treatments-xml/data/03/80/87/0380878FFFA4FF83FDD0FCEBC01EFE3A.xml

296 lines
67 KiB
XML

<document id="720EC2A8FDF1FD84D3DF48E9CC2ADE54" ID-DOI="10.5281/zenodo.12996779" ID-Zenodo-Dep="12996779" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="luisschmitz" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1721870935690" checkinUser="luisschmitz" docAuthor="Gerry Cassis &amp; Loren Vanags" docDate="2006" docId="0380878FFFA4FF83FDD0FCEBC01EFE3A" docLanguage="en" docName="Denisia.19.275-398.pdf.imf" docOrigin="Denisia 19" docTitle="Austrotichus GROSS 1975" docType="treatment" docVersion="8" lastPageNumber="303" masterDocId="FFB9FFF7FFBFFF9DFFC4FF8CC239FFED" masterDocTitle="Jewel Bugs of Australia (Insecta, Heteroptera, Scutelleridae)" masterLastPageNumber="398" masterPageNumber="275" pageNumber="302" updateTime="1725646846613" updateUser="ExternalLinkService">
<mods:mods id="37BE345A58B35E400AFE45B5E5E61FA5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="902732EEA33E46ADAA9D01E2F6B41DC3">
<mods:title id="626C0D2E1831B4A2AD29178359C3697E">Jewel Bugs of Australia (Insecta, Heteroptera, Scutelleridae)</mods:title>
</mods:titleInfo>
<mods:name id="0DCF934399529C11CB96EC862B545936" type="personal">
<mods:role id="88D7451A1ECB26E82BB52F4550DFF79E">
<mods:roleTerm id="85E728B70E8A2EBA53D2EC88BD50414A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DC24E2A690CC5A2EAA1A09FDD6B76DEF">Gerry Cassis</mods:namePart>
</mods:name>
<mods:name id="EC135A0505A3FC8AF02735BFE8AAF550" type="personal">
<mods:role id="EF60AE6AEBEA9638E05FDCEF66BFF34C">
<mods:roleTerm id="D1A7F755821C087A64F94087FEDBB5E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6623DDF329F0E9AF3AD4ACFEFF430F64">Loren Vanags</mods:namePart>
</mods:name>
<mods:typeOfResource id="8CA2A3611FC9EAD634832E5B609A7D80">text</mods:typeOfResource>
<mods:relatedItem id="1E338EB46749EA38BC26D7BCD85056B6" type="host">
<mods:titleInfo id="48450F71E69E2269A500DA93B422F4FE">
<mods:title id="53561A478A53DFFAD7DD3703E3A2453E">Denisia</mods:title>
</mods:titleInfo>
<mods:part id="B0526725F3C54B9EE7BAC428810439B3">
<mods:date id="8E3F4106A47821952640AC2F5E905CD1">2006</mods:date>
<mods:detail id="27EB8915D0F8A29662003D949A78296D" type="volume">
<mods:number id="6FA167F30603A59912961FE2FF00DC36">19</mods:number>
</mods:detail>
<mods:extent id="BD21E949931B753258FB8FC5012E29C9" unit="page">
<mods:start id="A86A66F93DC70E62E4D343618462A3EB">275</mods:start>
<mods:end id="4A123A8119206A0D8168FD2125AC9FB1">398</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B01EA47D7D79B576C7A2A085B5A71DA2">journal article</mods:classification>
<mods:identifier id="74C03B4E0429C1743438D19FB42D8C19" type="DOI">10.5281/zenodo.12996779</mods:identifier>
<mods:identifier id="0A547B481BDD7B49ABD8C53C15433B74" type="Zenodo-Dep">12996779</mods:identifier>
</mods:mods>
<treatment id="0380878FFFA4FF83FDD0FCEBC01EFE3A" LSID="urn:lsid:plazi:treatment:0380878FFFA4FF83FDD0FCEBC01EFE3A" httpUri="http://treatment.plazi.org/id/0380878FFFA4FF83FDD0FCEBC01EFE3A" lastPageId="30" lastPageNumber="303" pageId="27" pageNumber="302">
<subSubSection id="C3336512FFA4FF86FDD0FCEBC0FCFC72" pageId="27" pageNumber="302" type="nomenclature">
<paragraph id="8B963699FFA4FF86FDD0FCEBC0FCFC72" blockId="27.[532,966,110,1917]" pageId="27" pageNumber="302">
<taxonomicName id="4C294D1AFFA4FF86FDD0FCEBC11AFC92" authority="GROSS 1975" authorityName="GROSS" authorityYear="1975" box="[532,803,871,895]" class="Insecta" family="Scutelleridae" genus="Austrotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA4FF86FDD0FCEBC098FC92" box="[532,673,871,895]" italics="true" pageId="27" pageNumber="302">Austrotichus</emphasis>
<bibRefCitation id="EFB84B68FFA4FF86FD6DFCEBC11AFC92" author="GROSS G. F." box="[681,803,871,895]" journalOrPublisher="A. B. James, Adelaide" pageId="27" pageNumber="302" pagination="1 - 250" refId="ref84925" refString="GROSS G. F. (1975): Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1. A. B. James, Adelaide: 1 - 250, 3 colour pls." title="Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1" type="book chapter" year="1975">
<smallCapsWord id="8D70A045FFA4FF86FD6DFCEBC0DCFC93" baselines="889,889" box="[681,741,871,895]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gross" pageId="27" pageNumber="302">GROSS</smallCapsWord>
1975
</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13122A1CFFA4FF86FDDDFC0BC055FC72" box="[537,620,903,927]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="27" pageNumber="302">Figs 3a</figureCitation>
,
<figureCitation id="13122A1CFFA4FF86FDBCFC0BC0B0FC72" box="[632,649,903,927]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">4</figureCitation>
,
<figureCitation id="13122A1CFFA4FF86FD50FC0BC09CFC72" box="[660,677,903,927]" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="27" pageNumber="302">5</figureCitation>
,
<figureCitation id="13122A1CFFA4FF86FD6BFC0BC087FC72" box="[687,702,903,927]" captionStart="Fig" captionStartId="30.[1089,1125,859,878]" captionTargetBox="[632,1514,139,831]" captionTargetId="figure-558@30.[620,1532,106,834]" captionTargetPageId="30" captionText="Fig. 6: Distribution of Australian Elvisurinae species." figureDoi="http://doi.org/10.5281/zenodo.13713975" httpUri="https://zenodo.org/record/13713975/files/figure.png" pageId="27" pageNumber="302">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3336512FFA4FF86FDD0FC3CC06CFBCC" pageId="27" pageNumber="302" type="reference_group">
<paragraph id="8B963699FFA4FF86FDD0FC3CC06CFBCC" blockId="27.[532,966,110,1917]" pageId="27" pageNumber="302">
<taxonomicName id="4C294D1AFFA4FF86FDD0FC3CC127FC28" authority="GROSS 1975: 83" authorityName="GROSS" authorityPageNumber="83" authorityYear="1975" box="[532,798,944,965]" class="Insecta" family="Scutelleridae" genus="Austrotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA4FF86FDD0FC3CC045FC28" box="[532,636,944,965]" italics="true" pageId="27" pageNumber="302">Austrotichus</emphasis>
<bibRefCitation id="EFB84B68FFA4FF86FD43FC3CC127FC28" author="GROSS G. F." box="[647,798,944,965]" journalOrPublisher="A. B. James, Adelaide" pageId="27" pageNumber="302" pagination="1 - 250" refId="ref84925" refString="GROSS G. F. (1975): Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1. A. B. James, Adelaide: 1 - 250, 3 colour pls." title="Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1" type="book chapter" year="1975">
<smallCapsWord id="8D70A045FFA4FF86FD43FC3CC0F8FC2E" baselines="960,959" box="[647,705,944,965]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Gross" pageId="27" pageNumber="302">GROSS</smallCapsWord>
1975: 83
</bibRefCitation>
</taxonomicName>
(gen. nov.);
<smallCapsWord id="8D70A045FFA4FF86FC5BFC3CC067FC0F" baselines="960,959,960,991,990" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Mc-Donald" pageId="27" pageNumber="302">MC-DONALD</smallCapsWord>
&amp;
<smallCapsWord id="8D70A045FFA4FF86FDBBFC43C085FC0F" baselines="991,990" box="[639,700,975,996]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Cassis" pageId="27" pageNumber="302">CASSIS</smallCapsWord>
1984: 538, 545 (key description, genitalia);
<bibRefCitation id="EFB84B68FFA4FF86FD67FC61C1ACFBEF" author="CASSIS G. &amp; G. F. GROSS" box="[675,917,1005,1026]" editor="HOUSTON W. W. K. &amp; G. V. MAYNARD" journalOrPublisher="CSIRO Publishing, Melbourne: xiv" pageId="27" pageNumber="302" pagination="1 - 732" refId="ref83117" refString="CASSIS G. &amp; G. F. GROSS (2002): Hemiptera: Heteroptera (Pentatomomorpha). - In: HOUSTON W. W. K. &amp; G. V. MAYNARD (Eds), Zoological Catalogue of Australia. Vol. 27.3 B. CSIRO Publishing, Melbourne: xiv + 1 - 732." title="Hemiptera: Heteroptera (Pentatomomorpha)" type="book chapter" volumeTitle="Zoological Catalogue of Australia. Vol. 27.3 B" year="2002">
<smallCapsWord id="8D70A045FFA4FF86FD67FC61C0D9FBED" baselines="1021,1020" box="[675,736,1005,1026]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Cassis" pageId="27" pageNumber="302">CASSIS</smallCapsWord>
&amp;
<smallCapsWord id="8D70A045FFA4FF86FD3AFC61C101FBED" baselines="1021,1020" box="[766,824,1005,1026]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Gross" pageId="27" pageNumber="302">GROSS</smallCapsWord>
2002: 582
</bibRefCitation>
(catalogue)
</paragraph>
</subSubSection>
<subSubSection id="C3336512FFA4FF86FDD0FBA7C0FDFBB3" pageId="27" pageNumber="302" type="type_taxon">
<paragraph id="8B963699FFA4FF86FDD0FBA7C0FDFBB3" blockId="27.[532,966,110,1917]" pageId="27" pageNumber="302">
Type species:
<taxonomicName id="4C294D1AFFA4FF86FD4EFBA7C19EFBAD" authority="GROSS 1975" authorityName="GROSS" authorityYear="1975" box="[650,935,1067,1088]" class="Insecta" family="Scutelleridae" genus="Austrotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="302" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="B95DEA8BFFA4FF86FD4EFBA7C10FFBAD" box="[650,822,1067,1088]" italics="true" pageId="27" pageNumber="302">Austrotichus rugosus</emphasis>
<bibRefCitation id="EFB84B68FFA4FF86FCFFFBA7C19EFBAD" author="GROSS G. F." box="[827,935,1067,1088]" journalOrPublisher="A. B. James, Adelaide" pageId="27" pageNumber="302" pagination="1 - 250" refId="ref84925" refString="GROSS G. F. (1975): Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1. A. B. James, Adelaide: 1 - 250, 3 colour pls." title="Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1" type="book chapter" year="1975">
<smallCapsWord id="8D70A045FFA4FF86FCFFFBA7C14CFBD3" baselines="1083,1082" box="[827,885,1067,1088]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Gross" pageId="27" pageNumber="302">GROSS</smallCapsWord>
1975
</bibRefCitation>
</taxonomicName>
, by original designation
</paragraph>
</subSubSection>
<subSubSection id="C3336512FFA4FF86FDFEFBF4C054F96E" pageId="27" pageNumber="302" type="diagnosis">
<paragraph id="8B963699FFA4FF86FDFEFBF4C054F96E" blockId="27.[532,966,110,1917]" pageId="27" pageNumber="302">
Diagnosis:
<taxonomicName id="4C294D1AFFA4FF86FD63FBF4C122FB62" authorityName="GROSS" authorityYear="1975" box="[679,795,1144,1167]" class="Insecta" family="Scutelleridae" genus="Austrotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA4FF86FD63FBF4C122FB62" box="[679,795,1144,1167]" italics="true" pageId="27" pageNumber="302">Austrotichus</emphasis>
</taxonomicName>
is recognised by the following characters: posterior margin of pronotum rectilinear (
<figureCitation id="13122A1CFFA4FF86FCC0FB37C16FFB3F" box="[772,854,1211,1234]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="27" pageNumber="302">Fig. 3a</figureCitation>
); thoracic sterna carinate, not overlapping (
<figureCitation id="13122A1CFFA4FF86FCA9FB50C180FB1E" box="[877,953,1244,1267]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4c</figureCitation>
); anterior margin of proepisternum truncate (
<figureCitation id="13122A1CFFA4FF86FDDFFA93C05DFADB" box="[539,612,1311,1334]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4c</figureCitation>
); peritreme of metathoracic glands short, arcuate (
<figureCitation id="13122A1CFFA4FF86FD6EFACCC0CDFABA" box="[682,756,1344,1367]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4d</figureCitation>
); posterior margin of male pygophore medially emarginate (
<figureCitation id="13122A1CFFA4FF86FC5BFAEDC017FA77" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4e</figureCitation>
); CAI absent (
<figureCitation id="13122A1CFFA4FF86FD04FA0FC11BFA77" box="[704,802,1411,1434]" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="27" pageNumber="302">Figs 5c, d</figureCitation>
); CAII(L) without lobal sclerite (
<figureCitation id="13122A1CFFA4FF86FD07FA28C11AFA56" box="[707,803,1444,1467]" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="27" pageNumber="302">Figs 5c, d</figureCitation>
); CAII(M) with short lobal sclerite (
<figureCitation id="13122A1CFFA4FF86FD1CFA49C102FA31" box="[728,827,1477,1500]" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="27" pageNumber="302">Figs 5c, d</figureCitation>
); CAIII bifid, medially fused, CAIII(V) with short lobal sclerite (
<figureCitation id="13122A1CFFA4FF86FDACF984C0F5F9F2" box="[616,716,1544,1567]" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="27" pageNumber="302">Figs 5c, d</figureCitation>
); gonocoxae I not divid-ed, medial margins raised; and, spermathecal reservoir membraneous, weakly dilated, elliptoid.
</paragraph>
</subSubSection>
<subSubSection id="C3336512FFA4FF81FDFEF911C395F96D" lastPageId="28" lastPageNumber="303" pageId="27" pageNumber="302" type="description">
<paragraph id="8B963699FFA4FF81FDFEF911C395F96D" blockId="27.[532,966,110,1917]" lastBlockId="28.[154,587,1426,1907]" lastPageId="28" lastPageNumber="303" pageId="27" pageNumber="302">
Description: Body elongate-ovoid (
<figureCitation id="13122A1CFFA4FF86FC5BF911C014F938" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="27" pageNumber="302">Fig. 3a</figureCitation>
); dorsum and venter strongly convex; wing tip not extending beyond scutellum (
<figureCitation id="13122A1CFFA4FF86FDDFF88DC05DF8F5" box="[539,612,1793,1816]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="27" pageNumber="302">Fig. 3a</figureCitation>
). Head: mostly flattened, posterior aspect of vertex weakly convex (
<figureCitation id="13122A1CFFA4FF86FC95F8AFC1AEF8D7" box="[849,919,1827,1850]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4a</figureCitation>
); jugal margins thickened, subcarinate (
<figureCitation id="13122A1CFFA4FF86FCBCF8C9C01AF890" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Figs 4a, b</figureCitation>
); bucculae margins subparallel, weakly expanded anteriorly (
<figureCitation id="13122A1CFFA4FF86FB7AFFE2C72BFF68" box="[1214,1298,110,133]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4c</figureCitation>
). Pronotum: strongly transverse, subtrapezoidal; anterior margin weakly concave (
<figureCitation id="13122A1CFFA4FF86FB1AFF3DC71DFF25" box="[1246,1316,177,200]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="27" pageNumber="302">Fig. 3a</figureCitation>
); anterolateral margins subcarinate; posterior margin rectilinear (
<figureCitation id="13122A1CFFA4FF86FB98FF78C69DFEE6" box="[1116,1188,244,267]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="27" pageNumber="302">Fig. 3a</figureCitation>
); callosite region weakly demarcated; disc strongly convex, declivent posteriorly (
<figureCitation id="13122A1CFFA4FF86FBAFFEB4C686FEA2" box="[1131,1215,312,335]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="27" pageNumber="302">Fig. 3a</figureCitation>
). Scutellum: broad, strongly depressed beyond anterior 1/2; elliptoid tumescence on anterior margin, marked sublaterally by dark foveae (
<figureCitation id="13122A1CFFA4FF86FA83FE11C7B4FE59" box="[1351,1421,413,436]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="27" pageNumber="302">Fig. 3a</figureCitation>
); lateral margins convex (
<figureCitation id="13122A1CFFA4FF86FB2DFE32C70FFE38" box="[1257,1334,446,469]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="27" pageNumber="302">Fig. 3a</figureCitation>
); base of corium and clavus, and connexiva III-VII exposed (
<figureCitation id="13122A1CFFA4FF86FB81FD8EC6B2FDF4" box="[1093,1163,514,537]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="27" pageNumber="302">Fig. 3a</figureCitation>
). Thoracic pleura: external efferent system of metathoracic glands well-developed (
<figureCitation id="13122A1CFFA4FF86FB9EFDC9C69BFDB1" box="[1114,1186,581,604]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4d</figureCitation>
); evaporative areas occupying most of metepisternum, extending to mesepimeron (
<figureCitation id="13122A1CFFA4FF86FBB9FD04C6F2FD72" box="[1149,1227,648,671]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4d</figureCitation>
); ostiole small (
<figureCitation id="13122A1CFFA4FF86FAB7FD04C63DFD2C" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4d</figureCitation>
); peritreme short, raised, reniform, not medially sulcate (
<figureCitation id="13122A1CFFA4FF86FB5CFD40C6DBFD0E" box="[1176,1250,716,739]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4d</figureCitation>
). Thoracic sterna: lateral margins sulcate, prosternal and mesosternal keels not overlapping (
<figureCitation id="13122A1CFFA4FF86FA82FC83C7B4FCCB" box="[1350,1421,783,806]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4c</figureCitation>
); anterior margin of proepisternum truncate, not elevated (
<figureCitation id="13122A1CFFA4FF86FBB1FCDEC686FC84" box="[1141,1215,850,873]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Fig. 4c</figureCitation>
). Male Genitalia: pygophore (
<figureCitation id="13122A1CFFA4FF86FB8DFCF8C694FC66" box="[1097,1197,884,907]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Figs 4e, f</figureCitation>
,
<figureCitation id="13122A1CFFA4FF86FB7FFCF8C6EAFC66" box="[1211,1235,884,907]" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="27" pageNumber="302">5a</figureCitation>
) caudally oriented, moderately large, transverse; genital opening dorsal, narrow, suboval, lateral margins setose (
<figureCitation id="13122A1CFFA4FF86FBF6FC55C641FC1D" box="[1074,1144,985,1008]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="27" pageNumber="302">Figs 4f</figureCitation>
,
<figureCitation id="13122A1CFFA4FF86FB41FC55C6A6FC1D" box="[1157,1183,985,1008]" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="27" pageNumber="302">5a</figureCitation>
); parameres with columnar stem, moderately hook-shaped crown, flange at base of crown (
<figureCitation id="13122A1CFFA4FF86FB27FB90C717FBDE" box="[1251,1326,1052,1075]" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="27" pageNumber="302">Fig. 5b</figureCitation>
); aedeagus (
<figureCitation id="13122A1CFFA4FF86FC2BFBB2C65DFBB8" box="[1007,1124,1086,1109]" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="27" pageNumber="302">Figs 5c, d</figureCitation>
): phallotheca short, without processes; ductus seminis proximalis narrow, singular, membraneous, tubelike; ejaculatory apparatus moderately developed; ventral conducting canal with about eight paired convolutions, heavily sclerotized; ejaculatory reservoir small, subelliptical; dorsal conducting canal narrow; ductus seminis distalis and vesica basally incrassate, sinuate, apically truncate; CAI absent; paired CAII greatly enlarged, membraneous, distally bifurcate, lateral branch without sclerotization, medial branch with small, acute lobal sclerite (
<figureCitation id="13122A1CFFA4FF86FB85FA78C697F9E6" box="[1089,1198,1524,1547]" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="27" pageNumber="302">Figs 5c, d</figureCitation>
); paired CAIII mostly membraneous, with lateral margins sclerotized, bifurcate, lateral branch membraneous, medial branch with small acute lobal sclerite, medial pair fused along midline (
<figureCitation id="13122A1CFFA4FF86FC2BF910C676F95E" box="[1007,1103,1692,1715]" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="27" pageNumber="302">Figs 5c, d</figureCitation>
). Female Abdominal Venter: posterior margin of SVII weakly bisinuate, weakly thickened medially. Female Terminalia: caudally oriented, bi-planar; paratergites VIII subelliptoid, medially raised; paratergites IX small, narrowly elliptoid, medially rounded; gonocoxae I moderately large, medially depressed, posterior margin concave, medial margins elevated. Spermatheca: short proximal fecundation canal; spermathecal reservoir narrowly dilated, elliptoid, membraneous, without pronounced modifications; pump short, with dorsal and proximal flanges; bulb oval.
</paragraph>
</subSubSection>
<caption id="DF566611FFA3FF81FB30FFE7C796FCDE" ID-DOI="http://doi.org/10.5281/zenodo.13713969" ID-Zenodo-Dep="13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="28" pageNumber="303" startId="28.[1268,1304,107,126]" subCaptionStartIDs="28.[1268,1410,480,499]" subCaptionStarts="Abbr" targetBox="[151,1252,106,1409]" targetPageId="28" targetType="figure">
<paragraph id="8B963699FFA3FF81FB30FFE7C796FCDE" blockId="28.[1268,1526,107,819]" pageId="28" pageNumber="303">
<emphasis id="B95DEA8BFFA3FF81FB30FFE7C709FF93" bold="true" box="[1268,1328,107,126]" pageId="28" pageNumber="303">Fig. 4</emphasis>
: Scanning electron micrographs of key characters of
<taxonomicName id="4C294D1AFFA3FF81FB30FF37C7FDFF23" authorityName="GROSS" authorityYear="1975" box="[1268,1476,187,206]" class="Insecta" family="Scutelleridae" genus="Austrotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="303" phylum="Arthropoda" rank="species" species="rugosus">Austrotichus rugosus</taxonomicName>
(
<emphasis id="B95DEA8BFFA3FF81FB3FFF5AC731FF04" bold="true" box="[1275,1288,214,233]" pageId="28" pageNumber="303">a</emphasis>
) Head, dorsal view (
<emphasis id="B95DEA8BFFA3FF81FB3FFF7CC730FEEE" bold="true" box="[1275,1289,240,259]" pageId="28" pageNumber="303">b</emphasis>
) Head, lateral view (
<emphasis id="B95DEA8BFFA3FF81FB3FFE87C73FFEF3" bold="true" box="[1275,1286,267,286]" pageId="28" pageNumber="303">c</emphasis>
) Head and thorax, ventral view (
<emphasis id="B95DEA8BFFA3FF81FAB9FEAAC7B2FED4" bold="true" box="[1405,1419,294,313]" pageId="28" pageNumber="303">d</emphasis>
) External efferent system of metathoracic glands, ventral view (
<emphasis id="B95DEA8BFFA3FF81FAB9FEFAC7B3FE64" bold="true" box="[1405,1418,374,393]" pageId="28" pageNumber="303">e</emphasis>
) male pygophore, ventral view (
<emphasis id="B95DEA8BFFA3FF81FB3FFE27C73DFE53" bold="true" box="[1275,1284,427,446]" pageId="28" pageNumber="303">f</emphasis>
) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="C3336512FFA3FF81FF04F917C079F8F0" pageId="28" pageNumber="303" type="distribution">
<paragraph id="8B963699FFA3FF81FF04F917C079F8F0" blockId="28.[154,587,1426,1907]" pageId="28" pageNumber="303">
Diversity and distribution:
<taxonomicName id="4C294D1AFFA3FF81FE00F917C00FF95F" authorityName="GROSS" authorityYear="1975" box="[452,566,1691,1714]" class="Insecta" family="Scutelleridae" genus="Austrotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA3FF81FE00F917C00FF95F" box="[452,566,1691,1714]" italics="true" pageId="28" pageNumber="303">Austrotichus</emphasis>
</taxonomicName>
is a monotypic genus, which is broadly distributed in temperate Australia, including Victoria, South Australia and Western Australia.
</paragraph>
</subSubSection>
<subSubSection id="C3336512FFA3FF81FF5EF8B5C073F89E" pageId="28" pageNumber="303" type="taxon_list">
<paragraph id="8B963699FFA3FF81FF5EF8B5C073F89E" blockId="28.[154,587,1426,1907]" pageId="28" pageNumber="303">
<emphasis id="B95DEA8BFFA3FF81FF5EF8B5C378F8BD" bold="true" box="[154,321,1849,1872]" pageId="28" pageNumber="303">Included species</emphasis>
:
<taxonomicName id="4C294D1AFFA3FF81FF5EF8D0C347F89E" authority="GROSS 1975" authorityName="GROSS" authorityYear="1975" box="[154,382,1884,1907]" class="Insecta" family="Scutelleridae" genus="Austrotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="303" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="B95DEA8BFFA3FF81FF5EF8D0C2C6F89E" box="[154,255,1884,1907]" italics="true" pageId="28" pageNumber="303">A. rugosus</emphasis>
<bibRefCitation id="EFB84B68FFA3FF81FEC2F8D0C347F89E" author="GROSS G. F." box="[262,382,1884,1907]" journalOrPublisher="A. B. James, Adelaide" pageId="28" pageNumber="303" pagination="1 - 250" refId="ref84925" refString="GROSS G. F. (1975): Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1. A. B. James, Adelaide: 1 - 250, 3 colour pls." title="Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1" type="book chapter" year="1975">GROSS 1975</bibRefCitation>
</taxonomicName>
Australia
</paragraph>
</subSubSection>
<subSubSection id="C3336512FFA3FF83FD50FA1EC01EFE3A" lastPageId="30" lastPageNumber="305" pageId="28" pageNumber="303" type="discussion">
<paragraph id="8B963699FFA3FF83FD50FA1EC01EFE3A" blockId="28.[622,1056,1426,1917]" lastBlockId="30.[154,588,110,1917]" lastPageId="30" lastPageNumber="305" pageId="28" pageNumber="303">
Remarks:
<bibRefCitation id="EFB84B68FFA3FF81FD3CFA1EC1BDFA44" author="GROSS G. F." box="[760,900,1426,1449]" journalOrPublisher="A. B. James, Adelaide" pageId="28" pageNumber="303" pagination="1 - 250" refId="ref84925" refString="GROSS G. F. (1975): Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1. A. B. James, Adelaide: 1 - 250, 3 colour pls." title="Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1" type="book chapter" year="1975">
<smallCapsWord id="8D70A045FFA3FF81FD3CFA1EC100FA45" baselines="1443,1444" box="[760,825,1426,1449]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Gross" pageId="28" pageNumber="303">GROSS</smallCapsWord>
(1975)
</bibRefCitation>
described
<taxonomicName id="4C294D1AFFA3FF81FC36FA1EC082FA27" authorityName="GROSS" authorityYear="1975" class="Insecta" family="Scutelleridae" genus="Austrotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA3FF81FC36FA1EC082FA27" italics="true" pageId="28" pageNumber="303">Austrotichus</emphasis>
</taxonomicName>
as a monotypic genus, based on external characters alone.
<bibRefCitation id="EFB84B68FFA3FF81FC9DFA59C0EDF9E0" author="MCDONALD F. J. D. &amp; G. CASSIS" journalOrPublisher="Aust. J. Zool." pageId="28" pageNumber="303" pagination="537 - 572" part="32" refId="ref87212" refString="MCDONALD F. J. D. &amp; G. CASSIS (1984): Revision of the Australian Scutelleridae LEACH (Hemiptera). - Aust. J. Zool. 32: 537 - 572." title="Revision of the Australian Scutelleridae LEACH (Hemiptera)" type="journal article" year="1984">
<smallCapsWord id="8D70A045FFA3FF81FC9DFA59C142FA06" baselines="1510,1511" box="[857,891,1493,1516]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Mc" pageId="28" pageNumber="303">
<smallCapsWord id="8D70A045FFA3FF81FCBFFA59C1F7FA06" baselines="1510,1511" box="[891,974,1493,1516]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Donald" pageId="28" pageNumber="303">MCDONALD</smallCapsWord>
</smallCapsWord>
&amp;
<smallCapsWord id="8D70A045FFA3FF81FC2AFA59C0B0F9E1" baselines="1510,1511,1510,1544" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Cassis" pageId="28" pageNumber="303">CASSIS</smallCapsWord>
(1984)
</bibRefCitation>
supported its generic status, and described the male and female genitalia. It is a distinctive genus of jewel bug, with a robust body, apomorphic male genitalia and metathoracic glands, and specialized host plant associations. Unlike species of
<taxonomicName id="4C294D1AFFA3FF81FDABF933C0E0F93B" box="[623,729,1727,1750]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA3FF81FDABF933C0E0F93B" box="[623,729,1727,1750]" italics="true" pageId="28" pageNumber="303">Coleotichus</emphasis>
</taxonomicName>
, this genus has the posterior margin of the pronotum rectilinear (
<figureCitation id="13122A1CFFA3FF81FC6FF96CC1D7F91A" box="[939,1006,1760,1783]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="28" pageNumber="303">Fig. 3a</figureCitation>
) (also in
<taxonomicName id="4C294D1AFFA3FF81FD6EF88EC123F8F4" box="[682,794,1794,1817]" class="Insecta" family="Scutelleridae" genus="Solenotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA3FF81FD6EF88EC123F8F4" box="[682,794,1794,1817]" italics="true" pageId="28" pageNumber="303">Solenotichus</emphasis>
</taxonomicName>
and Afrotropical
<taxonomicName id="4C294D1AFFA3FF81FC17F88EC137F8D7" authority="SPINOLA" authorityName="SPINOLA" class="Insecta" family="Scutelleridae" genus="Solenosthedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA3FF81FC17F88EC08EF8D7" italics="true" pageId="28" pageNumber="303">Solenosthedium</emphasis>
<smallCapsWord id="8D70A045FFA3FF81FD79F8AFC137F8D7" baselines="1844,1846" box="[701,782,1827,1850]" lowerCaseFontSize="10" mainFontSize="10" normCase="title" normString="Spinola" pageId="28" pageNumber="303">SPINOLA</smallCapsWord>
</taxonomicName>
) and the anterior margin of the scutellum is moderately tumose, with sublateral dark foveae (
<figureCitation id="13122A1CFFA3FF81FC99F8EAC19EF890" box="[861,935,1894,1917]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="28" pageNumber="303">Fig. 3a</figureCitation>
). The male genitalia of
<taxonomicName id="4C294D1AFFA1FF83FEDFFFE2C3E6FF68" authorityName="GROSS" authorityYear="1975" box="[283,479,110,133]" class="Insecta" family="Scutelleridae" genus="Austrotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="305" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="B95DEA8BFFA1FF83FEDFFFE2C3E6FF68" box="[283,479,110,133]" italics="true" pageId="30" pageNumber="305">Austrotichus rugosus</emphasis>
</taxonomicName>
, with the paired CAIII mostly membraneous (
<figureCitation id="13122A1CFFA1FF83FDC4FF03C290FF25" captionStart="Fig" captionStartId="29.[1073,1109,107,126]" captionTargetBox="[68,1435,106,1920]" captionTargetId="figure-15@29.[68,1435,105,1921]" captionTargetPageId="29" captionText="Fig. 5: Male genitalia of Austrotichus rugosus (a) pygophore, dorsal view (b) right paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII(D) = third conjunctival appendage, dorsal branch; CAIII(V) = third conjunctival appendage, ventral branch; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713971" httpUri="https://zenodo.org/record/13713971/files/figure.png" pageId="30" pageNumber="305">Fig. 5c, d</figureCitation>
), differs from species of
<taxonomicName id="4C294D1AFFA1FF83FE1FFF3DC07CFF25" box="[475,581,177,200]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA1FF83FE1FFF3DC07CFF25" box="[475,581,177,200]" italics="true" pageId="30" pageNumber="305">Coleotichus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D1AFFA1FF83FF5EFF5FC3BAFF07" box="[154,387,211,234]" class="Insecta" family="Scutelleridae" genus="Solenotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="305" phylum="Arthropoda" rank="species" species="circuliferus">
<emphasis id="B95DEA8BFFA1FF83FF5EFF5FC3BAFF07" box="[154,387,211,234]" italics="true" pageId="30" pageNumber="305">Solenotichus circuliferus</emphasis>
</taxonomicName>
, and extralimital species of
<taxonomicName id="4C294D1AFFA1FF83FEC2FF79C34CFEE1" box="[262,373,245,268]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="300" phylum="Arthropoda" rank="subFamily" subFamily="Elvisurinae">Elvisurinae</taxonomicName>
, where the CAIII is sclerotized. In addition, the peritreme is short in
<taxonomicName id="4C294D1AFFA1FF83FF29FEB4C359FEA2" authorityName="GROSS" authorityYear="1975" box="[237,352,312,335]" class="Insecta" family="Scutelleridae" genus="Austrotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA1FF83FF29FEB4C359FEA2" box="[237,352,312,335]" italics="true" pageId="30" pageNumber="305">Austrotichus</emphasis>
</taxonomicName>
(
<figureCitation id="13122A1CFFA1FF83FEAAFEB4C38FFEA2" box="[366,438,312,335]" captionStart="Fig" captionStartId="28.[1268,1304,107,126]" captionTargetBox="[151,1252,106,1409]" captionTargetId="figure-299@28.[151,1253,106,1409]" captionTargetPageId="28" captionText="Fig. 4: Scanning electron micrographs of key characters of Austrotichus rugosus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Go = genital opening; J = juga; Me = mesepimeron; Met = metepisternum; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713969" httpUri="https://zenodo.org/record/13713969/files/figure.png" pageId="30" pageNumber="305">Fig. 4d</figureCitation>
), much shorter than in
<taxonomicName id="4C294D1AFFA1FF83FF28FED6C365FE9C" box="[236,348,346,369]" class="Insecta" family="Scutelleridae" genus="Solenotichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA1FF83FF28FED6C365FE9C" box="[236,348,346,369]" italics="true" pageId="30" pageNumber="305">Solenotichus</emphasis>
</taxonomicName>
(
<figureCitation id="13122A1CFFA1FF83FEAFFED6C38CFE9C" box="[363,437,346,369]" captionStart="Fig" captionStartId="42.[1274,1310,107,126]" captionTargetBox="[151,1258,106,1407]" captionTargetId="figure-263@42.[151,1258,106,1408]" captionTargetPageId="42" captionText="Fig. 9: Scanning electron micrographs of key characters of Solenotichus circuliferus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; P(r) = right paramere. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713983" httpUri="https://zenodo.org/record/13713983/files/figure.png" pageId="30" pageNumber="305">Fig. 9d</figureCitation>
), still apparent in comparison to
<taxonomicName id="4C294D1AFFA1FF83FE88FEF0C3E1FE7E" box="[332,472,380,403]" class="Insecta" family="Scutelleridae" genus="Solenosthedium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA1FF83FE88FEF0C3E1FE7E" box="[332,472,380,403]" italics="true" pageId="30" pageNumber="305">Solenosthedium</emphasis>
</taxonomicName>
, and much shorter than the consistently elongate peritreme of species of
<taxonomicName id="4C294D1AFFA1FF83FE9FFE4CC3FCFE3A" box="[347,453,448,471]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFA1FF83FE9FFE4CC3FCFE3A" box="[347,453,448,471]" italics="true" pageId="30" pageNumber="305">Coleotichus</emphasis>
</taxonomicName>
(
<figureCitation id="13122A1CFFA1FF83FE16FE4CC022FE3A" box="[466,539,448,471]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="30" pageNumber="305">Fig. 7d</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>