147 lines
16 KiB
XML
147 lines
16 KiB
XML
<document id="E6EC106EA2E3CDB1C6EFABE990DF94B9" ID-CLB-Dataset="25594" ID-DOI="10.11646/zootaxa.4646.3.3" ID-GBIF-Dataset="4a3d31ca-e16b-4ccf-8407-231aa27702f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3351058" ID-ZooBank="25CA4A18-955F-4EA7-8978-E93032B54A2E" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564032263204" checkinUser="plazi" docAuthor="Cranston, Peter S." docDate="2019" docId="B33701109B1D1E42C68CFF31FB513386" docLanguage="en" docName="zootaxao.4646.3.3.pdf" docOrigin="Zootaxa 4646 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Riethia Kieffer 1917" docType="treatment" docVersion="12" lastPageNumber="495" masterDocId="4F0E79689B3F1E60C61BFFD0FFA93021" masterDocTitle="Riethia (Kieffer 1917) (Diptera: Chironomidae) revised for the Austro-Pacific region" masterLastPageNumber="500" masterPageNumber="461" pageNumber="495" updateTime="1698747359191" updateUser="ExternalLinkService">
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<mods:titleInfo id="3694222F40314B2E7ADBFFA0F2195C8D">
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<mods:title id="2300346AFB2D9F8C5FA7CCC8BAC85916">Riethia (Kieffer 1917) (Diptera: Chironomidae) revised for the Austro-Pacific region</mods:title>
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<mods:name id="14534203B8D0D1FDF49841FE5537138E" type="personal">
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<mods:roleTerm id="2B5C82577AB6D8FE3B3B32CD4CCA4CAD">Author</mods:roleTerm>
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<mods:namePart id="49366709E1CCA2661163057EF171CDE2">Cranston, Peter S.</mods:namePart>
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<mods:typeOfResource id="4E06B813EB707962DBD641D222F248AC">text</mods:typeOfResource>
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<mods:title id="9E4A25EE9D23890ED9F864F87365AAB1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A167C085D96EB159031F5A9B69EFFF55">
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<mods:date id="9192B2CFB322910FA6CB9D1AE77692FE">2019</mods:date>
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<mods:detail id="573E8234FCCDF739AED35807A84185AF" type="pubDate">
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<mods:number id="5394FC3FCBA104EC64785A637A1E2479">2019-07-25</mods:number>
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<mods:number id="25E8A42037AB6B0880C355C93D0BF0CA">4646</mods:number>
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<mods:detail id="FAB482A3EFE58427606EFFE563DBFBCA" type="issue">
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<mods:number id="6611772D7FFF8C1264CC3E24E2BAC083">3</mods:number>
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<mods:end id="56C8AD3265E404D87BD5AB0F88DF56F8">500</mods:end>
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<mods:classification id="503ABBC4B7A800644D3432C760FD0BAE">journal article</mods:classification>
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<mods:identifier id="87834DFDDEF7C2B9C470A48B292587FF" type="CLB-Dataset">25594</mods:identifier>
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<mods:identifier id="88C0659EAF5FF4FC7898D46EE83CD169" type="DOI">10.11646/zootaxa.4646.3.3</mods:identifier>
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<mods:identifier id="869C9190711487324A0ECA5B5CD52D24" type="GBIF-Dataset">4a3d31ca-e16b-4ccf-8407-231aa27702f1</mods:identifier>
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<mods:identifier id="B0E36DC14E268FB648BC66F5FF621AB0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C19E799EE343AA1EA4C8559002D5F2C1" type="Zenodo-Dep">3351058</mods:identifier>
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<mods:identifier id="A1DF79384BB4E4E8426D6E713C0D8DF7" type="ZooBank">25CA4A18-955F-4EA7-8978-E93032B54A2E</mods:identifier>
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<treatment id="B33701109B1D1E42C68CFF31FB513386" ID-DOI="http://doi.org/10.5281/zenodo.5587174" ID-GBIF-Taxon="159413583" ID-Zenodo-Dep="5587174" LSID="urn:lsid:plazi:treatment:B33701109B1D1E42C68CFF31FB513386" httpUri="http://treatment.plazi.org/id/B33701109B1D1E42C68CFF31FB513386" lastPageNumber="495" pageId="34" pageNumber="495">
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<subSubSection id="7384E38D9B1D1E42C68CFF31FECC30DA" box="[151,357,225,252]" pageId="34" pageNumber="495" type="nomenclature">
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<paragraph id="3B21B0069B1D1E42C68CFF31FECC30DA" blockId="34.[151,357,225,252]" box="[151,357,225,252]" pageId="34" pageNumber="495">
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<heading id="6069076A9B1D1E42C68CFF31FECC30DA" bold="true" box="[151,357,225,252]" fontSize="11" level="1" pageId="34" pageNumber="495" reason="1">
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<emphasis id="09EA6C149B1D1E42C68CFF31FECC30DA" bold="true" box="[151,357,225,252]" pageId="34" pageNumber="495">
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<taxonomicName id="FC9ECB859B1D1E42C68CFF31FF4630DD" ID-CoL="79ZG" ID-ENA="391820" authorityName="Kieffer" authorityYear="1917" box="[151,239,225,252]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="495" phylum="Arthropoda" rank="genus">
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<emphasis id="09EA6C149B1D1E42C68CFF31FF4630DD" bold="true" box="[151,239,225,252]" italics="true" pageId="34" pageNumber="495">Riethia</emphasis>
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</taxonomicName>
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larva ‘A’
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7384E38D9B1D1E42C68CFEF9FB513386" pageId="34" pageNumber="495" type="materials_examined">
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<paragraph id="3B21B0069B1D1E42C68CFEF9FEE13191" blockId="34.[151,1437,297,936]" pageId="34" pageNumber="495">
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<emphasis id="09EA6C149B1D1E42C68CFEF9FED13165" bold="true" box="[151,376,297,324]" pageId="34" pageNumber="495">Material examined</emphasis>
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, larval head capsules only:
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<collectingCountry id="4389F0969B1D1E42C4B1FEF9FCE03162" box="[682,841,297,323]" name="Australia" pageId="34" pageNumber="495">AUSTRALIA</collectingCountry>
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, 4L,
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<collectingRegion id="F95A7EE49B1D1E42C599FEFAFBE33162" box="[898,1098,297,324]" country="Australia" name="New South Wales" pageId="34" pageNumber="495">New South Wales</collectingRegion>
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, Currowan S.F., Cabbage Tree Ck.,
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<geoCoordinate id="5EAAD6C19B1D1E42C6D6FE9DFE8C3149" box="[205,293,333,360]" degrees="35" direction="south" minutes="34" orientation="latitude" pageId="34" pageNumber="495" precision="925" value="-35.566666">35°34'S</geoCoordinate>
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<geoCoordinate id="5EAAD6C19B1D1E42C730FE9DFE3C3149" box="[299,405,333,360]" degrees="150" direction="east" minutes="02" orientation="longitude" pageId="34" pageNumber="495" precision="925" value="150.03334">150°02'E</geoCoordinate>
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,
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<date id="4F2096C69B1D1E42C7BAFE9DFE553146" box="[417,508,333,359]" pageId="34" pageNumber="495" value="2009-01-02">2.i.2009</date>
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(
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<collectionCode id="5D8F28C39B1D1E42C410FE9DFD903146" box="[523,569,333,359]" country="0" name="University of Montana Museum" pageId="34" pageNumber="495">MV</collectionCode>
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<emphasis id="09EA6C149B1D1E42C45BFE9DFC963149" bold="true" box="[576,831,333,360]" pageId="34" pageNumber="495">AU09CTCR1, 2, 4, 5)</emphasis>
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; L, same, except
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<date id="4F2096C69B1D1E42C21FFE9DFBC73146" box="[1028,1134,333,359]" pageId="34" pageNumber="495" value="2000-01-23">23.i.2000</date>
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(
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<emphasis id="09EA6C149B1D1E42C265FE9EFA223146" box="[1150,1419,333,360]" italics="true" pageId="34" pageNumber="495">Cranston & Dimitriadis</emphasis>
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);
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<materialsCitation id="8BF6BA5B9B1D1E42C68CFEA2FEEB3191" ID-GBIF-Occurrence="2382714436" collectingDate="2009-02-04" collectionCode="MV" collectorName="Glenbog SF & Rutherford Ck." elevation="909" latitude="-36.6" location="Brown Mountain" longLatPrecision="1186" longitude="149.78334" pageId="34" pageNumber="495" specimenCode="NSWRGCR4" specimenCount="1">
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L,
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<collectorName id="966BD5D09B1D1E42C6A2FEA1FEEF31AA" box="[185,326,369,395]" pageId="34" pageNumber="495">Glenbog SF</collectorName>
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.,
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<location id="3E41E6DD9B1D1E42C74CFEA2FDB431AA" LSID="urn:lsid:plazi:treatment:B33701109B1D1E42C68CFF31FB513386:3E41E6DD9B1D1E42C74CFEA2FDB431AA" box="[343,541,369,396]" latitude="-36.6" longLatPrecision="1186" longitude="149.78334" name="Brown Mountain" pageId="34" pageNumber="495">Brown Mountain</location>
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, Fastigata Rd,
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<collectorName id="966BD5D09B1D1E42C4D1FEA1FCD031AA" box="[714,889,369,395]" pageId="34" pageNumber="495">Rutherford Ck.</collectorName>
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,
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<geoCoordinate id="5EAAD6C19B1D1E42C59CFEA1FC4A31AA" box="[903,995,369,395]" degrees="36" direction="south" minutes="36" orientation="latitude" pageId="34" pageNumber="495" precision="925" value="-36.6">36°36’S</geoCoordinate>
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<geoCoordinate id="5EAAD6C19B1D1E42C5F6FEA1FBF231AA" box="[1005,1115,369,395]" degrees="149" direction="east" minutes="47" orientation="longitude" pageId="34" pageNumber="495" precision="925" value="149.78334">149°47’E</geoCoordinate>
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,
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<elevation id="B0B357359B1D1E42C272FEA1FB1B31AD" box="[1129,1202,369,396]" metricMagnitude="2" metricUnit="m" metricValue="9.09" pageId="34" pageNumber="495" unit="m" value="909.0">
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<quantity id="FC661DE39B1D1E42C272FEA1FB1B31AD" box="[1129,1202,369,396]" metricMagnitude="2" metricUnit="m" metricValue="9.09" pageId="34" pageNumber="495" unit="m" value="909.0">909 m</quantity>
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</elevation>
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a.s.l,
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<collectingDate id="5F646F2E9B1D1E42C2E2FEA1FAF531AA" box="[1273,1372,369,395]" pageId="34" pageNumber="495" value="2009-02-04">
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<date id="4F2096C69B1D1E42C2E2FEA1FAF531AA" box="[1273,1372,369,395]" pageId="34" pageNumber="495" value="2009-02-04">4.ii.2009</date>
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</collectingDate>
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(
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<collectionCode id="5D8F28C39B1D1E42C375FEA1FA3531AA" box="[1390,1436,369,395]" country="0" name="University of Montana Museum" pageId="34" pageNumber="495">MV</collectionCode>
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<specimenCode id="6B38187D9B1D1E42C68CFE45FE9E3191" box="[151,311,405,432]" collectionCode="MV" country="0" name="University of Montana Museum" pageId="34" pageNumber="495">
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<emphasis id="09EA6C149B1D1E42C68CFE45FE9E3191" bold="true" box="[151,311,405,432]" pageId="34" pageNumber="495">NSWRGCR4</emphasis>
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</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="3B21B0069B1D1E42C6DCFE69FB513386" blockId="34.[151,1437,297,936]" pageId="34" pageNumber="495">
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Molecular data shows that four larvae from Cabbage Tree Ck (
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<collectionCode id="5D8F28C39B1D1E42C56FFE69FC0431F2" box="[884,941,441,467]" country="China" name="Chongqing Normal University" pageId="34" pageNumber="495" type="Herbarium">CTC</collectionCode>
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) (collected
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<date id="4F2096C69B1D1E42C235FE69FB2231F5" box="[1070,1163,441,468]" pageId="34" pageNumber="495" value="2009-01-02">2.i.2009</date>
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) are essentially identical and belong to a clade distinct from any others with molecular sampling. These larvae are smaller than any congener measured from
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<collectingCountry id="4389F0969B1D1E42C75CFDD1FE07323A" box="[327,430,513,539]" name="Australia" pageId="34" pageNumber="495">Australia</collectingCountry>
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||
(
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<tableCitation id="761C85BD9B1D1E42C7A7FDD1FDBC323A" box="[444,533,513,539]" captionStart="TABLE 2" captionStartId="32.[149,242,622,649]" captionTargetBox="[164,1976,660,1609]" captionText="TABLE 2. Morphometric data for larvae of Riethia Kieffer" httpUri="http://table.plazi.org/id/6FE1E08E9B1F1E40C68EFDBEFC8432A8" pageId="34" pageNumber="495" tableUuid="6FE1E08E9B1F1E40C68EFDBEFC8432A8">Table 2</tableCitation>
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). Although it is possible they might be 3
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<superScript id="CCEB1D4E9B1D1E42C5FBFDD0FC47322F" attach="left" box="[992,1006,512,526]" fontSize="6" pageId="34" pageNumber="495">rd</superScript>
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instars, this seems unlikely given that in all other species in which 3
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<superScript id="CCEB1D4E9B1D1E42C7F1FDF4FE513213" attach="left" box="[490,504,548,562]" fontSize="6" pageId="34" pageNumber="495">rd</superScript>
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and 4
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<superScript id="CCEB1D4E9B1D1E42C426FDF4FDE33213" attach="left" box="[573,586,548,562]" fontSize="6" pageId="34" pageNumber="495">th</superScript>
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instars are known, the size ratio of 4
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<superScript id="CCEB1D4E9B1D1E42C5F7FDF4FC503213" attach="left" box="[1004,1017,548,562]" fontSize="6" pageId="34" pageNumber="495">th</superScript>
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to 3
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<superScript id="CCEB1D4E9B1D1E42C237FDF4FB933213" attach="left" box="[1068,1082,548,562]" fontSize="6" pageId="34" pageNumber="495">rd</superScript>
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instar for sclerotised structures (head, postmentum and mandible lengths) is 1.6 (that is 4
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<superScript id="CCEB1D4E9B1D1E42C539FD98FC863277" attach="left" box="[802,815,584,598]" fontSize="6" pageId="34" pageNumber="495">th</superScript>
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is 160% length of 3rd), and is nearer to 2 for the basal antennal segment. The small larva ‘A’ is only a negligible outlier to the smallest other 4
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<superScript id="CCEB1D4E9B1D1E42C246FDBCFBC3325B" attach="left" box="[1117,1130,620,634]" fontSize="6" pageId="34" pageNumber="495">th</superScript>
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instars described (
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<tableCitation id="761C85BD9B1D1E42C323FDBDFA2632A6" box="[1336,1423,621,647]" captionStart="TABLE 2" captionStartId="32.[149,242,622,649]" captionTargetBox="[164,1976,660,1609]" captionText="TABLE 2. Morphometric data for larvae of Riethia Kieffer" httpUri="http://table.plazi.org/id/6FE1E08E9B1F1E40C68EFDBEFC8432A8" pageId="34" pageNumber="495" tableUuid="6FE1E08E9B1F1E40C68EFDBEFC8432A8">Table 2</tableCitation>
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). The larvae show some unexpected variability in morphology: the antennal flagellum can extend to the base of the 4
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<superScript id="CCEB1D4E9B1D1E42C6BEFD64FF1B32E3" attach="left" box="[165,178,692,706]" fontSize="6" pageId="34" pageNumber="495">th</superScript>
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antennal segment or significantly beyond the terminal, and the clypeus approximates that of
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<taxonomicName id="FC9ECB859B1D1E42C2DDFD66FA3432EE" authorityName="Cranston" authorityYear="2019" box="[1222,1437,693,719]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="495" phylum="Arthropoda" rank="species" species="queenslandensis">
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<emphasis id="09EA6C149B1D1E42C2DDFD66FA3432EE" box="[1222,1437,693,719]" italics="true" pageId="34" pageNumber="495">R. queenslandensis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="A3A5AC839B1D1E42C6BBFD09FF5432D5" box="[160,253,729,756]" captionStart="FIGURE 6" captionStartId="15.[151,250,1892,1917]" captionTargetBox="[163,1417,176,1843]" captionTargetId="figure@15.[151,1436,166,1873]" captionTargetPageId="15" captionText="FIGURE 6. Riethia spp. Larva. Mid-dorsal head:A. R. donedwardi, sp.n., B. R. noongar sp.n.. Clypeus: C. R. larval sp. ‘C’, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. hamodivisa sp.n., H. R. neocaledonica sp.n., I. R. noongar sp.n., J. R. paluma sp.n., K. R. phengari sp.n., L. R. plumosa Freeman, M. R. queenslandensis sp.n., N. R. stictoptera Kieffer, O. R. wazeylandica sp.n., P. R. zeylandica Freeman." httpUri="https://zenodo.org/record/3351070/files/figure.png" pageId="34" pageNumber="495">Fig. 6M</figureCitation>
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) in shape and size, especially with regard to the curved posterior boundary but with weaker antero-lateral flare. Given that these 4th instars can be keyed, significant features from the antenna can be recognised as being the low AR of 0.9–1.1, 3
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<superScript id="CCEB1D4E9B1D1E42C7B1FCF0FE11330F" attach="left" box="[426,440,800,814]" fontSize="6" pageId="34" pageNumber="495">rd</superScript>
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||
segment shorter than the 2
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||
<superScript id="CCEB1D4E9B1D1E42C4FFFCF0FD5D330F" attach="left" box="[740,756,800,814]" fontSize="6" pageId="34" pageNumber="495">nd</superScript>
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and with an evenly and completely sclerotised wall to the 3
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<superScript id="CCEB1D4E9B1D1E42C394FCF0FA34330F" attach="none" box="[1423,1437,800,814]" fontSize="6" pageId="34" pageNumber="495">rd</superScript>
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segment. Morphology of this larval molecular cluster suggests 2 additional unreared larvae (lacking molecular data) belong to the clade. The occurrence of this larval
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<typeStatus id="E4250EA49B1D1E42C4D9FCBAFD5B33A5" box="[706,754,874,900]" pageId="34" pageNumber="495">type</typeStatus>
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in both Cabbage Tree and Rutherford Creeks, but seemingly nowhere else, cannot allow association by sympatry as both sites support several coexisting species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |