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<document ID-DOI="http://doi.org/10.5281/zenodo.3794353" ID-GBIF-Dataset="43234c4b-40c9-4d1b-aa5f-51d222a421b2" ID-GBIF-Taxon="163963895" ID-ISSN="1876-312X" ID-Zenodo-Dep="3794353" ID-ZooBank="217D79EE-A577-4B7F-8AB0-1107C9FC4857" approvalRequired="3" approvalRequired_for_textStreams="3" checkinTime="1588686095692" checkinUser="felipe" docAuthor="Cruz, Paulo Vilela, Salles, Frederico Falcão &amp; Hamada, Neusa" docDate="2020" docId="074FCA78FFBEFFD7FFE1A0B4FEDD3D10" docLanguage="en" docName="InsectSystEvol.51.2.256-275.pdf.imf" docOrigin="Insect Systematics &amp; Evolution 51 (2)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2a2c15f9" docStyleName="insectsystevol.2020.journal_article" docTitle="Rhopyscelis singuil Cruza &amp; Sallesb &amp; Hamada 2020, comb. n." docType="treatment" docVersion="4" lastPageNumber="272" masterDocId="FB76B200FFAEFFC7FF83A260FFAD382B" masterDocTitle="A cladistic approach for generic delimitation of Paracloeodes Day, Rivudiva Lugo-Ortiz &amp; McCafferty, and Varipes Lugo-Ortiz &amp; McCafferty (Ephemeroptera: Baetidae)" masterLastPageNumber="275" masterPageNumber="256" pageId="16" pageNumber="272" updateTime="1643438799729" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A cladistic approach for generic delimitation of Paracloeodes Day, Rivudiva Lugo-Ortiz &amp; McCafferty, and Varipes Lugo-Ortiz &amp; McCafferty (Ephemeroptera: Baetidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cruz, Paulo Vilela</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salles, Frederico Falcão</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hamada, Neusa</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Insect Systematics &amp; Evolution</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-03-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>51</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>256</mods:start>
<mods:end>275</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1163/1876312X-00002199</mods:identifier>
<mods:identifier type="GBIF-Dataset">43234c4b-40c9-4d1b-aa5f-51d222a421b2</mods:identifier>
<mods:identifier type="ISSN">1876-312X</mods:identifier>
<mods:identifier type="Zenodo-Dep">3786794</mods:identifier>
<mods:identifier type="ZooBank">217D79EE-A577-4B7F-8AB0-1107C9FC4857</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3794353" ID-GBIF-Taxon="163963895" ID-Zenodo-Dep="3794353" LSID="urn:lsid:plazi:treatment:074FCA78FFBEFFD7FFE1A0B4FEDD3D10" httpUri="http://treatment.plazi.org/id/074FCA78FFBEFFD7FFE1A0B4FEDD3D10" lastPageNumber="272" pageId="16" pageNumber="272">
<subSubSection box="[98,564,724,750]" pageId="16" pageNumber="272" type="nomenclature">
<paragraph blockId="16.[98,1066,724,1339]" box="[98,564,724,750]" pageId="16" pageNumber="272">
<heading box="[98,564,724,750]" fontSize="11" level="2" pageId="16" pageNumber="272" reason="3">
<taxonomicName authority="(Nieto 2004)" authorityName="Cruza &amp; Sallesb &amp; Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[98,451,724,750]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="singuil" status="comb. n.">
<emphasis box="[98,295,724,750]" italics="true" pageId="16" pageNumber="272">Rhopyscelis singuil</emphasis>
(
<bibRefCitation author="Nieto" box="[311,442,724,750]" firstAuthor="Nieto" pageId="16" pageNumber="272" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
)
</taxonomicName>
<emphasis bold="true" box="[458,564,724,750]" pageId="16" pageNumber="272">
<taxonomicNameLabel box="[458,564,724,750]" pageId="16" pageNumber="272" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="272" type="reference_group">
<paragraph blockId="16.[98,1066,724,1339]" pageId="16" pageNumber="272">
<treatmentCitationGroup pageId="16" pageNumber="272">
<treatmentCitation author="Nieto, C." box="[98,457,758,785]" page="169" pageId="16" pageNumber="272" year="2004">
<taxonomicName authority="Nieto 2004: 169" authorityName="Nieto" authorityPageNumber="169" authorityYear="2004" box="[98,457,758,785]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="singuil">
<emphasis box="[98,256,758,784]" italics="true" pageId="16" pageNumber="272">Varipes singuil</emphasis>
<bibRefCitation author="Nieto" box="[264,457,758,785]" firstAuthor="Nieto" pageId="16" pageNumber="272" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004: 169</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation author="Dominguez, E. &amp; Molineri, C. &amp; Pescador, M. L. &amp; Hubbard, M. D. &amp; Nieto, C." box="[469,794,758,785]" page="188" pageId="16" pageNumber="272" year="2006">
<bibRefCitation author="Dominguez" box="[469,794,758,785]" etAl="et al." firstAuthor="Dominguez" pageId="16" pageNumber="272" refId="ref10411" refString="Dominguez, E., Molineri, C., Pescador, M. L., Hubbard, M. D. &amp; Nieto, C. (2006) Aquatic Biodiversity of Latin America, Ephemeroptera of South America. Vol. 2. Pensoft, Sofia-Moscow, 646 pp." type="book" year="2006">Domínguez et al. 2006: 188</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Castillo, M. M. &amp; Perez, B." page="348" pageId="16" pageNumber="272" year="2011">
<bibRefCitation author="Castillo &amp; Perez" firstAuthor="Castillo" pageId="16" pageNumber="272" pagination="346 - 349" refId="ref10191" refString="Castillo, M. M. &amp; Perez, B. (2011). Varipes (Ephemeroptera: Baetidae) em Venezuela: descripcion de nueva especie. Revista Colombiana de Entomologia 37 (1): 346 - 349." type="journal article" year="2011">Castillo &amp; Pérez 2011: 348</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Gutierrez, Y. &amp; Dias, L. G. &amp; Salles, F. F." box="[157,454,793,820]" page="295" pageId="16" pageNumber="272" year="2013">
<bibRefCitation author="Gutierrez" box="[157,454,793,820]" etAl="et al." firstAuthor="Gutierrez" pageId="16" pageNumber="272" pagination="291 - 295" refId="ref10661" refString="Gutierrez, Y., Dias, L. G. &amp; Salles, F. F. (2013) Paracloeodes caldensis (Ephemeroptera: Baetidae), an atypical new species from the Colombian Andes. Zootaxa 3721: 291 - 295." type="journal article" year="2013">Gutiérrez et al. 2013: 295</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="272" type="materials_examined">
<paragraph blockId="16.[98,1066,724,1339]" pageId="16" pageNumber="272">
<emphasis box="[98,314,828,854]" italics="true" pageId="16" pageNumber="272">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="2610422307" box="[322,873,828,854]" collectorName="One" country="Argentina" location="ARGENTINA" pageId="16" pageNumber="272" specimenCount="1" typeStatus="paratype">
<collectorName box="[322,372,828,854]" pageId="16" pageNumber="272">One</collectorName>
nymph on slide (
<typeStatus box="[578,681,828,854]" pageId="16" pageNumber="272">paratype</typeStatus>
),
<collectingCountry box="[701,873,828,854]" name="Argentina" pageId="16" pageNumber="272">ARGENTINA</collectingCountry>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2610422306" collectingDate="1999-08-05" collectorName="Molineri C." country="Argentina" county="Rio Singuil" location="Escaba" municipality="State" pageId="16" pageNumber="272" specimenCount="1" stateProvince="Tucuman" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:074FCA78FFBEFFD7FFE1A0B4FEDD3D10:8A392DB5FFBEFFD7FCF5A15CFC6B3B7D" box="[886,966,828,854]" country="Argentina" county="Rio Singuil" municipality="State" name="Escaba" pageId="16" pageNumber="272" stateProvince="Tucuman">Escaba</location>
,
<collectingMunicipality box="[979,1034,828,854]" pageId="16" pageNumber="272">State</collectingMunicipality>
of
<collectingRegion box="[98,209,863,889]" country="Argentina" name="Tucuman" pageId="16" pageNumber="272">Tucuman</collectingRegion>
,
<collectingCounty box="[222,354,862,888]" pageId="16" pageNumber="272">Rio Singuil</collectingCounty>
,
<date box="[366,492,862,889]" pageId="16" pageNumber="272" value="1999-08-05">
<collectingDate box="[366,492,862,889]" pageId="16" pageNumber="272" value="1999-08-05">5.viii.1999</collectingDate>
</date>
,
<collectorName box="[505,640,862,889]" pageId="16" pageNumber="272">Molineri C.</collectorName>
col. One nymph on slide (
<typeStatus box="[951,1054,863,889]" pageId="16" pageNumber="272">paratype</typeStatus>
),
<collectingCountry box="[98,270,897,923]" name="Argentina" pageId="16" pageNumber="272">ARGENTINA</collectingCountry>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2610422302" collectingDate="1998-09-15" collectorName="Dominguez E. &amp; Molineri C. &amp; Ubero &amp; Museo Miguel Lillo" country="Argentina" location="Dpto. Guachipas" municipality="Rio Grande de El Sauce" pageId="16" pageNumber="272" specimenCount="1" stateProvince="Salta" typeStatus="paratype">
<collectingRegion box="[292,349,897,923]" country="Argentina" name="Salta" pageId="16" pageNumber="272">Salta</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:074FCA78FFBEFFD7FFE1A0B4FEDD3D10:8A392DB5FFBEFFD7FEF0A1E2FDEF3BB0" box="[371,578,897,924]" country="Argentina" municipality="Rio Grande de El Sauce" name="Dpto. Guachipas" pageId="16" pageNumber="272" stateProvince="Salta">Dpto. Guachipas</location>
,
<collectingMunicipality box="[600,914,897,923]" pageId="16" pageNumber="272">Rio Grande de El Sauce</collectingMunicipality>
,
<date box="[936,1061,897,924]" pageId="16" pageNumber="272" value="1998-09-15">
<collectingDate box="[936,1061,897,924]" pageId="16" pageNumber="272" value="1998-09-15">15.ix.1998</collectingDate>
</date>
,
<collectorName box="[98,271,932,958]" pageId="16" pageNumber="272">Domínguez E.</collectorName>
,
<collectorName box="[290,430,932,958]" pageId="16" pageNumber="272">Molineri C.</collectorName>
and
<collectorName box="[499,571,932,958]" pageId="16" pageNumber="272">Ubero</collectorName>
cols, deposited at
<collectorName box="[802,1039,932,958]" pageId="16" pageNumber="272">Museo Miguel Lillo</collectorName>
–
<collectingCountry box="[98,214,966,992]" name="Argentina" pageId="16" pageNumber="272">Argentina</collectingCountry>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2610422309" country="Argentina" location="Tucuman" pageId="16" pageNumber="272" specimenCount="2" stateProvince="Tucuman">
<emphasis box="[232,628,966,992]" italics="true" pageId="16" pageNumber="272">
<typeStatus box="[232,453,966,992]" isOtherTypeStatus="true" pageId="16" pageNumber="272">Additional material</typeStatus>
not examined:
</emphasis>
<typeStatus box="[642,749,966,992]" pageId="16" pageNumber="272">Holotype</typeStatus>
(male nymph),
<collectingRegion box="[951,1062,967,993]" country="Argentina" name="Tucuman" pageId="16" pageNumber="272">Tucuman</collectingRegion>
,
<collectingCountry box="[98,270,1001,1027]" name="Argentina" pageId="16" pageNumber="272">ARGENTINA</collectingCountry>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2610422304" collectingDate="2001-06-14" collectorName="Moiineri &amp; Manzo &amp; Orce &amp; Nieto &amp; Museo Miguel Lillo" country="Argentina" elevation="860" latitude="-26.540833" location="river Medina" longLatPrecision="20" longitude="-65.02917" pageId="16" pageNumber="272" specimenCount="860">
<location LSID="urn:lsid:plazi:treatment:074FCA78FFBEFFD7FFE1A0B4FEDD3D10:8A392DB5FFBEFFD7FE94A189FE063C28" box="[279,427,1001,1027]" country="Argentina" latitude="-26.540833" longLatPrecision="20" longitude="-65.02917" name="river Medina" pageId="16" pageNumber="272">river Medina</location>
,
<geoCoordinate box="[436,569,1001,1027]" degrees="26" direction="south" minutes="32" orientation="latitude" pageId="16" pageNumber="272" precision="15" seconds="27" value="-26.540833">S26°32’27”</geoCoordinate>
.
<geoCoordinate box="[577,724,1001,1028]" degrees="65" direction="west" minutes="01" orientation="longitude" pageId="16" pageNumber="272" precision="15" seconds="45" value="-65.02917">W65°01’45”</geoCoordinate>
,
<specimenCount box="[734,813,1001,1028]" pageId="16" pageNumber="272" type="generic">
<quantity box="[734,810,1001,1028]" metricMagnitude="2" metricUnit="m" metricValue="8.6" pageId="16" pageNumber="272" unit="m" value="860.0">
<elevation box="[734,810,1001,1028]" metricMagnitude="2" metricUnit="m" metricValue="8.6" pageId="16" pageNumber="272" unit="m" value="860.0">860 m</elevation>
</quantity>
.
</specimenCount>
<date box="[818,949,1001,1027]" pageId="16" pageNumber="272" value="2001-06-14">
<collectingDate box="[818,949,1001,1027]" pageId="16" pageNumber="272" value="2001-06-14">14.vi. 2001</collectingDate>
</date>
,
<collectorName box="[959,1062,1001,1027]" pageId="16" pageNumber="272">Moiineri</collectorName>
,
<collectorName box="[98,180,1036,1062]" pageId="16" pageNumber="272">Manzo</collectorName>
,
<collectorName box="[192,248,1036,1062]" pageId="16" pageNumber="272">Orce</collectorName>
&amp;
<collectorName box="[286,351,1036,1062]" pageId="16" pageNumber="272">Nieto</collectorName>
cols, deposited at
<collectorName box="[560,784,1036,1062]" pageId="16" pageNumber="272">Museo Miguel Lillo</collectorName>
-
<collectingCountry box="[808,924,1036,1062]" name="Argentina" pageId="16" pageNumber="272">Argentina</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph blockId="16.[98,1066,724,1339]" pageId="16" pageNumber="272">
<emphasis box="[98,211,1071,1096]" italics="true" pageId="16" pageNumber="272">Diagnosis.</emphasis>
<emphasis bold="true" box="[217,365,1070,1096]" pageId="16" pageNumber="272">Male imago.</emphasis>
The characters used in the generic section can also be used for specific identification.
</paragraph>
<paragraph blockId="16.[98,1066,724,1339]" pageId="16" pageNumber="272">
<emphasis bold="true" box="[128,319,1140,1166]" pageId="16" pageNumber="272">Mature nymph.</emphasis>
1) antennae long, 1.5 × length of head capsule; 2) presence of medial decline on distal margin of labrum; 3) maxillae with palp 1.5 × length of galealacinia; 4) lingua with a small rounded apical projection (
<figureCitation box="[762,842,1209,1236]" captionStart="Figs" captionStartId="13.[106,148,1086,1108]" captionTargetBox="[106,1062,131,1061]" captionTargetId="figure@13.[106,1074,131,1061]" captionTargetPageId="13" captionText="Figs. 31–36. 31, genitalia, black arrows indicate the unistyliger inner projection (Paracloeodes sp.) Chars 142–149; 32, genitalia, black arrow indicates the spine and convex subgenital plate (Paracloeodes atroari) Chars 142–149; 33, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis sp.) Chars 142–149; 34, genitalia, black arrow indicates the concave subgenital plate (Paracloeodes sp.) Chars 142–149; 35, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis capixaba) Chars 142–149; 36, genitalia, black arrow indicates the triangular subgenital plate (Apobaetis fiuzai) Chars 142–149, adapted from Cruz et al. 2011." figureDoi="http://doi.org/10.5281/zenodo.3786802" httpUri="https://zenodo.org/record/3786802/files/figure.png" pageId="16" pageNumber="272">
Fig.
<quantity box="[813,842,1209,1236]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="16" pageNumber="272" unit="in" value="35.0">35</quantity>
</figureCitation>
in
<bibRefCitation author="Nieto" box="[884,1016,1209,1235]" firstAuthor="Nieto" pageId="16" pageNumber="272" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
); 5) glossa subcircular (Fig.
<quantity box="[370,432,1244,1270]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" pageId="16" pageNumber="272" unit="in" value="37.0">37 in</quantity>
<bibRefCitation author="Nieto" box="[442,576,1244,1270]" firstAuthor="Nieto" pageId="16" pageNumber="272" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
); 6) tarsal claws 0.36 × length of tarsi; 7) abdominal terga I and
<collectionCode box="[355,405,1279,1305]" pageId="16" pageNumber="272">VIII</collectionCode>
dark brown.
</paragraph>
<paragraph blockId="16.[98,1066,724,1339]" box="[98,368,1313,1339]" pageId="16" pageNumber="272">
<emphasis box="[98,242,1313,1339]" italics="true" pageId="16" pageNumber="272">Distribution.</emphasis>
<collectingCountry box="[248,364,1313,1339]" name="Argentina" pageId="16" pageNumber="272">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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