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<document id="0DC0CA119FDFB1ECC27DCA86F80BD9D6" ID-CLB-Dataset="22391" ID-DOI="10.5852/ejt.2015.139" ID-GBIF-Dataset="8a8a87b6-7e06-4004-b74e-4bb7b0eca921" ID-ISSN="2118-9773" ID-Zenodo-Dep="3785506" ID-ZooBank="F7065B0E-FCCC-45EC-9C4F-19F6C9B972B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588622268591" checkinUser="carolina" docAuthor="Vandenspiegel, D., Golovatch, S. I. &amp; Fiemapong, A. R. Nzoko" docDate="2015" docId="69391077FFA79B06FD95F9E5FE5AA2E3" docLanguage="en" docName="ejt-2015-139.pdf.imf" docOrigin="European Journal of Taxonomy 139" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Campodesmus alobatus Vandenspiegel &amp; Golovatch &amp; Fiemapong 2015, sp. nov." docType="treatment" docUuid="5483AFA3-1222-4B5F-B7C7-B85EEE2FFB58" docUuidSource="ZooBank" docVersion="6" lastPageNumber="10" masterDocId="9500680FFFA29B0FFFCFFFFEFFF8A610" masterDocTitle="Two new species, including one representing a new genus, of the West African millipede family Campodesmidae (Diplopoda: Polydesmida)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="6" updateTime="1698838444074" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D2BECA1871E66C6A987F13A4DA86372F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2271D7593F28D7631EF1396C0F61F243">
<mods:title id="30303E2B72CE229C0790EF0A75805C22">Two new species, including one representing a new genus, of the West African millipede family Campodesmidae (Diplopoda: Polydesmida)</mods:title>
</mods:titleInfo>
<mods:name id="485B9506D3D2D0275BCDB2EF51ED036C" type="personal">
<mods:role id="3B8395BF1809EC034086AE7F617D26D6">
<mods:roleTerm id="4E2C50BC7DC6D0A31DAFFE1E3662646A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FB1BBBF7DE716B943B6B6E8AD09A68C4">Vandenspiegel, D.</mods:namePart>
<mods:nameIdentifier id="F73E47D1CFC97799DB38A8A7779DC7DA" type="LSID">CE8C3D01-28AD-43F7-9D4F-04802E68CB1A</mods:nameIdentifier>
<mods:affiliation id="EF8B941FB4B4ABE63149F9419ABB16FD">Musée Royal de l’Afrique Centrale, B- 3480 Tervuren, Belgique. Email: didier. van. den. spiegel @ africamuseum. be (corresponding author) &amp; urn: lsid: zoobank. org: author: CE 8 C 3 D 01 - 28 AD- 43 F 7 - 9 D 4 F- 04802 E 68 CB 1 A</mods:affiliation>
</mods:name>
<mods:name id="6A7397E911CB7254672A5E6BE507D62E" type="personal">
<mods:role id="0FBF8EC5150650B7FE354872C0D9B4F0">
<mods:roleTerm id="5AEFC6CE0337D3BA7D9B88923E1E4529">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="657EFA96294273D554E63BE2AE143CD2">Golovatch, S. I.</mods:namePart>
<mods:nameIdentifier id="079C3A96022D9F50413D1D284E37FD75" type="LSID">71532F45-BDD5-415D-BC54-86256E5D5D4A</mods:nameIdentifier>
<mods:affiliation id="073B8A027E84CE6C50E68C870B6436B2">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia. Email: sgolovatch @ yandex. ru (corresponding author) &amp; urn: lsid: zoobank. org: author: 71532 F 45 - BDD 5 - 415 D-BC 54 - 86256 E 5 D 5 D 4 A</mods:affiliation>
</mods:name>
<mods:name id="6E8B3B01BCB14912D5B2C838BD7A1F68" type="personal">
<mods:role id="6C3BC6A30D320FB5345945C43B12EF23">
<mods:roleTerm id="51205528D0D82B3883FED76C4C05F7E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="58116B679A7C820D5E514DB2D0D8FE9A">Fiemapong, A. R. Nzoko</mods:namePart>
<mods:nameIdentifier id="E61C3A6600D732DD160991386C503672" type="LSID">ECDC7821-D8DA-49D4-9D6A-13A6E732863E</mods:nameIdentifier>
<mods:affiliation id="DA6CDE7EB8515F6DC6BCC5639BBB9261">Laboratoire de Zoologie, Université Yaoundé 1, BP 812, Cameroon. Email: armandnzoko @ gmail. com &amp; urn: lsid: zoobank. org: author: ECDC 7821 - D 8 DA- 49 D 4 - 9 D 6 A- 13 A 6 E 732863 E</mods:affiliation>
</mods:name>
<mods:typeOfResource id="5673738AE019448D5634E07C71BCF543">text</mods:typeOfResource>
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<mods:titleInfo id="61F981491E3C19AEE0017DB9C85C64AC">
<mods:title id="1326E2A6A9E5FFCD657C6108545FADA0">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="C8E242FAD50997D9AED2FEBDEAEB2A87">
<mods:date id="044FD2806F1AC887F55994C3BD1033AB">2015</mods:date>
<mods:detail id="571ABC917E58E30E17BD75DC0BFFC6CA" type="pubDate">
<mods:number id="DF7E3077CCD60C6B6599107A305DA89D">2015-09-28</mods:number>
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<mods:number id="A38A4DCE5BD8A746BE80BEE46306480C">139</mods:number>
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<mods:start id="1D2695B618D67A89D5D837B28BEFB2BA">1</mods:start>
<mods:end id="898F32968BC7F6694BC668BB546ADDED">11</mods:end>
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<mods:classification id="20F7219945992981DDE903F6B5DEDF05">journal article</mods:classification>
<mods:identifier id="0D59ABF3C3617142C9E8C71C7F175293" type="CLB-Dataset">22391</mods:identifier>
<mods:identifier id="C207352910928CF3E204028A4AF75306" type="DOI">10.5852/ejt.2015.139</mods:identifier>
<mods:identifier id="763F024A31752B14E227736DA27A6BE9" type="GBIF-Dataset">8a8a87b6-7e06-4004-b74e-4bb7b0eca921</mods:identifier>
<mods:identifier id="41831B2F87E17F11BDA5F57AAEE6594C" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="F0C577D5AFD11CB2145A0D61EA6C04FC" type="Zenodo-Dep">3785506</mods:identifier>
<mods:identifier id="B4D71C269CD2E0E45B248173FCC4213D" type="ZooBank">F7065B0E-FCCC-45EC-9C4F-19F6C9B972B3</mods:identifier>
</mods:mods>
<treatment id="69391077FFA79B06FD95F9E5FE5AA2E3" ID-DOI="http://doi.org/10.5281/zenodo.3794595" ID-GBIF-Taxon="163943971" ID-Zenodo-Dep="3794595" LSID="urn:lsid:zoobank.org:act:5483AFA3-1222-4B5F-B7C7-B85EEE2FFB58" httpUri="http://treatment.plazi.org/id/69391077FFA79B06FD95F9E5FE5AA2E3" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6">
<subSubSection id="A98AF2EAFFA79B0AFD95F9E5FC22A026" box="[602,986,1563,1590]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="E12FA161FFA79B0AFD95F9E5FC22A026" blockId="5.[367,1220,1563,1660]" box="[602,986,1563,1590]" pageId="5" pageNumber="6">
<heading id="BA67160DFFA79B0AFD95F9E5FC22A026" box="[602,986,1563,1590]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
<taxonomicName id="2690DAE2FFA79B0AFD95F9E5FC80A026" authority="Vandenspiegel &amp; Golovatch &amp; Fiemapong, 2015" authorityName="Vandenspiegel &amp; Golovatch &amp; Fiemapong" authorityYear="2015" box="[602,888,1563,1590]" class="Diplopoda" family="Campodesmidae" genus="Campodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="alobatus" status="sp. nov.">
<emphasis id="D3E47D73FFA79B0AFD95F9E5FC80A026" bold="true" box="[602,888,1563,1590]" italics="true" pageId="5" pageNumber="6">Campodesmus alobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8D7C008FFA79B0AFCB0F9E2FC22A026" box="[895,986,1564,1590]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A98AF2EAFFA79B0AFEA0F9C1FCB5A06C" pageId="5" pageNumber="6" type="description">
<paragraph id="E12FA161FFA79B0AFEA0F9C1FB3CA04A" blockId="5.[367,1220,1563,1660]" box="[367,1220,1599,1626]" pageId="5" pageNumber="6">
<uri id="9501AD63FFA79B0AFEA0F9C1FB3CA04A" box="[367,1220,1599,1626]" pageId="5" pageNumber="6">
urn:lsid:zoobank.org:act:
<uuid id="95369BB4FFA79B0AFD40F9C1FB3CA04A" box="[655,1220,1599,1626]" pageId="5" pageNumber="6">5483AFA3-1222-4B5F-B7C7-B85EEE2FFB58</uuid>
</uri>
</paragraph>
<paragraph id="E12FA161FFA79B0AFD28F99CFCB5A06C" blockId="5.[367,1220,1563,1660]" box="[743,845,1634,1660]" pageId="5" pageNumber="6">
<figureCitation id="79ABBDE4FFA79B0AFD28F99CFCB5A06C" box="[743,845,1634,1660]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[189,232,1965,1991]" captionStartId-1="7.[189,232,1824,1850]" captionStartId-2="8.[189,232,1929,1955]" captionTargetBox-0="[189,1399,919,1925]" captionTargetBox-1="[257,1331,265,1784]" captionTargetBox-2="[189,1382,1355,1889]" captionTargetId-0="figure@6.[189,1399,919,1925]" captionTargetId-1="figure@7.[257,1331,265,1784]" captionTargetId-2="figure@8.[189,1399,1355,1889]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Fig. 4. Habitus of Campodesmus alobatus sp. nov., ♂, holotype, lateral view. Photo by J. Brecko. Scale bar = 1.0 mm." captionText-1="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." captionText-2="Fig. 6. Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A–B. Left gonopod, ventrolateral and submesal views. C. Right gonopod, mesal view.Scale bars:A–B = 0.1 mm; C = 0.2 mm.Abbreviations are explained in the text." figureDoi-0="http://doi.org/10.5281/zenodo.3785514" figureDoi-1="http://doi.org/10.5281/zenodo.3785516" figureDoi-2="http://doi.org/10.5281/zenodo.3785518" httpUri-0="https://zenodo.org/record/3785514/files/figure.png" httpUri-1="https://zenodo.org/record/3785516/files/figure.png" httpUri-2="https://zenodo.org/record/3785518/files/figure.png" pageId="5" pageNumber="6">Figs 4–6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A98AF2EAFFA79B0AFF72F95AFAF1A147" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="E12FA161FFA79B0AFF72F95AFECEA0AE" blockId="5.[189,310,1700,1726]" box="[189,310,1700,1726]" pageId="5" pageNumber="6">
<heading id="BA67160DFFA79B0AFF72F95AFECEA0AE" bold="true" box="[189,310,1700,1726]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="D3E47D73FFA79B0AFF72F95AFECEA0AE" bold="true" box="[189,310,1700,1726]" pageId="5" pageNumber="6">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="E12FA161FFA79B0AFF72F92CFAF1A147" blockId="5.[189,1399,1746,1879]" pageId="5" pageNumber="6">Differs from other members of the genus in the complete lack of a parabasal process/lobe on the gonopod telopodite which is not directed mesally, but held subparallel to the main body axis, coupled with much more numerous, nearly undifferentiated and low tubercles on metaterga and the presence of deeply incised and clearly separated antero- and caudolateral lobulations on the postcollum paranota.</paragraph>
</subSubSection>
<subSubSection id="A98AF2EAFFA79B0AFF72F881FE50A1FB" pageId="5" pageNumber="6" type="etymology">
<paragraph id="E12FA161FFA79B0AFF72F881FEBCA189" blockId="5.[189,324,1919,1945]" box="[189,324,1919,1945]" pageId="5" pageNumber="6">
<heading id="BA67160DFFA79B0AFF72F881FEBCA189" bold="true" box="[189,324,1919,1945]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="D3E47D73FFA79B0AFF72F881FEBCA189" bold="true" box="[189,324,1919,1945]" pageId="5" pageNumber="6">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="E12FA161FFA79B0AFF72F853FE50A1FB" blockId="5.[189,1399,1965,2027]" pageId="5" pageNumber="6">The name of this species emphasizes the absence of a dorsal/lateral parabasal process/lobe from the gonopod telopodite.</paragraph>
</subSubSection>
<subSubSection id="A98AF2EAFFA49B09FF72FEF5FB40A455" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="E12FA161FFA49B09FF72FEF5FE52A735" blockId="6.[189,426,267,293]" box="[189,426,267,293]" pageId="6" pageNumber="7">
<heading id="BA67160DFFA49B09FF72FEF5FE52A735" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="D3E47D73FFA49B09FF72FEF5FE52A735" bold="true" box="[189,426,267,293]" pageId="6" pageNumber="7">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="51F8AB3CFFA49B09FF25FEC4FC2DA78A" ID-GBIF-Occurrence="2610058768" collectingDate="1994-11-30" collectingMethod="pitfall" collectionCode="R" collectorName="Jocque &amp; N. Seabe" country="Ivory Coast" latitude="6.5833335" location="Appouesso" longLatPrecision="1304" longitude="-3.4666667" pageId="6" pageNumber="7" specimenCode="MRAC 17.264" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="E12FA161FFA49B09FF25FEC4FEA4A744" blockId="6.[189,1398,314,410]" box="[234,348,314,340]" pageId="6" pageNumber="7">
<emphasis id="D3E47D73FFA49B09FF25FEC4FEA4A744" bold="true" box="[234,348,314,340]" pageId="6" pageNumber="7">
<typeStatus id="3E2B1FC3FFA49B09FF25FEC4FEA4A744" box="[234,348,314,340]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="E12FA161FFA49B09FF72FEA3FC2DA78A" blockId="6.[189,1398,314,410]" pageId="6" pageNumber="7">
<collectingCountry id="9987E1F1FFA49B09FF72FEA3FE78A767" box="[189,384,349,375]" name="Ivory Coast" pageId="6" pageNumber="7">IVORY COAST</collectingCountry>
:
<specimenCount id="F7966AE8FFA49B09FE5DFEA3FE52A767" box="[402,426,349,375]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
,
<location id="E44FF7BAFFA49B09FE78FEA3FDC4A767" LSID="urn:lsid:plazi:treatment:69391077FFA79B06FD95F9E5FE5AA2E3:E44FF7BAFFA49B09FE78FEA3FDC4A767" box="[439,572,349,375]" country="Ivory Coast" latitude="6.5833335" longLatPrecision="1304" longitude="-3.4666667" name="Appouesso" pageId="6" pageNumber="7">Appouesso</location>
,
<geoCoordinate id="84A4C7A6FFA49B09FD84FEA3FD44A767" box="[587,700,348,375]" degrees="06" direction="north" minutes="35" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="6.5833335">N 06°35’</geoCoordinate>
,
<geoCoordinate id="84A4C7A6FFA49B09FD04FEA3FCA9A767" box="[715,849,349,375]" degrees="003" direction="west" minutes="28" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="-3.4666667">W 003°28’</geoCoordinate>
, Forêt Classée Bossematié, station 5A,
<collectingMethod id="38D1D976FFA49B09FAFFFEA2FA8EA766" box="[1328,1398,348,374]" pageId="6" pageNumber="7">pitfall</collectingMethod>
trapping,
<date id="952E87A1FFA49B09FEE2FE7EFE36A78A" box="[301,462,384,410]" pageId="6" pageNumber="7" value="1994-11-30">
<collectingDate id="856A7E49FFA49B09FEE2FE7EFE36A78A" box="[301,462,384,410]" pageId="6" pageNumber="7" value="1994-11-30">
<date id="952E87A1FFA49B09FEE2FE7EFE73A78A" box="[301,395,384,410]" day="30" month="11" monthName="Nov" pageId="6" pageNumber="7" year="1930">
30
<date id="952E87A1FFA49B09FE9DFE7EFE73A78A" box="[338,395,384,410]" month="11" monthName="Nov" pageId="6" pageNumber="7" year="1994">Nov.</date>
</date>
1994
</collectingDate>
</date>
, leg. R.
<collectorName id="4C65C4B7FFA49B09FDE0FE7EFD78A78A" box="[559,640,384,410]" pageId="6" pageNumber="7">Jocqué</collectorName>
&amp;
<collectorName id="4C65C4B7FFA49B09FD69FE7EFCEBA78A" box="[678,787,384,410]" pageId="6" pageNumber="7">N. Séabé</collectorName>
,
<specimenCode id="B136091AFFA49B09FCD0FE7EFC29A78A" box="[799,977,384,410]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="7" type="Museum">MRAC 17.264</specimenCode>
.
</paragraph>
</materialsCitation>
<paragraph id="E12FA161FFA49B09FF25FE3FFE9FA7CB" blockId="6.[189,1398,449,582]" box="[234,359,449,475]" pageId="6" pageNumber="7">
<emphasis id="D3E47D73FFA49B09FF25FE3FFE9FA7CB" bold="true" box="[234,359,449,475]" pageId="6" pageNumber="7">
<typeStatus id="3E2B1FC3FFA49B09FF25FE3FFE9FA7CB" box="[234,359,449,475]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="E12FA161FFA49B09FF72FE1BFB40A455" blockId="6.[189,1398,449,582]" pageId="6" pageNumber="7">
<collectingCountry id="9987E1F1FFA49B09FF72FE1BFE84A7EF" box="[189,380,485,511]" name="Ivory Coast" pageId="6" pageNumber="7">IVORY COAST</collectingCountry>
:
<specimenCount id="F7966AE8FFA49B09FE45FE1BFE4FA7EF" box="[394,439,485,511]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
, same locality, station 1A,
<date id="952E87A1FFA49B09FD22FE1BFC7DA7EF" box="[749,901,485,511]" pageId="6" pageNumber="7" value="1994-10-31">
<date id="952E87A1FFA49B09FD22FE1BFCBAA7EF" box="[749,834,485,511]" day="31" month="10" monthName="Oct" pageId="6" pageNumber="7" year="1931">
31
<date id="952E87A1FFA49B09FCDFFE1BFCBAA7EF" box="[784,834,485,511]" month="10" monthName="Oct" pageId="6" pageNumber="7" year="1994">Oct.</date>
</date>
1994
</date>
,
<collectionCode id="878139A4FFA49B09FC5FFE1BFC1EA7EF" box="[912,998,485,511]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="7" type="Museum">MRAC</collectionCode>
17.272;
<specimenCount id="F7966AE8FFA49B09FB84FE1BFB8FA7EF" box="[1099,1143,485,511]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
, same locality, station 5B,
<date id="952E87A1FFA49B09FF22FDF6FE7EA432" box="[237,390,520,546]" pageId="6" pageNumber="7" value="1994-10-30">
<date id="952E87A1FFA49B09FF22FDF6FEBBA432" box="[237,323,520,546]" day="30" month="10" monthName="Oct" pageId="6" pageNumber="7" year="1930">
30
<date id="952E87A1FFA49B09FEDEFDF6FEBBA432" box="[273,323,520,546]" month="10" monthName="Oct" pageId="6" pageNumber="7" year="1994">Oct.</date>
</date>
1994
</date>
,
<collectionCode id="878139A4FFA49B09FE5EFDF6FE1FA432" box="[401,487,520,546]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="7" type="Museum">MRAC</collectionCode>
17.276;
<specimenCount id="F7966AE8FFA49B09FD82FDF6FD80A432" box="[589,632,520,546]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
(
<collectionCode id="878139A4FFA49B09FD49FDF6FD3DA432" box="[646,709,520,546]" pageId="6" pageNumber="7">SEM</collectionCode>
) and
<specimenCount id="F7966AE8FFA49B09FCCAFDF6FCCAA432" box="[773,818,520,546]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
, same locality,
<date id="952E87A1FFA49B09FC2BFDF6FB85A432" box="[996,1149,520,546]" pageId="6" pageNumber="7" value="1994-10-31">
<date id="952E87A1FFA49B09FC2BFDF6FBC2A432" box="[996,1082,520,546]" day="31" month="10" monthName="Oct" pageId="6" pageNumber="7" year="1931">
31
<date id="952E87A1FFA49B09FBC7FDF6FBC2A432" box="[1032,1082,520,546]" month="10" monthName="Oct" pageId="6" pageNumber="7" year="1994">Oct.</date>
</date>
1994
</date>
,
<collectionCode id="878139A4FFA49B09FB47FDF6FB26A432" box="[1160,1246,520,546]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="7" type="Museum">MRAC</collectionCode>
17.278;
<specimenCount id="F7966AE8FFA49B09FA8BFDF6FA89A432" box="[1348,1393,520,546]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
, same locality, station 5D,
<date id="952E87A1FFA49B09FE22FDD5FD70A455" box="[493,648,555,581]" pageId="6" pageNumber="7" value="1994-10-31">31 Oct. 1994</date>
,
<collectionCode id="878139A4FFA49B09FD5BFDD5FD17A455" box="[660,751,555,581]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="6" pageNumber="7">ZMUM</collectionCode>
ρ2665. All leg.
<collectionCode id="878139A4FFA49B09FC62FDD5FC3CA455" box="[941,964,555,581]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="7">R</collectionCode>
. Jocqué &amp; N. Séabé.
</paragraph>
</subSubSection>
<subSubSection id="A98AF2EAFFA49B06FF72FD92FE95A50C" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7" type="description">
<paragraph id="E12FA161FFA49B09FF72FD92FEB6A496" blockId="6.[189,334,620,646]" box="[189,334,620,646]" pageId="6" pageNumber="7">
<heading id="BA67160DFFA49B09FF72FD92FEB6A496" bold="true" box="[189,334,620,646]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="D3E47D73FFA49B09FF72FD92FEB6A496" bold="true" box="[189,334,620,646]" pageId="6" pageNumber="7">Description</emphasis>
</heading>
</paragraph>
<paragraph id="E12FA161FFA49B09FF72FD65FADFA50F" blockId="6.[189,1399,667,799]" pageId="6" pageNumber="7">
Length of
<typeStatus id="3E2B1FC3FFA49B09FEF2FD65FE5CA4A5" box="[317,420,667,693]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
<emphasis id="D3E47D73FFA49B09FE61FD62FE3AA4A5" box="[430,450,668,693]" italics="true" pageId="6" pageNumber="7">c.</emphasis>
<quantity id="26680C84FFA49B09FE03FD65FDDDA4A5" box="[460,549,667,693]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="7" unit="mm" value="17.0">17 mm</quantity>
, width on midbody pro- and metazonae 1.7 and
<quantity id="26680C84FFA49B09FBBAFD65FB2DA4A5" box="[1141,1237,667,693]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="7" unit="mm" value="3.6">3.6 mm</quantity>
, respectively.
<typeStatus id="3E2B1FC3FFA49B09FF72FD41FED7A4C9" box="[189,303,703,729]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
(males) also
<emphasis id="D3E47D73FFA49B09FE19FD41FE12A4C8" box="[470,490,703,728]" italics="true" pageId="6" pageNumber="7">c.</emphasis>
<quantity id="26680C84FFA49B09FE3AFD40FDB3A4C9" box="[501,587,702,729]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="7" unit="mm" value="17.0">17 mm</quantity>
long, width on midbody pro- and metazonae 1.6–1.7 and 3.0 mm, respectively. Coloration from light grey-brown to brown (
<figureCitation id="79ABBDE4FFA49B09FC57FD1FFC10A4EB" box="[920,1000,737,763]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[189,1399,919,1925]" captionTargetId="figure@6.[189,1399,919,1925]" captionTargetPageId="6" captionText="Fig. 4. Habitus of Campodesmus alobatus sp. nov., ♂, holotype, lateral view. Photo by J. Brecko. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3785514" httpUri="https://zenodo.org/record/3785514/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
); clypeolabral region, antennae, prozonae, venter, legs and tip of epiproct pallid to light yellow; stricture as brown as metazonae.
</paragraph>
<caption id="B5EFF1E9FFA49B09FF72F853FE9BA1FB" ID-DOI="http://doi.org/10.5281/zenodo.3785514" ID-Zenodo-Dep="3785514" httpUri="https://zenodo.org/record/3785514/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1965,1991]" targetBox="[189,1399,919,1925]" targetPageId="6">
<paragraph id="E12FA161FFA49B09FF72F853FE9BA1FB" blockId="6.[189,1399,1965,2027]" pageId="6" pageNumber="7">
<emphasis id="D3E47D73FFA49B09FF72F853FEF2A1D7" bold="true" box="[189,266,1965,1991]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
Habitus of
<taxonomicName id="2690DAE2FFA49B09FE5CF853FD52A1D7" authority="Vandenspiegel &amp; Golovatch &amp; Fiemapong, 2015" authorityName="Vandenspiegel &amp; Golovatch &amp; Fiemapong" authorityYear="2015" box="[403,682,1965,1991]" class="Diplopoda" family="Campodesmidae" genus="Campodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="alobatus" status="sp. nov.">
<emphasis id="D3E47D73FFA49B09FE5CF853FD52A1D7" box="[403,682,1965,1991]" italics="true" pageId="6" pageNumber="7">Campodesmus alobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8D7C008FFA49B09FD7DF850FCF7A1D8" box="[690,783,1966,1992]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype, lateral view. Photo by J. Brecko. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="B5EFF1E9FFA59B08FF72F8DEFB25A1FB" ID-DOI="http://doi.org/10.5281/zenodo.3785516" ID-Zenodo-Dep="3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1824,1850]" targetBox="[257,1331,265,1784]" targetPageId="7">
<paragraph id="E12FA161FFA59B08FF72F8DEFB25A1FB" blockId="7.[189,1399,1824,2027]" pageId="7" pageNumber="8">
<emphasis id="D3E47D73FFA59B08FF72F8DEFEF5A12A" bold="true" box="[189,269,1824,1850]" pageId="7" pageNumber="8">Fig. 5.</emphasis>
SEM micrographs of
<taxonomicName id="2690DAE2FFA59B08FDD3F8DEFCCEA12A" authority="Vandenspiegel &amp; Golovatch &amp; Fiemapong, 2015" authorityName="Vandenspiegel &amp; Golovatch &amp; Fiemapong" authorityYear="2015" box="[540,822,1824,1850]" class="Diplopoda" family="Campodesmidae" genus="Campodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alobatus" status="sp. nov.">
<emphasis id="D3E47D73FFA59B08FDD3F8DEFCCEA12A" box="[540,822,1824,1850]" italics="true" pageId="7" pageNumber="8">Campodesmus alobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8D7C008FFA59B08FC8FF8DFFC67A12B" box="[832,927,1825,1851]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂, MRAC 17.278.
<emphasis id="D3E47D73FFA59B08FAC4F8DEFAD8A12A" bold="true" box="[1291,1312,1824,1850]" pageId="7" pageNumber="8">A</emphasis>
,
<emphasis id="D3E47D73FFA59B08FAFDF8DEFABFA12A" bold="true" box="[1330,1351,1824,1850]" pageId="7" pageNumber="8">D</emphasis>
,
<emphasis id="D3E47D73FFA59B08FA97F8DEFA97A12A" bold="true" box="[1368,1391,1824,1850]" pageId="7" pageNumber="8">G</emphasis>
. Anterior part of body, lateral, dorsal and ventral views, respectively.
<emphasis id="D3E47D73FFA59B08FC1BF8BDFC10A14D" bold="true" box="[980,1000,1859,1885]" pageId="7" pageNumber="8">B</emphasis>
,
<emphasis id="D3E47D73FFA59B08FC3AF8BDFBF1A14D" bold="true" box="[1013,1033,1859,1885]" pageId="7" pageNumber="8">E</emphasis>
,
<emphasis id="D3E47D73FFA59B08FBD9F8BDFBD5A14D" bold="true" box="[1046,1069,1859,1885]" pageId="7" pageNumber="8">H</emphasis>
. Midbody segments, lateral, dorsal and ventral views, respectively.
<emphasis id="D3E47D73FFA59B08FD48F898FD64A190" bold="true" box="[647,668,1894,1920]" pageId="7" pageNumber="8">C</emphasis>
,
<emphasis id="D3E47D73FFA59B08FD64F899FD45A191" bold="true" box="[683,701,1895,1921]" pageId="7" pageNumber="8">F</emphasis>
,
<emphasis id="D3E47D73FFA59B08FD02F899FD20A191" bold="true" box="[717,728,1895,1921]" pageId="7" pageNumber="8">I</emphasis>
. Posterior part of body, lateral, dorsal and ventral views, respectively.
<emphasis id="D3E47D73FFA59B08FE99F874FE9DA1B4" bold="true" box="[342,357,1930,1956]" pageId="7" pageNumber="8">J</emphasis>
. Cross-section of a midbody segment, caudal view.
<emphasis id="D3E47D73FFA59B08FC0FF874FC2FA1B4" bold="true" box="[960,983,1930,1956]" pageId="7" pageNumber="8">K</emphasis>
. Midbody paratergum, lateral view.
<emphasis id="D3E47D73FFA59B08FF72F853FF29A1D7" bold="true" box="[189,209,1965,1991]" pageId="7" pageNumber="8">L</emphasis>
. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view.
<emphasis id="D3E47D73FFA59B08FA9CF853FA97A1D7" bold="true" box="[1363,1391,1965,1991]" pageId="7" pageNumber="8">M</emphasis>
. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm.
</paragraph>
</caption>
<paragraph id="E12FA161FFAA9B07FF72FEF2FA8FA735" blockId="8.[189,1399,267,294]" box="[189,1399,267,294]" pageId="8" pageNumber="9">HEAD. Densely setose, clypeolabral region smooth, remaining parts densely microdenticulate-</paragraph>
<paragraph id="E12FA161FFAA9B07FF72FED0FEABA7A3" blockId="8.[189,1399,302,435]" pageId="8" pageNumber="9">
microgranulate, interantennal isthmus
<emphasis id="D3E47D73FFAA9B07FD48FED1FD63A758" box="[647,667,303,328]" italics="true" pageId="8" pageNumber="9">c.</emphasis>
1.5 times as broad as diameter of antennal socket. Antennae short and strongly clavate, C-shaped, antennomere 5 largest and longest, but only a little larger than 6
<superScript id="16E50C29FFAA9B07FAADFEAEFA97A74E" attach="none" box="[1378,1391,336,350]" fontSize="6" pageId="8" pageNumber="9">th</superScript>
, both with a dorso-apical group of tiny bacilliform sensilla; apical segment with usual four sensory cones (
<figureCitation id="79ABBDE4FFAA9B07FF0AFE66FEEAA7A2" box="[197,274,408,434]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[189,1399,919,1925]" captionTargetId="figure@6.[189,1399,919,1925]" captionTargetPageId="6" captionText="Fig. 4. Habitus of Campodesmus alobatus sp. nov., ♂, holotype, lateral view. Photo by J. Brecko. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3785514" httpUri="https://zenodo.org/record/3785514/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
,
<figureCitation id="79ABBDE4FFAA9B07FED1FE66FEBEA7A3" box="[286,326,408,435]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">5G</figureCitation>
).
</paragraph>
<paragraph id="E12FA161FFAA9B07FF72FE24FC33A30E" blockId="8.[189,1399,473,1310]" pageId="8" pageNumber="9">
BODY. With 20 segments (male). In width, head &lt;&lt; collum = segment 2 &lt;3 &lt;4 &lt;5–17; thereafter body rapidly tapering towards telson. Collum transversely ellipsoid, subflabellate, covering most of head from above, dorsal surface rather regularly and densely tuberculate, with about 4–5 transverse rows of round, setigerous, undifferentiated tubercles lying between a moderately crenulate/lobulate and regularly rounded anterior margin and a clearly ribbed and almost straight caudal edge; small, but discernible caudal paranota with a rather sharp lateral tooth (
<figureCitation id="79ABBDE4FFAA9B07FD3EFD77FCC4A4B4" box="[753,828,649,676]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[189,1399,919,1925]" captionTargetId="figure@6.[189,1399,919,1925]" captionTargetPageId="6" captionText="Fig. 4. Habitus of Campodesmus alobatus sp. nov., ♂, holotype, lateral view. Photo by J. Brecko. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3785514" httpUri="https://zenodo.org/record/3785514/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
,
<figureCitation id="79ABBDE4FFAA9B07FC88FD77FC4DA4B4" box="[839,949,649,676]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">5A, D, G</figureCitation>
). Prozonae smooth and poorly shining, stricture finely microgranulate; metazonae dull, densely tuberculate and microgranulate over entire surface; cerotegument well-developed, sometimes in places clothed with a crust of soil; dorsal surface usually with two regular, mixostictic (= not regularly longitudinal) transverse rows of normally similar round tubercles, much like on collum, lying between a fore row of denser and smaller knobs/tubercles behind stricture and a densely ribbed caudal edge represented by a flatter row of oblong tubercules (
<figureCitation id="79ABBDE4FFAA9B07FA87FCC7FF32A568" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
A–F). Paranota 2 not enlarged compared to following ones (
<figureCitation id="79ABBDE4FFAA9B07FC4BFCA3FC37A567" box="[900,975,861,887]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[189,1399,919,1925]" captionTargetId="figure@6.[189,1399,919,1925]" captionTargetPageId="6" captionText="Fig. 4. Habitus of Campodesmus alobatus sp. nov., ♂, holotype, lateral view. Photo by J. Brecko. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3785514" httpUri="https://zenodo.org/record/3785514/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
,
<figureCitation id="79ABBDE4FFAA9B07FC14FCA3FBFBA568" box="[987,1027,861,888]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">5A</figureCitation>
). Postcollum paranota set rather low (at about half of midbody height, thus dorsum being very strongly convex), strongly declivous, continuing the outline of dorsum, but their tips not reaching level of venter (
<figureCitation id="79ABBDE4FFAA9B07FBE6FC5DFB94A5AE" box="[1065,1132,931,958]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
A–C, J); lateral margin of paranota usually 5-lobulate, laterals continuing into even more deeply lobulate caudo- and anterolaterals both at caudal and fore margin, respectively; caudo- and anterolateral lobulations continuing both caudal and fore rows of tubercles at base of paranota, respectively (
<figureCitation id="79ABBDE4FFAA9B07FC40FBF3FC25A237" box="[911,989,1037,1063]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[189,1399,919,1925]" captionTargetId="figure@6.[189,1399,919,1925]" captionTargetPageId="6" captionText="Fig. 4. Habitus of Campodesmus alobatus sp. nov., ♂, holotype, lateral view. Photo by J. Brecko. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3785514" httpUri="https://zenodo.org/record/3785514/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
,
<figureCitation id="79ABBDE4FFAA9B07FC24FBF3FC01A238" box="[1003,1017,1037,1064]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">5</figureCitation>
A–I). Caudal corner of paranota increasingly strongly inclined caudad behind rear tergal margin only in segments 17–19 (
<figureCitation id="79ABBDE4FFAA9B07FB19FBCEFA90A25A" box="[1238,1384,1072,1099]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">Fig. 5C, F, I</figureCitation>
). Tergal setae very short and thin (
<figureCitation id="79ABBDE4FFAA9B07FDF7FBADFD84A27E" box="[568,636,1107,1134]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
A–L). Ozopores inconspicuous, opening flush on tergal surface at about midlength of paranota 5 and 7 off lateral margin. Pleurosternal carinae absent (
<figureCitation id="79ABBDE4FFAA9B07FB66FB88FAFFA281" box="[1193,1287,1142,1169]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">Fig. 5G</figureCitation>
). Limbus microcrenulate, with microserrate tips of crenulations, these latter clearly separated from one another (
<figureCitation id="79ABBDE4FFAA9B07FF09FB43FEDFA2C8" box="[198,295,1213,1240]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">Fig. 5M</figureCitation>
). Epiproct short, small, subtruncate, slightly flattened dorsoventrally, tuberculate dorsally, with a pair of rather large subapical papillae (
<figureCitation id="79ABBDE4FFAA9B07FDB6FB1EFD3CA2EA" box="[633,708,1248,1274]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[189,1399,919,1925]" captionTargetId="figure@6.[189,1399,919,1925]" captionTargetPageId="6" captionText="Fig. 4. Habitus of Campodesmus alobatus sp. nov., ♂, holotype, lateral view. Photo by J. Brecko. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3785514" httpUri="https://zenodo.org/record/3785514/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
,
<figureCitation id="79ABBDE4FFAA9B07FD1FFB1EFCD0A2EA" box="[720,808,1248,1275]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">5C, F, I</figureCitation>
). Hypoproct nearly semi-circular with 1 + 1 caudal setae very distinctly separated and borne on minute knobs (
<figureCitation id="79ABBDE4FFAA9B07FCA3FAFDFC45A30E" box="[876,957,1283,1310]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="8" pageNumber="9">Fig. 5I</figureCitation>
).
</paragraph>
<caption id="B5EFF1E9FFAA9B07FF72F877FE26A1FB" ID-DOI="http://doi.org/10.5281/zenodo.3785518" ID-Zenodo-Dep="3785518" httpUri="https://zenodo.org/record/3785518/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1929,1955]" targetBox="[189,1382,1355,1889]" targetPageId="8">
<paragraph id="E12FA161FFAA9B07FF72F877FE26A1FB" blockId="8.[189,1398,1929,2027]" pageId="8" pageNumber="9">
<emphasis id="D3E47D73FFAA9B07FF72F877FEF2A1B3" bold="true" box="[189,266,1929,1955]" pageId="8" pageNumber="9">Fig. 6.</emphasis>
<taxonomicName id="2690DAE2FFAA9B07FEDDF874FDD1A1B4" authority="Vandenspiegel &amp; Golovatch &amp; Fiemapong, 2015" authorityName="Vandenspiegel &amp; Golovatch &amp; Fiemapong" authorityYear="2015" box="[274,553,1930,1956]" class="Diplopoda" family="Campodesmidae" genus="Campodesmus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alobatus" status="sp. nov.">
<emphasis id="D3E47D73FFAA9B07FEDDF874FDD1A1B4" box="[274,553,1930,1956]" italics="true" pageId="8" pageNumber="9">Campodesmus alobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8D7C008FFAA9B07FDFFF874FD75A1B4" box="[560,653,1930,1956]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂, MRAC 17.278.
<emphasis id="D3E47D73FFAA9B07FC21F877FBDDA1B3" bold="true" box="[1006,1061,1929,1955]" pageId="8" pageNumber="9">A–B</emphasis>
. Left gonopod, ventrolateral and submesal views.
<emphasis id="D3E47D73FFAA9B07FE62F853FE3AA1D7" bold="true" box="[429,450,1965,1991]" pageId="8" pageNumber="9">C</emphasis>
. Right gonopod, mesal view.Scale bars:A–B = 0.1 mm; C = 0.2 mm.Abbreviations are explained in the text.
</paragraph>
</caption>
<paragraph id="E12FA161FFAB9B06FF72FEF5FAE7A782" blockId="9.[189,1399,267,403]" pageId="9" pageNumber="10">
STERNA. Broad, almost twice as broad as coxa length, with a very deep axial impression (
<figureCitation id="79ABBDE4FFAB9B06FB3FFEF5FACFA736" box="[1264,1335,267,294]" captionStart="Fig" captionStartId="7.[189,232,1824,1850]" captionTargetBox="[257,1331,265,1784]" captionTargetId="figure@7.[257,1331,265,1784]" captionTargetPageId="7" captionText="Fig. 5. SEM micrographs of Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Midbody paratergum, lateral view. L. Fine structure of pro- and metazonae, as well as of stricture between them, subdorsal view. M. Limbus and adjacent structures. Scale bars: A–J = 0.5 mm; K–L = 0.1 mm; M = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3785516" httpUri="https://zenodo.org/record/3785516/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
G–I). Gonapophysis on male coxae 2 a small tubercle. Legs short, even in male about 0.9–1.0 times as long as midbody height, very slender; in length, femur&gt; tarsus&gt; prefemur&gt; postfemur = tibia = coxa; claw very small, very slightly curved; ventral surface of tarsi densely setose, but forming no brushes.
</paragraph>
<paragraph id="E12FA161FFAB9B06FF72FE45FE95A50C" blockId="9.[189,1399,443,796]" pageId="9" pageNumber="10">
GONOPOD. Aperture transversely ovoid, large, its lateral and posterior edges slightly elevated, fully concealing gonocoxae and bases of telopodites. Gonopod coxites medium-sized, subcylindrical, fused at base to a small membranous sternal remnant, each with only one long distodorsal seta (
<figureCitation id="79ABBDE4FFAB9B06FB28FDFDFA90A40E" box="[1255,1384,515,542]" captionStart="Fig" captionStartId="8.[189,232,1929,1955]" captionTargetBox="[189,1382,1355,1889]" captionTargetId="figure@8.[189,1399,1355,1889]" captionTargetPageId="8" captionText="Fig. 6. Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A–B. Left gonopod, ventrolateral and submesal views. C. Right gonopod, mesal view.Scale bars:A–B = 0.1 mm; C = 0.2 mm.Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3785518" httpUri="https://zenodo.org/record/3785518/files/figure.png" pageId="9" pageNumber="10">Fig. 6A, B</figureCitation>
). Cannulae long and slender, without peculiarities. Telopodites rather regularly curved caudad,
<emphasis id="D3E47D73FFAB9B06FAE4FDD7FA8EA452" box="[1323,1398,553,578]" italics="true" pageId="9" pageNumber="10">in situ</emphasis>
held subparallel to each other, subcircular, long, slender and quite simple (
<figureCitation id="79ABBDE4FFAB9B06FBD7FDB2FBA7A477" box="[1048,1119,588,615]" captionStart="Fig" captionStartId="8.[189,232,1929,1955]" captionTargetBox="[189,1382,1355,1889]" captionTargetId="figure@8.[189,1399,1355,1889]" captionTargetPageId="8" captionText="Fig. 6. Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A–B. Left gonopod, ventrolateral and submesal views. C. Right gonopod, mesal view.Scale bars:A–B = 0.1 mm; C = 0.2 mm.Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3785518" httpUri="https://zenodo.org/record/3785518/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
). Prefemoral (= densely setose) portion (pf) subcylindrical, rather long, totally devoid of outgrowths. Femorite (fe) with a small parabasal denticle (d) on ventral face and a series of inconspicuous folds (f) on opposite side. Seminal groove (sg) running entirely on mesal side, terminating at about halfway along fe, at base of a small denticle (o), with neither a hairpad nor an accessory seminal chamber. Tip of fe slightly broadened and bifid (
<figureCitation id="79ABBDE4FFAB9B06FECCFCFFFE98A50C" box="[259,352,769,796]" captionStart="Fig" captionStartId="8.[189,232,1929,1955]" captionTargetBox="[189,1382,1355,1889]" captionTargetId="figure@8.[189,1399,1355,1889]" captionTargetPageId="8" captionText="Fig. 6. Campodesmus alobatus sp. nov., paratype, ♂, MRAC 17.278. A–B. Left gonopod, ventrolateral and submesal views. C. Right gonopod, mesal view.Scale bars:A–B = 0.1 mm; C = 0.2 mm.Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3785518" httpUri="https://zenodo.org/record/3785518/files/figure.png" pageId="9" pageNumber="10">Fig. 6C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A98AF2EAFFAB9B06FF72FCBAFE5AA2E3" pageId="9" pageNumber="10" type="discussion">
<paragraph id="E12FA161FFAB9B06FF72FCBAFED7A54E" blockId="9.[189,303,836,862]" box="[189,303,836,862]" pageId="9" pageNumber="10">
<heading id="BA67160DFFAB9B06FF72FCBAFED7A54E" bold="true" box="[189,303,836,862]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="D3E47D73FFAB9B06FF72FCBAFED7A54E" bold="true" box="[189,303,836,862]" pageId="9" pageNumber="10">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="E12FA161FFAB9B06FF72FC8AFAAEA20F" blockId="9.[189,1399,882,1056]" pageId="9" pageNumber="10">
The species described above is the 8
<superScript id="16E50C29FFAB9B06FDA3FC8CFD81A590" attach="left" box="[620,633,882,896]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
known species of
<taxonomicName id="2690DAE2FFAB9B06FC99FC8AFBF9A59E" authorityName="Cook" authorityYear="1896" box="[854,1025,884,910]" class="Diplopoda" family="Campodesmidae" genus="Campodesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="D3E47D73FFAB9B06FC99FC8AFBF9A59E" box="[854,1025,884,910]" italics="true" pageId="9" pageNumber="10">Campodesmus</emphasis>
</taxonomicName>
which, despite the peculiarities referred to in the diagnosis, remains a rather typical congener. This is primarily evidenced by the gonopod conformation which shows long and subcircular telopodites. Such non-gonopod characters as low paranota, a reduced pore formula, short antennae and legs etc. also attest to the unequivocal assignment of this new species to
<taxonomicName id="2690DAE2FFAB9B06FD83FBFBFD0FA20F" authorityName="Cook" authorityYear="1896" box="[588,759,1029,1055]" class="Diplopoda" family="Campodesmidae" genus="Campodesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="D3E47D73FFAB9B06FD83FBFBFD0FA20F" box="[588,759,1029,1055]" italics="true" pageId="9" pageNumber="10">Campodesmus</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="8501DC90FFAB9B06FCE1FBFBFC25A230" author="Hoffman R. L." box="[814,989,1029,1056]" pageId="9" pageNumber="10" pagination="209 - 212" refId="ref4511" refString="Hoffman R. L. 1951. The diplopod family Campodesmidae (Polydesmida). Journal of the Washington Academy of Sciences 41 (6): 209 - 212." type="journal article" year="1951">Hoffman 1951</bibRefCitation>
;
<bibRefCitation id="8501DC90FFAB9B06FC26FBFBFB7EA20F" author="Schiotz V." box="[1001,1158,1029,1055]" pageId="9" pageNumber="10" pagination="81 - 104" refId="ref4600" refString="Schiotz V. 1966. Notes on West African diplopods I. The family Campodesmidae (Polydesmida). V idensksbelige Meddelelser fra Dansk naturhistorisk Forening 129: 81 - 104." type="journal article" year="1966">Schiøtz 1966</bibRefCitation>
;
<bibRefCitation id="8501DC90FFAB9B06FB5DFBFBFAB1A20F" author="Demange J. M." box="[1170,1353,1029,1055]" pageId="9" pageNumber="10" pagination="191 - 240" refId="ref4439" refString="Demange J. M. 1971. Myriapodes Diplopodes. Le massif des Monts Loma (Sierra Leone). Fascicule I. Memoires de l'Institut Fondamental d'Afrique Noire 86: 191 - 240." type="journal article" year="1971">Demange 1971</bibRefCitation>
).
</paragraph>
<paragraph id="E12FA161FFAB9B06FF72FBB9FE5AA2E3" blockId="9.[189,1399,1095,1267]" pageId="9" pageNumber="10">
The clearly abbreviated seminal groove in
<taxonomicName id="2690DAE2FFAB9B06FD1AFBB6FC9AA271" authority="Vandenspiegel &amp; Golovatch &amp; Fiemapong, 2015" authorityName="Vandenspiegel &amp; Golovatch &amp; Fiemapong" authorityYear="2015" box="[725,866,1095,1122]" class="Diplopoda" family="Campodesmidae" genus="Campodesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="alobatus" status="sp. nov.">
<emphasis id="D3E47D73FFAB9B06FD1AFBB6FC9AA271" box="[725,866,1095,1122]" italics="true" pageId="9" pageNumber="10">C. alobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8D7C008FFAB9B06FCA0FBB6FC2AA272" box="[879,978,1096,1122]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, which only extends to about the midlength of an untwisted gonopod femorite, invites comparisons with other congeners. Regrettably, no such information is available in the literature.
<bibRefCitation id="8501DC90FFAB9B06FD31FB6EFC53A2BB" author="Schiotz V." box="[766,939,1168,1195]" pageId="9" pageNumber="10" pagination="81 - 104" refId="ref4600" refString="Schiotz V. 1966. Notes on West African diplopods I. The family Campodesmidae (Polydesmida). V idensksbelige Meddelelser fra Dansk naturhistorisk Forening 129: 81 - 104." type="journal article" year="1966">Schiøtz (1966)</bibRefCitation>
attempted to group most of the species of
<taxonomicName id="2690DAE2FFAB9B06FF12FB4AFE70A2DE" authorityName="Cook" authorityYear="1896" box="[221,392,1204,1230]" class="Diplopoda" family="Campodesmidae" genus="Campodesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="D3E47D73FFAB9B06FF12FB4AFE70A2DE" box="[221,392,1204,1230]" italics="true" pageId="9" pageNumber="10">Campodesmus</emphasis>
</taxonomicName>
, but more data are needed, including the course of the seminal groove, to reach more secure conclusions.
</paragraph>
</subSubSection>
</treatment>
</document>
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