666 lines
125 KiB
XML
666 lines
125 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3794349" ID-GBIF-Dataset="43234c4b-40c9-4d1b-aa5f-51d222a421b2" ID-GBIF-Taxon="163963894" ID-ISSN="1876-312X" ID-Zenodo-Dep="3794349" ID-ZooBank="217D79EE-A577-4B7F-8AB0-1107C9FC4857" approvalRequired="3" approvalRequired_for_textStreams="3" checkinTime="1588686095692" checkinUser="felipe" docAuthor="Cruz, Paulo Vilela, Salles, Frederico Falcão & Hamada, Neusa" docDate="2020" docId="074FCA78FFA3FFD7FFE9A724FF063A83" docLanguage="en" docName="InsectSystEvol.51.2.256-275.pdf.imf" docOrigin="Insect Systematics & Evolution 51 (2)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2a2c15f9" docStyleName="insectsystevol.2020.journal_article" docTitle="Rhopyscelis Cruza & Sallesb & Hamada 2020, gen. n." docType="treatment" docVersion="4" lastPageId="16" lastPageNumber="272" masterDocId="FB76B200FFAEFFC7FF83A260FFAD382B" masterDocTitle="A cladistic approach for generic delimitation of Paracloeodes Day, Rivudiva Lugo-Ortiz & McCafferty, and Varipes Lugo-Ortiz & McCafferty (Ephemeroptera: Baetidae)" masterLastPageNumber="275" masterPageNumber="256" pageId="13" pageNumber="269" updateTime="1643438799729" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A cladistic approach for generic delimitation of Paracloeodes Day, Rivudiva Lugo-Ortiz & McCafferty, and Varipes Lugo-Ortiz & McCafferty (Ephemeroptera: Baetidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cruz, Paulo Vilela</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salles, Frederico Falcão</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hamada, Neusa</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Insect Systematics & Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-03-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>51</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>256</mods:start>
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<mods:end>275</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1163/1876312X-00002199</mods:identifier>
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<mods:identifier type="GBIF-Dataset">43234c4b-40c9-4d1b-aa5f-51d222a421b2</mods:identifier>
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<mods:identifier type="ISSN">1876-312X</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3786794</mods:identifier>
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<mods:identifier type="ZooBank">217D79EE-A577-4B7F-8AB0-1107C9FC4857</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3794349" ID-GBIF-Taxon="163963894" ID-Zenodo-Dep="3794349" LSID="urn:lsid:plazi:treatment:074FCA78FFA3FFD7FFE9A724FF063A83" httpUri="http://treatment.plazi.org/id/074FCA78FFA3FFD7FFE9A724FF063A83" lastPageId="16" lastPageNumber="272" pageId="13" pageNumber="269">
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<subSubSection box="[106,311,1348,1374]" pageId="13" pageNumber="269" type="nomenclature">
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<paragraph blockId="13.[106,1074,1348,1617]" box="[106,311,1348,1374]" pageId="13" pageNumber="269">
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<heading box="[106,311,1348,1374]" fontSize="11" level="2" pageId="13" pageNumber="269" reason="3">
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<taxonomicName authority="Cruza & Sallesb & Hamada, 2020" authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" box="[106,223,1348,1374]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="269" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[106,223,1348,1374]" italics="true" pageId="13" pageNumber="269">Rhopyscelis</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[230,311,1348,1374]" pageId="13" pageNumber="269">
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<taxonomicNameLabel box="[230,311,1348,1374]" pageId="13" pageNumber="269" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[106,980,1382,1409]" pageId="13" pageNumber="269" type="description">
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<paragraph blockId="13.[106,1074,1348,1617]" box="[106,980,1382,1409]" pageId="13" pageNumber="269">
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ZooBank: http://zoobank.org/
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<uuid box="[448,980,1382,1409]" pageId="13" pageNumber="269">46753411-F21E-45AE-BC10-8397138F68A4</uuid>
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</paragraph>
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||
</subSubSection>
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<subSubSection box="[106,805,1417,1444]" pageId="13" pageNumber="269" type="type_taxon">
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<paragraph blockId="13.[106,1074,1348,1617]" box="[106,805,1417,1444]" pageId="13" pageNumber="269">
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<typeStatus box="[106,161,1418,1444]" pageId="13" pageNumber="269">Type</typeStatus>
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species:
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<taxonomicName authority="Nieto 2004" authorityName="Nieto" authorityYear="2004" box="[260,555,1417,1443]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="269" phylum="Arthropoda" rank="species" species="singuil">
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<emphasis box="[260,417,1417,1443]" italics="true" pageId="13" pageNumber="269">Varipes singuil</emphasis>
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<bibRefCitation author="Nieto" box="[424,555,1417,1443]" firstAuthor="Nieto" pageId="13" pageNumber="269" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
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</taxonomicName>
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(original designation)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="269" type="etymology">
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<paragraph blockId="13.[106,1074,1348,1617]" pageId="13" pageNumber="269">
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<emphasis box="[106,223,1452,1478]" italics="true" pageId="13" pageNumber="269">Etymology.</emphasis>
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The generic name is an arbitrary combination of the Greek words
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<emphasis box="[1025,1074,1452,1478]" italics="true" pageId="13" pageNumber="269">rhop</emphasis>
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(brush) and
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<emphasis box="[247,298,1486,1512]" italics="true" pageId="13" pageNumber="269">scelis</emphasis>
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(leg), an allusion to the many long femoral setae on both species allocated to the new genus. The gender is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="269" type="discussion">
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<paragraph blockId="13.[106,1074,1348,1617]" pageId="13" pageNumber="269">
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<emphasis box="[106,250,1556,1582]" italics="true" pageId="13" pageNumber="269">Composition.</emphasis>
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<taxonomicName authority="(Nieto 2004)" authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[259,617,1556,1582]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="269" phylum="Arthropoda" rank="species" species="singuil" status="comb. n.">
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<emphasis box="[259,457,1556,1582]" italics="true" pageId="13" pageNumber="269">Rhopyscelis singuil</emphasis>
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(
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<bibRefCitation author="Nieto" box="[475,608,1556,1582]" firstAuthor="Nieto" pageId="13" pageNumber="269" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
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)
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</taxonomicName>
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<emphasis bold="true" box="[627,734,1556,1582]" pageId="13" pageNumber="269">
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<taxonomicNameLabel box="[627,734,1556,1582]" pageId="13" pageNumber="269" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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(
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<typeStatus box="[751,801,1556,1582]" pageId="13" pageNumber="269">type</typeStatus>
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species) and
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<taxonomicName authority="(Gutierrez, Dias & Salles 2013)" authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="269" phylum="Arthropoda" rank="species" species="caldensis" status="comb. n.">
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<emphasis italics="true" pageId="13" pageNumber="269">Rhopyscelis caldensis</emphasis>
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(
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<bibRefCitation author="Gutierrez, Dias & Salles" box="[215,554,1590,1617]" firstAuthor="Gutierrez" pageId="13" pageNumber="269" pagination="291 - 295" refId="ref10661" refString="Gutierrez, Y., Dias, L. G. & Salles, F. F. (2013) Paracloeodes caldensis (Ephemeroptera: Baetidae), an atypical new species from the Colombian Andes. Zootaxa 3721: 291 - 295." type="journal article" year="2013">Gutiérrez, Dias & Salles 2013</bibRefCitation>
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)
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</taxonomicName>
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<emphasis bold="true" box="[571,676,1590,1616]" pageId="13" pageNumber="269">
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<taxonomicNameLabel box="[571,676,1590,1616]" pageId="13" pageNumber="269" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="270" type="diagnosis">
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<paragraph blockId="14.[98,1066,134,750]" pageId="14" pageNumber="270">
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<emphasis box="[98,211,135,160]" italics="true" pageId="14" pageNumber="270">Diagnosis.</emphasis>
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<emphasis bold="true" box="[218,368,134,160]" pageId="14" pageNumber="270">Male imago.</emphasis>
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1) forewing hyaline (
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<figureCitation box="[616,696,134,161]" captionStart="Figs" captionStartId="8.[99,141,1129,1151]" captionTargetId="figure@9.[106,1074,131,1523]" captionTargetPageId="9" captionText="Figs. 16–30. 16, detail of forefemur concave setae on dorsal margin (Paracloeodes atroari) Char. 77; 17, detail of forefemur, black arrow indicates the FLRNDM = forefemur longitudinal row of setae near dorsal margin on anterior surface (Paracloeodes eurybranchus) Char. 80; 18, detail of forefemur, black arrow indicates the FLRNML = forefemur longitudinal row of setae at middle on anterior surface (Paracloeodes sp.) Char. 82; 19, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis singuil comb. n.) Char. 85 and 86; 20, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis singuil comb. n.) Chars 89 and 90; 21, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis caldensis comb. n.) Chars 85 and 86; 22, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis caldensis comb. n.) Chars 89 and 90; 23, detail of transversal row of setae on hind femur, (Rivudiva trichobasis) Chars 89 and 91; 24, detail of transversal row of setae on hind femur, (Varipes helenae) Chars 89 and 91; 25, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis singuil comb. n.) Chars 96–101; 26, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96–101; 27, hind claw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96– 101; 28, hind claw, black arrows indicate the regularity of row of denticles, (Rivudiva trichobasis) Chars 99 and 102; 29, foreclaw, black arrow indicates the absence of denticles on base, red arrow indicates the conical and apical denticles, (Varipes lasiobrachius) Chars 106 and 107; 30, detail of foreclaw, black arrow indicates apical denticles on claw extremely higher than the others (Gen. A.) Char. 104." figureDoi="http://doi.org/10.5281/zenodo.3786800" httpUri="https://zenodo.org/record/3786800/files/figure.png" pageId="14" pageNumber="270">
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Fig.
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</figureCitation>
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in
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<bibRefCitation author="Nieto" box="[736,868,134,161]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
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); 2) marginal intercalary veins paired (
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<figureCitation box="[351,429,169,195]" captionStart="Figs" captionStartId="8.[99,141,1129,1151]" captionTargetId="figure@9.[106,1074,131,1523]" captionTargetPageId="9" captionText="Figs. 16–30. 16, detail of forefemur concave setae on dorsal margin (Paracloeodes atroari) Char. 77; 17, detail of forefemur, black arrow indicates the FLRNDM = forefemur longitudinal row of setae near dorsal margin on anterior surface (Paracloeodes eurybranchus) Char. 80; 18, detail of forefemur, black arrow indicates the FLRNML = forefemur longitudinal row of setae at middle on anterior surface (Paracloeodes sp.) Char. 82; 19, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis singuil comb. n.) Char. 85 and 86; 20, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis singuil comb. n.) Chars 89 and 90; 21, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis caldensis comb. n.) Chars 85 and 86; 22, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis caldensis comb. n.) Chars 89 and 90; 23, detail of transversal row of setae on hind femur, (Rivudiva trichobasis) Chars 89 and 91; 24, detail of transversal row of setae on hind femur, (Varipes helenae) Chars 89 and 91; 25, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis singuil comb. n.) Chars 96–101; 26, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96–101; 27, hind claw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96– 101; 28, hind claw, black arrows indicate the regularity of row of denticles, (Rivudiva trichobasis) Chars 99 and 102; 29, foreclaw, black arrow indicates the absence of denticles on base, red arrow indicates the conical and apical denticles, (Varipes lasiobrachius) Chars 106 and 107; 30, detail of foreclaw, black arrow indicates apical denticles on claw extremely higher than the others (Gen. A.) Char. 104." figureDoi="http://doi.org/10.5281/zenodo.3786800" httpUri="https://zenodo.org/record/3786800/files/figure.png" pageId="14" pageNumber="270">
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Fig.
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in
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<bibRefCitation author="Nieto" box="[466,597,169,195]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
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); 3) hind wing absent; 4) subgenital plate without apical spine (
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<figureCitation box="[350,431,204,230]" captionStart="Figs" captionStartId="13.[106,148,1086,1108]" captionTargetBox="[106,1062,131,1061]" captionTargetId="figure@13.[106,1074,131,1061]" captionTargetPageId="13" captionText="Figs. 31–36. 31, genitalia, black arrows indicate the unistyliger inner projection (Paracloeodes sp.) Chars 142–149; 32, genitalia, black arrow indicates the spine and convex subgenital plate (Paracloeodes atroari) Chars 142–149; 33, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis sp.) Chars 142–149; 34, genitalia, black arrow indicates the concave subgenital plate (Paracloeodes sp.) Chars 142–149; 35, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis capixaba) Chars 142–149; 36, genitalia, black arrow indicates the triangular subgenital plate (Apobaetis fiuzai) Chars 142–149, adapted from Cruz et al. 2011." figureDoi="http://doi.org/10.5281/zenodo.3786802" httpUri="https://zenodo.org/record/3786802/files/figure.png" pageId="14" pageNumber="270">
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Fig.
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</figureCitation>
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in
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<bibRefCitation author="Nieto" box="[474,607,204,230]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
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); 5) first segment of forceps with inner projection (
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<figureCitation box="[229,307,238,265]" captionStart="Figs" captionStartId="13.[106,148,1086,1108]" captionTargetBox="[106,1062,131,1061]" captionTargetId="figure@13.[106,1074,131,1061]" captionTargetPageId="13" captionText="Figs. 31–36. 31, genitalia, black arrows indicate the unistyliger inner projection (Paracloeodes sp.) Chars 142–149; 32, genitalia, black arrow indicates the spine and convex subgenital plate (Paracloeodes atroari) Chars 142–149; 33, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis sp.) Chars 142–149; 34, genitalia, black arrow indicates the concave subgenital plate (Paracloeodes sp.) Chars 142–149; 35, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis capixaba) Chars 142–149; 36, genitalia, black arrow indicates the triangular subgenital plate (Apobaetis fiuzai) Chars 142–149, adapted from Cruz et al. 2011." figureDoi="http://doi.org/10.5281/zenodo.3786802" httpUri="https://zenodo.org/record/3786802/files/figure.png" pageId="14" pageNumber="270">
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Fig.
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in
|
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<bibRefCitation author="Nieto" box="[343,473,238,265]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
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); 6) last segment of forceps rod-like (
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<figureCitation box="[888,966,238,265]" captionStart="Figs" captionStartId="13.[106,148,1086,1108]" captionTargetBox="[106,1062,131,1061]" captionTargetId="figure@13.[106,1074,131,1061]" captionTargetPageId="13" captionText="Figs. 31–36. 31, genitalia, black arrows indicate the unistyliger inner projection (Paracloeodes sp.) Chars 142–149; 32, genitalia, black arrow indicates the spine and convex subgenital plate (Paracloeodes atroari) Chars 142–149; 33, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis sp.) Chars 142–149; 34, genitalia, black arrow indicates the concave subgenital plate (Paracloeodes sp.) Chars 142–149; 35, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis capixaba) Chars 142–149; 36, genitalia, black arrow indicates the triangular subgenital plate (Apobaetis fiuzai) Chars 142–149, adapted from Cruz et al. 2011." figureDoi="http://doi.org/10.5281/zenodo.3786802" httpUri="https://zenodo.org/record/3786802/files/figure.png" pageId="14" pageNumber="270">
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Fig.
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</figureCitation>
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in
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<bibRefCitation author="Nieto" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
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).
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</paragraph>
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<paragraph blockId="14.[98,1066,134,750]" pageId="14" pageNumber="270">
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<emphasis bold="true" box="[128,321,308,334]" pageId="14" pageNumber="270">Mature nymph.</emphasis>
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1) prostheca of right mandible slender and bifid (
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<figureCitation box="[943,1029,308,334]" captionStart="Figs" captionStartId="13.[106,148,1086,1108]" captionTargetBox="[106,1062,131,1061]" captionTargetId="figure@13.[106,1074,131,1061]" captionTargetPageId="13" captionText="Figs. 31–36. 31, genitalia, black arrows indicate the unistyliger inner projection (Paracloeodes sp.) Chars 142–149; 32, genitalia, black arrow indicates the spine and convex subgenital plate (Paracloeodes atroari) Chars 142–149; 33, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis sp.) Chars 142–149; 34, genitalia, black arrow indicates the concave subgenital plate (Paracloeodes sp.) Chars 142–149; 35, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis capixaba) Chars 142–149; 36, genitalia, black arrow indicates the triangular subgenital plate (Apobaetis fiuzai) Chars 142–149, adapted from Cruz et al. 2011." figureDoi="http://doi.org/10.5281/zenodo.3786802" httpUri="https://zenodo.org/record/3786802/files/figure.png" pageId="14" pageNumber="270">
|
||
Fig.
|
||
<quantity box="[1000,1029,308,334]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="14" pageNumber="270" unit="in" value="34.0">34</quantity>
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</figureCitation>
|
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in
|
||
<bibRefCitation author="Nieto" box="[98,230,342,369]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
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); 2) mandibular incisors partially fused (
|
||
<figureCitation box="[698,778,342,369]" captionStart="Figs" captionStartId="13.[106,148,1086,1108]" captionTargetBox="[106,1062,131,1061]" captionTargetId="figure@13.[106,1074,131,1061]" captionTargetPageId="13" captionText="Figs. 31–36. 31, genitalia, black arrows indicate the unistyliger inner projection (Paracloeodes sp.) Chars 142–149; 32, genitalia, black arrow indicates the spine and convex subgenital plate (Paracloeodes atroari) Chars 142–149; 33, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis sp.) Chars 142–149; 34, genitalia, black arrow indicates the concave subgenital plate (Paracloeodes sp.) Chars 142–149; 35, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis capixaba) Chars 142–149; 36, genitalia, black arrow indicates the triangular subgenital plate (Apobaetis fiuzai) Chars 142–149, adapted from Cruz et al. 2011." figureDoi="http://doi.org/10.5281/zenodo.3786802" httpUri="https://zenodo.org/record/3786802/files/figure.png" pageId="14" pageNumber="270">
|
||
Fig.
|
||
<quantity box="[749,778,343,369]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="14" pageNumber="270" unit="in" value="34.0">34</quantity>
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</figureCitation>
|
||
in
|
||
<bibRefCitation author="Nieto" box="[817,949,342,369]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
||
); 3) inner set of incisors at right angle to the outer set; 4) distomedial projection of labial palp segment II rounded (
|
||
<figureCitation box="[344,411,412,438]" captionStart="Figs" captionStartId="6.[98,140,1100,1122]" captionTargetId="figure@7.[106,1074,131,1525]" captionTargetPageId="7" captionText="Figs. 2–15. 2, elevation of frons with two keels (Paracloeodes charrua), Chars 1 and 2, black arrow indicates frons; 3, elevation of frons with one keel (Paracloeodes atroari) Chars 1 and 2, black arrow indicates frons; 4, organization of setae on labrum (Callibaetis capixaba, left = dorsal, right = ventral) Chars 5, 6, 8, 15, 16, 19, 20, LAS = lateral arc of setae, DAS = distal arc of setae, DMA = distomedial arc of setae, SDA = subdistal arc of setae on ventral surface, MDA = mediodorsal arc of setae, RSVL = ventral row of spine-like setae on lateral margin of labrum, TSVALM = tuft of setae ventrally on anterolateral margin of labrum; 5, organization of setae on labrum (Apobaetis sp.) Chars 10, 14, DDA = distodorsal arc of setae, SDMA = sub-distomedial arc of setae on dorsal surface; 6, organization of setae on labrum (Waltzoyphius sp.) Char. 12, SDLA = sub-distolateral arc of setae on dorsal surface; 7, detail of deep cleft incisors of right mandible (Callibaetis sp.) Char. 30, black arrow indicates the incisors; 8, detail of partially fused incisors of right mandible (Paracloeodes sp.) Char. 30, black arrow indicates the incisors; 9, detail of a small lobe on apex of last segment of maxillary palp (Rivudiva sp.) Char. 43, back arrow indicates the lobe; 10, detail of basal projection of first dentiseta coupling with canine on maxilla (Varipes lasiobranchius) Char. 47; 11, detail of row of setae on outer margin of paraglossa (ROMP) (Callibaetis sp.) Char. 59; 12, detail of labial palp with red lines showing axes for angle measure (Apobaetis sp.) Char. 65; 13, detail of labial palp segment II and III, black arrow indicates the medial concavity on segment III (Callibaetis calloventer) Char. 68; 14, detail of trochanter inserted on ventral margin of femur, red line shows the connection (Varipes lasiobrachius) Char. 73; 15, trochanter inserted along entire base of femur, red line shows the connection (Rhopyscelis caldensis comb. n.) Char. 73." figureDoi="http://doi.org/10.5281/zenodo.3786798" httpUri="https://zenodo.org/record/3786798/files/figure.png" pageId="14" pageNumber="270">
|
||
Fig.
|
||
<quantity box="[396,411,412,438]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="14" pageNumber="270" unit="in" value="7.0">7</quantity>
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</figureCitation>
|
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in
|
||
<bibRefCitation author="Gutierrez" box="[452,694,412,438]" etAl="et al." firstAuthor="Gutierrez" pageId="14" pageNumber="270" pagination="291 - 295" refId="ref10661" refString="Gutierrez, Y., Dias, L. G. & Salles, F. F. (2013) Paracloeodes caldensis (Ephemeroptera: Baetidae), an atypical new species from the Colombian Andes. Zootaxa 3721: 291 - 295." type="journal article" year="2013">Gutiérrez et al. 2013</bibRefCitation>
|
||
; Fig.
|
||
<quantity box="[760,822,412,438]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" pageId="14" pageNumber="270" unit="in" value="37.0">37 in</quantity>
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<bibRefCitation author="Nieto" box="[831,964,412,438]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
||
); 5) segment III of labial palp conical (apex broadly pointed) (
|
||
<figureCitation box="[720,785,446,473]" captionStart="Figs" captionStartId="6.[98,140,1100,1122]" captionTargetId="figure@7.[106,1074,131,1525]" captionTargetPageId="7" captionText="Figs. 2–15. 2, elevation of frons with two keels (Paracloeodes charrua), Chars 1 and 2, black arrow indicates frons; 3, elevation of frons with one keel (Paracloeodes atroari) Chars 1 and 2, black arrow indicates frons; 4, organization of setae on labrum (Callibaetis capixaba, left = dorsal, right = ventral) Chars 5, 6, 8, 15, 16, 19, 20, LAS = lateral arc of setae, DAS = distal arc of setae, DMA = distomedial arc of setae, SDA = subdistal arc of setae on ventral surface, MDA = mediodorsal arc of setae, RSVL = ventral row of spine-like setae on lateral margin of labrum, TSVALM = tuft of setae ventrally on anterolateral margin of labrum; 5, organization of setae on labrum (Apobaetis sp.) Chars 10, 14, DDA = distodorsal arc of setae, SDMA = sub-distomedial arc of setae on dorsal surface; 6, organization of setae on labrum (Waltzoyphius sp.) Char. 12, SDLA = sub-distolateral arc of setae on dorsal surface; 7, detail of deep cleft incisors of right mandible (Callibaetis sp.) Char. 30, black arrow indicates the incisors; 8, detail of partially fused incisors of right mandible (Paracloeodes sp.) Char. 30, black arrow indicates the incisors; 9, detail of a small lobe on apex of last segment of maxillary palp (Rivudiva sp.) Char. 43, back arrow indicates the lobe; 10, detail of basal projection of first dentiseta coupling with canine on maxilla (Varipes lasiobranchius) Char. 47; 11, detail of row of setae on outer margin of paraglossa (ROMP) (Callibaetis sp.) Char. 59; 12, detail of labial palp with red lines showing axes for angle measure (Apobaetis sp.) Char. 65; 13, detail of labial palp segment II and III, black arrow indicates the medial concavity on segment III (Callibaetis calloventer) Char. 68; 14, detail of trochanter inserted on ventral margin of femur, red line shows the connection (Varipes lasiobrachius) Char. 73; 15, trochanter inserted along entire base of femur, red line shows the connection (Rhopyscelis caldensis comb. n.) Char. 73." figureDoi="http://doi.org/10.5281/zenodo.3786798" httpUri="https://zenodo.org/record/3786798/files/figure.png" pageId="14" pageNumber="270">
|
||
Fig.
|
||
<quantity box="[770,785,447,473]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="14" pageNumber="270" unit="in" value="7.0">7</quantity>
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</figureCitation>
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in
|
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<bibRefCitation author="Gutierrez" box="[823,1061,446,473]" etAl="et al." firstAuthor="Gutierrez" pageId="14" pageNumber="270" pagination="291 - 295" refId="ref10661" refString="Gutierrez, Y., Dias, L. G. & Salles, F. F. (2013) Paracloeodes caldensis (Ephemeroptera: Baetidae), an atypical new species from the Colombian Andes. Zootaxa 3721: 291 - 295." type="journal article" year="2013">Gutiérrez et al. 2013</bibRefCitation>
|
||
; Fig.
|
||
<quantity box="[152,214,481,507]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" pageId="14" pageNumber="270" unit="in" value="37.0">37 in</quantity>
|
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<bibRefCitation author="Nieto" box="[225,359,481,507]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
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); 6) trochanter inserted along entire base of femur (
|
||
<figureCitation box="[976,1060,481,508]" captionStart="Figs" captionStartId="6.[98,140,1100,1122]" captionTargetId="figure@7.[106,1074,131,1525]" captionTargetPageId="7" captionText="Figs. 2–15. 2, elevation of frons with two keels (Paracloeodes charrua), Chars 1 and 2, black arrow indicates frons; 3, elevation of frons with one keel (Paracloeodes atroari) Chars 1 and 2, black arrow indicates frons; 4, organization of setae on labrum (Callibaetis capixaba, left = dorsal, right = ventral) Chars 5, 6, 8, 15, 16, 19, 20, LAS = lateral arc of setae, DAS = distal arc of setae, DMA = distomedial arc of setae, SDA = subdistal arc of setae on ventral surface, MDA = mediodorsal arc of setae, RSVL = ventral row of spine-like setae on lateral margin of labrum, TSVALM = tuft of setae ventrally on anterolateral margin of labrum; 5, organization of setae on labrum (Apobaetis sp.) Chars 10, 14, DDA = distodorsal arc of setae, SDMA = sub-distomedial arc of setae on dorsal surface; 6, organization of setae on labrum (Waltzoyphius sp.) Char. 12, SDLA = sub-distolateral arc of setae on dorsal surface; 7, detail of deep cleft incisors of right mandible (Callibaetis sp.) Char. 30, black arrow indicates the incisors; 8, detail of partially fused incisors of right mandible (Paracloeodes sp.) Char. 30, black arrow indicates the incisors; 9, detail of a small lobe on apex of last segment of maxillary palp (Rivudiva sp.) Char. 43, back arrow indicates the lobe; 10, detail of basal projection of first dentiseta coupling with canine on maxilla (Varipes lasiobranchius) Char. 47; 11, detail of row of setae on outer margin of paraglossa (ROMP) (Callibaetis sp.) Char. 59; 12, detail of labial palp with red lines showing axes for angle measure (Apobaetis sp.) Char. 65; 13, detail of labial palp segment II and III, black arrow indicates the medial concavity on segment III (Callibaetis calloventer) Char. 68; 14, detail of trochanter inserted on ventral margin of femur, red line shows the connection (Varipes lasiobrachius) Char. 73; 15, trochanter inserted along entire base of femur, red line shows the connection (Rhopyscelis caldensis comb. n.) Char. 73." figureDoi="http://doi.org/10.5281/zenodo.3786798" httpUri="https://zenodo.org/record/3786798/files/figure.png" pageId="14" pageNumber="270">Fig. 15</figureCitation>
|
||
;
|
||
<figureCitation box="[98,179,516,542]" captionStart="Figs" captionStartId="6.[98,140,1100,1122]" captionTargetId="figure@7.[106,1074,131,1525]" captionTargetPageId="7" captionText="Figs. 2–15. 2, elevation of frons with two keels (Paracloeodes charrua), Chars 1 and 2, black arrow indicates frons; 3, elevation of frons with one keel (Paracloeodes atroari) Chars 1 and 2, black arrow indicates frons; 4, organization of setae on labrum (Callibaetis capixaba, left = dorsal, right = ventral) Chars 5, 6, 8, 15, 16, 19, 20, LAS = lateral arc of setae, DAS = distal arc of setae, DMA = distomedial arc of setae, SDA = subdistal arc of setae on ventral surface, MDA = mediodorsal arc of setae, RSVL = ventral row of spine-like setae on lateral margin of labrum, TSVALM = tuft of setae ventrally on anterolateral margin of labrum; 5, organization of setae on labrum (Apobaetis sp.) Chars 10, 14, DDA = distodorsal arc of setae, SDMA = sub-distomedial arc of setae on dorsal surface; 6, organization of setae on labrum (Waltzoyphius sp.) Char. 12, SDLA = sub-distolateral arc of setae on dorsal surface; 7, detail of deep cleft incisors of right mandible (Callibaetis sp.) Char. 30, black arrow indicates the incisors; 8, detail of partially fused incisors of right mandible (Paracloeodes sp.) Char. 30, black arrow indicates the incisors; 9, detail of a small lobe on apex of last segment of maxillary palp (Rivudiva sp.) Char. 43, back arrow indicates the lobe; 10, detail of basal projection of first dentiseta coupling with canine on maxilla (Varipes lasiobranchius) Char. 47; 11, detail of row of setae on outer margin of paraglossa (ROMP) (Callibaetis sp.) Char. 59; 12, detail of labial palp with red lines showing axes for angle measure (Apobaetis sp.) Char. 65; 13, detail of labial palp segment II and III, black arrow indicates the medial concavity on segment III (Callibaetis calloventer) Char. 68; 14, detail of trochanter inserted on ventral margin of femur, red line shows the connection (Varipes lasiobrachius) Char. 73; 15, trochanter inserted along entire base of femur, red line shows the connection (Rhopyscelis caldensis comb. n.) Char. 73." figureDoi="http://doi.org/10.5281/zenodo.3786798" httpUri="https://zenodo.org/record/3786798/files/figure.png" pageId="14" pageNumber="270">
|
||
Fig.
|
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</figureCitation>
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in
|
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<bibRefCitation author="Gutierrez" box="[221,463,516,542]" etAl="et al." firstAuthor="Gutierrez" pageId="14" pageNumber="270" pagination="291 - 295" refId="ref10661" refString="Gutierrez, Y., Dias, L. G. & Salles, F. F. (2013) Paracloeodes caldensis (Ephemeroptera: Baetidae), an atypical new species from the Colombian Andes. Zootaxa 3721: 291 - 295." type="journal article" year="2013">Gutiérrez et al. 2013</bibRefCitation>
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; Fig.
|
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<quantity box="[529,591,516,542]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" pageId="14" pageNumber="270" unit="in" value="38.0">38 in</quantity>
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<bibRefCitation author="Nieto" box="[600,733,516,542]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
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); 7) many long thin femoral setae (
|
||
<figureCitation box="[167,299,550,577]" captionStart="Figs" captionStartId="8.[99,141,1129,1151]" captionTargetId="figure@9.[106,1074,131,1523]" captionTargetPageId="9" captionText="Figs. 16–30. 16, detail of forefemur concave setae on dorsal margin (Paracloeodes atroari) Char. 77; 17, detail of forefemur, black arrow indicates the FLRNDM = forefemur longitudinal row of setae near dorsal margin on anterior surface (Paracloeodes eurybranchus) Char. 80; 18, detail of forefemur, black arrow indicates the FLRNML = forefemur longitudinal row of setae at middle on anterior surface (Paracloeodes sp.) Char. 82; 19, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis singuil comb. n.) Char. 85 and 86; 20, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis singuil comb. n.) Chars 89 and 90; 21, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis caldensis comb. n.) Chars 85 and 86; 22, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis caldensis comb. n.) Chars 89 and 90; 23, detail of transversal row of setae on hind femur, (Rivudiva trichobasis) Chars 89 and 91; 24, detail of transversal row of setae on hind femur, (Varipes helenae) Chars 89 and 91; 25, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis singuil comb. n.) Chars 96–101; 26, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96–101; 27, hind claw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96– 101; 28, hind claw, black arrows indicate the regularity of row of denticles, (Rivudiva trichobasis) Chars 99 and 102; 29, foreclaw, black arrow indicates the absence of denticles on base, red arrow indicates the conical and apical denticles, (Varipes lasiobrachius) Chars 106 and 107; 30, detail of foreclaw, black arrow indicates apical denticles on claw extremely higher than the others (Gen. A.) Char. 104." figureDoi="http://doi.org/10.5281/zenodo.3786800" httpUri="https://zenodo.org/record/3786800/files/figure.png" pageId="14" pageNumber="270">Figs. 19–22</figureCitation>
|
||
); 8) vestigial transverse row of long setae on proximal third of forefemur (
|
||
<figureCitation box="[164,340,585,611]" captionStart="Figs" captionStartId="8.[99,141,1129,1151]" captionTargetId="figure@9.[106,1074,131,1523]" captionTargetPageId="9" captionText="Figs. 16–30. 16, detail of forefemur concave setae on dorsal margin (Paracloeodes atroari) Char. 77; 17, detail of forefemur, black arrow indicates the FLRNDM = forefemur longitudinal row of setae near dorsal margin on anterior surface (Paracloeodes eurybranchus) Char. 80; 18, detail of forefemur, black arrow indicates the FLRNML = forefemur longitudinal row of setae at middle on anterior surface (Paracloeodes sp.) Char. 82; 19, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis singuil comb. n.) Char. 85 and 86; 20, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis singuil comb. n.) Chars 89 and 90; 21, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis caldensis comb. n.) Chars 85 and 86; 22, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis caldensis comb. n.) Chars 89 and 90; 23, detail of transversal row of setae on hind femur, (Rivudiva trichobasis) Chars 89 and 91; 24, detail of transversal row of setae on hind femur, (Varipes helenae) Chars 89 and 91; 25, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis singuil comb. n.) Chars 96–101; 26, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96–101; 27, hind claw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96– 101; 28, hind claw, black arrows indicate the regularity of row of denticles, (Rivudiva trichobasis) Chars 99 and 102; 29, foreclaw, black arrow indicates the absence of denticles on base, red arrow indicates the conical and apical denticles, (Varipes lasiobrachius) Chars 106 and 107; 30, detail of foreclaw, black arrow indicates apical denticles on claw extremely higher than the others (Gen. A.) Char. 104." figureDoi="http://doi.org/10.5281/zenodo.3786800" httpUri="https://zenodo.org/record/3786800/files/figure.png" pageId="14" pageNumber="270">Figs. 19 and 20</figureCitation>
|
||
); 9) mid and hind femora with transversal row of long setae on proximal third (
|
||
<figureCitation box="[278,451,620,646]" captionStart="Figs" captionStartId="8.[99,141,1129,1151]" captionTargetId="figure@9.[106,1074,131,1523]" captionTargetPageId="9" captionText="Figs. 16–30. 16, detail of forefemur concave setae on dorsal margin (Paracloeodes atroari) Char. 77; 17, detail of forefemur, black arrow indicates the FLRNDM = forefemur longitudinal row of setae near dorsal margin on anterior surface (Paracloeodes eurybranchus) Char. 80; 18, detail of forefemur, black arrow indicates the FLRNML = forefemur longitudinal row of setae at middle on anterior surface (Paracloeodes sp.) Char. 82; 19, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis singuil comb. n.) Char. 85 and 86; 20, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis singuil comb. n.) Chars 89 and 90; 21, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis caldensis comb. n.) Chars 85 and 86; 22, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis caldensis comb. n.) Chars 89 and 90; 23, detail of transversal row of setae on hind femur, (Rivudiva trichobasis) Chars 89 and 91; 24, detail of transversal row of setae on hind femur, (Varipes helenae) Chars 89 and 91; 25, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis singuil comb. n.) Chars 96–101; 26, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96–101; 27, hind claw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96– 101; 28, hind claw, black arrows indicate the regularity of row of denticles, (Rivudiva trichobasis) Chars 99 and 102; 29, foreclaw, black arrow indicates the absence of denticles on base, red arrow indicates the conical and apical denticles, (Varipes lasiobrachius) Chars 106 and 107; 30, detail of foreclaw, black arrow indicates apical denticles on claw extremely higher than the others (Gen. A.) Char. 104." figureDoi="http://doi.org/10.5281/zenodo.3786800" httpUri="https://zenodo.org/record/3786800/files/figure.png" pageId="14" pageNumber="270">Figs. 20 and 22</figureCitation>
|
||
); 10) sockets of transverse row of setae on the mid and hind femora not touching each other (
|
||
<figureCitation box="[532,620,654,681]" captionStart="Figs" captionStartId="8.[99,141,1129,1151]" captionTargetId="figure@9.[106,1074,131,1523]" captionTargetPageId="9" captionText="Figs. 16–30. 16, detail of forefemur concave setae on dorsal margin (Paracloeodes atroari) Char. 77; 17, detail of forefemur, black arrow indicates the FLRNDM = forefemur longitudinal row of setae near dorsal margin on anterior surface (Paracloeodes eurybranchus) Char. 80; 18, detail of forefemur, black arrow indicates the FLRNML = forefemur longitudinal row of setae at middle on anterior surface (Paracloeodes sp.) Char. 82; 19, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis singuil comb. n.) Char. 85 and 86; 20, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis singuil comb. n.) Chars 89 and 90; 21, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis caldensis comb. n.) Chars 85 and 86; 22, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis caldensis comb. n.) Chars 89 and 90; 23, detail of transversal row of setae on hind femur, (Rivudiva trichobasis) Chars 89 and 91; 24, detail of transversal row of setae on hind femur, (Varipes helenae) Chars 89 and 91; 25, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis singuil comb. n.) Chars 96–101; 26, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96–101; 27, hind claw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96– 101; 28, hind claw, black arrows indicate the regularity of row of denticles, (Rivudiva trichobasis) Chars 99 and 102; 29, foreclaw, black arrow indicates the absence of denticles on base, red arrow indicates the conical and apical denticles, (Varipes lasiobrachius) Chars 106 and 107; 30, detail of foreclaw, black arrow indicates apical denticles on claw extremely higher than the others (Gen. A.) Char. 104." figureDoi="http://doi.org/10.5281/zenodo.3786800" httpUri="https://zenodo.org/record/3786800/files/figure.png" pageId="14" pageNumber="270">Figs. 23</figureCitation>
|
||
); 11) two rows of denticles on foreclaw apically directed (
|
||
<figureCitation box="[299,432,689,716]" captionStart="Figs" captionStartId="8.[99,141,1129,1151]" captionTargetId="figure@9.[106,1074,131,1523]" captionTargetPageId="9" captionText="Figs. 16–30. 16, detail of forefemur concave setae on dorsal margin (Paracloeodes atroari) Char. 77; 17, detail of forefemur, black arrow indicates the FLRNDM = forefemur longitudinal row of setae near dorsal margin on anterior surface (Paracloeodes eurybranchus) Char. 80; 18, detail of forefemur, black arrow indicates the FLRNML = forefemur longitudinal row of setae at middle on anterior surface (Paracloeodes sp.) Char. 82; 19, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis singuil comb. n.) Char. 85 and 86; 20, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis singuil comb. n.) Chars 89 and 90; 21, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis caldensis comb. n.) Chars 85 and 86; 22, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis caldensis comb. n.) Chars 89 and 90; 23, detail of transversal row of setae on hind femur, (Rivudiva trichobasis) Chars 89 and 91; 24, detail of transversal row of setae on hind femur, (Varipes helenae) Chars 89 and 91; 25, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis singuil comb. n.) Chars 96–101; 26, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96–101; 27, hind claw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96– 101; 28, hind claw, black arrows indicate the regularity of row of denticles, (Rivudiva trichobasis) Chars 99 and 102; 29, foreclaw, black arrow indicates the absence of denticles on base, red arrow indicates the conical and apical denticles, (Varipes lasiobrachius) Chars 106 and 107; 30, detail of foreclaw, black arrow indicates apical denticles on claw extremely higher than the others (Gen. A.) Char. 104." figureDoi="http://doi.org/10.5281/zenodo.3786800" httpUri="https://zenodo.org/record/3786800/files/figure.png" pageId="14" pageNumber="270">Figs. 25–27</figureCitation>
|
||
); 12) hind wing pads absent; 13) gill I-VII not racquetshaped; 14) abdominal terga surface creased, without scale-like setae/socket.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="15" lastPageNumber="271" pageId="14" pageNumber="270" type="description">
|
||
<paragraph blockId="14.[98,222,794,819]" box="[98,222,794,819]" pageId="14" pageNumber="270">
|
||
<emphasis box="[98,222,794,819]" italics="true" pageId="14" pageNumber="270">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="14.[98,1066,845,1634]" pageId="14" pageNumber="270">
|
||
<emphasis bold="true" box="[98,328,845,871]" pageId="14" pageNumber="270">Male imago. Head.</emphasis>
|
||
Dorsal portion of compound eyes oval (
|
||
<figureCitation box="[796,876,845,871]" captionStart="Figs" captionStartId="8.[99,141,1129,1151]" captionTargetId="figure@9.[106,1074,131,1523]" captionTargetPageId="9" captionText="Figs. 16–30. 16, detail of forefemur concave setae on dorsal margin (Paracloeodes atroari) Char. 77; 17, detail of forefemur, black arrow indicates the FLRNDM = forefemur longitudinal row of setae near dorsal margin on anterior surface (Paracloeodes eurybranchus) Char. 80; 18, detail of forefemur, black arrow indicates the FLRNML = forefemur longitudinal row of setae at middle on anterior surface (Paracloeodes sp.) Char. 82; 19, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis singuil comb. n.) Char. 85 and 86; 20, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis singuil comb. n.) Chars 89 and 90; 21, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis caldensis comb. n.) Chars 85 and 86; 22, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis caldensis comb. n.) Chars 89 and 90; 23, detail of transversal row of setae on hind femur, (Rivudiva trichobasis) Chars 89 and 91; 24, detail of transversal row of setae on hind femur, (Varipes helenae) Chars 89 and 91; 25, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis singuil comb. n.) Chars 96–101; 26, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96–101; 27, hind claw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96– 101; 28, hind claw, black arrows indicate the regularity of row of denticles, (Rivudiva trichobasis) Chars 99 and 102; 29, foreclaw, black arrow indicates the absence of denticles on base, red arrow indicates the conical and apical denticles, (Varipes lasiobrachius) Chars 106 and 107; 30, detail of foreclaw, black arrow indicates apical denticles on claw extremely higher than the others (Gen. A.) Char. 104." figureDoi="http://doi.org/10.5281/zenodo.3786800" httpUri="https://zenodo.org/record/3786800/files/figure.png" pageId="14" pageNumber="270">
|
||
Fig.
|
||
<quantity box="[847,876,845,871]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" pageId="14" pageNumber="270" unit="in" value="29.0">29</quantity>
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</figureCitation>
|
||
in
|
||
<bibRefCitation author="Nieto" box="[917,1049,845,871]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
||
); stalk in lateral view of dorsal portion of compound eye divergent.
|
||
<emphasis bold="true" box="[859,952,880,906]" pageId="14" pageNumber="270">Thorax.</emphasis>
|
||
Forewing hyaline with paired marginal intercalary veins (
|
||
<figureCitation box="[632,712,914,941]" captionStart="Figs" captionStartId="8.[99,141,1129,1151]" captionTargetId="figure@9.[106,1074,131,1523]" captionTargetPageId="9" captionText="Figs. 16–30. 16, detail of forefemur concave setae on dorsal margin (Paracloeodes atroari) Char. 77; 17, detail of forefemur, black arrow indicates the FLRNDM = forefemur longitudinal row of setae near dorsal margin on anterior surface (Paracloeodes eurybranchus) Char. 80; 18, detail of forefemur, black arrow indicates the FLRNML = forefemur longitudinal row of setae at middle on anterior surface (Paracloeodes sp.) Char. 82; 19, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis singuil comb. n.) Char. 85 and 86; 20, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis singuil comb. n.) Chars 89 and 90; 21, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis caldensis comb. n.) Chars 85 and 86; 22, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis caldensis comb. n.) Chars 89 and 90; 23, detail of transversal row of setae on hind femur, (Rivudiva trichobasis) Chars 89 and 91; 24, detail of transversal row of setae on hind femur, (Varipes helenae) Chars 89 and 91; 25, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis singuil comb. n.) Chars 96–101; 26, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96–101; 27, hind claw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96– 101; 28, hind claw, black arrows indicate the regularity of row of denticles, (Rivudiva trichobasis) Chars 99 and 102; 29, foreclaw, black arrow indicates the absence of denticles on base, red arrow indicates the conical and apical denticles, (Varipes lasiobrachius) Chars 106 and 107; 30, detail of foreclaw, black arrow indicates apical denticles on claw extremely higher than the others (Gen. A.) Char. 104." figureDoi="http://doi.org/10.5281/zenodo.3786800" httpUri="https://zenodo.org/record/3786800/files/figure.png" pageId="14" pageNumber="270">
|
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Fig.
|
||
<quantity box="[683,712,915,941]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="14" pageNumber="270" unit="in" value="30.0">30</quantity>
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</figureCitation>
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in
|
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<bibRefCitation author="Nieto" box="[750,882,914,941]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
||
); length of each intercalary vein 0.5 × distance between adjacent longitudinal veins (
|
||
<figureCitation box="[879,960,949,975]" captionStart="Figs" captionStartId="8.[99,141,1129,1151]" captionTargetId="figure@9.[106,1074,131,1523]" captionTargetPageId="9" captionText="Figs. 16–30. 16, detail of forefemur concave setae on dorsal margin (Paracloeodes atroari) Char. 77; 17, detail of forefemur, black arrow indicates the FLRNDM = forefemur longitudinal row of setae near dorsal margin on anterior surface (Paracloeodes eurybranchus) Char. 80; 18, detail of forefemur, black arrow indicates the FLRNML = forefemur longitudinal row of setae at middle on anterior surface (Paracloeodes sp.) Char. 82; 19, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis singuil comb. n.) Char. 85 and 86; 20, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis singuil comb. n.) Chars 89 and 90; 21, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis caldensis comb. n.) Chars 85 and 86; 22, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis caldensis comb. n.) Chars 89 and 90; 23, detail of transversal row of setae on hind femur, (Rivudiva trichobasis) Chars 89 and 91; 24, detail of transversal row of setae on hind femur, (Varipes helenae) Chars 89 and 91; 25, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis singuil comb. n.) Chars 96–101; 26, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96–101; 27, hind claw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96– 101; 28, hind claw, black arrows indicate the regularity of row of denticles, (Rivudiva trichobasis) Chars 99 and 102; 29, foreclaw, black arrow indicates the absence of denticles on base, red arrow indicates the conical and apical denticles, (Varipes lasiobrachius) Chars 106 and 107; 30, detail of foreclaw, black arrow indicates apical denticles on claw extremely higher than the others (Gen. A.) Char. 104." figureDoi="http://doi.org/10.5281/zenodo.3786800" httpUri="https://zenodo.org/record/3786800/files/figure.png" pageId="14" pageNumber="270">
|
||
Fig.
|
||
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</figureCitation>
|
||
in
|
||
<bibRefCitation author="Nieto" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
||
). Hind wing absent.
|
||
<emphasis bold="true" box="[401,525,983,1009]" pageId="14" pageNumber="270">Abdomen.</emphasis>
|
||
Genitalia (
|
||
<figureCitation box="[654,733,984,1010]" captionStart="Figs" captionStartId="13.[106,148,1086,1108]" captionTargetBox="[106,1062,131,1061]" captionTargetId="figure@13.[106,1074,131,1061]" captionTargetPageId="13" captionText="Figs. 31–36. 31, genitalia, black arrows indicate the unistyliger inner projection (Paracloeodes sp.) Chars 142–149; 32, genitalia, black arrow indicates the spine and convex subgenital plate (Paracloeodes atroari) Chars 142–149; 33, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis sp.) Chars 142–149; 34, genitalia, black arrow indicates the concave subgenital plate (Paracloeodes sp.) Chars 142–149; 35, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis capixaba) Chars 142–149; 36, genitalia, black arrow indicates the triangular subgenital plate (Apobaetis fiuzai) Chars 142–149, adapted from Cruz et al. 2011." figureDoi="http://doi.org/10.5281/zenodo.3786802" httpUri="https://zenodo.org/record/3786802/files/figure.png" pageId="14" pageNumber="270">
|
||
Fig.
|
||
<quantity box="[704,733,984,1010]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" pageId="14" pageNumber="270" unit="in" value="31.0">31</quantity>
|
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</figureCitation>
|
||
in
|
||
<bibRefCitation author="Nieto" box="[772,904,984,1010]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
||
). Forceps segment II narrow medially; segment III rod-like.
|
||
</paragraph>
|
||
<paragraph blockId="14.[98,1066,845,1634]" lastBlockId="15.[106,1074,134,403]" lastPageId="15" lastPageNumber="271" pageId="14" pageNumber="270">
|
||
<emphasis bold="true" box="[128,402,1053,1079]" pageId="14" pageNumber="270">Mature nymph. Head.</emphasis>
|
||
Frons with two longitudinal keels. Antenna with spines and simple setae. Labrum (
|
||
<figureCitation box="[403,482,1088,1114]" captionStart="Figs" captionStartId="13.[106,148,1086,1108]" captionTargetBox="[106,1062,131,1061]" captionTargetId="figure@13.[106,1074,131,1061]" captionTargetPageId="13" captionText="Figs. 31–36. 31, genitalia, black arrows indicate the unistyliger inner projection (Paracloeodes sp.) Chars 142–149; 32, genitalia, black arrow indicates the spine and convex subgenital plate (Paracloeodes atroari) Chars 142–149; 33, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis sp.) Chars 142–149; 34, genitalia, black arrow indicates the concave subgenital plate (Paracloeodes sp.) Chars 142–149; 35, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis capixaba) Chars 142–149; 36, genitalia, black arrow indicates the triangular subgenital plate (Apobaetis fiuzai) Chars 142–149, adapted from Cruz et al. 2011." figureDoi="http://doi.org/10.5281/zenodo.3786802" httpUri="https://zenodo.org/record/3786802/files/figure.png" pageId="14" pageNumber="270">
|
||
Fig.
|
||
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</figureCitation>
|
||
in
|
||
<bibRefCitation author="Nieto" box="[519,650,1088,1114]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
||
;
|
||
<figureCitation box="[664,729,1088,1114]" captionStart="Figs" captionStartId="6.[98,140,1100,1122]" captionTargetId="figure@7.[106,1074,131,1525]" captionTargetPageId="7" captionText="Figs. 2–15. 2, elevation of frons with two keels (Paracloeodes charrua), Chars 1 and 2, black arrow indicates frons; 3, elevation of frons with one keel (Paracloeodes atroari) Chars 1 and 2, black arrow indicates frons; 4, organization of setae on labrum (Callibaetis capixaba, left = dorsal, right = ventral) Chars 5, 6, 8, 15, 16, 19, 20, LAS = lateral arc of setae, DAS = distal arc of setae, DMA = distomedial arc of setae, SDA = subdistal arc of setae on ventral surface, MDA = mediodorsal arc of setae, RSVL = ventral row of spine-like setae on lateral margin of labrum, TSVALM = tuft of setae ventrally on anterolateral margin of labrum; 5, organization of setae on labrum (Apobaetis sp.) Chars 10, 14, DDA = distodorsal arc of setae, SDMA = sub-distomedial arc of setae on dorsal surface; 6, organization of setae on labrum (Waltzoyphius sp.) Char. 12, SDLA = sub-distolateral arc of setae on dorsal surface; 7, detail of deep cleft incisors of right mandible (Callibaetis sp.) Char. 30, black arrow indicates the incisors; 8, detail of partially fused incisors of right mandible (Paracloeodes sp.) Char. 30, black arrow indicates the incisors; 9, detail of a small lobe on apex of last segment of maxillary palp (Rivudiva sp.) Char. 43, back arrow indicates the lobe; 10, detail of basal projection of first dentiseta coupling with canine on maxilla (Varipes lasiobranchius) Char. 47; 11, detail of row of setae on outer margin of paraglossa (ROMP) (Callibaetis sp.) Char. 59; 12, detail of labial palp with red lines showing axes for angle measure (Apobaetis sp.) Char. 65; 13, detail of labial palp segment II and III, black arrow indicates the medial concavity on segment III (Callibaetis calloventer) Char. 68; 14, detail of trochanter inserted on ventral margin of femur, red line shows the connection (Varipes lasiobrachius) Char. 73; 15, trochanter inserted along entire base of femur, red line shows the connection (Rhopyscelis caldensis comb. n.) Char. 73." figureDoi="http://doi.org/10.5281/zenodo.3786798" httpUri="https://zenodo.org/record/3786798/files/figure.png" pageId="14" pageNumber="270">
|
||
Fig.
|
||
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|
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</figureCitation>
|
||
in
|
||
<bibRefCitation author="Gutierrez" box="[765,1000,1088,1114]" etAl="et al." firstAuthor="Gutierrez" pageId="14" pageNumber="270" pagination="291 - 295" refId="ref10661" refString="Gutierrez, Y., Dias, L. G. & Salles, F. F. (2013) Paracloeodes caldensis (Ephemeroptera: Baetidae), an atypical new species from the Colombian Andes. Zootaxa 3721: 291 - 295." type="journal article" year="2013">Gutiérrez et al. 2013</bibRefCitation>
|
||
) with bifid setae on distoventral arc; bifid setae on distomedial arc; mediodorsal arc of setae present and extending laterally; and absence of pair of denticles on the medial emargination. Right mandible (
|
||
<figureCitation box="[401,480,1192,1218]" captionStart="Figs" captionStartId="13.[106,148,1086,1108]" captionTargetBox="[106,1062,131,1061]" captionTargetId="figure@13.[106,1074,131,1061]" captionTargetPageId="13" captionText="Figs. 31–36. 31, genitalia, black arrows indicate the unistyliger inner projection (Paracloeodes sp.) Chars 142–149; 32, genitalia, black arrow indicates the spine and convex subgenital plate (Paracloeodes atroari) Chars 142–149; 33, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis sp.) Chars 142–149; 34, genitalia, black arrow indicates the concave subgenital plate (Paracloeodes sp.) Chars 142–149; 35, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis capixaba) Chars 142–149; 36, genitalia, black arrow indicates the triangular subgenital plate (Apobaetis fiuzai) Chars 142–149, adapted from Cruz et al. 2011." figureDoi="http://doi.org/10.5281/zenodo.3786802" httpUri="https://zenodo.org/record/3786802/files/figure.png" pageId="14" pageNumber="270">
|
||
Fig.
|
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</figureCitation>
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in
|
||
<bibRefCitation author="Nieto" box="[517,648,1192,1218]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
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;
|
||
<figureCitation box="[662,727,1192,1218]" captionStart="Figs" captionStartId="6.[98,140,1100,1122]" captionTargetId="figure@7.[106,1074,131,1525]" captionTargetPageId="7" captionText="Figs. 2–15. 2, elevation of frons with two keels (Paracloeodes charrua), Chars 1 and 2, black arrow indicates frons; 3, elevation of frons with one keel (Paracloeodes atroari) Chars 1 and 2, black arrow indicates frons; 4, organization of setae on labrum (Callibaetis capixaba, left = dorsal, right = ventral) Chars 5, 6, 8, 15, 16, 19, 20, LAS = lateral arc of setae, DAS = distal arc of setae, DMA = distomedial arc of setae, SDA = subdistal arc of setae on ventral surface, MDA = mediodorsal arc of setae, RSVL = ventral row of spine-like setae on lateral margin of labrum, TSVALM = tuft of setae ventrally on anterolateral margin of labrum; 5, organization of setae on labrum (Apobaetis sp.) Chars 10, 14, DDA = distodorsal arc of setae, SDMA = sub-distomedial arc of setae on dorsal surface; 6, organization of setae on labrum (Waltzoyphius sp.) Char. 12, SDLA = sub-distolateral arc of setae on dorsal surface; 7, detail of deep cleft incisors of right mandible (Callibaetis sp.) Char. 30, black arrow indicates the incisors; 8, detail of partially fused incisors of right mandible (Paracloeodes sp.) Char. 30, black arrow indicates the incisors; 9, detail of a small lobe on apex of last segment of maxillary palp (Rivudiva sp.) Char. 43, back arrow indicates the lobe; 10, detail of basal projection of first dentiseta coupling with canine on maxilla (Varipes lasiobranchius) Char. 47; 11, detail of row of setae on outer margin of paraglossa (ROMP) (Callibaetis sp.) Char. 59; 12, detail of labial palp with red lines showing axes for angle measure (Apobaetis sp.) Char. 65; 13, detail of labial palp segment II and III, black arrow indicates the medial concavity on segment III (Callibaetis calloventer) Char. 68; 14, detail of trochanter inserted on ventral margin of femur, red line shows the connection (Varipes lasiobrachius) Char. 73; 15, trochanter inserted along entire base of femur, red line shows the connection (Rhopyscelis caldensis comb. n.) Char. 73." figureDoi="http://doi.org/10.5281/zenodo.3786798" httpUri="https://zenodo.org/record/3786798/files/figure.png" pageId="14" pageNumber="270">
|
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Fig.
|
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in
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<bibRefCitation author="Gutierrez" box="[764,1000,1192,1218]" etAl="et al." firstAuthor="Gutierrez" pageId="14" pageNumber="270" pagination="291 - 295" refId="ref10661" refString="Gutierrez, Y., Dias, L. G. & Salles, F. F. (2013) Paracloeodes caldensis (Ephemeroptera: Baetidae), an atypical new species from the Colombian Andes. Zootaxa 3721: 291 - 295." type="journal article" year="2013">Gutiérrez et al. 2013</bibRefCitation>
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) with external margin convex; prostheca slender and bifid; incisors partially fused; and inner set of incisors at right angle with outer set. Left mandible (
|
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<figureCitation box="[746,824,1261,1287]" captionStart="Figs" captionStartId="13.[106,148,1086,1108]" captionTargetBox="[106,1062,131,1061]" captionTargetId="figure@13.[106,1074,131,1061]" captionTargetPageId="13" captionText="Figs. 31–36. 31, genitalia, black arrows indicate the unistyliger inner projection (Paracloeodes sp.) Chars 142–149; 32, genitalia, black arrow indicates the spine and convex subgenital plate (Paracloeodes atroari) Chars 142–149; 33, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis sp.) Chars 142–149; 34, genitalia, black arrow indicates the concave subgenital plate (Paracloeodes sp.) Chars 142–149; 35, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis capixaba) Chars 142–149; 36, genitalia, black arrow indicates the triangular subgenital plate (Apobaetis fiuzai) Chars 142–149, adapted from Cruz et al. 2011." figureDoi="http://doi.org/10.5281/zenodo.3786802" httpUri="https://zenodo.org/record/3786802/files/figure.png" pageId="14" pageNumber="270">
|
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Fig.
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</figureCitation>
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in
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<bibRefCitation author="Nieto" box="[860,989,1261,1287]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
|
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;
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<figureCitation box="[1002,1066,1261,1287]" captionStart="Figs" captionStartId="6.[98,140,1100,1122]" captionTargetId="figure@7.[106,1074,131,1525]" captionTargetPageId="7" captionText="Figs. 2–15. 2, elevation of frons with two keels (Paracloeodes charrua), Chars 1 and 2, black arrow indicates frons; 3, elevation of frons with one keel (Paracloeodes atroari) Chars 1 and 2, black arrow indicates frons; 4, organization of setae on labrum (Callibaetis capixaba, left = dorsal, right = ventral) Chars 5, 6, 8, 15, 16, 19, 20, LAS = lateral arc of setae, DAS = distal arc of setae, DMA = distomedial arc of setae, SDA = subdistal arc of setae on ventral surface, MDA = mediodorsal arc of setae, RSVL = ventral row of spine-like setae on lateral margin of labrum, TSVALM = tuft of setae ventrally on anterolateral margin of labrum; 5, organization of setae on labrum (Apobaetis sp.) Chars 10, 14, DDA = distodorsal arc of setae, SDMA = sub-distomedial arc of setae on dorsal surface; 6, organization of setae on labrum (Waltzoyphius sp.) Char. 12, SDLA = sub-distolateral arc of setae on dorsal surface; 7, detail of deep cleft incisors of right mandible (Callibaetis sp.) Char. 30, black arrow indicates the incisors; 8, detail of partially fused incisors of right mandible (Paracloeodes sp.) Char. 30, black arrow indicates the incisors; 9, detail of a small lobe on apex of last segment of maxillary palp (Rivudiva sp.) Char. 43, back arrow indicates the lobe; 10, detail of basal projection of first dentiseta coupling with canine on maxilla (Varipes lasiobranchius) Char. 47; 11, detail of row of setae on outer margin of paraglossa (ROMP) (Callibaetis sp.) Char. 59; 12, detail of labial palp with red lines showing axes for angle measure (Apobaetis sp.) Char. 65; 13, detail of labial palp segment II and III, black arrow indicates the medial concavity on segment III (Callibaetis calloventer) Char. 68; 14, detail of trochanter inserted on ventral margin of femur, red line shows the connection (Varipes lasiobrachius) Char. 73; 15, trochanter inserted along entire base of femur, red line shows the connection (Rhopyscelis caldensis comb. n.) Char. 73." figureDoi="http://doi.org/10.5281/zenodo.3786798" httpUri="https://zenodo.org/record/3786798/files/figure.png" pageId="14" pageNumber="270">
|
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Fig.
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in
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<bibRefCitation author="Gutierrez" box="[130,370,1296,1322]" etAl="et al." firstAuthor="Gutierrez" pageId="14" pageNumber="270" pagination="291 - 295" refId="ref10661" refString="Gutierrez, Y., Dias, L. G. & Salles, F. F. (2013) Paracloeodes caldensis (Ephemeroptera: Baetidae), an atypical new species from the Colombian Andes. Zootaxa 3721: 291 - 295." type="journal article" year="2013">Gutiérrez et al. 2013</bibRefCitation>
|
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) with external margin of mandibles convex; prostheca stout; molars with constriction; incisors partially fused; inner set of incisors at right angle with outer set. Maxilla (
|
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<figureCitation box="[373,452,1365,1391]" captionStart="Figs" captionStartId="13.[106,148,1086,1108]" captionTargetBox="[106,1062,131,1061]" captionTargetId="figure@13.[106,1074,131,1061]" captionTargetPageId="13" captionText="Figs. 31–36. 31, genitalia, black arrows indicate the unistyliger inner projection (Paracloeodes sp.) Chars 142–149; 32, genitalia, black arrow indicates the spine and convex subgenital plate (Paracloeodes atroari) Chars 142–149; 33, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis sp.) Chars 142–149; 34, genitalia, black arrow indicates the concave subgenital plate (Paracloeodes sp.) Chars 142–149; 35, genitalia, black arrow indicates the rounded apex of subgenital plate (Callibaetis capixaba) Chars 142–149; 36, genitalia, black arrow indicates the triangular subgenital plate (Apobaetis fiuzai) Chars 142–149, adapted from Cruz et al. 2011." figureDoi="http://doi.org/10.5281/zenodo.3786802" httpUri="https://zenodo.org/record/3786802/files/figure.png" pageId="14" pageNumber="270">
|
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Fig.
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in
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<bibRefCitation author="Nieto" box="[492,623,1365,1391]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
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;
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<figureCitation box="[638,704,1365,1391]" captionStart="Figs" captionStartId="6.[98,140,1100,1122]" captionTargetId="figure@7.[106,1074,131,1525]" captionTargetPageId="7" captionText="Figs. 2–15. 2, elevation of frons with two keels (Paracloeodes charrua), Chars 1 and 2, black arrow indicates frons; 3, elevation of frons with one keel (Paracloeodes atroari) Chars 1 and 2, black arrow indicates frons; 4, organization of setae on labrum (Callibaetis capixaba, left = dorsal, right = ventral) Chars 5, 6, 8, 15, 16, 19, 20, LAS = lateral arc of setae, DAS = distal arc of setae, DMA = distomedial arc of setae, SDA = subdistal arc of setae on ventral surface, MDA = mediodorsal arc of setae, RSVL = ventral row of spine-like setae on lateral margin of labrum, TSVALM = tuft of setae ventrally on anterolateral margin of labrum; 5, organization of setae on labrum (Apobaetis sp.) Chars 10, 14, DDA = distodorsal arc of setae, SDMA = sub-distomedial arc of setae on dorsal surface; 6, organization of setae on labrum (Waltzoyphius sp.) Char. 12, SDLA = sub-distolateral arc of setae on dorsal surface; 7, detail of deep cleft incisors of right mandible (Callibaetis sp.) Char. 30, black arrow indicates the incisors; 8, detail of partially fused incisors of right mandible (Paracloeodes sp.) Char. 30, black arrow indicates the incisors; 9, detail of a small lobe on apex of last segment of maxillary palp (Rivudiva sp.) Char. 43, back arrow indicates the lobe; 10, detail of basal projection of first dentiseta coupling with canine on maxilla (Varipes lasiobranchius) Char. 47; 11, detail of row of setae on outer margin of paraglossa (ROMP) (Callibaetis sp.) Char. 59; 12, detail of labial palp with red lines showing axes for angle measure (Apobaetis sp.) Char. 65; 13, detail of labial palp segment II and III, black arrow indicates the medial concavity on segment III (Callibaetis calloventer) Char. 68; 14, detail of trochanter inserted on ventral margin of femur, red line shows the connection (Varipes lasiobrachius) Char. 73; 15, trochanter inserted along entire base of femur, red line shows the connection (Rhopyscelis caldensis comb. n.) Char. 73." figureDoi="http://doi.org/10.5281/zenodo.3786798" httpUri="https://zenodo.org/record/3786798/files/figure.png" pageId="14" pageNumber="270">
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Fig.
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in
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<bibRefCitation author="Gutierrez" box="[743,981,1365,1392]" etAl="et al." firstAuthor="Gutierrez" pageId="14" pageNumber="270" pagination="291 - 295" refId="ref10661" refString="Gutierrez, Y., Dias, L. G. & Salles, F. F. (2013) Paracloeodes caldensis (Ephemeroptera: Baetidae), an atypical new species from the Colombian Andes. Zootaxa 3721: 291 - 295." type="journal article" year="2013">Gutiérrez et al. 2013</bibRefCitation>
|
||
) innerdorsal margin with two rows of setae; first dentiseta more robust than others; first dentiseta not coupling with canine at base (
|
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<figureCitation box="[591,670,1434,1461]" captionStart="Figs" captionStartId="6.[98,140,1100,1122]" captionTargetId="figure@7.[106,1074,131,1525]" captionTargetPageId="7" captionText="Figs. 2–15. 2, elevation of frons with two keels (Paracloeodes charrua), Chars 1 and 2, black arrow indicates frons; 3, elevation of frons with one keel (Paracloeodes atroari) Chars 1 and 2, black arrow indicates frons; 4, organization of setae on labrum (Callibaetis capixaba, left = dorsal, right = ventral) Chars 5, 6, 8, 15, 16, 19, 20, LAS = lateral arc of setae, DAS = distal arc of setae, DMA = distomedial arc of setae, SDA = subdistal arc of setae on ventral surface, MDA = mediodorsal arc of setae, RSVL = ventral row of spine-like setae on lateral margin of labrum, TSVALM = tuft of setae ventrally on anterolateral margin of labrum; 5, organization of setae on labrum (Apobaetis sp.) Chars 10, 14, DDA = distodorsal arc of setae, SDMA = sub-distomedial arc of setae on dorsal surface; 6, organization of setae on labrum (Waltzoyphius sp.) Char. 12, SDLA = sub-distolateral arc of setae on dorsal surface; 7, detail of deep cleft incisors of right mandible (Callibaetis sp.) Char. 30, black arrow indicates the incisors; 8, detail of partially fused incisors of right mandible (Paracloeodes sp.) Char. 30, black arrow indicates the incisors; 9, detail of a small lobe on apex of last segment of maxillary palp (Rivudiva sp.) Char. 43, back arrow indicates the lobe; 10, detail of basal projection of first dentiseta coupling with canine on maxilla (Varipes lasiobranchius) Char. 47; 11, detail of row of setae on outer margin of paraglossa (ROMP) (Callibaetis sp.) Char. 59; 12, detail of labial palp with red lines showing axes for angle measure (Apobaetis sp.) Char. 65; 13, detail of labial palp segment II and III, black arrow indicates the medial concavity on segment III (Callibaetis calloventer) Char. 68; 14, detail of trochanter inserted on ventral margin of femur, red line shows the connection (Varipes lasiobrachius) Char. 73; 15, trochanter inserted along entire base of femur, red line shows the connection (Rhopyscelis caldensis comb. n.) Char. 73." figureDoi="http://doi.org/10.5281/zenodo.3786798" httpUri="https://zenodo.org/record/3786798/files/figure.png" pageId="14" pageNumber="270">Fig. 10</figureCitation>
|
||
); palp with two segments. Labium (Fig.
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<bibRefCitation author="Nieto" box="[229,362,1469,1495]" firstAuthor="Nieto" pageId="14" pageNumber="270" pagination="161 - 173" refId="ref10968" refString="Nieto, C. (2004) Redescription of Varipes (Ephemeroptera: Baetidae), with the description of new species from Bolivia and Argentina. Aquatic Insects 26: 161 - 173." type="journal article" year="2004">Nieto 2004</bibRefCitation>
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;
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<figureCitation box="[378,444,1469,1496]" captionStart="Figs" captionStartId="6.[98,140,1100,1122]" captionTargetId="figure@7.[106,1074,131,1525]" captionTargetPageId="7" captionText="Figs. 2–15. 2, elevation of frons with two keels (Paracloeodes charrua), Chars 1 and 2, black arrow indicates frons; 3, elevation of frons with one keel (Paracloeodes atroari) Chars 1 and 2, black arrow indicates frons; 4, organization of setae on labrum (Callibaetis capixaba, left = dorsal, right = ventral) Chars 5, 6, 8, 15, 16, 19, 20, LAS = lateral arc of setae, DAS = distal arc of setae, DMA = distomedial arc of setae, SDA = subdistal arc of setae on ventral surface, MDA = mediodorsal arc of setae, RSVL = ventral row of spine-like setae on lateral margin of labrum, TSVALM = tuft of setae ventrally on anterolateral margin of labrum; 5, organization of setae on labrum (Apobaetis sp.) Chars 10, 14, DDA = distodorsal arc of setae, SDMA = sub-distomedial arc of setae on dorsal surface; 6, organization of setae on labrum (Waltzoyphius sp.) Char. 12, SDLA = sub-distolateral arc of setae on dorsal surface; 7, detail of deep cleft incisors of right mandible (Callibaetis sp.) Char. 30, black arrow indicates the incisors; 8, detail of partially fused incisors of right mandible (Paracloeodes sp.) Char. 30, black arrow indicates the incisors; 9, detail of a small lobe on apex of last segment of maxillary palp (Rivudiva sp.) Char. 43, back arrow indicates the lobe; 10, detail of basal projection of first dentiseta coupling with canine on maxilla (Varipes lasiobranchius) Char. 47; 11, detail of row of setae on outer margin of paraglossa (ROMP) (Callibaetis sp.) Char. 59; 12, detail of labial palp with red lines showing axes for angle measure (Apobaetis sp.) Char. 65; 13, detail of labial palp segment II and III, black arrow indicates the medial concavity on segment III (Callibaetis calloventer) Char. 68; 14, detail of trochanter inserted on ventral margin of femur, red line shows the connection (Varipes lasiobrachius) Char. 73; 15, trochanter inserted along entire base of femur, red line shows the connection (Rhopyscelis caldensis comb. n.) Char. 73." figureDoi="http://doi.org/10.5281/zenodo.3786798" httpUri="https://zenodo.org/record/3786798/files/figure.png" pageId="14" pageNumber="270">
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Fig.
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in
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<bibRefCitation author="Gutierrez" box="[485,725,1469,1496]" etAl="et al." firstAuthor="Gutierrez" pageId="14" pageNumber="270" pagination="291 - 295" refId="ref10661" refString="Gutierrez, Y., Dias, L. G. & Salles, F. F. (2013) Paracloeodes caldensis (Ephemeroptera: Baetidae), an atypical new species from the Colombian Andes. Zootaxa 3721: 291 - 295." type="journal article" year="2013">Gutiérrez et al. 2013</bibRefCitation>
|
||
) with elongate mentum; base of glossa not overlapping the base of paraglossa; paraglossa subtriangular; distomedial projection of labial palp segment II rounded; segment III of labial palp conical (apex broadly pointed).
|
||
<emphasis bold="true" box="[299,392,1573,1599]" pageId="14" pageNumber="270">Thorax.</emphasis>
|
||
Hind wing pad absent. Legs (
|
||
<figureCitation box="[722,799,1573,1599]" captionStart="Figs" captionStartId="8.[99,141,1129,1151]" captionTargetId="figure@9.[106,1074,131,1523]" captionTargetPageId="9" captionText="Figs. 16–30. 16, detail of forefemur concave setae on dorsal margin (Paracloeodes atroari) Char. 77; 17, detail of forefemur, black arrow indicates the FLRNDM = forefemur longitudinal row of setae near dorsal margin on anterior surface (Paracloeodes eurybranchus) Char. 80; 18, detail of forefemur, black arrow indicates the FLRNML = forefemur longitudinal row of setae at middle on anterior surface (Paracloeodes sp.) Char. 82; 19, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis singuil comb. n.) Char. 85 and 86; 20, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis singuil comb. n.) Chars 89 and 90; 21, anterior surface of forefemur, black arrow indicates the vestigial row of setae, (Rhopyscelis caldensis comb. n.) Chars 85 and 86; 22, anterior surface of hind femur, black arrow indicates the transversal row of setae, (Rhopyscelis caldensis comb. n.) Chars 89 and 90; 23, detail of transversal row of setae on hind femur, (Rivudiva trichobasis) Chars 89 and 91; 24, detail of transversal row of setae on hind femur, (Varipes helenae) Chars 89 and 91; 25, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis singuil comb. n.) Chars 96–101; 26, foreclaw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96–101; 27, hind claw, black arrow indicates the denticles apically directed (Rhopyscelis caldensis comb. n.) Chars 96– 101; 28, hind claw, black arrows indicate the regularity of row of denticles, (Rivudiva trichobasis) Chars 99 and 102; 29, foreclaw, black arrow indicates the absence of denticles on base, red arrow indicates the conical and apical denticles, (Varipes lasiobrachius) Chars 106 and 107; 30, detail of foreclaw, black arrow indicates apical denticles on claw extremely higher than the others (Gen. A.) Char. 104." figureDoi="http://doi.org/10.5281/zenodo.3786800" httpUri="https://zenodo.org/record/3786800/files/figure.png" pageId="14" pageNumber="270">Fig. 22</figureCitation>
|
||
) with trochanter insert- ed along entire base of femur; many long and thin setae on femora; vestigial transverse row of long setae on proximal third of forefemur; mid and hind femora with transverse row of long setae on proximal third; sockets of setae on the mid and hind transverse rows not touching each other; two rows of denticles on the claws; foreclaw denticles apically directed; denticles triangular.
|
||
<emphasis bold="true" box="[582,706,238,264]" pageId="15" pageNumber="271">Abdomen.</emphasis>
|
||
Gills on segment I–VII without folds, gill I not racquet-shaped; posterior margin of terga with spines; abdominal terga surface creased, without scale-like setae/socket; terminal filament subequal in length than cerci; caudal filaments with spines on each segment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[106,546,377,403]" pageId="15" pageNumber="271" type="distribution">
|
||
<paragraph blockId="15.[106,1074,134,403]" box="[106,546,377,403]" pageId="15" pageNumber="271">
|
||
<emphasis box="[106,248,377,403]" italics="true" pageId="15" pageNumber="271">Distribution:</emphasis>
|
||
<collectingCountry box="[256,368,377,403]" name="Argentina" pageId="15" pageNumber="271">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry box="[425,542,377,403]" name="Colombia" pageId="15" pageNumber="271">Colombia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="16" lastPageNumber="272" pageId="15" pageNumber="271" type="discussion">
|
||
<paragraph blockId="15.[106,1074,462,1633]" box="[106,237,462,488]" pageId="15" pageNumber="271">
|
||
<emphasis bold="true" box="[106,237,462,488]" pageId="15" pageNumber="271">Comments</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="15.[106,1074,462,1633]" pageId="15" pageNumber="271">
|
||
Considering the topology, the position of (
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[610,730,497,523]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">
|
||
<emphasis box="[610,730,497,523]" italics="true" pageId="15" pageNumber="271">P. caldensis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[767,871,497,523]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">
|
||
<emphasis box="[767,871,497,523]" italics="true" pageId="15" pageNumber="271">V. singuil</emphasis>
|
||
</taxonomicName>
|
||
) corroborated its placement in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[268,346,533,558]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[268,346,533,558]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
as subgenus or a new genus; in a broad sense, Gen. A could also belong to
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[280,358,567,592]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[280,358,567,592]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
. However, evaluating the stability of the clades,
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[929,1007,567,592]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[929,1007,567,592]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
(narrow sense) + (
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[263,381,601,627]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">
|
||
<emphasis box="[263,381,601,627]" italics="true" pageId="15" pageNumber="271">P. caldensis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[412,514,601,627]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">
|
||
<emphasis box="[412,514,601,627]" italics="true" pageId="15" pageNumber="271">V. singuil</emphasis>
|
||
</taxonomicName>
|
||
) is a weakly supported clade, while (
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[932,1050,601,627]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">
|
||
<emphasis box="[932,1050,601,627]" italics="true" pageId="15" pageNumber="271">P. caldensis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[106,210,636,662]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">
|
||
<emphasis box="[106,210,636,662]" italics="true" pageId="15" pageNumber="271">V. singuil</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[282,360,637,662]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[282,360,637,662]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
have high values of the indices and, in the case of
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[966,1044,637,662]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[966,1044,637,662]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
, a large number of synapomorphies. Gen. A + (
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[633,753,670,696]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">
|
||
<emphasis box="[633,753,670,696]" italics="true" pageId="15" pageNumber="271">P. caldensis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[789,893,670,696]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">
|
||
<emphasis box="[789,893,670,696]" italics="true" pageId="15" pageNumber="271">V. singuil</emphasis>
|
||
</taxonomicName>
|
||
) +
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[937,1015,671,696]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[937,1015,671,696]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
) is a weakly supported clade, while Gen. A, (
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[566,685,705,731]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">
|
||
<emphasis box="[566,685,705,731]" italics="true" pageId="15" pageNumber="271">P. caldensis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[717,820,705,731]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">
|
||
<emphasis box="[717,820,705,731]" italics="true" pageId="15" pageNumber="271">V. singuil</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[888,966,706,731]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[888,966,706,731]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
are three well-supported groups. Analytical settings were varied to verify the robustness of these monophyletic groups (see Materials and Methods section). All topologies resulting from these exploratory searches recovered Gen. A, (
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[691,809,809,835]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">
|
||
<emphasis box="[691,809,809,835]" italics="true" pageId="15" pageNumber="271">P. caldensis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[840,942,809,835]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">
|
||
<emphasis box="[840,942,809,835]" italics="true" pageId="15" pageNumber="271">V. singuil</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName box="[967,1066,809,835]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[967,1066,809,835]" italics="true" pageId="15" pageNumber="271">Rivudiva</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[106,184,845,870]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[106,184,845,870]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authorityName="Day" authorityYear="1955" box="[248,381,844,870]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[248,381,844,870]" italics="true" pageId="15" pageNumber="271">Paracloeodes</emphasis>
|
||
</taxonomicName>
|
||
as well-supported and stable monophyletic groups.
|
||
</paragraph>
|
||
<paragraph blockId="15.[106,1074,462,1633]" lastBlockId="16.[98,1066,134,680]" lastPageId="16" lastPageNumber="272" pageId="15" pageNumber="271">
|
||
In addition to the highly stable monophyletic groups and the understanding that the long setae state is an independent acquisition between groups, a large number of generic characteristics was obtained in morphological analyses (some of them treated as characters and states in the matrix). Those attributed to
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[789,908,982,1008]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">
|
||
<emphasis box="[789,908,982,1008]" italics="true" pageId="15" pageNumber="271">P. caldensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[970,1074,982,1008]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">
|
||
<emphasis box="[970,1074,982,1008]" italics="true" pageId="15" pageNumber="271">V. singuil</emphasis>
|
||
</taxonomicName>
|
||
have proved to be very effective in differentiating both of these species, not only from
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[106,184,1053,1078]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[106,184,1053,1078]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
, but also from all other Neotropical and Nearctic genera as follows: scape and pedicel without scale-like tubercles in
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[537,655,1086,1112]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">
|
||
<emphasis box="[537,655,1086,1112]" italics="true" pageId="15" pageNumber="271">P. caldensis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[685,786,1086,1112]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">
|
||
<emphasis box="[685,786,1086,1112]" italics="true" pageId="15" pageNumber="271">V. singuil</emphasis>
|
||
</taxonomicName>
|
||
, while present in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[988,1066,1087,1112]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[988,1066,1087,1112]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
; mandibular incisors partially fused in
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[544,663,1121,1147]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">
|
||
<emphasis box="[544,663,1121,1147]" italics="true" pageId="15" pageNumber="271">P. caldensis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[697,801,1121,1147]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">
|
||
<emphasis box="[697,801,1121,1147]" italics="true" pageId="15" pageNumber="271">V. singuil</emphasis>
|
||
</taxonomicName>
|
||
, while fused in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[988,1066,1122,1147]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[988,1066,1122,1147]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
; inner set of incisors at right angle to the outer set in
|
||
<emphasis box="[712,966,1156,1182]" italics="true" pageId="15" pageNumber="271">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[712,831,1156,1182]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[863,966,1156,1182]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while in the same plane in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[314,392,1191,1216]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[314,392,1191,1216]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
; maxilla with the first dentiseta with the same robustness as the others and with its basal projection not coupling with canine in
|
||
<emphasis italics="true" pageId="15" pageNumber="271">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[898,1017,1225,1251]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while the first dentiseta is more robust than the others and with base coupling with canine in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[271,349,1295,1320]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[271,349,1295,1320]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
; segment III of labial palp semi conical (apex broadly pointed) in
|
||
<emphasis box="[106,355,1329,1355]" italics="true" pageId="15" pageNumber="271">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[106,223,1329,1355]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[254,355,1329,1355]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while globose in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[560,638,1330,1355]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[560,638,1330,1355]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
; base of glossa not overlapping base of paraglossa in
|
||
<emphasis box="[255,500,1364,1390]" italics="true" pageId="15" pageNumber="271">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[255,372,1364,1390]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[400,500,1364,1390]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while overlapping in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[749,827,1365,1390]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[749,827,1365,1390]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
; lingua with distomedial projection in
|
||
<emphasis box="[318,580,1398,1424]" italics="true" pageId="15" pageNumber="271">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[318,439,1398,1424]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[475,580,1398,1424]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while absent in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[785,863,1399,1424]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[785,863,1399,1424]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
; dorsal margin of femora parallel to ventral in
|
||
<emphasis box="[423,669,1433,1459]" italics="true" pageId="15" pageNumber="271">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[423,540,1433,1459]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[568,669,1433,1459]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while convex in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[864,942,1434,1459]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[864,942,1434,1459]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
; trochanter inserted along entire base of femur in
|
||
<emphasis box="[527,773,1468,1494]" italics="true" pageId="15" pageNumber="271">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[527,644,1468,1494]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[672,773,1468,1494]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while at the base of femur in
|
||
<emphasis box="[135,219,1503,1528]" italics="true" pageId="15" pageNumber="271">
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[135,214,1503,1528]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">Varipes</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
vestigial transverse row of long setae on proximal third of forefemur in
|
||
<emphasis italics="true" pageId="15" pageNumber="271">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[199,301,1537,1563]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while not vestigial in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[560,638,1538,1563]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[560,638,1538,1563]" italics="true" pageId="15" pageNumber="271">Varipes</emphasis>
|
||
</taxonomicName>
|
||
; mid and hind femora with transverse row of long setae on proximal third with sockets not touching each other in
|
||
<emphasis italics="true" pageId="15" pageNumber="271">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[936,1051,1572,1598]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="caldensis">P.caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[106,208,1606,1632]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while touching each other in
|
||
<emphasis box="[558,642,1607,1632]" italics="true" pageId="15" pageNumber="271">
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[558,637,1607,1632]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="271" phylum="Arthropoda" rank="genus">Varipes</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
tibio-patelar suture with length equal or less than one third of tibia in
|
||
<emphasis box="[457,703,134,160]" italics="true" pageId="16" pageNumber="272">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[457,574,134,160]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[603,703,134,160]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while equal or more than a half of tibia in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[216,294,170,195]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[216,294,170,195]" italics="true" pageId="16" pageNumber="272">Varipes</emphasis>
|
||
</taxonomicName>
|
||
; claws with triangular and apically directed denticles starting at base in
|
||
<emphasis box="[129,380,204,230]" italics="true" pageId="16" pageNumber="272">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[129,247,204,230]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[278,380,204,230]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while conical, perpendicular and starting at medial third in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[98,176,239,264]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[98,176,239,264]" italics="true" pageId="16" pageNumber="272">Varipes</emphasis>
|
||
</taxonomicName>
|
||
; gill I symmetrical in
|
||
<emphasis box="[425,674,238,264]" italics="true" pageId="16" pageNumber="272">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[425,543,238,264]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[573,674,238,264]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, while racquet-shaped in
|
||
<taxonomicName authorityName="Lugo-Ortiz & McCafferty" authorityYear="1998" box="[965,1043,239,264]" class="Insecta" family="Baetidae" genus="Varipes" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[965,1043,239,264]" italics="true" pageId="16" pageNumber="272">Varipes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="16.[98,1066,134,680]" pageId="16" pageNumber="272">
|
||
<emphasis box="[128,374,273,299]" italics="true" pageId="16" pageNumber="272">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[128,244,273,299]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[274,374,273,299]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
could also be confused with the Nearctic genus
|
||
<taxonomicName authority="McCafferty" authorityName="McCafferty" authorityYear="2011" class="Insecta" family="Baetidae" genus="Kirmaushenkreena" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="16" pageNumber="272">Kirmaushenkreena</emphasis>
|
||
McCafferty
|
||
</taxonomicName>
|
||
, mainly based on long setae on forefemur and claws with symmetrical rows of denticles (key of
|
||
<bibRefCitation author="Waltz & Burian" box="[381,636,342,369]" firstAuthor="Waltz" pageId="16" pageNumber="272" pagination="181 - 236" refId="ref11476" refString="Waltz, R. D. & Burian, S. K. (2008) Ephemeroptera. In: Merritt, R. W., Cummins, K. W. & Berg, M. B. (Eds.), An Introduction to the Aquatic Insects of North America. Kendall / Hunt, Dubuque, Iowa, pp. 181 - 236." type="book chapter" year="2008">Waltz & Burian, 2008</bibRefCitation>
|
||
). The genus
|
||
<taxonomicName authorityName="McCafferty" authorityYear="2011" box="[783,986,342,368]" class="Insecta" family="Baetidae" genus="Kirmaushenkreena" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[783,986,342,368]" italics="true" pageId="16" pageNumber="272">Kirmaushenkreena</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from
|
||
<emphasis box="[317,562,377,403]" italics="true" pageId="16" pageNumber="272">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[317,433,377,403]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[462,562,377,403]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
, among other characteristics, by the presence of a subapical pair of setae on claws, while is absent in
|
||
<emphasis box="[715,965,412,438]" italics="true" pageId="16" pageNumber="272">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[715,832,412,438]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[864,965,412,438]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
; the first dentiseta with its base coupling with canine, while not coupling in
|
||
<emphasis box="[821,1059,446,472]" italics="true" pageId="16" pageNumber="272">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[821,936,446,472]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="caldensis">P.caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[961,1059,446,472]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
; and transverse row of setae on femora absent, while present in
|
||
<emphasis box="[784,1035,481,507]" italics="true" pageId="16" pageNumber="272">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[784,900,481,507]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[928,1031,481,507]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="16.[98,1066,134,680]" pageId="16" pageNumber="272">
|
||
Taking into account the phylogenetic position, the number of synapomorphies, clade stability, and the great number of generic characteristics that differentiate the
|
||
<emphasis box="[98,345,585,611]" italics="true" pageId="16" pageNumber="272">
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Gutierrez, Dias & Salles" baseAuthorityYear="2013" box="[98,215,585,611]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="caldensis">P. caldensis</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" baseAuthorityName="Nieto" baseAuthorityYear="2004" box="[245,345,585,611]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="species" species="singuil">V. singuil</taxonomicName>
|
||
</emphasis>
|
||
clade from other genera in the Nearctic and Neotropical realms, we propose this clade as a new genus,
|
||
<taxonomicName authority="Cruza & Sallesb & Hamada, 2020" authorityName="Cruza & Sallesb & Hamada" authorityYear="2020" box="[553,670,620,646]" class="Insecta" family="Baetidae" genus="Rhopyscelis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="272" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[553,670,620,646]" italics="true" pageId="16" pageNumber="272">Rhopyscelis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[681,765,620,646]" pageId="16" pageNumber="272">
|
||
<taxonomicNameLabel box="[681,765,620,646]" pageId="16" pageNumber="272" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, described and diagnosed above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |