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<document id="3F0C129B8079E7E77B2CFEC766CC2FEC" ID-DOI="10.5281/zenodo.5356276" ID-ISSN="0253-116X" ID-Zenodo-Dep="5356276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630216915060" checkinUser="marcus" docAuthor="W, David &amp; M, Boris" docDate="2016" docId="A31A87A70D0CFFF452C5EF4BFC4E3824" docLanguage="en" docName="LinzerbiolBeitr.2016.48.2.1783-1806.pdf" docOrigin="Linzer biologische Beiträge 48 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Acinopus (Acinopus) arabicus W &amp; M 2016, nov.sp." docType="treatment" docVersion="4" lastPageNumber="1803" masterDocId="5F23FFDF0D1DFFE05277EA3CFF833D7C" masterDocTitle="Four new species of genus Acinopus DEJEAN, 1821, subgenus Acinopus from southern Iran, from Sinai, and from western Saudi Arabia, and faunistic and taxonomic notes on species previously described (Coleoptera, Carabidae, Harpalini, Harpalina)" masterLastPageNumber="1806" masterPageNumber="1783" pageNumber="1800" updateTime="1699190026385" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1C0B1013EF2973C6C8A22F13E1B535F8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E61F6DFF37F3B779DD0589FBD268036C">
<mods:title id="83D9439ECBFC216B3F48459295B1BFFC">Four new species of genus Acinopus DEJEAN, 1821, subgenus Acinopus from southern Iran, from Sinai, and from western Saudi Arabia, and faunistic and taxonomic notes on species previously described (Coleoptera, Carabidae, Harpalini, Harpalina)</mods:title>
</mods:titleInfo>
<mods:name id="F6221E4CF572F9B3AD1E04D98D80BEDA" type="personal">
<mods:role id="CF42853F2EE231F33E3C16AD931B50AD">
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<mods:namePart id="3DC7BC6CE953B682B3E4B2DE6D4E9750">W, David</mods:namePart>
</mods:name>
<mods:name id="3F6F0D09D8D37C3CF22CC538DF0E29F0" type="personal">
<mods:role id="E85FB51876170A6BCFF698493775F3B7">
<mods:roleTerm id="890C77A6B73D6F20DF7A70A87F21F100">Author</mods:roleTerm>
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<mods:namePart id="05DFAB5A5470680B0EF8ACE60601BB5C">M, Boris</mods:namePart>
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<mods:typeOfResource id="5057D109404C37E121F25A329626313A">text</mods:typeOfResource>
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<mods:title id="1378D0FE0BB05B8CF8103713EBACB99F">Linzer biologische Beiträge</mods:title>
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<mods:part id="21D5EFDAEE4D280508556C617F6F6A1A">
<mods:date id="3F982915E0CE33704D1F1DEEC7F67293">2016</mods:date>
<mods:detail id="F6AC33633252AC98692EBA26FC9B9EF4" type="pubDate">
<mods:number id="F615490C6DB1E3D9A54E6A407823DA52">2016-12-19</mods:number>
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<mods:detail id="E7382B8D4855D4914DDFFCFDCA6C6EEA" type="volume">
<mods:number id="34579A20228EB6B787D7D471C66A5C6A">48</mods:number>
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<mods:detail id="BD2B8D9A9FEBDD692FE49E80B1AD5358" type="issue">
<mods:number id="5BDEA84E655B3D574F823FCC75CD1EC2">2</mods:number>
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<mods:extent id="69DCACEB3A3794729515C12D976D879C" unit="page">
<mods:start id="293A7B8CA05B52400575400DC1C20765">1783</mods:start>
<mods:end id="14A5085C76FD0EA8CE74DA2CE8BEF1FA">1806</mods:end>
</mods:extent>
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<mods:classification id="9D1CFB9262EF9339BB464DA95F5418BB">journal article</mods:classification>
<mods:identifier id="C9E22EA9866739244CD68E5CC2754B60" type="DOI">10.5281/zenodo.5356276</mods:identifier>
<mods:identifier id="847507168992A471E171E1DA44B78521" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="1774C2D5147AB336A8592BA5B9D10714" type="Zenodo-Dep">5356276</mods:identifier>
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<treatment id="A31A87A70D0CFFF452C5EF4BFC4E3824" ID-DOI="http://doi.org/10.5281/zenodo.5452545" ID-Zenodo-Dep="5452545" LSID="urn:lsid:plazi:treatment:A31A87A70D0CFFF452C5EF4BFC4E3824" httpUri="http://treatment.plazi.org/id/A31A87A70D0CFFF452C5EF4BFC4E3824" lastPageId="20" lastPageNumber="1803" pageId="17" pageNumber="1800">
<subSubSection id="63A9653A0D0CFFF152C5EF4BFDF4389E" pageId="17" pageNumber="1800" type="nomenclature">
<paragraph id="2B0C36B10D0CFFF152C5EF4BFC8938F0" blockId="17.[178,778,1398,1420]" box="[178,778,1398,1420]" pageId="17" pageNumber="1800">
<heading id="704481DD0D0CFFF152C5EF4BFC8938F0" box="[178,778,1398,1420]" fontSize="8" level="2" pageId="17" pageNumber="1800" reason="2">
<emphasis id="19C7EAA30D0CFFF152C5EF4BFC8938F0" bold="true" box="[178,778,1398,1420]" pageId="17" pageNumber="1800">
<taxonomicName id="ECB34D320D0CFFF152C5EF4BFE6C38F0" ID-CoL="64KDY" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[178,495,1398,1420]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1800" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov." subGenus="Acinopus">
<emphasis id="19C7EAA30D0CFFF152C5EF4BFE9738F0" bold="true" box="[178,276,1399,1420]" italics="true" pageId="17" pageNumber="1800">Acinopus</emphasis>
(
<emphasis id="19C7EAA30D0CFFF15354EF4BFE0638F0" bold="true" box="[291,389,1399,1420]" italics="true" pageId="17" pageNumber="1800">Acinopus</emphasis>
)
<emphasis id="19C7EAA30D0CFFF153E2EF4AFE6C38F0" bold="true" box="[405,495,1398,1420]" italics="true" pageId="17" pageNumber="1800">arabicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D0CFFF15382EF4BFDC238F0" box="[501,577,1399,1420]" pageId="17" pageNumber="1800" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="B3882A340D0CFFF15039EF4AFD1238F0" box="[590,657,1398,1420]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="17" pageNumber="1800">Figs 4</figureCitation>
,
<figureCitation id="B3882A340D0CFFF150EAEF4BFD5E38F0" box="[669,733,1399,1420]" captionStart="Figs 9-12" captionStartId="8.[178,218,1513,1535]" captionTargetBox="[209,1036,218,1469]" captionTargetId="figure-0@8.[178,1085,204,1527]" captionTargetPageId="8" captionText="Figs 9-12: Acinopus DEJEAN, 1821. Median lobe of male. (9, 10) A. brittoni nov.sp. (9) Lateral view (holotype). (10) dorsal view (paratype). (11, 12) A. arabicus nov.sp. (11) Lateral view (paratype). (12) dorsal view (holotype). Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5356282" httpUri="https://zenodo.org/record/5356282/files/figure.png" pageId="17" pageNumber="1800">11, 12</figureCitation>
,
<figureCitation id="B3882A340D0CFFF1509EEF4AFC8038F0" box="[745,771,1398,1420]" captionStart="Figs 13-16" captionStartId="9.[178,218,1509,1531]" captionTargetBox="[257,975,222,1478]" captionTargetId="figure-0@9.[178,1085,204,1527]" captionTargetPageId="9" captionText="Figs 13-16: Acinopus DEJEAN, 1821. Female genitalia, (right hemisternite, gonocoxite1, 2, ventral view). (13) A. orszuliki nov.sp. (paratype, type locality). (14) A. sinaiticus nov.sp. (paratype, Mt. Katherine). (15) A. brittoni nov.sp. (paratype, type locality). (16) A. arabicus nov.sp. (paratype, type locality). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5356284" httpUri="https://zenodo.org/record/5356284/files/figure.png" pageId="17" pageNumber="1800">16</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="2B0C36B10D0CFFF152C5EFA1FDF4389E" blockId="17.[178,1087,1436,1511]" pageId="17" pageNumber="1800">
T y p e m a t e r i a l:
<typeStatus id="F40888130D0CFFF153DEEFA0FE7838CE" box="[425,507,1436,1458]" pageId="17" pageNumber="1800" type="holotype">Holotype</typeStatus>
6, labelled: &quot;
<collectingCountry id="53A476210D0CFFF1500CEFA1FC9E38CE" box="[635,797,1437,1458]" name="Saudi Arabia" pageId="17" pageNumber="1800">SAUDI ARABIA</collectingCountry>
/
<collectingRegion id="E977F8530D0CFFF15145EFA0FC1238CE" box="[818,913,1436,1458]" country="Saudi Arabia" name="'Asir" pageId="17" pageNumber="1800">Asir Prov.</collectingRegion>
,
<quantity id="EC4B9B540D0CFFF151EDEFA0FB9038CE" box="[922,1043,1436,1458]" metricMagnitude="3" metricUnit="m" metricValue="2.925" metricValueMax="3.05" metricValueMin="2.8" pageId="17" pageNumber="1800" unit="m" value="2925.0" valueMax="3050.0" valueMin="2800.0">2800-3050 m</quantity>
/ Al Soudah, ca.
<quantity id="EC4B9B540D0CFFF1533DEF88FE0038B6" box="[330,387,1460,1482]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="1800" unit="km" value="50.0">50 km</quantity>
NW Abha /
<date id="5F0D10710D0CFFF15383EF89FDC238B6" box="[500,577,1461,1482]" pageId="17" pageNumber="1800" value="1994-01-04">4.1.1994</date>
/ H.J. Bremer leg.&quot; (black print on white label), (MFNB).
<typeStatus id="F40888130D0CFFF152A8EFF1FEB4389E" box="[223,311,1485,1506]" pageId="17" pageNumber="1800" type="paratype">Paratypes</typeStatus>
: 16,
<specimenCount id="3DB5FD380D0CFFF15310EFF1FEFD389B" box="[359,382,1485,1511]" pageId="17" pageNumber="1800" type="female">1♀</specimenCount>
: same data (MFNB, cWR).
</paragraph>
</subSubSection>
<subSubSection id="63A9653A0D0FFFF452C5EAE7FC4E3824" lastPageId="20" lastPageNumber="1803" pageId="18" pageNumber="1801" type="description">
<paragraph id="2B0C36B10D0FFFF252C5EAE7FDC13C71" blockId="18.[178,1085,218,269]" pageId="18" pageNumber="1801">E t y m o l o g y: The specific epithet is a latinizid adjective, based on the name of the region in which this species was found.</paragraph>
<paragraph id="2B0C36B10D0FFFF252C5EB20FE983F4D" blockId="18.[178,1085,283,562]" pageId="18" pageNumber="1801">
D i a g n o s i s: A micropterous species of small to medium size for
<taxonomicName id="ECB34D320D0FFFF251D6EB27FC7C3C4C" authorityName="DEJEAN" authorityYear="1821" box="[929,1023,283,304]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1801" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D0FFFF251D6EB27FC7C3C4C" box="[929,1023,283,304]" italics="true" pageId="18" pageNumber="1801">Acinopus</emphasis>
</taxonomicName>
, with excision of dorsal edge of right mandible (subgenus
<taxonomicName id="ECB34D320D0FFFF250A8EB04FCBA3C31" authorityName="DEJEAN" authorityYear="1821" box="[735,825,312,333]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1801" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D0FFFF250A8EB04FCBA3C31" box="[735,825,312,333]" italics="true" pageId="18" pageNumber="1801">Acinopus</emphasis>
</taxonomicName>
), black, tarsomeres and antennae dark reddish-brown, labrum moderately incised at anterior margin, clypeus with one setiferous pore puncture at anterior angles, pronotum laterally almost rectilinearly or very weakly convexly narrowed toward the obtuse-angled posterior angles and weakly convexly narrowed apicad, anterior angles somewhat protruding, narrowly rounded at tip, posterior angles widely rounded, elytra short-oval, moderately convex on disc, metepisternum very short, pro- and mesotarsi in males moderately widened, apical lamella of median lobe short, about somewhat elongate-triangular (dorsal view). Habitus see
<figureCitation id="B3882A340D0FFFF252AEE820FE983F4D" box="[217,283,540,562]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="18" pageNumber="1801">Fig. 4.</figureCitation>
</paragraph>
<paragraph id="2B0C36B10D0FFFF252C5E87DFEE73FF3" blockId="18.[178,1085,577,655]" pageId="18" pageNumber="1801">
D e s c r i p t i o n: Body length in males
<quantity id="EC4B9B540D0FFFF2500FE87DFC8F3F2A" box="[632,780,577,598]" metricMagnitude="-2" metricUnit="m" metricValue="1.455" metricValueMax="1.47" metricValueMin="1.44" pageId="18" pageNumber="1801" unit="mm" value="14.55" valueMax="14.7" valueMin="14.4">14.4-14.7 mm</quantity>
, in
<typeStatus id="F40888130D0FFFF25142E87DFC0D3F2B" box="[821,910,577,599]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
<quantity id="EC4B9B540D0FFFF251E1E87DFC773F2A" box="[918,1012,577,598]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="18" pageNumber="1801" unit="mm" value="14.7">14.7 mm</quantity>
, in the female
<typeStatus id="F40888130D0FFFF2528AE862FED73F0F" box="[253,340,606,627]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
<quantity id="EC4B9B540D0FFFF2532CE861FE3A3F0F" box="[347,441,605,627]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="18" pageNumber="1801" unit="mm" value="12.9">12.9 mm</quantity>
; width in males
<quantity id="EC4B9B540D0FFFF25017E861FD5B3F0F" box="[608,728,605,627]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" metricValueMax="5.5" metricValueMin="5.4" pageId="18" pageNumber="1801" unit="mm" value="5.45" valueMax="5.5" valueMin="5.4">5.4-5.5 mm</quantity>
, in
<typeStatus id="F40888130D0FFFF2508AE861FCD53F0F" box="[765,854,605,627]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
<quantity id="EC4B9B540D0FFFF2512AE861FC2E3F0F" box="[861,941,605,627]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="1801" unit="mm" value="5.5">5.5 mm</quantity>
; in the female
<typeStatus id="F40888130D0FFFF252C5E846FE8A3FF3" box="[178,265,634,655]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
<quantity id="EC4B9B540D0FFFF25367E845FEE23FF3" box="[272,353,633,655]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="18" pageNumber="1801" unit="mm" value="5.1">5.1 mm</quantity>
.
</paragraph>
<paragraph id="2B0C36B10D0FFFF252C5E8A2FD6B3FC8" blockId="18.[178,744,670,692]" box="[178,744,670,692]" pageId="18" pageNumber="1801">Colour: Black, tarsomeres and antennae reddish-brown.</paragraph>
<paragraph id="2B0C36B10D0FFFF252C5E8FFFBBD3EA5" blockId="18.[178,1086,707,985]" pageId="18" pageNumber="1801">
Head: Wide (as normal in
<taxonomicName id="ECB34D320D0FFFF253B2E8FFFDA03FA4" authorityName="DEJEAN" authorityYear="1821" box="[453,547,707,728]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1801" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D0FFFF253B2E8FFFDA03FA4" box="[453,547,707,728]" italics="true" pageId="18" pageNumber="1801">Acinopus</emphasis>
</taxonomicName>
), only somewhat narrower than pronotum (PW/HW in males 1.18-1.21, in
<typeStatus id="F40888130D0FFFF253EBE8E3FE763F89" box="[412,501,735,757]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
1.18; in the female
<typeStatus id="F40888130D0FFFF250B0E8DCFC9D3F89" box="[711,798,736,757]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
1.24). Eyes small and relatively flat, tempora oblique, about three fourth as long as diameter of eye or somewhat shorter in female
<typeStatus id="F40888130D0FFFF25310E925FE3D3E52" box="[359,446,793,814]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
(dorsally seen), rectilinearly converging to neck. Disc smooth, laterad with small, weakly impressed and widely spaced punctures. Labrum strongly and somewhat angulately incised at anterior margin, clypeus weakly semicircularly incised, with anterior angles with one setiferous pore puncture each. Mentum tooth small, submentum with one long seta laterally each side. Excision of dorsal lateral edge of right mandible well developed, almost semicircular, relatively close to base of mandible, basal angle of excision situated distinctly anterior to level of anterior margin of clypeus (with mandibles closed).
</paragraph>
<paragraph id="2B0C36B10D0FFFF252C5E9D4FC1D380D" blockId="18.[178,1085,1000,1393]" pageId="18" pageNumber="1801">
Pronotum (
<figureCitation id="B3882A340D0FFFF2535EE9D4FEED3E82" box="[297,366,1000,1022]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="18" pageNumber="1801">Fig. 4</figureCitation>
): Transverse (PW/PL in males 1.52-1.58, in
<typeStatus id="F40888130D0FFFF2512DE9D4FC303E82" box="[858,947,1000,1022]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
1.52; in the female
<typeStatus id="F40888130D0FFFF25374EE39FED93966" box="[259,346,1029,1050]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
1.50), widest at about end of anterior third, where the lateral seta inserted. Anterior margin a little wider than posterior margin, only weakly excavate, anterior angles moderately projecting forward, relatively narrowly rounded at tip. Sides moderately curved apicad, basad from widest point almost rectilinear at a short distance, and anterior to posterior angles gently convexly narrowed. Posterior angles somewhat obtuse-angled, widely rounded at tip. Basal edge almost rectlinear between the basal foveae, posterior angles very weakly shifted backwards. Basal foveae short, elongate, distinctly converging basad, shallowly or more strongly impressed, some punctures of unequal size in and around fovae, more or less widely spaced and continuing to the area of posterior angles. Disc convex, area of posterior angles more or less strongly flattened, anterior and posterior transverse impressions very shallow or only indicated, median line weakly impressed, terminated anterior to apical margin and hardly reaching basal margin. Lateral gutter narrow, somewhat widened toward anterior angles.
</paragraph>
<paragraph id="2B0C36B10D0FFFF352C5EFBCFC6C3C1E" blockId="18.[178,1086,1408,1601]" lastBlockId="19.[178,1085,218,354]" lastPageId="19" lastPageNumber="1802" pageId="18" pageNumber="1801">
Elytra (
<figureCitation id="B3882A340D0FFFF25288EFBCFEBC38EA" box="[255,319,1408,1430]" captionStart="Figs 1-4" captionStartId="4.[178,218,1514,1536]" captionTargetBox="[277,985,216,1490]" captionTargetId="figure-0@4.[178,1085,204,1527]" captionTargetPageId="4" captionText="Figs 1-4: Acinopus DEJEAN, 1821. Habitus. (1) A. orszuliki nov.sp. (male paratype, type locality). (2) A. sinaiticus nov.sp. (male paratype, Mt. Katherine). (3) A. brittoni nov.sp. (female paratype, type locality). (4) A. arabicus nov.sp. (male paratype, type locality)." figureDoi="http://doi.org/10.5281/zenodo.5356278" httpUri="https://zenodo.org/record/5356278/files/figure.png" pageId="18" pageNumber="1801">Fig. 4</figureCitation>
): short-oval (EL/EW in males 1.39-1.43, in
<typeStatus id="F40888130D0FFFF25171EFBCFCDC38EA" box="[774,863,1408,1430]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
1.39; in female
<typeStatus id="F40888130D0FFFF2567EEFBCFF5D38CE" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
1.42), somewhat wider than pronotum (EW/PW in males 1.08-1.11, in
<typeStatus id="F40888130D0FFFF25193EFA0FBBE38CE" box="[996,1085,1436,1458]" pageId="18" pageNumber="1801" type="holotype">holotype</typeStatus>
1.11; in female
<typeStatus id="F40888130D0FFFF25324EF85FE2938B2" box="[339,426,1465,1486]" pageId="18" pageNumber="1801" type="paratype">paratype</typeStatus>
1.12). On disc moderately convex, humeri distinctly developed, rounded at tip, without humeral tooth, not fused at suture; toward behind somewhat convexely enlarged, widest about in middle. Basal bead weakly sinuate, arcuately curving inside humerus and weakly angled or round towards lateral margin. Scutellar pore puncture present, scutellar stria long. Striae fine, smooth, intervals flat on disc, becoming convex toward elytral declivity, interval 3 without pore puncture (
<specimenCount id="3DB5FD380D0EFFF351AFEAE6FF683C70" pageId="19" pageNumber="1802" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
),
<typeStatus id="F40888130D0EFFF35377EACBFEDB3C71" box="[256,344,247,269]" pageId="19" pageNumber="1802" type="holotype">holotype</typeStatus>
with pore puncture on right elytron only at about beginning of apical fourth close to stria 2. Interval 5 with one to tree pore punctures apically, interval 7 with two to four pore punctures apically (
<typeStatus id="F40888130D0EFFF35051EB0CFD073C3A" box="[550,644,304,326]" pageId="19" pageNumber="1802" type="holotype">holotype</typeStatus>
), or such punctures lacking (remaining
<specimenCount id="3DB5FD380D0EFFF35660EB0CFE9B3C1E" pageId="19" pageNumber="1802" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
). Preapical sinuation weak in males, only suggested in female
<typeStatus id="F40888130D0EFFF351E6EB71FC6F3C1E" box="[913,1004,333,354]" pageId="19" pageNumber="1802" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5EB4DFCAF3CFB" blockId="19.[178,812,369,391]" box="[178,812,369,391]" pageId="19" pageNumber="1802">Hindwings: Reduced, about one third as long as elytral length.</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5EBA9FDC83F2A" blockId="19.[178,1085,405,599]" pageId="19" pageNumber="1802">Ventral surface: Prosternum with short setae, apex of prosternal process with long setae, proepisternum and anterior part of mesepisternum with sparse, fine setae, metepisternum and metasternum (laterally) with sparse, coarse seta-bearing punctures. Metepisternum very short, ratio of anterior margin/internal margin (visible parts) about 0.91, moderately narrowed behind. Abdominal sternites III-V with scattered setae of unequal length. Last sternite, beside some fine scattered setae, with two pore punctures bearing a long seta on each side at apical margin in both sexes.</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5E85AFB8D3E75" blockId="19.[178,1086,613,778]" pageId="19" pageNumber="1802">
Legs: Normal for
<taxonomicName id="ECB34D320D0EFFF35303E859FDAE3F07" box="[372,557,613,635]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1802" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="19C7EAA30D0EFFF35303E859FE513F06" box="[372,466,613,634]" italics="true" pageId="19" pageNumber="1802">Acinopus</emphasis>
species.
</taxonomicName>
Pro- and mesotarsi moderately widened in males (less wider than in
<taxonomicName id="ECB34D320D0EFFF353F6E8BEFD8B3FEB" authorityName="MENETRIES" authorityYear="1832" box="[385,520,642,663]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1802" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="19C7EAA30D0EFFF353F6E8BEFD8B3FEB" box="[385,520,642,663]" italics="true" pageId="19" pageNumber="1802">A. laevigatus</emphasis>
</taxonomicName>
), pro- and mesotarsomere 1 with two small adhesive scales apically, pro- and mesotarsomere 2-4 with biseriate adhesive vestiture. Ventroapical tubercle of protibia with three to five spines, arranged in a transverse row. Outer distal margin of fore tibia with five to eight spines. Eight to nine spines on lower surface of protibia arranged in one row. Tarsi smooth on superior surface except obligatory setae.
</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5E924FCC13EF8" blockId="19.[178,1085,792,900]" pageId="19" pageNumber="1802">Microsculpture of surface: Head and pronotum in males with transverse to almost isodiametric meshes weakly engraved and somewhat irregular, elytra with about the same kind of meshes somewhat more strongly engraved as on the forebody, surface very shiny, in females meshes somewhat stronger engraved, surface less shiny.</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5E9AEFC2C3E82" blockId="19.[178,1085,914,1022]" pageId="19" pageNumber="1802">
Median lobe of aedeagus (
<figureCitation id="B3882A340D0EFFF353BBE9AEFDC93ED5" box="[460,586,914,937]" captionStart="Figs 9-12" captionStartId="8.[178,218,1513,1535]" captionTargetBox="[209,1036,218,1469]" captionTargetId="figure-0@8.[178,1085,204,1527]" captionTargetPageId="8" captionText="Figs 9-12: Acinopus DEJEAN, 1821. Median lobe of male. (9, 10) A. brittoni nov.sp. (9) Lateral view (holotype). (10) dorsal view (paratype). (11, 12) A. arabicus nov.sp. (11) Lateral view (paratype). (12) dorsal view (holotype). Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5356282" httpUri="https://zenodo.org/record/5356282/files/figure.png" pageId="19" pageNumber="1802">Figs 11, 12</figureCitation>
): Of normal construction for
<taxonomicName id="ECB34D320D0EFFF351F2E9AFFBB93ED4" box="[901,1082,914,936]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1802" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="19C7EAA30D0EFFF351F2E9AFFC603ED4" box="[901,995,915,936]" italics="true" pageId="19" pageNumber="1802">Acinopus</emphasis>
species
</taxonomicName>
, middle and apical part somewhat bent to the left, terminal lamella wide and flat, somewhat elongate-triangular (dorsal view), and somewhat reflexed apically (lateral view), internal sac with a small, elongate group of about ten medium-sized spines.
</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5EE31FD343904" blockId="19.[178,1085,1037,1144]" pageId="19" pageNumber="1802">
Female genitalia (
<figureCitation id="B3882A340D0EFFF3531CEE31FE3B395F" box="[363,440,1037,1059]" captionStart="Figs 13-16" captionStartId="9.[178,218,1509,1531]" captionTargetBox="[257,975,222,1478]" captionTargetId="figure-0@9.[178,1085,204,1527]" captionTargetPageId="9" captionText="Figs 13-16: Acinopus DEJEAN, 1821. Female genitalia, (right hemisternite, gonocoxite1, 2, ventral view). (13) A. orszuliki nov.sp. (paratype, type locality). (14) A. sinaiticus nov.sp. (paratype, Mt. Katherine). (15) A. brittoni nov.sp. (paratype, type locality). (16) A. arabicus nov.sp. (paratype, type locality). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5356284" httpUri="https://zenodo.org/record/5356284/files/figure.png" pageId="19" pageNumber="1802">Fig. 16</figureCitation>
): Hemisternite asymmetrical, with numerous scattered setae in its internal, somewhat hyaline part. Gonocoxite 1 elongate, apically with numerous setae, gonocoxite 2 scoop-shaped, apically somewhat blunt, with a double-sensilla in a furrow, and with numerous fine long setae at concave side.
</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5EEBBFCAC3872" blockId="19.[178,1085,1159,1295]" pageId="19" pageNumber="1802">
I n t r a s p e c i f i c v a r i a b i l i t y: The material is too small to make an extended statement on this item. Concerning the
<specimenCount id="3DB5FD380D0EFFF350B7EE9FFCE539C5" box="[704,870,1187,1209]" pageId="19" pageNumber="1802" type="generic">three specimens</specimenCount>
examined it can be noted that the body size varies somewhat and that the elytra can have or have not one pore puncture in interval 3, with the possibility that only one elytron has this puncture, also the number of pore punctures in interval 5 and 7 can vary.
</paragraph>
<paragraph id="2B0C36B10D0EFFF352C5EF22FCD33B48" blockId="19.[178,1086,1309,1588]" pageId="19" pageNumber="1802">
C o m p a r i s o n s: The species is closest to
<taxonomicName id="ECB34D320D0EFFF350DCEF22FC94384E" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[683,791,1309,1331]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1802" phylum="Arthropoda" rank="species" species="brittoni" status="sp. nov.">
<emphasis id="19C7EAA30D0EFFF350DCEF22FC94384E" box="[683,791,1309,1331]" italics="true" pageId="19" pageNumber="1802">A. brittoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D0EFFF35156EF22FCE8384F" box="[801,875,1310,1331]" pageId="19" pageNumber="1802" rank="species">nov.sp.</taxonomicNameLabel>
in having the same body colour, a somewhat flattened body, a clypeus with anterior angles with one setiferous pore puncture each, and the same position of the excision of the dorsal lateral edge of the right mandible, but differs in smaller body size, a wider head (compared with pronotum), a longer pronotum with a somewhat narrower lateral gutter and with less protruding anterior angles, and with a less strongly flattened area of the posterior angles. In addition, the elytra of the new species are somewhat longer and a little less oval, the median lobe of the aedeagus is somewhat shorter and stouter, the apical lamella a little longer and almost evenly narrowed, apically more acute (dorsal view), somewhat reflexed apically (lateral view), the spines of the internal sac are lower in number (compare
<figureCitation id="B3882A340D0EFFF35011EC22FD5A3B4F" box="[614,729,1566,1588]" captionStart="Figs 9-12" captionStartId="8.[178,218,1513,1535]" captionTargetBox="[209,1036,218,1469]" captionTargetId="figure-0@8.[178,1085,204,1527]" captionTargetPageId="8" captionText="Figs 9-12: Acinopus DEJEAN, 1821. Median lobe of male. (9, 10) A. brittoni nov.sp. (9) Lateral view (holotype). (10) dorsal view (paratype). (11, 12) A. arabicus nov.sp. (11) Lateral view (paratype). (12) dorsal view (holotype). Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5356282" httpUri="https://zenodo.org/record/5356282/files/figure.png" pageId="19" pageNumber="1802">Figs 11, 12</figureCitation>
, and 9, 10).
</paragraph>
<paragraph id="2B0C36B10D09FFF452C5EAE7FDC43CE7" blockId="20.[178,1085,218,411]" pageId="20" pageNumber="1803">
H a b i t a t: Nothing is known about biotope and circumstances of collecting. Al Soudah is situated west of Abah City on the Sirwat Mountains, a massif with heights of more than
<quantity id="EC4B9B540D09FFF45352EB2FFEF73C55" box="[293,372,275,297]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="1803" unit="m" value="3000.0">3000 m</quantity>
a.s.l. After a source in the internet (http://www.the-saudi.net/saudiarabia/abha/abha_city.htm) the Abha region has the highest level of rainfall of any part of
<collectingCountry id="53A476210D09FFF452A3EB70FEE13C1E" box="[212,354,332,354]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
, the mountains are covered with stands of Araar trees,
<taxonomicName id="ECB34D320D09FFF451B8EB70FE613C02" authority="(VAHL) MASTERS" authorityName="MASTERS" baseAuthorityName="VAHL" class="Pinopsida" family="Cupressaceae" genus="Tetraclinis" kingdom="Plantae" order="Pinales" pageId="20" pageNumber="1803" phylum="Tracheophyta" rank="species" species="articulata">
<emphasis id="19C7EAA30D09FFF451B8EB70FE953C02" italics="true" pageId="20" pageNumber="1803">Tetraclinis articulata</emphasis>
(VAHL) MASTERS
</taxonomicName>
, an evergreen coniferous tree of the cypress (family
<taxonomicName id="ECB34D320D09FFF452C5EBB9FEBD3CE7" box="[178,318,389,411]" class="Pinopsida" family="Cupressaceae" kingdom="Plantae" order="Pinales" pageId="20" pageNumber="1803" phylum="Tracheophyta" rank="family">Cupressaceae</taxonomicName>
) and other natural forests.
</paragraph>
<paragraph id="2B0C36B10D09FFF452C5EB96FCD73C85" blockId="20.[178,1085,426,505]" pageId="20" pageNumber="1803">
D i s t r i b u t i o n: Up to present only known from one locality in the Sirwat Mountains (
<collectingCountry id="53A476210D09FFF45283EBFAFE073CA0" box="[244,388,454,476]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
), situated in the Sarat '
<collectingRegion id="E977F8530D09FFF450F7EBFAFD793CA0" box="[640,762,454,476]" country="Saudi Arabia" name="'Asir" pageId="20" pageNumber="1803">Asir region</collectingRegion>
in the middle of the Sarawat Mountain range, but surely more widely distributed in that massif.
</paragraph>
<paragraph id="2B0C36B10D09FFF452C5E834FC4E3824" blockId="20.[178,1086,519,1369]" pageId="20" pageNumber="1803">
R e m a r k s: The Sarawat mountain range, a massif running parallel to the western coast of the Arabian Peninsula and is among the Peninsula's most prominent geographical features, starts from the border of
<collectingCountry id="53A476210D09FFF45038E87CFD113F2A" box="[591,658,576,598]" name="Jordan" pageId="20" pageNumber="1803">Jordan</collectingCountry>
in the north to the Gulf of Aden in the south, running through
<collectingCountry id="53A476210D09FFF453DEE861FDB13F0F" box="[425,562,605,627]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
and
<collectingCountry id="53A476210D09FFF4501EE861FD363F0E" box="[617,693,605,626]" name="Yemen" pageId="20" pageNumber="1803">Yemen</collectingCountry>
. The range's northern half, known as Sarat al-Hejaz rarely rises about 2100 meters, while the middle and southern portions (Sarat '
<collectingRegion id="E977F8530D09FFF4528DE8AAFEAB3FD0" box="[250,296,662,684]" country="Saudi Arabia" name="'Asir" pageId="20" pageNumber="1803">Asir</collectingRegion>
and Sarat al-Yemen, respectively) can reach heights of over 3,300 meters. The discovery of two members of the subgenus
<taxonomicName id="ECB34D320D09FFF450D3E88FFC813FB4" authorityName="DEJEAN" authorityYear="1821" box="[676,770,691,712]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="genus">
<emphasis id="19C7EAA30D09FFF450D3E88FFC813FB4" box="[676,770,691,712]" italics="true" pageId="20" pageNumber="1803">Acinopus</emphasis>
</taxonomicName>
in western
<collectingCountry id="53A476210D09FFF4510CE88FFB873FB5" box="[891,1028,691,713]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
, both similar to the North African
<taxonomicName id="ECB34D320D09FFF453ACE8F3FDDF3F98" baseAuthorityName="FABRICIUS" baseAuthorityYear="1792" box="[475,604,719,740]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="species" species="sabulosus">
<emphasis id="19C7EAA30D09FFF453ACE8F3FDDF3F98" box="[475,604,719,740]" italics="true" pageId="20" pageNumber="1803">A. sabulosus</emphasis>
</taxonomicName>
, is very interesting in a zoogeographical point of view. There is a wide gap of about thousand kilometres between eastern populations of
<taxonomicName id="ECB34D320D09FFF452A5E935FED53E61" baseAuthorityName="FABRICIUS" baseAuthorityYear="1792" box="[210,342,776,798]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="species" species="sabulosus">
<emphasis id="19C7EAA30D09FFF452A5E935FED53E61" box="[210,342,776,798]" italics="true" pageId="20" pageNumber="1803">A. sabulosus</emphasis>
</taxonomicName>
from
<collectingCountry id="53A476210D09FFF453EBE934FE543E62" box="[412,471,776,798]" name="Libya" pageId="20" pageNumber="1803">Libya</collectingCountry>
and the populations of two new species in
<collectingCountry id="53A476210D09FFF451C3E934FBBE3E62" box="[948,1085,776,798]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
(from
<collectingCountry id="53A476210D09FFF45284E919FEB13E46" box="[243,306,805,826]" name="Egypt" pageId="20" pageNumber="1803">Egypt</collectingCountry>
, in fact from the
<collectingRegion id="E977F8530D09FFF4539FE919FD0A3E47" box="[488,649,805,827]" country="Egypt" name="Shamal Sina'" pageId="20" pageNumber="1803">Sinai Peninsula</collectingRegion>
, only one rather dissimilar and apparently unrelated species,
<taxonomicName id="ECB34D320D09FFF45318E97EFE6E3E2B" authority="W &amp; M, 2016" authorityName="W &amp; M" authorityYear="2016" box="[367,493,834,855]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="species" species="sinaiticus" status="sp. nov.">
<emphasis id="19C7EAA30D09FFF45318E97EFE6E3E2B" box="[367,493,834,855]" italics="true" pageId="20" pageNumber="1803">A. sinaiticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02F457D80D09FFF45383E97EFDC33E2B" box="[500,576,834,855]" pageId="20" pageNumber="1803" rank="species">nov.sp.</taxonomicNameLabel>
, is known). It can probably be hypothetizied, due to the relative similarity of
<taxonomicName id="ECB34D320D09FFF453B8E962FDD33E0F" baseAuthorityName="FABRICIUS" baseAuthorityYear="1792" box="[463,592,862,883]" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="species" species="sabulosus">
<emphasis id="19C7EAA30D09FFF453B8E962FDD33E0F" box="[463,592,862,883]" italics="true" pageId="20" pageNumber="1803">A. sabulosus</emphasis>
</taxonomicName>
and the two species from
<collectingCountry id="53A476210D09FFF45113E962FC6D3E08" box="[868,1006,862,884]" name="Saudi Arabia" pageId="20" pageNumber="1803">Saudi Arabia</collectingCountry>
, that
<taxonomicName id="ECB34D320D09FFF45650E962FE953EF3" baseAuthorityName="FABRICIUS" baseAuthorityYear="1792" class="Insecta" family="Carabidae" genus="Acinopus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1803" phylum="Arthropoda" rank="species" species="sabulosus">
<emphasis id="19C7EAA30D09FFF45650E962FE953EF3" italics="true" pageId="20" pageNumber="1803">A. sabulosus</emphasis>
</taxonomicName>
on the one hand and the two Arabian species on the other hand are adelphotaxa with an ancestor widely distributed over North Africa and the Arabian Peninsula in the Neogene, most likely not earlier than the middle Miocene, because in the Paleogene and the early Miocene, North Africa was dominated by tropical forests; by the middle Miocene grass-dominated ecosystems begin to spread and vegetation changed into a more open grassland vegetation towards the end of the Miocene (e.g.,
<bibRefCitation id="4F224B400D09FFF451EFEE35FE9F3947" author="MICHEELS A &amp; ERONEN J" pageId="20" pageNumber="1803" pagination="193 - 204" refId="ref14796" refString="MICHEELS A., ERONEN J. &amp; V. MOSBRUGGER (2009): The Late Miocene climate response to a modern Sahara desert. - Global and Planetary Change 67: 193 - 204." type="journal article" year="2009">MICHEELS et al. 2009: 193</bibRefCitation>
;
<bibRefCitation id="4F224B400D09FFF45350EE1AFD913947" author="JACOBS B &amp; PAN A" box="[295,530,1061,1083]" pageId="20" pageNumber="1803" pagination="57 - 72" refId="ref14487" refString="JACOBS B. F., PAN A. D. &amp; C. R. SCOTESE (2010): A review of the Cenozoic vegetation history of Africa. Pp. 57 - 72. - In: WERDELIN L. &amp; W. SANDERS (eds), Cenozoic Mammals of Africa. Berkeley: University of California Press, 986 pp." type="book chapter" year="2010">JACOBS et al. 2010: 69</bibRefCitation>
). The continuous distribution of the common ancestor of the species discussed was probably divided into the African and Arabian parts by lowland desert areas appeared for the first time in North Africa at around the Miocene/Pliocene boundary (e.g.,
<bibRefCitation id="4F224B400D09FFF45064EE40FD5B39ED" author="LE HOUEROU H" box="[531,728,1147,1169]" pageId="20" pageNumber="1803" pagination="619 - 647" refId="ref14711" refString="LE HOUEROU H. N. (1997): Climate, flora and fauna changes in the Sahara over the past 500 million years. - Journal of Arid Environments 37: 619 - 647." type="journal article" year="1997">LE HOUEROU 1997</bibRefCitation>
). This separation was aided by the Red Sea which formation began in the Eocene and became a sea in the Miocene, causing the isolation of the mountains of South-West Arabia from those of Africa (
<bibRefCitation id="4F224B400D09FFF45195EE88FE8D399B" author="BRITTON E" pageId="20" pageNumber="1803" pagination="126 - 131" refId="ref14287" refString="BRITTON E. B. (1948): Coleoptera: Cicindelidae and Carabidae. Pp. 87 - 125, pl. 7 - 9. - In: Expedition to South-west Arabia 1937 - 8, Vol. 1. London: British Museum (Natural History), pp. 67 - 178, pl. 4 - 9 [with an appendix by P. BASILEWSKY: Trois nouveaux Carabiques du Sud-Ouest de l'Arabie, pp. 126 - 131]." type="book chapter" year="1948">BRITTON 1948: 89</bibRefCitation>
). The result of alternating dry and damp climatic periods in a region today drier than in the past, caused a further splitting in two species with a relict-like occurrence, which need for surviving plants and herewith connected moisture, in the high mountains of western Arabia, in one species in the Sarat al-Hejaz in the northern portion of the Sarawat mountain range, and a second one in the Sarat '
<collectingRegion id="E977F8530D09FFF45091EF7FFC973825" box="[742,788,1347,1369]" country="Saudi Arabia" name="'Asir" pageId="20" pageNumber="1803">Asir</collectingRegion>
in its middle part.
</paragraph>
</subSubSection>
</treatment>
</document>
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