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<document id="BEE8D07BEDDA5D4A57A069978D7E105D" ID-CLB-Dataset="25594" ID-DOI="10.11646/zootaxa.4646.3.3" ID-GBIF-Dataset="4a3d31ca-e16b-4ccf-8407-231aa27702f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3351058" ID-ZooBank="25CA4A18-955F-4EA7-8978-E93032B54A2E" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564032263204" checkinUser="plazi" docAuthor="Cranston, Peter S." docDate="2019" docId="B33701109B231E7DC68CFA21FED537B9" docLanguage="en" docName="zootaxao.4646.3.3.pdf" docOrigin="Zootaxa 4646 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Riethia zeylandica Freeman 1959" docType="treatment" docVersion="8" lastPageNumber="490" masterDocId="4F0E79689B3F1E60C61BFFD0FFA93021" masterDocTitle="Riethia (Kieffer 1917) (Diptera: Chironomidae) revised for the Austro-Pacific region" masterLastPageNumber="500" masterPageNumber="461" pageNumber="489" updateTime="1698747359191" updateUser="ExternalLinkService">
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<mods:title id="D997F2C06457A7AE4DEC4C1D6CA7E8AA">Riethia (Kieffer 1917) (Diptera: Chironomidae) revised for the Austro-Pacific region</mods:title>
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<mods:namePart id="AD634E43F50F0A52D26AAAD97CE20FE9">Cranston, Peter S.</mods:namePart>
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<mods:date id="FDEBB695FAC0170E973B63E018393F41">2019</mods:date>
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<treatment id="B33701109B231E7DC68CFA21FED537B9" ID-DOI="http://doi.org/10.5281/zenodo.5587172" ID-GBIF-Taxon="159413599" ID-Zenodo-Dep="5587172" LSID="urn:lsid:plazi:treatment:B33701109B231E7DC68CFA21FED537B9" httpUri="http://treatment.plazi.org/id/B33701109B231E7DC68CFA21FED537B9" lastPageId="29" lastPageNumber="490" pageId="28" pageNumber="489">
<subSubSection id="7384E38D9B231E7CC68CFA21FD86362A" box="[151,559,1521,1548]" pageId="28" pageNumber="489" type="nomenclature">
<paragraph id="3B21B0069B231E7CC68CFA21FD86362A" blockId="28.[151,559,1521,1584]" box="[151,559,1521,1548]" pageId="28" pageNumber="489">
<heading id="6069076A9B231E7CC68CFA21FD86362A" bold="true" box="[151,559,1521,1548]" fontSize="11" level="1" pageId="28" pageNumber="489" reason="1">
<taxonomicName id="FC9ECB859B231E7CC68CFA21FD86362A" ID-CoL="4T32R" authority="Freeman 1959" authorityName="Freeman" authorityYear="1959" box="[151,559,1521,1548]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="489" phylum="Arthropoda" rank="species" species="zeylandica">
<emphasis id="09EA6C149B231E7CC68CFA21FD86362A" bold="true" box="[151,559,1521,1548]" pageId="28" pageNumber="489">
<emphasis id="09EA6C149B231E7CC68CFA21FEDF362D" bold="true" box="[151,374,1521,1548]" italics="true" pageId="28" pageNumber="489">Riethia zeylandica</emphasis>
<bibRefCitation id="5F0FCDF79B231E7CC766FA21FD86362A" author="Freeman, P." box="[381,559,1521,1547]" pageId="28" pageNumber="489" pagination="395 - 437" refId="ref40130" refString="Freeman, P. (1959) A study of the New Zealand Chironomidae (Diptera, Nematocera). Bulletin of the British Museum (Natural History), Entomology, 7, 395 - 437." type="journal article" year="1959">Freeman 1959</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7384E38D9B231E7CC68CF9C5FE3D3611" box="[151,404,1557,1584]" pageId="28" pageNumber="489" type="description">
<paragraph id="3B21B0069B231E7CC68CF9C5FE3D3611" blockId="28.[151,559,1521,1584]" box="[151,404,1557,1584]" pageId="28" pageNumber="489">
(
<figureCitation id="A3A5AC839B231E7CC684F9C5FF55360E" box="[159,252,1557,1583]" captionStart="FIGURE 1" captionStartId="10.[151,250,1970,1995]" captionTargetBox="[237,1393,192,1878]" captionTargetId="figure@10.[178,1410,166,1950]" captionTargetPageId="10" captionText="FIGURE 1. Riethia spp. Male hypopygia L. side dorsal, R. side, ventral/internal, stylised: A. R. donedwardi sp.n., B. R. hamodivisa sp.n., C. R. phengari, sp.n. D. R. plumosa Freeman, E. R. stictoptera Keffer, F. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351060/files/figure.png" pageId="28" pageNumber="489">Figs. 1F</figureCitation>
,
<figureCitation id="A3A5AC839B231E7CC713F9C5FE98360E" box="[264,305,1557,1583]" captionStart="FIGURE 2" captionStartId="11.[151,250,1925,1950]" captionTargetBox="[177,1375,200,1857]" captionTargetId="figure@11.[160,1427,166,1905]" captionTargetPageId="11" captionText="FIGURE 2. Riethia spp. Volsellae: A. R. azeylandica sp.n., B. R. cinctipes Freeman, C. R. donedwardi sp.n. D. R. hamodivisa sp.n., E. R. kakadu sp.n., F. R. neocaledonica sp.n., G. R. noongar sp.n., H. R. phengari sp.n., I. R. plumosa Freeman, J. R. queenslandensis sp.n., K. R. stictoptera Keffer, L. R. wazeylandica sp.n., M. R. zeylandica sp.n." httpUri="https://zenodo.org/record/3351062/files/figure.png" pageId="28" pageNumber="489">2M</figureCitation>
,
<figureCitation id="A3A5AC839B231E7CC726F9C5FEC8360E" box="[317,353,1557,1583]" captionStart="FIGURE 4" captionStartId="13.[151,250,1897,1922]" captionTargetBox="[199,1399,178,1821]" captionTargetId="figure@13.[175,1412,166,1877]" captionTargetPageId="13" captionText="FIGURE 4. Riethia spp. Pupa. Abdominal tergites: A. R. noongar sp.n., B. R. paluma sp.n., C. R. plumosa Freeman, D. R. stictoptera Kieffer. Posterolateral corner of VIII (‘comb’): E. R. noongar, F. R. paluma, G. R. plumosa, H. R. stictoptera I. R. cinctipes (var), J. R. phengari sp.n., K. R. neocaledonica sp.n.. Tergites: L, IV-V of R. neocaledonica; M. Tergites VI-VIII of R. queenslandensis sp.n., N. Sternites VI-VIII R. queenslandensis, O. Tergites V-AL of R. zeylandica." httpUri="https://zenodo.org/record/3351066/files/figure.png" pageId="28" pageNumber="489">4O</figureCitation>
,
<figureCitation id="A3A5AC839B231E7CC776F9C5FE253611" box="[365,396,1557,1584]" captionStart="FIGURE 6" captionStartId="15.[151,250,1892,1917]" captionTargetBox="[163,1417,176,1843]" captionTargetId="figure@15.[151,1436,166,1873]" captionTargetPageId="15" captionText="FIGURE 6. Riethia spp. Larva. Mid-dorsal head:A. R. donedwardi, sp.n., B. R. noongar sp.n.. Clypeus: C. R. larval sp. ‘C’, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. neocaledonica sp.n., I. R. noongar sp.n., J. R. paluma sp.n., K. R. phengari sp.n., L. R. plumosa Freeman, M. R. queenslandensis sp.n., N. R. stictoptera Kieffer, O. R. wazeylandica sp.n., P. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351070/files/figure.png" pageId="28" pageNumber="489">6P</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7384E38D9B231E7CC68CF989FDFF36B1" pageId="28" pageNumber="489" type="reference_group">
<paragraph id="3B21B0069B231E7CC68CF989FD8C3650" blockId="28.[151,598,1624,1680]" box="[151,549,1624,1649]" pageId="28" pageNumber="489">
<treatmentCitationGroup id="1B8E97289B231E7CC68CF989FD8C3650" box="[151,549,1624,1649]" pageId="28" pageNumber="489">
<taxonomicName id="FC9ECB859B231E7CC68CF989FD8C3650" ID-CoL="4T32R" authority="Freeman, 1959: 422" authorityName="Freeman" authorityPageNumber="422" authorityYear="1959" box="[151,549,1624,1649]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="489" phylum="Arthropoda" rank="species" species="zeylandica">
<emphasis id="09EA6C149B231E7CC68CF989FEFA3650" box="[151,339,1625,1649]" italics="true" pageId="28" pageNumber="489">Riethia zeylandica</emphasis>
<treatmentCitation id="BA3F96179B231E7CC742F98AFD8C3650" author="Freeman, P." box="[345,549,1624,1649]" page="422" pageId="28" pageNumber="489" year="1959">
<bibRefCitation id="5F0FCDF79B231E7CC742F98AFD8C3650" author="Freeman, P." box="[345,549,1624,1649]" pageId="28" pageNumber="489" pagination="395 - 437" refId="ref40130" refString="Freeman, P. (1959) A study of the New Zealand Chironomidae (Diptera, Nematocera). Bulletin of the British Museum (Natural History), Entomology, 7, 395 - 437." type="journal article" year="1959">Freeman, 1959: 422</bibRefCitation>
</treatmentCitation>
</taxonomicName>
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</paragraph>
<paragraph id="3B21B0069B231E7CC68CF9A8FDFF36B1" blockId="28.[151,598,1624,1680]" box="[151,598,1655,1680]" pageId="28" pageNumber="489">
[not
<taxonomicName id="FC9ECB859B231E7CC6DDF9A8FDF836B1" ID-CoL="4T32R" authority="Freeman 1961: 680" authorityName="Freeman" authorityPageNumber="680" authorityYear="1961" box="[198,593,1655,1680]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="489" phylum="Arthropoda" rank="species" species="zeylandica">
<emphasis id="09EA6C149B231E7CC6DDF9A8FE2B36B1" box="[198,386,1656,1680]" italics="true" pageId="28" pageNumber="489">Riethia zeylandica</emphasis>
<bibRefCitation id="5F0FCDF79B231E7CC793F9A8FDF836B1" author="Freeman, P." box="[392,593,1655,1680]" pageId="28" pageNumber="489" pagination="611 - 737" refId="ref40168" refString="Freeman, P. (1961) The Chironomidae of Australia. Australian Journal of Zoology, 9, 611 - 737. https: // doi. org / 10.1071 / ZO 9610611" type="journal article" year="1961">Freeman 1961: 680</bibRefCitation>
</taxonomicName>
]
</paragraph>
</subSubSection>
<subSubSection id="7384E38D9B231E7DC68CF96EFED537B9" lastPageId="29" lastPageNumber="490" pageId="28" pageNumber="489" type="materials_examined">
<paragraph id="3B21B0069B231E7CC68CF96EFB8D36DD" blockId="28.[151,1437,1725,2041]" pageId="28" pageNumber="489">
<emphasis id="09EA6C149B231E7CC68CF96EFEEC36F9" bold="true" box="[151,325,1725,1752]" pageId="28" pageNumber="489">Type material.</emphasis>
<materialsCitation id="8BF6BA5B9B231E7CC757F96EFB8D36DD" ID-GBIF-Occurrence="2382714928" collectingDate="1922-11" collectingDateMax="1998-09" collectingDateMin="1922-11" collectionCode="BMNH" collectorName="T. K. Harris" country="New Zealand" location="Ohakune" pageId="28" pageNumber="489" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="E4250EA49B231E7CC757F96EFE1036F9" box="[332,441,1726,1752]" pageId="28" pageNumber="489" type="holotype">
<emphasis id="09EA6C149B231E7CC757F96EFE1036F9" bold="true" box="[332,441,1726,1752]" pageId="28" pageNumber="489">Holotype</emphasis>
</typeStatus>
<specimenCount id="2D987B8F9B231E7CC7DAF96FFE7136F8" box="[449,472,1727,1753]" pageId="28" pageNumber="489" type="male">♂</specimenCount>
, pinned, hypopygium on celluloid mount,
<collectingCountry id="4389F0969B231E7CC5AEF96FFB2A36F9" box="[949,1155,1726,1753]" name="New Zealand" pageId="28" pageNumber="489">NEW ZEALAND</collectingCountry>
,
<location id="3E41E6DD9B231E7CC295F96EFB5F36F9" LSID="urn:lsid:plazi:treatment:B33701109B231E7DC68CFA21FED537B9:3E41E6DD9B231E7CC295F96EFB5F36F9" box="[1166,1270,1726,1752]" country="New Zealand" name="Ohakune" pageId="28" pageNumber="489">Ohakune</location>
,.
<collectingDate id="5F646F2E9B231E7CC31EF96EFAF736F9" box="[1285,1374,1726,1752]" pageId="28" pageNumber="489" value="1922-11">
<date id="4F2096C69B231E7CC31EF96EFAF736F9" box="[1285,1374,1726,1752]" pageId="28" pageNumber="489" value="1922-11">xi.1922</date>
</collectingDate>
(
<collectorName id="966BD5D09B231E7CC375F96FFF4B36DD" pageId="28" pageNumber="489">
<emphasis id="09EA6C149B231E7CC375F96FFF4B36DD" italics="true" pageId="28" pageNumber="489">T.K. Harris</emphasis>
</collectorName>
) BM 1923-13 (
<collectionCode id="5D8F28C39B231E7CC78AF932FE4236DD" box="[401,491,1762,1788]" pageId="28" pageNumber="489">BMNH</collectionCode>
). Examined, hypopygium drawn,
<collectingDate id="5F646F2E9B231E7CC57DF932FB8936DD" box="[870,1056,1762,1788]" pageId="28" pageNumber="489" value="1998-09">
<date id="4F2096C69B231E7CC57DF932FB8936DD" box="[870,1056,1762,1788]" pageId="28" pageNumber="489" value="1998-09">September 1998</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="3B21B0069B231E7CC68CF8D5FD9C3749" blockId="28.[151,1437,1725,2041]" pageId="28" pageNumber="489">
<emphasis id="09EA6C149B231E7CC68CF8D5FEE33701" bold="true" box="[151,330,1797,1824]" pageId="28" pageNumber="489">Other material</emphasis>
(all
<emphasis id="09EA6C149B231E7CC767F8D7FE4D3700" box="[380,484,1799,1825]" italics="true" pageId="28" pageNumber="489">Cranston</emphasis>
ANIC /
<collectionCode id="5D8F28C39B231E7CC45FF8D6FD3F3701" box="[580,662,1798,1824]" country="New Zealand" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="28" pageNumber="489" type="Other institutional type">NZAC</collectionCode>
):
<materialsCitation id="8BF6BA5B9B231E7CC4B3F8D7FD1E3765" ID-GBIF-Occurrence="2382714930" collectingDate="2000-01-12" collectorName="N. Island &amp; Pe" country="New Zealand" latitude="-35.208668" location="Pukete Forest" longLatPrecision="11" longitude="173.7905" pageId="28" pageNumber="489" specimenCount="1">
<collectingCountry id="4389F0969B231E7CC4B3F8D7FCDE3701" box="[680,887,1798,1825]" name="New Zealand" pageId="28" pageNumber="489">NEW ZEALAND</collectingCountry>
:
<collectorName id="966BD5D09B231E7CC59BF8D7FC403701" box="[896,1001,1798,1825]" pageId="28" pageNumber="489">N. Island</collectorName>
, 24
<collectorName id="966BD5D09B231E7CC20BF8D6FB873701" box="[1040,1070,1798,1824]" pageId="28" pageNumber="489">Pe</collectorName>
,
<location id="3E41E6DD9B231E7CC222F8D6FB7C3700" LSID="urn:lsid:plazi:treatment:B33701109B231E7DC68CFA21FED537B9:3E41E6DD9B231E7CC222F8D6FB7C3700" box="[1081,1237,1798,1825]" country="New Zealand" latitude="-35.208668" longLatPrecision="11" longitude="173.7905" name="Pukete Forest" pageId="28" pageNumber="489">Pukete Forest</location>
, 1–2 order stream (unnamed),
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,
<collectingDate id="5F646F2E9B231E7CC451F8FAFD1E3765" box="[586,695,1834,1860]" pageId="28" pageNumber="489" value="2000-01-12">
<date id="4F2096C69B231E7CC451F8FAFD1E3765" box="[586,695,1834,1860]" pageId="28" pageNumber="489" value="2000-01-12">12.i.2000</date>
</collectingDate>
</materialsCitation>
; Le/P
<specimenCount id="2D987B8F9B231E7CC4E4F8FAFCBB3765" box="[767,786,1834,1860]" pageId="28" pageNumber="489" type="male">♂</specimenCount>
,
<geoCoordinate id="5EAAD6C19B231E7CC53BF8F9FC363765" box="[800,927,1833,1860]" degrees="35" direction="south" minutes="12.41" orientation="latitude" pageId="28" pageNumber="489" precision="9" value="-35.206833">35°12.41’S</geoCoordinate>
<geoCoordinate id="5EAAD6C19B231E7CC5B2F8FAFB923765" box="[937,1083,1834,1860]" degrees="173" direction="east" minutes="47.37" orientation="longitude" pageId="28" pageNumber="489" precision="9" value="173.7895">173°47.37’E</geoCoordinate>
,
<date id="4F2096C69B231E7CC252F8FAFB1C3765" box="[1097,1205,1834,1860]" pageId="28" pageNumber="489" value="2006-01-30">30.i.2006</date>
;
<materialsCitation id="8BF6BA5B9B231E7CC2DFF8FAFE363748" ID-GBIF-Occurrence="2382714929" collectionCode="MV" country="New Zealand" location="NZ" pageId="28" pageNumber="489" specimenCount="1" specimenCount-male="1">
2L, P
<specimenCount id="2D987B8F9B231E7CC31EF8FBFAB03764" box="[1285,1305,1835,1861]" pageId="28" pageNumber="489" type="male">♂</specimenCount>
same (
<collectionCode id="5D8F28C39B231E7CC375F8FAFA353765" box="[1390,1436,1834,1860]" country="0" name="University of Montana Museum" pageId="28" pageNumber="489">MV</collectionCode>
<emphasis id="09EA6C149B231E7CC68CF89DFE3F3749" bold="true" box="[151,406,1869,1896]" pageId="28" pageNumber="489">
<collectingCountry id="4389F0969B231E7CC68CF89DFF163749" box="[151,191,1869,1896]" name="New Zealand" pageId="28" pageNumber="489">NZ</collectingCountry>
10PSR1-2, PSRP1
</emphasis>
)
</materialsCitation>
27.xiii.2010.
</paragraph>
<paragraph id="3B21B0069B231E7DC6DCF8A1FA393162" blockId="28.[151,1437,1725,2041]" lastBlockId="29.[151,1437,153,1944]" lastPageId="29" lastPageNumber="490" pageId="28" pageNumber="489">
<emphasis id="09EA6C149B231E7CC6DCF8A1FE3637AD" bold="true" box="[199,415,1905,1932]" pageId="28" pageNumber="489">Description. Male</emphasis>
, n=1–3 [based on
<bibRefCitation id="5F0FCDF79B231E7CC463F8A3FCB637AD" author="Freeman, P." box="[632,799,1905,1933]" pageId="28" pageNumber="489" pagination="395 - 437" refId="ref40130" refString="Freeman, P. (1959) A study of the New Zealand Chironomidae (Diptera, Nematocera). Bulletin of the British Museum (Natural History), Entomology, 7, 395 - 437." type="journal article" year="1959">Freeman 1959</bibRefCitation>
, re-examination of
<typeStatus id="E4250EA49B231E7CC21BF8A2FBCB37AD" box="[1024,1122,1906,1932]" pageId="28" pageNumber="489" type="holotype">holotype</typeStatus>
by Cranston; the only new specimens are teneral within pupa]. Body pale green with light brown thoracic bands; wing and legs pale, without markings. Antennal ratio 1.05 [Freeman states 1.3]. Legs with all tarsomeres missing, ratios unable to be calculated [Freeman ‘slightly more than 1]. Hypopygium (
<figureCitation id="A3A5AC839B231E7CC4A9F80EFCAC37D9" box="[690,773,2014,2040]" captionStart="FIGURE 1" captionStartId="10.[151,250,1970,1995]" captionTargetBox="[237,1393,192,1878]" captionTargetId="figure@10.[178,1410,166,1950]" captionTargetPageId="10" captionText="FIGURE 1. Riethia spp. Male hypopygia L. side dorsal, R. side, ventral/internal, stylised: A. R. donedwardi sp.n., B. R. hamodivisa sp.n., C. R. phengari, sp.n. D. R. plumosa Freeman, E. R. stictoptera Keffer, F. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351060/files/figure.png" pageId="28" pageNumber="489">Fig. 1F</figureCitation>
) [
<bibRefCitation id="5F0FCDF79B231E7CC507F80FFC6C37D9" author="Freeman, P." box="[796,965,2014,2041]" pageId="28" pageNumber="489" pagination="611 - 737" refId="ref40168" refString="Freeman, P. (1961) The Chironomidae of Australia. Australian Journal of Zoology, 9, 611 - 737. https: // doi. org / 10.1071 / ZO 9610611" type="journal article" year="1961">Freeman 1961</bibRefCitation>
: fig. 3a] with 4–6 clustered stout setae on anal tergite near posterior margin and interspersed with regular setae. Gonocoxite slightly narrowed at junction to gonostylus, gonostylus tapering to point. Superior volsella (
<figureCitation id="A3A5AC839B221E7DC520FF6DFC3130F6" box="[827,920,189,215]" captionStart="FIGURE 2" captionStartId="11.[151,250,1925,1950]" captionTargetBox="[177,1375,200,1857]" captionTargetId="figure@11.[160,1427,166,1905]" captionTargetPageId="11" captionText="FIGURE 2. Riethia spp. Volsellae: A. R. azeylandica sp.n., B. R. cinctipes Freeman, C. R. donedwardi sp.n. D. R. hamodivisa sp.n., E. R. kakadu sp.n., F. R. neocaledonica sp.n., G. R. noongar sp.n., H. R. phengari sp.n., I. R. plumosa Freeman, J. R. queenslandensis sp.n., K. R. stictoptera Keffer, L. R. wazeylandica sp.n., M. R. zeylandica sp.n." httpUri="https://zenodo.org/record/3351062/files/figure.png" pageId="29" pageNumber="490">Fig. 2M</figureCitation>
) with basal ‘heel’ prior to non-microtrichiose elongate, non-tapered, bare lobe bearing 4–6 simple setae and without pectinate scales. Pseudovolsella an angular protrusion (not a tubercle) bearing 3 slightly approximated long setae. Inferior volsella well developed, microtrichiose, curved medially, subapically with 7–8 pectinate setae and several simple setae. Mensural details, see
<tableCitation id="761C85BD9B221E7DC32CFEF9FA233162" box="[1335,1418,297,323]" captionStart="TABLE 1" captionStartId="30.[149,242,517,544]" captionTargetBox="[164,1998,555,1659]" captionText="TABLE 1. Morphometric data for males of Riethia Kieffer" httpUri="http://table.plazi.org/id/6FE1E08E9B211E7EC68EFDD5FC833201" pageId="29" pageNumber="490" tableUuid="6FE1E08E9B211E7EC68EFDD5FC833201">Table 1</tableCitation>
.
</paragraph>
<paragraph id="3B21B0069B221E7DC6DCFE9DFE3C3146" blockId="29.[151,1437,153,1944]" box="[199,405,333,359]" pageId="29" pageNumber="490">
<emphasis id="09EA6C149B221E7DC6DCFE9DFEB73146" bold="true" box="[199,286,333,359]" pageId="29" pageNumber="490">Female</emphasis>
unknown.
</paragraph>
<paragraph id="3B21B0069B221E7DC6DCFEA1FBB332EE" blockId="29.[151,1437,153,1944]" pageId="29" pageNumber="490">
<emphasis id="09EA6C149B221E7DC6DCFEA1FEAC31AA" bold="true" box="[199,261,369,395]" pageId="29" pageNumber="490">Pupa</emphasis>
. Almost hyaline except for yellowish dorsal thorax, anterior wing sheaths; tergal spinules golden, apophyses almost invisible except on VIII; comb yellow-brown. Frons with short warts, cephalothorax smooth, with one row of tubercles each side of dorsal ecdysial line. Prealar tubercle modestly developed as rectangular to elongate rounded lobe. Continuous hook row on II occupying 30–35% width of segment II; continuous conjunctival spinule band on III, IV and sparser on V. Pedes spurii B absent, vortex very weak. Tergite I bare, II–V with quadrate armament of quite dense spinules, without strongly delimited anterior or posterior transverse band; VI with more patchily distributed spinules although essentially quadrate; VII and VIII bare. Segment V with 3L setae, all non-taeniate, VI–VIII with 0, 4, 4, 5 taeniate LS, on VIII more or less evenly spaced (
<figureCitation id="A3A5AC839B221E7DC5D3FDBDFB8A32A6" box="[968,1059,621,647]" captionStart="FIGURE 4" captionStartId="13.[151,250,1897,1922]" captionTargetBox="[199,1399,178,1821]" captionTargetId="figure@13.[175,1412,166,1877]" captionTargetPageId="13" captionText="FIGURE 4. Riethia spp. Pupa. Abdominal tergites: A. R. noongar sp.n., B. R. paluma sp.n., C. R. plumosa Freeman, D. R. stictoptera Kieffer. Posterolateral corner of VIII (‘comb’): E. R. noongar, F. R. paluma, G. R. plumosa, H. R. stictoptera I. R. cinctipes (var), J. R. phengari sp.n., K. R. neocaledonica sp.n.. Tergites: L, IV-V of R. neocaledonica; M. Tergites VI-VIII of R. queenslandensis sp.n., N. Sternites VI-VIII R. queenslandensis, O. Tergites V-AL of R. zeylandica." httpUri="https://zenodo.org/record/3351066/files/figure.png" pageId="29" pageNumber="490">Fig. 4O</figureCitation>
). Sternites bare. Tergite VIII posteriorly with yellow apophyses meeting comb comprising few golden spines, the strongest directed posteriorly, and innermost directed more posteromedially. Anal lobe with 40–50 biserial taeniae.
</paragraph>
<paragraph id="3B21B0069B221E7DC6DCFD09FF0533EA" blockId="29.[151,1437,153,1944]" pageId="29" pageNumber="490">
<emphasis id="09EA6C149B221E7DC6DCFD09FEB932D2" bold="true" box="[199,272,729,755]" pageId="29" pageNumber="490">Larva</emphasis>
. The head is evenly pale pigmented with postoccipital margin broad and dark, and mentum and inner mandibular teeth distinctly brown. The clypeus (
<figureCitation id="A3A5AC839B221E7DC4A2FD2DFCA53339" box="[697,780,765,792]" captionStart="FIGURE 6" captionStartId="15.[151,250,1892,1917]" captionTargetBox="[163,1417,176,1843]" captionTargetId="figure@15.[151,1436,166,1873]" captionTargetPageId="15" captionText="FIGURE 6. Riethia spp. Larva. Mid-dorsal head:A. R. donedwardi, sp.n., B. R. noongar sp.n.. Clypeus: C. R. larval sp. ‘C’, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. neocaledonica sp.n., I. R. noongar sp.n., J. R. paluma sp.n., K. R. phengari sp.n., L. R. plumosa Freeman, M. R. queenslandensis sp.n., N. R. stictoptera Kieffer, O. R. wazeylandica sp.n., P. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351070/files/figure.png" pageId="29" pageNumber="490">Fig. 6P</figureCitation>
) has an indented lateral margin with clypeal (S3) setae located midway between anterior and posterior margins. Antenna with short pedestal; AR c. 1.4, each segment shorter than preceding (thus 3
<superScript id="CCEB1D4E9B221E7DC78DFC94FE0D3373" attach="left" box="[406,420,836,850]" fontSize="6" pageId="29" pageNumber="490">rd</superScript>
shorter than 2
<superScript id="CCEB1D4E9B221E7DC451FC94FDF33373" attach="none" box="[586,602,836,850]" fontSize="6" pageId="29" pageNumber="490">nd</superScript>
). Premandible with very small basal-most tooth, essentially with 4 teeth. Mandible with gold apical tooth and 4 dark inner mandibular teeth, seta subdentalis arising from pointed lobe, with 1–2 molar spines. Mentum relatively narrow with ventromental plates of similar width. Mensural details, see
<tableCitation id="761C85BD9B221E7DC345FC5DFF0133EA" captionStart="TABLE 2" captionStartId="32.[149,242,622,649]" captionTargetBox="[164,1976,660,1609]" captionText="TABLE 2. Morphometric data for larvae of Riethia Kieffer" httpUri="http://table.plazi.org/id/6FE1E08E9B1F1E40C68EFDBEFC8432A8" pageId="29" pageNumber="490" tableUuid="6FE1E08E9B1F1E40C68EFDBEFC8432A8">Table 2</tableCitation>
.
</paragraph>
<paragraph id="3B21B0069B221E7DC6DCFC04FAF73416" blockId="29.[151,1437,153,1944]" pageId="29" pageNumber="490">
<emphasis id="09EA6C149B221E7DC6DCFC04FEEB33CE" bold="true" box="[199,322,980,1007]" pageId="29" pageNumber="490">Diagnosis.</emphasis>
The adult male closely resembles the Australian
<taxonomicName id="FC9ECB859B221E7DC572FC06FBA733CE" authorityName="Cranston" authorityYear="2019" box="[873,1038,981,1007]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="490" phylum="Arthropoda" rank="species" species="azeylandica">
<emphasis id="09EA6C149B221E7DC572FC06FBA733CE" box="[873,1038,981,1007]" italics="true" pageId="29" pageNumber="490">R. azeylandica</emphasis>
</taxonomicName>
, differing mainly in the pseudovolsella. In both species it bears 3 setae, but in
<taxonomicName id="FC9ECB859B221E7DC462FC2AFCA63432" authorityName="Freeman" authorityYear="1961" box="[633,783,1017,1043]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="490" phylum="Arthropoda" rank="species" species="zeylandica">
<emphasis id="09EA6C149B221E7DC462FC2AFCA63432" box="[633,783,1017,1043]" italics="true" pageId="29" pageNumber="490">R. zeylandica</emphasis>
</taxonomicName>
these arise from a distinct angular lobe (
<figureCitation id="A3A5AC839B221E7DC2CBFC29FA873432" box="[1232,1326,1017,1043]" captionStart="FIGURE 2" captionStartId="11.[151,250,1925,1950]" captionTargetBox="[177,1375,200,1857]" captionTargetId="figure@11.[160,1427,166,1905]" captionTargetPageId="11" captionText="FIGURE 2. Riethia spp. Volsellae: A. R. azeylandica sp.n., B. R. cinctipes Freeman, C. R. donedwardi sp.n. D. R. hamodivisa sp.n., E. R. kakadu sp.n., F. R. neocaledonica sp.n., G. R. noongar sp.n., H. R. phengari sp.n., I. R. plumosa Freeman, J. R. queenslandensis sp.n., K. R. stictoptera Keffer, L. R. wazeylandica sp.n., M. R. zeylandica sp.n." httpUri="https://zenodo.org/record/3351062/files/figure.png" pageId="29" pageNumber="490">Fig. 2M</figureCitation>
), whereas there is no such lobe in
<taxonomicName id="FC9ECB859B221E7DC7BAFBCEFDEF3416" authorityName="Cranston" authorityYear="2019" box="[417,582,1053,1079]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="490" phylum="Arthropoda" rank="species" species="azeylandica">
<emphasis id="09EA6C149B221E7DC7BAFBCEFDEF3416" box="[417,582,1053,1079]" italics="true" pageId="29" pageNumber="490">R. azeylandica</emphasis>
</taxonomicName>
(
<figureCitation id="A3A5AC839B221E7DC44EFBCDFD063416" box="[597,687,1053,1079]" captionStart="FIGURE 2" captionStartId="11.[151,250,1925,1950]" captionTargetBox="[177,1375,200,1857]" captionTargetId="figure@11.[160,1427,166,1905]" captionTargetPageId="11" captionText="FIGURE 2. Riethia spp. Volsellae: A. R. azeylandica sp.n., B. R. cinctipes Freeman, C. R. donedwardi sp.n. D. R. hamodivisa sp.n., E. R. kakadu sp.n., F. R. neocaledonica sp.n., G. R. noongar sp.n., H. R. phengari sp.n., I. R. plumosa Freeman, J. R. queenslandensis sp.n., K. R. stictoptera Keffer, L. R. wazeylandica sp.n., M. R. zeylandica sp.n." httpUri="https://zenodo.org/record/3351062/files/figure.png" pageId="29" pageNumber="490">Fig. 2A</figureCitation>
), with setae arising from the inner contour of the gonocoxite.
</paragraph>
<paragraph id="3B21B0069B221E7DC6DCFB91FF16352E" blockId="29.[151,1437,153,1944]" pageId="29" pageNumber="490">
The pupa, with continuous hook row and conjunctival spinule bands on II, IV and V, is unlike any taxon in the ‘zeylandica’ grouping from
<collectingCountry id="4389F0969B221E7DC7CAFBB5FD92345E" box="[465,571,1125,1151]" name="Australia" pageId="29" pageNumber="490">Australia</collectingCountry>
. The conjunctive of V has a sparse, perhaps narrowly medially-divided band of spinules, well separated from posterior tergal spinules and weaker than in possibly conspecific Australian specimens, all of which have a denser band on V. The absence of taeniae on the lateral segment V is unique in the genus, and led to early inability to recognise the pupal exuviae as belonging to
<emphasis id="09EA6C149B221E7DC5A7FB01FBAC34CA" box="[956,1029,1233,1259]" italics="true" pageId="29" pageNumber="490">Rietha</emphasis>
due to (0), 4, 4, 5 pattern of taeniate LS.
</paragraph>
<paragraph id="3B21B0069B221E7DC6DCFAC9FB7D3576" blockId="29.[151,1437,153,1944]" pageId="29" pageNumber="490">
The larva with an AR of 1.4 with consecutively shorter segments and smooth clypeus probably is only confidently identifiable by its restriction to
<collectingCountry id="4389F0969B221E7DC45EFAEDFD743576" box="[581,733,1341,1367]" name="New Zealand" pageId="29" pageNumber="490">New Zealand</collectingCountry>
, whereas look-alikes are Western Australian.
</paragraph>
<paragraph id="3B21B0069B221E7DC6DCFAB1FEAE362A" blockId="29.[151,1437,153,1944]" pageId="29" pageNumber="490">
<emphasis id="09EA6C149B221E7DC6DCFAB1FE9D355A" bold="true" box="[199,308,1377,1403]" pageId="29" pageNumber="490">Remarks</emphasis>
. The resemblance of the adult male of
<taxonomicName id="FC9ECB859B221E7DC4E8FAB2FC22355A" authorityName="Freeman" authorityYear="1961" box="[755,907,1377,1403]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="490" phylum="Arthropoda" rank="species" species="zeylandica">
<emphasis id="09EA6C149B221E7DC4E8FAB2FC22355A" box="[755,907,1377,1403]" italics="true" pageId="29" pageNumber="490">R. zeylandica</emphasis>
</taxonomicName>
to Australian specimens allocated by
<bibRefCitation id="5F0FCDF79B221E7DC321FAB1FF4B3581" author="Freeman, P." pageId="29" pageNumber="490" pagination="611 - 737" refId="ref40168" refString="Freeman, P. (1961) The Chironomidae of Australia. Australian Journal of Zoology, 9, 611 - 737. https: // doi. org / 10.1071 / ZO 9610611" type="journal article" year="1961">Freeman (1961)</bibRefCitation>
to this species is evident and misidentication is quite understandable.Although molecular data shows distinction, and the pupae provided strong morphological confirmation, particularly in the non-taeniate LS on segment V and weak to absent vortices on IV, differentiation of the male and larva from some Australian congeners remains uncertain.
</paragraph>
<paragraph id="3B21B0069B221E7DC6DCF9C5FDC036C2" blockId="29.[151,1437,153,1944]" pageId="29" pageNumber="490">
Intensive searches around the
<typeStatus id="E4250EA49B221E7DC40FF9C5FDED360E" box="[532,580,1557,1583]" pageId="29" pageNumber="490">type</typeStatus>
locality of Ohakune revealed no
<taxonomicName id="FC9ECB859B221E7DC5A9F9C5FBAA360E" box="[946,1027,1557,1583]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="490" phylum="Arthropoda" rank="genus">
<emphasis id="09EA6C149B221E7DC5A9F9C5FBAA360E" box="[946,1027,1557,1583]" italics="true" pageId="29" pageNumber="490">Riethia</emphasis>
</taxonomicName>
in any life stage. Since Harris’s original collection in 1922, the local volcano Mount Ruapehu has erupted several times, and conceivably local stream conditions no longer suit
<taxonomicName id="FC9ECB859B221E7DC7A8F98DFDAD3656" box="[435,516,1629,1655]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="490" phylum="Arthropoda" rank="genus">
<emphasis id="09EA6C149B221E7DC7A8F98DFDAD3656" box="[435,516,1629,1655]" italics="true" pageId="29" pageNumber="490">Riethia</emphasis>
</taxonomicName>
. The species was found much further north in Pukete Forest (
<collectingRegion id="F95A7EE49B221E7DC2AFF98DFA833656" box="[1204,1322,1629,1655]" country="New Zealand" name="Northland" pageId="29" pageNumber="490">Northland</collectingRegion>
, Kerikeri) from drifting pupal exuviae in 2000, with larvae and pharate adults found in 2006 and larval molecular material collected in 2010. The life history association derives from molecular data of 2 unreared larvae and a pharate pupa with clearly visible male genitalia within.
</paragraph>
<paragraph id="3B21B0069B221E7DC6DCF93CFED537B9" blockId="29.[151,1437,153,1944]" pageId="29" pageNumber="490">
<emphasis id="09EA6C149B221E7DC6DCF93CFE473726" bold="true" box="[199,494,1772,1799]" pageId="29" pageNumber="490">Distribution and ecology</emphasis>
. Pukete Forest in northern
<collectingCountry id="4389F0969B221E7DC53BF93DFC1F3726" box="[800,950,1773,1799]" name="New Zealand" pageId="29" pageNumber="490">New Zealand</collectingCountry>
is a large intact tract of native forest dominated by ancient
<taxonomicName id="FC9ECB859B221E7DC74AF8C1FE57370D" authority="(Kauri)" authorityName="Kauri" baseAuthorityName="Kauri" box="[337,510,1809,1836]" class="Insecta" family="Braconidae" genus="Agathis" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="490" phylum="Arthropoda" rank="genus">
<emphasis id="09EA6C149B221E7DC74AF8C1FE0E370A" box="[337,423,1809,1835]" italics="true" pageId="29" pageNumber="490">Agathis</emphasis>
(Kauri)
</taxonomicName>
trees. The low order streams where
<taxonomicName id="FC9ECB859B221E7DC592F8C1FC73370A" box="[905,986,1809,1835]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="490" phylum="Arthropoda" rank="genus">
<emphasis id="09EA6C149B221E7DC592F8C1FC73370A" box="[905,986,1809,1835]" italics="true" pageId="29" pageNumber="490">Riethia</emphasis>
</taxonomicName>
occur are of modest gradient, with rocky substrate and filled with leaf litter. Apparently represented by the sole species,
<taxonomicName id="FC9ECB859B221E7DC218F8E5FBFD376E" box="[1027,1108,1845,1871]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="490" phylum="Arthropoda" rank="genus">
<emphasis id="09EA6C149B221E7DC218F8E5FBFD376E" box="[1027,1108,1845,1871]" italics="true" pageId="29" pageNumber="490">Riethia</emphasis>
</taxonomicName>
has been found nowhere else in the country despite widespread sampling across both North and South islands, including in other ancient forested streams (see above).
</paragraph>
</subSubSection>
</treatment>
</document>
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