270 lines
36 KiB
XML
270 lines
36 KiB
XML
<document id="0D2425A66C0808FD5C95D25BFF84A263" ID-CLB-Dataset="25594" ID-DOI="10.11646/zootaxa.4646.3.3" ID-GBIF-Dataset="4a3d31ca-e16b-4ccf-8407-231aa27702f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3351058" ID-ZooBank="25CA4A18-955F-4EA7-8978-E93032B54A2E" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564032263204" checkinUser="plazi" docAuthor="Cranston, Peter S." docDate="2019" docId="B33701109B2A1E76C68CFB25FD1F3483" docLanguage="en" docName="zootaxao.4646.3.3.pdf" docOrigin="Zootaxa 4646 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Riethia phengari Cranston 2019, sp.n." docType="treatment" docVersion="12" lastPageNumber="483" masterDocId="4F0E79689B3F1E60C61BFFD0FFA93021" masterDocTitle="Riethia (Kieffer 1917) (Diptera: Chironomidae) revised for the Austro-Pacific region" masterLastPageNumber="500" masterPageNumber="461" pageNumber="482" updateTime="1698747359191" updateUser="ExternalLinkService">
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<mods:title id="53B4D6B0F7FFB0C086023D38D4342390">Riethia (Kieffer 1917) (Diptera: Chironomidae) revised for the Austro-Pacific region</mods:title>
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<mods:namePart id="6D9BE15BF1E87E625BF827A0B16460FF">Cranston, Peter S.</mods:namePart>
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<mods:title id="86E97A78D754BF368E2AB45126247872">Zootaxa</mods:title>
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<mods:part id="28F27EFFDC0A9ED02F36BD804A430E1A">
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<mods:date id="A3390B058C759BC9A2671C382A9A7D2E">2019</mods:date>
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<mods:number id="12923296BAA4901C1D39C1534A7991F5">2019-07-25</mods:number>
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<treatment id="B33701109B2A1E76C68CFB25FD1F3483" ID-DOI="http://doi.org/10.5281/zenodo.5587168" ID-GBIF-Taxon="159413594" ID-Zenodo-Dep="5587168" LSID="urn:lsid:plazi:treatment:B33701109B2A1E76C68CFB25FD1F3483" httpUri="http://treatment.plazi.org/id/B33701109B2A1E76C68CFB25FD1F3483" lastPageId="22" lastPageNumber="483" pageId="21" pageNumber="482">
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<subSubSection id="7384E38D9B2A1E75C68CFB25FD8A352E" box="[151,547,1269,1296]" pageId="21" pageNumber="482" type="nomenclature">
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<paragraph id="3B21B0069B2A1E75C68CFB25FD8A352E" blockId="21.[151,547,1269,1367]" box="[151,547,1269,1296]" pageId="21" pageNumber="482">
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<heading id="6069076A9B2A1E75C68CFB25FD8A352E" bold="true" box="[151,547,1269,1296]" fontSize="11" level="1" pageId="21" pageNumber="482" reason="1">
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<emphasis id="09EA6C149B2A1E75C68CFB25FD8A352E" bold="true" box="[151,547,1269,1296]" pageId="21" pageNumber="482">
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<taxonomicName id="FC9ECB859B2A1E75C68CFB25FE49352E" ID-CoL="9HZ6T" authority="Cranston" authorityName="Cranston" authorityYear="2019" box="[151,480,1269,1296]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="482" phylum="Arthropoda" rank="species" species="phengari" status="sp. nov.">
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<emphasis id="09EA6C149B2A1E75C68CFB25FECD3531" bold="true" box="[151,356,1269,1296]" italics="true" pageId="21" pageNumber="482">Riethia phengari</emphasis>
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Cranston
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</taxonomicName>
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<taxonomicNameLabel id="12D9D16F9B2A1E75C7F3FB25FD8A352E" box="[488,547,1269,1295]" pageId="21" pageNumber="482" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="7384E38D9B2A1E75C68CFAC9FDE335B8" pageId="21" pageNumber="482" type="description">
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<paragraph id="3B21B0069B2A1E75C68CFAC9FE393515" blockId="21.[151,547,1269,1367]" box="[151,400,1305,1332]" pageId="21" pageNumber="482">
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(
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<figureCitation id="A3A5AC839B2A1E75C684FAC9FEA83512" box="[159,257,1305,1331]" captionStart="FIGURE 1" captionStartId="10.[151,250,1970,1995]" captionTargetBox="[237,1393,192,1878]" captionTargetId="figure@10.[178,1410,166,1950]" captionTargetPageId="10" captionText="FIGURE 1. Riethia spp. Male hypopygia L. side dorsal, R. side, ventral/internal, stylised: A. R. donedwardi sp.n., B. R. hamodivisa sp.n., C. R. phengari, sp.n. D. R. plumosa Freeman, E. R. stictoptera Keffer, F. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351060/files/figure.png" pageId="21" pageNumber="482">Figs. 1C</figureCitation>
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,
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<figureCitation id="A3A5AC839B2A1E75C716FAC9FE9B3512" box="[269,306,1305,1331]" captionStart="FIGURE 2" captionStartId="11.[151,250,1925,1950]" captionTargetBox="[177,1375,200,1857]" captionTargetId="figure@11.[160,1427,166,1905]" captionTargetPageId="11" captionText="FIGURE 2. Riethia spp. Volsellae: A. R. azeylandica sp.n., B. R. cinctipes Freeman, C. R. donedwardi sp.n. D. R. hamodivisa sp.n., E. R. kakadu sp.n., F. R. neocaledonica sp.n., G. R. noongar sp.n., H. R. phengari sp.n., I. R. plumosa Freeman, J. R. queenslandensis sp.n., K. R. stictoptera Keffer, L. R. wazeylandica sp.n., M. R. zeylandica sp.n." httpUri="https://zenodo.org/record/3351062/files/figure.png" pageId="21" pageNumber="482">2H</figureCitation>
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,
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<figureCitation id="A3A5AC839B2A1E75C725FAC9FEF03512" box="[318,345,1305,1331]" captionStart="FIGURE 4" captionStartId="13.[151,250,1897,1922]" captionTargetBox="[199,1399,178,1821]" captionTargetId="figure@13.[175,1412,166,1877]" captionTargetPageId="13" captionText="FIGURE 4. Riethia spp. Pupa. Abdominal tergites: A. R. noongar sp.n., B. R. paluma sp.n., C. R. plumosa Freeman, D. R. stictoptera Kieffer. Posterolateral corner of VIII (‘comb’): E. R. noongar, F. R. paluma, G. R. plumosa, H. R. stictoptera I. R. cinctipes (var), J. R. phengari sp.n., K. R. neocaledonica sp.n.. Tergites: L, IV-V of R. neocaledonica; M. Tergites VI-VIII of R. queenslandensis sp.n., N. Sternites VI-VIII R. queenslandensis, O. Tergites V-AL of R. zeylandica." httpUri="https://zenodo.org/record/3351066/files/figure.png" pageId="21" pageNumber="482">4J</figureCitation>
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,
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<figureCitation id="A3A5AC839B2A1E75C77EFAC9FE213515" box="[357,392,1305,1332]" captionStart="FIGURE 6" captionStartId="15.[151,250,1892,1917]" captionTargetBox="[163,1417,176,1843]" captionTargetId="figure@15.[151,1436,166,1873]" captionTargetPageId="15" captionText="FIGURE 6. Riethia spp. Larva. Mid-dorsal head:A. R. donedwardi, sp.n., B. R. noongar sp.n.. Clypeus: C. R. larval sp. ‘C’, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. neocaledonica sp.n., I. R. noongar sp.n., J. R. paluma sp.n., K. R. phengari sp.n., L. R. plumosa Freeman, M. R. queenslandensis sp.n., N. R. stictoptera Kieffer, O. R. wazeylandica sp.n., P. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351070/files/figure.png" pageId="21" pageNumber="482">6K</figureCitation>
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)
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</paragraph>
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<paragraph id="3B21B0069B2A1E75C68CFAEDFE193576" blockId="21.[151,547,1269,1367]" box="[151,432,1341,1367]" pageId="21" pageNumber="482">urn:lsid:zoobank.org:act:</paragraph>
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<paragraph id="3B21B0069B2A1E75C68CFA51FDE335B8" blockId="21.[151,586,1408,1433]" box="[151,586,1408,1433]" pageId="21" pageNumber="482">
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<emphasis id="09EA6C149B2A1E75C68CFA51FEC035B8" box="[151,361,1409,1433]" italics="true" pageId="21" pageNumber="482">
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<taxonomicName id="FC9ECB859B2A1E75C68CFA51FF4935B8" box="[151,224,1409,1433]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="482" phylum="Arthropoda" rank="genus">Riethia</taxonomicName>
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||
“moonlight”
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||
</emphasis>
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||
Cranston, 1966, 2000
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="7384E38D9B2A1E76C68CFA18FD1F3483" lastPageId="22" lastPageNumber="483" pageId="21" pageNumber="482" type="materials_examined">
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<paragraph id="3B21B0069B2A1E75C68CFA18FD88366F" blockId="21.[151,1437,1479,2010]" pageId="21" pageNumber="482">
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<emphasis id="09EA6C149B2A1E75C68CFA18FEE935C3" bold="true" box="[151,320,1479,1506]" pageId="21" pageNumber="482">Type material</emphasis>
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.
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<materialsCitation id="8BF6BA5B9B2A1E75C74AFA18FE403627" ID-GBIF-Occurrence="2382714494" collectingDate="1991-10-20" collectionCode="ACT, ANIC" latitude="-35.416668" location="Moonlight Hollow" longLatPrecision="925" longitude="148.78833" municipality="Le" pageId="21" pageNumber="482" specimenCount="1" specimenCount-male="1" stateProvince="Australian Capital Territory" typeStatus="holotype">
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<emphasis id="09EA6C149B2A1E75C74AFA18FE1735C3" bold="true" box="[337,446,1480,1506]" pageId="21" pageNumber="482">Holotype</emphasis>
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</typeStatus>
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||
<collectingMunicipality id="DB452A7C9B2A1E75C7DCFA18FE4E35C3" box="[455,487,1480,1506]" pageId="21" pageNumber="482">Le</collectingMunicipality>
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/P
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||
<specimenCount id="2D987B8F9B2A1E75C41BFA18FDBA35C3" box="[512,531,1480,1506]" pageId="21" pageNumber="482" type="male">♂</specimenCount>
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||
,
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||
<collectingRegion id="F95A7EE49B2A1E75C43BFA17FCF335C0" box="[544,858,1479,1505]" country="Australia" name="Australian Capital Territory" pageId="21" pageNumber="482">Australian Capital Territory</collectingRegion>
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(
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<collectionCode id="5D8F28C39B2A1E75C577FA18FC0035C3" box="[876,937,1480,1506]" pageId="21" pageNumber="482">ACT</collectionCode>
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),
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<location id="3E41E6DD9B2A1E75C5A4FA17FB3D35C0" LSID="urn:lsid:plazi:treatment:B33701109B2A1E76C68CFB25FD1F3483:3E41E6DD9B2A1E75C5A4FA17FB3D35C0" box="[959,1172,1479,1505]" latitude="-35.416668" longLatPrecision="925" longitude="148.78833" municipality="Le" name="Moonlight Hollow" pageId="21" pageNumber="482" stateProvince="Australian Capital Territory">Moonlight Hollow</location>
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,
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<geoCoordinate id="5EAAD6C19B2A1E75C31CFA18FA3135C3" box="[1287,1432,1480,1506]" degrees="148" direction="east" minutes="47.30" orientation="longitude" pageId="21" pageNumber="482" precision="9" value="148.78833">148°47.30’E</geoCoordinate>
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,
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<collectingDate id="5F646F2E9B2A1E75C68CFA3CFEAE3627" box="[151,263,1516,1542]" pageId="21" pageNumber="482" value="1991-10-20">
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<date id="4F2096C69B2A1E75C68CFA3CFEAE3627" box="[151,263,1516,1542]" pageId="21" pageNumber="482" value="1991-10-20">20.x.1991</date>
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</collectingDate>
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(
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<emphasis id="09EA6C149B2A1E75C70CFA3CFED63627" box="[279,383,1516,1542]" italics="true" pageId="21" pageNumber="482">Cranston</emphasis>
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) (
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<collectionCode id="5D8F28C39B2A1E75C78CFA3CFE4B3627" box="[407,482,1516,1542]" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="482">ANIC</collectionCode>
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)
|
||
</materialsCitation>
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||
.
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||
<materialsCitation id="8BF6BA5B9B2A1E75C7E8FA3CFCEB360B" ID-GBIF-Occurrence="2382714530" collectingDate="1988-08-24" collectingDateMax="1988-08-27" collectingDateMin="1988-08-24" collectionCode="ANIC" collectorName="Le & Pe & Brindabellas & Blundell's Ck." latitude="-35.366665" longLatPrecision="1193" longitude="148.83333" pageId="21" pageNumber="482" specimenCount="1" specimenCount-male="1" stateProvince="Australian Capital Territory" typeStatus="paratype">
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<typeStatus id="E4250EA49B2A1E75C7E8FA3CFDC93627" box="[499,608,1516,1542]" pageId="21" pageNumber="482" type="paratype">Paratypes</typeStatus>
|
||
(all collected
|
||
<emphasis id="09EA6C149B2A1E75C4E6FA3CFCCC3627" box="[765,869,1516,1542]" italics="true" pageId="21" pageNumber="482">Cranston</emphasis>
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||
, deposited
|
||
<collectionCode id="5D8F28C39B2A1E75C5F9FA3CFB853627" box="[994,1068,1516,1542]" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="482">ANIC</collectionCode>
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||
):
|
||
<collectorName id="966BD5D09B2A1E75C225FA3CFBF53627" box="[1086,1116,1516,1542]" pageId="21" pageNumber="482">Le</collectorName>
|
||
/
|
||
<collectorName id="966BD5D09B2A1E75C27EFA3CFB2B3627" box="[1125,1154,1516,1542]" pageId="21" pageNumber="482">Pe</collectorName>
|
||
, as
|
||
<typeStatus id="E4250EA49B2A1E75C2B1FA3BFAB93624" box="[1194,1296,1515,1541]" pageId="21" pageNumber="482" type="holotype">holotype</typeStatus>
|
||
; 2L,
|
||
<collectorName id="966BD5D09B2A1E75C35CFA3BFF723608" pageId="21" pageNumber="482">Brindabellas</collectorName>
|
||
,
|
||
<collectorName id="966BD5D09B2A1E75C6FCF9DFFE203608" box="[231,393,1551,1577]" pageId="21" pageNumber="482">Blundell's Ck.</collectorName>
|
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,
|
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<geoCoordinate id="5EAAD6C19B2A1E75C78EF9DFFE44360B" box="[405,493,1551,1578]" degrees="35" direction="south" minutes="22" orientation="latitude" pageId="21" pageNumber="482" precision="925" value="-35.366665">35°22'S</geoCoordinate>
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<geoCoordinate id="5EAAD6C19B2A1E75C7EFF9DFFDF7360B" box="[500,606,1551,1578]" degrees="148" direction="east" minutes="50" orientation="longitude" pageId="21" pageNumber="482" precision="925" value="148.83333">148°50'E</geoCoordinate>
|
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,
|
||
<collectingDate id="5F646F2E9B2A1E75C471F9DFFCB63608" box="[618,799,1551,1577]" pageId="21" pageNumber="482" value="1988-08-24" valueMax="1988-08-27" valueMin="1988-08-24">
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<date id="4F2096C69B2A1E75C471F9DFFCB63608" box="[618,799,1551,1577]" pageId="21" pageNumber="482" value="1988-08-24" valueMax="1988-08-27" valueMin="1988-08-24">24–27.viii.1988</date>
|
||
</collectingDate>
|
||
;
|
||
<specimenCount id="2D987B8F9B2A1E75C530F9C0FCEB360B" box="[811,834,1552,1578]" pageId="21" pageNumber="482" type="male">♂</specimenCount>
|
||
</materialsCitation>
|
||
, Blundell's Ck., no further data (
|
||
<emphasis id="09EA6C149B2A1E75C2A8F9C0FAAC360B" box="[1203,1285,1552,1578]" italics="true" pageId="21" pageNumber="482">Colless</emphasis>
|
||
); Pe, Condor Ck.,
|
||
<geoCoordinate id="5EAAD6C19B2A1E75C6D6F9E3FE80366F" box="[205,297,1587,1614]" degrees="35" direction="south" minutes="22" orientation="latitude" pageId="21" pageNumber="482" precision="925" value="-35.366665">35°22’S</geoCoordinate>
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<geoCoordinate id="5EAAD6C19B2A1E75C72BF9E3FE37366F" box="[304,414,1587,1614]" degrees="148" direction="east" minutes="51" orientation="longitude" pageId="21" pageNumber="482" precision="925" value="148.85">148°51’E</geoCoordinate>
|
||
,
|
||
<date id="4F2096C69B2A1E75C7B1F9E4FDB4366F" box="[426,541,1588,1614]" pageId="21" pageNumber="482" value="1991-10-27">27.x.1991</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B2A1E75C6DCF987FD5636B7" blockId="21.[151,1437,1479,2010]" pageId="21" pageNumber="482">
|
||
<materialsCitation id="8BF6BA5B9B2A1E75C6DCF987FD5236B7" ID-GBIF-Occurrence="2382714497" collectionCode="MV" collectorName="South George R." location="South George" pageId="21" pageNumber="482" specimenCount="2" stateProvince="Tasmania" typeStatus="paratype">
|
||
<emphasis id="09EA6C149B2A1E75C6DCF987FED43653" bold="true" box="[199,381,1623,1650]" pageId="21" pageNumber="482">Other material</emphasis>
|
||
. Unreared larva, excluded from
|
||
<typeStatus id="E4250EA49B2A1E75C4E3F988FCF13653" box="[760,856,1624,1650]" pageId="21" pageNumber="482" type="paratype">paratype</typeStatus>
|
||
series.
|
||
<collectingRegion id="F95A7EE49B2A1E75C5A9F987FB893650" box="[946,1056,1623,1649]" country="Australia" name="Tasmania" pageId="21" pageNumber="482">Tasmania</collectingRegion>
|
||
,
|
||
<collectorName id="966BD5D09B2A1E75C235F987FB593653" box="[1070,1264,1623,1650]" pageId="21" pageNumber="482">
|
||
<location id="3E41E6DD9B2A1E75C235F987FB623653" LSID="urn:lsid:plazi:treatment:B33701109B2A1E76C68CFB25FD1F3483:3E41E6DD9B2A1E75C235F987FB623653" box="[1070,1227,1623,1650]" name="South George" pageId="21" pageNumber="482" stateProvince="Tasmania">South George</location>
|
||
R.
|
||
</collectorName>
|
||
, intake bridge, undated, 41°18.39”S 147°56.02”E (
|
||
<collectionCode id="5D8F28C39B2A1E75C43CF9ACFDFC36B7" box="[551,597,1660,1686]" country="0" name="University of Montana Museum" pageId="21" pageNumber="482">MV</collectionCode>
|
||
, failed, TAS3)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B2A1E75C6DCF94FFD2F37FB" blockId="21.[151,1437,1479,2010]" pageId="21" pageNumber="482">
|
||
<emphasis id="09EA6C149B2A1E75C6DCF94FFEF8369B" bold="true" box="[199,337,1695,1722]" pageId="21" pageNumber="482">Description</emphasis>
|
||
.
|
||
<emphasis id="09EA6C149B2A1E75C77BF970FE34369B" bold="true" box="[352,413,1696,1722]" pageId="21" pageNumber="482">Male</emphasis>
|
||
. Thorax yellow-brown without distinct vittae; legs yellow-with slightly darker tarsomeres, without banding. Wing pale, unmarked, with yellow veins. TIX with fine setae dense across complete tergite. Gonostylus broad at junction with gonocoxite, abruptly attenuated, bearing dense brush of broad, apically plumose setae on inner margin (
|
||
<figureCitation id="A3A5AC839B2A1E75C743F8DBFE063707" box="[344,431,1803,1830]" captionStart="FIGURE 1" captionStartId="10.[151,250,1970,1995]" captionTargetBox="[237,1393,192,1878]" captionTargetId="figure@10.[178,1410,166,1950]" captionTargetPageId="10" captionText="FIGURE 1. Riethia spp. Male hypopygia L. side dorsal, R. side, ventral/internal, stylised: A. R. donedwardi sp.n., B. R. hamodivisa sp.n., C. R. phengari, sp.n. D. R. plumosa Freeman, E. R. stictoptera Keffer, F. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351060/files/figure.png" pageId="21" pageNumber="482">Fig. 1C</figureCitation>
|
||
). Superior volsella with basal area bearing proximally 4 long setae, remainder bare except for posterior setose projection; medially-directed digitus virtually devoid of microtrichia, with 3 long setae; inferior volsella very small, almost totally appressed to inner contour of gonocoxite and linked to posterior part of superior volsella, with long simple setae; junction between superior volsella and inferior volsella bears 1 pectinate scale (
|
||
<figureCitation id="A3A5AC839B2A1E75C374F8A7FF143797" captionStart="FIGURE 2" captionStartId="11.[151,250,1925,1950]" captionTargetBox="[177,1375,200,1857]" captionTargetId="figure@11.[160,1427,166,1905]" captionTargetPageId="11" captionText="FIGURE 2. Riethia spp. Volsellae: A. R. azeylandica sp.n., B. R. cinctipes Freeman, C. R. donedwardi sp.n. D. R. hamodivisa sp.n., E. R. kakadu sp.n., F. R. neocaledonica sp.n., G. R. noongar sp.n., H. R. phengari sp.n., I. R. plumosa Freeman, J. R. queenslandensis sp.n., K. R. stictoptera Keffer, L. R. wazeylandica sp.n., M. R. zeylandica sp.n." httpUri="https://zenodo.org/record/3351062/files/figure.png" pageId="21" pageNumber="482">Fig. 2H</figureCitation>
|
||
). No pseudovolsella, inner gonocoxal setae aligned, arising from weak tubercles, more dorsal than internal on gonocoxite. Mensural features as in
|
||
<tableCitation id="761C85BD9B2A1E75C436F86FFD2937FB" box="[557,640,1983,2010]" captionStart="TABLE 1" captionStartId="30.[149,242,517,544]" captionTargetBox="[164,1998,555,1659]" captionText="TABLE 1. Morphometric data for males of Riethia Kieffer" httpUri="http://table.plazi.org/id/6FE1E08E9B211E7EC68EFDD5FC833201" pageId="21" pageNumber="482" tableUuid="6FE1E08E9B211E7EC68EFDD5FC833201">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B291E76C6DCFF49FE3C3093" blockId="22.[151,1437,152,1187]" box="[199,405,152,179]" pageId="22" pageNumber="483">
|
||
<emphasis id="09EA6C149B291E76C6DCFF49FEB73092" bold="true" box="[199,286,153,179]" pageId="22" pageNumber="483">Female</emphasis>
|
||
unknown.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B291E76C6DCFF6DFDA831D7" blockId="22.[151,1437,152,1187]" pageId="22" pageNumber="483">
|
||
<emphasis id="09EA6C149B291E76C6DCFF6DFEAC30F6" bold="true" box="[199,261,189,215]" pageId="22" pageNumber="483">Pupa</emphasis>
|
||
. Very pale, almost translucent, with pale yellow dorsal thorax, anterior wing sheaths and lateral apophyses on abdominal segments V–VIII; comb pale yellow. Cephalothorax with almost smooth frons, thorax weakly rugulose with bi-triserial row of small tubercles along dorsal thorax each side of eclosion line. Hook row continuous on II, 45–50 % width of tergite; conjunctival spinule band on III and IV continuous. Pedes spurii B absent, vortex weak. Abdomen with tergite II with armament full width of hookrow posteriorly, tapering anteriorly and then expanded to form quite strong anterior transverse band of stronger spines; TIII–V similar to
|
||
<taxonomicName id="FC9ECB859B291E76C22CFEA1FB7531AB" authorityName="Cranston" authorityYear="2019" box="[1079,1244,368,394]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="483" phylum="Arthropoda" rank="species" species="azeylandica">
|
||
<emphasis id="09EA6C149B291E76C22CFEA1FB7531AB" box="[1079,1244,368,394]" italics="true" pageId="22" pageNumber="483">R. azeylandica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="A3A5AC839B291E76C2F7FEA0FAEE31AA" box="[1260,1351,368,395]" captionStart="FIGURE 3" captionStartId="12.[151,250,1862,1887]" captionTargetBox="[151,1400,192,1819]" captionTargetId="figure@12.[151,1436,166,1842]" captionTargetPageId="12" captionText="FIGURE 3. Riethia spp. Pupa. Cephalic tubules: A. R. azeylandica sp.n., B. R. noongar sp.n., C. Dorsal thorax R. kakadu sp.n.. Abdominal tergites: D. R. azeylandica, E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. kakadu. Posterolateral corner of VIII (‘comb’): I. R. azeylandica sp.n., J. R. cinctipes, K. R. donedwardi, L. R. hamodivisa, M. R. kakadu." httpUri="https://zenodo.org/record/3351064/files/figure.png" pageId="22" pageNumber="483">Fig. 3D</figureCitation>
|
||
). TVIII and anal lobe bare. Taeniate setae V–VIII 3, 4, 4, 5, on VIII evenly distributed along segment. Comb with 1 strong postero-laterally directed tooth (
|
||
<figureCitation id="A3A5AC839B291E76C41FFE68FDFC31F2" box="[516,597,440,467]" captionStart="FIGURE 4" captionStartId="13.[151,250,1897,1922]" captionTargetBox="[199,1399,178,1821]" captionTargetId="figure@13.[175,1412,166,1877]" captionTargetPageId="13" captionText="FIGURE 4. Riethia spp. Pupa. Abdominal tergites: A. R. noongar sp.n., B. R. paluma sp.n., C. R. plumosa Freeman, D. R. stictoptera Kieffer. Posterolateral corner of VIII (‘comb’): E. R. noongar, F. R. paluma, G. R. plumosa, H. R. stictoptera I. R. cinctipes (var), J. R. phengari sp.n., K. R. neocaledonica sp.n.. Tergites: L, IV-V of R. neocaledonica; M. Tergites VI-VIII of R. queenslandensis sp.n., N. Sternites VI-VIII R. queenslandensis, O. Tergites V-AL of R. zeylandica." httpUri="https://zenodo.org/record/3351066/files/figure.png" pageId="22" pageNumber="483">Fig. 4J</figureCitation>
|
||
), with or without 1–2 very small lateral teeth. Anal lobe with c 55–60 bidisorganised multiserial taeniae.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B291E76C6DCFDD1FE55328A" blockId="22.[151,1437,152,1187]" pageId="22" pageNumber="483">
|
||
<emphasis id="09EA6C149B291E76C6DCFDD1FEBE323A" bold="true" box="[199,279,513,539]" pageId="22" pageNumber="483">Larva.</emphasis>
|
||
Head capsule golden, slightly browner on postmentum, occipital margin broad, dark brown, mentum and four inner mandibular teeth similarly golden-brown. Clypeus (
|
||
<figureCitation id="A3A5AC839B291E76C59DFDF4FC77321E" box="[902,990,548,575]" captionStart="FIGURE 6" captionStartId="15.[151,250,1892,1917]" captionTargetBox="[163,1417,176,1843]" captionTargetId="figure@15.[151,1436,166,1873]" captionTargetPageId="15" captionText="FIGURE 6. Riethia spp. Larva. Mid-dorsal head:A. R. donedwardi, sp.n., B. R. noongar sp.n.. Clypeus: C. R. larval sp. ‘C’, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. neocaledonica sp.n., I. R. noongar sp.n., J. R. paluma sp.n., K. R. phengari sp.n., L. R. plumosa Freeman, M. R. queenslandensis sp.n., N. R. stictoptera Kieffer, O. R. wazeylandica sp.n., P. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351070/files/figure.png" pageId="22" pageNumber="483">Fig. 6K</figureCitation>
|
||
) trapezoid, stippled at least in posterior half. Inner margin of mandible with 2–3 strong serrations on mola commencing close to insertion of seta subdentalis. Each antennal segment shorter than preceding. The ventromental plates are 10–20% wider than the mentum. Mensural features as in
|
||
<tableCitation id="761C85BD9B291E76C7B9FD40FE51328A" box="[418,504,656,683]" captionStart="TABLE 2" captionStartId="32.[149,242,622,649]" captionTargetBox="[164,1976,660,1609]" captionText="TABLE 2. Morphometric data for larvae of Riethia Kieffer" httpUri="http://table.plazi.org/id/6FE1E08E9B1F1E40C68EFDBEFC8432A8" pageId="22" pageNumber="483" tableUuid="6FE1E08E9B1F1E40C68EFDBEFC8432A8">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B291E76C6DCFD65FAF632EF" blockId="22.[151,1437,152,1187]" box="[199,1375,692,719]" pageId="22" pageNumber="483">
|
||
<emphasis id="09EA6C149B291E76C6DCFD65FE5432EE" bold="true" box="[199,509,692,719]" pageId="22" pageNumber="483">Etymology. From φεγγάρι</emphasis>
|
||
(phengári) Greek for moonlight, after the
|
||
<typeStatus id="E4250EA49B291E76C5CCFD65FBAE32EE" box="[983,1031,693,719]" pageId="22" pageNumber="483">type</typeStatus>
|
||
locality of Moonlight Hollow.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B291E76C6DCFD08FD173387" blockId="22.[151,1437,152,1187]" pageId="22" pageNumber="483">
|
||
<emphasis id="09EA6C149B291E76C6DCFD08FEEB32D2" bold="true" box="[199,322,728,755]" pageId="22" pageNumber="483">Diagnosis.</emphasis>
|
||
In the male the attenuated gonostylus (
|
||
<figureCitation id="A3A5AC839B291E76C4E2FD08FCE732D2" box="[761,846,728,755]" captionStart="FIGURE 1" captionStartId="10.[151,250,1970,1995]" captionTargetBox="[237,1393,192,1878]" captionTargetId="figure@10.[178,1410,166,1950]" captionTargetPageId="10" captionText="FIGURE 1. Riethia spp. Male hypopygia L. side dorsal, R. side, ventral/internal, stylised: A. R. donedwardi sp.n., B. R. hamodivisa sp.n., C. R. phengari, sp.n. D. R. plumosa Freeman, E. R. stictoptera Keffer, F. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351060/files/figure.png" pageId="22" pageNumber="483">Fig. 1C</figureCitation>
|
||
) is unique. The pupa of
|
||
<taxonomicName id="FC9ECB859B291E76C27BFD09FB4D32D3" authorityName="Cranston" authorityYear="2019" box="[1120,1252,728,754]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="483" phylum="Arthropoda" rank="species" species="phengari">
|
||
<emphasis id="09EA6C149B291E76C27BFD09FB4D32D3" box="[1120,1252,728,754]" italics="true" pageId="22" pageNumber="483">R. phengari</emphasis>
|
||
</taxonomicName>
|
||
is near identical to that of
|
||
<taxonomicName id="FC9ECB859B291E76C713FD2DFE073337" authorityName="Cranston" authorityYear="2019" box="[264,430,764,790]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="483" phylum="Arthropoda" rank="species" species="azeylandica">
|
||
<emphasis id="09EA6C149B291E76C713FD2DFE073337" box="[264,430,764,790]" italics="true" pageId="22" pageNumber="483">R. azeylandica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="A3A5AC839B291E76C7A4FD2CFDA53336" box="[447,524,764,791]" captionStart="FIGURE 3" captionStartId="12.[151,250,1862,1887]" captionTargetBox="[151,1400,192,1819]" captionTargetId="figure@12.[151,1436,166,1842]" captionTargetPageId="12" captionText="FIGURE 3. Riethia spp. Pupa. Cephalic tubules: A. R. azeylandica sp.n., B. R. noongar sp.n., C. Dorsal thorax R. kakadu sp.n.. Abdominal tergites: D. R. azeylandica, E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. kakadu. Posterolateral corner of VIII (‘comb’): I. R. azeylandica sp.n., J. R. cinctipes, K. R. donedwardi, L. R. hamodivisa, M. R. kakadu." httpUri="https://zenodo.org/record/3351064/files/figure.png" pageId="22" pageNumber="483">
|
||
Fig.
|
||
<date id="4F2096C69B291E76C7EEFD2DFDA53336" box="[501,524,765,791]" pageId="22" pageNumber="483">3I</date>
|
||
</figureCitation>
|
||
) perhaps more nearly transparent without darkening and with fewer teeth in the ‘comb’ (
|
||
<figureCitation id="A3A5AC839B291E76C6EEFCF0FED0331A" box="[245,377,800,827]" captionStart="FIGURE 4" captionStartId="13.[151,250,1897,1922]" captionTargetBox="[199,1399,178,1821]" captionTargetId="figure@13.[175,1412,166,1877]" captionTargetPageId="13" captionText="FIGURE 4. Riethia spp. Pupa. Abdominal tergites: A. R. noongar sp.n., B. R. paluma sp.n., C. R. plumosa Freeman, D. R. stictoptera Kieffer. Posterolateral corner of VIII (‘comb’): E. R. noongar, F. R. paluma, G. R. plumosa, H. R. stictoptera I. R. cinctipes (var), J. R. phengari sp.n., K. R. neocaledonica sp.n.. Tergites: L, IV-V of R. neocaledonica; M. Tergites VI-VIII of R. queenslandensis sp.n., N. Sternites VI-VIII R. queenslandensis, O. Tergites V-AL of R. zeylandica." httpUri="https://zenodo.org/record/3351066/files/figure.png" pageId="22" pageNumber="483">
|
||
Fig 4J v.
|
||
<date id="4F2096C69B291E76C745FCF1FED0331A" box="[350,377,801,827]" pageId="22" pageNumber="483">4I</date>
|
||
</figureCitation>
|
||
). The larva of
|
||
<taxonomicName id="FC9ECB859B291E76C405FCF1FD0A331B" authorityName="Cranston" authorityYear="2019" box="[542,675,800,826]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="483" phylum="Arthropoda" rank="species" species="phengari">
|
||
<emphasis id="09EA6C149B291E76C405FCF1FD0A331B" box="[542,675,800,826]" italics="true" pageId="22" pageNumber="483">R. phengari</emphasis>
|
||
</taxonomicName>
|
||
has a pale mentum and inner mandibular teeth scarcely darker than the uniformly pale-golden head. The antennal pedestal is very short and the AR is between 1.6 and 1.8 with the 3
|
||
<superScript id="CCEB1D4E9B291E76C394FC93FA343370" attach="none" box="[1423,1437,835,849]" fontSize="6" pageId="22" pageNumber="483">rd</superScript>
|
||
segment shorter than 2
|
||
<superScript id="CCEB1D4E9B291E76C78FFCB7FE0D3354" attach="none" box="[404,420,871,885]" fontSize="6" pageId="22" pageNumber="483">nd</superScript>
|
||
. The clypeus (
|
||
<figureCitation id="A3A5AC839B291E76C453FCB8FD0933A2" box="[584,672,872,899]" captionStart="FIGURE 6" captionStartId="15.[151,250,1892,1917]" captionTargetBox="[163,1417,176,1843]" captionTargetId="figure@15.[151,1436,166,1873]" captionTargetPageId="15" captionText="FIGURE 6. Riethia spp. Larva. Mid-dorsal head:A. R. donedwardi, sp.n., B. R. noongar sp.n.. Clypeus: C. R. larval sp. ‘C’, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. neocaledonica sp.n., I. R. noongar sp.n., J. R. paluma sp.n., K. R. phengari sp.n., L. R. plumosa Freeman, M. R. queenslandensis sp.n., N. R. stictoptera Kieffer, O. R. wazeylandica sp.n., P. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351070/files/figure.png" pageId="22" pageNumber="483">Fig. 6K</figureCitation>
|
||
) resembles that of
|
||
<taxonomicName id="FC9ECB859B291E76C568FCB9FBB133A3" authorityName="Cranston" authorityYear="2019" box="[883,1048,872,898]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="483" phylum="Arthropoda" rank="species" species="azeylandica">
|
||
<emphasis id="09EA6C149B291E76C568FCB9FBB133A3" box="[883,1048,872,898]" italics="true" pageId="22" pageNumber="483">R. azeylandica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="A3A5AC839B291E76C23DFCB8FBD433A2" box="[1062,1149,872,899]" captionStart="FIGURE 6" captionStartId="15.[151,250,1892,1917]" captionTargetBox="[163,1417,176,1843]" captionTargetId="figure@15.[151,1436,166,1873]" captionTargetPageId="15" captionText="FIGURE 6. Riethia spp. Larva. Mid-dorsal head:A. R. donedwardi, sp.n., B. R. noongar sp.n.. Clypeus: C. R. larval sp. ‘C’, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. neocaledonica sp.n., I. R. noongar sp.n., J. R. paluma sp.n., K. R. phengari sp.n., L. R. plumosa Freeman, M. R. queenslandensis sp.n., N. R. stictoptera Kieffer, O. R. wazeylandica sp.n., P. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351070/files/figure.png" pageId="22" pageNumber="483">Fig. 6D</figureCitation>
|
||
) but may be more rectangular and can have stippling in the posterior half.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B291E76C6DCFC61FC453417" blockId="22.[151,1437,152,1187]" pageId="22" pageNumber="483">
|
||
<emphasis id="09EA6C149B291E76C6DCFC61FE9233EA" bold="true" box="[199,315,945,971]" pageId="22" pageNumber="483">Remarks.</emphasis>
|
||
Based on the immature stages alone,
|
||
<taxonomicName id="FC9ECB859B291E76C4F8FC61FCCE33EB" authorityName="Cranston" authorityYear="2019" box="[739,871,944,970]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="483" phylum="Arthropoda" rank="species" species="phengari">
|
||
<emphasis id="09EA6C149B291E76C4F8FC61FCCE33EB" box="[739,871,944,970]" italics="true" pageId="22" pageNumber="483">R. phengari</emphasis>
|
||
</taxonomicName>
|
||
would be poorly differentiated from
|
||
<taxonomicName id="FC9ECB859B291E76C313FC61FF1833CF" authorityName="Cranston" authorityYear="2019" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="483" phylum="Arthropoda" rank="species" species="azeylandica">
|
||
<emphasis id="09EA6C149B291E76C313FC61FF1833CF" italics="true" pageId="22" pageNumber="483">R. azeylandica</emphasis>
|
||
</taxonomicName>
|
||
. The attenuated gonostylus of the reared
|
||
<typeStatus id="E4250EA49B291E76C497FC04FD4733CF" box="[652,750,980,1006]" pageId="22" pageNumber="483" type="holotype">holotype</typeStatus>
|
||
is seen also in a prior-collected male from close-by, housed in the ANIC. The genitalia are totally symmetrical so evidently this gonostylus is not teratological but represents a distinctive new species. The species is unrepresented by molecular material.
|
||
</paragraph>
|
||
<paragraph id="3B21B0069B291E76C6DCFB90FD1F3483" blockId="22.[151,1437,152,1187]" pageId="22" pageNumber="483">
|
||
<emphasis id="09EA6C149B291E76C6DCFB90FE52347A" bold="true" box="[199,507,1088,1115]" pageId="22" pageNumber="483">DIstribution and Ecology.</emphasis>
|
||
<taxonomicName id="FC9ECB859B291E76C41AFB90FD14347B" authorityName="Cranston" authorityYear="2019" box="[513,701,1088,1114]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="483" phylum="Arthropoda" rank="species" species="phengari">
|
||
<emphasis id="09EA6C149B291E76C41AFB90FD14347B" box="[513,701,1088,1114]" italics="true" pageId="22" pageNumber="483">Riethia phengari</emphasis>
|
||
</taxonomicName>
|
||
is limited to a very small area of eastern
|
||
<collectingCountry id="4389F0969B291E76C298FB90FB43347B" box="[1155,1258,1088,1114]" name="Australia" pageId="22" pageNumber="483">Australia</collectingCountry>
|
||
coincident with the mountains of the
|
||
<collectingRegion id="F95A7EE49B291E76C765FBB4FD1D345F" box="[382,692,1124,1150]" country="Australia" name="Australian Capital Territory" pageId="22" pageNumber="483">Australian Capital Territory</collectingRegion>
|
||
. The streams are 2nd or 3rd order, in a region very widely surveyed including by drift netting over extended periods.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |