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<document id="E50A7E2FCC88B37F1467CD079FB99F19" ID-DOI="10.11646/zootaxa.5403.3.10" ID-ISSN="1175-5326" ID-Zenodo-Dep="10561929" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1706104449991" checkinUser="plazi" docAuthor="Henríquez, Carlos M., Sarmiento-Marina, Yessenia, Salgado-Maldonado, Guillermo &amp; Matamoros, Wilfredo A." docDate="2024" docId="F1142849FFAE9B50FEB1F9DDFB9FFD0C" docLanguage="en" docName="zootaxa.5403.3.10.pdf" docOrigin="Zootaxa 5403 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5403.3.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tlaloc portillorum" docType="treatment" docVersion="1" lastPageNumber="399" masterDocId="0D2D5031FFAE9B53FE76FFD4FFFFFFC7" masterDocTitle="Range extension of the Honduran endemic killifish Tlaloc portillorum (Matamoros &amp; Schaefer 2010) (Cyprinodontiformes: Profundulidae): new records from the upper reaches of the Patuca and Choluteca Rivers" masterLastPageNumber="400" masterPageNumber="396" pageNumber="396" updateTime="1727897985641" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="EAB1BAED92648F90E8077331B17DEC09">Range extension of the Honduran endemic killifish Tlaloc portillorum (Matamoros &amp; Schaefer 2010) (Cyprinodontiformes: Profundulidae): new records from the upper reaches of the Patuca and Choluteca Rivers</mods:title>
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<mods:namePart id="B07E5775FD82140FED7296E61DCD0539">Henríquez, Carlos M.</mods:namePart>
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<mods:namePart id="39D9031077166688522A1ECF6717D47B">Sarmiento-Marina, Yessenia</mods:namePart>
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<mods:affiliation id="E3781BC526B732C99EBA2217C7C48150">Doctorado en Ciencias en Biodiversidad y Conservación de Ecosistemas Tropicales, Instituto de Ciencias Biológicas, Universidad de Ciencias y Artes de Chiapas, Tuxtla Gutiérrez, Chiapas, México.</mods:affiliation>
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<mods:affiliation id="07D11B5AF4C9B6C13926613EA665AC6A">Universidad Nacional Autónoma de México, Instituto de Biología Av. Universidad 3000, Coyoacán, C. P. 04510 Ciudad de México.</mods:affiliation>
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<mods:affiliation id="3A61FC1529C23222662A0CC695E3B7D9">Doctorado en Ciencias en Biodiversidad y Conservación de Ecosistemas Tropicales, Instituto de Ciencias Biológicas, Universidad de Ciencias y Artes de Chiapas, Tuxtla Gutiérrez, Chiapas, México.</mods:affiliation>
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<emphasis id="4BC9454DFFAE9B53FEB1F9DDFE72F9E6" box="[199,397,1545,1569]" italics="true" pageId="0" pageNumber="396">Tlaloc portillorum</emphasis>
(
<bibRefCitation id="1D2CE4AEFFAE9B53FFEDF9DDFD2BF9E6" author="Matamoros, W. A. &amp; Schaefer, J. F." box="[411,724,1545,1569]" pageId="0" pageNumber="396" pagination="1498 - 507" refId="ref3234" refString="Matamoros, W. A. &amp; Schaefer, J. F. (2010) A new species of Profundulus (Cyprinodontiformes: Profundulidae) from the Honduran central highlands. Journal of Fish Biology, 76, 1498 - 507. https: // doi. org / 10.1111 / j. 1095 - 8649.2010.02596. x" type="journal article" year="2010">Matamoros &amp; Schaefer 2010</bibRefCitation>
)
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(
<figureCitation id="E18685DAFFAE9B53FC9DF9DDFCC3F9E6" box="[747,828,1545,1569]" captionStart="FIGURE 1" captionStartId="2.[152,255,596,620]" captionTargetBox="[349,1239,180,571]" captionTargetId="figure-23@2.[349,1239,180,571]" captionTargetPageId="2" captionText="FIGURE 1. (A) Map of Honduras showing localities where Tlalocportillorum hasbeen collected.Dashed lines depict majorriver basin borders (basin names in blue font with white background). Major city names in uppercase and bold font. (B) Photograph of live specimen of T. portillorum (5.5 cm standard length) from the Guacerique River (longitude= -87.109, latitude= 14.033), a tributary of the Choluteca River basin. (C) Habitat of T. portillorum. From the Guacerique River, tributary of the Choluteca River basin (longitude -87.111, latitude 14.032)." figureDoi="http://doi.org/10.5281/zenodo.10561935" httpUri="https://zenodo.org/record/10561935/files/figure.png" pageId="0" pageNumber="396">Fig. 1B</figureCitation>
)
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<paragraph id="7902995FFFAE9B53FD3AF9DEFB7AF93E" blockId="0.[151,1437,933,2037]" pageId="0" pageNumber="396">
was described with material from the Ulúa River basin. Until now the only known localities for the species came from the 
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series reported in the original description (
<figureCitation id="E18685DAFFAE9B53FB07F9F9FF46F9AE" captionStart="FIGURE 1" captionStartId="2.[152,255,596,620]" captionTargetBox="[349,1239,180,571]" captionTargetId="figure-23@2.[349,1239,180,571]" captionTargetPageId="2" captionText="FIGURE 1. (A) Map of Honduras showing localities where Tlalocportillorum hasbeen collected.Dashed lines depict majorriver basin borders (basin names in blue font with white background). Major city names in uppercase and bold font. (B) Photograph of live specimen of T. portillorum (5.5 cm standard length) from the Guacerique River (longitude= -87.109, latitude= 14.033), a tributary of the Choluteca River basin. (C) Habitat of T. portillorum. From the Guacerique River, tributary of the Choluteca River basin (longitude -87.111, latitude 14.032)." figureDoi="http://doi.org/10.5281/zenodo.10561935" httpUri="https://zenodo.org/record/10561935/files/figure.png" pageId="0" pageNumber="396">Fig. 1A</figureCitation>
, 
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), a few additional localities in the Nacaome River reported by 
<bibRefCitation id="1D2CE4AEFFAE9B53FDB2F985FADCF9AE" author="Salgado-Maldonado, G. &amp; Matamoros, W. A. &amp; Caspeta-Mandujano, J. M. &amp; Martinez-Ramirez, E. &amp; Mendoza-Franco, E. F. &amp; Velazquez-Velazquez, E." box="[964,1315,1617,1641]" pageId="0" pageNumber="396" pagination="1507 - 1513" refId="ref3571" refString="Salgado-Maldonado, G., Matamoros, W. A., Caspeta-Mandujano, J. M., Martinez-Ramirez, E., Mendoza-Franco, E. F. &amp; Velazquez-Velazquez, E. (2014) Range extension of helminth parasites of Profundulus spp. (Teleostei: Profundulidae) from southern Mexico and Central America. Check List, 10, 1507 - 1513. https: // doi. org / 10.15560 / 10.6.1507" type="journal article" year="2014">
Salgado-Maldonado 
<emphasis id="4BC9454DFFAE9B53FAD2F986FB2BF9AE" box="[1188,1236,1617,1641]" italics="true" pageId="0" pageNumber="396">et al</emphasis>
. (2014
</bibRefCitation>
, 
<bibRefCitation id="1D2CE4AEFFAE9B53FB46F985FA98F9AE" author="Salgado-Maldonado, G. &amp; Matamoros, W. A. &amp; Kreiser, B. R. &amp; Caspeta-Mandujano, J. M. &amp; Mendoza-Franco, E. F." box="[1328,1383,1617,1641]" pageId="0" pageNumber="396" pagination="5" refId="ref3649" refString="Salgado-Maldonado, G., Matamoros, W. A., Kreiser, B. R., Caspeta-Mandujano, J. M. &amp; Mendoza-Franco, E. F. (2015) First record of the invasive Asian fish tapeworm Bothriocephalus acheilognathi in Honduras, Central America. Parasite, 22, 5. https: // doi. org / 10.1051 / parasite / 2015007" type="journal article" year="2015">2015</bibRefCitation>
) and one more locality very close to the 
<typeStatus id="A60627FDFFAE9B53FC79F9A2FDC2F949" box="[527,573,1654,1678]" pageId="0" pageNumber="396">type</typeStatus>
locality reported by 
<bibRefCitation id="1D2CE4AEFFAE9B53FD6DF9A1FBA3F94A" author="Pinacho-Pinacho, C. D. &amp; Garcia-Varela, M. &amp; Mendoza-Palmero, C. &amp; Sereno-Uribe, A. L. &amp; Martinez-Ramirez, E. &amp; Andrade-Gomez, L. &amp; Hernandez-Cruz, E. &amp; Perez Ponce de Leon, G." box="[795,1116,1653,1677]" pageId="0" pageNumber="396" pagination="1 - 30" refId="ref3466" refString="Pinacho-Pinacho, C. D., Garcia-Varela, M., Hernandez-Orts, J. S., Mendoza-Palmero, C. a., Sereno-Uribe, A. L., Martinez-Ramirez, E., Andrade-Gomez, L., Hernandez-Cruz, E. &amp; Perez Ponce de Leon, G. (2015) Checklist of the helminth parasites of the genus Profundulus Hubbs, 1924 (Cyprinodontiformes, Profundulidae), an endemic family of freshwater fishes in Middle-America. ZooKeys, 523, 1 - 30. https: // doi. org / 10.3897 / zookeys. 523.6088" type="journal article" year="2015">
Pinacho-Pinacho 
<emphasis id="4BC9454DFFAE9B53FDA0F9A2FBF3F94A" box="[982,1036,1653,1677]" italics="true" pageId="0" pageNumber="396">et al.</emphasis>
(2015)
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. Although, 
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<emphasis id="4BC9454DFFAE9B53FAA1F9A2FA93F94A" box="[1239,1388,1653,1677]" italics="true" pageId="0" pageNumber="396">T. portillorum</emphasis>
</taxonomicName>
was described more than a decade ago, basic knowledge about its biology, ecology and life history are still lacking. To date, there are only a few papers in the scientific literature dealing with the species (
<emphasis id="4BC9454DFFAE9B53FDAEF96AFC01F912" box="[984,1022,1726,1749]" italics="true" pageId="0" pageNumber="396">e.g.</emphasis>
<bibRefCitation id="1D2CE4AEFFAE9B53FA73F969FACBF912" author="Pinacho-Pinacho, C. D. &amp; Garcia-Varela, M. &amp; Mendoza-Palmero, C. &amp; Sereno-Uribe, A. L. &amp; Martinez-Ramirez, E. &amp; Andrade-Gomez, L. &amp; Hernandez-Cruz, E. &amp; Perez Ponce de Leon, G." box="[1029,1332,1725,1749]" pageId="0" pageNumber="396" pagination="1 - 30" refId="ref3466" refString="Pinacho-Pinacho, C. D., Garcia-Varela, M., Hernandez-Orts, J. S., Mendoza-Palmero, C. a., Sereno-Uribe, A. L., Martinez-Ramirez, E., Andrade-Gomez, L., Hernandez-Cruz, E. &amp; Perez Ponce de Leon, G. (2015) Checklist of the helminth parasites of the genus Profundulus Hubbs, 1924 (Cyprinodontiformes, Profundulidae), an endemic family of freshwater fishes in Middle-America. ZooKeys, 523, 1 - 30. https: // doi. org / 10.3897 / zookeys. 523.6088" type="journal article" year="2015">
Pinacho-Pinacho 
<emphasis id="4BC9454DFFAE9B53FAB6F96AFB0AF912" box="[1216,1269,1725,1749]" italics="true" pageId="0" pageNumber="396">et al.</emphasis>
2015
</bibRefCitation>
; 
<bibRefCitation id="1D2CE4AEFFAE9B53FB4BF969FE74F93E" author="Salgado-Maldonado, G. &amp; Matamoros, W. A. &amp; Caspeta-Mandujano, J. M. &amp; Martinez-Ramirez, E. &amp; Mendoza-Franco, E. F. &amp; Velazquez-Velazquez, E." pageId="0" pageNumber="396" pagination="1507 - 1513" refId="ref3571" refString="Salgado-Maldonado, G., Matamoros, W. A., Caspeta-Mandujano, J. M., Martinez-Ramirez, E., Mendoza-Franco, E. F. &amp; Velazquez-Velazquez, E. (2014) Range extension of helminth parasites of Profundulus spp. (Teleostei: Profundulidae) from southern Mexico and Central America. Check List, 10, 1507 - 1513. https: // doi. org / 10.15560 / 10.6.1507" type="journal article" year="2014">
Salgado-Maldonado 
<emphasis id="4BC9454DFFAE9B53FF61F936FEB2F93E" box="[279,333,1761,1785]" italics="true" pageId="0" pageNumber="396">et al.</emphasis>
2014
</bibRefCitation>
, 
<bibRefCitation id="1D2CE4AEFFAE9B53FFE0F935FE30F93E" author="Salgado-Maldonado, G. &amp; Matamoros, W. A. &amp; Kreiser, B. R. &amp; Caspeta-Mandujano, J. M. &amp; Mendoza-Franco, E. F." box="[406,463,1761,1785]" pageId="0" pageNumber="396" pagination="5" refId="ref3649" refString="Salgado-Maldonado, G., Matamoros, W. A., Kreiser, B. R., Caspeta-Mandujano, J. M. &amp; Mendoza-Franco, E. F. (2015) First record of the invasive Asian fish tapeworm Bothriocephalus acheilognathi in Honduras, Central America. Parasite, 22, 5. https: // doi. org / 10.1051 / parasite / 2015007" type="journal article" year="2015">2015</bibRefCitation>
), and these are focused on describing their parasitic associations.
</paragraph>
<paragraph id="7902995FFFAE9B53FEB1F8D2FABCF832" blockId="0.[151,1437,933,2037]" pageId="0" pageNumber="396">
<collectingCounty id="9063E1D3FFAE9B53FEB1F8D2FE69F8DA" box="[199,406,1797,1821]" pageId="0" pageNumber="396">
In 
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</collectingCounty>
, Honduras is one of the countries with the lowest number of endemic freshwater fish species (4), and 
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<emphasis id="4BC9454DFFAE9B53FEB1F8FEFF29F886" box="[199,214,1834,1857]" italics="true" pageId="0" pageNumber="396">T</emphasis>
. 
<emphasis id="4BC9454DFFAE9B53FE90F8FDFE9FF886" box="[230,352,1833,1857]" italics="true" pageId="0" pageNumber="396">portillorum</emphasis>
</taxonomicName>
is the only endemic profundulid (
<bibRefCitation id="1D2CE4AEFFAE9B53FCAEF8FDFBC6F886" author="Contreras-MacBeath, T. &amp; Ardon, D. A. &amp; Quintana, Y. &amp; Angulo, A. &amp; Lyons, T. &amp; Lardizabal, C. C. &amp; Mcmahan, C. D. &amp; Elias, D. J. &amp; Matamoros, W. A. &amp; Barraza, J. E. &amp; Gonzalez, R. &amp; Fuentes-Montejo, C. E. &amp; Ambruster, J. W. &amp; Carrasco, J. C. &amp; Brito Rodriguez, M." box="[728,1081,1833,1857]" pageId="0" pageNumber="396" pagination="1 - 24" refId="ref2739" refString="Contreras-MacBeath, T., Ardon, D. A., Quintana, Y., Angulo, A., Lyons, T., Lardizabal, C. C., Mcmahan, C. D., Elias, D. J., Matamoros, W. A., Barraza, J. E., Gonzalez, R., Fuentes-Montejo, C. E., Ambruster, J. W., Carrasco, J. C. &amp; Brito Rodriguez, M. (2022) Freshwater fishes of Central America: distribution, assessment, and major threats. Diversity, 14, 1 - 24. https: // doi. org / 10.3390 / d 14100793" type="journal article" year="2022">
Contreras-MacBeath 
<emphasis id="4BC9454DFFAE9B53FDB6F8FEFC07F886" box="[960,1016,1833,1857]" italics="true" pageId="0" pageNumber="396">et al.</emphasis>
2022
</bibRefCitation>
; 
<bibRefCitation id="1D2CE4AEFFAE9B53FA31F8FEFABBF886" author="Matamoros, W. A. &amp; Schaefer, J. F. &amp; Kreiser, B. R." box="[1095,1348,1833,1857]" pageId="0" pageNumber="396" pagination="1 - 38" refId="ref3293" refString="Matamoros, W. A., Schaefer, J. F. &amp; Kreiser, B. R. (2009) Annotated checklist of the freshwater fishes of continental and insular Honduras. Zootaxa, 2307 (1), 1 - 38. https: // doi. org / 10.11646 / zootaxa. 2307.1.1" type="journal article" year="2009">
Matamoros 
<emphasis id="4BC9454DFFAE9B53FABFF8FEFAFDF886" box="[1225,1282,1833,1857]" italics="true" pageId="0" pageNumber="396">et al.</emphasis>
2009
</bibRefCitation>
). 
<taxonomicName id="BEBDE2DCFFAE9B53FB2FF8FDFEEEF8A2" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="0" pageNumber="396" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAE9B53FB2FF8FDFEEEF8A2" italics="true" pageId="0" pageNumber="396">Tlaloc portillorum</emphasis>
</taxonomicName>
is known for localities in the upper reaches of the 
<location id="7C62CF84FFAE9B53FD52F899FC66F8A2" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAE9B53FD52F899FC66F8A2" box="[804,921,1869,1893]" country="Honduras" county="In Central America" municipality="Central America" name="Ulua River" pageId="0" pageNumber="396" stateProvince="Comayagua">Ulúa River</location>
near the city of 
<location id="7C62CF84FFAE9B53FA35F899FB2BF8A2" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAE9B53FA35F899FB2BF8A2" box="[1091,1236,1869,1893]" country="Honduras" county="In Central America" municipality="Central America" name="Siguatepeque" pageId="0" pageNumber="396" stateProvince="Comayagua">Siguatepeque</location>
, in the 
<location id="7C62CF84FFAE9B53FB56F89AFA62F8A2" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAE9B53FB56F89AFA62F8A2" box="[1312,1437,1870,1893]" country="Honduras" county="In Central America" municipality="Central America" name="Department" pageId="0" pageNumber="396" stateProvince="Comayagua">Department</location>
of Comayagua (
<figureCitation id="E18685DAFFAE9B53FF31F8A5FE61F84E" box="[327,414,1905,1929]" captionStart="FIGURE 1" captionStartId="2.[152,255,596,620]" captionTargetBox="[349,1239,180,571]" captionTargetId="figure-23@2.[349,1239,180,571]" captionTargetPageId="2" captionText="FIGURE 1. (A) Map of Honduras showing localities where Tlalocportillorum hasbeen collected.Dashed lines depict majorriver basin borders (basin names in blue font with white background). Major city names in uppercase and bold font. (B) Photograph of live specimen of T. portillorum (5.5 cm standard length) from the Guacerique River (longitude= -87.109, latitude= 14.033), a tributary of the Choluteca River basin. (C) Habitat of T. portillorum. From the Guacerique River, tributary of the Choluteca River basin (longitude -87.111, latitude 14.032)." figureDoi="http://doi.org/10.5281/zenodo.10561935" httpUri="https://zenodo.org/record/10561935/files/figure.png" pageId="0" pageNumber="396">Fig. 1A</figureCitation>
), as well as localities of the 
<location id="7C62CF84FFAE9B53FC97F8A6FC75F84E" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAE9B53FC97F8A6FC75F84E" box="[737,906,1905,1929]" country="Honduras" county="In Central America" municipality="Central America" name="Nacaome River" pageId="0" pageNumber="396" stateProvince="Comayagua">Nacaome River</location>
in the towns of 
<location id="7C62CF84FFAE9B53FA30F8A5FB3BF84E" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAE9B53FA30F8A5FB3BF84E" box="[1094,1220,1905,1929]" country="Honduras" county="In Central America" municipality="Central America" name="Lepaterique" pageId="0" pageNumber="396" stateProvince="Comayagua">Lepaterique</location>
and 
<location id="7C62CF84FFAE9B53FA88F8A5FAACF84E" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAE9B53FA88F8A5FAACF84E" box="[1278,1363,1905,1929]" country="Honduras" county="In Central America" municipality="Central America" name="Ojojona" pageId="0" pageNumber="396" stateProvince="Comayagua">Ojojona</location>
in the 
<location id="7C62CF84FFAE9B53FEE1F842FEEBF86A" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAE9B53FEE1F842FEEBF86A" box="[151,276,1942,1965]" country="Honduras" county="In Central America" municipality="Central America" name="Department" pageId="0" pageNumber="396" stateProvince="Comayagua">Department</location>
of Francisco Morazán. 
<location id="7C62CF84FFAE9B53FC7CF842FDDFF86A" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAE9B53FC7CF842FDDFF86A" box="[522,544,1942,1965]" country="Honduras" county="In Central America" municipality="Central America" name="In" pageId="0" pageNumber="396" stateProvince="Comayagua">In</location>
the original description, 
<taxonomicName id="BEBDE2DCFFAE9B53FD52F842FC48F86A" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[804,951,1941,1965]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="0" pageNumber="396" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAE9B53FD52F842FC48F86A" box="[804,951,1941,1965]" italics="true" pageId="0" pageNumber="396">T. portillorum</emphasis>
</taxonomicName>
is reported to inhabit shallow creeks with very small width (
<elevation id="F2907E6CFFAE9B53FF56F86DFE9BF816" box="[288,356,1977,2001]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="0" pageNumber="396" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">3–5 m</elevation>
) and bottoms dominated by large cobblestone (
<bibRefCitation id="1D2CE4AEFFAE9B53FD24F86DFB78F816" author="Matamoros, W. A. &amp; Schaefer, J. F." box="[850,1159,1977,2001]" pageId="0" pageNumber="396" pagination="1498 - 507" refId="ref3234" refString="Matamoros, W. A. &amp; Schaefer, J. F. (2010) A new species of Profundulus (Cyprinodontiformes: Profundulidae) from the Honduran central highlands. Journal of Fish Biology, 76, 1498 - 507. https: // doi. org / 10.1111 / j. 1095 - 8649.2010.02596. x" type="journal article" year="2010">Matamoros &amp; Schaefer 2010</bibRefCitation>
). It was found in sympatry with 
<taxonomicName id="BEBDE2DCFFAE9B53FEBBF809FD93F832" authority="(Linnaeus 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[205,620,2013,2037]" family="Cichlidae" genus="Oreochromis" kingdom="Animalia" order="Perciformes" pageId="0" pageNumber="396" phylum="Chordata" rank="species" species="niloticus">
<emphasis id="4BC9454DFFAE9B53FEBBF809FE47F832" box="[205,440,2013,2037]" italics="true" pageId="0" pageNumber="396">Oreochromis niloticus</emphasis>
(Linnaeus 1758)
</taxonomicName>
and 
<taxonomicName id="BEBDE2DCFFAE9B53FCE9F809FAC0F832" authority="(Kner 1858) (Matamoros &amp; Schaefer 2010)" authorityName="Matamoros &amp; Schaefer" authorityYear="2010" baseAuthorityName="Kner" baseAuthorityYear="1858" box="[671,1343,2013,2037]" family="Heptapteridae" genus="Rhamdia" kingdom="Animalia" order="Siluriformes" pageId="0" pageNumber="396" phylum="Chordata" rank="species" species="laticauda">
<emphasis id="4BC9454DFFAE9B53FCE9F809FC96F832" box="[671,873,2013,2037]" italics="true" pageId="0" pageNumber="396">Rhamdia laticauda</emphasis>
(Kner 1858) (
<bibRefCitation id="1D2CE4AEFFAE9B53FD89F809FAC8F832" author="Matamoros, W. A. &amp; Schaefer, J. F." box="[1023,1335,2013,2037]" pageId="0" pageNumber="396" pagination="1498 - 507" refId="ref3234" refString="Matamoros, W. A. &amp; Schaefer, J. F. (2010) A new species of Profundulus (Cyprinodontiformes: Profundulidae) from the Honduran central highlands. Journal of Fish Biology, 76, 1498 - 507. https: // doi. org / 10.1111 / j. 1095 - 8649.2010.02596. x" type="journal article" year="2010">Matamoros &amp; Schaefer 2010</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="2DC2C9D7FFAF9B52FEEEFF43FC78FF30" ID-Table-UUID="2DC2C9D7FFAF9B52FEEEFF43FC78FF30" httpUri="http://table.plazi.org/id/2DC2C9D7FFAF9B52FEEEFF43FC78FF30" pageId="1" pageNumber="397" startId="1.[152,245,151,175]" targetBox="[165,1418,266,1892]" targetIsTable="true" targetPageId="1" targetType="table">
<paragraph id="7902995FFFAF9B52FEEEFF43FC78FF30" blockId="1.[151,1436,151,248]" pageId="1" pageNumber="397">
<emphasis id="4BC9454DFFAF9B52FEEEFF43FEEFFF68" bold="true" box="[152,272,151,175]" pageId="1" pageNumber="397">TABLE 1.</emphasis>
Table with geographic information, for each sample included in Figure one. Type series as in 
<bibRefCitation id="1D2CE4AEFFAF9B52FB71FF4CFEC3FF13" author="Matamoros, W. A. &amp; Schaefer, J. F." pageId="1" pageNumber="397" pagination="1498 - 507" refId="ref3234" refString="Matamoros, W. A. &amp; Schaefer, J. F. (2010) A new species of Profundulus (Cyprinodontiformes: Profundulidae) from the Honduran central highlands. Journal of Fish Biology, 76, 1498 - 507. https: // doi. org / 10.1111 / j. 1095 - 8649.2010.02596. x" type="journal article" year="2010">Matamoros &amp; Schaefer, 2010</bibRefCitation>
; The source Museum collections LSUMZ= Louisiana State University Museum Zoology, collection of fishes; USM= University of Southern Mississippi, collection of fishes.
</paragraph>
</caption>
<paragraph id="7902995FFFAF9B52FED3FEDFFB0EF880" pageId="1" pageNumber="397">
<table id="0BBD6BFFFFAF64ACFED3FEDEFA75F8A3" box="[165,1418,266,1892]" gridcols="7" gridrows="40" pageId="1" pageNumber="397">
<tr id="C78D9B1DFFAF64ACFED3FEDEFA75FEE4" box="[165,1418,266,291]" gridrow="0" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FEDEFF3EFEE4" box="[165,193,266,291]" gridcol="0" gridrow="0" pageId="1" pageNumber="397">
<emphasis id="4BC9454DFFAF9B52FED3FEDFFF3EFEE4" bold="true" box="[165,193,267,291]" pageId="1" pageNumber="397">ID</emphasis>
</th>
<th id="845CF261FFAF64ACFEACFEDEFE97FEE4" box="[218,360,266,291]" gridcol="1" gridrow="0" pageId="1" pageNumber="397">
<emphasis id="4BC9454DFFAF9B52FEACFEDEFE97FEE4" bold="true" box="[218,360,266,291]" pageId="1" pageNumber="397">City or Town</emphasis>
</th>
<th id="845CF261FFAF64ACFFFFFEDEFD92FEE4" box="[393,621,266,291]" gridcol="2" gridrow="0" pageId="1" pageNumber="397">
<emphasis id="4BC9454DFFAF9B52FFFFFEDEFDC6FEE5" bold="true" box="[393,569,266,290]" pageId="1" pageNumber="397">Basin/Sub-basin</emphasis>
</th>
<th id="845CF261FFAF64ACFCE6FEDEFD00FEE4" box="[656,767,266,291]" gridcol="3" gridrow="0" pageId="1" pageNumber="397">
<emphasis id="4BC9454DFFAF9B52FCE6FEDEFD00FEE5" bold="true" box="[656,767,266,290]" pageId="1" pageNumber="397">Longitude</emphasis>
</th>
<th id="845CF261FFAF64ACFD52FEDEFC7EFEE4" box="[804,897,266,291]" gridcol="4" gridrow="0" pageId="1" pageNumber="397">
<emphasis id="4BC9454DFFAF9B52FD52FEDEFC7EFEE5" bold="true" box="[804,897,266,290]" pageId="1" pageNumber="397">Latitude</emphasis>
</th>
<th id="845CF261FFAF64ACFDDCFEDEFBCFFEE4" box="[938,1072,266,291]" gridcol="5" gridrow="0" pageId="1" pageNumber="397">
<emphasis id="4BC9454DFFAF9B52FDDCFEDEFBCFFEE5" bold="true" box="[938,1072,266,290]" pageId="1" pageNumber="397">Altitude (m)</emphasis>
</th>
<th id="845CF261FFAF64ACFA3CFEDEFA75FEE4" box="[1098,1418,266,291]" gridcol="6" gridrow="0" pageId="1" pageNumber="397">
<emphasis id="4BC9454DFFAF9B52FA3CFEDEFB6BFEE5" bold="true" box="[1098,1172,266,290]" pageId="1" pageNumber="397">Source</emphasis>
</th>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FEE2FA75FE89" box="[165,1418,310,334]" gridrow="1" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FEE2FF3EFE89" box="[165,193,310,334]" gridcol="0" gridrow="1" pageId="1" pageNumber="397">1</th>
<td id="845CF261FFAF64ACFEACFEE2FE97FE89" box="[218,360,310,334]" gridcol="1" gridrow="1" pageId="1" pageNumber="397">Siguatepeque</td>
<td id="845CF261FFAF64ACFFFFFEE2FD92FE89" box="[393,621,310,334]" gridcol="2" gridrow="1" pageId="1" pageNumber="397">Ulúa-Calam</td>
<td id="845CF261FFAF64ACFCE6FEE2FD00FE89" box="[656,767,310,334]" gridcol="3" gridrow="1" pageId="1" pageNumber="397">-87.876</td>
<td id="845CF261FFAF64ACFD52FEE2FC7EFE89" box="[804,897,310,334]" gridcol="4" gridrow="1" pageId="1" pageNumber="397">14.533</td>
<td id="845CF261FFAF64ACFDDCFEE2FBCFFE89" box="[938,1072,310,334]" gridcol="5" gridrow="1" pageId="1" pageNumber="397">1368</td>
<td id="845CF261FFAF64ACFA3CFEE2FA75FE89" box="[1098,1418,310,334]" gridcol="6" gridrow="1" pageId="1" pageNumber="397">Holotype/USM31400</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FE8AFA75FEB1" box="[165,1418,350,374]" gridrow="2" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FE8AFF3EFEB1" box="[165,193,350,374]" gridcol="0" gridrow="2" pageId="1" pageNumber="397">2</th>
<td id="845CF261FFAF64ACFEACFE8AFE97FEB1" box="[218,360,350,374]" gridcol="1" gridrow="2" pageId="1" pageNumber="397">Siguatepeque</td>
<td id="845CF261FFAF64ACFFFFFE8AFD92FEB1" box="[393,621,350,374]" gridcol="2" gridrow="2" pageId="1" pageNumber="397">Ulúa-Calam</td>
<td id="845CF261FFAF64ACFCE6FE8AFD00FEB1" box="[656,767,350,374]" gridcol="3" gridrow="2" pageId="1" pageNumber="397">-87.876</td>
<td id="845CF261FFAF64ACFD52FE8AFC7EFEB1" box="[804,897,350,374]" gridcol="4" gridrow="2" pageId="1" pageNumber="397">14.533</td>
<td id="845CF261FFAF64ACFDDCFE8AFBCFFEB1" box="[938,1072,350,374]" gridcol="5" gridrow="2" pageId="1" pageNumber="397">1368</td>
<td id="845CF261FFAF64ACFA3CFE8AFA75FEB1" box="[1098,1418,350,374]" gridcol="6" gridrow="2" pageId="1" pageNumber="397">Paratypes/USM31589</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FE52FA75FE59" box="[165,1418,390,414]" gridrow="3" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FE52FF3EFE59" box="[165,193,390,414]" gridcol="0" gridrow="3" pageId="1" pageNumber="397">3</th>
<td id="845CF261FFAF64ACFEACFE52FE97FE59" box="[218,360,390,414]" gridcol="1" gridrow="3" pageId="1" pageNumber="397">Siguatepeque</td>
<td id="845CF261FFAF64ACFFFFFE52FD92FE59" box="[393,621,390,414]" gridcol="2" gridrow="3" pageId="1" pageNumber="397">Ulúa-Calam</td>
<td id="845CF261FFAF64ACFCE6FE52FD00FE59" box="[656,767,390,414]" gridcol="3" gridrow="3" pageId="1" pageNumber="397">-87.878</td>
<td id="845CF261FFAF64ACFD52FE52FC7EFE59" box="[804,897,390,414]" gridcol="4" gridrow="3" pageId="1" pageNumber="397">14.554</td>
<td id="845CF261FFAF64ACFDDCFE52FBCFFE59" box="[938,1072,390,414]" gridcol="5" gridrow="3" pageId="1" pageNumber="397">1368</td>
<td id="845CF261FFAF64ACFA3CFE52FA75FE59" box="[1098,1418,390,414]" gridcol="6" gridrow="3" pageId="1" pageNumber="397">Paratypes/USM31598</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FE7AFA75FE00" box="[165,1418,430,455]" gridrow="4" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FE7AFF3EFE00" box="[165,193,430,455]" gridcol="0" gridrow="4" pageId="1" pageNumber="397">4</th>
<td id="845CF261FFAF64ACFEACFE7AFE97FE00" box="[218,360,430,455]" gridcol="1" gridrow="4" pageId="1" pageNumber="397">Siguatepeque</td>
<td id="845CF261FFAF64ACFFFFFE7AFD92FE00" box="[393,621,430,455]" gridcol="2" gridrow="4" pageId="1" pageNumber="397">Ulúa-Calam</td>
<td id="845CF261FFAF64ACFCE6FE7AFD00FE00" box="[656,767,430,455]" gridcol="3" gridrow="4" pageId="1" pageNumber="397">-87.878</td>
<td id="845CF261FFAF64ACFD52FE7AFC7EFE00" box="[804,897,430,455]" gridcol="4" gridrow="4" pageId="1" pageNumber="397">14.554</td>
<td id="845CF261FFAF64ACFDDCFE7AFBCFFE00" box="[938,1072,430,455]" gridcol="5" gridrow="4" pageId="1" pageNumber="397">1368</td>
<td id="845CF261FFAF64ACFA3CFE7AFA75FE00" box="[1098,1418,430,455]" gridcol="6" gridrow="4" pageId="1" pageNumber="397">Paratypes/USM31592</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FE03FA75FE28" box="[165,1418,471,495]" gridrow="5" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FE03FF3EFE28" box="[165,193,471,495]" gridcol="0" gridrow="5" pageId="1" pageNumber="397">5</th>
<td id="845CF261FFAF64ACFEACFE03FE97FE28" box="[218,360,471,495]" gridcol="1" gridrow="5" pageId="1" pageNumber="397">Siguatepeque</td>
<td id="845CF261FFAF64ACFFFFFE03FD92FE28" box="[393,621,471,495]" gridcol="2" gridrow="5" pageId="1" pageNumber="397">Ulúa-Calam</td>
<td id="845CF261FFAF64ACFCE6FE03FD00FE28" box="[656,767,471,495]" gridcol="3" gridrow="5" pageId="1" pageNumber="397">-87.881</td>
<td id="845CF261FFAF64ACFD52FE03FC7EFE28" box="[804,897,471,495]" gridcol="4" gridrow="5" pageId="1" pageNumber="397">14.541</td>
<td id="845CF261FFAF64ACFDDCFE03FBCFFE28" box="[938,1072,471,495]" gridcol="5" gridrow="5" pageId="1" pageNumber="397">1414</td>
<td id="845CF261FFAF64ACFA3CFE03FA75FE28" box="[1098,1418,471,495]" gridcol="6" gridrow="5" pageId="1" pageNumber="397">Paratypes/USM31597</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FE2BFA75FDD0" box="[165,1418,511,535]" gridrow="6" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FE2BFF3EFDD0" box="[165,193,511,535]" gridcol="0" gridrow="6" pageId="1" pageNumber="397">6</th>
<td id="845CF261FFAF64ACFEACFE2BFE97FDD0" box="[218,360,511,535]" gridcol="1" gridrow="6" pageId="1" pageNumber="397">Ojojona</td>
<td id="845CF261FFAF64ACFFFFFE2BFD92FDD0" box="[393,621,511,535]" gridcol="2" gridrow="6" pageId="1" pageNumber="397">Nacaome-Verdugo</td>
<td id="845CF261FFAF64ACFCE6FE2BFD00FDD0" box="[656,767,511,535]" gridcol="3" gridrow="6" pageId="1" pageNumber="397">-87.294</td>
<td id="845CF261FFAF64ACFD52FE2BFC7EFDD0" box="[804,897,511,535]" gridcol="4" gridrow="6" pageId="1" pageNumber="397">13.928</td>
<td id="845CF261FFAF64ACFDDCFE2BFBCFFDD0" box="[938,1072,511,535]" gridcol="5" gridrow="6" pageId="1" pageNumber="397">1391</td>
<td id="845CF261FFAF64ACFA3CFE2BFA75FDD0" box="[1098,1418,511,535]" gridcol="6" gridrow="6" pageId="1" pageNumber="397">
<bibRefCitation id="1D2CE4AEFFAF9B52FA3CFE2BFA75FDD0" author="Salgado-Maldonado, G. &amp; Matamoros, W. A. &amp; Kreiser, B. R. &amp; Caspeta-Mandujano, J. M. &amp; Mendoza-Franco, E. F." box="[1098,1418,511,535]" pageId="1" pageNumber="397" pagination="5" refId="ref3649" refString="Salgado-Maldonado, G., Matamoros, W. A., Kreiser, B. R., Caspeta-Mandujano, J. M. &amp; Mendoza-Franco, E. F. (2015) First record of the invasive Asian fish tapeworm Bothriocephalus acheilognathi in Honduras, Central America. Parasite, 22, 5. https: // doi. org / 10.1051 / parasite / 2015007" type="journal article" year="2015">
Salgado-Maldonado 
<emphasis id="4BC9454DFFAF9B52FB68FDD4FAB4FDD0" box="[1310,1355,511,535]" italics="true" pageId="1" pageNumber="397">et al</emphasis>
. 2015
</bibRefCitation>
</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FDF3FA75FD87" box="[165,1418,551,576]" gridrow="7" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FDF3FF3EFD87" box="[165,193,551,576]" gridcol="0" gridrow="7" pageId="1" pageNumber="397">7</th>
<td id="845CF261FFAF64ACFEACFDF3FE97FD87" box="[218,360,551,576]" gridcol="1" gridrow="7" pageId="1" pageNumber="397">Lepaterique</td>
<td id="845CF261FFAF64ACFFFFFDF3FD92FD87" box="[393,621,551,576]" gridcol="2" gridrow="7" pageId="1" pageNumber="397">Nacaome-Verdugo</td>
<td id="845CF261FFAF64ACFCE6FDF3FD00FD87" box="[656,767,551,576]" gridcol="3" gridrow="7" pageId="1" pageNumber="397">-87.466</td>
<td id="845CF261FFAF64ACFD52FDF3FC7EFD87" box="[804,897,551,576]" gridcol="4" gridrow="7" pageId="1" pageNumber="397">14.061</td>
<td id="845CF261FFAF64ACFDDCFDF3FBCFFD87" box="[938,1072,551,576]" gridcol="5" gridrow="7" pageId="1" pageNumber="397">1462</td>
<td id="845CF261FFAF64ACFA3CFDF3FA75FD87" box="[1098,1418,551,576]" gridcol="6" gridrow="7" pageId="1" pageNumber="397">
<bibRefCitation id="1D2CE4AEFFAF9B52FA3CFDF3FA75FD87" author="Salgado-Maldonado, G. &amp; Matamoros, W. A. &amp; Caspeta-Mandujano, J. M. &amp; Martinez-Ramirez, E. &amp; Mendoza-Franco, E. F. &amp; Velazquez-Velazquez, E." box="[1098,1418,551,576]" pageId="1" pageNumber="397" pagination="1507 - 1513" refId="ref3571" refString="Salgado-Maldonado, G., Matamoros, W. A., Caspeta-Mandujano, J. M., Martinez-Ramirez, E., Mendoza-Franco, E. F. &amp; Velazquez-Velazquez, E. (2014) Range extension of helminth parasites of Profundulus spp. (Teleostei: Profundulidae) from southern Mexico and Central America. Check List, 10, 1507 - 1513. https: // doi. org / 10.15560 / 10.6.1507" type="journal article" year="2014">
Salgado-Maldonado 
<emphasis id="4BC9454DFFAF9B52FB68FDFCFAB4FDF8" box="[1310,1355,551,575]" italics="true" pageId="1" pageNumber="397">et al</emphasis>
. 2014
</bibRefCitation>
</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FD9BFA75FDAF" box="[165,1418,591,616]" gridrow="8" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FD9BFF3EFDAF" box="[165,193,591,616]" gridcol="0" gridrow="8" pageId="1" pageNumber="397">8</th>
<td id="845CF261FFAF64ACFEACFD9BFE97FDAF" box="[218,360,591,616]" gridcol="1" gridrow="8" pageId="1" pageNumber="397">Lepaterique</td>
<td id="845CF261FFAF64ACFFFFFD9BFD92FDAF" box="[393,621,591,616]" gridcol="2" gridrow="8" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FD9BFD00FDAF" box="[656,767,591,616]" gridcol="3" gridrow="8" pageId="1" pageNumber="397">-87.432</td>
<td id="845CF261FFAF64ACFD52FD9BFC7EFDAF" box="[804,897,591,616]" gridcol="4" gridrow="8" pageId="1" pageNumber="397">14.070</td>
<td id="845CF261FFAF64ACFDDCFD9BFBCFFDAF" box="[938,1072,591,616]" gridcol="5" gridrow="8" pageId="1" pageNumber="397">1656</td>
<td id="845CF261FFAF64ACFA3CFD9BFA75FDAF" box="[1098,1418,591,616]" gridcol="6" gridrow="8" pageId="1" pageNumber="397">
<bibRefCitation id="1D2CE4AEFFAF9B52FA3CFD84FA75FDAF" author="Salgado-Maldonado, G. &amp; Matamoros, W. A. &amp; Caspeta-Mandujano, J. M. &amp; Martinez-Ramirez, E. &amp; Mendoza-Franco, E. F. &amp; Velazquez-Velazquez, E." box="[1098,1418,592,616]" pageId="1" pageNumber="397" pagination="1507 - 1513" refId="ref3571" refString="Salgado-Maldonado, G., Matamoros, W. A., Caspeta-Mandujano, J. M., Martinez-Ramirez, E., Mendoza-Franco, E. F. &amp; Velazquez-Velazquez, E. (2014) Range extension of helminth parasites of Profundulus spp. (Teleostei: Profundulidae) from southern Mexico and Central America. Check List, 10, 1507 - 1513. https: // doi. org / 10.15560 / 10.6.1507" type="journal article" year="2014">
Salgado-Maldonado 
<emphasis id="4BC9454DFFAF9B52FB68FD84FAB4FDAF" box="[1310,1355,592,616]" italics="true" pageId="1" pageNumber="397">et al</emphasis>
. 2014
</bibRefCitation>
</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FDACFA75FD57" box="[165,1418,632,656]" gridrow="9" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FDACFF3EFD57" box="[165,193,632,656]" gridcol="0" gridrow="9" pageId="1" pageNumber="397">9</th>
<td id="845CF261FFAF64ACFEACFDACFE97FD57" box="[218,360,632,656]" gridcol="1" gridrow="9" pageId="1" pageNumber="397">Ojojona</td>
<td id="845CF261FFAF64ACFFFFFDACFD92FD57" box="[393,621,632,656]" gridcol="2" gridrow="9" pageId="1" pageNumber="397">Nacaome-Verdugo</td>
<td id="845CF261FFAF64ACFCE6FDACFD00FD57" box="[656,767,632,656]" gridcol="3" gridrow="9" pageId="1" pageNumber="397">-87.294</td>
<td id="845CF261FFAF64ACFD52FDACFC7EFD57" box="[804,897,632,656]" gridcol="4" gridrow="9" pageId="1" pageNumber="397">13.928</td>
<td id="845CF261FFAF64ACFDDCFDACFBCFFD57" box="[938,1072,632,656]" gridcol="5" gridrow="9" pageId="1" pageNumber="397">1391</td>
<td id="845CF261FFAF64ACFA3CFDACFA75FD57" box="[1098,1418,632,656]" gridcol="6" gridrow="9" pageId="1" pageNumber="397">
<bibRefCitation id="1D2CE4AEFFAF9B52FA3CFDACFA75FD57" author="Salgado-Maldonado, G. &amp; Matamoros, W. A. &amp; Caspeta-Mandujano, J. M. &amp; Martinez-Ramirez, E. &amp; Mendoza-Franco, E. F. &amp; Velazquez-Velazquez, E." box="[1098,1418,632,656]" pageId="1" pageNumber="397" pagination="1507 - 1513" refId="ref3571" refString="Salgado-Maldonado, G., Matamoros, W. A., Caspeta-Mandujano, J. M., Martinez-Ramirez, E., Mendoza-Franco, E. F. &amp; Velazquez-Velazquez, E. (2014) Range extension of helminth parasites of Profundulus spp. (Teleostei: Profundulidae) from southern Mexico and Central America. Check List, 10, 1507 - 1513. https: // doi. org / 10.15560 / 10.6.1507" type="journal article" year="2014">
Salgado-Maldonado 
<emphasis id="4BC9454DFFAF9B52FB68FDADFAB4FD57" box="[1310,1355,632,656]" italics="true" pageId="1" pageNumber="397">et al</emphasis>
. 2014
</bibRefCitation>
</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FD74FA75FD7F" box="[165,1418,672,696]" gridrow="10" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FD74FF3EFD7F" box="[165,193,672,696]" gridcol="0" gridrow="10" pageId="1" pageNumber="397">10</th>
<td id="845CF261FFAF64ACFEACFD74FE97FD7F" box="[218,360,672,696]" gridcol="1" gridrow="10" pageId="1" pageNumber="397">Potrerillos</td>
<td id="845CF261FFAF64ACFFFFFD74FD92FD7F" box="[393,621,672,696]" gridcol="2" gridrow="10" pageId="1" pageNumber="397">Ulúa-Selguapa</td>
<td id="845CF261FFAF64ACFCE6FD74FD00FD7F" box="[656,767,672,696]" gridcol="3" gridrow="10" pageId="1" pageNumber="397">-87.881</td>
<td id="845CF261FFAF64ACFD52FD74FC7EFD7F" box="[804,897,672,696]" gridcol="4" gridrow="10" pageId="1" pageNumber="397">14.541</td>
<td id="845CF261FFAF64ACFDDCFD74FBCFFD7F" box="[938,1072,672,696]" gridcol="5" gridrow="10" pageId="1" pageNumber="397">1450</td>
<td id="845CF261FFAF64ACFA3CFD74FA75FD7F" box="[1098,1418,672,696]" gridcol="6" gridrow="10" pageId="1" pageNumber="397">
<bibRefCitation id="1D2CE4AEFFAF9B52FA3CFD74FA97FD7F" author="Pinacho-Pinacho, C. D. &amp; Garcia-Varela, M. &amp; Mendoza-Palmero, C. &amp; Sereno-Uribe, A. L. &amp; Martinez-Ramirez, E. &amp; Andrade-Gomez, L. &amp; Hernandez-Cruz, E. &amp; Perez Ponce de Leon, G." box="[1098,1384,672,696]" pageId="1" pageNumber="397" pagination="1 - 30" refId="ref3466" refString="Pinacho-Pinacho, C. D., Garcia-Varela, M., Hernandez-Orts, J. S., Mendoza-Palmero, C. a., Sereno-Uribe, A. L., Martinez-Ramirez, E., Andrade-Gomez, L., Hernandez-Cruz, E. &amp; Perez Ponce de Leon, G. (2015) Checklist of the helminth parasites of the genus Profundulus Hubbs, 1924 (Cyprinodontiformes, Profundulidae), an endemic family of freshwater fishes in Middle-America. ZooKeys, 523, 1 - 30. https: // doi. org / 10.3897 / zookeys. 523.6088" type="journal article" year="2015">
Pinacho-Pinacho 
<emphasis id="4BC9454DFFAF9B52FA8AFD75FAD6FD7F" box="[1276,1321,672,696]" italics="true" pageId="1" pageNumber="397">et al</emphasis>
. 2015
</bibRefCitation>
</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FD1CFA75FD26" box="[165,1418,712,737]" gridrow="11" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FD1CFF3EFD26" box="[165,193,712,737]" gridcol="0" gridrow="11" pageId="1" pageNumber="397">11</th>
<td id="845CF261FFAF64ACFEACFD1CFE97FD26" box="[218,360,712,737]" gridcol="1" gridrow="11" pageId="1" pageNumber="397">Ojojona</td>
<td id="845CF261FFAF64ACFFFFFD1CFD92FD26" box="[393,621,712,737]" gridcol="2" gridrow="11" pageId="1" pageNumber="397">Nacaome-Verdugo</td>
<td id="845CF261FFAF64ACFCE6FD1CFD00FD26" box="[656,767,712,737]" gridcol="3" gridrow="11" pageId="1" pageNumber="397">-87.318</td>
<td id="845CF261FFAF64ACFD52FD1CFC7EFD26" box="[804,897,712,737]" gridcol="4" gridrow="11" pageId="1" pageNumber="397">13.930</td>
<td id="845CF261FFAF64ACFDDCFD1CFBCFFD26" box="[938,1072,712,737]" gridcol="5" gridrow="11" pageId="1" pageNumber="397">1575</td>
<td id="845CF261FFAF64ACFA3CFD1CFA75FD26" box="[1098,1418,712,737]" gridcol="6" gridrow="11" pageId="1" pageNumber="397">LSUMZ16071</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FD25FA75FCCE" box="[165,1418,753,777]" gridrow="12" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FD25FF3EFCCE" box="[165,193,753,777]" gridcol="0" gridrow="12" pageId="1" pageNumber="397">12</th>
<td id="845CF261FFAF64ACFEACFD25FE97FCCE" box="[218,360,753,777]" gridcol="1" gridrow="12" pageId="1" pageNumber="397">San Sebastian</td>
<td id="845CF261FFAF64ACFFFFFD25FD92FCCE" box="[393,621,753,777]" gridcol="2" gridrow="12" pageId="1" pageNumber="397">Nacaome-Verdugo</td>
<td id="845CF261FFAF64ACFCE6FD25FD00FCCE" box="[656,767,753,777]" gridcol="3" gridrow="12" pageId="1" pageNumber="397">-87.360</td>
<td id="845CF261FFAF64ACFD52FD25FC7EFCCE" box="[804,897,753,777]" gridcol="4" gridrow="12" pageId="1" pageNumber="397">13.962</td>
<td id="845CF261FFAF64ACFDDCFD25FBCFFCCE" box="[938,1072,753,777]" gridcol="5" gridrow="12" pageId="1" pageNumber="397">1673</td>
<td id="845CF261FFAF64ACFA3CFD25FA75FCCE" box="[1098,1418,753,777]" gridcol="6" gridrow="12" pageId="1" pageNumber="397">LSUMZ16073</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FCCDFA75FCF6" box="[165,1418,793,817]" gridrow="13" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FCCDFF3EFCF6" box="[165,193,793,817]" gridcol="0" gridrow="13" pageId="1" pageNumber="397">13</th>
<td id="845CF261FFAF64ACFEACFCCDFE97FCF6" box="[218,360,793,817]" gridcol="1" gridrow="13" pageId="1" pageNumber="397">Ojojona</td>
<td id="845CF261FFAF64ACFFFFFCCDFD92FCF6" box="[393,621,793,817]" gridcol="2" gridrow="13" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FCCDFD00FCF6" box="[656,767,793,817]" gridcol="3" gridrow="13" pageId="1" pageNumber="397">-87.373</td>
<td id="845CF261FFAF64ACFD52FCCDFC7EFCF6" box="[804,897,793,817]" gridcol="4" gridrow="13" pageId="1" pageNumber="397">13.989</td>
<td id="845CF261FFAF64ACFDDCFCCDFBCFFCF6" box="[938,1072,793,817]" gridcol="5" gridrow="13" pageId="1" pageNumber="397">1415</td>
<td id="845CF261FFAF64ACFA3CFCCDFA75FCF6" box="[1098,1418,793,817]" gridcol="6" gridrow="13" pageId="1" pageNumber="397">LSUMZ16075</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FC95FA75FC9E" box="[165,1418,833,857]" gridrow="14" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FC95FF3EFC9E" box="[165,193,833,857]" gridcol="0" gridrow="14" pageId="1" pageNumber="397">14</th>
<td id="845CF261FFAF64ACFEACFC95FE97FC9E" box="[218,360,833,857]" gridcol="1" gridrow="14" pageId="1" pageNumber="397">Lepaterique</td>
<td id="845CF261FFAF64ACFFFFFC95FD92FC9E" box="[393,621,833,857]" gridcol="2" gridrow="14" pageId="1" pageNumber="397">Nacaome-Verdugo</td>
<td id="845CF261FFAF64ACFCE6FC95FD00FC9E" box="[656,767,833,857]" gridcol="3" gridrow="14" pageId="1" pageNumber="397">-87.466</td>
<td id="845CF261FFAF64ACFD52FC95FC7EFC9E" box="[804,897,833,857]" gridcol="4" gridrow="14" pageId="1" pageNumber="397">14.064</td>
<td id="845CF261FFAF64ACFDDCFC95FBCFFC9E" box="[938,1072,833,857]" gridcol="5" gridrow="14" pageId="1" pageNumber="397">1464</td>
<td id="845CF261FFAF64ACFA3CFC95FA75FC9E" box="[1098,1418,833,857]" gridcol="6" gridrow="14" pageId="1" pageNumber="397">USM31628</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FCBDFA75FC45" box="[165,1418,873,898]" gridrow="15" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FCBDFF3EFC45" box="[165,193,873,898]" gridcol="0" gridrow="15" pageId="1" pageNumber="397">15</th>
<td id="845CF261FFAF64ACFEACFCBDFE97FC45" box="[218,360,873,898]" gridcol="1" gridrow="15" pageId="1" pageNumber="397">Lepaterique</td>
<td id="845CF261FFAF64ACFFFFFCBDFD92FC45" box="[393,621,873,898]" gridcol="2" gridrow="15" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FCBDFD00FC45" box="[656,767,873,898]" gridcol="3" gridrow="15" pageId="1" pageNumber="397">-87.432</td>
<td id="845CF261FFAF64ACFD52FCBDFC7EFC45" box="[804,897,873,898]" gridcol="4" gridrow="15" pageId="1" pageNumber="397">14.071</td>
<td id="845CF261FFAF64ACFDDCFCBDFBCFFC45" box="[938,1072,873,898]" gridcol="5" gridrow="15" pageId="1" pageNumber="397">1675</td>
<td id="845CF261FFAF64ACFA3CFCBDFA75FC45" box="[1098,1418,873,898]" gridcol="6" gridrow="15" pageId="1" pageNumber="397">USM31631</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FC45FA75FC6D" box="[165,1418,913,938]" gridrow="16" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FC45FF3EFC6D" box="[165,193,913,938]" gridcol="0" gridrow="16" pageId="1" pageNumber="397">16</th>
<td id="845CF261FFAF64ACFEACFC45FE97FC6D" box="[218,360,913,938]" gridcol="1" gridrow="16" pageId="1" pageNumber="397">Siguatepeque</td>
<td id="845CF261FFAF64ACFFFFFC45FD92FC6D" box="[393,621,913,938]" gridcol="2" gridrow="16" pageId="1" pageNumber="397">Ulúa-Selguapa</td>
<td id="845CF261FFAF64ACFCE6FC45FD00FC6D" box="[656,767,913,938]" gridcol="3" gridrow="16" pageId="1" pageNumber="397">-87.870</td>
<td id="845CF261FFAF64ACFD52FC45FC7EFC6D" box="[804,897,913,938]" gridcol="4" gridrow="16" pageId="1" pageNumber="397">14.535</td>
<td id="845CF261FFAF64ACFDDCFC45FBCFFC6D" box="[938,1072,913,938]" gridcol="5" gridrow="16" pageId="1" pageNumber="397">1347</td>
<td id="845CF261FFAF64ACFA3CFC45FA75FC6D" box="[1098,1418,913,938]" gridcol="6" gridrow="16" pageId="1" pageNumber="397">USM45512</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FC6EFA75FC15" box="[165,1418,954,978]" gridrow="17" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FC6EFF3EFC15" box="[165,193,954,978]" gridcol="0" gridrow="17" pageId="1" pageNumber="397">17</th>
<td id="845CF261FFAF64ACFEACFC6EFE97FC15" box="[218,360,954,978]" gridcol="1" gridrow="17" pageId="1" pageNumber="397">Siguatepeque</td>
<td id="845CF261FFAF64ACFFFFFC6EFD92FC15" box="[393,621,954,978]" gridcol="2" gridrow="17" pageId="1" pageNumber="397">Ulúa-Calam</td>
<td id="845CF261FFAF64ACFCE6FC6EFD00FC15" box="[656,767,954,978]" gridcol="3" gridrow="17" pageId="1" pageNumber="397">-87.875</td>
<td id="845CF261FFAF64ACFD52FC6EFC7EFC15" box="[804,897,954,978]" gridcol="4" gridrow="17" pageId="1" pageNumber="397">14.550</td>
<td id="845CF261FFAF64ACFDDCFC6EFBCFFC15" box="[938,1072,954,978]" gridcol="5" gridrow="17" pageId="1" pageNumber="397">1337</td>
<td id="845CF261FFAF64ACFA3CFC6EFA75FC15" box="[1098,1418,954,978]" gridcol="6" gridrow="17" pageId="1" pageNumber="397">USM31652</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FC36FA75FC3D" box="[165,1418,994,1018]" gridrow="18" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FC36FF3EFC3D" box="[165,193,994,1018]" gridcol="0" gridrow="18" pageId="1" pageNumber="397">18</th>
<td id="845CF261FFAF64ACFEACFC36FE97FC3D" box="[218,360,994,1018]" gridcol="1" gridrow="18" pageId="1" pageNumber="397">Potrerillos</td>
<td id="845CF261FFAF64ACFFFFFC36FD92FC3D" box="[393,621,994,1018]" gridcol="2" gridrow="18" pageId="1" pageNumber="397">Ulúa-Canoas</td>
<td id="845CF261FFAF64ACFCE6FC36FD00FC3D" box="[656,767,994,1018]" gridcol="3" gridrow="18" pageId="1" pageNumber="397">-87.878</td>
<td id="845CF261FFAF64ACFD52FC36FC7EFC3D" box="[804,897,994,1018]" gridcol="4" gridrow="18" pageId="1" pageNumber="397">14.554</td>
<td id="845CF261FFAF64ACFDDCFC36FBCFFC3D" box="[938,1072,994,1018]" gridcol="5" gridrow="18" pageId="1" pageNumber="397">1332</td>
<td id="845CF261FFAF64ACFA3CFC36FA75FC3D" box="[1098,1418,994,1018]" gridcol="6" gridrow="18" pageId="1" pageNumber="397">USM31590</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FBDEFA75FBE5" box="[165,1418,1034,1058]" gridrow="19" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FBDEFF3EFBE5" box="[165,193,1034,1058]" gridcol="0" gridrow="19" pageId="1" pageNumber="397">19</th>
<td id="845CF261FFAF64ACFEACFBDEFE97FBE5" box="[218,360,1034,1058]" gridcol="1" gridrow="19" pageId="1" pageNumber="397">Ojojona</td>
<td id="845CF261FFAF64ACFFFFFBDEFD92FBE5" box="[393,621,1034,1058]" gridcol="2" gridrow="19" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FBDEFD00FBE5" box="[656,767,1034,1058]" gridcol="3" gridrow="19" pageId="1" pageNumber="397">-87.287</td>
<td id="845CF261FFAF64ACFD52FBDEFC7EFBE5" box="[804,897,1034,1058]" gridcol="4" gridrow="19" pageId="1" pageNumber="397">13.954</td>
<td id="845CF261FFAF64ACFDDCFBDEFBCFFBE5" box="[938,1072,1034,1058]" gridcol="5" gridrow="19" pageId="1" pageNumber="397">1286</td>
<td id="845CF261FFAF64ACFA3CFBDEFA75FBE5" box="[1098,1418,1034,1058]" gridcol="6" gridrow="19" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FBE7FA75FB8C" box="[165,1418,1075,1099]" gridrow="20" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FBE7FF3EFB8C" box="[165,193,1075,1099]" gridcol="0" gridrow="20" pageId="1" pageNumber="397">20</th>
<td id="845CF261FFAF64ACFEACFBE7FE97FB8C" box="[218,360,1075,1099]" gridcol="1" gridrow="20" pageId="1" pageNumber="397">El Aguacatal</td>
<td id="845CF261FFAF64ACFFFFFBE7FD92FB8C" box="[393,621,1075,1099]" gridcol="2" gridrow="20" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FBE7FD00FB8C" box="[656,767,1075,1099]" gridcol="3" gridrow="20" pageId="1" pageNumber="397">-87.362</td>
<td id="845CF261FFAF64ACFD52FBE7FC7EFB8C" box="[804,897,1075,1099]" gridcol="4" gridrow="20" pageId="1" pageNumber="397">14.012</td>
<td id="845CF261FFAF64ACFDDCFBE7FBCFFB8C" box="[938,1072,1075,1099]" gridcol="5" gridrow="20" pageId="1" pageNumber="397">1409</td>
<td id="845CF261FFAF64ACFA3CFBE7FA75FB8C" box="[1098,1418,1075,1099]" gridcol="6" gridrow="20" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FB8FFA75FBB4" box="[165,1418,1115,1139]" gridrow="21" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FB8FFF3EFBB4" box="[165,193,1115,1139]" gridcol="0" gridrow="21" pageId="1" pageNumber="397">21</th>
<td id="845CF261FFAF64ACFEACFB8FFE97FBB4" box="[218,360,1115,1139]" gridcol="1" gridrow="21" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFFB8FFD92FBB4" box="[393,621,1115,1139]" gridcol="2" gridrow="21" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FB8FFD00FBB4" box="[656,767,1115,1139]" gridcol="3" gridrow="21" pageId="1" pageNumber="397">-87.101</td>
<td id="845CF261FFAF64ACFD52FB8FFC7EFBB4" box="[804,897,1115,1139]" gridcol="4" gridrow="21" pageId="1" pageNumber="397">14.013</td>
<td id="845CF261FFAF64ACFDDCFB8FFBCFFBB4" box="[938,1072,1115,1139]" gridcol="5" gridrow="21" pageId="1" pageNumber="397">1309</td>
<td id="845CF261FFAF64ACFA3CFB8FFA75FBB4" box="[1098,1418,1115,1139]" gridcol="6" gridrow="21" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FB57FA75FB5C" box="[165,1418,1155,1179]" gridrow="22" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FB57FF3EFB5C" box="[165,193,1155,1179]" gridcol="0" gridrow="22" pageId="1" pageNumber="397">22</th>
<td id="845CF261FFAF64ACFEACFB57FE97FB5C" box="[218,360,1155,1179]" gridcol="1" gridrow="22" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFFB57FD92FB5C" box="[393,621,1155,1179]" gridcol="2" gridrow="22" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FB57FD00FB5C" box="[656,767,1155,1179]" gridcol="3" gridrow="22" pageId="1" pageNumber="397">-87.096</td>
<td id="845CF261FFAF64ACFD52FB57FC7EFB5C" box="[804,897,1155,1179]" gridcol="4" gridrow="22" pageId="1" pageNumber="397">14.015</td>
<td id="845CF261FFAF64ACFDDCFB57FBCFFB5C" box="[938,1072,1155,1179]" gridcol="5" gridrow="22" pageId="1" pageNumber="397">1295</td>
<td id="845CF261FFAF64ACFA3CFB57FA75FB5C" box="[1098,1418,1155,1179]" gridcol="6" gridrow="22" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FB7FFA75FB04" box="[165,1418,1195,1219]" gridrow="23" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FB7FFF3EFB04" box="[165,193,1195,1219]" gridcol="0" gridrow="23" pageId="1" pageNumber="397">23</th>
<td id="845CF261FFAF64ACFEACFB7FFE97FB04" box="[218,360,1195,1219]" gridcol="1" gridrow="23" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFFB7FFD92FB04" box="[393,621,1195,1219]" gridcol="2" gridrow="23" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FB7FFD00FB04" box="[656,767,1195,1219]" gridcol="3" gridrow="23" pageId="1" pageNumber="397">-87.098</td>
<td id="845CF261FFAF64ACFD52FB7FFC7EFB04" box="[804,897,1195,1219]" gridcol="4" gridrow="23" pageId="1" pageNumber="397">14.016</td>
<td id="845CF261FFAF64ACFDDCFB7FFBCFFB04" box="[938,1072,1195,1219]" gridcol="5" gridrow="23" pageId="1" pageNumber="397">1314</td>
<td id="845CF261FFAF64ACFA3CFB7FFA75FB04" box="[1098,1418,1195,1219]" gridcol="6" gridrow="23" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FB07FA75FB2B" box="[165,1418,1235,1260]" gridrow="24" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FB07FF3EFB2B" box="[165,193,1235,1260]" gridcol="0" gridrow="24" pageId="1" pageNumber="397">24</th>
<td id="845CF261FFAF64ACFEACFB07FE97FB2B" box="[218,360,1235,1260]" gridcol="1" gridrow="24" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFFB07FD92FB2B" box="[393,621,1235,1260]" gridcol="2" gridrow="24" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FB07FD00FB2B" box="[656,767,1235,1260]" gridcol="3" gridrow="24" pageId="1" pageNumber="397">-87.097</td>
<td id="845CF261FFAF64ACFD52FB07FC7EFB2B" box="[804,897,1235,1260]" gridcol="4" gridrow="24" pageId="1" pageNumber="397">14.016</td>
<td id="845CF261FFAF64ACFDDCFB07FBCFFB2B" box="[938,1072,1235,1260]" gridcol="5" gridrow="24" pageId="1" pageNumber="397">1319</td>
<td id="845CF261FFAF64ACFA3CFB07FA75FB2B" box="[1098,1418,1235,1260]" gridcol="6" gridrow="24" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FB28FA75FAD3" box="[165,1418,1276,1300]" gridrow="25" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FB28FF3EFAD3" box="[165,193,1276,1300]" gridcol="0" gridrow="25" pageId="1" pageNumber="397">25</th>
<td id="845CF261FFAF64ACFEACFB28FE97FAD3" box="[218,360,1276,1300]" gridcol="1" gridrow="25" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFFB28FD92FAD3" box="[393,621,1276,1300]" gridcol="2" gridrow="25" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FB28FD00FAD3" box="[656,767,1276,1300]" gridcol="3" gridrow="25" pageId="1" pageNumber="397">-87.096</td>
<td id="845CF261FFAF64ACFD52FB28FC7EFAD3" box="[804,897,1276,1300]" gridcol="4" gridrow="25" pageId="1" pageNumber="397">14.017</td>
<td id="845CF261FFAF64ACFDDCFB28FBCFFAD3" box="[938,1072,1276,1300]" gridcol="5" gridrow="25" pageId="1" pageNumber="397">1301</td>
<td id="845CF261FFAF64ACFA3CFB28FA75FAD3" box="[1098,1418,1276,1300]" gridcol="6" gridrow="25" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FAF0FA75FAFB" box="[165,1418,1316,1340]" gridrow="26" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FAF0FF3EFAFB" box="[165,193,1316,1340]" gridcol="0" gridrow="26" pageId="1" pageNumber="397">26</th>
<td id="845CF261FFAF64ACFEACFAF0FE97FAFB" box="[218,360,1316,1340]" gridcol="1" gridrow="26" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFFAF0FD92FAFB" box="[393,621,1316,1340]" gridcol="2" gridrow="26" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FAF0FD00FAFB" box="[656,767,1316,1340]" gridcol="3" gridrow="26" pageId="1" pageNumber="397">-87.111</td>
<td id="845CF261FFAF64ACFD52FAF0FC7EFAFB" box="[804,897,1316,1340]" gridcol="4" gridrow="26" pageId="1" pageNumber="397">14.032</td>
<td id="845CF261FFAF64ACFDDCFAF0FBCFFAFB" box="[938,1072,1316,1340]" gridcol="5" gridrow="26" pageId="1" pageNumber="397">1259</td>
<td id="845CF261FFAF64ACFA3CFAF0FA75FAFB" box="[1098,1418,1316,1340]" gridcol="6" gridrow="26" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FA98FA75FAA3" box="[165,1418,1356,1380]" gridrow="27" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FA98FF3EFAA3" box="[165,193,1356,1380]" gridcol="0" gridrow="27" pageId="1" pageNumber="397">27</th>
<td id="845CF261FFAF64ACFEACFA98FE97FAA3" box="[218,360,1356,1380]" gridcol="1" gridrow="27" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFFA98FD92FAA3" box="[393,621,1356,1380]" gridcol="2" gridrow="27" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FA98FD00FAA3" box="[656,767,1356,1380]" gridcol="3" gridrow="27" pageId="1" pageNumber="397">-87.110</td>
<td id="845CF261FFAF64ACFD52FA98FC7EFAA3" box="[804,897,1356,1380]" gridcol="4" gridrow="27" pageId="1" pageNumber="397">14.033</td>
<td id="845CF261FFAF64ACFDDCFA98FBCFFAA3" box="[938,1072,1356,1380]" gridcol="5" gridrow="27" pageId="1" pageNumber="397">1314</td>
<td id="845CF261FFAF64ACFA3CFA98FA75FAA3" box="[1098,1418,1356,1380]" gridcol="6" gridrow="27" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FAA0FA75FA4A" box="[165,1418,1396,1421]" gridrow="28" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FAA0FF3EFA4A" box="[165,193,1396,1421]" gridcol="0" gridrow="28" pageId="1" pageNumber="397">28</th>
<td id="845CF261FFAF64ACFEACFAA0FE97FA4A" box="[218,360,1396,1421]" gridcol="1" gridrow="28" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFFAA0FD92FA4A" box="[393,621,1396,1421]" gridcol="2" gridrow="28" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FAA0FD00FA4A" box="[656,767,1396,1421]" gridcol="3" gridrow="28" pageId="1" pageNumber="397">-87.109</td>
<td id="845CF261FFAF64ACFD52FAA0FC7EFA4A" box="[804,897,1396,1421]" gridcol="4" gridrow="28" pageId="1" pageNumber="397">14.033</td>
<td id="845CF261FFAF64ACFDDCFAA0FBCFFA4A" box="[938,1072,1396,1421]" gridcol="5" gridrow="28" pageId="1" pageNumber="397">1314</td>
<td id="845CF261FFAF64ACFA3CFAA0FA75FA4A" box="[1098,1418,1396,1421]" gridcol="6" gridrow="28" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FA49FA75FA72" box="[165,1418,1437,1461]" gridrow="29" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FA49FF3EFA72" box="[165,193,1437,1461]" gridcol="0" gridrow="29" pageId="1" pageNumber="397">29</th>
<td id="845CF261FFAF64ACFEACFA49FE97FA72" box="[218,360,1437,1461]" gridcol="1" gridrow="29" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFFA49FD92FA72" box="[393,621,1437,1461]" gridcol="2" gridrow="29" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FA49FD00FA72" box="[656,767,1437,1461]" gridcol="3" gridrow="29" pageId="1" pageNumber="397">-87.108</td>
<td id="845CF261FFAF64ACFD52FA49FC7EFA72" box="[804,897,1437,1461]" gridcol="4" gridrow="29" pageId="1" pageNumber="397">14.034</td>
<td id="845CF261FFAF64ACFDDCFA49FBCFFA72" box="[938,1072,1437,1461]" gridcol="5" gridrow="29" pageId="1" pageNumber="397">1265</td>
<td id="845CF261FFAF64ACFA3CFA49FA75FA72" box="[1098,1418,1437,1461]" gridcol="6" gridrow="29" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FA11FA75FA1A" box="[165,1418,1477,1501]" gridrow="30" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FA11FF3EFA1A" box="[165,193,1477,1501]" gridcol="0" gridrow="30" pageId="1" pageNumber="397">30</th>
<td id="845CF261FFAF64ACFEACFA11FE97FA1A" box="[218,360,1477,1501]" gridcol="1" gridrow="30" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFFA11FD92FA1A" box="[393,621,1477,1501]" gridcol="2" gridrow="30" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FA11FD00FA1A" box="[656,767,1477,1501]" gridcol="3" gridrow="30" pageId="1" pageNumber="397">-87.107</td>
<td id="845CF261FFAF64ACFD52FA11FC7EFA1A" box="[804,897,1477,1501]" gridcol="4" gridrow="30" pageId="1" pageNumber="397">14.034</td>
<td id="845CF261FFAF64ACFDDCFA11FBCFFA1A" box="[938,1072,1477,1501]" gridcol="5" gridrow="30" pageId="1" pageNumber="397">1274</td>
<td id="845CF261FFAF64ACFA3CFA11FA75FA1A" box="[1098,1418,1477,1501]" gridcol="6" gridrow="30" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3FA39FA75F9C2" box="[165,1418,1517,1541]" gridrow="31" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3FA39FF3EF9C2" box="[165,193,1517,1541]" gridcol="0" gridrow="31" pageId="1" pageNumber="397">31</th>
<td id="845CF261FFAF64ACFEACFA39FE97F9C2" box="[218,360,1517,1541]" gridcol="1" gridrow="31" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFFA39FD92F9C2" box="[393,621,1517,1541]" gridcol="2" gridrow="31" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6FA39FD00F9C2" box="[656,767,1517,1541]" gridcol="3" gridrow="31" pageId="1" pageNumber="397">-87.106</td>
<td id="845CF261FFAF64ACFD52FA39FC7EF9C2" box="[804,897,1517,1541]" gridcol="4" gridrow="31" pageId="1" pageNumber="397">14.035</td>
<td id="845CF261FFAF64ACFDDCFA39FBCFF9C2" box="[938,1072,1517,1541]" gridcol="5" gridrow="31" pageId="1" pageNumber="397">1277</td>
<td id="845CF261FFAF64ACFA3CFA39FA75F9C2" box="[1098,1418,1517,1541]" gridcol="6" gridrow="31" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3F9C1FA75F9E9" box="[165,1418,1557,1582]" gridrow="32" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3F9C1FF3EF9E9" box="[165,193,1557,1582]" gridcol="0" gridrow="32" pageId="1" pageNumber="397">32</th>
<td id="845CF261FFAF64ACFEACF9C1FE97F9E9" box="[218,360,1557,1582]" gridcol="1" gridrow="32" pageId="1" pageNumber="397">Tatumbla</td>
<td id="845CF261FFAF64ACFFFFF9C1FD92F9E9" box="[393,621,1557,1582]" gridcol="2" gridrow="32" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6F9C1FD00F9E9" box="[656,767,1557,1582]" gridcol="3" gridrow="32" pageId="1" pageNumber="397">-87.107</td>
<td id="845CF261FFAF64ACFD52F9C1FC7EF9E9" box="[804,897,1557,1582]" gridcol="4" gridrow="32" pageId="1" pageNumber="397">14.036</td>
<td id="845CF261FFAF64ACFDDCF9C1FBCFF9E9" box="[938,1072,1557,1582]" gridcol="5" gridrow="32" pageId="1" pageNumber="397">1276</td>
<td id="845CF261FFAF64ACFA3CF9C1FA75F9E9" box="[1098,1418,1557,1582]" gridcol="6" gridrow="32" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3F9EAFA75F991" box="[165,1418,1598,1622]" gridrow="33" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3F9EAFF3EF991" box="[165,193,1598,1622]" gridcol="0" gridrow="33" pageId="1" pageNumber="397">33</th>
<td id="845CF261FFAF64ACFEACF9EAFE97F991" box="[218,360,1598,1622]" gridcol="1" gridrow="33" pageId="1" pageNumber="397">Lepaterique</td>
<td id="845CF261FFAF64ACFFFFF9EAFD92F991" box="[393,621,1598,1622]" gridcol="2" gridrow="33" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6F9EAFD00F991" box="[656,767,1598,1622]" gridcol="3" gridrow="33" pageId="1" pageNumber="397">-87.412</td>
<td id="845CF261FFAF64ACFD52F9EAFC7EF991" box="[804,897,1598,1622]" gridcol="4" gridrow="33" pageId="1" pageNumber="397">14.054</td>
<td id="845CF261FFAF64ACFDDCF9EAFBCFF991" box="[938,1072,1598,1622]" gridcol="5" gridrow="33" pageId="1" pageNumber="397">1516</td>
<td id="845CF261FFAF64ACFA3CF9EAFA75F991" box="[1098,1418,1598,1622]" gridcol="6" gridrow="33" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3F9B2FA75F9B9" box="[165,1418,1638,1662]" gridrow="34" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3F9B2FF3EF9B9" box="[165,193,1638,1662]" gridcol="0" gridrow="34" pageId="1" pageNumber="397">34</th>
<td id="845CF261FFAF64ACFEACF9B2FE97F9B9" box="[218,360,1638,1662]" gridcol="1" gridrow="34" pageId="1" pageNumber="397">Lepaterique</td>
<td id="845CF261FFAF64ACFFFFF9B2FD92F9B9" box="[393,621,1638,1662]" gridcol="2" gridrow="34" pageId="1" pageNumber="397">Nacaome-Verdugo</td>
<td id="845CF261FFAF64ACFCE6F9B2FD00F9B9" box="[656,767,1638,1662]" gridcol="3" gridrow="34" pageId="1" pageNumber="397">-87.462</td>
<td id="845CF261FFAF64ACFD52F9B2FC7EF9B9" box="[804,897,1638,1662]" gridcol="4" gridrow="34" pageId="1" pageNumber="397">14.065</td>
<td id="845CF261FFAF64ACFDDCF9B2FBCFF9B9" box="[938,1072,1638,1662]" gridcol="5" gridrow="34" pageId="1" pageNumber="397">1487</td>
<td id="845CF261FFAF64ACFA3CF9B2FA75F9B9" box="[1098,1418,1638,1662]" gridcol="6" gridrow="34" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3F95AFA75F961" box="[165,1418,1678,1702]" gridrow="35" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3F95AFF3EF961" box="[165,193,1678,1702]" gridcol="0" gridrow="35" pageId="1" pageNumber="397">35</th>
<td id="845CF261FFAF64ACFEACF95AFE97F961" box="[218,360,1678,1702]" gridcol="1" gridrow="35" pageId="1" pageNumber="397">Tegucigalpa</td>
<td id="845CF261FFAF64ACFFFFF95AFD92F961" box="[393,621,1678,1702]" gridcol="2" gridrow="35" pageId="1" pageNumber="397">Choluteca-Guacerique</td>
<td id="845CF261FFAF64ACFCE6F95AFD00F961" box="[656,767,1678,1702]" gridcol="3" gridrow="35" pageId="1" pageNumber="397">-87.205</td>
<td id="845CF261FFAF64ACFD52F95AFC7EF961" box="[804,897,1678,1702]" gridcol="4" gridrow="35" pageId="1" pageNumber="397">14.084</td>
<td id="845CF261FFAF64ACFDDCF95AFBCFF961" box="[938,1072,1678,1702]" gridcol="5" gridrow="35" pageId="1" pageNumber="397">942</td>
<td id="845CF261FFAF64ACFA3CF95AFA75F961" box="[1098,1418,1678,1702]" gridcol="6" gridrow="35" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3F962FA75F908" box="[165,1418,1718,1743]" gridrow="36" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3F962FF3EF908" box="[165,193,1718,1743]" gridcol="0" gridrow="36" pageId="1" pageNumber="397">36</th>
<td id="845CF261FFAF64ACFEACF962FE97F908" box="[218,360,1718,1743]" gridcol="1" gridrow="36" pageId="1" pageNumber="397">Tamara</td>
<td id="845CF261FFAF64ACFFFFF962FD92F908" box="[393,621,1718,1743]" gridcol="2" gridrow="36" pageId="1" pageNumber="397">Choluteca-El Hombre</td>
<td id="845CF261FFAF64ACFCE6F962FD00F908" box="[656,767,1718,1743]" gridcol="3" gridrow="36" pageId="1" pageNumber="397">-87.293</td>
<td id="845CF261FFAF64ACFD52F962FC7EF908" box="[804,897,1718,1743]" gridcol="4" gridrow="36" pageId="1" pageNumber="397">14.175</td>
<td id="845CF261FFAF64ACFDDCF962FBCFF908" box="[938,1072,1718,1743]" gridcol="5" gridrow="36" pageId="1" pageNumber="397">1317</td>
<td id="845CF261FFAF64ACFA3CF962FA75F908" box="[1098,1418,1718,1743]" gridcol="6" gridrow="36" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3F90BFA75F930" box="[165,1418,1759,1783]" gridrow="37" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3F90BFF3EF930" box="[165,193,1759,1783]" gridcol="0" gridrow="37" pageId="1" pageNumber="397">37</th>
<td id="845CF261FFAF64ACFEACF90BFE97F930" box="[218,360,1759,1783]" gridcol="1" gridrow="37" pageId="1" pageNumber="397">Comayagua</td>
<td id="845CF261FFAF64ACFFFFF90BFD92F930" box="[393,621,1759,1783]" gridcol="2" gridrow="37" pageId="1" pageNumber="397">Ulúa-Humuya Alto</td>
<td id="845CF261FFAF64ACFCE6F90BFD00F930" box="[656,767,1759,1783]" gridcol="3" gridrow="37" pageId="1" pageNumber="397">-87.654</td>
<td id="845CF261FFAF64ACFD52F90BFC7EF930" box="[804,897,1759,1783]" gridcol="4" gridrow="37" pageId="1" pageNumber="397">14.445</td>
<td id="845CF261FFAF64ACFDDCF90BFBCFF930" box="[938,1072,1759,1783]" gridcol="5" gridrow="37" pageId="1" pageNumber="397">576</td>
<td id="845CF261FFAF64ACFA3CF90BFA75F930" box="[1098,1418,1759,1783]" gridcol="6" gridrow="37" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3F8D3FA75F8D8" box="[165,1418,1799,1823]" gridrow="38" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3F8D3FF3EF8D8" box="[165,193,1799,1823]" gridcol="0" gridrow="38" pageId="1" pageNumber="397">38</th>
<td id="845CF261FFAF64ACFEACF8D3FE97F8D8" box="[218,360,1799,1823]" gridcol="1" gridrow="38" pageId="1" pageNumber="397">Comayagua</td>
<td id="845CF261FFAF64ACFFFFF8D3FD92F8D8" box="[393,621,1799,1823]" gridcol="2" gridrow="38" pageId="1" pageNumber="397">Ulúa-Humuya Alto</td>
<td id="845CF261FFAF64ACFCE6F8D3FD00F8D8" box="[656,767,1799,1823]" gridcol="3" gridrow="38" pageId="1" pageNumber="397">-87.654</td>
<td id="845CF261FFAF64ACFD52F8D3FC7EF8D8" box="[804,897,1799,1823]" gridcol="4" gridrow="38" pageId="1" pageNumber="397">14.467</td>
<td id="845CF261FFAF64ACFDDCF8D3FBCFF8D8" box="[938,1072,1799,1823]" gridcol="5" gridrow="38" pageId="1" pageNumber="397">573</td>
<td id="845CF261FFAF64ACFA3CF8D3FA75F8D8" box="[1098,1418,1799,1823]" gridcol="6" gridrow="38" pageId="1" pageNumber="397">This Publication</td>
</tr>
<tr id="C78D9B1DFFAF64ACFED3F8FBFA75F8A3" box="[165,1418,1839,1892]" gridrow="39" pageId="1" pageNumber="397">
<th id="845CF261FFAF64ACFED3F8FBFF3EF8A3" box="[165,193,1839,1892]" gridcol="0" gridrow="39" pageId="1" pageNumber="397">39</th>
<td id="845CF261FFAF64ACFEACF8FBFE97F8A3" box="[218,360,1839,1892]" gridcol="1" gridrow="39" pageId="1" pageNumber="397">San Francisco de Orica</td>
<td id="845CF261FFAF64ACFFFFF8FBFD92F8A3" box="[393,621,1839,1892]" gridcol="2" gridrow="39" pageId="1" pageNumber="397">Patuca-Guayape</td>
<td id="845CF261FFAF64ACFCE6F8FBFD00F8A3" box="[656,767,1839,1892]" gridcol="3" gridrow="39" pageId="1" pageNumber="397">-86.977</td>
<td id="845CF261FFAF64ACFD52F8FBFC7EF8A3" box="[804,897,1839,1892]" gridcol="4" gridrow="39" pageId="1" pageNumber="397">14.908</td>
<td id="845CF261FFAF64ACFDDCF8FBFBCFF8A3" box="[938,1072,1839,1892]" gridcol="5" gridrow="39" pageId="1" pageNumber="397">1570</td>
<td id="845CF261FFAF64ACFA3CF8FBFA75F8A3" box="[1098,1418,1839,1892]" gridcol="6" gridrow="39" pageId="1" pageNumber="397">This Publication</td>
</tr>
</table>
</paragraph>
<caption id="2DC2C9D7FFAC9B51FEEEFD80FD7EFD3B" ID-DOI="http://doi.org/10.5281/zenodo.10561935" ID-Zenodo-Dep="10561935" httpUri="https://zenodo.org/record/10561935/files/figure.png" pageId="2" pageNumber="398" startId="2.[152,255,596,620]" targetBox="[349,1239,180,571]" targetPageId="2" targetType="figure">
<paragraph id="7902995FFFAC9B51FEEEFD80FD7EFD3B" blockId="2.[151,1436,596,764]" pageId="2" pageNumber="398">
<emphasis id="4BC9454DFFAC9B51FEEEFD80FEE9FDAB" bold="true" box="[152,278,596,620]" pageId="2" pageNumber="398">FIGURE 1.</emphasis>
(A) Map of Honduras showing localities where 
<taxonomicName id="BEBDE2DCFFAC9B51FC9BFD80FC59FDAB" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[749,934,596,620]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FC9BFD80FC59FDAB" box="[749,934,596,620]" italics="true" pageId="2" pageNumber="398">Tlaloc portillorum</emphasis>
</taxonomicName>
has been collected.Dashed lines depict major river basin borders (basin names in blue font with white background). Major city names in uppercase and bold font. (B) Photograph of live specimen of 
<taxonomicName id="BEBDE2DCFFAC9B51FF14FD49FE0FFD73" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[354,496,668,692]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FF14FD49FE0FFD73" box="[354,496,668,692]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
(5.5 cm standard length) from the Guacerique River (longitude= -87.109, latitude= 14.033), a tributary of the Choluteca River basin. (C) Habitat of 
<taxonomicName id="BEBDE2DCFFAC9B51FCA5FD15FC9DFD1F" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[723,866,704,728]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FCA5FD15FC9DFD1F" box="[723,866,704,728]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
. From the Guacerique River, tributary of the Choluteca River basin (longitude -87.111, latitude 14.032).
</paragraph>
</caption>
<subSubSection id="31A7CAD4FFAC9B51FEB1FC9CFE9EFBD5" pageId="2" pageNumber="398" type="conservation">
<paragraph id="7902995FFFAC9B51FEB1FC9CFE9EFBD5" blockId="2.[151,1437,840,2043]" pageId="2" pageNumber="398">
Regarding its conservation status, 
<taxonomicName id="BEBDE2DCFFAC9B51FC4CFC9DFD2EFCA7" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[570,721,840,864]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FC4CFC9DFD2EFCA7" box="[570,721,840,864]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
is listed as endangered in the IUCN red list (
<bibRefCitation id="1D2CE4AEFFAC9B51FACBFC9CFF2EFC43" author="Lyons, T. J. &amp; Matamoros, W. A." pageId="2" pageNumber="398" pagination="164692340" refId="ref2997" refString="Lyons, T. J. &amp; Matamoros, W. A. (2020) Tlaloc portillorum. The IUCN Red List of Threatened Species, 2020, e. T 125347429 A 164692340." type="journal article" year="2020">Lyons &amp; Matamoros 2020</bibRefCitation>
). The criteria for placing the species under this category is that at the time the IUCN report was made, there was a limited number of known localities for the species (
<bibRefCitation id="1D2CE4AEFFAC9B51FCAFFC44FC09FC60" author="Lyons, T. J. &amp; Matamoros, W. A." box="[729,1014,911,936]" pageId="2" pageNumber="398" pagination="164692340" refId="ref2997" refString="Lyons, T. J. &amp; Matamoros, W. A. (2020) Tlaloc portillorum. The IUCN Red List of Threatened Species, 2020, e. T 125347429 A 164692340." type="journal article" year="2020">Lyons &amp; Matamoros 2020</bibRefCitation>
) and some of these localities are under critical anthropogenic pressure (W.A. Matamoros pers. Obs.). It is paramount for conservation planning to know in the most comprehensible way the actual species distributional range, which will allow conservation managers to take more accurate decisions.
</paragraph>
</subSubSection>
<subSubSection id="31A7CAD4FFAC9B51FEB1FBCAFD99FBBA" pageId="2" pageNumber="398" type="discussion">
<paragraph id="7902995FFFAC9B51FEB1FBCAFD99FBBA" blockId="2.[151,1437,840,2043]" pageId="2" pageNumber="398">
The aims of this study are twofold: First, to build the most comprehensive distribution map of 
<taxonomicName id="BEBDE2DCFFAC9B51FAD6FBCBFACBFBF1" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[1184,1332,1054,1078]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FAD6FBCBFACBFBF1" box="[1184,1332,1054,1078]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
, based on data from the scientific literature, collection records and our own surveys. Second, to provide a more complete description of the habitats where 
<taxonomicName id="BEBDE2DCFFAC9B51FF0BFBB2FDEDFBB9" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[381,530,1126,1150]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FF0BFBB2FDEDFBB9" box="[381,530,1126,1150]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
occurs.
</paragraph>
</subSubSection>
<subSubSection id="31A7CAD4FFAC9B51FEB1FB5DFB81F9EC" pageId="2" pageNumber="398" type="materials_examined">
<paragraph id="7902995FFFAC9B51FEB1FB5DFB81F9EC" blockId="2.[151,1437,840,2043]" pageId="2" pageNumber="398">
<collectingCounty id="9063E1D3FFAC9B51FEB1FB5DFEC4FB66" box="[199,315,1161,1185]" pageId="2" pageNumber="398">Specimens</collectingCounty>
were collected as part of surveys performed by the first author in order to complete his undergraduate professional practice in 2014 (
<bibRefCitation id="1D2CE4AEFFAC9B51FFABFB79FD6FFB02" author="Henriquez, C. M." box="[477,656,1197,1221]" pageId="2" pageNumber="398" refId="ref2963" refString="Henriquez, C. M. (2014) Diversidad de peces en dos tributarios de la cuenca alta de rio Patuca. Universidad Nacional Autonoma de Honduras, Tegucigalpa, 28 pp." type="book" year="2014">Henríquez 2014</bibRefCitation>
). 
<collectingMunicipality id="99660325FFAC9B51FCD4FB79FCF7FB02" box="[674,776,1197,1221]" pageId="2" pageNumber="398">Sampling</collectingMunicipality>
was carried out in the upper reaches of the 
<location id="7C62CF84FFAC9B51FA95FB79FAA7FB02" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAC9B51FA95FB79FAA7FB02" box="[1251,1368,1197,1221]" county="Specimens" municipality="Sampling" name="Rio Patuca" pageId="2" pageNumber="398" stateProvince="Francisco Morazan">Río Patuca</location>
in the 
<location id="7C62CF84FFAC9B51FEE1FB05FEFDFB2E" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAC9B51FEE1FB05FEFDFB2E" box="[151,258,1233,1257]" county="Specimens" municipality="Sampling" name="Honduran" pageId="2" pageNumber="398" stateProvince="Francisco Morazan">Honduran</location>
department of Francisco Morazán. 
<location id="7C62CF84FFAC9B51FCF8FB05FCE9FB2E" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAC9B51FCF8FB05FCE9FB2E" box="[654,790,1233,1257]" county="Specimens" municipality="Sampling" name="Additionally" pageId="2" pageNumber="398" stateProvince="Francisco Morazan">Additionally</location>
, as part of a 
<taxonomicName id="BEBDE2DCFFAC9B51FDD8FB06FBB9FB2E" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[942,1094,1233,1257]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FDD8FB06FBB9FB2E" box="[942,1094,1233,1257]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
life history and diet study, we sampled specimens of 
<taxonomicName id="BEBDE2DCFFAC9B51FFF0FB21FDE5FACA" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[390,538,1269,1293]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FFF0FB21FDE5FACA" box="[390,538,1269,1293]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
between the months of 
<location id="7C62CF84FFAC9B51FD64FB21FCB5FACA" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAC9B51FD64FB21FCB5FACA" box="[786,842,1269,1293]" county="Specimens" municipality="Sampling" name="April" pageId="2" pageNumber="398" stateProvince="Francisco Morazan">April</location>
to 
<location id="7C62CF84FFAC9B51FD1FFB21FC6AFACA" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAC9B51FD1FFB21FC6AFACA" box="[873,917,1269,1293]" county="Specimens" municipality="Sampling" name="July" pageId="2" pageNumber="398" stateProvince="Francisco Morazan">July</location>
of 2023 in tributaries of the Choluteca, 
<location id="7C62CF84FFAC9B51FB41FB21FA63FACB" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAC9B51FB41FB21FA63FACB" box="[1335,1436,1269,1292]" county="Specimens" municipality="Sampling" name="Nacaome" pageId="2" pageNumber="398" stateProvince="Francisco Morazan">Nacaome</location>
and 
<location id="7C62CF84FFAC9B51FEB5FACCFF08FAF7" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAC9B51FEB5FACCFF08FAF7" box="[195,247,1304,1328]" county="Specimens" municipality="Sampling" name="Ulua" pageId="2" pageNumber="398" stateProvince="Francisco Morazan">Ulúa</location>
basins (
<figureCitation id="E18685DAFFAC9B51FF3BFACCFE5EFAF7" box="[333,417,1304,1328]" captionStart="FIGURE 1" captionStartId="2.[152,255,596,620]" captionTargetBox="[349,1239,180,571]" captionTargetId="figure-23@2.[349,1239,180,571]" captionTargetPageId="2" captionText="FIGURE 1. (A) Map of Honduras showing localities where Tlalocportillorum hasbeen collected.Dashed lines depict majorriver basin borders (basin names in blue font with white background). Major city names in uppercase and bold font. (B) Photograph of live specimen of T. portillorum (5.5 cm standard length) from the Guacerique River (longitude= -87.109, latitude= 14.033), a tributary of the Choluteca River basin. (C) Habitat of T. portillorum. From the Guacerique River, tributary of the Choluteca River basin (longitude -87.111, latitude 14.032)." figureDoi="http://doi.org/10.5281/zenodo.10561935" httpUri="https://zenodo.org/record/10561935/files/figure.png" pageId="2" pageNumber="398">Fig. 1A</figureCitation>
; 
<tableCitation id="343FACE4FFAC9B51FFDDFACCFE06FAF7" box="[427,505,1304,1328]" captionStart="TABLE 1" captionStartId="1.[152,245,151,175]" captionTargetPageId="1" captionText="TABLE 1. Table with geographic information, for each sample included in Figure one. Type series as in Matamoros &amp; Schaefer, 2010; The source Museum collections LSUMZ= Louisiana State University Museum Zoology, collection of fishes; USM= University of Southern Mississippi, collection of fishes." httpUri="http://table.plazi.org/id/2DC2C9D7FFAF9B52FEEEFF43FC78FF30" pageId="2" pageNumber="398" tableUuid="2DC2C9D7FFAF9B52FEEEFF43FC78FF30">Table 1</tableCitation>
). 
<location id="7C62CF84FFAC9B51FC78FACDFDCEFAF7" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAC9B51FC78FACDFDCEFAF7" box="[526,561,1305,1328]" county="Specimens" municipality="Sampling" name="We" pageId="2" pageNumber="398" stateProvince="Francisco Morazan">We</location>
also gathered distributional data from the scientific literature and a limited number of records were added from the 
<location id="7C62CF84FFAC9B51FC76FAE8FDAFFA93" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAC9B51FC76FAE8FDAFFA93" box="[512,592,1340,1364]" county="Specimens" municipality="Sampling" name="Fishnet" pageId="2" pageNumber="398" stateProvince="Francisco Morazan">Fishnet</location>
2 database aggregator (www.fishnet2.net, accessed 
<date id="0D03BF9FFFAC9B51FAF2FAE8FAFAFA93" box="[1156,1285,1340,1364]" pageId="2" pageNumber="398" value="2023-04-08">
<collectingDate id="1D474677FFAC9B51FAF2FAE8FAFAFA93" box="[1156,1285,1340,1364]" pageId="2" pageNumber="398" value="2023-04-08">2023-08-04</collectingDate>
</date>
). In the field, fish were collected with the use of dip-nets and a three m long and 1.80 m high seine with a 5 mm mesh size. In each locality, fish were sampled in all possible habitats including currents, riffles, and pools. Since we knew that 
<taxonomicName id="BEBDE2DCFFAC9B51FB7EFA50FA63FA5C" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[1288,1436,1411,1435]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FB7EFA50FA63FA5C" box="[1288,1436,1411,1435]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
specimens tend to hide underneath boulders, rocks and vegetation in the river undercut bank, we put special efforts into sampling this type of habitat. The sampling effort consisted of 30–40 minutes at each site. After capture, individuals were euthanized in a clove oil solution and then transferred to 10% formalin or to pure ethanol for tissue preservation. After 10–12 days, formalin-fixed specimens were transferred to 70% ethanol for long-term storage.
</paragraph>
</subSubSection>
<subSubSection id="31A7CAD4FFAC9B50FEB1F9E2FB62FF14" lastPageId="3" lastPageNumber="399" pageId="2" pageNumber="398" type="description">
<paragraph id="7902995FFFAC9B50FEB1F9E2FB62FF14" blockId="2.[151,1437,840,2043]" lastBlockId="3.[151,1437,151,823]" lastPageId="3" lastPageNumber="399" pageId="2" pageNumber="398">
Species identification was based on characters provided by 
<bibRefCitation id="1D2CE4AEFFAC9B51FD2AF9E3FB4FF989" author="Matamoros, W. A. &amp; Schaefer, J. F." box="[860,1200,1590,1614]" pageId="2" pageNumber="398" pagination="1498 - 507" refId="ref3234" refString="Matamoros, W. A. &amp; Schaefer, J. F. (2010) A new species of Profundulus (Cyprinodontiformes: Profundulidae) from the Honduran central highlands. Journal of Fish Biology, 76, 1498 - 507. https: // doi. org / 10.1111 / j. 1095 - 8649.2010.02596. x" type="journal article" year="2010">Matamoros &amp; Schaefer (2010)</bibRefCitation>
. The specimens were determined to belong to 
<taxonomicName id="BEBDE2DCFFAC9B51FFE0F98FFDD6F9B5" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[406,553,1626,1650]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FFE0F98FFDD6F9B5" box="[406,553,1626,1650]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
(
<figureCitation id="E18685DAFFAC9B51FC40F98EFD79F9B5" box="[566,646,1626,1650]" captionStart="FIGURE 1" captionStartId="2.[152,255,596,620]" captionTargetBox="[349,1239,180,571]" captionTargetId="figure-23@2.[349,1239,180,571]" captionTargetPageId="2" captionText="FIGURE 1. (A) Map of Honduras showing localities where Tlalocportillorum hasbeen collected.Dashed lines depict majorriver basin borders (basin names in blue font with white background). Major city names in uppercase and bold font. (B) Photograph of live specimen of T. portillorum (5.5 cm standard length) from the Guacerique River (longitude= -87.109, latitude= 14.033), a tributary of the Choluteca River basin. (C) Habitat of T. portillorum. From the Guacerique River, tributary of the Choluteca River basin (longitude -87.111, latitude 14.032)." figureDoi="http://doi.org/10.5281/zenodo.10561935" httpUri="https://zenodo.org/record/10561935/files/figure.png" pageId="2" pageNumber="398">Fig. 1B</figureCitation>
) due to the lack of a humeral spot and having less than half of the caudal fin covered by scales, with a prominent lower jaw that protrudes and covers the upper jaw. The new records of 
<taxonomicName id="BEBDE2DCFFAC9B51FB7EF9AAFA63F951" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[1288,1436,1662,1686]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FB7EF9AAFA63F951" box="[1288,1436,1662,1686]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
in tributaries of the Choluteca River and the Patuca River in conjunction represent a substantial range expansion of at least ~120 km (
<figureCitation id="E18685DAFFAC9B51FF4BF911FE6DF91A" box="[317,402,1733,1757]" captionStart="FIGURE 1" captionStartId="2.[152,255,596,620]" captionTargetBox="[349,1239,180,571]" captionTargetId="figure-23@2.[349,1239,180,571]" captionTargetPageId="2" captionText="FIGURE 1. (A) Map of Honduras showing localities where Tlalocportillorum hasbeen collected.Dashed lines depict majorriver basin borders (basin names in blue font with white background). Major city names in uppercase and bold font. (B) Photograph of live specimen of T. portillorum (5.5 cm standard length) from the Guacerique River (longitude= -87.109, latitude= 14.033), a tributary of the Choluteca River basin. (C) Habitat of T. portillorum. From the Guacerique River, tributary of the Choluteca River basin (longitude -87.111, latitude 14.032)." figureDoi="http://doi.org/10.5281/zenodo.10561935" httpUri="https://zenodo.org/record/10561935/files/figure.png" pageId="2" pageNumber="398">Fig. 1A</figureCitation>
). We recorded 
<taxonomicName id="BEBDE2DCFFAC9B51FC45F912FD36F91A" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[563,713,1733,1757]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FC45F912FD36F91A" box="[563,713,1733,1757]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
specimens at lower stream order creeks and never in the main river channels, at altitudes ranging from ~570 to ~1670 masl. The lower altitude localities were found in the Comayagua valley, close to the city of Comayagua (
<tableCitation id="343FACE4FFAC9B51FF8AF8D9FDB2F8E2" box="[508,589,1805,1829]" captionStart="TABLE 1" captionStartId="1.[152,245,151,175]" captionTargetPageId="1" captionText="TABLE 1. Table with geographic information, for each sample included in Figure one. Type series as in Matamoros &amp; Schaefer, 2010; The source Museum collections LSUMZ= Louisiana State University Museum Zoology, collection of fishes; USM= University of Southern Mississippi, collection of fishes." httpUri="http://table.plazi.org/id/2DC2C9D7FFAF9B52FEEEFF43FC78FF30" pageId="2" pageNumber="398" tableUuid="2DC2C9D7FFAF9B52FEEEFF43FC78FF30">Table 1</tableCitation>
). The highest localities are reported from the upper reaches of the Choluteca, Nacaome and Patuca Rivers (
<tableCitation id="343FACE4FFAC9B51FFA1F8E4FDD8F88F" box="[471,551,1840,1864]" captionStart="TABLE 1" captionStartId="1.[152,245,151,175]" captionTargetPageId="1" captionText="TABLE 1. Table with geographic information, for each sample included in Figure one. Type series as in Matamoros &amp; Schaefer, 2010; The source Museum collections LSUMZ= Louisiana State University Museum Zoology, collection of fishes; USM= University of Southern Mississippi, collection of fishes." httpUri="http://table.plazi.org/id/2DC2C9D7FFAF9B52FEEEFF43FC78FF30" pageId="2" pageNumber="398" tableUuid="2DC2C9D7FFAF9B52FEEEFF43FC78FF30">Table 1</tableCitation>
). These high-altitude lower stream order creeks where 
<taxonomicName id="BEBDE2DCFFAC9B51FA08F8E5FAEBF88F" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[1150,1300,1840,1864]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FA08F8E5FAEBF88F" box="[1150,1300,1840,1864]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
were found, are usually shallow (from a few centimeters of water depth to up to 50–70 cm of water depth). Creeks are generally very narrow, ranging from canals of less than one meter in width to more than 10 meters (
<figureCitation id="E18685DAFFAC9B51FA6BF8ACFB90F857" box="[1053,1135,1912,1936]" captionStart="FIGURE 1" captionStartId="2.[152,255,596,620]" captionTargetBox="[349,1239,180,571]" captionTargetId="figure-23@2.[349,1239,180,571]" captionTargetPageId="2" captionText="FIGURE 1. (A) Map of Honduras showing localities where Tlalocportillorum hasbeen collected.Dashed lines depict majorriver basin borders (basin names in blue font with white background). Major city names in uppercase and bold font. (B) Photograph of live specimen of T. portillorum (5.5 cm standard length) from the Guacerique River (longitude= -87.109, latitude= 14.033), a tributary of the Choluteca River basin. (C) Habitat of T. portillorum. From the Guacerique River, tributary of the Choluteca River basin (longitude -87.111, latitude 14.032)." figureDoi="http://doi.org/10.5281/zenodo.10561935" httpUri="https://zenodo.org/record/10561935/files/figure.png" pageId="2" pageNumber="398">Fig. 1C</figureCitation>
), with sediments dominated by large boulders, rocks and cobblestone. Frequently 
<taxonomicName id="BEBDE2DCFFAC9B51FCA1F848FC92F874" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[727,877,1947,1971]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FCA1F848FC92F874" box="[727,877,1947,1971]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
specimens are found underneath large boulders and rocks in both flowing current and pools (
<figureCitation id="E18685DAFFAC9B51FC22F86BFD58F810" box="[596,679,1983,2007]" captionStart="FIGURE 1" captionStartId="2.[152,255,596,620]" captionTargetBox="[349,1239,180,571]" captionTargetId="figure-23@2.[349,1239,180,571]" captionTargetPageId="2" captionText="FIGURE 1. (A) Map of Honduras showing localities where Tlalocportillorum hasbeen collected.Dashed lines depict majorriver basin borders (basin names in blue font with white background). Major city names in uppercase and bold font. (B) Photograph of live specimen of T. portillorum (5.5 cm standard length) from the Guacerique River (longitude= -87.109, latitude= 14.033), a tributary of the Choluteca River basin. (C) Habitat of T. portillorum. From the Guacerique River, tributary of the Choluteca River basin (longitude -87.111, latitude 14.032)." figureDoi="http://doi.org/10.5281/zenodo.10561935" httpUri="https://zenodo.org/record/10561935/files/figure.png" pageId="2" pageNumber="398">Fig. 1C</figureCitation>
), as well as hidden in vegetation along the riverbank undercut. In our surveys, 
<taxonomicName id="BEBDE2DCFFAC9B51FE80F830FE75F83C" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[246,394,2019,2043]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAC9B51FE80F830FE75F83C" box="[246,394,2019,2043]" italics="true" pageId="2" pageNumber="398">T. portillorum</emphasis>
</taxonomicName>
was found in sympatry with other species including 
<taxonomicName id="BEBDE2DCFFAC9B51FDCCF837FAB0F83C" authority="Steindachner 1863" authorityName="Steindachner" authorityYear="1863" box="[954,1359,2019,2043]" family="Poeciliidae" genus="Poecilia" kingdom="Animalia" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="mexicana">
<emphasis id="4BC9454DFFAC9B51FDCCF837FB80F83C" box="[954,1151,2019,2043]" italics="true" pageId="2" pageNumber="398">Poecilia mexicana</emphasis>
Steindachner 1863
</taxonomicName>
, 
<taxonomicName id="BEBDE2DCFFAC9B50FB2FF837FE7CFF77" authority="(Fowler 1923)" baseAuthorityName="Fowler" baseAuthorityYear="1923" family="Poeciliidae" genus="Alfaro" kingdom="Animalia" lastPageId="3" lastPageNumber="399" order="Cyprinodontiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="huberi">
<emphasis id="4BC9454DFFAC9B50FB2FF837FF22FF68" italics="true" lastPageId="3" lastPageNumber="399" pageId="2" pageNumber="398">Alfaro huberi</emphasis>
(Fowler 1923)
</taxonomicName>
, 
<taxonomicName id="BEBDE2DCFFAD9B50FFE6FF43FD11FF68" authority="(Kner 1858)" authorityName="Matamoros &amp; Schaefer" authorityYear="2010" baseAuthorityName="Kner" baseAuthorityYear="1858" box="[400,750,151,176]" family="Heptapteridae" genus="Rhamdia" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="399" phylum="Chordata" rank="species" species="laticauda">
<emphasis id="4BC9454DFFAD9B50FFE6FF43FDA1FF68" box="[400,606,151,175]" italics="true" pageId="3" pageNumber="399">Rhamdia laticauda</emphasis>
(Kner 1858)
</taxonomicName>
, 
<taxonomicName id="BEBDE2DCFFAD9B50FC8DFF43FB8DFF77" authority="(Miller 1907)" baseAuthorityName="Miller" baseAuthorityYear="1907" box="[763,1138,151,176]" family="Poeciliidae" genus="Phallichthys" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="399" phylum="Chordata" rank="species" species="amates">
<emphasis id="4BC9454DFFAD9B50FC8DFF43FC2AFF68" box="[763,981,151,175]" italics="true" pageId="3" pageNumber="399">Phallichthys amates</emphasis>
(Miller 1907)
</taxonomicName>
, 
<taxonomicName id="BEBDE2DCFFAD9B50FA09FF4CFF27FF14" authority="(Gunther 1860)" baseAuthorityName="Gunther" baseAuthorityYear="1860" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="399" phylum="Chordata" rank="species" species="aeneus">
<emphasis id="4BC9454DFFAD9B50FA09FF4CFACCFF68" box="[1151,1331,152,175]" italics="true" pageId="3" pageNumber="399">Astyanax aeneus</emphasis>
(Günther 1860)
</taxonomicName>
, 
<taxonomicName id="BEBDE2DCFFAD9B50FE95FF6FFD5EFF14" authority="(Gunther 1867)" baseAuthorityName="Gunther" baseAuthorityYear="1867" box="[227,673,187,212]" family="Cichlidae" genus="Amatitlania" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="399" phylum="Chordata" rank="species" species="nigrofasciata">
<emphasis id="4BC9454DFFAD9B50FE95FF6FFE0BFF13" box="[227,500,187,212]" italics="true" pageId="3" pageNumber="399">Amatitlania nigrofasciata</emphasis>
(Günther 1867)
</taxonomicName>
and 
<taxonomicName id="BEBDE2DCFFAD9B50FCA2FF6FFB66FF14" authority="(Gunther 1867)" baseAuthorityName="Gunther" baseAuthorityYear="1867" box="[724,1177,187,211]" family="Cichlidae" genus="Parachromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="399" phylum="Chordata" rank="species" species="motaguensis">
<emphasis id="4BC9454DFFAD9B50FCA2FF6FFC15FF14" box="[724,1002,187,211]" italics="true" pageId="3" pageNumber="399">Parachromis motaguensis</emphasis>
(Günther 1867)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="31A7CAD4FFAD9B50FEB1FF34FB3FFDD0" pageId="3" pageNumber="399" type="distribution">
<paragraph id="7902995FFFAD9B50FEB1FF34FB3FFDD0" blockId="3.[151,1437,151,823]" pageId="3" pageNumber="399">
<collectorName id="D448FC89FFAD9B50FEB1FF34FEE6FF30" box="[199,281,224,248]" pageId="3" pageNumber="399">Carr Jr.</collectorName>
&amp; 
<collectorName id="D448FC89FFAD9B50FF36FF0BFE47FF30" box="[320,440,223,247]" pageId="3" pageNumber="399">Giovannoli</collectorName>
(1950) surveyed the Choluteca River and did not report any 
<taxonomicName id="BEBDE2DCFFAD9B50FA16FF0BFB08FF30" box="[1120,1271,223,247]" family="Profundulidae" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="399" phylum="Chordata" rank="family">Profundulidae</taxonomicName>
species in this river. Later on 
<bibRefCitation id="1D2CE4AEFFAD9B50FF49FED0FDB3FEDB" author="Matamoros, W. A. &amp; Schaefer, J. F. &amp; Kreiser, B. R." box="[319,588,259,284]" pageId="3" pageNumber="399" pagination="1 - 38" refId="ref3293" refString="Matamoros, W. A., Schaefer, J. F. &amp; Kreiser, B. R. (2009) Annotated checklist of the freshwater fishes of continental and insular Honduras. Zootaxa, 2307 (1), 1 - 38. https: // doi. org / 10.11646 / zootaxa. 2307.1.1" type="journal article" year="2009">
Matamoros 
<emphasis id="4BC9454DFFAD9B50FFB5FED0FE0AFEDC" box="[451,501,259,283]" italics="true" pageId="3" pageNumber="399">et al</emphasis>
. (2009)
</bibRefCitation>
in a major river drainage checklist of Honduran freshwater fishes, surveyed the Choluteca and the 
<location id="7C62CF84FFAD9B50FFE2FEFCFDDEFEF8" LSID="urn:lsid:plazi:treatment:F1142849FFAE9B50FEB1F9DDFB9FFD0C:7C62CF84FFAD9B50FFE2FEFCFDDEFEF8" box="[404,545,295,319]" country="Honduras" county="Central America" municipality="Mosquitia-San Juan Ichthyological province" name="Patuca River" pageId="3" pageNumber="399" stateProvince="Choluteca">Patuca River</location>
and at that time they also fail to detect any 
<taxonomicName id="BEBDE2DCFFAD9B50FA6DFEF3FB4DFEF8" box="[1051,1202,295,319]" family="Profundulidae" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="399" phylum="Chordata" rank="family">Profundulidae</taxonomicName>
species in these two rivers. Although the Choluteca and Patuca Rivers are among the most important rivers in Honduras, their ichthyofauna remains poorly known. Besides the above-mentioned publications, no other attempts have been made to increase the knowledge of the distributional patterns and ecology of the fish communities of these rivers. Our results demonstrate that the distributional range of 
<taxonomicName id="BEBDE2DCFFAD9B50FFCEFE6CFDB1FE08" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[440,590,439,463]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="399" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAD9B50FFCEFE6CFDB1FE08" box="[440,590,439,463]" italics="true" pageId="3" pageNumber="399">T. portillorum</emphasis>
</taxonomicName>
is larger than previously thought, and that it extends to localities approaching drainages in eastern Honduras in the 
<collectingMunicipality id="99660325FFAD9B50FC44FE0FFBEDFE34" box="[562,1042,475,499]" pageId="3" pageNumber="399">Mosquitia-San Juan Ichthyological province</collectingMunicipality>
(
<bibRefCitation id="1D2CE4AEFFAD9B50FA55FE08FADFFE33" author="Matamoros, W. A. &amp; Kreiser, B. R. &amp; Schaefer, J. F." box="[1059,1312,475,500]" pageId="3" pageNumber="399" pagination="351 - 365" refId="ref3093" refString="Matamoros, W. A., Kreiser, B. R. &amp; Schaefer, J. F. (2012) A delineation of Nuclear Middle America biogeographical provinces based on river basin faunistic similarities. Reviews in Fish Biology and Fisheries, 22, 351 - 365. https: // doi. org / 10.1007 / s 11160 - 011 - 9232 - 8" type="journal article" year="2012">
Matamoros 
<emphasis id="4BC9454DFFAD9B50FAD1FE08FB1FFE34" box="[1191,1248,475,499]" italics="true" pageId="3" pageNumber="399">et al.</emphasis>
2012
</bibRefCitation>
, 2015), the Atlantic slope of 
<collectingCounty id="9063E1D3FFAD9B50FF26FE2BFE00FDD0" box="[336,511,511,535]" pageId="3" pageNumber="399">Central America</collectingCounty>
and the upper reaches of the Choluteca River in the Pacific slope.
</paragraph>
</subSubSection>
<subSubSection id="31A7CAD4FFAD9B50FEB1FDF7FB9FFD0C" pageId="3" pageNumber="399" type="biology_ecology">
<paragraph id="7902995FFFAD9B50FEB1FDF7FB9FFD0C" blockId="3.[151,1437,151,823]" pageId="3" pageNumber="399">
Regarding the type of habitats in which 
<taxonomicName id="BEBDE2DCFFAD9B50FC04FDF0FCF8FDFC" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[626,775,547,571]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="399" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAD9B50FC04FDF0FCF8FDFC" box="[626,775,547,571]" italics="true" pageId="3" pageNumber="399">T. portillorum</emphasis>
</taxonomicName>
is found, they generally coincide with the habitats reported in 
<bibRefCitation id="1D2CE4AEFFAD9B50FEE1FD9CFE0BFDA7" author="Matamoros, W. A. &amp; Schaefer, J. F." box="[151,500,583,608]" pageId="3" pageNumber="399" pagination="1498 - 507" refId="ref3234" refString="Matamoros, W. A. &amp; Schaefer, J. F. (2010) A new species of Profundulus (Cyprinodontiformes: Profundulidae) from the Honduran central highlands. Journal of Fish Biology, 76, 1498 - 507. https: // doi. org / 10.1111 / j. 1095 - 8649.2010.02596. x" type="journal article" year="2010">Matamoros and Schaefer (2010)</bibRefCitation>
. This fish is found at higher altitudes lower order streams, ranging from ~570 to ~1670 masl. These creeks are all characterized by having smaller widths, from less than a meter to ~ 10 meters, with bottoms dominated by boulders, rocks, gravel and cobblestone. The number of species that share habitats with 
<taxonomicName id="BEBDE2DCFFAD9B50FAAAFD44FA8DFD60" baseAuthorityName="Matamoros &amp; Schaefer" baseAuthorityYear="2010" box="[1244,1394,655,679]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="3" pageNumber="399" phylum="Chordata" rank="species" species="portillorum">
<emphasis id="4BC9454DFFAD9B50FAAAFD44FA8DFD60" box="[1244,1394,655,679]" italics="true" pageId="3" pageNumber="399">T. portillorum</emphasis>
</taxonomicName>
has been increased from what was previously known and includes more poecilids and cichlids.
</paragraph>
</subSubSection>
</treatment>
</document>