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<document id="2FCBD16B92EED7AA5C6AC7F8B03FABE1" ID-DOI="10.5281/zenodo.203245" ID-GBIF-Dataset="8484b438-d520-4985-bccf-f914612a5798" ID-ISSN="1175-5326" ID-Zenodo-Dep="203245" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460235874530" checkinUser="plazi" docAuthor="Sites, Robert W. &amp; Zettel, Herbert" docDate="2011" docId="513487E73660FF93FF4BD56EFD0283E6" docLanguage="en" docName="zt02760p017.pdf" docOrigin="Zootaxa 2760" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pogonocaudina Sites and Zettel, NEW GENUS" docType="treatment" docVersion="11" lastPageNumber="6" masterDocId="AD0DFF9F3661FF96FFDCD10FFFA98557" masterDocTitle="Waterfall-inhabiting Naucoridae (Hemiptera: Heteroptera) of southern India and Sri Lanka: Pogonocaudina Sites and Zettel, n. gen., and a review of Diaphorocoris with descriptions of two new species" masterLastPageNumber="17" masterPageNumber="1" pageNumber="2" updateTime="1698239580061" updateUser="plazi">
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<mods:title id="2EEFEB3E696807CA0C2FB508FBF0E269">Waterfall-inhabiting Naucoridae (Hemiptera: Heteroptera) of southern India and Sri Lanka: Pogonocaudina Sites and Zettel, n. gen., and a review of Diaphorocoris with descriptions of two new species</mods:title>
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<mods:namePart id="479C2DC2F47A7A9B8E6FD738DF9E8657">Sites, Robert W.</mods:namePart>
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<mods:date id="AF1CA561007083231D49AD0C8B3AC4FE">2011</mods:date>
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<treatment id="513487E73660FF93FF4BD56EFD0283E6" ID-DOI="http://doi.org/10.5281/zenodo.5670822" ID-GBIF-Taxon="119388196" ID-Zenodo-Dep="5670822" LSID="urn:lsid:plazi:treatment:513487E73660FF93FF4BD56EFD0283E6" httpUri="http://treatment.plazi.org/id/513487E73660FF93FF4BD56EFD0283E6" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2">
<subSubSection id="9187657A3660FF97FF4BD56EFF5681CB" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="D92236F13660FF97FF4BD56EFD76812D" blockId="1.[151,735,1120,1180]" box="[151,735,1120,1147]" pageId="1" pageNumber="2">
<heading id="826A819D3660FF97FF4BD56EFD76812D" bold="true" box="[151,735,1120,1147]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="EBE9EAE33660FF97FF4BD56EFD76812D" bold="true" box="[151,735,1120,1147]" pageId="1" pageNumber="2">
<taxonomicName id="1E9D4D723660FF97FF4BD56EFD89812D" ID-CoL="6TMW" authority="Sites and Zettel" authorityName="Sites and Zettel" box="[151,544,1120,1147]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33660FF97FF4BD56EFEFF812C" bold="true" box="[151,342,1121,1147]" italics="true" pageId="1" pageNumber="2">Pogonocaudina</emphasis>
Sites and Zettel
</taxonomicName>
,
<taxonomicNameLabel id="F0DA57983660FF97FDF0D56FFD76812D" box="[556,735,1120,1146]" pageId="1" pageNumber="2" rank="genus">NEW GENUS</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D92236F13660FF97FF4BD58BFF5681CB" blockId="1.[151,735,1120,1180]" box="[151,255,1156,1180]" pageId="1" pageNumber="2">
<figureCitation id="41A62A743660FF97FF4BD58BFF5681CB" box="[151,255,1156,1180]" captionStart-0="FIGURES 1 3" captionStart-1="FIGURES 4 5" captionStart-2="FIGURE 6" captionStartId-0="2.[151,264,1769,1792]" captionStartId-1="4.[151,264,1958,1981]" captionStartId-2="5.[151,250,1082,1105]" captionTargetBox-0="[163,1411,754,1714]" captionTargetBox-1="[218,1373,193,1938]" captionTargetBox-2="[151,1436,193,1060]" captionTargetId-0="figure@2.[151,1436,735,1745]" captionTargetId-1="figure@4.[218,1374,193,1939]" captionTargetId-2="figure@5.[151,1436,193,1061]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1 3. Features of Pogonocaudina, n. gen.: (1) habitus (dry specimen), (2) single segmented front tarsus with paired claws of male appressed to anterior margin of femur, (3) dense fringe of long setae on outer margin of posterior abdominal segments (specimen in alcohol)." captionText-1="FIGURES 4 5. (4) Kiliyur Waterfall, the type locality of Pogonocaudina indica n. sp. and Diaphorocoris kiliyur n. sp., (5) habitat of Pogonocaudina indica n. sp., Diaphorocoris arunachalami n. sp. and Diaphorocoris kiliyur n. sp." captionText-2="FIGURE 6. Live specimen of Pogonocaudina indica n. sp." httpUri-0="https://zenodo.org/record/203246/files/figure.png" httpUri-1="https://zenodo.org/record/203247/files/figure.png" httpUri-2="https://zenodo.org/record/203248/files/figure.png" pageId="1" pageNumber="2">Figs. 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="9187657A3660FF97FF4BD5C2FCF681B6" box="[151,863,1227,1250]" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="D92236F13660FF97FF4BD5C2FCF681B6" blockId="1.[151,863,1227,1250]" box="[151,863,1227,1250]" pageId="1" pageNumber="2">
<typeStatus id="062688533660FF97FF4BD5C2FF6181B5" box="[151,200,1229,1250]" pageId="1" pageNumber="2">Type</typeStatus>
species:
<taxonomicName id="1E9D4D723660FF97FEFAD5C3FD1B81B6" authority="Sites and Zettel" authorityName="Sites and Zettel" box="[294,690,1227,1250]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="EBE9EAE33660FF97FEFAD5C3FDA281B5" box="[294,523,1228,1250]" italics="true" pageId="1" pageNumber="2">Pogonocaudina indica</emphasis>
Sites and Zettel
</taxonomicName>
,
<taxonomicNameLabel id="F0DA57983660FF97FD60D5C2FCF681B6" box="[700,863,1228,1250]" pageId="1" pageNumber="2" rank="species">
<collectionCode id="BF8CAE343660FF97FD60D5C2FD5F81B5" box="[700,758,1229,1250]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="1" pageNumber="2">NEW</collectionCode>
<collectionCode id="BF8CAE343660FF97FD20D5C3FCF681B6" box="[764,863,1228,1249]" pageId="1" pageNumber="2">SPECIES</collectionCode>
</taxonomicNameLabel>
</paragraph>
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<subSubSection id="9187657A3660FF95FF4BD401FEF385A0" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
<paragraph id="D92236F13660FF97FF4BD401FDA680C5" blockId="1.[151,1437,1294,2006]" pageId="1" pageNumber="2">
<emphasis id="EBE9EAE33660FF97FF4BD401FE808070" bold="true" box="[151,297,1294,1319]" pageId="1" pageNumber="2">Description.</emphasis>
Length 10.4510.56 (mean = 10.50), maximum width 6.006.13 (mean = 6.06). Body elongate, parallel-sided, broadly rounded anteriorly and posteriorly, macropterous; widest across anterior half of embolia; dorsally head and pronotum light colored but heavily covered with brown punctures; scutellum and wings dark (
<figureCitation id="41A62A743660FF97FAB2D459FF0D80C5" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1769,1792]" captionTargetBox="[163,1411,754,1714]" captionTargetId="figure@2.[151,1436,735,1745]" captionTargetPageId="2" captionText="FIGURES 1 3. Features of Pogonocaudina, n. gen.: (1) habitus (dry specimen), (2) single segmented front tarsus with paired claws of male appressed to anterior margin of femur, (3) dense fringe of long setae on outer margin of posterior abdominal segments (specimen in alcohol)." httpUri="https://zenodo.org/record/203246/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
). Coarsely punctate throughout.
</paragraph>
<paragraph id="D92236F13660FF97FF19D4AEFAFB838E" blockId="1.[151,1437,1294,2006]" pageId="1" pageNumber="2">Head angled ventrad ca. 45o with respect to pronotum; compound eyes divergent, mesal and lateral margins slightly convergent anteriorly; lateral hyperoche widening anteriorly, anterior margin of eye subequal in length to head laterad of eye, black margin of head laterad and caudad of eye; series of four setal rosettes paralleling inner margin of eye; posterior margin of head slightly convex between eyes; apex of head deflexed, oriented posteriorly; broad, yellow, impunctate band along anterior margin immediately above deflexion; labrum broad, rounded; rostrum short, segment three (first visible anteriorly) partially concealed behind labrum, segment four narrower, reaching to near middle of prothoracic coxae; maxillary plates with brush of long hairs distally, hairs reaching apex of rostrum; antennae pale, segments one and two short and inconspicuous, three longest and gradually widened apically, four 3/4 length of three; antennae set behind brushes of long setae ventrally on head and proepimeron.</paragraph>
<paragraph id="D92236F13660FF94FF19D7E9FDAE87F0" blockId="1.[151,1437,1294,2006]" lastBlockId="2.[151,1436,151,680]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
Pronotum concave behind eyes; dark punctation heavy throughout, dark pigmentation diminishing laterally; short, recumbent setae throughout, more evident laterally in absence of dark pigmentation; lateral margins smooth, convex; posterolateral corners broadly rounded; posterior margin straight to slightly convex; incomplete transverse sulcus setting off posterior one fourth more evident laterad of midline; transverse depression in anterior third. Propleura meeting prosternum; prosternal midline with wedge-shaped carina between coxae. Scutellum large, granulate, raised above level of wings, without paired protuberances, posterolateral margins sinuate. Mesosternum with broad tumescence and two rows of pegs at midline. Metaxyphus broadly triangular and slightly acuminate posteriorly. Forelegs with femur broad, strongly widened posterobasally; tibia narrow; tarsus single-segmented in both sexes (
<figureCitation id="41A62A743663FF94FEFAD1B3FED98583" box="[294,368,188,212]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1769,1792]" captionTargetBox="[163,1411,754,1714]" captionTargetId="figure@2.[151,1436,735,1745]" captionTargetPageId="2" captionText="FIGURES 1 3. Features of Pogonocaudina, n. gen.: (1) habitus (dry specimen), (2) single segmented front tarsus with paired claws of male appressed to anterior margin of femur, (3) dense fringe of long setae on outer margin of posterior abdominal segments (specimen in alcohol)." httpUri="https://zenodo.org/record/203246/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
); paired, articulated claws in both sexes (
<figureCitation id="41A62A743663FF94FCB9D1B3FC198583" box="[869,944,188,212]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1769,1792]" captionTargetBox="[163,1411,754,1714]" captionTargetId="figure@2.[151,1436,735,1745]" captionTargetPageId="2" captionText="FIGURES 1 3. Features of Pogonocaudina, n. gen.: (1) habitus (dry specimen), (2) single segmented front tarsus with paired claws of male appressed to anterior margin of femur, (3) dense fringe of long setae on outer margin of posterior abdominal segments (specimen in alcohol)." httpUri="https://zenodo.org/record/203246/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
). Middle and hind femora flattened, with anteroventral and posteroventral sparse rows of short spines; middle femur slightly curved dorsad; tibiae with numerous stout spines. Middle tibia gradually widening apically; male with dense pad of hairs beginning at middle of ventral surface and widening distally, pad reduced on female; single row of posteroventral heavy spines, multiple rows of anterior heavy spines, row of posterodorsal swimming hairs.
<collectingCountry id="A18A76613663FF94FC1FD043FC528433" box="[963,1019,332,356]" name="India" pageId="2" pageNumber="3">Hind</collectingCountry>
leg with tibia elongate, thin, multiple rows of spines on anterior and posterior margin, posterodorsal swimming hairs on tibia becoming more sparse on tarsi; tarsomere one short and produced beneath base of tarsomere two, two long, three longer. Mid- and hind pretarsi with empodia; claws symmetrical, long, evenly curved, each with basal tooth. Meso- and metathoracic coxae strongly recessed within thoracic venter; coxae with ventromesal series of stout hairs. Hemelytra and scutellum finely punctate with short setae, coarsely granulate. Hemelytra with distinct clavus and embolium; clavus with longitudinal suture between claval suture and scutellum, claval commisure with longitudinal ridge; embolium with proximal brush of long hairs; costal fracture at level of anterior 1/3 of abdominal segment IV; membrane evident by absence of punctation and coarse granulation. Hindwing with well-developed veins C, R, M, Cu, m-Cu, pCu, pCu- A, 1A, 2A; cells R, M; anal fold.
</paragraph>
<caption id="8DE266793663FF94FF4BD7E6FE15826A" httpUri="https://zenodo.org/record/203246/files/figure.png" pageId="2" pageNumber="3" targetBox="[163,1411,754,1714]" targetPageId="2">
<paragraph id="D92236F13663FF94FF4BD7E6FE15826A" blockId="2.[151,1436,1769,1853]" pageId="2" pageNumber="3">
<emphasis id="EBE9EAE33663FF94FF4BD7E6FE9383A8" bold="true" box="[151,314,1769,1792]" pageId="2" pageNumber="3">FIGURES 13.</emphasis>
Features of
<taxonomicName id="1E9D4D723663FF94FE65D7E5FDF08257" box="[441,601,1770,1792]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33663FF94FE65D7E5FDF08257" box="[441,601,1770,1792]" italics="true" pageId="2" pageNumber="3">Pogonocaudina</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F0DA57983663FF94FDBAD7E5FD0783A8" box="[614,686,1770,1791]" pageId="2" pageNumber="3" rank="genus">
<emphasis id="EBE9EAE33663FF94FDBAD7E5FD0783A8" bold="true" box="[614,686,1770,1791]" pageId="2" pageNumber="3">n. gen.</emphasis>
</taxonomicNameLabel>
: (1) habitus (dry specimen), (2) single segmented front tarsus with paired claws of male appressed to anterior margin of femur, (3) dense fringe of long setae on outer margin of posterior abdominal segments (specimen in alcohol).
</paragraph>
</caption>
<paragraph id="D92236F13663FF95FF19D666FEF385A0" blockId="2.[151,1436,1897,2031]" lastBlockId="3.[151,1437,151,1148]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Abdomen dorsally dark brown; males with terga IIV entire, VVI dissociated into medio- and laterotergites, VII entire; all terga entire in females. Connexival margin II smooth, IIIIV set with stout peg-like setae, posterior ¼ of IV and entire length of VVIII with extremely long, dense fringe of brown hairs in both sexes (
<figureCitation id="41A62A743663FF94FB09D6BDFAB0829D" box="[1237,1305,1970,1994]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1769,1792]" captionTargetBox="[163,1411,754,1714]" captionTargetId="figure@2.[151,1436,735,1745]" captionTargetPageId="2" captionText="FIGURES 1 3. Features of Pogonocaudina, n. gen.: (1) habitus (dry specimen), (2) single segmented front tarsus with paired claws of male appressed to anterior margin of femur, (3) dense fringe of long setae on outer margin of posterior abdominal segments (specimen in alcohol)." httpUri="https://zenodo.org/record/203246/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
). Abdomen ventrally generally covered with fine, light colored setae; with glabrous yellowish lateral band; strong brush of midventral brown hairs beginning on III, widening and continuing to posterior end of abdomen. Spiracle on II at anterior margin, concealed beneath metepimeron; spiracles IIIVI evident between and slightly anterior to paired glabrous patches.
</paragraph>
</subSubSection>
<subSubSection id="9187657A3662FF95FF1BD00CFC768740" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="D92236F13662FF95FF1BD00CFC768740" blockId="3.[151,1437,151,1148]" pageId="3" pageNumber="4">
<emphasis id="EBE9EAE33662FF95FF1BD00CFEE8844B" bold="true" box="[199,321,259,284]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
The most striking unique attribute of this insect is the dense fringe of long hairs around the perimeter of the posterior abdominal segments in both sexes (
<figureCitation id="41A62A743662FF95FCD6D028FCE68468" box="[778,847,295,319]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1769,1792]" captionTargetBox="[163,1411,754,1714]" captionTargetId="figure@2.[151,1436,735,1745]" captionTargetPageId="2" captionText="FIGURES 1 3. Features of Pogonocaudina, n. gen.: (1) habitus (dry specimen), (2) single segmented front tarsus with paired claws of male appressed to anterior margin of femur, (3) dense fringe of long setae on outer margin of posterior abdominal segments (specimen in alcohol)." httpUri="https://zenodo.org/record/203246/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
). This genus can be recognized further by the male forelegs having one tarsal segment, which is similar to the condition of
<taxonomicName id="1E9D4D723662FF95FC6AD043FBFA8433" box="[950,1107,332,356]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33662FF95FC6AD043FBFA8433" box="[950,1107,332,356]" italics="true" pageId="3" pageNumber="4">Namtokocoris</emphasis>
</taxonomicName>
, but with two pretarsal claws (
<figureCitation id="41A62A743662FF95FF43D060FF4B84D0" box="[159,226,367,391]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1769,1792]" captionTargetBox="[163,1411,754,1714]" captionTargetId="figure@2.[151,1436,735,1745]" captionTargetPageId="2" captionText="FIGURES 1 3. Features of Pogonocaudina, n. gen.: (1) habitus (dry specimen), (2) single segmented front tarsus with paired claws of male appressed to anterior margin of femur, (3) dense fringe of long setae on outer margin of posterior abdominal segments (specimen in alcohol)." httpUri="https://zenodo.org/record/203246/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
) as in males of all other known
<taxonomicName id="1E9D4D723662FF95FD94D060FD4B84D0" box="[584,738,367,391]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
. The scutellum is elevated slightly above the level of the hemelytra as in
<taxonomicName id="1E9D4D723662FF95FEC1D09BFE1384FB" box="[285,442,404,428]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33662FF95FEC1D09BFE1384FB" box="[285,442,404,428]" italics="true" pageId="3" pageNumber="4">Namtokocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="1E9D4D723662FF95FE28D09BFD3184FB" box="[500,664,404,428]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33662FF95FE28D09BFD3184FB" box="[500,664,404,428]" italics="true" pageId="3" pageNumber="4">Diaphorocoris</emphasis>
</taxonomicName>
. However, unlike
<taxonomicName id="1E9D4D723662FF95FCB6D09BFBAE84FB" box="[874,1031,404,428]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33662FF95FCB6D09BFBAE84FB" box="[874,1031,404,428]" italics="true" pageId="3" pageNumber="4">Namtokocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="1E9D4D723662FF95FB9DD09BFB4C84FB" box="[1089,1253,404,428]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33662FF95FB9DD09BFB4C84FB" box="[1089,1253,404,428]" italics="true" pageId="3" pageNumber="4">Diaphorocoris</emphasis>
</taxonomicName>
, linear series of stout hairs on the hemelytra and scutellar protuberances are lacking. Other subfamily attributes of
<taxonomicName id="1E9D4D723662FF95FB01D0B8FADD8498" box="[1245,1396,439,463]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
are present, including a sexually dimorphic pad of hairs on the mesotibia, a dense midventral abdominal band of hairs, the front of the head folded posteroventrally, and a well-developed labrum.
</paragraph>
</subSubSection>
<subSubSection id="9187657A3662FF93FF1BD32CFE638312" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="discussion">
<paragraph id="D92236F13662FF95FF1BD32CFB01860B" blockId="3.[151,1437,151,1148]" pageId="3" pageNumber="4">
<emphasis id="EBE9EAE33662FF95FF1BD32CFEE5876B" bold="true" box="[199,332,547,572]" pageId="3" pageNumber="4">Discussion.</emphasis>
The front tarsal condition differs among the genera of Asian
<taxonomicName id="1E9D4D723662FF95FC21D32BFB31876B" box="[1021,1176,548,572]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
: Although most genera are sexually dimorphic, where the male has two tarsal segments and the female one, both sexes of
<emphasis id="EBE9EAE33662FF95FAD7D347FA358708" box="[1291,1436,584,607]" italics="true" pageId="3" pageNumber="4">Ctenipocoris</emphasis>
have two segments and both sexes of
<taxonomicName id="1E9D4D723662FF95FD95D363FD4F87D3" box="[585,742,620,644]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33662FF95FD95D363FD4F87D3" box="[585,742,620,644]" italics="true" pageId="3" pageNumber="4">Namtokocoris</emphasis>
</taxonomicName>
have one segment. Thus,
<taxonomicName id="1E9D4D723662FF95FBCFD363FB6C87D3" box="[1043,1221,620,644]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33662FF95FBCFD363FB6C87D3" box="[1043,1221,620,644]" italics="true" pageId="3" pageNumber="4">Pogonocaudina</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="1E9D4D723662FF95FA82D362FF5687FF" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33662FF95FA82D362FF5687FF" italics="true" pageId="3" pageNumber="4">Namtokocoris</emphasis>
</taxonomicName>
in the possession of single-segmented front tarsi in both sexes but differs in having paired pretarsal claws, among other features: a straight posterior margin of the pronotum instead of convex; no scutellar protuberances; mesotibia with a row of posterodorsal swimming hairs instead of a dorsal row of spines; the legs are proportionately longer and less spinose; the dense midventral band of hairs begins on III instead of V; the lack of a long row of erect hairs setting off the glabrous ventral lateral abdominal band; the female has mediosternite VI transverse instead of triangular and narrowly pointed; and the digitate lobe of laterosternite V is unique.
</paragraph>
<paragraph id="D92236F13662FF95FF1BD268FB7686F3" blockId="3.[151,1437,151,1148]" pageId="3" pageNumber="4">The secondary longitudinal suture within the clavus was observed in all specimens examined, and appears to correspond with macroptery. All specimens examined for hindwing condition were macropterous.</paragraph>
<paragraph id="D92236F13662FF95FF1BD2A0FD01812B" blockId="3.[151,1437,151,1148]" pageId="3" pageNumber="4">
Based on the character states in common with other members of the subfamily,
<taxonomicName id="1E9D4D723662FF95FBE0D2BFFB44869F" box="[1084,1261,944,968]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33662FF95FBE0D2BFFB44869F" box="[1084,1261,944,968]" italics="true" pageId="3" pageNumber="4">Pogonocaudina</emphasis>
</taxonomicName>
clearly appears to be most closely related to
<taxonomicName id="1E9D4D723662FF95FE07D2DBFDD186BB" box="[475,632,980,1004]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33662FF95FE07D2DBFDD186BB" box="[475,632,980,1004]" italics="true" pageId="3" pageNumber="4">Namtokocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="1E9D4D723662FF95FD6CD2DBFCFD86BB" box="[688,852,980,1004]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33662FF95FD6CD2DBFCFD86BB" box="[688,852,980,1004]" italics="true" pageId="3" pageNumber="4">Diaphorocoris</emphasis>
</taxonomicName>
. The latter genera previously were considered to be closely related (Sites &amp; Vitheepradit 2007), and
<taxonomicName id="1E9D4D723662FF95FD1DD2F7FCDA8147" box="[705,883,1016,1040]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33662FF95FD1DD2F7FCDA8147" box="[705,883,1016,1040]" italics="true" pageId="3" pageNumber="4">Pogonocaudina</emphasis>
</taxonomicName>
has features in common with each of them, and elevation of the scutellum and waterfall habitation with both of them (
<tableCitation id="941F034A3662FF95FC61D513FBBA8163" box="[957,1043,1052,1076]" captionStart="TABLE 1" captionStartId="3.[151,235,1192,1213]" captionTargetBox="[151,1426,1242,1822]" captionTargetPageId="3" captionText="TABLE 1. Character comparison of Pogonocaudina with other Asian genera of Laccocorinae." httpUri="http://table.plazi.org/id/8DE266793662FF95FF4BD5A7FBFF81E9" pageId="3" pageNumber="4" tableUuid="8DE266793662FF95FF4BD5A7FBFF81E9">Table 1</tableCitation>
). All three of these genera have a dense midventral hair band in common with
<emphasis id="EBE9EAE33662FF95FD41D54FFC878100" box="[669,814,1088,1111]" italics="true" pageId="3" pageNumber="4">Ctenipocoris</emphasis>
. Analysis including molecular data is needed to more effectively evaluate phylogenetic relationships.
</paragraph>
<caption id="8DE266793662FF95FF4BD5A7FBFF81E9" ID-Table-UUID="8DE266793662FF95FF4BD5A7FBFF81E9" box="[151,1110,1192,1214]" httpUri="http://table.plazi.org/id/8DE266793662FF95FF4BD5A7FBFF81E9" pageId="3" pageNumber="4" targetBox="[151,1426,1242,1822]" targetIsTable="true" targetPageId="3">
<paragraph id="D92236F13662FF95FF4BD5A7FBFF81E9" blockId="3.[151,1110,1192,1214]" box="[151,1110,1192,1214]" pageId="3" pageNumber="4">
<emphasis id="EBE9EAE33662FF95FF4BD5A7FF5781EA" bold="true" box="[151,254,1192,1213]" pageId="3" pageNumber="4">TABLE 1</emphasis>
. Character comparison of
<taxonomicName id="1E9D4D723662FF95FDD7D5A7FD0281E9" box="[523,683,1192,1214]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33662FF95FDD7D5A7FD0281E9" box="[523,683,1192,1214]" italics="true" pageId="3" pageNumber="4">Pogonocaudina</emphasis>
</taxonomicName>
with other Asian genera of
<taxonomicName id="1E9D4D723662FF95FC1AD5A7FBFB81E9" box="[966,1106,1192,1214]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="D92236F13662FF95FF43D5D5FAED8249" pageId="3" pageNumber="4">
<table id="AB9DC45136620069FF4BD5D5FA3B8249" box="[151,1426,1242,1822]" gridcols="6" gridrows="13" pageId="3" pageNumber="4">
<tr id="67AD34B336620069FF4BD5D5FA3B81A7" box="[151,1426,1242,1264]" gridrow="0" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD5D5FE4781A7" box="[151,494,1242,1264]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">Character</th>
<th id="247C5DCF36620069FDDAD5D5FD3D81A7" box="[518,660,1242,1264]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="1E9D4D723662FF95FDDAD5D5FD3D81A7" box="[518,660,1242,1264]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33662FF95FDDAD5D5FD3D81A7" box="[518,660,1242,1264]" italics="true" pageId="3" pageNumber="4">Namtokocoris</emphasis>
</taxonomicName>
</th>
<th id="247C5DCF36620069FD75D5D5FCE081A7" box="[681,841,1242,1264]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="1E9D4D723662FF95FD75D5D5FCE081A7" box="[681,841,1242,1264]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33662FF95FD75D5D5FCE081A7" box="[681,841,1242,1264]" italics="true" pageId="3" pageNumber="4">Pogonocaudina</emphasis>
</taxonomicName>
</th>
<th id="247C5DCF36620069FC83D5D5FB2E81A7" box="[863,1159,1242,1264]" gridcol="3" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="EBE9EAE33662FF95FC83D5D5FB2E81A7" box="[863,1159,1242,1264]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="1E9D4D723662FF95FC83D5D5FC5A81A7" box="[863,1011,1242,1264]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">Diaphorocoris</taxonomicName>
Ctenipocoris
</emphasis>
</th>
<th id="247C5DCF36620069FB41D5D5FAA781A7" box="[1181,1294,1242,1264]" gridcol="4" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="1E9D4D723662FF95FB41D5D4FAA781A7" box="[1181,1294,1243,1264]" class="Insecta" family="Naucoridae" genus="Laccocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33662FF95FB41D5D4FAA781A7" box="[1181,1294,1243,1264]" italics="true" pageId="3" pageNumber="4">Laccocoris</emphasis>
</taxonomicName>
</th>
<th id="247C5DCF36620069FAFFD5D5FA3B81A7" box="[1315,1426,1242,1264]" gridcol="5" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="1E9D4D723662FF95FAFFD5D5FA3B81A7" box="[1315,1426,1242,1264]" class="Insecta" family="Naucoridae" genus="Heleocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33662FF95FAFFD5D5FA3B81A7" box="[1315,1426,1242,1264]" italics="true" pageId="3" pageNumber="4">Heleocoris</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="67AD34B336620069FF4BD40CFA3B804D" box="[151,1426,1283,1306]" gridrow="1" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD40CFE47804D" box="[151,494,1283,1306]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">dense caudal fringe of hairs</th>
<td id="247C5DCF36620069FDDAD40CFD3D804D" box="[518,660,1283,1306]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">no</td>
<td id="247C5DCF36620069FD75D40CFCE0804D" box="[681,841,1283,1306]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FC83D40CFB2E804D" box="[863,1159,1283,1306]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">no no</td>
<td id="247C5DCF36620069FB41D40CFAA7804D" box="[1181,1294,1283,1306]" gridcol="4" gridrow="1" pageId="3" pageNumber="4">no</td>
<td id="247C5DCF36620069FAFFD40CFA3B804D" box="[1315,1426,1283,1306]" gridcol="5" gridrow="1" pageId="3" pageNumber="4">no</td>
</tr>
<tr id="67AD34B336620069FF4BD422FA3B8014" box="[151,1426,1325,1347]" gridrow="2" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD422FE478014" box="[151,494,1325,1347]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">male protarsal segments</th>
<td id="247C5DCF36620069FDDAD422FD3D8014" box="[518,660,1325,1347]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">1</td>
<td id="247C5DCF36620069FD75D422FCE08014" box="[681,841,1325,1347]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">1</td>
<td id="247C5DCF36620069FC83D422FB2E8014" box="[863,1159,1325,1347]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">2 2</td>
<td id="247C5DCF36620069FB41D422FAA78014" box="[1181,1294,1325,1347]" gridcol="4" gridrow="2" pageId="3" pageNumber="4">2</td>
<td id="247C5DCF36620069FAFFD422FA3B8014" box="[1315,1426,1325,1347]" gridcol="5" gridrow="2" pageId="3" pageNumber="4">2</td>
</tr>
<tr id="67AD34B336620069FF4BD459FA3B803B" box="[151,1426,1366,1388]" gridrow="3" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD459FE47803B" box="[151,494,1366,1388]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">pretarsal claws on foreleg</th>
<td id="247C5DCF36620069FDDAD459FD3D803B" box="[518,660,1366,1388]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">1</td>
<td id="247C5DCF36620069FD75D459FCE0803B" box="[681,841,1366,1388]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">2</td>
<td id="247C5DCF36620069FC83D459FB2E803B" box="[863,1159,1366,1388]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">2 2</td>
<td id="247C5DCF36620069FB41D459FAA7803B" box="[1181,1294,1366,1388]" gridcol="4" gridrow="3" pageId="3" pageNumber="4">2</td>
<td id="247C5DCF36620069FAFFD459FA3B803B" box="[1315,1426,1366,1388]" gridcol="5" gridrow="3" pageId="3" pageNumber="4">2</td>
</tr>
<tr id="67AD34B336620069FF4BD48FFA3B80C1" box="[151,1426,1408,1430]" gridrow="4" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD48FFE4780C1" box="[151,494,1408,1430]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">expansive, thin, prosternal carina</th>
<td id="247C5DCF36620069FDDAD48FFD3D80C1" box="[518,660,1408,1430]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FD75D48FFCE080C1" box="[681,841,1408,1430]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">no</td>
<td id="247C5DCF36620069FC83D48FFB2E80C1" box="[863,1159,1408,1430]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">no no</td>
<td id="247C5DCF36620069FB41D48FFAA780C1" box="[1181,1294,1408,1430]" gridcol="4" gridrow="4" pageId="3" pageNumber="4">no</td>
<td id="247C5DCF36620069FAFFD48FFA3B80C1" box="[1315,1426,1408,1430]" gridcol="5" gridrow="4" pageId="3" pageNumber="4">no</td>
</tr>
<tr id="67AD34B336620069FF4BD4A6FA3B80E8" box="[151,1426,1449,1471]" gridrow="5" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD4A6FE4780E8" box="[151,494,1449,1471]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">scutellar protuberances</th>
<td id="247C5DCF36620069FDDAD4A6FD3D80E8" box="[518,660,1449,1471]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">distinct</td>
<td id="247C5DCF36620069FD75D4A6FCE080E8" box="[681,841,1449,1471]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">no</td>
<td id="247C5DCF36620069FC83D4A6FB2E80E8" box="[863,1159,1449,1471]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">weak no</td>
<td id="247C5DCF36620069FB41D4A6FAA780E8" box="[1181,1294,1449,1471]" gridcol="4" gridrow="5" pageId="3" pageNumber="4">no</td>
<td id="247C5DCF36620069FAFFD4A6FA3B80E8" box="[1315,1426,1449,1471]" gridcol="5" gridrow="5" pageId="3" pageNumber="4">no</td>
</tr>
<tr id="67AD34B336620069FF4BD4DDFA3B80BF" box="[151,1426,1490,1512]" gridrow="6" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD4DDFE4780BF" box="[151,494,1490,1512]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">scutellum elevated</th>
<td id="247C5DCF36620069FDDAD4DDFD3D80BF" box="[518,660,1490,1512]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FD75D4DDFCE080BF" box="[681,841,1490,1512]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FC83D4DDFB2E80BF" box="[863,1159,1490,1512]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">yes no</td>
<td id="247C5DCF36620069FB41D4DDFAA780BF" box="[1181,1294,1490,1512]" gridcol="4" gridrow="6" pageId="3" pageNumber="4">no</td>
<td id="247C5DCF36620069FAFFD4DDFA3B80BF" box="[1315,1426,1490,1512]" gridcol="5" gridrow="6" pageId="3" pageNumber="4">no</td>
</tr>
<tr id="67AD34B336620069FF4BD4F4FA3B8345" box="[151,1426,1531,1554]" gridrow="7" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD4F4FE478345" box="[151,494,1531,1554]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">hair lines on hemelytra</th>
<td id="247C5DCF36620069FDDAD4F4FD3D8345" box="[518,660,1531,1554]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FD75D4F4FCE08345" box="[681,841,1531,1554]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">no</td>
<td id="247C5DCF36620069FC83D4F4FB2E8345" box="[863,1159,1531,1554]" gridcol="3" gridrow="7" pageId="3" pageNumber="4">yes no</td>
<td id="247C5DCF36620069FB41D4F4FAA78345" box="[1181,1294,1531,1554]" gridcol="4" gridrow="7" pageId="3" pageNumber="4">no</td>
<td id="247C5DCF36620069FAFFD4F4FA3B8345" box="[1315,1426,1531,1554]" gridcol="5" gridrow="7" pageId="3" pageNumber="4">no</td>
</tr>
<tr id="67AD34B336620069FF4BD72AFA3B836C" box="[151,1426,1573,1595]" gridrow="8" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD72AFE47836C" box="[151,494,1573,1595]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">dense midventral band of hairs</th>
<td id="247C5DCF36620069FDDAD72AFD3D836C" box="[518,660,1573,1595]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FD75D72AFCE0836C" box="[681,841,1573,1595]" gridcol="2" gridrow="8" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FC83D72AFB2E836C" box="[863,1159,1573,1595]" gridcol="3" gridrow="8" pageId="3" pageNumber="4">yes yes</td>
<td id="247C5DCF36620069FB41D72AFAA7836C" box="[1181,1294,1573,1595]" gridcol="4" gridrow="8" pageId="3" pageNumber="4">no</td>
<td id="247C5DCF36620069FAFFD72AFA3B836C" box="[1315,1426,1573,1595]" gridcol="5" gridrow="8" pageId="3" pageNumber="4">no</td>
</tr>
<tr id="67AD34B336620069FF4BD741FA3B8333" box="[151,1426,1614,1636]" gridrow="9" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="247C5DCF36620069FF4BD741FE478333" box="[151,494,1614,1636]" gridcol="0" gridrow="9" pageId="3" pageNumber="4">on abdominal sterna VVII/VIII</th>
</tr>
<tr id="67AD34B336620069FF4BD777FA3B83FB" box="[151,1426,1656,1708]" gridrow="10" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD777FE4783FB" box="[151,494,1656,1708]" gridcol="0" gridrow="10" pageId="3" pageNumber="4">dimorphic pad of hairs on mesotibia</th>
<td id="247C5DCF36620069FDDAD777FD3D83FB" box="[518,660,1656,1708]" gridcol="1" gridrow="10" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FD75D777FCE083FB" box="[681,841,1656,1708]" gridcol="2" gridrow="10" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FC83D777FB2E83FB" box="[863,1159,1656,1708]" gridcol="3" gridrow="10" pageId="3" pageNumber="4">yes yes</td>
<td id="247C5DCF36620069FB41D777FAA783FB" box="[1181,1294,1656,1708]" gridcol="4" gridrow="10" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FAFFD777FA3B83FB" box="[1315,1426,1656,1708]" gridcol="5" gridrow="10" pageId="3" pageNumber="4">yes</td>
</tr>
<tr id="67AD34B336620069FF4BD7CFFA3B83A3" box="[151,1426,1728,1780]" gridrow="11" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD7CFFE4783A3" box="[151,494,1728,1780]" gridcol="0" gridrow="11" pageId="3" pageNumber="4">front of head folded posteroven- trally</th>
<td id="247C5DCF36620069FDDAD7CFFD3D83A3" box="[518,660,1728,1780]" gridcol="1" gridrow="11" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FD75D7CFFCE083A3" box="[681,841,1728,1780]" gridcol="2" gridrow="11" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FC83D7CFFB2E83A3" box="[863,1159,1728,1780]" gridcol="3" gridrow="11" pageId="3" pageNumber="4">yes yes</td>
<td id="247C5DCF36620069FB41D7CFFAA783A3" box="[1181,1294,1728,1780]" gridcol="4" gridrow="11" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FAFFD7CFFA3B83A3" box="[1315,1426,1728,1780]" gridcol="5" gridrow="11" pageId="3" pageNumber="4">yes</td>
</tr>
<tr id="67AD34B336620069FF4BD607FA3B8249" box="[151,1426,1800,1822]" gridrow="12" pageId="3" pageNumber="4">
<th id="247C5DCF36620069FF4BD607FE478249" box="[151,494,1800,1822]" gridcol="0" gridrow="12" pageId="3" pageNumber="4">well-developed labrum</th>
<td id="247C5DCF36620069FDDAD607FD3D8249" box="[518,660,1800,1822]" gridcol="1" gridrow="12" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FD75D607FCE08249" box="[681,841,1800,1822]" gridcol="2" gridrow="12" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FC83D607FB2E8249" box="[863,1159,1800,1822]" gridcol="3" gridrow="12" pageId="3" pageNumber="4">yes yes</td>
<td id="247C5DCF36620069FB41D607FAA78249" box="[1181,1294,1800,1822]" gridcol="4" gridrow="12" pageId="3" pageNumber="4">yes</td>
<td id="247C5DCF36620069FAFFD607FA3B8249" box="[1315,1426,1800,1822]" gridcol="5" gridrow="12" pageId="3" pageNumber="4">yes</td>
</tr>
</table>
</paragraph>
<paragraph id="D92236F13662FF93FF19D66AFE988072" blockId="3.[151,1436,1893,2025]" lastBlockId="5.[151,1436,1149,1713]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
<emphasis id="EBE9EAE33662FF95FF19D66AFE25822A" bold="true" box="[197,396,1893,1918]" pageId="3" pageNumber="4">Ecological notes.</emphasis>
Specimens of
<taxonomicName id="1E9D4D723662FF95FDECD66AFD48822A" box="[560,737,1893,1917]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33662FF95FDECD66AFD48822A" box="[560,737,1893,1917]" italics="true" pageId="3" pageNumber="4">Pogonocaudina</emphasis>
</taxonomicName>
were observed on smooth surfaces of a granite waterfall at the
<typeStatus id="062688533662FF95FF4BD685FF6E82F5" box="[151,199,1930,1954]" pageId="3" pageNumber="4">type</typeStatus>
locality (
<figureCitation id="41A62A743662FF95FEEDD686FEDC82F6" box="[305,373,1929,1953]" captionStart="FIGURES 4 5" captionStartId="4.[151,264,1958,1981]" captionTargetBox="[218,1373,193,1938]" captionTargetId="figure@4.[218,1374,193,1939]" captionTargetPageId="4" captionText="FIGURES 4 5. (4) Kiliyur Waterfall, the type locality of Pogonocaudina indica n. sp. and Diaphorocoris kiliyur n. sp., (5) habitat of Pogonocaudina indica n. sp., Diaphorocoris arunachalami n. sp. and Diaphorocoris kiliyur n. sp." httpUri="https://zenodo.org/record/203247/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
), usually associated with a film of water and in contact with fallen leaves (
<figureCitation id="41A62A743662FF95FB6BD686FAAF82F6" box="[1207,1286,1929,1953]" captionStart="FIGURES 4 5" captionStartId="4.[151,264,1958,1981]" captionTargetBox="[218,1373,193,1938]" captionTargetId="figure@4.[218,1374,193,1939]" captionTargetPageId="4" captionText="FIGURES 4 5. (4) Kiliyur Waterfall, the type locality of Pogonocaudina indica n. sp. and Diaphorocoris kiliyur n. sp., (5) habitat of Pogonocaudina indica n. sp., Diaphorocoris arunachalami n. sp. and Diaphorocoris kiliyur n. sp." httpUri="https://zenodo.org/record/203247/files/figure.png" pageId="3" pageNumber="4">Figs. 5</figureCitation>
,
<figureCitation id="41A62A743662FF95FACDD686FA8982F6" box="[1297,1312,1929,1953]" captionStart="FIGURE 6" captionStartId="5.[151,250,1082,1105]" captionTargetBox="[151,1436,193,1060]" captionTargetId="figure@5.[151,1436,193,1061]" captionTargetPageId="5" captionText="FIGURE 6. Live specimen of Pogonocaudina indica n. sp." httpUri="https://zenodo.org/record/203248/files/figure.png" pageId="3" pageNumber="4">6</figureCitation>
) or at least on wet rock surfaces and algae. Some individuals were associated with small organic debris dams comprising sticks and leaves, where water trickled through but most water was held back in small pools; thus, some current was present at this microsite. Other insects were present on the rock surfaces of the waterfall, including mayflies, hydrophilids (including
<taxonomicName id="1E9D4D723664FF93FE79D5AEFDBE81EE" box="[421,535,1185,1209]" class="Insecta" family="Hydrophilidae" genus="Scoliopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33664FF93FE79D5AEFDBE81EE" box="[421,535,1185,1209]" italics="true" pageId="5" pageNumber="6">Scoliopsis</emphasis>
</taxonomicName>
), gerrids, and the naucorid
<taxonomicName id="1E9D4D723664FF93FC96D5AEFC4781EE" box="[842,1006,1185,1209]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33664FF93FC96D5AEFC4781EE" box="[842,1006,1185,1209]" italics="true" pageId="5" pageNumber="6">Diaphorocoris</emphasis>
</taxonomicName>
. Detailed accounts of waterfall-inhabiting insects in
<collectingCountry id="A18A76613664FF93FE9CD5CAFE0A818A" box="[320,419,1221,1245]" name="Thailand" pageId="5" pageNumber="6">Thailand</collectingCountry>
and elsewhere in Indochina were given for
<taxonomicName id="1E9D4D723664FF93FC57D5CAFB07818A" box="[907,1198,1221,1245]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33664FF93FC57D5CAFC5A818A" box="[907,1011,1221,1245]" italics="true" pageId="5" pageNumber="6">Oocyclus</emphasis>
(Hydrophilidae)
</taxonomicName>
by
<bibRefCitation id="BD0C4B003664FF93FB0BD5CAFF4D8056" author="Short" pageId="5" pageNumber="17" refString="Short, A. E. Z. &amp; Swanson, A. P. (2005) A revision of the Oocyclus Sharp of Thailand with description of six new species (Coleoptera: Hydrophilidae). Zootaxa, 1078, 1 - 24." type="journal article" year="2005">Short &amp; Swanson (2005)</bibRefCitation>
,
<taxonomicName id="1E9D4D723664FF93FF2DD5E6FE768056" box="[241,479,1257,1281]" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33664FF93FF2DD5E6FEC88056" box="[241,353,1257,1281]" italics="true" pageId="5" pageNumber="6">Eotrechus</emphasis>
(Gerridae)
</taxonomicName>
by Vitheepradit &amp; Sites (2007), and
<taxonomicName id="1E9D4D723664FF93FC5AD5E6FB168056" box="[902,1215,1257,1281]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33664FF93FC5AD5E6FB8A8056" box="[902,1059,1257,1281]" italics="true" pageId="5" pageNumber="6">Namtokocoris</emphasis>
(Naucoridae)
</taxonomicName>
by Sites &amp; Vitheepradit (2007).
</paragraph>
<caption id="8DE266793665FF92FF4BD6A9FB4D828D" httpUri="https://zenodo.org/record/203247/files/figure.png" pageId="4" pageNumber="5" targetBox="[218,1373,193,1938]" targetPageId="4">
<paragraph id="D92236F13665FF92FF4BD6A9FB4D828D" blockId="4.[151,1436,1958,2011]" pageId="4" pageNumber="5">
<emphasis id="EBE9EAE33665FF92FF4BD6A9FE9582EB" bold="true" box="[151,316,1958,1981]" pageId="4" pageNumber="5">FIGURES 45.</emphasis>
(4)
<taxonomicName id="1E9D4D723665FF92FEB5D6A9FE1B82EB" box="[361,434,1958,1980]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="16" phylum="Arthropoda" rank="species" species="kiliyur" status="sp. nov.">Kiliyur</taxonomicName>
Waterfall, the type locality of
<taxonomicName id="1E9D4D723665FF92FD2DD6A8FC7182EA" box="[753,984,1959,1981]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="EBE9EAE33665FF92FD2DD6A8FC7182EA" box="[753,984,1959,1981]" italics="true" pageId="4" pageNumber="5">Pogonocaudina indica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0DA57983665FF92FC3CD6A8FBB282EB" box="[992,1051,1959,1980]" pageId="4" pageNumber="5" rank="species">
<emphasis id="EBE9EAE33665FF92FC3CD6A8FBB282EB" bold="true" box="[992,1051,1959,1980]" pageId="4" pageNumber="5">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="1E9D4D723665FF92FB93D6A8FA8782EA" box="[1103,1326,1959,1981]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kiliyur" status="sp. nov.">
<emphasis id="EBE9EAE33665FF92FB93D6A8FA8782EA" box="[1103,1326,1959,1981]" italics="true" pageId="4" pageNumber="5">Diaphorocoris kiliyur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0DA57983665FF92FAE9D6A8FAD982EB" box="[1333,1392,1959,1980]" pageId="4" pageNumber="5" rank="species">
<emphasis id="EBE9EAE33665FF92FAE9D6A8FAD982EB" bold="true" box="[1333,1392,1959,1980]" pageId="4" pageNumber="5">n. sp.</emphasis>
</taxonomicNameLabel>
, (5) habitat of
<taxonomicName id="1E9D4D723665FF92FF21D6CAFE4A828C" box="[253,483,1989,2011]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="EBE9EAE33665FF92FF21D6CAFE4A828C" box="[253,483,1989,2011]" italics="true" pageId="4" pageNumber="5">Pogonocaudina indica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0DA57983665FF92FE36D6CAFD8A828D" box="[490,547,1989,2010]" pageId="4" pageNumber="5" rank="species">
<emphasis id="EBE9EAE33665FF92FE36D6CAFD8A828D" bold="true" box="[490,547,1989,2010]" pageId="4" pageNumber="5">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="1E9D4D723665FF92FDF3D6CAFCF1828C" box="[559,856,1989,2011]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="arunachalami" status="sp. nov.">
<emphasis id="EBE9EAE33665FF92FDF3D6CAFCF1828C" box="[559,856,1989,2011]" italics="true" pageId="4" pageNumber="5">Diaphorocoris arunachalami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0DA57983665FF92FC82D6CAFC3E828D" box="[862,919,1989,2010]" pageId="4" pageNumber="5" rank="species">
<emphasis id="EBE9EAE33665FF92FC82D6CAFC3E828D" bold="true" box="[862,919,1989,2010]" pageId="4" pageNumber="5">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="1E9D4D723665FF92FC14D6CAFB0C828C" box="[968,1189,1989,2011]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kiliyur" status="sp. nov.">
<emphasis id="EBE9EAE33665FF92FC14D6CAFB0C828C" box="[968,1189,1989,2011]" italics="true" pageId="4" pageNumber="5">Diaphorocoris kiliyur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0DA57983665FF92FB77D6CAFB4D828D" box="[1195,1252,1989,2010]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="8DE266793664FF93FF4BD535FD5D8107" box="[151,756,1082,1105]" httpUri="https://zenodo.org/record/203248/files/figure.png" pageId="5" pageNumber="6" targetBox="[151,1436,193,1060]" targetPageId="5">
<paragraph id="D92236F13664FF93FF4BD535FD5D8107" blockId="5.[151,756,1082,1105]" box="[151,756,1082,1105]" pageId="5" pageNumber="6">
<emphasis id="EBE9EAE33664FF93FF4BD535FEBB8107" bold="true" box="[151,274,1082,1105]" pageId="5" pageNumber="6">FIGURE 6.</emphasis>
Live specimen of
<taxonomicName id="1E9D4D723664FF93FE13D535FD1D8107" box="[463,692,1082,1104]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="EBE9EAE33664FF93FE13D535FD1D8107" box="[463,692,1082,1104]" italics="true" pageId="5" pageNumber="6">Pogonocaudina indica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0DA57983664FF93FD67D534FD5D8107" box="[699,756,1083,1104]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="D92236F13664FF93FF1BD43EFE638312" blockId="5.[151,1436,1149,1713]" pageId="5" pageNumber="6">
The discovery of
<taxonomicName id="1E9D4D723664FF93FE50D43EFD94801E" box="[396,573,1329,1353]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33664FF93FE50D43EFD94801E" box="[396,573,1329,1353]" italics="true" pageId="5" pageNumber="6">Pogonocaudina</emphasis>
</taxonomicName>
adds to a growing number of naucorid genera known to occur exclusively or facultatively in waterfalls. In Indochina,
<taxonomicName id="1E9D4D723664FF93FD82D45AFD52803A" box="[606,763,1365,1389]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33664FF93FD82D45AFD52803A" box="[606,763,1365,1389]" italics="true" pageId="5" pageNumber="6">Namtokocoris</emphasis>
</taxonomicName>
is restricted to waterfalls, although individuals occasionally drift downstream (Sites &amp; Vitheepradit 2007). Many nymphs of
<taxonomicName id="1E9D4D723664FF93FCAFD476FC5480C6" box="[883,1021,1401,1425]" class="Insecta" family="Naucoridae" genus="Cheirochela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="EBE9EAE33664FF93FCAFD476FC5480C6" box="[883,1021,1401,1425]" italics="true" pageId="5" pageNumber="6">Cheirochela</emphasis>
</taxonomicName>
have been collected on waterfalls in
<collectingCountry id="A18A76613664FF93FF4BD492FF5280E2" box="[151,251,1437,1461]" name="Thailand" pageId="5" pageNumber="6">Thailand</collectingCountry>
and
<collectingCountry id="A18A76613664FF93FEEBD492FE3080E2" box="[311,409,1437,1461]" name="Vietnam" pageId="5" pageNumber="6">Vietnam</collectingCountry>
, although adults have been collected on large rocks in fast current (Sites &amp; Vitheepradit 2007). In
<collectingCountry id="A18A76613664FF93FEDAD4CEFEEB808E" box="[262,322,1473,1497]" name="India" pageId="5" pageNumber="6">India</collectingCountry>
,
<taxonomicName id="1E9D4D723664FF93FE92D4CEFE5B808E" box="[334,498,1473,1497]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33664FF93FE92D4CEFE5B808E" box="[334,498,1473,1497]" italics="true" pageId="5" pageNumber="6">Diaphorocoris</emphasis>
</taxonomicName>
was found on wet rock surfaces of waterfalls, including with
<taxonomicName id="1E9D4D723664FF93FB70D4CEFAF4808E" box="[1196,1373,1473,1497]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBE9EAE33664FF93FB70D4CEFAF4808E" box="[1196,1373,1473,1497]" italics="true" pageId="5" pageNumber="6">Pogonocaudina</emphasis>
</taxonomicName>
at its
<typeStatus id="062688533664FF93FF4BD4E9FF6E80A9" box="[151,199,1510,1534]" pageId="5" pageNumber="6">type</typeStatus>
locality. From
<collectingCountry id="A18A76613664FF93FEA9D4EAFE6280AA" box="[373,459,1509,1533]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
to Central
<collectingCountry id="A18A76613664FF93FD97D4EAFD0280AA" box="[587,683,1509,1533]" name="United States of America" pageId="5" pageNumber="6">America</collectingCountry>
in the New World,
<emphasis id="EBE9EAE33664FF93FC50D4E9FB9C80AA" box="[908,1077,1510,1533]" italics="true" pageId="5" pageNumber="6">Cataractocoris</emphasis>
can be found in waterfall habitats, including in sheeting water on vertical rock surfaces, the waterfall splash zone, and fast current in turbulent, rocky streams (
<bibRefCitation id="BD0C4B003664FF93FE9ED722FE178312" author="Sites" box="[322,446,1581,1605]" pageId="5" pageNumber="17" refString="Sites, R. W. (2004) Distributional records of Cataractocoris (Heteroptera: Naucoridae) in Mesoamerica. Entomological News, 115, 173 - 174." type="journal article" year="2004">Sites 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9187657A3664FF93FF1BD75EFB7B83DA" pageId="5" pageNumber="6" type="etymology">
<paragraph id="D92236F13664FF93FF1BD75EFB7B83DA" blockId="5.[151,1436,1149,1713]" pageId="5" pageNumber="6">
<emphasis id="EBE9EAE33664FF93FF1BD75EFEE7833E" bold="true" box="[199,334,1617,1641]" pageId="5" pageNumber="6">Etymology.</emphasis>
This genus is named for the dense fringe of long hairs around the perimeter of the posterior abdominal segments. &quot;Pogono-&quot; is Greek for beard, and &quot;caud-&quot; is Latin for tail or posterior end.
</paragraph>
</subSubSection>
<subSubSection id="9187657A3664FF93FF1BD797FD0283E6" box="[199,683,1688,1713]" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="D92236F13664FF93FF1BD797FD0283E6" blockId="5.[151,1436,1149,1713]" box="[199,683,1688,1713]" pageId="5" pageNumber="6">
<emphasis id="EBE9EAE33664FF93FF1BD797FE1983E6" bold="true" box="[199,432,1688,1713]" pageId="5" pageNumber="6">Material examined.</emphasis>
See species localities.
</paragraph>
</subSubSection>
</treatment>
</document>