226 lines
28 KiB
XML
226 lines
28 KiB
XML
<document id="49A67B2630A4A34FBCE0E236F1B238F1" ID-CLB-Dataset="25594" ID-DOI="10.11646/zootaxa.4646.3.3" ID-GBIF-Dataset="4a3d31ca-e16b-4ccf-8407-231aa27702f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3351058" ID-ZooBank="25CA4A18-955F-4EA7-8978-E93032B54A2E" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564032263204" checkinUser="plazi" docAuthor="Cranston, Peter S." docDate="2019" docId="B33701109B2B1E75C68CFB59FBE13482" docLanguage="en" docName="zootaxao.4646.3.3.pdf" docOrigin="Zootaxa 4646 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Riethia paluma Cranston 2019, sp. n." docType="treatment" docUuid="1D698677-C7ED-401E-A5D3-2AD11708C222" docUuidSource="ZooBank" docVersion="13" lastPageNumber="482" masterDocId="4F0E79689B3F1E60C61BFFD0FFA93021" masterDocTitle="Riethia (Kieffer 1917) (Diptera: Chironomidae) revised for the Austro-Pacific region" masterLastPageNumber="500" masterPageNumber="461" pageNumber="481" updateTime="1698747359191" updateUser="ExternalLinkService">
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<mods:title id="24D799CFF0957A9A60422704119CF0C9">Riethia (Kieffer 1917) (Diptera: Chironomidae) revised for the Austro-Pacific region</mods:title>
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<mods:name id="3A668A6C51C7C368BC12A06317D2C9A8" type="personal">
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<mods:roleTerm id="DAE56109466EE31B61E26637EB25FA18">Author</mods:roleTerm>
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<mods:namePart id="F4207C6FD454568640D088E355338E69">Cranston, Peter S.</mods:namePart>
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<mods:typeOfResource id="AD053D8456495BB37EC40B5A0F291D23">text</mods:typeOfResource>
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<mods:title id="57B615A82615823D6EA0A16A188D81F6">Zootaxa</mods:title>
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<mods:part id="183B8BC3B05318257BBDF3833510C255">
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<mods:date id="1A9BCB11E5BE05FE9E3E274ECC4CD977">2019</mods:date>
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<mods:number id="CF41918CDA7711323436002F143C882B">2019-07-25</mods:number>
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<mods:number id="97DCD6ED3AE77FF27D113CBEA0969202">4646</mods:number>
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<mods:number id="4B4ECD162FA326689DC0AE83B723A864">3</mods:number>
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<mods:classification id="AF407C3E5229E85DEA61717BE4958CB5">journal article</mods:classification>
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<mods:identifier id="2473CBD60B72256B890CD91060F54458" type="DOI">10.11646/zootaxa.4646.3.3</mods:identifier>
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<treatment id="B33701109B2B1E75C68CFB59FBE13482" ID-DOI="http://doi.org/10.5281/zenodo.5587166" ID-GBIF-Taxon="159413593" ID-Zenodo-Dep="5587166" LSID="urn:lsid:zoobank.org:act:1D698677-C7ED-401E-A5D3-2AD11708C222" httpUri="http://treatment.plazi.org/id/B33701109B2B1E75C68CFB59FBE13482" lastPageId="21" lastPageNumber="482" pageId="20" pageNumber="481">
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<subSubSection id="7384E38D9B2B1E74C68CFB59FDB13482" box="[151,536,1161,1188]" pageId="20" pageNumber="481" type="nomenclature">
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<paragraph id="3B21B0069B2B1E74C68CFB59FDB13482" blockId="20.[151,536,1161,1259]" box="[151,536,1161,1188]" pageId="20" pageNumber="481">
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<heading id="6069076A9B2B1E74C68CFB59FDB13482" bold="true" box="[151,536,1161,1188]" fontSize="11" level="1" pageId="20" pageNumber="481" reason="1">
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<emphasis id="09EA6C149B2B1E74C68CFB59FDB13482" bold="true" box="[151,536,1161,1188]" pageId="20" pageNumber="481">
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<taxonomicName id="FC9ECB859B2B1E74C68CFB59FE673482" ID-CoL="9HNXY" authority="Cranston" authorityName="Cranston" authorityYear="2019" box="[151,462,1161,1188]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="481" phylum="Arthropoda" rank="species" species="paluma" status="sp. nov.">
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<emphasis id="09EA6C149B2B1E74C68CFB59FEFB3485" bold="true" box="[151,338,1161,1188]" italics="true" pageId="20" pageNumber="481">Riethia paluma</emphasis>
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Cranston
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</taxonomicName>
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<taxonomicNameLabel id="12D9D16F9B2B1E74C7CDFB59FDB13482" box="[470,536,1161,1187]" pageId="20" pageNumber="481" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="7384E38D9B2B1E74C68CFB7DFE1934CA" pageId="20" pageNumber="481" type="description">
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<paragraph id="3B21B0069B2B1E74C68CFB7DFEF834E9" blockId="20.[151,536,1161,1259]" box="[151,337,1197,1224]" pageId="20" pageNumber="481">
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(
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<figureCitation id="A3A5AC839B2B1E74C6BBFB7DFE8D34E6" box="[160,292,1197,1223]" captionStart="FIGURE 4" captionStartId="13.[151,250,1897,1922]" captionTargetBox="[199,1399,178,1821]" captionTargetId="figure@13.[175,1412,166,1877]" captionTargetPageId="13" captionText="FIGURE 4. Riethia spp. Pupa. Abdominal tergites: A. R. noongar sp.n., B. R. paluma sp.n., C. R. plumosa Freeman, D. R. stictoptera Kieffer. Posterolateral corner of VIII (‘comb’): E. R. noongar, F. R. paluma, G. R. plumosa, H. R. stictoptera I. R. cinctipes (var), J. R. phengari sp.n., K. R. neocaledonica sp.n.. Tergites: L, IV-V of R. neocaledonica; M. Tergites VI-VIII of R. queenslandensis sp.n., N. Sternites VI-VIII R. queenslandensis, O. Tergites V-AL of R. zeylandica." httpUri="https://zenodo.org/record/3351066/files/figure.png" pageId="20" pageNumber="481">Fig. 4B, 4F</figureCitation>
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;
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<figureCitation id="A3A5AC839B2B1E74C734FB7DFEE034E9" box="[303,329,1197,1224]" captionStart="FIGURE 6" captionStartId="15.[151,250,1892,1917]" captionTargetBox="[163,1417,176,1843]" captionTargetId="figure@15.[151,1436,166,1873]" captionTargetPageId="15" captionText="FIGURE 6. Riethia spp. Larva. Mid-dorsal head:A. R. donedwardi, sp.n., B. R. noongar sp.n.. Clypeus: C. R. larval sp. ‘C’, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. neocaledonica sp.n., I. R. noongar sp.n., J. R. paluma sp.n., K. R. phengari sp.n., L. R. plumosa Freeman, M. R. queenslandensis sp.n., N. R. stictoptera Kieffer, O. R. wazeylandica sp.n., P. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351070/files/figure.png" pageId="20" pageNumber="481">6J</figureCitation>
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)
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</paragraph>
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<paragraph id="3B21B0069B2B1E74C68CFB01FE1934CA" blockId="20.[151,536,1161,1259]" box="[151,432,1233,1259]" pageId="20" pageNumber="481">urn:lsid:zoobank.org:act:</paragraph>
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</subSubSection>
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<subSubSection id="7384E38D9B2B1E75C68CFAC9FBE13482" lastPageId="21" lastPageNumber="482" pageId="20" pageNumber="481" type="materials_examined">
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<paragraph id="3B21B0069B2B1E74C68CFAC9FDE23576" blockId="20.[151,1437,1304,2015]" pageId="20" pageNumber="481">
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||
<emphasis id="09EA6C149B2B1E74C68CFAC9FEED3512" bold="true" box="[151,324,1304,1331]" pageId="20" pageNumber="481">Type material</emphasis>
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.
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<materialsCitation id="8BF6BA5B9B2B1E74C743FAC9FDE23576" ID-GBIF-Occurrence="2382714531" collectingDate="1998-10-24" collectionCode="ANIC" collectorName="Birthday Ck." country="Australia" elevation="800" latitude="-18.8" location="Queensland" longLatPrecision="1274" longitude="146.15" pageId="20" pageNumber="481" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="holotype">
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||
<typeStatus id="E4250EA49B2B1E74C743FAC9FE6C3512" box="[344,453,1305,1331]" pageId="20" pageNumber="481" type="holotype">
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||
<emphasis id="09EA6C149B2B1E74C743FAC9FE6C3512" bold="true" box="[344,453,1305,1331]" pageId="20" pageNumber="481">Holotype</emphasis>
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</typeStatus>
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Le/Pe/
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<specimenCount id="2D987B8F9B2B1E74C43FFAC9FD9B3512" box="[548,562,1305,1331]" pageId="20" pageNumber="481" type="female">♀</specimenCount>
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,
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<collectingCountry id="4389F0969B2B1E74C459FAC9FD4D3512" box="[578,740,1305,1331]" name="Australia" pageId="20" pageNumber="481">AUSTRALIA</collectingCountry>
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,
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<collectingRegion id="F95A7EE49B2B1E74C4EFFAC9FCD23512" box="[756,891,1305,1331]" country="Australia" name="Queensland" pageId="20" pageNumber="481">Queensland</collectingRegion>
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, Paluma,
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<collectorName id="966BD5D09B2B1E74C5EFFAC9FB243512" box="[1012,1165,1305,1331]" pageId="20" pageNumber="481">Birthday Ck.</collectorName>
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,
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||
<elevation id="B0B357359B2B1E74C284FAC9FA873512" box="[1183,1326,1305,1331]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="481" unit="m" value="800.0">
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||
<quantity id="FC661DE39B2B1E74C284FAC9FB423512" box="[1183,1259,1305,1331]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="481" unit="m" value="800.0">800 m</quantity>
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a.s.l.
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</elevation>
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,
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||
<geoCoordinate id="5EAAD6C19B2B1E74C35BFAC9FA343512" box="[1344,1437,1305,1331]" degrees="18" direction="south" minutes="48" orientation="latitude" pageId="20" pageNumber="481" precision="925" value="-18.8">18°48’S</geoCoordinate>
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||
<geoCoordinate id="5EAAD6C19B2B1E74C68CFAEDFEAC3576" box="[151,261,1341,1367]" degrees="146" direction="east" minutes="09" orientation="longitude" pageId="20" pageNumber="481" precision="925" value="146.15">146°09’E</geoCoordinate>
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,
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||
<collectingDate id="5F646F2E9B2B1E74C70AFAEDFE293576" box="[273,384,1341,1367]" pageId="20" pageNumber="481" value="1998-10-24">
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||
<date id="4F2096C69B2B1E74C70AFAEDFE293576" box="[273,384,1341,1367]" pageId="20" pageNumber="481" value="1998-10-24">24.x.1998</date>
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||
</collectingDate>
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||
(
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<emphasis id="09EA6C149B2B1E74C78AFAEEFE753576" box="[401,476,1342,1367]" italics="true" pageId="20" pageNumber="481">McKie</emphasis>
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||
) (
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||
<collectionCode id="5D8F28C39B2B1E74C7EFFAEDFD963576" box="[500,575,1341,1367]" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="20" pageNumber="481">ANIC</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="3B21B0069B2B1E74C6DCFAB0FDBA355A" blockId="20.[151,1437,1304,2015]" box="[199,531,1376,1403]" pageId="20" pageNumber="481">
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<emphasis id="09EA6C149B2B1E74C6DCFAB0FEF8355A" bold="true" box="[199,337,1376,1403]" pageId="20" pageNumber="481">Description</emphasis>
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.
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<emphasis id="09EA6C149B2B1E74C77BFAB1FE34355A" bold="true" box="[352,413,1377,1403]" pageId="20" pageNumber="481">Male</emphasis>
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unknown.
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</paragraph>
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<paragraph id="3B21B0069B2B1E74C6DCFA55FF4D36C2" blockId="20.[151,1437,1304,2015]" pageId="20" pageNumber="481">
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<emphasis id="09EA6C149B2B1E74C6DCFA55FEB735BE" bold="true" box="[199,286,1413,1439]" pageId="20" pageNumber="481">Female</emphasis>
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. Colour (teneral) median vitta an anterior brown oval area, with lateral vittae brown, extending from midscutum to just anterior to scutellum; scutellum pale, postnotum brown, anepisternum with broad brown ‘smudge’. Wing pale at base, infuscate from cross-veins to apex. Legs darkened at distal apices of femora and proximal base of tibia (‘knees’); tibial apices darkened near spurs; apices of successive tarsomeres brown with distalmost all dark. B.l.
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<quantity id="FC661DE39B2B1E74C6D3F9C5FE88360E" box="[200,289,1557,1583]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="20" pageNumber="481" unit="mm" value="6.2">6.2 mm</quantity>
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, W.l.
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<quantity id="FC661DE39B2B1E74C77FF9C5FE17360E" box="[356,446,1557,1583]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="20" pageNumber="481" unit="mm" value="2.4">2.4 mm</quantity>
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. Head: Antenna flagellomere 137, 106, 100, 95, 165, AR 0.29; 14 clypeals, 3 clustered verticals. Palpomeres 50, 40, 205, 230, 240. Thorax. Ac?9, Dc 11, Pa 3, Scts 15. Wing V.R. 1.14. Vein setation: R with 24, R
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<subScript id="A71AB2439B2B1E74C714F9BEFEBE365D" attach="left" box="[271,279,1646,1660]" fontSize="6" pageId="20" pageNumber="481">1</subScript>
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with 25 and R
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<subScript id="A71AB2439B2B1E74C7D9F9BEFE75365D" attach="left" box="[450,476,1646,1660]" fontSize="6" pageId="20" pageNumber="481">4+5</subScript>
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47–60, sq. 11. Leg lengths: P
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<subScript id="A71AB2439B2B1E74C52BF9BEFC91365D" attach="left" box="[816,824,1646,1660]" fontSize="6" pageId="20" pageNumber="481">1</subScript>
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1100, 1120, 1500, 600, 500, 400, 200; P
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||
<subScript id="A71AB2439B2B1E74C311F9BEFABB365D" attach="left" box="[1290,1298,1646,1660]" fontSize="6" pageId="20" pageNumber="481">2</subScript>
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||
1200, 1160, 700, 400, 260, 200, 120; P
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||
<subScript id="A71AB2439B2B1E74C7DEF942FE643681" attach="left" box="[453,461,1682,1696]" fontSize="6" pageId="20" pageNumber="481">3</subScript>
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1200, 1240, 900, 540, 380, 260, 140. Ratios: LR
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<subScript id="A71AB2439B2B1E74C5E1F942FBAB3681" attach="left" box="[1018,1026,1682,1696]" fontSize="6" pageId="20" pageNumber="481">1</subScript>
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1.34, LR
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<subScript id="A71AB2439B2B1E74C270F942FBDA3681" attach="left" box="[1131,1139,1682,1696]" fontSize="6" pageId="20" pageNumber="481">2</subScript>
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||
0.44, LR
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<subScript id="A71AB2439B2B1E74C2C7F942FB4D3681" attach="left" box="[1244,1252,1682,1696]" fontSize="6" pageId="20" pageNumber="481">3</subScript>
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||
0.72;, BV
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<subScript id="A71AB2439B2B1E74C34CF942FAF63681" attach="left" box="[1367,1375,1682,1696]" fontSize="6" pageId="20" pageNumber="481">1</subScript>
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2.18, BV
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<subScript id="A71AB2439B2B1E74C6A5F966FF6F36E5" attach="left" box="[190,198,1718,1732]" fontSize="6" pageId="20" pageNumber="481">2</subScript>
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3.12, BV
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<subScript id="A71AB2439B2B1E74C734F966FE9E36E5" attach="left" box="[303,311,1718,1732]" fontSize="6" pageId="20" pageNumber="481">3</subScript>
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2.46; SV
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<subScript id="A71AB2439B2B1E74C7BBF966FE0136E5" attach="left" box="[416,424,1718,1732]" fontSize="6" pageId="20" pageNumber="481">1</subScript>
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1.35, SV
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<subScript id="A71AB2439B2B1E74C415F966FDBF36E5" attach="left" box="[526,534,1718,1732]" fontSize="6" pageId="20" pageNumber="481">2</subScript>
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3.37, SV
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<subScript id="A71AB2439B2B1E74C467F966FD2D36E5" attach="left" box="[636,644,1718,1732]" fontSize="6" pageId="20" pageNumber="481">3</subScript>
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2.62. Foretibial spur short, nearly simple 30 long; midtibial 55, 55, rear 85, 75.
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</paragraph>
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<paragraph id="3B21B0069B2B1E75C6DCF93DFED130DA" blockId="20.[151,1437,1304,2015]" lastBlockId="21.[151,1437,152,1187]" lastPageId="21" lastPageNumber="482" pageId="20" pageNumber="481">
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<emphasis id="09EA6C149B2B1E74C6DCF93DFEA43726" bold="true" box="[199,269,1773,1799]" pageId="20" pageNumber="481">Pupa.</emphasis>
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Pale yellow with darker dorsal thorax, anterior wing sheaths and lateral apophyses on posterior abdominal segments; comb brown. Cephalic area without warts, dorsal thorax rugose with multiserial rows of tubercle. Hook row on II discontinuous, occupying c. 40% of tergite width. Conjunctival spinule bands on III – IV separated from posterior tergal armament, 4–5 spinules wide, medially absent (‘divided’). Pedes spurii B absent, vortex well developed. Abdomen (
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<figureCitation id="A3A5AC839B2B1E74C78EF8ADFE4537B6" box="[405,492,1917,1943]" captionStart="FIGURE 4" captionStartId="13.[151,250,1897,1922]" captionTargetBox="[199,1399,178,1821]" captionTargetId="figure@13.[175,1412,166,1877]" captionTargetPageId="13" captionText="FIGURE 4. Riethia spp. Pupa. Abdominal tergites: A. R. noongar sp.n., B. R. paluma sp.n., C. R. plumosa Freeman, D. R. stictoptera Kieffer. Posterolateral corner of VIII (‘comb’): E. R. noongar, F. R. paluma, G. R. plumosa, H. R. stictoptera I. R. cinctipes (var), J. R. phengari sp.n., K. R. neocaledonica sp.n.. Tergites: L, IV-V of R. neocaledonica; M. Tergites VI-VIII of R. queenslandensis sp.n., N. Sternites VI-VIII R. queenslandensis, O. Tergites V-AL of R. zeylandica." httpUri="https://zenodo.org/record/3351066/files/figure.png" pageId="20" pageNumber="481">Fig. 4B</figureCitation>
|
||
) with tergites I and II essentially bare with few very fine spinules only immediately anterior to and narrower than hookrow. TIII–VI anteriorly with transverse band of spinules, broadest on II, diminishing in lateral extent to VI, always well separated from paired posterolateral spinule patches; TVII and VIII with few finest spinules. Taeniate setae V–VIII 3, 4, 4, 5; L
|
||
<subScript id="A71AB2439B2A1E75C4EAFF7AFCA33099" attach="left" box="[753,778,170,184]" fontSize="6" pageId="21" pageNumber="482">3–5</subScript>
|
||
on VIII clustered with respect to L
|
||
<subScript id="A71AB2439B2A1E75C28BFF7AFB013099" attach="left" box="[1168,1192,170,184]" fontSize="6" pageId="21" pageNumber="482">1–2</subScript>
|
||
. Comb (
|
||
<figureCitation id="A3A5AC839B2A1E75C312FF49FAF53092" box="[1289,1372,153,179]" captionStart="FIGURE 4" captionStartId="13.[151,250,1897,1922]" captionTargetBox="[199,1399,178,1821]" captionTargetId="figure@13.[175,1412,166,1877]" captionTargetPageId="13" captionText="FIGURE 4. Riethia spp. Pupa. Abdominal tergites: A. R. noongar sp.n., B. R. paluma sp.n., C. R. plumosa Freeman, D. R. stictoptera Kieffer. Posterolateral corner of VIII (‘comb’): E. R. noongar, F. R. paluma, G. R. plumosa, H. R. stictoptera I. R. cinctipes (var), J. R. phengari sp.n., K. R. neocaledonica sp.n.. Tergites: L, IV-V of R. neocaledonica; M. Tergites VI-VIII of R. queenslandensis sp.n., N. Sternites VI-VIII R. queenslandensis, O. Tergites V-AL of R. zeylandica." httpUri="https://zenodo.org/record/3351066/files/figure.png" pageId="21" pageNumber="482">Fig. 4F</figureCitation>
|
||
) with 1 strong, inner spine directed postero-medially and 3–4 subordinate spines. Anal lobe with 60–65 taeniae in tightly packed biserial row.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B2A1E75C6DCFED5FAF1318E" blockId="21.[151,1437,152,1187]" pageId="21" pageNumber="482">
|
||
<emphasis id="09EA6C149B2A1E75C6DCFED5FEBE313E" bold="true" box="[199,279,261,287]" pageId="21" pageNumber="482">Larva.</emphasis>
|
||
Head capsule yellow with golden brown mental and mandibular teeth (inner only slightly darker than apical) and occipital margin narrow and dark brown. Dorsal head with clypeus ovoid-rhomboid, rounded posteriorly, clypeal setae inserted near to lateral margins (
|
||
<figureCitation id="A3A5AC839B2A1E75C4E9FE9DFCEC3149" box="[754,837,333,360]" captionStart="FIGURE 6" captionStartId="15.[151,250,1892,1917]" captionTargetBox="[163,1417,176,1843]" captionTargetId="figure@15.[151,1436,166,1873]" captionTargetPageId="15" captionText="FIGURE 6. Riethia spp. Larva. Mid-dorsal head:A. R. donedwardi, sp.n., B. R. noongar sp.n.. Clypeus: C. R. larval sp. ‘C’, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. neocaledonica sp.n., I. R. noongar sp.n., J. R. paluma sp.n., K. R. phengari sp.n., L. R. plumosa Freeman, M. R. queenslandensis sp.n., N. R. stictoptera Kieffer, O. R. wazeylandica sp.n., P. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351070/files/figure.png" pageId="21" pageNumber="482">Fig. 6J</figureCitation>
|
||
). Antennal pedestal short. Antenna (n=1, not readily interpretable) with AR less than 1.0, segments 2-4 apparently subequal. Ventromental plate 15% longer than width of mentum. mandible with almost smooth mola, with only 1 very fine spine. Mensural features as in
|
||
<tableCitation id="761C85BD9B2A1E75C2E5FE45FAFD318E" box="[1278,1364,405,431]" captionStart="TABLE 2" captionStartId="32.[149,242,622,649]" captionTargetBox="[164,1976,660,1609]" captionText="TABLE 2. Morphometric data for larvae of Riethia Kieffer" httpUri="http://table.plazi.org/id/6FE1E08E9B1F1E40C68EFDBEFC8432A8" pageId="21" pageNumber="482" tableUuid="6FE1E08E9B1F1E40C68EFDBEFC8432A8">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B2A1E75C6DCFE69FBDF31D6" blockId="21.[151,1437,152,1187]" pageId="21" pageNumber="482">
|
||
<emphasis id="09EA6C149B2A1E75C6DCFE69FEE731F2" bold="true" box="[199,334,441,467]" pageId="21" pageNumber="482">Etymology.</emphasis>
|
||
From the
|
||
<typeStatus id="E4250EA49B2A1E75C7DDFE69FE5031F2" box="[454,505,441,467]" pageId="21" pageNumber="482">type</typeStatus>
|
||
, and so-far only known locality, Paluma, a high elevation well-studied location in the southernmost wet tropics of north
|
||
<collectingRegion id="F95A7EE49B2A1E75C45AFE0DFD6031D6" box="[577,713,477,503]" country="Australia" name="Queensland" pageId="21" pageNumber="482">Queensland</collectingRegion>
|
||
. To be treated as a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B2A1E75C6DCFDD0FEE632A6" blockId="21.[151,1437,152,1187]" pageId="21" pageNumber="482">
|
||
<emphasis id="09EA6C149B2A1E75C6DCFDD0FE93323A" bold="true" box="[199,314,512,539]" pageId="21" pageNumber="482">Diagnosis</emphasis>
|
||
. Although the male is unknown, matching to the female should show a broadly infuscate wing, with darker band across the wing at the cross veins, and with legs darkened at the ‘knees’—the distal apex of femur and proximal apex of the tibia. Although the female
|
||
<typeStatus id="E4250EA49B2A1E75C4B4FD99FCB83242" box="[687,785,585,611]" pageId="21" pageNumber="482" type="holotype">holotype</typeStatus>
|
||
is quite teneral, the colour pattern will be stronger in fully hardened adults.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B2A1E75C6DCFD41FDF33336" blockId="21.[151,1437,152,1187]" pageId="21" pageNumber="482">
|
||
The pupa has a unique combination of features, although each state is shared with one or more other species yet never in combination. The hookrow on TII is medially divided (otherwise seen only in
|
||
<taxonomicName id="FC9ECB859B2A1E75C289FD66FA9A32EE" authorityName="Cranston" authorityYear="2019" box="[1170,1331,693,719]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="482" phylum="Arthropoda" rank="species" species="hamodivisa">
|
||
<emphasis id="09EA6C149B2A1E75C289FD66FA9A32EE" box="[1170,1331,693,719]" italics="true" pageId="21" pageNumber="482">R. hamodivisa</emphasis>
|
||
</taxonomicName>
|
||
), and the conjunctival bands, present on tergites II and IV, also are medially divided (otherwise only in
|
||
<taxonomicName id="FC9ECB859B2A1E75C2A4FD0AFA9932D2" authorityName="Cranston" authorityYear="2019" box="[1215,1328,729,755]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="482" phylum="Arthropoda" rank="species" species="kakadu">
|
||
<emphasis id="09EA6C149B2A1E75C2A4FD0AFA9932D2" box="[1215,1328,729,755]" italics="true" pageId="21" pageNumber="482">R. kakadu</emphasis>
|
||
</taxonomicName>
|
||
). The essentially bare TII is unique in the genus.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B2A1E75C6DCFCF1FDA33386" blockId="21.[151,1437,152,1187]" pageId="21" pageNumber="482">
|
||
The larval head is uniformly pale, with mandibular and mental teeth golden brown. The antenna (only 1 of which is present, poorly laid out) can be determined to have a weak pedestal, a very low AR of 0.9, and ventromental plates 15% longer than the width of the mentum. The large clypeus (
|
||
<figureCitation id="A3A5AC839B2A1E75C580FCB9FC4233A5" box="[923,1003,873,900]" captionStart="FIGURE 6" captionStartId="15.[151,250,1892,1917]" captionTargetBox="[163,1417,176,1843]" captionTargetId="figure@15.[151,1436,166,1873]" captionTargetPageId="15" captionText="FIGURE 6. Riethia spp. Larva. Mid-dorsal head:A. R. donedwardi, sp.n., B. R. noongar sp.n.. Clypeus: C. R. larval sp. ‘C’, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. neocaledonica sp.n., I. R. noongar sp.n., J. R. paluma sp.n., K. R. phengari sp.n., L. R. plumosa Freeman, M. R. queenslandensis sp.n., N. R. stictoptera Kieffer, O. R. wazeylandica sp.n., P. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351070/files/figure.png" pageId="21" pageNumber="482">Fig. 6J</figureCitation>
|
||
), curved posteriorly with widely separated clypeal setae, is distinctive.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B2A1E75C6DCFC61FECC347A" blockId="21.[151,1437,152,1187]" pageId="21" pageNumber="482">
|
||
<emphasis id="09EA6C149B2A1E75C6DCFC61FE9D33EA" bold="true" box="[199,308,945,971]" pageId="21" pageNumber="482">Remarks</emphasis>
|
||
. Allocation of this taxon proved problematic, partially because of the lack of a male, but also because larval, pupal and female morphology do not converge on a single otherwise known species. The location, Birthday Creek near Paluma in north
|
||
<collectingRegion id="F95A7EE49B2A1E75C420FC29FD6B3432" box="[571,706,1017,1043]" country="Australia" name="Queensland" pageId="21" pageNumber="482">Queensland</collectingRegion>
|
||
, is well studied, and has three other
|
||
<taxonomicName id="FC9ECB859B2A1E75C242FC29FB033432" box="[1113,1194,1017,1043]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="482" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="09EA6C149B2A1E75C242FC29FB033432" box="[1113,1194,1017,1043]" italics="true" pageId="21" pageNumber="482">Riethia</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<emphasis id="09EA6C149B2A1E75C308FC2AFEC63416" italics="true" pageId="21" pageNumber="482">
|
||
<taxonomicName id="FC9ECB859B2A1E75C308FC2AFA313432" box="[1299,1432,1018,1043]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="482" phylum="Arthropoda" rank="species" species="cinctipes">R. cinctipes</taxonomicName>
|
||
,
|
||
<taxonomicName id="FC9ECB859B2A1E75C68CFBCEFEC63416" authorityName="Cranston" authorityYear="2019" box="[151,367,1053,1079]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="482" phylum="Arthropoda" rank="species" species="queenslandensis">R. queenslandensis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="FC9ECB859B2A1E75C7BEFBCEFD953416" box="[421,572,1054,1079]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="482" phylum="Arthropoda" rank="species" species="stictoptera">
|
||
<emphasis id="09EA6C149B2A1E75C7BEFBCEFD953416" box="[421,572,1054,1079]" italics="true" pageId="21" pageNumber="482">R. stictoptera</emphasis>
|
||
</taxonomicName>
|
||
). This species cannot be allocated in any stage to these (or indeed any other) species of
|
||
<taxonomicName id="FC9ECB859B2A1E75C716FB91FEF7347A" box="[269,350,1089,1115]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="482" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="09EA6C149B2A1E75C716FB91FEF7347A" box="[269,350,1089,1115]" italics="true" pageId="21" pageNumber="482">Riethia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B2A1E75C6DCFBB4FBE13482" blockId="21.[151,1437,152,1187]" pageId="21" pageNumber="482">
|
||
<emphasis id="09EA6C149B2A1E75C6DCFBB4FE56345E" bold="true" box="[199,511,1124,1151]" pageId="21" pageNumber="482">DIstribution and Ecology.</emphasis>
|
||
<taxonomicName id="FC9ECB859B2A1E75C41CFBB5FD1C345E" authorityName="Cranston" authorityYear="2019" box="[519,693,1125,1151]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="482" phylum="Arthropoda" rank="species" species="paluma">
|
||
<emphasis id="09EA6C149B2A1E75C41CFBB5FD1C345E" box="[519,693,1125,1151]" italics="true" pageId="21" pageNumber="482">Riethia paluma</emphasis>
|
||
</taxonomicName>
|
||
is known only from a single reared specimen from well-sampled creeks in the mountains of the southern wet tropics immediately north of Townsville
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |