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<document id="073E0AB9C406E1007EC3885CC632B947" ID-DOI="10.26107/RBZ-2019-0034" ID-ISSN="2345-7600" ID-Zenodo-Dep="4576271" ID-ZooBank="4537754B-0BBE-4388-84E1-5A51DCD507AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614777929123" checkinUser="carolina" docAuthor="Okanishi, Masanori, Oba, Yuichi &amp; Fujita, Yoshihisa" docDate="2019" docId="511E87A92A084F6CFE91905920D8F78B" docLanguage="en" docName="RafflesBZool.67.421-439.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Ophiomora elegans Koehler 1907" docType="treatment" docVersion="4" lastPageNumber="436" masterDocId="AD27FFD12A064F63FF949365212FFF94" masterDocTitle="Brittle stars from a submarine cave of Christmas Island, northwestern Australia, with description of a new bioluminescent species Ophiopsila xmasilluminans (Echinodermata: Ophiuroidea) and notes on its behaviour" masterLastPageNumber="439" masterPageNumber="421" pageNumber="435" updateTime="1698878073412" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="25DFA5A167AC6953AA22B9204AB2CDFE">Brittle stars from a submarine cave of Christmas Island, northwestern Australia, with description of a new bioluminescent species Ophiopsila xmasilluminans (Echinodermata: Ophiuroidea) and notes on its behaviour</mods:title>
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<mods:namePart id="E6A7A7C23074FF91002A7C97091AF172">Okanishi, Masanori</mods:namePart>
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<mods:namePart id="D621B02460B7EC32D0F26916537AE9BB">Oba, Yuichi</mods:namePart>
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<treatment id="511E87A92A084F6CFE91905920D8F78B" ID-DOI="http://doi.org/10.5281/zenodo.4575948" ID-Zenodo-Dep="4575948" LSID="urn:lsid:plazi:treatment:511E87A92A084F6CFE91905920D8F78B" httpUri="http://treatment.plazi.org/id/511E87A92A084F6CFE91905920D8F78B" lastPageId="15" lastPageNumber="436" pageId="14" pageNumber="435">
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<taxonomicName id="1EB74D3C2A084F6DFE91905923A5FCC0" authority="Koehler, 1907" authorityName="Koehler" authorityYear="1907" box="[261,650,828,853]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiomora" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="435" phylum="Echinodermata" rank="species" species="elegans">
<emphasis id="EBC3EAAD2A084F6DFE91905923A5FCC0" bold="true" box="[261,650,828,853]" pageId="14" pageNumber="435">
<emphasis id="EBC3EAAD2A084F6DFE91905920F0FCC0" bold="true" box="[261,479,828,852]" italics="true" pageId="14" pageNumber="435">Ophiomora elegans</emphasis>
<bibRefCitation id="BD264B4E2A084F6DFE7D905823A5FCC0" author="Koehler R" box="[489,650,828,853]" pageId="14" pageNumber="435" pagination="279 - 351" refId="ref9688" refString="Koehler R (1907) Revision de la collection des Ophiures du Museum d'histoire Naturelle Paris. Bulletin Scientifique de La Franca et de La Belgique, 41: 279 - 351." type="journal article" year="1907">Koehler, 1907</bibRefCitation>
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<paragraph id="D90836BF2A084F6DFECB90392301FCE1" blockId="14.[261,650,828,885]" box="[351,558,860,885]" pageId="14" pageNumber="435">
(
<figureCitation id="418C2A3A2A084F6DFEF3903920FBFCE1" box="[359,468,860,885]" captionStart="Fig" captionStartId="13.[813,847,1163,1185]" captionTargetBox="[813,1452,188,1148]" captionTargetId="figure-443@13.[813,1452,188,1148]" captionTargetPageId="13" captionText="Fig.11. Aboral views ofOphioconis claviculata (RUMF-ZE-00155) (A) and Ophiomora elegans (RUMF-ZE-00142) (B), anesthetised living specimens. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576304" httpUri="https://zenodo.org/record/4576304/files/figure.png" pageId="14" pageNumber="435">Figs. 11B</figureCitation>
,
<figureCitation id="418C2A3A2A084F6DFE75903820D1FCE1" box="[481,510,861,885]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">12</figureCitation>
,
<figureCitation id="418C2A3A2A084F6DFD9F90382309FCE1" box="[523,550,861,885]" captionStart="Fig" captionStartId="16.[135,170,2006,2028]" captionTargetBox="[151,1456,190,1990]" captionTargetId="figure-82@16.[148,1456,190,1990]" captionTargetPageId="16" captionText="Fig. 13. Ophiomora elegans (RUMF-ZE-00142). A, aboral proximal portion of the arm; B, aboral distal portion of the arm; C, oral proximal portion of the arm; D, oral middle portion of the arm; E, oro-lateral distal portion of the arm. F, lateral proximal portion of the arm; G, lateral middle portion of the arm. Abbreviations: AS, arm spine; LAP, lateral arm plate; TS, tentacle scale; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576310" httpUri="https://zenodo.org/record/4576310/files/figure.png" pageId="14" pageNumber="435">13</figureCitation>
)
</paragraph>
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<treatmentCitation id="581610AE2A084F6DFF1390FE2321FC24" author="Koehler R" box="[135,526,923,945]" page="342" pageId="14" pageNumber="435" year="1907">
<taxonomicName id="1EB74D3C2A084F6DFF1390FE2321FC24" authority="Koehler, 1907: 342" authorityName="Koehler" authorityPageNumber="342" authorityYear="1907" box="[135,526,923,945]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiomora" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="435" phylum="Echinodermata" rank="species" species="elegans">
<emphasis id="EBC3EAAD2A084F6DFF1390FE2069FC24" box="[135,326,923,944]" italics="true" pageId="14" pageNumber="435">Ophiomora elegans</emphasis>
<bibRefCitation id="BD264B4E2A084F6DFEDB90FE2321FC24" author="Koehler R" box="[335,526,923,945]" pageId="14" pageNumber="435" pagination="279 - 351" refId="ref9688" refString="Koehler R (1907) Revision de la collection des Ophiures du Museum d'histoire Naturelle Paris. Bulletin Scientifique de La Franca et de La Belgique, 41: 279 - 351." type="journal article" year="1907">Koehler, 1907: 342</bibRefCitation>
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, 343, pl. 14; H. L.
<treatmentCitation id="581610AE2A084F6DFD5890FE203DFC59" author="Clark HL" page="174" pageId="14" pageNumber="435" year="1915">
<bibRefCitation id="BD264B4E2A084F6DFD5890FE203DFC59" author="Clark HL" pageId="14" pageNumber="435" pagination="163 - 376" refId="ref9126" refString="Clark HL (1915) Catalogue of recent ophiurans: based on the collection of the Museum of Comparative Zoology. Memoirs of the Museum of Comparative Zoology at Havard College, 25 (4): 163 - 376." type="journal article" year="1915">Clark, 1915: 174</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="581610AE2A084F6DFE8A90DD20C0FC59" author="Tan HH &amp; Naruse T &amp; Fujita Y &amp; Kiat TS" box="[286,495,952,974]" page="413" pageId="14" pageNumber="435" year="2014">
<bibRefCitation id="BD264B4E2A084F6DFE8A90DD20C0FC59" author="Tan HH &amp; Naruse T &amp; Fujita Y &amp; Kiat TS" box="[286,495,952,974]" pageId="14" pageNumber="435" pagination="406 - 418" refId="ref10651" refString="Tan HH, Naruse T, Fujita Y &amp; Kiat TS (2014) Observations on the fauna from submarine and associated anchialine caves in Christmas Island, Indian Ocean Territory, Australia. Raffles Bulletin of Zoology, 7600 (30): 406 - 418." type="journal article" year="2014">Tan et al., 2014: 413</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="581610AE2A084F6DFE6E90DD23FEFC5A" author="Fujita Y &amp; Irimura S &amp; Kogure Y &amp; Okanishi M &amp; Francois M" box="[506,721,952,974]" page="50" pageId="14" pageNumber="435" year="2015">
<bibRefCitation id="BD264B4E2A084F6DFE6E90DD23FEFC5A" author="Fujita Y &amp; Irimura S &amp; Kogure Y &amp; Okanishi M &amp; Francois M" box="[506,721,952,974]" pageId="14" pageNumber="435" pagination="1 - 106" refId="ref9376" refString="Fujita Y, Irimura S, Kogure Y, Okanishi M &amp; Francois M (2015) Catalogue of Echinodermata specimens deposited in the University Museum (Fujukan), University of the Ryukyus. In: Catalogue of materials deposited in the University Museum (Fujukan), University of the Ryukyus. University Museum (Fujukan), University of the Ryukyus, Okinawa. Pp. 1 - 106. [In Japanese]" type="book chapter" year="2015">Fujita et al., 2015: 50</bibRefCitation>
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, 97.
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</paragraph>
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<subSubSection id="91AD65342A084F6DFF139093203EFB1A" pageId="14" pageNumber="435" type="materials_examined">
<paragraph id="D90836BF2A084F6DFF139093203EFB1A" blockId="14.[135,775,1014,1166]" pageId="14" pageNumber="435">
<emphasis id="EBC3EAAD2A084F6DFF139093205DFB9A" bold="true" box="[135,370,1014,1038]" pageId="14" pageNumber="435">Material examined.</emphasis>
<materialsCitation id="69DF3CE22A084F6DFEE890932022FB1A" collectingDate="2011-03-27" collectionCode="SCUBA" collectorName="Tan Heok Hui &amp; Partly" country="Christmas Island" elevation="10" location="Thunderdome Cave" pageId="14" pageNumber="435" specimenCount="1">
RUMF-ZE-00142, submarine cave “
<location id="DC6860642A084F6DFF059773205CFBBA" LSID="urn:lsid:plazi:treatment:511E87A92A084F6CFE91905920D8F78B:DC6860642A084F6DFF059773205CFBBA" box="[145,371,1046,1070]" country="Christmas Island" name="Thunderdome Cave" pageId="14" pageNumber="435">Thunderdome Cave</location>
” at
<collectingCountry id="A1A0762F2A084F6DFE3297732348FBBA" box="[422,615,1046,1070]" name="Christmas Island" pageId="14" pageNumber="435">Christmas Island</collectingCountry>
, northwestern
<collectingCountry id="A1A0762F2A084F6DFF13975321C3FBDA" box="[135,236,1078,1102]" name="Australia" pageId="14" pageNumber="435">Australia</collectingCountry>
, depth approximately
<quantity id="1E4F9B5A2A084F6DFE4997532338FBDA" box="[477,535,1078,1102]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="435" unit="m" value="10.0">
<elevation id="529AD18C2A084F6DFE4997532338FBDA" box="[477,535,1078,1102]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="435" unit="m" value="10.0">10 m</elevation>
</quantity>
,
<collectionCode id="BFA6AE7A2A084F6DFDB797532353FBDA" box="[547,636,1078,1102]" pageId="14" pageNumber="435">SCUBA</collectionCode>
diving, coll.
<collectorName id="744253692A084F6DFF1D97322007FBFA" box="[137,296,1110,1134]" pageId="14" pageNumber="435">Tan Heok Hui</collectorName>
,
<date id="AD09107F2A084F6DFEA0973320F0FBFA" box="[308,479,1110,1134]" pageId="14" pageNumber="435" value="2011-03-27">
<collectingDate id="BD4DE9972A084F6DFEA0973320F0FBFA" box="[308,479,1110,1134]" pageId="14" pageNumber="435" value="2011-03-27">27 March 2011</collectingDate>
</date>
.
<collectorName id="744253692A084F6DFE7E97332305FBFA" box="[490,554,1110,1134]" pageId="14" pageNumber="435">Partly</collectorName>
bleached to remove integuments
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="91AD65342A084F6DFF1397D3232CFB5B" box="[135,515,1206,1231]" pageId="14" pageNumber="435" type="diagnosis">
<paragraph id="D90836BF2A084F6DFF1397D3232CFB5B" blockId="14.[135,515,1206,1231]" box="[135,515,1206,1231]" pageId="14" pageNumber="435">
<emphasis id="EBC3EAAD2A084F6DFF1397D321D3FB5A" bold="true" box="[135,252,1206,1230]" pageId="14" pageNumber="435">Diagnosis.</emphasis>
The same as the genus.
</paragraph>
</subSubSection>
<subSubSection id="91AD65342A084F6DFF139793246CFAC0" pageId="14" pageNumber="435" type="description">
<paragraph id="D90836BF2A084F6DFF13979320C2F7BA" blockId="14.[134,774,1270,2094]" pageId="14" pageNumber="435">
<emphasis id="EBC3EAAD2A084F6DFF1397932229FA9A" bold="true" box="[135,774,1270,1294]" pageId="14" pageNumber="435">Description of external morphology (RUMF-ZE-00142).</emphasis>
<emphasis id="EBC3EAAD2A084F6DFF1C967221EFFABA" box="[136,192,1303,1326]" italics="true" pageId="14" pageNumber="435">Disc.</emphasis>
Circular,
<quantity id="1E4F9B5A2A084F6DFEA496732052FABA" box="[304,381,1302,1326]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="14" pageNumber="435" unit="mm" value="22.0">22 mm</quantity>
in diameter, covered by thick skin, embedding small granules, approximately 200400 μm in length (
<figureCitation id="418C2A3A2A084F6DFF4196332014FAFA" box="[213,315,1366,1390]" captionStart="Fig" captionStartId="13.[813,847,1163,1185]" captionTargetBox="[813,1452,188,1148]" captionTargetId="figure-443@13.[813,1452,188,1148]" captionTargetPageId="13" captionText="Fig.11. Aboral views ofOphioconis claviculata (RUMF-ZE-00155) (A) and Ophiomora elegans (RUMF-ZE-00142) (B), anesthetised living specimens. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576304" httpUri="https://zenodo.org/record/4576304/files/figure.png" pageId="14" pageNumber="435">Figs. 11B</figureCitation>
,
<figureCitation id="418C2A3A2A084F6DFED196332059FAFA" box="[325,374,1366,1390]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">12A</figureCitation>
). On aboral surface, removing the skin, radial shields exposing, bar-like, 3 to 4 times longer than wide, almost reaching to disc center (
<figureCitation id="418C2A3A2A084F6DFE7496F3236BFA3A" box="[480,580,1430,1454]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12B</figureCitation>
). Stout polygonal plates also exposing and forming a single transverse row on lateral disc margin (
<figureCitation id="418C2A3A2A084F6DFECD96B32096FA7A" box="[345,441,1494,1518]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12B</figureCitation>
). On oral surface, removing the skin, adoral shields trapezoid, wider than long (
<figureCitation id="418C2A3A2A084F6DFD00969323D6F99A" box="[660,761,1526,1550]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12D</figureCitation>
), approximately
<quantity id="1E4F9B5A2A084F6DFEB39573204BF9BA" box="[295,356,1558,1582]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="435" unit="mm" value="1.0">1 mm</quantity>
at outer edge and 700 μm at inner edge in length,
<quantity id="1E4F9B5A2A084F6DFF6495532003F9DA" box="[240,300,1590,1614]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="435" unit="mm" value="1.0">1 mm</quantity>
in width, in contact with the first ventral arm plates (
<figureCitation id="418C2A3A2A084F6DFF4095332018F9FA" box="[212,311,1622,1646]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12D</figureCitation>
). Oral plates thin, approximately 800 μm in length, 400 μm in width, contacting each other. Oral shields triangular, approximately as long as wide (
<figureCitation id="418C2A3A2A084F6DFDDC95F323E8F93A" box="[584,711,1686,1710]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12C, D</figureCitation>
). One circular oral shield serving as madreporite and having at least three pores (
<figureCitation id="418C2A3A2A084F6DFE9F95B32042F97A" box="[267,365,1750,1774]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12C</figureCitation>
). Interradial oral disc covered only by skin (
<figureCitation id="418C2A3A2A084F6DFF5095932007F89A" box="[196,296,1782,1806]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12C</figureCitation>
). Genital slits small, along between 3rd and 4th arm segments (
<figureCitation id="418C2A3A2A084F6DFEC2947320EFF8BA" box="[342,448,1814,1838]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Figs. 12F</figureCitation>
,
<figureCitation id="418C2A3A2A084F6DFE58947320D4F8BA" box="[460,507,1814,1838]" captionStart="Fig" captionStartId="16.[135,170,2006,2028]" captionTargetBox="[151,1456,190,1990]" captionTargetId="figure-82@16.[148,1456,190,1990]" captionTargetPageId="16" captionText="Fig. 13. Ophiomora elegans (RUMF-ZE-00142). A, aboral proximal portion of the arm; B, aboral distal portion of the arm; C, oral proximal portion of the arm; D, oral middle portion of the arm; E, oro-lateral distal portion of the arm. F, lateral proximal portion of the arm; G, lateral middle portion of the arm. Abbreviations: AS, arm spine; LAP, lateral arm plate; TS, tentacle scale; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576310" httpUri="https://zenodo.org/record/4576310/files/figure.png" pageId="14" pageNumber="435">13B</figureCitation>
), approximately 200 μm in length (
<figureCitation id="418C2A3A2A084F6DFF6294532075F8DA" box="[246,346,1846,1870]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12C</figureCitation>
). Abradial genital plates visible on side of genital slit, bar-like, approximately
<quantity id="1E4F9B5A2A084F6DFDB69433235CF8FA" box="[546,627,1878,1902]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="435" unit="mm" value="1.5">1.5 mm</quantity>
in length and
<quantity id="1E4F9B5A2A084F6DFF13941321F5F81A" box="[135,218,1910,1934]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="435" unit="mm" value="0.3">0.3 mm</quantity>
in width (
<figureCitation id="418C2A3A2A084F6DFEDA9413209FF81A" box="[334,432,1910,1934]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12E</figureCitation>
). Thorny granules inside of the genital slit, near the abradial genital plates (
<figureCitation id="418C2A3A2A084F6DFDCB94F323EFF83A" box="[607,704,1942,1966]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12E</figureCitation>
). Four flat, subequal trapezoidal oral papillae at each adoral shield horizontally (
<figureCitation id="418C2A3A2A084F6DFE8394B320B4F87A" box="[279,411,2006,2030]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12C, D</figureCitation>
). Teeth triangular, forming single vertical row on dental plate (
<figureCitation id="418C2A3A2A084F6DFE2194932337F79A" box="[437,536,2038,2062]" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Fig. 12D</figureCitation>
). Second tentacle pore completely inside the mouth slit.
</paragraph>
<paragraph id="D90836BF2A084F6DFCD793D8250DFEA1" blockId="14.[835,1477,188,309]" pageId="14" pageNumber="435">
<emphasis id="EBC3EAAD2A084F6DFCD793D822ACFF40" box="[835,899,189,212]" italics="true" pageId="14" pageNumber="435">Arms.</emphasis>
Five, all arms entire almost entire length,
<quantity id="1E4F9B5A2A084F6DFADA93D924EDFF40" box="[1358,1474,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="14" pageNumber="435" unit="mm" value="62.5" valueMax="75.0" valueMin="50.0">5075 mm</quantity>
long. Arms approximately 4.0 mm wide and 2.0 mm high, flatten. Arms tapering and flatting gradually distally (
<figureCitation id="418C2A3A2A084F6DFA1A9399225DFEA1" captionStart="Fig" captionStartId="13.[813,847,1163,1185]" captionTargetBox="[813,1452,188,1148]" captionTargetId="figure-443@13.[813,1452,188,1148]" captionTargetPageId="13" captionText="Fig.11. Aboral views ofOphioconis claviculata (RUMF-ZE-00155) (A) and Ophiomora elegans (RUMF-ZE-00142) (B), anesthetised living specimens. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576304" httpUri="https://zenodo.org/record/4576304/files/figure.png" pageId="14" pageNumber="435">Figs. 11B</figureCitation>
,
<figureCitation id="418C2A3A2A084F6DFCEB927822FFFEA0" box="[895,976,285,309]" captionStart="Fig" captionStartId="16.[135,170,2006,2028]" captionTargetBox="[151,1456,190,1990]" captionTargetId="figure-82@16.[148,1456,190,1990]" captionTargetPageId="16" captionText="Fig. 13. Ophiomora elegans (RUMF-ZE-00142). A, aboral proximal portion of the arm; B, aboral distal portion of the arm; C, oral proximal portion of the arm; D, oral middle portion of the arm; E, oro-lateral distal portion of the arm. F, lateral proximal portion of the arm; G, lateral middle portion of the arm. Abbreviations: AS, arm spine; LAP, lateral arm plate; TS, tentacle scale; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576310" httpUri="https://zenodo.org/record/4576310/files/figure.png" pageId="14" pageNumber="435">13A, B</figureCitation>
, DG).
</paragraph>
<paragraph id="D90836BF2A084F6DFCD7923922F6FBA1" blockId="14.[834,1477,348,1077]" pageId="14" pageNumber="435">
Throughout the arm, dorsal arm plates absent and small fragmented plates, approximately 300500 μm in length on aboral side (
<figureCitation id="418C2A3A2A084F6DFC5E92F92501FE21" box="[970,1070,412,437]" captionStart="Fig" captionStartId="16.[135,170,2006,2028]" captionTargetBox="[151,1456,190,1990]" captionTargetId="figure-82@16.[148,1456,190,1990]" captionTargetPageId="16" captionText="Fig. 13. Ophiomora elegans (RUMF-ZE-00142). A, aboral proximal portion of the arm; B, aboral distal portion of the arm; C, oral proximal portion of the arm; D, oral middle portion of the arm; E, oro-lateral distal portion of the arm. F, lateral proximal portion of the arm; G, lateral middle portion of the arm. Abbreviations: AS, arm spine; LAP, lateral arm plate; TS, tentacle scale; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576310" httpUri="https://zenodo.org/record/4576310/files/figure.png" pageId="14" pageNumber="435">Fig. 13A</figureCitation>
). On distal portion of the arm, lateral arm plates contacting on mid line each other (
<figureCitation id="418C2A3A2A084F6DFAB992D924A1FE41" box="[1325,1422,444,469]" captionStart="Fig" captionStartId="16.[135,170,2006,2028]" captionTargetBox="[151,1456,190,1990]" captionTargetId="figure-82@16.[148,1456,190,1990]" captionTargetPageId="16" captionText="Fig. 13. Ophiomora elegans (RUMF-ZE-00142). A, aboral proximal portion of the arm; B, aboral distal portion of the arm; C, oral proximal portion of the arm; D, oral middle portion of the arm; E, oro-lateral distal portion of the arm. F, lateral proximal portion of the arm; G, lateral middle portion of the arm. Abbreviations: AS, arm spine; LAP, lateral arm plate; TS, tentacle scale; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576310" httpUri="https://zenodo.org/record/4576310/files/figure.png" pageId="14" pageNumber="435">Fig. 13B</figureCitation>
). On oral side, removing the skin, ventral arm plates polygonal, slightly wider than long, lateral sides concave, contiguous on proximal portion of the arm (
<figureCitation id="418C2A3A2A084F6DFBE8917925F1FDA1" box="[1148,1246,540,565]" captionStart="Fig" captionStartId="16.[135,170,2006,2028]" captionTargetBox="[151,1456,190,1990]" captionTargetId="figure-82@16.[148,1456,190,1990]" captionTargetPageId="16" captionText="Fig. 13. Ophiomora elegans (RUMF-ZE-00142). A, aboral proximal portion of the arm; B, aboral distal portion of the arm; C, oral proximal portion of the arm; D, oral middle portion of the arm; E, oro-lateral distal portion of the arm. F, lateral proximal portion of the arm; G, lateral middle portion of the arm. Abbreviations: AS, arm spine; LAP, lateral arm plate; TS, tentacle scale; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576310" httpUri="https://zenodo.org/record/4576310/files/figure.png" pageId="14" pageNumber="435">Fig. 13C</figureCitation>
). On middle to distal portion of the arm, ventral arm plates triangular, pointing to proximal portion, and separated by each other (
<figureCitation id="418C2A3A2A084F6DFA009139225AFD01" captionStart="Fig" captionStartId="16.[135,170,2006,2028]" captionTargetBox="[151,1456,190,1990]" captionTargetId="figure-82@16.[148,1456,190,1990]" captionTargetPageId="16" captionText="Fig. 13. Ophiomora elegans (RUMF-ZE-00142). A, aboral proximal portion of the arm; B, aboral distal portion of the arm; C, oral proximal portion of the arm; D, oral middle portion of the arm; E, oro-lateral distal portion of the arm. F, lateral proximal portion of the arm; G, lateral middle portion of the arm. Abbreviations: AS, arm spine; LAP, lateral arm plate; TS, tentacle scale; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576310" httpUri="https://zenodo.org/record/4576310/files/figure.png" pageId="14" pageNumber="435">Fig. 13D</figureCitation>
). Lateral arm plates bearing arm spines, on proximal portion, 3 or 4 spines, long and pointed, and aboral-most spine approximately 1.5 times as long as corresponding arm segment, and other spines as long as corresponding arm segment (
<figureCitation id="418C2A3A2A084F6DFC7791992545FC81" box="[995,1130,764,789]" captionStart="Fig" captionStartId="16.[135,170,2006,2028]" captionTargetBox="[151,1456,190,1990]" captionTargetId="figure-82@16.[148,1456,190,1990]" captionTargetPageId="16" captionText="Fig. 13. Ophiomora elegans (RUMF-ZE-00142). A, aboral proximal portion of the arm; B, aboral distal portion of the arm; C, oral proximal portion of the arm; D, oral middle portion of the arm; E, oro-lateral distal portion of the arm. F, lateral proximal portion of the arm; G, lateral middle portion of the arm. Abbreviations: AS, arm spine; LAP, lateral arm plate; TS, tentacle scale; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576310" httpUri="https://zenodo.org/record/4576310/files/figure.png" pageId="14" pageNumber="435">Fig. 13D, F</figureCitation>
). On middle portion of arm, 3 subequal long and pointed spines, approximately as long as the corresponding arm segment (
<figureCitation id="418C2A3A2A084F6DFB4F9059246BFCC1" box="[1243,1348,828,853]" captionStart="Fig" captionStartId="16.[135,170,2006,2028]" captionTargetBox="[151,1456,190,1990]" captionTargetId="figure-82@16.[148,1456,190,1990]" captionTargetPageId="16" captionText="Fig. 13. Ophiomora elegans (RUMF-ZE-00142). A, aboral proximal portion of the arm; B, aboral distal portion of the arm; C, oral proximal portion of the arm; D, oral middle portion of the arm; E, oro-lateral distal portion of the arm. F, lateral proximal portion of the arm; G, lateral middle portion of the arm. Abbreviations: AS, arm spine; LAP, lateral arm plate; TS, tentacle scale; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576310" httpUri="https://zenodo.org/record/4576310/files/figure.png" pageId="14" pageNumber="435">Fig. 13G</figureCitation>
). On distal portion of arms, 2 subequal cylindrical and oval arm spines, approximately half to one-thirds length of corresponding arm segment (
<figureCitation id="418C2A3A2A084F6DFC7290F92563FC21" box="[998,1100,924,949]" captionStart="Fig" captionStartId="16.[135,170,2006,2028]" captionTargetBox="[151,1456,190,1990]" captionTargetId="figure-82@16.[148,1456,190,1990]" captionTargetPageId="16" captionText="Fig. 13. Ophiomora elegans (RUMF-ZE-00142). A, aboral proximal portion of the arm; B, aboral distal portion of the arm; C, oral proximal portion of the arm; D, oral middle portion of the arm; E, oro-lateral distal portion of the arm. F, lateral proximal portion of the arm; G, lateral middle portion of the arm. Abbreviations: AS, arm spine; LAP, lateral arm plate; TS, tentacle scale; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576310" httpUri="https://zenodo.org/record/4576310/files/figure.png" pageId="14" pageNumber="435">Fig. 13E</figureCitation>
). Tentacle scales 1, flat and fanshaped, two-thirds to the same length of the corresponding arm segment, and outer one half to one-fourth length of the corresponding arm segment throughout the arms (
<figureCitation id="418C2A3A2A084F6DFA1990992240FBA1" captionStart="Fig" captionStartId="15.[113,148,1575,1597]" captionTargetBox="[113,1451,191,1558]" captionTargetId="figure-11@15.[111,1454,186,1559]" captionTargetPageId="15" captionText="Fig. 12. Ophiomora elegans (RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="14" pageNumber="435">Figs. 12F</figureCitation>
,
<figureCitation id="418C2A3A2A084F6DFCE8977822E2FBA1" box="[892,973,1053,1077]" captionStart="Fig" captionStartId="16.[135,170,2006,2028]" captionTargetBox="[151,1456,190,1990]" captionTargetId="figure-82@16.[148,1456,190,1990]" captionTargetPageId="16" captionText="Fig. 13. Ophiomora elegans (RUMF-ZE-00142). A, aboral proximal portion of the arm; B, aboral distal portion of the arm; C, oral proximal portion of the arm; D, oral middle portion of the arm; E, oro-lateral distal portion of the arm. F, lateral proximal portion of the arm; G, lateral middle portion of the arm. Abbreviations: AS, arm spine; LAP, lateral arm plate; TS, tentacle scale; VAP, ventral arm plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576310" httpUri="https://zenodo.org/record/4576310/files/figure.png" pageId="14" pageNumber="435">13D, E</figureCitation>
).
</paragraph>
<paragraph id="D90836BF2A084F6DFCD797392484FA80" blockId="14.[835,1475,1116,1300]" pageId="14" pageNumber="435">
<emphasis id="EBC3EAAD2A084F6DFCD7973922CFFBE0" bold="true" box="[835,992,1116,1140]" pageId="14" pageNumber="435">Colour in life.</emphasis>
Body basic creamy white. On aboral surface, dense orange small spots appear except proximal portion of the arm where the spots are sparsely scattered (
<figureCitation id="418C2A3A2A084F6DFAC697F92499FB21" box="[1362,1462,1180,1205]" captionStart="Fig" captionStartId="13.[813,847,1163,1185]" captionTargetBox="[813,1452,188,1148]" captionTargetId="figure-443@13.[813,1452,188,1148]" captionTargetPageId="13" captionText="Fig.11. Aboral views ofOphioconis claviculata (RUMF-ZE-00155) (A) and Ophiomora elegans (RUMF-ZE-00142) (B), anesthetised living specimens. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576304" httpUri="https://zenodo.org/record/4576304/files/figure.png" pageId="14" pageNumber="435">Fig. 11B</figureCitation>
). Deep purple bands present at intervals of 6 to 8 arm segments. Oral disc also creamy white with the deep purple bands on arms continuous from aboral surface (photo not shown).
</paragraph>
<paragraph id="D90836BF2A084F6DFCD79659246CFAC0" blockId="14.[835,1347,1340,1365]" box="[835,1347,1340,1365]" pageId="14" pageNumber="435">
<emphasis id="EBC3EAAD2A084F6DFCD796592544FAC1" bold="true" box="[835,1131,1340,1365]" pageId="14" pageNumber="435">Common Japanese name.</emphasis>
Gama-kumohitode.
</paragraph>
</subSubSection>
<subSubSection id="91AD65342A084F6DFCD7961922F2F9E1" pageId="14" pageNumber="435" type="distribution">
<paragraph id="D90836BF2A084F6DFCD7961922F2F9E1" blockId="14.[835,1474,1404,1653]" pageId="14" pageNumber="435">
<emphasis id="EBC3EAAD2A084F6DFCD7961922F9FA00" bold="true" box="[835,982,1404,1428]" pageId="14" pageNumber="435">Distribution.</emphasis>
Known from the
<typeStatus id="060C881D2A084F6DFB09961825E4FA01" box="[1181,1227,1405,1429]" pageId="14" pageNumber="435">type</typeStatus>
locality, the Fernando Veloso, near Inhaca Island, southern
<collectingCountry id="A1A0762F2A084F6DFB2396F92491FA20" box="[1207,1470,1436,1460]" name="Mozambique" pageId="14" pageNumber="435">Republic of Mozambique</collectingCountry>
, eastern coast of Africa (
<bibRefCitation id="BD264B4E2A084F6DFBDF96D925C4FA41" author="Koehler R" box="[1099,1259,1468,1493]" pageId="14" pageNumber="435" pagination="279 - 351" refId="ref9688" refString="Koehler R (1907) Revision de la collection des Ophiures du Museum d'histoire Naturelle Paris. Bulletin Scientifique de La Franca et de La Belgique, 41: 279 - 351." type="journal article" year="1907">Koehler, 1907</bibRefCitation>
); Kumejima Island, the Ryukyu Islands, southeastern
<collectingCountry id="A1A0762F2A084F6DFB2B96B825D0FA60" box="[1215,1279,1501,1524]" name="Japan" pageId="14" pageNumber="435">Japan</collectingCountry>
, submarine cave, approximately
<quantity id="1E4F9B5A2A084F6DFC739698250EF980" box="[999,1057,1533,1557]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="435" unit="m" value="18.0">18 m</quantity>
depth (
<bibRefCitation id="BD264B4E2A084F6DFBE29699246EF980" author="Fujita Y &amp; Irimura S &amp; Kogure Y &amp; Okanishi M &amp; Francois M" box="[1142,1345,1532,1556]" pageId="14" pageNumber="435" pagination="1 - 106" refId="ref9376" refString="Fujita Y, Irimura S, Kogure Y, Okanishi M &amp; Francois M (2015) Catalogue of Echinodermata specimens deposited in the University Museum (Fujukan), University of the Ryukyus. In: Catalogue of materials deposited in the University Museum (Fujukan), University of the Ryukyus. University Museum (Fujukan), University of the Ryukyus, Okinawa. Pp. 1 - 106. [In Japanese]" type="book chapter" year="2015">Fujita et al., 2015</bibRefCitation>
);
<collectingCountry id="A1A0762F2A084F6DFAC2969922A7F9A0" name="Christmas Island" pageId="14" pageNumber="435">Christmas Island</collectingCountry>
, northwestern
<collectingCountry id="A1A0762F2A084F6DFBBA957925BBF9A0" box="[1070,1172,1564,1588]" name="Australia" pageId="14" pageNumber="435">Australia</collectingCountry>
, submarine cave known as “Thunderdome Cave”, approximately
<quantity id="1E4F9B5A2A084F6DFB6B95582414F9C0" box="[1279,1339,1597,1621]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="435" unit="m" value="10.0">10 m</quantity>
depth (this study,
<figureCitation id="418C2A3A2A084F6DFC19953922E2F9E1" box="[909,973,1628,1653]" captionStart="Fig" captionStartId="1.[112,147,1168,1190]" captionTargetBox="[115,1450,187,1153]" captionTargetId="figure-395@1.[101,1451,177,1163]" captionTargetPageId="1" captionText="Fig. 1. Sampling location of three ophiuroid species examined in this study." figureDoi="http://doi.org/10.5281/zenodo.4576273" httpUri="https://zenodo.org/record/4576273/files/figure.png" pageId="14" pageNumber="435">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="91AD65342A084F6CFCD795F820D8F78B" lastPageId="15" lastPageNumber="436" pageId="14" pageNumber="435" type="discussion">
<paragraph id="D90836BF2A084F6CFCD795F823DEF8AB" blockId="14.[835,1475,1692,2100]" lastBlockId="15.[113,753,1767,1856]" lastPageId="15" lastPageNumber="436" pageId="14" pageNumber="435">
<emphasis id="EBC3EAAD2A084F6DFCD795F82298F921" bold="true" box="[835,951,1693,1717]" pageId="14" pageNumber="435">Remarks.</emphasis>
In original description, Koehlers examined single specimen had a disc diameter of
<quantity id="1E4F9B5A2A084F6DFB6E95D82465F940" box="[1274,1354,1725,1749]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="435" unit="mm" value="13.0">13 mm</quantity>
, almost
<quantity id="1E4F9B5A2A084F6DFA3395D82242F960" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="435" unit="mm" value="10.0">10 mm</quantity>
smaller than our examined specimen (
<quantity id="1E4F9B5A2A084F6DFA8A95B82440F960" box="[1310,1391,1757,1781]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="14" pageNumber="435" unit="mm" value="22.0">22 mm</quantity>
in disc diameter).
<bibRefCitation id="BD264B4E2A084F6DFC2F9599254BF881" author="Koehler R" box="[955,1124,1788,1813]" pageId="14" pageNumber="435" pagination="279 - 351" refId="ref9688" refString="Koehler R (1907) Revision de la collection des Ophiures du Museum d'histoire Naturelle Paris. Bulletin Scientifique de La Franca et de La Belgique, 41: 279 - 351." type="journal article" year="1907">Koehler (1907)</bibRefCitation>
distinguished
<taxonomicName id="1EB74D3C2A084F6DFA9C959924ABF880" authorityName="Koehler" authorityYear="1907" box="[1288,1412,1788,1812]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiomora" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="435" phylum="Echinodermata" rank="genus">
<emphasis id="EBC3EAAD2A084F6DFA9C959924ABF880" box="[1288,1412,1788,1812]" italics="true" pageId="14" pageNumber="435">Ophiomora</emphasis>
</taxonomicName>
from other genera in the
<taxonomicName id="1EB74D3C2A084F6DFB8994792593F8A0" authorityName="Ljungman" authorityYear="1867" box="[1053,1212,1820,1844]" class="Ophiuroidea" family="Ophiomyxidae" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="435" phylum="Echinodermata" rank="family">Ophiomyxidae</taxonomicName>
in having small genital slits with carrying few calcareous ossicles beside them and one tentacle scale on each tentacle pore (
<bibRefCitation id="BD264B4E2A084F6DFA8094392497F8E1" author="Koehler R" box="[1300,1464,1884,1909]" pageId="14" pageNumber="435" pagination="279 - 351" refId="ref9688" refString="Koehler R (1907) Revision de la collection des Ophiures du Museum d'histoire Naturelle Paris. Bulletin Scientifique de La Franca et de La Belgique, 41: 279 - 351." type="journal article" year="1907">Koehler, 1907</bibRefCitation>
). These characters are also recognised in our specimen (
<figureCitation id="418C2A3A2A084F6DFA03941922BEF821" captionStart="Fig" captionStartId="13.[813,847,1163,1185]" captionTargetBox="[813,1452,188,1148]" captionTargetId="figure-443@13.[813,1452,188,1148]" captionTargetPageId="13" captionText="Fig.11. Aboral views ofOphioconis claviculata (RUMF-ZE-00155) (A) and Ophiomora elegans (RUMF-ZE-00142) (B), anesthetised living specimens. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4576304" httpUri="https://zenodo.org/record/4576304/files/figure.png" pageId="14" pageNumber="435">Fig. 11E, F</figureCitation>
). Considering the distinct characters, although this genus is now
<emphasis id="EBC3EAAD2A084F6DFC7094D825AFF840" box="[996,1152,1980,2004]" italics="true" pageId="14" pageNumber="435">incertae sedis</emphasis>
in the order
<taxonomicName id="1EB74D3C2A084F6DFA8F94D92491F840" box="[1307,1470,1980,2004]" class="Ophiuroidea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="435" phylum="Echinodermata" rank="order">Ophiacanthida</taxonomicName>
, we maintain
<taxonomicName id="1EB74D3C2A084F6DFC4394B9257BF860" authorityName="Koehler" authorityYear="1907" box="[983,1108,2012,2036]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiomora" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="435" phylum="Echinodermata" rank="genus">
<emphasis id="EBC3EAAD2A084F6DFC4394B9257BF860" box="[983,1108,2012,2036]" italics="true" pageId="14" pageNumber="435">Ophiomora</emphasis>
</taxonomicName>
as a valid genus. Our extensive second description of this species revealed that this species has radial shields, which were stated in the original
<bibRefCitation id="BD264B4E2A084F6CFAF99B792194F96B" author="Koehler R" lastPageId="15" lastPageNumber="436" pageId="14" pageNumber="435" pagination="279 - 351" refId="ref9688" refString="Koehler R (1907) Revision de la collection des Ophiures du Museum d'histoire Naturelle Paris. Bulletin Scientifique de La Franca et de La Belgique, 41: 279 - 351." type="journal article" year="1907">Koehler (1907)</bibRefCitation>
as absent. However, of course, to determine the precise taxonomic position of this species, reexamination of the single
<typeStatus id="060C881D2A094F6CFF4B944D2022F8D4" box="[223,269,1832,1856]" pageId="15" pageNumber="436">type</typeStatus>
specimen of
<taxonomicName id="1EB74D3C2A094F6CFE0A9442235EF8AB" authorityName="Koehler" authorityYear="1907" box="[414,625,1831,1855]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiomora" kingdom="Animalia" order="Ophiurida" pageId="15" pageNumber="436" phylum="Echinodermata" rank="species" species="elegans">
<emphasis id="EBC3EAAD2A094F6CFE0A9442235EF8AB" box="[414,625,1831,1855]" italics="true" pageId="15" pageNumber="436">Ophiomora elegans</emphasis>
</taxonomicName>
is required.
</paragraph>
<caption id="8DC866372A094F6CFFE59542201CF93B" ID-DOI="http://doi.org/10.5281/zenodo.4576306" ID-Zenodo-Dep="4576306" httpUri="https://zenodo.org/record/4576306/files/figure.png" pageId="15" pageNumber="436" startId="15.[113,148,1575,1597]" targetBox="[113,1451,191,1558]" targetPageId="15">
<paragraph id="D90836BF2A094F6CFFE59542201CF93B" blockId="15.[113,1454,1575,1712]" pageId="15" pageNumber="436">
Fig. 12.
<taxonomicName id="1EB74D3C2A094F6CFF50954220ACF9A8" authorityName="Koehler" authorityYear="1907" box="[196,387,1575,1596]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiomora" kingdom="Animalia" order="Ophiurida" pageId="15" pageNumber="436" phylum="Echinodermata" rank="species" species="elegans">
<emphasis id="EBC3EAAD2A094F6CFF50954220ACF9A8" box="[196,387,1575,1596]" italics="true" pageId="15" pageNumber="436">Ophiomora elegans</emphasis>
</taxonomicName>
(RUMF-ZE-00142). A, aboral periphery part of the disc; B, aboral periphery part of the disc, skin removed; C, oral disc, skin partly removed; D, jaws, skin partly removed; E, interradial oral disc, genital slit, skin partly removed; F, genital slit and proximal portion of the arm. Abbreviations: AbGP, abradial genital plate; AdS, adradial shield; GS, genital slit; MP, marginal plate; OS, oral shield; OP, oral papillae; OrP, oral plate; T, teeth; TG, thorny granule; TS, tentacle scale; Te, tentacle; VAP, ventral arm plate. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="D90836BF2A094F6CFFE5940220D8F78B" blockId="15.[113,752,1895,2079]" pageId="15" pageNumber="436">
The sampling locality of the present specimen (
<collectingCountry id="A1A0762F2A094F6CFD169402219AF80B" name="Christmas Island" pageId="15" pageNumber="436">Christmas Island</collectingCountry>
, northwestern
<collectingCountry id="A1A0762F2A094F6CFEC294E22095F80B" box="[342,442,1927,1951]" name="Australia" pageId="15" pageNumber="436">Australia</collectingCountry>
) is located midway between the
<typeStatus id="060C881D2A094F6CFF0894CD21E5F854" box="[156,202,1960,1984]" pageId="15" pageNumber="436">type</typeStatus>
locality (East Africa) and the recent record from
<collectingCountry id="A1A0762F2A094F6CFFE594AD219CF84B" box="[113,179,1992,2015]" name="Japan" pageId="15" pageNumber="436">Japan</collectingCountry>
. This geographical evidence indicates this species is widely distributed in the Indo-Western Pacific region (
<bibRefCitation id="BD264B4E2A094F6CFFED9B622037F78B" author="Koehler R" box="[121,280,2055,2079]" pageId="15" pageNumber="436" pagination="279 - 351" refId="ref9688" refString="Koehler R (1907) Revision de la collection des Ophiures du Museum d'histoire Naturelle Paris. Bulletin Scientifique de La Franca et de La Belgique, 41: 279 - 351." type="journal article" year="1907">Koehler, 1907</bibRefCitation>
;
<bibRefCitation id="BD264B4E2A094F6CFEB19B6220C4F78B" author="Fujita Y &amp; Irimura S &amp; Kogure Y &amp; Okanishi M &amp; Francois M" box="[293,491,2055,2079]" pageId="15" pageNumber="436" pagination="1 - 106" refId="ref9376" refString="Fujita Y, Irimura S, Kogure Y, Okanishi M &amp; Francois M (2015) Catalogue of Echinodermata specimens deposited in the University Museum (Fujukan), University of the Ryukyus. In: Catalogue of materials deposited in the University Museum (Fujukan), University of the Ryukyus. University Museum (Fujukan), University of the Ryukyus, Okinawa. Pp. 1 - 106. [In Japanese]" type="book chapter" year="2015">Fujita et al., 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>