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<document id="BFB251C8A7737D9510E53441C861D098" ID-CLB-Dataset="32729" ID-DOI="10.11646/zootaxa.4286.4.6" ID-GBIF-Dataset="9d2b00be-244a-4203-962b-1144b3a728ea" ID-ISSN="1175-5326" ID-Zenodo-Dep="828773" ID-ZooBank="692B2C15-8D7C-4577-B68D-62A34CA20396" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500109652088" checkinUser="plazi" docAuthor="Duarte, Adriane Da Fonseca, Moreira, Grazielle Furtado, Cunha, Uemerson Silva Da &amp; Moraes, Gilberto José De" docDate="2017" docId="03D5304D916BFFFCFF79F9EA77F702C9" docLanguage="en" docName="zootaxa.4286.4.6.pdf" docOrigin="Zootaxa 4286 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cosmolaelaps guttulatus Karg 1978" docType="treatment" docVersion="9" lastPageNumber="538" masterDocId="FFEC4835916AFFFFFFEEFFD57261073F" masterDocTitle="Cosmolaelaps Berlese (Mesostigmata: Laelapidae) from southern Brazil, with a new record of a heteromorphic male, description of a new species, and a key to the species reported from that country" masterLastPageNumber="544" masterPageNumber="535" pageNumber="536" updateTime="1698465293361" updateUser="ExternalLinkService">
<mods:mods id="1ED2B31E04A453F5D5427B2E2A6525B2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BE66E8CC4069DF261A113813EF33C2B0">
<mods:title id="F8E194DB2652556FD5F1F454239DAEE9">Cosmolaelaps Berlese (Mesostigmata: Laelapidae) from southern Brazil, with a new record of a heteromorphic male, description of a new species, and a key to the species reported from that country</mods:title>
</mods:titleInfo>
<mods:name id="D2D337E3CC9DA05168D4D6B2B446FD5A" type="personal">
<mods:role id="F3D977DE071783514F1DFE01E7D524D2">
<mods:roleTerm id="F6D87516E5588315EB8BC0B521158DF2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5892696A64062B23B0484833C69640F7">Duarte, Adriane Da Fonseca</mods:namePart>
</mods:name>
<mods:name id="40244D8F26C201E5E79D32057FCC228B" type="personal">
<mods:role id="991366FB46CC13C71D2E3C6951AC1C17">
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</mods:role>
<mods:namePart id="A61687825111D7246C139A77BD01C1EF">Moreira, Grazielle Furtado</mods:namePart>
</mods:name>
<mods:name id="F41F245560A279D60CB7419ED7111BF0" type="personal">
<mods:role id="F11430DDF210452F94D430CF26DB8F99">
<mods:roleTerm id="7DFEB92FE76E1C40DBCDA6E3840E0C3B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="43D1CE77573ACD3D49C27539A97A2984">Cunha, Uemerson Silva Da</mods:namePart>
</mods:name>
<mods:name id="F4E0F0FA9939CF35E0485424B75206AF" type="personal">
<mods:role id="D66EBE2829162AB0B3370303572015F9">
<mods:roleTerm id="D058AEBF3B10A156BC9576989FA7D02F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4CFC683F652FD1F486F943A8817A4935">Moraes, Gilberto José De</mods:namePart>
</mods:name>
<mods:typeOfResource id="87DA844FCDC2A77C81F686ED370B03CE">text</mods:typeOfResource>
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<mods:titleInfo id="D474F277D770B09A6863AEFA9D10B59E">
<mods:title id="601267F86B4365231E210F3BD441067F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="188B70F76CBAEA6988B63E64071CBFAA">
<mods:date id="E7F5DBEB2F33B4AD1B30D805D34285EA">2017</mods:date>
<mods:detail id="76A593212A9A53EDB214D336879565EE" type="volume">
<mods:number id="8A45757161378191B6F719D42AAF3A1B">4286</mods:number>
</mods:detail>
<mods:detail id="602ADE3491DD1AD4B3A11B722CE83E73" type="issue">
<mods:number id="3FA2A0270F7A39DF6AA65264ED1B07FC">4</mods:number>
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<mods:start id="D3F005BA54F6318E4823D4F1002DB438">535</mods:start>
<mods:end id="4E5A87AD6B2F248D6BC6EDF1757B6DA3">544</mods:end>
</mods:extent>
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<mods:classification id="B48CA068DB64FDCACFC128080FC67663">journal article</mods:classification>
<mods:identifier id="C98C4F237434CFAA6783F34BDB9BB071" type="CLB-Dataset">32729</mods:identifier>
<mods:identifier id="EE05A0A60258788EE9C5B5CED48BE5B4" type="DOI">10.11646/zootaxa.4286.4.6</mods:identifier>
<mods:identifier id="32DF6A3E46F81F3696F8240921A8FA7C" type="GBIF-Dataset">9d2b00be-244a-4203-962b-1144b3a728ea</mods:identifier>
<mods:identifier id="42505B0A3C79299A7CA553B04ECDCC59" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C27330A3F2F52B4AD5DFDC95DE0BB204" type="Zenodo-Dep">828773</mods:identifier>
<mods:identifier id="8087B35C73E611BC590D6695D6E2AE56" type="ZooBank">692B2C15-8D7C-4577-B68D-62A34CA20396</mods:identifier>
</mods:mods>
<treatment id="03D5304D916BFFFCFF79F9EA77F702C9" ID-DOI="http://doi.org/10.5281/zenodo.6015523" ID-GBIF-Taxon="132549051" ID-Zenodo-Dep="6015523" LSID="urn:lsid:plazi:treatment:03D5304D916BFFFCFF79F9EA77F702C9" httpUri="http://treatment.plazi.org/id/03D5304D916BFFFCFF79F9EA77F702C9" lastPageId="3" lastPageNumber="538" pageId="1" pageNumber="536">
<subSubSection id="C366D2D0916BFFFEFF79F9EA70040165" box="[151,613,1599,1626]" pageId="1" pageNumber="536" type="nomenclature">
<paragraph id="8BC3815B916BFFFEFF79F9EA70040165" blockId="1.[151,613,1599,1626]" box="[151,613,1599,1626]" pageId="1" pageNumber="536">
<heading id="D08B3637916BFFFEFF79F9EA70040165" bold="true" box="[151,613,1599,1626]" fontSize="11" level="1" pageId="1" pageNumber="536" reason="1">
<taxonomicName id="4C7CFAD8916BFFFEFF79F9EA70040165" authority="Karg, 1978" authorityName="Karg" authorityYear="1978" box="[151,613,1599,1626]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="536" phylum="Arthropoda" rank="species" species="guttulatus">
<emphasis id="B9085D49916BFFFEFF79F9EA70040165" bold="true" box="[151,613,1599,1626]" pageId="1" pageNumber="536">
<emphasis id="B9085D49916BFFFEFF79F9EA73A10166" bold="true" box="[151,448,1599,1625]" italics="true" pageId="1" pageNumber="536">Cosmolaelaps guttulatus</emphasis>
(
<bibRefCitation id="EFEDFCAA916BFFFEFE3EF995703C0165" author="Karg" box="[464,605,1600,1626]" pageId="1" pageNumber="536" refString="Karg, W. (1978) Zur Kenntnis der Milbengattungen Hypoaspis, Androlaelaps und Reticulolaelaps (Acarina, Parasitiformes, Dermanyssidae). Zoologisches Jahrbuch der Systematik, 105, 1 - 32." type="journal article" year="1978">Karg, 1978</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C366D2D0916BFFFEFF79F95D71760028" pageId="1" pageNumber="536" type="reference_group">
<paragraph id="8BC3815B916BFFFEFF79F95D70C301A3" blockId="1.[151,1424,1670,1815]" box="[151,674,1670,1693]" pageId="1" pageNumber="536">
<treatmentCitationGroup id="AB6CA675916BFFFEFF79F95D70C301A3" box="[151,674,1670,1693]" pageId="1" pageNumber="536">
<taxonomicName id="4C7CFAD8916BFFFEFF79F95D70FF01A3" authority="Karg, 1978: 5" authorityName="Karg" authorityPageNumber="5" authorityYear="1978" box="[151,670,1670,1693]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="536" phylum="Arthropoda" rank="species" species="guttulata" subGenus="Cosmolaelaps">
<emphasis id="B9085D49916BFFFEFF79F95D736301A2" box="[151,258,1672,1693]" italics="true" pageId="1" pageNumber="536">Hypoaspis</emphasis>
(
<emphasis id="B9085D49916BFFFEFEFEF95373FE01A3" box="[272,415,1670,1692]" italics="true" pageId="1" pageNumber="536">Cosmolaelaps</emphasis>
)
<emphasis id="B9085D49916BFFFEFE40F953706B01A3" box="[430,522,1670,1692]" italics="true" pageId="1" pageNumber="536">guttulata</emphasis>
<treatmentCitation id="0ADDA74A916BFFFEFDFEF95270FF01A3" author="Karg" box="[528,670,1670,1692]" page="5" pageId="1" pageNumber="536" year="1978">
<bibRefCitation id="EFEDFCAA916BFFFEFDFEF95270E401A3" author="Karg" box="[528,645,1670,1692]" pageId="1" pageNumber="536" refString="Karg, W. (1978) Zur Kenntnis der Milbengattungen Hypoaspis, Androlaelaps und Reticulolaelaps (Acarina, Parasitiformes, Dermanyssidae). Zoologisches Jahrbuch der Systematik, 105, 1 - 32." type="journal article" year="1978">Karg, 1978</bibRefCitation>
: 5
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC3815B916BFFFEFF79F97370D401E5" blockId="1.[151,1424,1670,1815]" pageId="1" pageNumber="536">
<taxonomicName id="4C7CFAD8916BFFFEFF79F973706B0184" box="[151,522,1701,1723]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="536" phylum="Arthropoda" rank="species" species="guttulata" subGenus="Cosmolaelaps">
<emphasis id="B9085D49916BFFFEFF79F97373630184" box="[151,258,1702,1723]" italics="true" pageId="1" pageNumber="536">Hypoaspis</emphasis>
(
<emphasis id="B9085D49916BFFFEFEFEF97073FE0184" box="[272,415,1701,1723]" italics="true" pageId="1" pageNumber="536">Cosmolaelaps</emphasis>
)
<emphasis id="B9085D49916BFFFEFE40F970706B0184" box="[430,522,1701,1723]" italics="true" pageId="1" pageNumber="536">guttulata</emphasis>
</taxonomicName>
.—Karg, 1979: 72; 1981: 218; 1985: 236; 1987: 291; 1988: 516; Xu &amp; Liang, 1996: 193.
<taxonomicName id="4C7CFAD8916BFFFEFF79F91173F001E5" box="[151,401,1732,1754]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="536" phylum="Arthropoda" rank="species" species="guttulatus">
<emphasis id="B9085D49916BFFFEFF79F91173F001E5" box="[151,401,1732,1754]" italics="true" pageId="1" pageNumber="536">Cosmolaelaps guttulatus</emphasis>
</taxonomicName>
.—Moreira
<emphasis id="B9085D49916BFFFEFDE7F910705701E5" box="[521,566,1732,1754]" italics="true" pageId="1" pageNumber="536">et al</emphasis>
., 2014: 319.
</paragraph>
<paragraph id="8BC3815B916BFFFEFF79F93771760028" blockId="1.[151,1424,1670,1815]" pageId="1" pageNumber="536">
<treatmentCitationGroup id="AB6CA675916BFFFEFF79F93771760028" pageId="1" pageNumber="536">
<taxonomicName id="4C7CFAD8916BFFFEFF79F937710701C7" authority="Moreira, Klompen &amp; Moraes, 2014: 333" authorityName="Moreira, Klompen &amp; Moraes" authorityPageNumber="333" authorityYear="2014" box="[151,870,1762,1784]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="536" phylum="Arthropoda" rank="species" species="confinisetarum">
<emphasis id="B9085D49916BFFFEFF79F93773A501C7" box="[151,452,1762,1784]" italics="true" pageId="1" pageNumber="536">Cosmolaelaps confinisetarum</emphasis>
<treatmentCitation id="0ADDA74A916BFFFEFE25F937710701C7" ID-DOI="http://doi.org/10.5281/zenodo.3500636" ID-Zenodo-Dep="3500636" author="Moreira" box="[459,870,1762,1784]" httpUri="http://treatment.plazi.org/id/03DF87B5FFC1FF80FF10F8DAFD28F909" page="333" pageId="1" pageNumber="536" year="2014">
<bibRefCitation id="EFEDFCAA916BFFFEFE25F937715601C7" author="Moreira" box="[459,823,1762,1784]" pageId="1" pageNumber="536" refString="Moreira, G. F., Klompen, H. &amp; Moraes, G. J. de (2014) Redefinition of Cosmolaelaps Berlese (Acari: Laelapidae) and description of five new species from Brazil. Zootaxa, 3764 (3), 317 - 346." type="journal article" year="2014">Moreira, Klompen &amp; Moraes, 2014</bibRefCitation>
: 333
</treatmentCitation>
</taxonomicName>
(synonymy by
<treatmentCitation id="0ADDA74A916BFFFEFBEAF937771001C7" author="Nemati" box="[1028,1393,1762,1784]" page="546" pageId="1" pageNumber="536" year="2016">
<bibRefCitation id="EFEDFCAA916BFFFEFBEAF937775F01C7" author="Nemati" box="[1028,1342,1762,1784]" pageId="1" pageNumber="536" refString="Nemati, A. &amp; Gwiazdowicz, D. J. (2016) Description of a new species of Cosmolaelaps Berlese and the male of C. brevipedestra (Karg) from Iran, with notes on some other species of Cosmolaelaps Berlese (Acari: Laelapidae). Zootaxa, 4066 (5), 535 - 551." type="journal article" year="2016">Nemati &amp; Gwiazdowicz, 2016</bibRefCitation>
: 546
</treatmentCitation>
).
<taxonomicName id="4C7CFAD8916BFFFEFF79F8D473E90028" box="[151,392,1793,1815]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="536" phylum="Arthropoda" rank="species" species="guttulata">
<emphasis id="B9085D49916BFFFEFF79F8D473E90028" box="[151,392,1793,1815]" italics="true" pageId="1" pageNumber="536">Cosmolaelaps guttulata</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0ADDA74A916BFFFEFE4BF8D471720028" author="Nemati" box="[421,787,1793,1815]" page="546" pageId="1" pageNumber="536" year="2016">
<bibRefCitation id="EFEDFCAA916BFFFEFE4BF8D470810028" author="Nemati" box="[421,736,1793,1815]" pageId="1" pageNumber="536" refString="Nemati, A. &amp; Gwiazdowicz, D. J. (2016) Description of a new species of Cosmolaelaps Berlese and the male of C. brevipedestra (Karg) from Iran, with notes on some other species of Cosmolaelaps Berlese (Acari: Laelapidae). Zootaxa, 4066 (5), 535 - 551." type="journal article" year="2016">Nemati &amp; Gwiazdowicz, 2016</bibRefCitation>
: 546
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C366D2D0916BFFFEFF79F89176A200D3" pageId="1" pageNumber="536" type="materials_examined">
<paragraph id="8BC3815B916BFFFEFF79F89176A200D3" blockId="1.[151,1436,1860,2029]" pageId="1" pageNumber="536">
<emphasis id="B9085D49916BFFFEFF79F89173E00062" bold="true" box="[151,385,1860,1885]" pageId="1" pageNumber="536">Material examined.</emphasis>
ACeguá-RS,
<collectingCountry id="F36BC1CB916BFFFEFDF7F89170010062" box="[537,608,1860,1885]" name="Brazil" pageId="1" pageNumber="536">Brazil</collectingCountry>
, soil, five females (
<date id="FFC2A79B916BFFFEFCD1F89176110063" box="[831,1136,1860,1885]" pageId="1" pageNumber="536" value="2012-03-05" valueMax="2012-06-26" valueMin="2012-03-05">5 MarCh and 26 June 2012</date>
,
<date id="FFC2A79B916BFFFEFB92F89177530063" box="[1148,1330,1860,1884]" pageId="1" pageNumber="536" value="2013-08-20">20 August 2013</date>
) and two heteromorphiC males (
<date id="FFC2A79B916BFFFEFE7EF8B2704B00BF" box="[400,554,1895,1920]" pageId="1" pageNumber="536" value="2012-06-26">26 June 2012</date>
), ColleCted by A. da F. Duarte. Rodovia Geraldo de Barros, km 204, São Pedro- SP,
<collectingCountry id="F36BC1CB916BFFFEFF28F859736C009A" box="[198,269,1932,1957]" name="Brazil" pageId="1" pageNumber="536">Brazil</collectingCountry>
, litter under
<taxonomicName id="4C7CFAD8916BFFFEFE44F85970AF009B" authority="Mart." authorityName="Mart." box="[426,718,1932,1956]" class="Liliopsida" family="Arecaceae" genus="Syagrus" kingdom="Plantae" order="Arecales" pageId="1" pageNumber="536" phylum="Tracheophyta" rank="species" species="oleracea">
<emphasis id="B9085D49916BFFFEFE44F8597013009B" box="[426,626,1932,1956]" italics="true" pageId="1" pageNumber="536">Syagrus oleracea</emphasis>
(Mart.)
</taxonomicName>
(
<taxonomicName id="4C7CFAD8916BFFFEFD0FF859713C009B" box="[737,861,1932,1956]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="1" pageNumber="536" phylum="Tracheophyta" rank="family">AreCaCeae</taxonomicName>
), one male (
<date id="FFC2A79B916BFFFEFC1AF85976D4009B" box="[1012,1205,1932,1956]" pageId="1" pageNumber="536" value="2000-08-10">10 August 2000</date>
), ColleCted by A.R. Oliveira. NúCleo AgríCola Vale Ribeira,
<collectionCode id="ED6D199E916BFFFEFD7FF86570A500F7" box="[657,708,1968,1992]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15566" name="Instituto Agron<6F>mico de Campinas" pageId="1" pageNumber="536">IAC</collectionCode>
, Pariquera-Açu-SP,
<collectingCountry id="F36BC1CB916BFFFEFC55F865766300F6" box="[955,1026,1968,1993]" name="Brazil" pageId="1" pageNumber="536">Brazil</collectingCountry>
, soil under
<taxonomicName id="4C7CFAD8916BFFFEFB4AF86477FD00F7" authority="Mart." authorityName="Mart." box="[1188,1436,1969,1992]" class="Liliopsida" family="Arecaceae" genus="Bactris" kingdom="Plantae" order="Arecales" pageId="1" pageNumber="536" phylum="Tracheophyta" rank="species" species="setosa">
<emphasis id="B9085D49916BFFFEFB4AF864772C00F7" box="[1188,1357,1969,1992]" italics="true" pageId="1" pageNumber="536">Bactris setosa</emphasis>
Mart.
</taxonomicName>
(
<taxonomicName id="4C7CFAD8916BFFFEFF70F801737A00D3" box="[158,283,2004,2028]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="1" pageNumber="536" phylum="Tracheophyta" rank="family">AreCaCeae</taxonomicName>
), one male (
<date id="FFC2A79B916BFFFEFE4CF801702200D3" box="[418,579,2004,2029]" pageId="1" pageNumber="536" value="2000-04-18">18 April 2000</date>
), ColleCted by A.R. Oliveira (deposited at ESALQ-USP).
</paragraph>
</subSubSection>
<subSubSection id="C366D2D09168FFFDFF29FF4273C3009D" pageId="2" pageNumber="537" type="description">
<paragraph id="8BC3815B9168FFFDFF29FF4276C707C7" blockId="2.[151,1436,151,429]" pageId="2" pageNumber="537">
<emphasis id="B9085D499168FFFDFF29FF42713E078F" bold="true" box="[199,863,151,176]" pageId="2" pageNumber="537">Description of the heteromorphic male. Diagnosis.</emphasis>
Dorsal idiosoma and gnathosoma similar to homeomorphiC male, sternogenitiventral shield with seven pairs of setae (
<emphasis id="B9085D499168FFFDFC14FF69762807EC" box="[1018,1097,188,212]" italics="true" pageId="2" pageNumber="537">st1–st5</emphasis>
,
<emphasis id="B9085D499168FFFDFBB2FF6976E307EB" box="[1116,1154,188,212]" italics="true" pageId="2" pageNumber="537">Jv1</emphasis>
and
<emphasis id="B9085D499168FFFDFB2CFF69768D07EB" box="[1218,1260,188,212]" italics="true" pageId="2" pageNumber="537">Zv1</emphasis>
); unsClerotised CutiCle with eight pairs of setae (
<emphasis id="B9085D499168FFFDFDEAFF35700107C7" box="[516,608,224,248]" italics="true" pageId="2" pageNumber="537">Jv2–Jv5</emphasis>
,
<emphasis id="B9085D499168FFFDFD80FF3570AE07C7" box="[622,719,224,248]" italics="true" pageId="2" pageNumber="537">Zv2–Zv5</emphasis>
). Two pairs of elongate metapodal plates.
</paragraph>
<paragraph id="8BC3815B9168FFFDFF29FED172900693" blockId="2.[151,1436,151,429]" pageId="2" pageNumber="537">
<emphasis id="B9085D499168FFFDFF29FED173E80622" bold="true" box="[199,393,260,285]" pageId="2" pageNumber="537">Dorsal idiosoma</emphasis>
. Dorsal shield 387–415 long and 272–275 wide (at level of
<emphasis id="B9085D499168FFFDFBDBFED1762C0623" box="[1077,1101,260,284]" italics="true" pageId="2" pageNumber="537">r6</emphasis>
), 314–384 long and 231–280 wide in homeomorphiC male; mostly with transverse elongate retiCules, with 39 pairs of setae, inCluding two pairs of extra setae, and one (
<emphasis id="B9085D499168FFFDFE49FE9973AF065B" box="[423,462,332,356]" italics="true" pageId="2" pageNumber="537">Jx3</emphasis>
) or two (
<emphasis id="B9085D499168FFFDFDD4FE997000065B" box="[570,609,332,356]" italics="true" pageId="2" pageNumber="537">Jx2</emphasis>
and
<emphasis id="B9085D499168FFFDFD77FE9970DE065B" box="[665,703,332,356]" italics="true" pageId="2" pageNumber="537">Jx3</emphasis>
) unpaired setae. UnsClerotised lateral CutiCle with seven pairs of setae (
<emphasis id="B9085D499168FFFDFF31FEA5729906B7" box="[223,248,368,392]" italics="true" pageId="2" pageNumber="537">r6</emphasis>
,
<emphasis id="B9085D499168FFFDFEE8FEA5734706B7" box="[262,294,368,392]" italics="true" pageId="2" pageNumber="537">R1</emphasis>
,
<emphasis id="B9085D499168FFFDFEDAFEA5733206B7" box="[308,339,368,392]" italics="true" pageId="2" pageNumber="537">R2</emphasis>
,
<emphasis id="B9085D499168FFFDFE8FFEA573CC06B7" box="[353,429,368,392]" italics="true" pageId="2" pageNumber="537">R4–R6</emphasis>
, one
<emphasis id="B9085D499168FFFDFE05FEA4707106B7" box="[491,528,369,392]" italics="true" pageId="2" pageNumber="537">UR</emphasis>
). All setae sCimitar-shaped, exCept
<emphasis id="B9085D499168FFFDFC4CFEA571D606B7" box="[930,951,368,392]" italics="true" pageId="2" pageNumber="537">j1</emphasis>
and
<emphasis id="B9085D499168FFFDFC00FEA5766606B7" box="[1006,1031,368,392]" italics="true" pageId="2" pageNumber="537">z1</emphasis>
, setiform. Length of setae shown in
<tableCitation id="C6FEB4E09168FFFDFF79FE41728A0693" box="[151,235,404,429]" captionStart="TABLE 1" captionStartId="2.[151,239,476,499]" captionText="TABLE 1. Length of dorsal idiosomal setae of males of Cosmolaelaps guttulatus (Karg, 1978); for homeomorphic males, mean (minimum – maximum); for heteromorphic males, minimum – maximum; * Brazilian specimens, paratypes of C. confinisetarum Moreira, Klompen and Moraes, 2014 (junior synonym); – = seta absent." pageId="2" pageNumber="537">Table 1</tableCitation>
.
</paragraph>
<caption id="DF03D1D39168FFFDFF79FE09763C050C" pageId="2" pageNumber="537">
<paragraph id="8BC3815B9168FFFDFF79FE09763C050C" blockId="2.[151,1435,475,563]" pageId="2" pageNumber="537">
<emphasis id="B9085D499168FFFDFF79FE09736F06CC" bold="true" box="[151,270,476,499]" pageId="2" pageNumber="537">TABLE 1.</emphasis>
Length of dorsal idiosomal setae of males of
<taxonomicName id="4C7CFAD89168FFFDFCF4FE0E76A306CC" authority="Karg, 1978" authorityName="Karg" authorityYear="1978" box="[794,1218,475,499]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="537" phylum="Arthropoda" rank="species" species="guttulatus">
<emphasis id="B9085D499168FFFDFCF4FE0E764606CC" box="[794,1063,475,499]" italics="true" pageId="2" pageNumber="537">Cosmolaelaps guttulatus</emphasis>
(Karg, 1978)
</taxonomicName>
; for homeomorphic males, mean (minimum–maximum); for heteromorphic males, minimum–maximum; *Brazilian specimens, paratypes of
<taxonomicName id="4C7CFAD89168FFFDFF79FDCE7088050C" authority="Moreira, Klompen and Moraes, 2014" authorityName="Moreira, Klompen and Moraes" authorityYear="2014" box="[151,745,539,563]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="537" phylum="Arthropoda" rank="species" species="confinisetarum">
<emphasis id="B9085D499168FFFDFF79FDCE7337050C" box="[151,342,539,563]" italics="true" pageId="2" pageNumber="537">C. confinisetarum</emphasis>
Moreira, Klompen and Moraes, 2014
</taxonomicName>
(junior synonym); – = seta absent.
</paragraph>
</caption>
<paragraph id="8BC3815B9168FFFDFF71FD9B76BC01B3" pageId="2" pageNumber="537">
<table id="F97C73FB91680000FF71FD9B773D01B3" box="[159,1372,590,1676]" gridcols="6" gridrows="9" pageId="2" pageNumber="537">
<tr id="354C831991680000FF71FD9B773D0593" box="[159,1372,590,684]" gridrow="0" pageId="2" pageNumber="537">
<th id="769DEA6591680000FF71FD9B72AB0593" box="[159,202,590,684]" gridcol="0" gridrow="0" pageId="2" pageNumber="537">Seta j1</th>
<th id="769DEA6591680000FEFEFD9B739C0593" box="[272,509,590,684]" gridcol="1" gridrow="0" pageId="2" pageNumber="537">Homeomorphic males* 19 (18–21)</th>
<th id="769DEA6591680000FDD0FD9B70B60593" box="[574,727,590,684]" gridcol="2" gridrow="0" pageId="2" pageNumber="537">Heteromorphic males 14–16</th>
<th id="769DEA6591680000FC83FD9B71F90593" box="[877,920,590,684]" gridcol="3" gridrow="0" pageId="2" pageNumber="537">Seta J1</th>
<th id="769DEA6591680000FC30FD9B761C0593" box="[990,1149,590,684]" gridcol="4" gridrow="0" pageId="2" pageNumber="537">Homeomorphic males* 41 (39–45)</th>
<th id="769DEA6591680000FB2DFD9B773D0593" box="[1219,1372,590,684]" gridcol="5" gridrow="0" pageId="2" pageNumber="537">Heteromorphic males 41–42</th>
</tr>
<tr id="354C831991680000FF71FD15773D0417" box="[159,1372,704,808]" gridrow="1" pageId="2" pageNumber="537">
<th id="769DEA6591680000FF71FD1572AB0417" box="[159,202,704,808]" gridcol="0" gridrow="1" pageId="2" pageNumber="537">j2 j3 j4</th>
<td id="769DEA6591680000FEFEFD15739C0417" box="[272,509,704,808]" gridcol="1" gridrow="1" pageId="2" pageNumber="537">32 (31–32) 40 (37–44) 43 (38–50)</td>
<td id="769DEA6591680000FDD0FD1570B60417" box="[574,727,704,808]" gridcol="2" gridrow="1" pageId="2" pageNumber="537">38–42 44–46 46–49</td>
<td id="769DEA6591680000FC83FD1571F90417" box="[877,920,704,808]" gridcol="3" gridrow="1" pageId="2" pageNumber="537">J2 J3 J4</td>
<td id="769DEA6591680000FC30FD15761C0417" box="[990,1149,704,808]" gridcol="4" gridrow="1" pageId="2" pageNumber="537">40 (38–43) 36 (35–38) 33 (30–37)</td>
<td id="769DEA6591680000FB2DFD15773D0417" box="[1219,1372,704,808]" gridcol="5" gridrow="1" pageId="2" pageNumber="537">39–41 38–39 32</td>
</tr>
<tr id="354C831991680000FF71FCE9773D049B" box="[159,1372,828,932]" gridrow="2" pageId="2" pageNumber="537">
<th id="769DEA6591680000FF71FCE972AB049B" box="[159,202,828,932]" gridcol="0" gridrow="2" pageId="2" pageNumber="537">j5 j6 z1</th>
<td id="769DEA6591680000FEFEFCE9739C049B" box="[272,509,828,932]" gridcol="1" gridrow="2" pageId="2" pageNumber="537">45 (41–48) 43 (40–46) 18 (16–20)</td>
<td id="769DEA6591680000FDD0FCE970B6049B" box="[574,727,828,932]" gridcol="2" gridrow="2" pageId="2" pageNumber="537">29–51 45–50 24</td>
<td id="769DEA6591680000FC83FCE971F9049B" box="[877,920,828,932]" gridcol="3" gridrow="2" pageId="2" pageNumber="537">J5 Jx2 Jx3</td>
<td id="769DEA6591680000FC30FCE9761C049B" box="[990,1149,828,932]" gridcol="4" gridrow="2" pageId="2" pageNumber="537">24 (21–29) 36 (32–39) 34 (32–40)</td>
<td id="769DEA6591680000FB2DFCE9773D049B" box="[1219,1372,828,932]" gridcol="5" gridrow="2" pageId="2" pageNumber="537">21–27 33 29–33</td>
</tr>
<tr id="354C831991680000FF71FC6D773D031F" box="[159,1372,952,1056]" gridrow="3" pageId="2" pageNumber="537">
<th id="769DEA6591680000FF71FC6D72AB031F" box="[159,202,952,1056]" gridcol="0" gridrow="3" pageId="2" pageNumber="537">z2 z3 z4</th>
<td id="769DEA6591680000FEFEFC6D739C031F" box="[272,509,952,1056]" gridcol="1" gridrow="3" pageId="2" pageNumber="537">37 (29–44) 42 (37–45) 43 (40–47)</td>
<td id="769DEA6591680000FDD0FC6D70B6031F" box="[574,727,952,1056]" gridcol="2" gridrow="3" pageId="2" pageNumber="537">49 42–45 51–53</td>
<td id="769DEA6591680000FC83FC6D71F9031F" box="[877,920,952,1056]" gridcol="3" gridrow="3" pageId="2" pageNumber="537">Jx4 Z1 Z2</td>
<td id="769DEA6591680000FC30FC6D761C031F" box="[990,1149,952,1056]" gridcol="4" gridrow="3" pageId="2" pageNumber="537">31 (28–35) 42 (39–46) 41 (38–46)</td>
<td id="769DEA6591680000FB2DFC6D773D031F" box="[1219,1372,952,1056]" gridcol="5" gridrow="3" pageId="2" pageNumber="537">– 44–45 40–43</td>
</tr>
<tr id="354C831991680000FF71FBE1773D03A3" box="[159,1372,1076,1180]" gridrow="4" pageId="2" pageNumber="537">
<th id="769DEA6591680000FF71FBE172AB03A3" box="[159,202,1076,1180]" gridcol="0" gridrow="4" pageId="2" pageNumber="537">z5 z6 s1</th>
<td id="769DEA6591680000FEFEFBE1739C03A3" box="[272,509,1076,1180]" gridcol="1" gridrow="4" pageId="2" pageNumber="537">43 (40–47) 44 (40–48) 33 (30–36)</td>
<td id="769DEA6591680000FDD0FBE170B603A3" box="[574,727,1076,1180]" gridcol="2" gridrow="4" pageId="2" pageNumber="537">49–50 47–49 37</td>
<td id="769DEA6591680000FC83FBE171F903A3" box="[877,920,1076,1180]" gridcol="3" gridrow="4" pageId="2" pageNumber="537">Z3 Z4 Z5</td>
<td id="769DEA6591680000FC30FBE1761C03A3" box="[990,1149,1076,1180]" gridcol="4" gridrow="4" pageId="2" pageNumber="537">38 (34–42) 34 (31–39) 40 (37–42)</td>
<td id="769DEA6591680000FB2DFBE1773D03A3" box="[1219,1372,1076,1180]" gridcol="5" gridrow="4" pageId="2" pageNumber="537">40–41 39–40 45–48</td>
</tr>
<tr id="354C831991680000FF71FB65773D0227" box="[159,1372,1200,1304]" gridrow="5" pageId="2" pageNumber="537">
<th id="769DEA6591680000FF71FB6572AB0227" box="[159,202,1200,1304]" gridcol="0" gridrow="5" pageId="2" pageNumber="537">s2 s3 s4</th>
<td id="769DEA6591680000FEFEFB65739C0227" box="[272,509,1200,1304]" gridcol="1" gridrow="5" pageId="2" pageNumber="537">34 (27–38) 39 (36–45) 45 (43–51)</td>
<td id="769DEA6591680000FDD0FB6570B60227" box="[574,727,1200,1304]" gridcol="2" gridrow="5" pageId="2" pageNumber="537">41 43–49 50–51</td>
<td id="769DEA6591680000FC83FB6571F90227" box="[877,920,1200,1304]" gridcol="3" gridrow="5" pageId="2" pageNumber="537">S1 S2 S3</td>
<td id="769DEA6591680000FC30FB65761C0227" box="[990,1149,1200,1304]" gridcol="4" gridrow="5" pageId="2" pageNumber="537">34 (32–36) 37 (34–41) 37 (36–38)</td>
<td id="769DEA6591680000FB2DFB65773D0227" box="[1219,1372,1200,1304]" gridcol="5" gridrow="5" pageId="2" pageNumber="537">41–42 41–42 37–41</td>
</tr>
<tr id="354C831991680000FF71FAF9773D02AB" box="[159,1372,1324,1428]" gridrow="6" pageId="2" pageNumber="537">
<th id="769DEA6591680000FF71FAF972AB02AB" box="[159,202,1324,1428]" gridcol="0" gridrow="6" pageId="2" pageNumber="537">s5 s6 r2</th>
<td id="769DEA6591680000FEFEFAF9739C02AB" box="[272,509,1324,1428]" gridcol="1" gridrow="6" pageId="2" pageNumber="537">47 (44–49) 38 (37–39) 30 (27–35)</td>
<td id="769DEA6591680000FDD0FAF970B602AB" box="[574,727,1324,1428]" gridcol="2" gridrow="6" pageId="2" pageNumber="537">53–54 42–44 34–37</td>
<td id="769DEA6591680000FC83FAF971F902AB" box="[877,920,1324,1428]" gridcol="3" gridrow="6" pageId="2" pageNumber="537">S4 S5 px2</td>
<td id="769DEA6591680000FC30FAF9761C02AB" box="[990,1149,1324,1428]" gridcol="4" gridrow="6" pageId="2" pageNumber="537">34 (32–36) 33 (30–37) 39 (34–47)</td>
<td id="769DEA6591680000FB2DFAF9773D02AB" box="[1219,1372,1324,1428]" gridcol="5" gridrow="6" pageId="2" pageNumber="537">37–39 38–40 31–36</td>
</tr>
<tr id="354C831991680000FF71FA7D773D012F" box="[159,1372,1448,1552]" gridrow="7" pageId="2" pageNumber="537">
<th id="769DEA6591680000FF71FA7D72AB012F" box="[159,202,1448,1552]" gridcol="0" gridrow="7" pageId="2" pageNumber="537">r3 r4 r5</th>
<td id="769DEA6591680000FEFEFA7D739C012F" box="[272,509,1448,1552]" gridcol="1" gridrow="7" pageId="2" pageNumber="537">37 (35–38) 37 (36–38) 40 (39–41)</td>
<td id="769DEA6591680000FDD0FA7D70B6012F" box="[574,727,1448,1552]" gridcol="2" gridrow="7" pageId="2" pageNumber="537">40 39–40 40–42</td>
<td id="769DEA6591680000FC83FA7D71F9012F" box="[877,920,1448,1552]" gridcol="3" gridrow="7" pageId="2" pageNumber="537">px3 R1 R2</td>
<td id="769DEA6591680000FC30FA7D761C012F" box="[990,1149,1448,1552]" gridcol="4" gridrow="7" pageId="2" pageNumber="537">36 (34–38) 18 (17–19) 19 (18–19)</td>
<td id="769DEA6591680000FB2DFA7D773D012F" box="[1219,1372,1448,1552]" gridcol="5" gridrow="7" pageId="2" pageNumber="537">35 26–28 27–28</td>
</tr>
<tr id="354C831991680000FF71F9F1773D01B3" box="[159,1372,1572,1676]" gridrow="8" pageId="2" pageNumber="537">
<th id="769DEA6591680000FF71F9F172AB01B3" box="[159,202,1572,1676]" gridcol="0" gridrow="8" pageId="2" pageNumber="537">r6</th>
<td id="769DEA6591680000FEFEF9F1739C01B3" box="[272,509,1572,1676]" gridcol="1" gridrow="8" pageId="2" pageNumber="537">18 (17–19)</td>
<td id="769DEA6591680000FDD0F9F170B601B3" box="[574,727,1572,1676]" gridcol="2" gridrow="8" pageId="2" pageNumber="537">26–30</td>
<td id="769DEA6591680000FC83F9F171F901B3" box="[877,920,1572,1676]" gridcol="3" gridrow="8" pageId="2" pageNumber="537">R4 R5 R6</td>
<td id="769DEA6591680000FC30F9F1761C01B3" box="[990,1149,1572,1676]" gridcol="4" gridrow="8" pageId="2" pageNumber="537">17 (17–17) 17 (16–17) –</td>
<td id="769DEA6591680000FB2DF9F1773D01B3" box="[1219,1372,1572,1676]" gridcol="5" gridrow="8" pageId="2" pageNumber="537">27–28 26–28 30</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC3815B9168FFFDFC83F97573C3009D" blockId="2.[877,1282,1696,1718]" lastBlockId="2.[151,1437,1785,1954]" pageId="2" pageNumber="537">
UR1 18 (17–19) 28–30
<emphasis id="B9085D499168FFFDFF29F92C73F2002D" bold="true" box="[199,403,1785,1810]" pageId="2" pageNumber="537">Ventral idiosoma</emphasis>
(
<figureCitation id="13479DDE9168FFFDFE4AF92C7386002D" box="[420,487,1785,1810]" captionStart="FIGURES 1 – 5. 1" captionStartId="4.[151,264,1970,1993]" captionTargetBox="[249,1331,198,1937]" captionTargetId="figure@4.[249,1334,193,1949]" captionTargetPageId="4" captionText="FIGURES 1 – 5. 1. Cosmolaelaps guttulatus, heteromorphic male; ventral idiosoma. 2. C. pampaensis, male, ventral idiosoma; 3. C. pampaensis, male, chelicera; 4. C. pampaensis, male, leg III; 5. C. pampaensis, male, leg IV." httpUri="https://zenodo.org/record/828775/files/figure.png" pageId="2" pageNumber="537">Fig. 1</figureCitation>
). Sternogenitiventral shield 240–242 long and 64–70 wide (behind Coxae IV), with seven pairs of setae (
<emphasis id="B9085D499168FFFDFE7EF8C87381000A" box="[400,480,1821,1845]" italics="true" pageId="2" pageNumber="537">st1–st5</emphasis>
,
<emphasis id="B9085D499168FFFDFE1FF8C87079000A" box="[497,536,1821,1845]" italics="true" pageId="2" pageNumber="537">Jv1</emphasis>
and
<emphasis id="B9085D499168FFFDFDB8F8C870E1000A" box="[598,640,1821,1845]" italics="true" pageId="2" pageNumber="537">Zv1</emphasis>
). UnsClerotised CutiCle with eight pairs of setae (
<emphasis id="B9085D499168FFFDFB2DF8CB76BD000A" box="[1219,1244,1822,1845]" italics="true" pageId="2" pageNumber="537">Jv</emphasis>
2–
<emphasis id="B9085D499168FFFDFB16F8CB7770000A" box="[1272,1297,1822,1845]" italics="true" pageId="2" pageNumber="537">Jv</emphasis>
5,
<emphasis id="B9085D499168FFFDFADFF8CB772C000A" box="[1329,1357,1822,1845]" italics="true" pageId="2" pageNumber="537">Zv</emphasis>
2–
<emphasis id="B9085D499168FFFDFA87F8CB77E4000A" box="[1385,1413,1822,1845]" italics="true" pageId="2" pageNumber="537">Zv</emphasis>
5) around anal shield. Length of setae shown in
<tableCitation id="C6FEB4E09168FFFDFD61F89470840065" box="[655,741,1857,1882]" captionStart="TABLE 2" captionStartId="3.[151,239,152,175]" captionText="TABLE 2. Length of ventral idiosomal setae of males of Cosmolaelaps guttulatus (Karg, 1978); for homeomorphic males, mean (minimum – maximum); for heteromorphic males, minimum – maximum. * Brazilian specimens, paratypes of C. confinisetarum Moreira, Klompen and Moraes, 2014 (junior synonym)." pageId="2" pageNumber="537">Table 2</tableCitation>
. Two anterior lobes of sternogenitiventral shield as sClerotised as remainder of the shield. Two pairs of elongate metapodal plates. Peritreme, gnathosoma and legs similar to those in homeomorphiC male.
</paragraph>
</subSubSection>
<caption id="DF03D1D39169FFFCFF79FF4D71D007D0" pageId="3" pageNumber="538">
<paragraph id="8BC3815B9169FFFCFF79FF4D71D007D0" blockId="3.[151,1435,151,239]" pageId="3" pageNumber="538">
<emphasis id="B9085D499169FFFCFF79FF4D736C0790" bold="true" box="[151,269,152,175]" pageId="3" pageNumber="538">TABLE 2.</emphasis>
Length of ventral idiosomal setae of males of
<taxonomicName id="4C7CFAD89169FFFCFCF3FF4276A20790" authority="Karg, 1978" authorityName="Karg" authorityYear="1978" box="[797,1219,151,175]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="guttulatus">
<emphasis id="B9085D499169FFFCFCF3FF4276490790" box="[797,1064,151,175]" italics="true" pageId="3" pageNumber="538">Cosmolaelaps guttulatus</emphasis>
(Karg, 1978)
</taxonomicName>
; for homeomorphic males, mean (minimum–maximum); for heteromorphic males, minimum–maximum. *Brazilian specimens, paratypes of
<taxonomicName id="4C7CFAD89169FFFCFF79FF0D708807D0" authority="Moreira, Klompen and Moraes, 2014" authorityName="Moreira, Klompen and Moraes" authorityYear="2014" box="[151,745,215,239]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="confinisetarum">
<emphasis id="B9085D499169FFFCFF79FF0D733707D0" box="[151,342,215,239]" italics="true" pageId="3" pageNumber="538">C. confinisetarum</emphasis>
Moreira, Klompen and Moraes, 2014
</taxonomicName>
(junior synonym).
</paragraph>
</caption>
<paragraph id="8BC3815B9169FFFCFF71FEDF718103BB" pageId="3" pageNumber="538">
<table id="F97C73FB91690000FF71FEDF761A03BB" box="[159,1147,266,1156]" gridcols="3" gridrows="22" pageId="3" pageNumber="538">
<tr id="354C831991690000FF71FEDF761A061F" box="[159,1147,266,288]" gridrow="0" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FEDF729E061F" box="[159,255,266,288]" gridcol="0" gridrow="0" pageId="3" pageNumber="538">Seta</th>
<th id="769DEA6591690000FE93FEDF700B061F" box="[381,618,266,288]" gridcol="1" gridrow="0" pageId="3" pageNumber="538">Homeomorphic males*</th>
<th id="769DEA6591690000FC4EFEDF761A061F" box="[928,1147,266,288]" gridcol="2" gridrow="0" pageId="3" pageNumber="538">Heteromorphic males</th>
</tr>
<tr id="354C831991690000FF71FEE1761A0675" box="[159,1147,308,330]" gridrow="1" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FEE1729E0675" box="[159,255,308,330]" gridcol="0" gridrow="1" pageId="3" pageNumber="538">h1</th>
<td id="769DEA6591690000FE93FEE1700B0675" box="[381,618,308,330]" gridcol="1" gridrow="1" pageId="3" pageNumber="538">21 (20–23)</td>
<td id="769DEA6591690000FC4EFEE1761A0675" box="[928,1147,308,330]" gridcol="2" gridrow="1" pageId="3" pageNumber="538">12–15</td>
</tr>
<tr id="354C831991690000FF71FE88761A064C" box="[159,1147,349,371]" gridrow="2" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FE88729E064C" box="[159,255,349,371]" gridcol="0" gridrow="2" pageId="3" pageNumber="538">h2</th>
<td id="769DEA6591690000FE93FE88700B064C" box="[381,618,349,371]" gridcol="1" gridrow="2" pageId="3" pageNumber="538">11 (10–13)</td>
<td id="769DEA6591690000FC4EFE88761A064C" box="[928,1147,349,371]" gridcol="2" gridrow="2" pageId="3" pageNumber="538">13–14</td>
</tr>
<tr id="354C831991690000FF71FE53761A06A3" box="[159,1147,390,412]" gridrow="3" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FE53729E06A3" box="[159,255,390,412]" gridcol="0" gridrow="3" pageId="3" pageNumber="538">h3</th>
<td id="769DEA6591690000FE93FE53700B06A3" box="[381,618,390,412]" gridcol="1" gridrow="3" pageId="3" pageNumber="538">24 (23–25)</td>
<td id="769DEA6591690000FC4EFE53761A06A3" box="[928,1147,390,412]" gridcol="2" gridrow="3" pageId="3" pageNumber="538">17–23</td>
</tr>
<tr id="354C831991690000FF71FE65761A06F9" box="[159,1147,432,454]" gridrow="4" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FE65729E06F9" box="[159,255,432,454]" gridcol="0" gridrow="4" pageId="3" pageNumber="538">Pc</th>
<td id="769DEA6591690000FE93FE65700B06F9" box="[381,618,432,454]" gridcol="1" gridrow="4" pageId="3" pageNumber="538">15 (15–18)</td>
<td id="769DEA6591690000FC4EFE65761A06F9" box="[928,1147,432,454]" gridcol="2" gridrow="4" pageId="3" pageNumber="538">19–20</td>
</tr>
<tr id="354C831991690000FF71FE0C761A06D0" box="[159,1147,473,495]" gridrow="5" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FE0C729E06D0" box="[159,255,473,495]" gridcol="0" gridrow="5" pageId="3" pageNumber="538">st1</th>
<td id="769DEA6591690000FE93FE0C700B06D0" box="[381,618,473,495]" gridcol="1" gridrow="5" pageId="3" pageNumber="538">26 (25–27)</td>
<td id="769DEA6591690000FC4EFE0C761A06D0" box="[928,1147,473,495]" gridcol="2" gridrow="5" pageId="3" pageNumber="538">30–31</td>
</tr>
<tr id="354C831991690000FF71FDD7761A0527" box="[159,1147,514,536]" gridrow="6" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FDD7729E0527" box="[159,255,514,536]" gridcol="0" gridrow="6" pageId="3" pageNumber="538">st2</th>
<td id="769DEA6591690000FE93FDD7700B0527" box="[381,618,514,536]" gridcol="1" gridrow="6" pageId="3" pageNumber="538">25 (23–28)</td>
<td id="769DEA6591690000FC4EFDD7761A0527" box="[928,1147,514,536]" gridcol="2" gridrow="6" pageId="3" pageNumber="538">25–27</td>
</tr>
<tr id="354C831991690000FF71FDF9761A057D" box="[159,1147,556,578]" gridrow="7" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FDF9729E057D" box="[159,255,556,578]" gridcol="0" gridrow="7" pageId="3" pageNumber="538">st3</th>
<td id="769DEA6591690000FE93FDF9700B057D" box="[381,618,556,578]" gridcol="1" gridrow="7" pageId="3" pageNumber="538">24 (22–26)</td>
<td id="769DEA6591690000FC4EFDF9761A057D" box="[928,1147,556,578]" gridcol="2" gridrow="7" pageId="3" pageNumber="538">24–28</td>
</tr>
<tr id="354C831991690000FF71FD80761A0554" box="[159,1147,597,619]" gridrow="8" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FD80729E0554" box="[159,255,597,619]" gridcol="0" gridrow="8" pageId="3" pageNumber="538">st4</th>
<td id="769DEA6591690000FE93FD80700B0554" box="[381,618,597,619]" gridcol="1" gridrow="8" pageId="3" pageNumber="538">24 (22–28)</td>
<td id="769DEA6591690000FC4EFD80761A0554" box="[928,1147,597,619]" gridcol="2" gridrow="8" pageId="3" pageNumber="538">27–29</td>
</tr>
<tr id="354C831991690000FF71FDAB761A05AB" box="[159,1147,638,660]" gridrow="9" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FDAB729E05AB" box="[159,255,638,660]" gridcol="0" gridrow="9" pageId="3" pageNumber="538">st5</th>
<td id="769DEA6591690000FE93FDAB700B05AB" box="[381,618,638,660]" gridcol="1" gridrow="9" pageId="3" pageNumber="538">23 (21–25)</td>
<td id="769DEA6591690000FC4EFDAB761A05AB" box="[928,1147,638,660]" gridcol="2" gridrow="9" pageId="3" pageNumber="538">27</td>
</tr>
<tr id="354C831991690000FF71FD7D761A0581" box="[159,1147,680,702]" gridrow="10" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FD7D729E0581" box="[159,255,680,702]" gridcol="0" gridrow="10" pageId="3" pageNumber="538">Jv1</th>
<td id="769DEA6591690000FE93FD7D700B0581" box="[381,618,680,702]" gridcol="1" gridrow="10" pageId="3" pageNumber="538">31 (25–34)</td>
<td id="769DEA6591690000FC4EFD7D761A0581" box="[928,1147,680,702]" gridcol="2" gridrow="10" pageId="3" pageNumber="538">14–27</td>
</tr>
<tr id="354C831991690000FF71FD04761A05D8" box="[159,1147,721,743]" gridrow="11" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FD04729E05D8" box="[159,255,721,743]" gridcol="0" gridrow="11" pageId="3" pageNumber="538">Jv2</th>
<td id="769DEA6591690000FE93FD04700B05D8" box="[381,618,721,743]" gridcol="1" gridrow="11" pageId="3" pageNumber="538">33 (28–38)</td>
<td id="769DEA6591690000FC4EFD04761A05D8" box="[928,1147,721,743]" gridcol="2" gridrow="11" pageId="3" pageNumber="538">32</td>
</tr>
<tr id="354C831991690000FF71FD2F761A042F" box="[159,1147,762,784]" gridrow="12" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FD2F729E042F" box="[159,255,762,784]" gridcol="0" gridrow="12" pageId="3" pageNumber="538">Jv3</th>
<td id="769DEA6591690000FE93FD2F700B042F" box="[381,618,762,784]" gridcol="1" gridrow="12" pageId="3" pageNumber="538">26 (22–31)</td>
<td id="769DEA6591690000FC4EFD2F761A042F" box="[928,1147,762,784]" gridcol="2" gridrow="12" pageId="3" pageNumber="538">29–31</td>
</tr>
<tr id="354C831991690000FF71FCF1761A0405" box="[159,1147,804,826]" gridrow="13" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FCF1729E0405" box="[159,255,804,826]" gridcol="0" gridrow="13" pageId="3" pageNumber="538">Jv4</th>
<td id="769DEA6591690000FE93FCF1700B0405" box="[381,618,804,826]" gridcol="1" gridrow="13" pageId="3" pageNumber="538">20 (18–22)</td>
<td id="769DEA6591690000FC4EFCF1761A0405" box="[928,1147,804,826]" gridcol="2" gridrow="13" pageId="3" pageNumber="538">26–30</td>
</tr>
<tr id="354C831991690000FF71FC98761A045C" box="[159,1147,845,867]" gridrow="14" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FC98729E045C" box="[159,255,845,867]" gridcol="0" gridrow="14" pageId="3" pageNumber="538">Jv5</th>
<td id="769DEA6591690000FE93FC98700B045C" box="[381,618,845,867]" gridcol="1" gridrow="14" pageId="3" pageNumber="538">25 (22–27)</td>
<td id="769DEA6591690000FC4EFC98761A045C" box="[928,1147,845,867]" gridcol="2" gridrow="14" pageId="3" pageNumber="538">35</td>
</tr>
<tr id="354C831991690000FF71FCA3761A04B3" box="[159,1147,886,908]" gridrow="15" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FCA3729E04B3" box="[159,255,886,908]" gridcol="0" gridrow="15" pageId="3" pageNumber="538">Zv1</th>
<td id="769DEA6591690000FE93FCA3700B04B3" box="[381,618,886,908]" gridcol="1" gridrow="15" pageId="3" pageNumber="538">27 (23–32)</td>
<td id="769DEA6591690000FC4EFCA3761A04B3" box="[928,1147,886,908]" gridcol="2" gridrow="15" pageId="3" pageNumber="538">25</td>
</tr>
<tr id="354C831991690000FF71FC75761A0489" box="[159,1147,928,950]" gridrow="16" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FC75729E0489" box="[159,255,928,950]" gridcol="0" gridrow="16" pageId="3" pageNumber="538">Zv2</th>
<td id="769DEA6591690000FE93FC75700B0489" box="[381,618,928,950]" gridcol="1" gridrow="16" pageId="3" pageNumber="538">29 (25–36)</td>
<td id="769DEA6591690000FC4EFC75761A0489" box="[928,1147,928,950]" gridcol="2" gridrow="16" pageId="3" pageNumber="538">29–31</td>
</tr>
<tr id="354C831991690000FF71FC1C761A04E0" box="[159,1147,969,991]" gridrow="17" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FC1C729E04E0" box="[159,255,969,991]" gridcol="0" gridrow="17" pageId="3" pageNumber="538">Zv3</th>
<td id="769DEA6591690000FE93FC1C700B04E0" box="[381,618,969,991]" gridcol="1" gridrow="17" pageId="3" pageNumber="538">17 (17–18)</td>
<td id="769DEA6591690000FC4EFC1C761A04E0" box="[928,1147,969,991]" gridcol="2" gridrow="17" pageId="3" pageNumber="538">24</td>
</tr>
<tr id="354C831991690000FF71FC27761A0337" box="[159,1147,1010,1032]" gridrow="18" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FC27729E0337" box="[159,255,1010,1032]" gridcol="0" gridrow="18" pageId="3" pageNumber="538">Zv4</th>
<td id="769DEA6591690000FE93FC27700B0337" box="[381,618,1010,1032]" gridcol="1" gridrow="18" pageId="3" pageNumber="538">18 (16–20)</td>
<td id="769DEA6591690000FC4EFC27761A0337" box="[928,1147,1010,1032]" gridcol="2" gridrow="18" pageId="3" pageNumber="538">25–26</td>
</tr>
<tr id="354C831991690000FF71FBC9761A030D" box="[159,1147,1052,1074]" gridrow="19" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FBC9729E030D" box="[159,255,1052,1074]" gridcol="0" gridrow="19" pageId="3" pageNumber="538">Zv5</th>
<td id="769DEA6591690000FE93FBC9700B030D" box="[381,618,1052,1074]" gridcol="1" gridrow="19" pageId="3" pageNumber="538">18 (17–19)</td>
<td id="769DEA6591690000FC4EFBC9761A030D" box="[928,1147,1052,1074]" gridcol="2" gridrow="19" pageId="3" pageNumber="538">20–27</td>
</tr>
<tr id="354C831991690000FF71FB90761A0364" box="[159,1147,1093,1115]" gridrow="20" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FB90729E0364" box="[159,255,1093,1115]" gridcol="0" gridrow="20" pageId="3" pageNumber="538">Para-anal</th>
<td id="769DEA6591690000FE93FB90700B0364" box="[381,618,1093,1115]" gridcol="1" gridrow="20" pageId="3" pageNumber="538">16 (13–19)</td>
<td id="769DEA6591690000FC4EFB90761A0364" box="[928,1147,1093,1115]" gridcol="2" gridrow="20" pageId="3" pageNumber="538">20–21</td>
</tr>
<tr id="354C831991690000FF71FBBB761A03BB" box="[159,1147,1134,1156]" gridrow="21" pageId="3" pageNumber="538">
<th id="769DEA6591690000FF71FBBB729E03BB" box="[159,255,1134,1156]" gridcol="0" gridrow="21" pageId="3" pageNumber="538">Post-anal</th>
<td id="769DEA6591690000FE93FBBB700B03BB" box="[381,618,1134,1156]" gridcol="1" gridrow="21" pageId="3" pageNumber="538">22 (19–24)</td>
<td id="769DEA6591690000FC4EFBBB761A03BB" box="[928,1147,1134,1156]" gridcol="2" gridrow="21" pageId="3" pageNumber="538">28–31</td>
</tr>
</table>
</paragraph>
<subSubSection id="C366D2D09169FFFCFF29FB1277F702C9" pageId="3" pageNumber="538" type="discussion">
<paragraph id="8BC3815B9169FFFCFF29FB1277F702C9" blockId="3.[151,1436,1223,1526]" pageId="3" pageNumber="538">
<emphasis id="B9085D499169FFFCFF29FB12735A03DF" bold="true" box="[199,315,1223,1248]" pageId="3" pageNumber="538">Remarks.</emphasis>
HeteromorphiC males have been reported for
<taxonomicName id="4C7CFAD89169FFFCFC70FB1D770A03DF" authority="MiChael, 1891" authorityName="MiChael" authorityYear="1891" box="[926,1387,1223,1248]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="B9085D499169FFFCFC70FB1D76F503DF" box="[926,1172,1224,1248]" italics="true" pageId="3" pageNumber="538">Cosmolaelaps vacua</emphasis>
(MiChael, 1891)
</taxonomicName>
by
<bibRefCitation id="EFEDFCAA9169FFFCFF79FB3F73E4023C" author="GwiazdowiCz" box="[151,389,1258,1283]" pageId="3" pageNumber="538" refString="Gwiazdowicz, D. J. (2004) Record of heteromorphic males of Hypoaspis (Cosmolaelaps) vacua (Michael, 1891) (Acari, Mesostigmata, Laelapidae) from Poland. Journal of the Acarological Society of Japan, 13, 181 - 184. https: // doi. org / 10.2300 / acari. 13.181" type="journal article" year="2004">GwiazdowiCz (2004)</bibRefCitation>
,
<taxonomicName id="4C7CFAD89169FFFCFE7EFB3E70A6023C" authority="MiChael, 1891" authorityName="MiChael" authorityYear="1891" box="[400,711,1258,1283]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="cuneifer">
<emphasis id="B9085D499169FFFCFE7EFB3E73C2023D" box="[400,419,1259,1282]" italics="true" pageId="3" pageNumber="538">C</emphasis>
.
<emphasis id="B9085D499169FFFCFE5EFB3E706D023C" box="[432,524,1259,1283]" italics="true" pageId="3" pageNumber="538">cuneifer</emphasis>
(MiChael, 1891)
</taxonomicName>
by wisniewski (1982) and
<taxonomicName id="4C7CFAD89169FFFCFC14FB3E7718023C" authority="Huhta &amp; Karg, 2010" authorityName="Huhta &amp; Karg" authorityYear="2010" box="[1018,1401,1258,1283]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="michaeli">
<emphasis id="B9085D499169FFFCFC14FB3E766C023D" box="[1018,1037,1259,1282]" italics="true" pageId="3" pageNumber="538">C</emphasis>
.
<emphasis id="B9085D499169FFFCFBF5FB3E761A023C" box="[1051,1147,1259,1283]" italics="true" pageId="3" pageNumber="538">michaeli</emphasis>
(Huhta &amp; Karg, 2010)
</taxonomicName>
by
<bibRefCitation id="EFEDFCAA9169FFFCFF79FAD870250219" author="Khalili-Moghdam" box="[151,580,1293,1318]" pageId="3" pageNumber="538" refString="Khalili-Moghadam, A. &amp; Saboori, A. (2015) Some mesostigmatic mites (Acari: Mesostigmata) associated with ants in Shahrekord region, Iran. Ecologica Montenegrina, 2, 315 - 326." type="journal article" year="2015">Khalili-Moghdam &amp; Saboori (2015)</bibRefCitation>
. In
<taxonomicName id="4C7CFAD89169FFFCFD97FADB708D021A" box="[633,748,1294,1317]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="vaccus">
<emphasis id="B9085D499169FFFCFD97FADB70ED021A" box="[633,652,1294,1317]" italics="true" pageId="3" pageNumber="538">C</emphasis>
.
<emphasis id="B9085D499169FFFCFD4EFADB708D021A" box="[672,748,1294,1317]" italics="true" pageId="3" pageNumber="538">vaccus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7CFAD89169FFFCFCC1FADB71D7021A" box="[815,950,1293,1317]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="michaeli">
<emphasis id="B9085D499169FFFCFCC1FADB7123021A" box="[815,834,1294,1317]" italics="true" pageId="3" pageNumber="538">C</emphasis>
.
<emphasis id="B9085D499169FFFCFCB8FAD871D7021A" box="[854,950,1293,1317]" italics="true" pageId="3" pageNumber="538">michaeli</emphasis>
</taxonomicName>
, the most distinCtive differenCe between homeomorphiC and heteromorphiC males is the prominent outgrowth in the ventrianal region of the holoventral shield of heteromorphiC males. Additionally, the heteromorphiC male of
<taxonomicName id="4C7CFAD89169FFFCFC27FA86762D0254" box="[969,1100,1363,1387]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="michaeli">
<emphasis id="B9085D499169FFFCFC27FA8671BD0255" box="[969,988,1363,1386]" italics="true" pageId="3" pageNumber="538">C</emphasis>
.
<emphasis id="B9085D499169FFFCFC05FA86762D0254" box="[1003,1100,1363,1387]" italics="true" pageId="3" pageNumber="538">michaeli</emphasis>
</taxonomicName>
has leg IV distinCtly stronger than the others and with two spur-like ventral struCtures (
<bibRefCitation id="EFEDFCAA9169FFFCFCA4FAA0769902B1" author="Khalili-Moghadam" box="[842,1272,1397,1422]" pageId="3" pageNumber="538" refString="Khalili-Moghadam, A. &amp; Saboori, A. (2015) Some mesostigmatic mites (Acari: Mesostigmata) associated with ants in Shahrekord region, Iran. Ecologica Montenegrina, 2, 315 - 326." type="journal article" year="2015">Khalili-Moghadam &amp; Saboori, 2015</bibRefCitation>
), whereas the heteromorphiC male of
<taxonomicName id="4C7CFAD89169FFFCFE70FA4D706F028F" box="[414,526,1432,1456]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="vacuus">
<emphasis id="B9085D499169FFFCFE70FA4D706F028F" box="[414,526,1432,1456]" italics="true" pageId="3" pageNumber="538">C. vacuus</emphasis>
</taxonomicName>
has dorsal shield longer than the respeCtive homeomorphiC male (
<bibRefCitation id="EFEDFCAA9169FFFCFB17FA4272B502EC" author="GwiazdowiCz" pageId="3" pageNumber="538" refString="Gwiazdowicz, D. J. (2004) Record of heteromorphic males of Hypoaspis (Cosmolaelaps) vacua (Michael, 1891) (Acari, Mesostigmata, Laelapidae) from Poland. Journal of the Acarological Society of Japan, 13, 181 - 184. https: // doi. org / 10.2300 / acari. 13.181" type="journal article" year="2004">GwiazdowiCz, 2004</bibRefCitation>
). In the
<taxonomicName id="4C7CFAD89169FFFCFEDEFA6E73A002EC" box="[304,449,1467,1491]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="guttulatus">
<emphasis id="B9085D499169FFFCFEDEFA6E732202ED" box="[304,323,1467,1490]" italics="true" pageId="3" pageNumber="538">C</emphasis>
.
<emphasis id="B9085D499169FFFCFEBFFA6E73A002EC" box="[337,449,1467,1491]" italics="true" pageId="3" pageNumber="538">guttulatus</emphasis>
</taxonomicName>
heteromorphiC male, the tarsus of leg IV is similar to the tarsi of the other legs and the dorsal shield of the homeomorphiC male is longer [463 (419–535)] than that in the heteromorphiC male [387–415].
</paragraph>
</subSubSection>
</treatment>
</document>
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