1BCFF69EC92" box="[159,230,188,213]" captionStart="FIGURE 3" captionStartId="3.[151,250,1960,1982]" captionTargetBox="[187,1399,194,1922]" captionTargetId="figure@3.[177,1403,193,1923]" captionTargetPageId="3" captionText="FIGURE 3. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242149/files/figure.png" pageId="6" pageNumber="221">Fig. 3</figureCitation>
D). First pleomere (
<figureCitation id="13122A33FFFFFF815A8CB1BCFD99EC92" box="[464,534,188,213]" captionStart="FIGURE 3" captionStartId="3.[151,250,1960,1982]" captionTargetBox="[187,1399,194,1922]" captionTargetId="figure@3.[177,1403,193,1923]" captionTargetPageId="3" captionText="FIGURE 3. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242149/files/figure.png" pageId="6" pageNumber="221">Fig. 3</figureCitation>
D) narrowing anteriorly in dorsal view; pleuron with defined ventral margin. Second pleomere (
<figureCitation id="13122A33FFFFFF815A33B1DFFE4EECBE" box="[367,449,223,248]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[169,1418,193,1940]" captionTargetId="figure@2.[169,1419,193,1941]" captionTargetPageId="2" captionText="FIGURE 2. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: whole body, lateral view; B: eyestalk and rostrum; C: large cheliped (lateral); D: small cheliped (mesial); E: second pereopod (lateral). Scale bar: 1 mm." httpUri="https://zenodo.org/record/242148/files/figure.png" pageId="6" pageNumber="221">Figs. 2</figureCitation>
A, 3D) approximately as long as wide; pleuron having posterolateral margin slightly expanded, with few setae on ventro-distal margin. Third to fifth pleura each with posterior tuft of long setae and few dorsal setae; posterolateral margins slightly expanded. Sixth pleomere (
<figureCitation id="13122A33FFFFFF8158B0B027FBB6ED06" box="[1004,1081,295,320]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[169,1418,193,1940]" captionTargetId="figure@2.[169,1419,193,1941]" captionTargetPageId="2" captionText="FIGURE 2. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: whole body, lateral view; B: eyestalk and rostrum; C: large cheliped (lateral); D: small cheliped (mesial); E: second pereopod (lateral). Scale bar: 1 mm." httpUri="https://zenodo.org/record/242148/files/figure.png" pageId="6" pageNumber="221">Figs. 2</figureCitation>
A, 3D, E) 1.2 times longer than wide, subquadrate, with parallel lateral margins and a row of setae (
<figureCitation id="13122A33FFFFFF8158CCB04CFC5CED22" box="[912,979,332,357]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[169,1418,193,1940]" captionTargetId="figure@2.[169,1419,193,1941]" captionTargetPageId="2" captionText="FIGURE 2. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: whole body, lateral view; B: eyestalk and rostrum; C: large cheliped (lateral); D: small cheliped (mesial); E: second pereopod (lateral). Scale bar: 1 mm." httpUri="https://zenodo.org/record/242148/files/figure.png" pageId="6" pageNumber="221">Fig. 2</figureCitation>
A); without conspicuous notch on lateral margins.
</paragraph>
<paragraph id="8B9636B6FFFFFF815B9BB094FCB7EE7B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="221">
Telson (
<figureCitation id="13122A33FFFFFF815A7FB094FEE8EDEA" box="[291,359,404,429]" captionStart="FIGURE 3" captionStartId="3.[151,250,1960,1982]" captionTargetBox="[187,1399,194,1922]" captionTargetId="figure@3.[177,1403,193,1923]" captionTargetPageId="3" captionText="FIGURE 3. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242149/files/figure.png" pageId="6" pageNumber="221">Fig. 3</figureCitation>
D, E) subtrapezoidal, slightly narrowing posteriorly, width approximately equal to length (1.2 times wider than long), lateral margins each with low convexity proximally and 2 pairs of movable spines near posterolateral angle; posterior margin deeply concave and bilobed, with small medial tooth and some spines and long spiniform setae. Dorsal surface slightly convex, armed with 1 pair of movable spines, proximomedian tuft of long setae and 2 tufts of long setae on each side of mid-line.
</paragraph>
<paragraph id="8B9636B6FFFFFF815B9BB347FE8AEF1A" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="221">
Uropodal protopod bearing bilobed posterodorsal margin. Endopod elongate, sub-semicircular, with distinctly convex inner margin and nearly straight outer margin (
<figureCitation id="13122A33FFFFFF81584CB36CFCDBEEC2" box="[784,852,620,645]" captionStart="FIGURE 3" captionStartId="3.[151,250,1960,1982]" captionTargetBox="[187,1399,194,1922]" captionTargetId="figure@3.[177,1403,193,1923]" captionTargetPageId="3" captionText="FIGURE 3. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242149/files/figure.png" pageId="6" pageNumber="221">Fig. 3</figureCitation>
D, E), about twice as long as broad, and distinctly longer than telson, lateral margin unarmed; mesial margin fringed with setae, distal margin with cluster of 4 movable spines (spiniform setae) mixed with flexible setae; dorsal surface armed with 2 movable spines on either side of obtuse median ridge. Exopod larger than endopod, rounded oblong, 2.5 times as long as broad; dorsal surface slightly elevated along midline; dorsal plate with distal row of 4 spiniform setae distinctly separated from setal row of posterior margin; posterior margin roundly truncate with row of stiff setae of various lengths and stoutness.
</paragraph>
<paragraph id="8B9636B6FFFFFF815B9BB267FB82EF8E" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="221">
Eyestalks (
<figureCitation id="13122A33FFFFFF815A1FB267FE09EFC6" box="[323,390,871,896]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[169,1418,193,1940]" captionTargetId="figure@2.[169,1419,193,1941]" captionTargetPageId="2" captionText="FIGURE 2. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: whole body, lateral view; B: eyestalk and rostrum; C: large cheliped (lateral); D: small cheliped (mesial); E: second pereopod (lateral). Scale bar: 1 mm." httpUri="https://zenodo.org/record/242148/files/figure.png" pageId="6" pageNumber="221">Fig. 2</figureCitation>
B, 3A, 3D) mildly elongate, subtriangular, contiguous, armed each with prominent spine on distal margin, tip overreaching distal end of basal antennal article; cornea located subterminally and laterally, large and occupying more than half eyestalk width in dorsal view, darkly pigmented.
</paragraph>
<paragraph id="8B9636B6FFFFFF815B9BB2D4FCD1E811" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="221">
Antennule peduncle (
<figureCitation id="13122A33FFFFFF815AE2B2D4FD8DEFAA" box="[446,514,980,1005]" captionStart="FIGURE 3" captionStartId="3.[151,250,1960,1982]" captionTargetBox="[187,1399,194,1922]" captionTargetId="figure@3.[177,1403,193,1923]" captionTargetPageId="3" captionText="FIGURE 3. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242149/files/figure.png" pageId="6" pageNumber="221">Fig. 3</figureCitation>
B) slightly shorter than antennal peduncle, ultimate segment 3 times as long as penultimate segment, with long setae, penultimate segment shorter than basal segment, with long setae ventrally; flagella unequal and dissimilar, upper flagellum composed of 14 articles and thicker and shorter than lower flagellum, lower flagellum with 19 articles, bearing short setae.
</paragraph>
<paragraph id="8B9636B6FFFFFF815B9BB564FB36E8E6" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="221">
Antennal peduncle (
<figureCitation id="13122A33FFFFFF815AF1B564FE7FE83A" box="[429,496,1124,1149]" captionStart="FIGURE 3" captionStartId="3.[151,250,1960,1982]" captionTargetBox="[187,1399,194,1922]" captionTargetId="figure@3.[177,1403,193,1923]" captionTargetPageId="3" captionText="FIGURE 3. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242149/files/figure.png" pageId="6" pageNumber="221">Fig. 3</figureCitation>
C) with third segment as long as second segment, fourth segment longest, about 2 times as long as fifth segment. Scaphocerite tiny, subovate, terminating in sharp, curved spine.
</paragraph>
<paragraph id="8B9636B6FFFFFF815B9BB5ACFC13E882" blockId="6.[151,1437,151,2013]" box="[199,924,1196,1221]" pageId="6" pageNumber="221">
Small process on either side of epistome bearing setae (
<figureCitation id="13122A33FFFFFF815869B5ACFCF8E882" box="[821,887,1196,1221]" captionStart="FIGURE 3" captionStartId="3.[151,250,1960,1982]" captionTargetBox="[187,1399,194,1922]" captionTargetId="figure@3.[177,1403,193,1923]" captionTargetPageId="3" captionText="FIGURE 3. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: lateral view of carapace and cephalic appendages; B: antennule; C: antennal peduncle; D: dorsal view of body; E: sixth abdominal somite and telson. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242149/files/figure.png" pageId="6" pageNumber="221">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="8B9636B6FFFFFF815B9BB5CFFE9FEAA6" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="221">
Mandible (
<figureCitation id="13122A33FFFFFF815A1AB5CFFE02E8AE" box="[326,397,1231,1256]" captionStart="FIGURE 4" captionStartId="4.[151,250,1958,1980]" captionTargetBox="[177,1407,196,1925]" captionTargetId="figure@4.[176,1408,193,1925]" captionTargetPageId="4" captionText="FIGURE 4. Cheramus iranicus sp. nov. Allotype female, INIOC- 1 - 16 S. A: mandible; B: first maxillae; C: second maxillae; D: first maxilliped; E: second maxilliped; F and G: third maxilliped. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242150/files/figure.png" pageId="6" pageNumber="221">Fig. 4</figureCitation>
A) with smooth molar process. Incisor process with 7 small unequal teeth, with slightly concave inner surface. Mandibular palp 3-segmented, curved mesially, third segment larger than two proximal segments, bearing many stiff setae; basal article 0.8 times as long as second article. First maxilla (
<figureCitation id="13122A33FFFFFF815E5CB417FAC9E976" box="[1280,1350,1303,1328]" captionStart="FIGURE 4" captionStartId="4.[151,250,1958,1980]" captionTargetBox="[177,1407,196,1925]" captionTargetId="figure@4.[176,1408,193,1925]" captionTargetPageId="4" captionText="FIGURE 4. Cheramus iranicus sp. nov. Allotype female, INIOC- 1 - 16 S. A: mandible; B: first maxillae; C: second maxillae; D: first maxilliped; E: second maxilliped; F and G: third maxilliped. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242150/files/figure.png" pageId="6" pageNumber="221">Fig. 4</figureCitation>
B) with basal endite wide, bearing dense setation; distal endite proximally rounded; endopod long and slender, terminal part deflected. Second maxilla (
<figureCitation id="13122A33FFFFFF815AA3B45FFDCBE93E" box="[511,580,1375,1400]" captionStart="FIGURE 4" captionStartId="4.[151,250,1958,1980]" captionTargetBox="[177,1407,196,1925]" captionTargetId="figure@4.[176,1408,193,1925]" captionTargetPageId="4" captionText="FIGURE 4. Cheramus iranicus sp. nov. Allotype female, INIOC- 1 - 16 S. A: mandible; B: first maxillae; C: second maxillae; D: first maxilliped; E: second maxilliped; F and G: third maxilliped. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242150/files/figure.png" pageId="6" pageNumber="221">Fig. 4</figureCitation>
C) with exopod of broad distal lobe, 2 times as long as wide, endopod long and slender, tip digitiform; proximal and distal endites each longitudinally subdivided and with numerous setae on mesial margin. First maxilliped (
<figureCitation id="13122A33FFFFFF81595BB4A7FDC5E986" box="[519,586,1447,1472]" captionStart="FIGURE 4" captionStartId="4.[151,250,1958,1980]" captionTargetBox="[177,1407,196,1925]" captionTargetId="figure@4.[176,1408,193,1925]" captionTargetPageId="4" captionText="FIGURE 4. Cheramus iranicus sp. nov. Allotype female, INIOC- 1 - 16 S. A: mandible; B: first maxillae; C: second maxillae; D: first maxilliped; E: second maxilliped; F and G: third maxilliped. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242150/files/figure.png" pageId="6" pageNumber="221">Fig. 4</figureCitation>
D) with exopod crest-shaped with setose external margins; endopod greatly reduced to bud-like process; distal endite elongate with curved lateral margins, minutely denticulate, covered with long setae. Second maxilliped with small epipod (
<figureCitation id="13122A33FFFFFF815989B4EFFC94EA4E" box="[725,795,1519,1544]" captionStart="FIGURE 4" captionStartId="4.[151,250,1958,1980]" captionTargetBox="[177,1407,196,1925]" captionTargetId="figure@4.[176,1408,193,1925]" captionTargetPageId="4" captionText="FIGURE 4. Cheramus iranicus sp. nov. Allotype female, INIOC- 1 - 16 S. A: mandible; B: first maxillae; C: second maxillae; D: first maxilliped; E: second maxilliped; F and G: third maxilliped. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242150/files/figure.png" pageId="6" pageNumber="221">Fig. 4</figureCitation>
E). Dactylus short, about half as long as propodus, with dense setation on distal and dorsal margins. Propodus slightly arcuate, length 2 times width. Carpus short and triangular. Merus length exceeding 4 times width. Exopod unsegmented, short, reaching nearly to midlength of merus. Third maxilliped (
<figureCitation id="13122A33FFFFFF815AEAB75CFE76EA32" box="[438,505,1628,1653]" captionStart="FIGURE 4" captionStartId="4.[151,250,1958,1980]" captionTargetBox="[177,1407,196,1925]" captionTargetId="figure@4.[176,1408,193,1925]" captionTargetPageId="4" captionText="FIGURE 4. Cheramus iranicus sp. nov. Allotype female, INIOC- 1 - 16 S. A: mandible; B: first maxillae; C: second maxillae; D: first maxilliped; E: second maxilliped; F and G: third maxilliped. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242150/files/figure.png" pageId="6" pageNumber="221">Fig. 4</figureCitation>
F, G) without exopod. Ischium rectangular,1.5 times as long as wide; crista dentata on mesial face composed of 15 sharp denticles; merus wider than long, combined length of ischium-merus about 3.2 times width. Carpus subovate, 2 times as long as broad. Dactylus narrow and digitiform, with stiff setae terminally.
</paragraph>
<paragraph id="8B9636B6FFFFFF805B9BB7ECFB1EED5B" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="222" pageId="6" pageNumber="221">
First pereopods (chelipeds) unequal and dissimilar. Larger cheliped 
<emphasis id="B95DEAA4FFFFFF81589BB7ECFC5FEB43" bold="true" box="[967,976,1772,1797]" pageId="6" pageNumber="221">(</emphasis>
<figureCitation id="13122A33FFFFFF81588CB7ECFB9DEB42" box="[976,1042,1772,1797]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[169,1418,193,1940]" captionTargetId="figure@2.[169,1419,193,1941]" captionTargetPageId="2" captionText="FIGURE 2. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: whole body, lateral view; B: eyestalk and rostrum; C: large cheliped (lateral); D: small cheliped (mesial); E: second pereopod (lateral). Scale bar: 1 mm." httpUri="https://zenodo.org/record/242148/files/figure.png" pageId="6" pageNumber="221">Fig. 2</figureCitation>
C) with ischium slender (about 3.0 times as long as broad), proximo-dorsal margin straight, disto-dorsal margin conspicuously convex (sinuous dorsal margin), with series of 7 acute spines on ventral margin slightly increasing in size distally. Merus about 1.5 times as long as wide; dorsal margin strongly convex, smoothly rounded; ventral margin also convex, with a small spine proximally, remainder of ventral margin with row of minute rounded denticles of similar size, and with row of setae. Carpus short, stout, 0.9 times as long as wide, with dense setation on ventral margin; disto-ventral angle rounded; mesial surface nearly flat; carinae present on dorsal and ventral margins. Propodus heavy, about 1.6 times as long as broad (including fixed finger); carinae present on dorsal and ventral margins. Palm sub-rectangular in lateral view; dactylus hooked distally, about 0.8 times as long as palm, with dense setae on dorsal margin and along cutting edge; fixed finger curved distally, cutting edge with hook-like tooth distally, remainder with row of small teeth, dorsal and ventral margins bearing row of long setae; narrow hiatus between fingers.
</paragraph>
<paragraph id="8B9636B6FFFEFF805B9BB027FD03EE5E" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="222">
Smaller cheliped (
<figureCitation id="13122A33FFFEFF805AC4B027FE52ED06" box="[408,477,295,320]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[169,1418,193,1940]" captionTargetId="figure@2.[169,1419,193,1941]" captionTargetPageId="2" captionText="FIGURE 2. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: whole body, lateral view; B: eyestalk and rostrum; C: large cheliped (lateral); D: small cheliped (mesial); E: second pereopod (lateral). Scale bar: 1 mm." httpUri="https://zenodo.org/record/242148/files/figure.png" pageId="7" pageNumber="222">Fig. 2</figureCitation>
D) shorter and much more slender than larger cheliped. Ischium about 4.0 times as long as wide; dorsal margin nearly straight, unarmed; ventral margin with row of 8 subequal spines. Merus about 2 times as long as broad; dorsal margin slightly convex with minute spine proximally; ventral margin with small proximal spine preceded by row of rounded denticles or granules. Carpus about 2.5 times as long as broad and 1.3 times as long as merus, widened distally; carinae present on dorsal and ventral margins. Propodus straight ventrally with numerous setae; carinae present on dorsal and ventral margins; dactylus almost as long as palm, unarmed on cutting edge; distinct hiatus between fingers.
</paragraph>
<paragraph id="8B9636B6FFFEFF805B9BB324FE6BEE8B" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="222">
Second pereopod (
<figureCitation id="13122A33FFFEFF805AC6B324FE52EE7A" box="[410,477,548,573]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[169,1418,193,1940]" captionTargetId="figure@2.[169,1419,193,1941]" captionTargetPageId="2" captionText="FIGURE 2. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: whole body, lateral view; B: eyestalk and rostrum; C: large cheliped (lateral); D: small cheliped (mesial); E: second pereopod (lateral). Scale bar: 1 mm." httpUri="https://zenodo.org/record/242148/files/figure.png" pageId="7" pageNumber="222">Fig. 2</figureCitation>
E) chelate. Ischium short, ventral margin gently concave. Merus 2.5 times as long as ischium, about 3.0 times as long as greatest width; ventral margin sinuous, with row of long setae. Carpus subtriangular, 1.8 times as long as wide, concave distally with dense setae proximoventrally. Propodus triangular with dense setae on ventral margin, fixed finger margin smooth. Dactylus with dense setae on dorsal margin; cutting edge straight, smooth.
</paragraph>
<paragraph id="8B9636B6FFFEFF805B9BB3D7FAE1EF1B" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="222">
Third pereopod (
<figureCitation id="13122A33FFFEFF805ADFB3D7FE4AEEB6" box="[387,453,727,752]" captionStart="FIGURE 5" captionStartId="5.[151,250,1477,1499]" captionTargetBox="[157,1427,193,1434]" captionTargetId="figure@5.[151,1436,193,1456]" captionTargetPageId="5" captionText="FIGURE 5. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: third pereopod; B: fifth pereopod; C: first pleopod; D: third pleopod; E and F: pleopod 1 and 2 in allotype female INIOC- 1 - 16 S. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242151/files/figure.png" pageId="7" pageNumber="222">Fig. 5</figureCitation>
A) strongly compressed laterally, non-chelate. Ischium 2 times as long as wide, ischium and merus with same width. Merus 1.7 times as long as ischium, 3.4 times as long as wide. Carpus, subtriangular, distal margin with two low lobes, and tufts of long setae at each angle. Propodus wider than carpus, subovate, with row of numerous setae, becoming longer proximally, on ventral margin. Dactylus much shorter than propodus.
</paragraph>
<paragraph id="8B9636B6FFFEFF805B9BB267FE69EFC6" blockId="7.[151,1437,151,2013]" box="[199,486,871,896]" pageId="7" pageNumber="222">Fourth pereopod missing.</paragraph>
<paragraph id="8B9636B6FFFEFF805B9BB28CFA94EFAA" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="222">
Fifth pereopod (
<figureCitation id="13122A33FFFEFF805ADEB28CFE48EFE3" box="[386,455,908,933]" captionStart="FIGURE 5" captionStartId="5.[151,250,1477,1499]" captionTargetBox="[157,1427,193,1434]" captionTargetId="figure@5.[151,1436,193,1456]" captionTargetPageId="5" captionText="FIGURE 5. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: third pereopod; B: fifth pereopod; C: first pleopod; D: third pleopod; E and F: pleopod 1 and 2 in allotype female INIOC- 1 - 16 S. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242151/files/figure.png" pageId="7" pageNumber="222">Fig. 5</figureCitation>
B) slender, chelate, longer than second to fourth pereopods. Coxa 2 times as long as wide. Ischium about as long as coxa. Merus about 2 times as long as ischium. Carpus slightly shorter than propodus. Propodus with dense setae on distal part. Dactylus curved, partially obscured by dense setae.
</paragraph>
<paragraph id="8B9636B6FFFEFF805B9BB2F7FDF5E873" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="222">
Male first pleopod 
<emphasis id="B95DEAA4FFFEFF805AC0B2F7FE2AE856" bold="true" box="[412,421,1015,1040]" pageId="7" pageNumber="222">(</emphasis>
<figureCitation id="13122A33FFFEFF805AF9B2F7FE6AE856" box="[421,485,1015,1040]" captionStart="FIGURE 5" captionStartId="5.[151,250,1477,1499]" captionTargetBox="[157,1427,193,1434]" captionTargetId="figure@5.[151,1436,193,1456]" captionTargetPageId="5" captionText="FIGURE 5. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: third pereopod; B: fifth pereopod; C: first pleopod; D: third pleopod; E and F: pleopod 1 and 2 in allotype female INIOC- 1 - 16 S. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242151/files/figure.png" pageId="7" pageNumber="222">Fig. 5</figureCitation>
C) uniramus, two segmented, proximal segment thick at base, distal segment (ramus) tapered to blunt tip, with few setae distally.
</paragraph>
<paragraph id="8B9636B6FFFEFF805B9BB53FFD82E81E" blockId="7.[151,1437,151,2013]" box="[199,525,1087,1112]" pageId="7" pageNumber="222">Male second pleopod absent.</paragraph>
<paragraph id="8B9636B6FFFEFF805B9BB564FEE1E8E6" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="222">
Appendix interna of third through fifth pleopods slender, finger–like; each exopod longer and broader than endopod (
<figureCitation id="13122A33FFFEFF805A5AB587FEC6E8E6" box="[262,329,1159,1184]" captionStart="FIGURE 5" captionStartId="5.[151,250,1477,1499]" captionTargetBox="[157,1427,193,1434]" captionTargetId="figure@5.[151,1436,193,1456]" captionTargetPageId="5" captionText="FIGURE 5. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: third pereopod; B: fifth pereopod; C: first pleopod; D: third pleopod; E and F: pleopod 1 and 2 in allotype female INIOC- 1 - 16 S. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242151/files/figure.png" pageId="7" pageNumber="222">Fig. 5</figureCitation>
D).
</paragraph>
<paragraph id="8B9636B6FFFEFF805B9BB5ACFBA8E94A" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="222">
Female first pleopod (
<figureCitation id="13122A33FFFEFF805A9AB5ACFD84E883" box="[454,523,1196,1221]" captionStart="FIGURE 5" captionStartId="5.[151,250,1477,1499]" captionTargetBox="[157,1427,193,1434]" captionTargetId="figure@5.[151,1436,193,1456]" captionTargetPageId="5" captionText="FIGURE 5. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: third pereopod; B: fifth pereopod; C: first pleopod; D: third pleopod; E and F: pleopod 1 and 2 in allotype female INIOC- 1 - 16 S. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242151/files/figure.png" pageId="7" pageNumber="222">Fig. 5</figureCitation>
E), uniramous, composed of three segments, proximal segment about 2.3 times longer than distal, with 5 long plumose setae. Female second pleopod (
<figureCitation id="13122A33FFFEFF80589DB5CFFB8BE8AE" box="[961,1028,1231,1256]" captionStart="FIGURE 5" captionStartId="5.[151,250,1477,1499]" captionTargetBox="[157,1427,193,1434]" captionTargetId="figure@5.[151,1436,193,1456]" captionTargetPageId="5" captionText="FIGURE 5. Cheramus iranicus sp. nov. Holotype male, INIOC- 1 - 15 S. A: third pereopod; B: fifth pereopod; C: first pleopod; D: third pleopod; E and F: pleopod 1 and 2 in allotype female INIOC- 1 - 16 S. Scale bar: 1 mm." httpUri="https://zenodo.org/record/242151/files/figure.png" pageId="7" pageNumber="222">Fig. 5</figureCitation>
F) biramous; endopod straight, with two terminal plumose setae, exopod longer than endopod, with 14 plumose setae.
</paragraph>
<paragraph id="8B9636B6FFFEFF805B9BB417FC9CE976" blockId="7.[151,1437,151,2013]" box="[199,787,1303,1328]" pageId="7" pageNumber="222">
<emphasis id="B95DEAA4FFFEFF805B9BB417FEACE976" bold="true" box="[199,291,1303,1328]" pageId="7" pageNumber="222">Habitat</emphasis>
. sandy-muddy bottom with shell fragments.
</paragraph>
</subSubSection>
<subSubSection id="C333653DFFFEFF8F5B9BB43CFE34EEEE" lastPageId="8" lastPageNumber="223" pageId="7" pageNumber="222" type="discussion">
<paragraph id="8B9636B6FFFEFF805B9BB43CFB37EADE" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="222">
<emphasis id="B95DEAA4FFFEFF805B9BB43CFEB4E913" bold="true" box="[199,315,1340,1365]" pageId="7" pageNumber="222">Remarks.</emphasis>
The new species is unique in the possession of a sharp distal spine on the eyestalk and the telson with two pairs of lateral movable spines and a deep median concavity and a small but distinct median spine on the posterior margin. Among other 
<taxonomicName id="4C294D35FFFEFF805979B484FD17E9DB" box="[549,664,1412,1437]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA4FFFEFF805979B484FD17E9DB" box="[549,664,1412,1437]" italics="true" pageId="7" pageNumber="222">Cheramus</emphasis>
</taxonomicName>
-like species, it is also characterized by a spiniform rostrum, suboperculiform merus-ischium of the third maxilliped and slender, digitiform appendices internae on the third to fifth pleopods. Among the known 
<taxonomicName id="4C294D35FFFEFF80594AB4CCFD06E9A3" box="[534,649,1484,1509]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA4FFFEFF80594AB4CCFD06E9A3" box="[534,649,1484,1509]" italics="true" pageId="7" pageNumber="222">Cheramus</emphasis>
</taxonomicName>
-like species, 
<taxonomicName id="4C294D35FFFEFF80587EB4CCFB16E9A3" authority="Sakai, 2005" authorityName="Sakai" authorityYear="2005" box="[802,1177,1484,1509]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="222" phylum="Arthropoda" rank="species" species="thorsoni">
<emphasis id="B95DEAA4FFFEFF80587EB4CCFB84E9A3" box="[802,1035,1484,1509]" italics="true" pageId="7" pageNumber="222">Callianassa thorsoni</emphasis>
<bibRefCitation id="EFB84B47FFFEFF805F4FB4CCFB16E9A3" author="Sakai" box="[1043,1177,1484,1509]" pageId="7" pageNumber="222" refString="Sakai, K. (2005) Callianassoidea of the world (Decapoda, Thalassinidea). Crustaceana Monographs, 4, 1 - 285." type="journal article" year="2005">Sakai, 2005</bibRefCitation>
</taxonomicName>
, transferred to 
<taxonomicName id="4C294D35FFFEFF805E1CB4CDFA13E9A2" box="[1344,1436,1485,1508]" class="Malacostraca" family="Callianassidae" genus="Trypaea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA4FFFEFF805E1CB4CDFA13E9A2" box="[1344,1436,1485,1508]" italics="true" pageId="7" pageNumber="222">Trypaea</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFB84B47FFFEFF805BE6B4EFFEC4EA4E" author="Sakai" box="[186,331,1519,1544]" pageId="7" pageNumber="222" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
, appears closest to the present new species in sharing the sharp distal spine on the eyestalk and the general configuration of the telson. However, 
<taxonomicName id="4C294D35FFFEFF8059C2B715FC93EA6B" box="[670,796,1556,1581]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="222" phylum="Arthropoda" rank="species" species="thorsoni">
<emphasis id="B95DEAA4FFFEFF8059C2B715FC93EA6B" box="[670,796,1556,1581]" italics="true" pageId="7" pageNumber="222">C. thorsoni</emphasis>
</taxonomicName>
is readily distinguished from the present new species by the only faintly concave posterior margin of the telson. In 
<taxonomicName id="4C294D35FFFEFF805841B738FC14EA16" box="[797,923,1591,1616]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="222" phylum="Arthropoda" rank="species" species="thorsoni">
<emphasis id="B95DEAA4FFFEFF805841B738FC14EA16" box="[797,923,1591,1616]" italics="true" pageId="7" pageNumber="222">C. thorsoni</emphasis>
</taxonomicName>
, only the minor cheliped of the male 
<typeStatus id="54928814FFFEFF805E60B739FA13EA17" box="[1340,1436,1593,1617]" pageId="7" pageNumber="222" type="paratype">paratype</typeStatus>
was preserved, in which there are only three tiny teeth on the ventral margin of the ischium. In the present new species, the ischium ventral margin is serrated by seven or eight small teeth on both chelipeds.
</paragraph>
<paragraph id="8B9636B6FFFEFF805B9BB7A4FC70EBFF" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="222">
As previously mentioned, the composition of the genus 
<taxonomicName id="4C294D35FFFEFF8058D5B7A4FC73EAFB" box="[905,1020,1700,1725]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA4FFFEFF8058D5B7A4FC73EAFB" box="[905,1020,1700,1725]" italics="true" pageId="7" pageNumber="222">Cheramus</emphasis>
</taxonomicName>
has not been consistent between taxonomists. In the present study, we accept the concept discussed by 
<bibRefCitation id="EFB84B47FFFEFF8058F4B7C7FB0AEAA6" box="[936,1157,1735,1760]" pageId="7" pageNumber="222" refString="Komai, T., Maenosono, T. &amp; Fujita, Y. (2014) Two new species of ghost shrimp assigned to the genus Cheramus Bate, 1888 (Crustacea: Decapoda: Axiidea: Callianassidae) from the Ryukyu Islands, Japan. Zootaxa, 3895 (4), 503 - 524. http: // dx. doi. org / 10.11646 / zootaxa. 3895.4.3" type="journal article">
Komai 
<emphasis id="B95DEAA4FFFEFF8058A7B7C9FBA3EAA6" box="[1019,1068,1735,1760]" italics="true" pageId="7" pageNumber="222">et al</emphasis>
. (2014)
</bibRefCitation>
and provisionally assign our new species to 
<taxonomicName id="4C294D35FFFEFF805A2EB7ECFE6AEB43" box="[370,485,1772,1797]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA4FFFEFF805A2EB7ECFE6AEB43" box="[370,485,1772,1797]" italics="true" pageId="7" pageNumber="222">Cheramus</emphasis>
</taxonomicName>
based on the following characters: 1) the long, slender body with a relatively small cephalothorax, 2) carapace with rostral spine, 3) corneas subterminal and lateral, 4) chelipeds unequal, major without distinct meral hook, 5) antennular peduncle not much longer or stouter than antennal peduncle, 6) distal margin of merus of third maxilliped not strongly produced, 7) plp1 slender and uniramous in both sexes; plp2 absent in male, slender and biramous in female; rami of plp3–5 relatively slender, with slender, finger-like appendix interna, 8) uropodal exopod elongate, length more than twice width.
</paragraph>
<paragraph id="8B9636B6FFFEFF8F5B9BB6C4FE34EEEE" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1436,151,717]" lastPageId="8" lastPageNumber="223" pageId="7" pageNumber="222">
<bibRefCitation id="EFB84B47FFFEFF805B9BB6C4FE7DEB9A" author="Manning" box="[199,498,1988,2013]" pageId="7" pageNumber="222" refString="Manning, R. B. &amp; Felder, D. L. (1991) Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington, 104, 764 - 792." type="journal article" year="1991">Manning &amp; Felder (1991)</bibRefCitation>
assigned the following species to 
<taxonomicName id="4C294D35FFFEFF8058D9B6C4FC77EB9B" box="[901,1016,1988,2013]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA4FFFEFF8058D9B6C4FC77EB9B" box="[901,1016,1988,2013]" italics="true" pageId="7" pageNumber="222">Cheramus</emphasis>
</taxonomicName>
: 
<taxonomicName id="4C294D35FFFEFF805F57B6C4FA1AEB9A" authority="Biffar, 1973" authorityName="Biffar" authorityYear="1973" box="[1035,1429,1988,2013]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="222" phylum="Arthropoda" rank="species" species="profunda">
<emphasis id="B95DEAA4FFFEFF805F57B6C4FA8EEB9B" box="[1035,1281,1988,2013]" italics="true" pageId="7" pageNumber="222">Callianassa profunda</emphasis>
<bibRefCitation id="EFB84B47FFFEFF805E57B6C4FA1AEB9A" author="Biffar" box="[1291,1429,1988,2013]" pageId="7" pageNumber="222" refString="Biffar, T. A. (1973) The taxonomic status of Callianassa occidentalis Bate, 1888, and C. batei Borradaile, 1903 (Decapoda, Callianassidae). Crustaceana, 24, 224 - 230. http: // dx. doi. org / 10.1163 / 156854073 X 00399" type="journal article" year="1973">Biffar, 1973</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C294D35FFF1FF8F5BCBB197FDC0ECF6" authority="Rathbun, 1901" authorityName="Rathbun" authorityYear="1901" box="[151,591,151,176]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="B95DEAA4FFF1FF8F5BCBB197FE14ECF6" box="[151,411,151,176]" italics="true" pageId="8" pageNumber="223">Callianassa marginata</emphasis>
<bibRefCitation id="EFB84B47FFF1FF8F5AF9B197FDC0ECF6" author="Rathbun" box="[421,591,151,176]" pageId="8" pageNumber="223" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Porto Rico. Bulletin of the United States Fish Commission, 20 (2), 1 - 127. Rodrigues, S. de A. &amp; Manning, R. B. (1992) Poti gaucho, a new genus and species of ghost shrimp from southern Brazil (Crustacea: Decapoda: Callianassidae). Bulletin of Marine Science, 51 (1), 9 - 13." type="journal article" year="1901">Rathbun, 1901</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C294D35FFF1FF8F5902B197FC5EECF6" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" box="[606,977,151,176]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B95DEAA4FFF1FF8F5902B197FCC8ECF6" box="[606,839,151,176]" italics="true" pageId="8" pageNumber="223">Cheramus orientalis</emphasis>
<bibRefCitation id="EFB84B47FFF1FF8F580DB198FC5EECF6" author="Bate" box="[849,977,152,176]" pageId="8" pageNumber="223" refString="Bate, C. S. (1888) Report on the Crustacea Macrura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. &quot; Challenger &quot; during the years 1837 - 76. Zoology, 24, 1 - 942." type="journal article" year="1888">Bate, 1888</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C294D35FFF1FF8F5F4EB197FE94EC92" authority="Le Loeuff &amp; Intes, 1974" authorityName="Le Loeuff &amp; Intes" authorityYear="1974" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="oblonga">
<emphasis id="B95DEAA4FFF1FF8F5F4EB197FB71ECF6" box="[1042,1278,151,176]" italics="true" pageId="8" pageNumber="223">Callianassa oblonga</emphasis>
<bibRefCitation id="EFB84B47FFF1FF8F5E54B198FE94EC92" author="Le" pageId="8" pageNumber="223" refString="Le Loeuff, P. &amp; Intes, A. (1974) Les Thalassinidea (Crustacea, Decapoda) du Golfe de Guinee. Systematique Ecologie. Cahiers O. R. S. T. O. M., Serie Oceanographique, 12 (l), 17 - 69." type="journal article" year="1974">Le Loeuff &amp; Intès, 1974</bibRefCitation>
</taxonomicName>
. 
<bibRefCitation id="EFB84B47FFF1FF8F5A77B1BCFD9EEC92" box="[299,529,188,213]" pageId="8" pageNumber="223" refString="Tudge, C. C., Poore, G. C. B. &amp; Lemaitre, R. (2000) Preliminary phylogenetic analysis of generic relationships within the Callianassidae and Ctenochelidae (Decapoda: Thalassinidea: Callianassoidea). Journal of Crustacean Biology, 20, 129 - 149." type="journal article">
Tudge 
<emphasis id="B95DEAA4FFF1FF8F5A21B1BDFE3CEC93" box="[381,435,188,213]" italics="true" pageId="8" pageNumber="223">et al</emphasis>
. (2000)
</bibRefCitation>
added 
<taxonomicName id="4C294D35FFF1FF8F5930B1BCFB9CEC93" authority="Sakai, 1967" authorityName="Sakai" authorityYear="1967" box="[620,1043,188,213]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="longicauda">
<emphasis id="B95DEAA4FFF1FF8F5930B1BCFCF4EC93" box="[620,891,188,213]" italics="true" pageId="8" pageNumber="223">Callianassa longicauda</emphasis>
<bibRefCitation id="EFB84B47FFF1FF8F58DBB1BCFB9CEC93" author="Sakai" box="[903,1043,188,213]" pageId="8" pageNumber="223" refString="Sakai, K. (1967) Three new species of Thalassinidea (Decapod Crustacea) from south west Japan. Publications of the Seto Marine Biological Laboratory, 15 (4), 319 - 328." type="journal article" year="1967">Sakai, 1967</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C294D35FFF1FF8F5F7FB1BDFB30EC93" box="[1059,1215,188,213]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="praedatrix">
<emphasis id="B95DEAA4FFF1FF8F5F7FB1BDFB30EC93" box="[1059,1215,188,213]" italics="true" pageId="8" pageNumber="223">C. praedatrix</emphasis>
</taxonomicName>
de 
<bibRefCitation id="EFB84B47FFF1FF8F5FADB1BDFAFDEC93" author="Man" box="[1265,1394,188,213]" pageId="8" pageNumber="223" refString="Man, J. G. de (1905) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeiniging, 9 (2), 587 - 614." type="journal article" year="1905">Man, 1905</bibRefCitation>
, 
<taxonomicName id="4C294D35FFF1FF8F5EDEB1BDFE84ECBE" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="propinqua">
<emphasis id="B95DEAA4FFF1FF8F5EDEB1BDFE84ECBE" italics="true" pageId="8" pageNumber="223">C. propinqua</emphasis>
</taxonomicName>
de 
<bibRefCitation id="EFB84B47FFF1FF8F5A6FB1E0FE20ECBE" author="Man" box="[307,431,223,248]" pageId="8" pageNumber="223" refString="Man, J. G. de (1905) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeiniging, 9 (2), 587 - 614." type="journal article" year="1905">Man, 1905</bibRefCitation>
, 
<taxonomicName id="4C294D35FFF1FF8F5AE6B1E0FDFFECBE" box="[442,624,223,248]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="rectangularis">
<emphasis id="B95DEAA4FFF1FF8F5AE6B1E0FDFFECBE" box="[442,624,223,248]" italics="true" pageId="8" pageNumber="223">C. rectangularis</emphasis>
</taxonomicName>
<bibRefCitation id="EFB84B47FFF1FF8F592BB1E0FCA8ECBE" author="Ngoc-Ho" box="[631,807,224,248]" pageId="8" pageNumber="223" refString="Ngoc-Ho, N. (1991) Sur quelques Callianassidae et Upogebiidae de Nouvelle Caledonie (Crustacea, Thalassinidea). In: Richer de Forges, B. (ed.), Le benthos des fonds meubles des lagons de Nouvelle Caledonie, 1, 281 - 311." type="journal article" year="1991">Ngoc-Ho, 1991</bibRefCitation>
, 
<taxonomicName id="4C294D35FFF1FF8F586FB1E0FC25ECBE" box="[819,938,223,248]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="B95DEAA4FFF1FF8F586FB1E0FC25ECBE" box="[819,938,223,248]" italics="true" pageId="8" pageNumber="223">C. sibogae</emphasis>
</taxonomicName>
de 
<bibRefCitation id="EFB84B47FFF1FF8F588FB1E0FBC1ECBE" author="Man" box="[979,1102,223,248]" pageId="8" pageNumber="223" refString="Man, J. G. de (1905) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeiniging, 9 (2), 587 - 614." type="journal article" year="1905">Man, 1905</bibRefCitation>
, and 
<taxonomicName id="4C294D35FFF1FF8F5FD5B1E0FF40ED5A" authority="Sakai, 1970" authorityName="Sakai" authorityYear="1970" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="spinophthalma">
<emphasis id="B95DEAA4FFF1FF8F5FD5B1E0FAC0ECBE" box="[1161,1359,223,248]" italics="true" pageId="8" pageNumber="223">C. spinophthalma</emphasis>
<bibRefCitation id="EFB84B47FFF1FF8F5E0BB1DFFF40ED5A" author="Sakai" pageId="8" pageNumber="223" refString="Sakai, K. (1970) A small collection of thalassinids from the waters around Tsushima islands, Japan, including a new species of Callianassa (Crustacea, Anomura). Publications of the Seto Marine Biological Laboratory, 18 (1), 37 - 47." type="journal article" year="1970">Sakai, 1970</bibRefCitation>
</taxonomicName>
to 
<taxonomicName id="4C294D35FFF1FF8F5BA0B004FEE0ED5B" box="[252,367,260,285]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA4FFF1FF8F5BA0B004FEE0ED5B" box="[252,367,260,285]" italics="true" pageId="8" pageNumber="223">Cheramus</emphasis>
</taxonomicName>
. 
<bibRefCitation id="EFB84B47FFF1FF8F5ADEB004FD9AED5A" author="Sakai" box="[386,533,260,285]" pageId="8" pageNumber="223" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
later transferred 
<taxonomicName id="4C294D35FFF1FF8F59B8B004FC65ED5B" box="[740,1002,260,285]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="anoploura">
<emphasis id="B95DEAA4FFF1FF8F59B8B004FC65ED5B" box="[740,1002,260,285]" italics="true" pageId="8" pageNumber="223">Callianassa anoploura</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C294D35FFF1FF8F58A1B005FB2FED5B" box="[1021,1184,260,285]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="longicauda">
<emphasis id="B95DEAA4FFF1FF8F58A1B005FB2FED5B" box="[1021,1184,260,285]" italics="true" pageId="8" pageNumber="223">C. longicauda</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C294D35FFF1FF8F5FEFB005FAFFED5B" box="[1203,1392,260,285]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="malaccaensis">
<emphasis id="B95DEAA4FFF1FF8F5FEFB005FAFFED5B" box="[1203,1392,260,285]" italics="true" pageId="8" pageNumber="223">C. malaccaensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C294D35FFF1FF8F5EDEB005FE96ED06" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="nigroculata">
<emphasis id="B95DEAA4FFF1FF8F5EDEB005FE96ED06" italics="true" pageId="8" pageNumber="223">C. nigroculata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C294D35FFF1FF8F5A7BB028FDB9ED06" authority="Sakai, 2002" authorityName="Sakai" authorityYear="2002" box="[295,566,295,320]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="contipes">
<emphasis id="B95DEAA4FFF1FF8F5A7BB028FE29ED06" box="[295,422,296,320]" italics="true" pageId="8" pageNumber="223">C. contipes</emphasis>
<bibRefCitation id="EFB84B47FFF1FF8F5AF1B027FDB9ED06" author="Sakai" box="[429,566,295,320]" pageId="8" pageNumber="223" refString="Sakai, K. (2002) Callianassidae (Decapoda, Thalassinidea) in the Andaman Sea, Thailand. In: Bruce, N. L., Berggren, M. &amp; Bussawarit, S. (Eds.), Proceedings of the International Workshop on the Biodiversity of Crustacea of the Andaman Sea. Phuket Marine Biological Center Special Publication, 23, 461 - 532." type="journal article" year="2002">Sakai, 2002</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C294D35FFF1FF8F591DB028FC3EED06" authority="Dworschak, 2003" authorityName="Dworschak" authorityYear="2003" box="[577,945,295,320]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="aqabaensis">
<emphasis id="B95DEAA4FFF1FF8F591DB028FD6EED06" box="[577,737,295,320]" italics="true" pageId="8" pageNumber="223">C. aqabaensis</emphasis>
<bibRefCitation id="EFB84B47FFF1FF8F59B4B027FC3EED06" author="Dworschak" box="[744,945,295,320]" pageId="8" pageNumber="223" refString="Dworschak, P. C. (2003) A new species of ghost shrimp from the Gulf of Aqaba, Red Sea (Crustacea: Decapoda: Callianassidae). Annalen des Naturhistorisches Museums in Wien, 104 (B), 415 - 428." type="journal article" year="2003">Dworschak, 2003</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C294D35FFF1FF8F58E1B028FBE4ED06" box="[957,1131,295,320]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="lobetobensis">
<emphasis id="B95DEAA4FFF1FF8F58E1B028FBE4ED06" box="[957,1131,295,320]" italics="true" pageId="8" pageNumber="223">C. lobetobensis</emphasis>
</taxonomicName>
de 
<bibRefCitation id="EFB84B47FFF1FF8F5FC9B028FA9EED06" author="Man" box="[1173,1297,295,320]" pageId="8" pageNumber="223" refString="Man, J. G. de (1905) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeiniging, 9 (2), 587 - 614." type="journal article" year="1905">Man, 1905</bibRefCitation>
, 
<taxonomicName id="4C294D35FFF1FF8F5E41B028FA13ED06" box="[1309,1436,295,320]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B95DEAA4FFF1FF8F5E41B028FA13ED06" box="[1309,1436,295,320]" italics="true" pageId="8" pageNumber="223">C. modesta</emphasis>
</taxonomicName>
de 
<bibRefCitation id="EFB84B47FFF1FF8F5BE4B04DFEBAED23" author="Man" box="[184,309,332,357]" pageId="8" pageNumber="223" refString="Man, J. G. de (1905) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeiniging, 9 (2), 587 - 614." type="journal article" year="1905">Man, 1905</bibRefCitation>
, C. 
<emphasis id="B95DEAA4FFF1FF8F5A3DB04CFE37ED23" box="[353,440,332,357]" italics="true" pageId="8" pageNumber="223">parvula</emphasis>
<bibRefCitation id="EFB84B47FFF1FF8F5AE3B04CFDC8ED22" author="Sakai" box="[447,583,332,357]" pageId="8" pageNumber="223" refString="Sakai, K. (1988) A new genus and five new species of Callianassidae (Crustacea: Decapoda: Thalassinidea) from northern Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences, 5, 51 - 69." type="journal article" year="1988">Sakai, 1988</bibRefCitation>
, and 
<taxonomicName id="4C294D35FFF1FF8F59DDB04DFBDCED22" authority="Rodrigues &amp; Manning, 1992" authorityName="Rodrigues &amp; Manning" authorityYear="1992" box="[641,1107,332,357]" class="Malacostraca" family="Callianassidae" genus="Poti" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="gaucho">
<emphasis id="B95DEAA4FFF1FF8F59DDB04DFC86ED23" box="[641,777,332,357]" italics="true" pageId="8" pageNumber="223">Poti gaucho</emphasis>
Rodrigues &amp; Manning, 1992
</taxonomicName>
to 
<taxonomicName id="4C294D35FFF1FF8F5F2BB04CFB65ED23" box="[1143,1258,332,357]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA4FFF1FF8F5F2BB04CFB65ED23" box="[1143,1258,332,357]" italics="true" pageId="8" pageNumber="223">Cheramus</emphasis>
</taxonomicName>
. In more recent studies, 
<taxonomicName id="4C294D35FFF1FF8F5BA8B06FFC8BEDCE" authority="Komai &amp; Fujiwara, 2012" authorityName="Komai &amp; Fujiwara" authorityYear="2012" box="[244,772,367,392]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="cavifrons">
<emphasis id="B95DEAA4FFF1FF8F5BA8B06FFE57EDCE" box="[244,472,367,392]" italics="true" pageId="8" pageNumber="223">Cheramus cavifrons</emphasis>
<bibRefCitation id="EFB84B47FFF1FF8F5ABDB06FFC8BEDCE" author="Komai" box="[481,772,367,392]" pageId="8" pageNumber="223" refString="Komai, T. &amp; Fujiwara, Y. (2012) New records of callianassid ghost shrimp (Crustacea: Decapoda: Axiidea) from reducing environments in Kyushu, southwestern Japan. Zootaxa, 3271, 55 - 67." type="journal article" year="2012">Komai &amp; Fujiwara, 2012</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C294D35FFF1FF8F584DB070FAC7EDCE" authority="Komai, Maenosono &amp; Fujita, 2014" authorityName="Komai, Maenosono &amp; Fujita" authorityYear="2014" box="[785,1352,367,392]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="spinicauda">
<emphasis id="B95DEAA4FFF1FF8F584DB070FC20EDCE" box="[785,943,367,392]" italics="true" pageId="8" pageNumber="223">C. spinicauda</emphasis>
<bibRefCitation id="EFB84B47FFF1FF8F58EBB06FFAC7EDCE" author="Komai" box="[951,1352,367,392]" pageId="8" pageNumber="223" refString="Komai, T., Maenosono, T. &amp; Fujita, Y. (2014) Two new species of ghost shrimp assigned to the genus Cheramus Bate, 1888 (Crustacea: Decapoda: Axiidea: Callianassidae) from the Ryukyu Islands, Japan. Zootaxa, 3895 (4), 503 - 524. http: // dx. doi. org / 10.11646 / zootaxa. 3895.4.3" type="journal article" year="2014">Komai, Maenosono &amp; Fujita, 2014</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C294D35FFF1FF8F5EDEB070FD18EDEA" authority="Komai, Maenosono &amp; Fujita, 2014" authorityName="Komai, Maenosono &amp; Fujita" authorityYear="2014" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="ohuranus">
<emphasis id="B95DEAA4FFF1FF8F5EDEB070FE8EEDEB" italics="true" pageId="8" pageNumber="223">C. ohuranus</emphasis>
<bibRefCitation id="EFB84B47FFF1FF8F5A55B094FD18EDEA" author="Komai" box="[265,663,404,429]" pageId="8" pageNumber="223" refString="Komai, T., Maenosono, T. &amp; Fujita, Y. (2014) Two new species of ghost shrimp assigned to the genus Cheramus Bate, 1888 (Crustacea: Decapoda: Axiidea: Callianassidae) from the Ryukyu Islands, Japan. Zootaxa, 3895 (4), 503 - 524. http: // dx. doi. org / 10.11646 / zootaxa. 3895.4.3" type="journal article" year="2014">Komai, Maenosono &amp; Fujita, 2014</bibRefCitation>
</taxonomicName>
were all described from Japanese waters (
<bibRefCitation id="EFB84B47FFF1FF8F5F27B094FA18EDEA" author="Komai" box="[1147,1431,404,429]" pageId="8" pageNumber="223" refString="Komai, T. &amp; Fujiwara, Y. (2012) New records of callianassid ghost shrimp (Crustacea: Decapoda: Axiidea) from reducing environments in Kyushu, southwestern Japan. Zootaxa, 3271, 55 - 67." type="journal article" year="2012">Komai &amp; Fujiwara 2012</bibRefCitation>
; 
<bibRefCitation id="EFB84B47FFF1FF8F5BCBB0B7FEE0ED96" author="Komai" box="[151,367,439,464]" pageId="8" pageNumber="223" refString="Komai, T., Maenosono, T. &amp; Fujita, Y. (2014) Two new species of ghost shrimp assigned to the genus Cheramus Bate, 1888 (Crustacea: Decapoda: Axiidea: Callianassidae) from the Ryukyu Islands, Japan. Zootaxa, 3895 (4), 503 - 524. http: // dx. doi. org / 10.11646 / zootaxa. 3895.4.3" type="journal article" year="2014">
Komai 
<emphasis id="B95DEAA4FFF1FF8F5BB1B0B9FEAEED96" box="[237,289,439,464]" italics="true" pageId="8" pageNumber="223">et al</emphasis>
. 2014
</bibRefCitation>
). However, according to the on-line World Register of Marine Species (
<bibRefCitation id="EFB84B47FFF1FF8F5FEFB0B8FAB3ED96" author="Poore" box="[1203,1340,439,464]" pageId="8" pageNumber="223" refString="Poore, G. C. B. (2015) World Register of Marine Species (WORMS) Available from: http: // www. marinespecies. org / aphia. php? p = taxdetails &amp; id = 415191 (Accessed on 2 September 2015)" type="journal article" year="2015">Poore 2015</bibRefCitation>
), only 9 species referred to 
<taxonomicName id="4C294D35FFF1FF8F5A33B0DCFE6DEDB3" box="[367,482,476,501]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA4FFF1FF8F5A33B0DCFE6DEDB3" box="[367,482,476,501]" italics="true" pageId="8" pageNumber="223">Cheramus</emphasis>
</taxonomicName>
are listed. 
<taxonomicName id="4C294D35FFF1FF8F593DB0DCFCB6EDB2" box="[609,825,476,501]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="iranicus" status="sp. nov.">
<emphasis id="B95DEAA4FFF1FF8F593DB0DCFCB6EDB2" box="[609,825,476,501]" italics="true" pageId="8" pageNumber="223">Cheramus iranicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57DFFFF1FF8F581DB0DCFC13EDB3" box="[833,924,476,501]" pageId="8" pageNumber="223" rank="species">
<emphasis id="B95DEAA4FFF1FF8F581DB0DCFC13EDB3" bold="true" box="[833,924,476,501]" pageId="8" pageNumber="223">sp. nov.</emphasis>
</taxonomicNameLabel>
is easily distinguished from these species by the deeply bilobed telson with a prominent medial tooth in combination with the two pairs of lateral movable spines. This combination of telson armature is found only in 
<taxonomicName id="4C294D35FFF1FF8F5806B324FC42EE7B" box="[858,973,548,573]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAA4FFF1FF8F5806B324FC42EE7B" box="[858,973,548,573]" italics="true" pageId="8" pageNumber="223">Cheramus</emphasis>
</taxonomicName>
in the species, 
<taxonomicName id="4C294D35FFF1FF8F5FDFB325FEF8EE26" authority="(De Man, 1905) (Sakai 2011)" authorityName="(De Man, 1905) (Sakai" authorityYear="2011" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="propinquus">
<emphasis id="B95DEAA4FFF1FF8F5FDFB325FAABEE7A" box="[1155,1316,549,572]" italics="true" pageId="8" pageNumber="223">C. propinquus</emphasis>
(De 
<bibRefCitation id="EFB84B47FFF1FF8F5E3EB325FF5FEE26" author="Man" pageId="8" pageNumber="223" refString="Man, J. G. de (1905) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeiniging, 9 (2), 587 - 614." type="journal article" year="1905">Man, 1905</bibRefCitation>
) (
<bibRefCitation id="EFB84B47FFF1FF8F5BB7B347FEE0EE26" author="Sakai" box="[235,367,583,608]" pageId="8" pageNumber="223" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai 2011</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C294D35FFF1FF8F5AEBB348FC02EE26" authority="Komai &amp; Fujiwara, 2012" authorityName="Komai &amp; Fujiwara" authorityYear="2012" box="[439,909,583,608]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="cavifrons">
<emphasis id="B95DEAA4FFF1FF8F5AEBB348FDCCEE26" box="[439,579,584,608]" italics="true" pageId="8" pageNumber="223">C. cavifrons</emphasis>
(
<bibRefCitation id="EFB84B47FFF1FF8F590BB347FC0AEE26" author="Komai" box="[599,901,583,608]" pageId="8" pageNumber="223" refString="Komai, T. &amp; Fujiwara, Y. (2012) New records of callianassid ghost shrimp (Crustacea: Decapoda: Axiidea) from reducing environments in Kyushu, southwestern Japan. Zootaxa, 3271, 55 - 67." type="journal article" year="2012">Komai &amp; Fujiwara, 2012</bibRefCitation>
)
</taxonomicName>
. Also, the additional characters of the major cheliped (7 ischial spines and a single proximal meral spine) and the distinctive post-corneal spine on each eyestalk are considered diagnostic.
</paragraph>
</subSubSection>
<subSubSection id="C333653DFFF1FF8F5B9BB3B4FB8AEE8A" box="[199,1029,692,717]" pageId="8" pageNumber="223" type="etymology">
<paragraph id="8B9636B6FFF1FF8F5B9BB3B4FB8AEE8A" blockId="8.[151,1436,151,717]" box="[199,1029,692,717]" pageId="8" pageNumber="223">
<emphasis id="B95DEAA4FFF1FF8F5B9BB3B4FEC9EE8B" bold="true" box="[199,326,692,717]" pageId="8" pageNumber="223">Etymology</emphasis>
. the species name “ 
<taxonomicName id="4C294D35FFF1FF8F5979B3B5FD0EEE8A" box="[549,641,693,716]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="B95DEAA4FFF1FF8F5979B3B5FD0EEE8A" box="[549,641,693,716]" italics="true" pageId="8" pageNumber="223">iranicus</emphasis>
</taxonomicName>
” refers to the 
<typeStatus id="54928814FFF1FF8F587DB3B5FCDEEE8B" box="[801,849,693,717]" pageId="8" pageNumber="223">type</typeStatus>
locality in 
<collectingCountry id="F33E7626FFF1FF8F588CB3B5FB8EEE8A" box="[976,1025,693,716]" name="Iran" pageId="8" pageNumber="223">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>