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<document id="42CE62C79BEAF9F07CBFAD0CBFCD244A" ID-CLB-Dataset="6214" ID-DOI="10.11646/zootaxa.4974.3.2" ID-GBIF-Dataset="5b4eaeb3-5722-4021-84ad-125ccd80855f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4777839" ID-ZooBank="1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621588618725" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="4772526A004DFFC7FF182D8F11883229" docLanguage="en" docName="zootaxa.4974.3.2.pdf" docOrigin="Zootaxa 4974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus biexcavatus Perkins 2021, new species" docType="treatment" docVersion="6" lastPageNumber="467" masterDocId="BB4B2A12004AFFCFFF8F2D18151D3030" masterDocTitle="Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="503" masterPageNumber="459" pageNumber="466" updateTime="1698940277359" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="981C119B966935C67470E68958172CB4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="624D0E09FB33BA6100F9BDEA185FF764">
<mods:title id="39B4CF833CB5DDEF8646BD4896301C88">Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
<mods:name id="EA4644EC55569CA1A0E325000B8BB5EE" type="personal">
<mods:role id="CA75D92CB83081FD61877336F5696781">
<mods:roleTerm id="65417EB2B10291FFA2AB2C66542635D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="760A1E9FFDFED1AC95710BCE280AF19A">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="BBFBDEA3674AAED25E2DFCE711676286">text</mods:typeOfResource>
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<mods:title id="16A7680013E56F5FE6E3523972EBDF0A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="96D132255E5637D5D848EB995607929A">
<mods:date id="7479D1544490CE3BF418FE22CD7CDD4B">2021</mods:date>
<mods:detail id="306E79B9278885365E1B9CDFCEE18D20" type="pubDate">
<mods:number id="B3716D4EC050AE43F936F1D337EED113">2021-05-21</mods:number>
</mods:detail>
<mods:detail id="A47CE17015DFE2D1463BC67B5E8239FC" type="volume">
<mods:number id="58E7577BAB6C7E2CA061A4BD83D20500">4974</mods:number>
</mods:detail>
<mods:detail id="44D1F1E23FCCC2A39852EFFC6EB13CE5" type="issue">
<mods:number id="91323D8E67545C60A1E7E8253633BC0D">3</mods:number>
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<mods:extent id="E4885D872334F4D8E41122E07319DCFC" unit="page">
<mods:start id="7A0085215A4F51DFBD8BA93CA3CEB3AB">459</mods:start>
<mods:end id="5A8E3334C0F40BA97C4892838BE5FE05">503</mods:end>
</mods:extent>
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<mods:classification id="EA5BAF244E0E3D5DD8C0A717BD559658">journal article</mods:classification>
<mods:identifier id="0467DEB3D5AEB3FA0186506C46F1169B" type="CLB-Dataset">6214</mods:identifier>
<mods:identifier id="8D66A9E8C1FD07A866DCDD94B8CA87BE" type="DOI">10.11646/zootaxa.4974.3.2</mods:identifier>
<mods:identifier id="BE1EC8852CF5BBC9011FEDF84C8A6C7B" type="GBIF-Dataset">5b4eaeb3-5722-4021-84ad-125ccd80855f</mods:identifier>
<mods:identifier id="64D002D3CE7EFA528040A61604EA3354" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0478A5C54BC5BF50ADAF2C7604CD1FBB" type="Zenodo-Dep">4777839</mods:identifier>
<mods:identifier id="07F328B575E851984D69CF3033B30683" type="ZooBank">1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8</mods:identifier>
</mods:mods>
<treatment id="4772526A004DFFC7FF182D8F11883229" ID-DOI="http://doi.org/10.5281/zenodo.4777853" ID-GBIF-Taxon="182642562" ID-Zenodo-Dep="4777853" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229" httpUri="http://treatment.plazi.org/id/4772526A004DFFC7FF182D8F11883229" lastPageId="8" lastPageNumber="467" pageId="7" pageNumber="466">
<subSubSection id="87C1B0F7004DFFC8FF182D8F174D3081" box="[151,592,151,178]" pageId="7" pageNumber="466" type="nomenclature">
<paragraph id="CF64E37C004DFFC8FF182D8F174D3081" blockId="7.[151,592,151,214]" box="[151,592,151,178]" pageId="7" pageNumber="466">
<heading id="942C5410004DFFC8FF182D8F174D3081" bold="true" box="[151,592,151,178]" fontSize="11" level="1" pageId="7" pageNumber="466" reason="1">
<emphasis id="FDAF3F6E004DFFC8FF182D8F174D3081" bold="true" box="[151,592,151,178]" pageId="7" pageNumber="466">
<taxonomicName id="08DB98FF004DFFC8FF182D8F14AF3082" ID-CoL="8Y4PD" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,434,151,178]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="466" phylum="Arthropoda" rank="species" species="biexcavatus" status="sp. nov.">
<emphasis id="FDAF3F6E004DFFC8FF182D8F14AF3082" bold="true" box="[151,434,151,178]" italics="true" pageId="7" pageNumber="466">Hydrochus biexcavatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E69C8215004DFFC8FE4E2D8F174D3081" box="[449,592,151,177]" pageId="7" pageNumber="466" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7004DFFC8FF182DA3143430E5" box="[151,297,187,214]" pageId="7" pageNumber="466" type="description">
<paragraph id="CF64E37C004DFFC8FF182DA3143430E5" blockId="7.[151,592,151,214]" box="[151,297,187,214]" pageId="7" pageNumber="466">
<figureCitation id="57E0FFF9004DFFC8FF182DA315FE30E5" box="[151,227,187,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,1863,1887]" captionTargetBox="[186,1395,194,1808]" captionTargetId="figure-22@4.[151,1435,179,1837]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of Central American Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4777843" httpUri="https://zenodo.org/record/4777843/files/figure.png" pageId="7" pageNumber="466">Figs. 1</figureCitation>
,
<figureCitation id="57E0FFF9004DFFC8FF7E2DA3141C30E6" box="[241,257,187,214]" captionStart="FIGURES 5–6" captionStartId="24.[151,264,1934,1958]" captionTargetBox="[161,1425,192,1893]" captionTargetId="figure-22@24.[151,1435,181,1910]" captionTargetPageId="24" captionText="FIGURES 5–6. Habitus and male genitalia of holotypes. 5. Hydrochus biexcavatus n. sp. 6. Hydrochus mesoamericanus n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777855" httpUri="https://zenodo.org/record/4777855/files/figure.png" pageId="7" pageNumber="466">5</figureCitation>
,
<figureCitation id="57E0FFF9004DFFC8FE822DA3143430E5" box="[269,297,187,213]" captionStart="FIGURES 28–30" captionStartId="35.[151,264,1944,1968]" captionTargetBox="[158,1429,181,1920]" captionTargetId="figure-15@35.[158,1429,181,1920]" captionTargetPageId="35" captionText="FIGURES 28–30. Geographical distribution maps." figureDoi="http://doi.org/10.5281/zenodo.4777904" httpUri="https://zenodo.org/record/4777904/files/figure.png" pageId="7" pageNumber="466">29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7004DFFC8FF182C1B14A9346A" pageId="7" pageNumber="466" type="materials_examined">
<paragraph id="CF64E37C004DFFC8FF182C1B14A9346A" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="466">
<emphasis id="FDAF3F6E004DFFC8FF182C1B145B312D" bold="true" box="[151,326,259,285]" pageId="7" pageNumber="466">Type material.</emphasis>
<materialsCitation id="7FB3E921004DFFC8FEC12C1B14723171" ID-GBIF-Occurrence="3128194533" collectingDate="2008-07-13" collectionCode="NMW" collectorName="Wewalka" country="Mexico" elevation="50" location="Campeche" pageId="7" pageNumber="466" specimenCount="1" stateProvince="Campeche" typeStatus="holotype">
<typeStatus id="10605DDE004DFFC8FEC12C1B14AB312D" box="[334,438,259,285]" pageId="7" pageNumber="466" type="holotype">Holotype</typeStatus>
(male): “
<emphasis id="FDAF3F6E004DFFC8FDAC2C1B1767312D" bold="true" box="[547,634,259,285]" pageId="7" pageNumber="466">
<collectingCountry id="B7CCA3EC004DFFC8FDAC2C1B1767312D" box="[547,634,259,285]" name="Mexico" pageId="7" pageNumber="466">Mexico</collectingCountry>
</emphasis>
: Prov.
<collectingRegion id="0D1F2D9E004DFFC8FD432C1B1658312D" box="[716,837,259,285]" country="Mexico" name="Campeche" pageId="7" pageNumber="466">Campeche</collectingRegion>
,
<quantity id="08234E99004DFFC8FCDE2C1B1684312D" box="[849,921,259,285]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="466" unit="km" value="10.0">10 km</quantity>
S
<collectingRegion id="0D1F2D9E004DFFC8FC372C1B112C312D" box="[952,1073,259,285]" country="Mexico" name="Campeche" pageId="7" pageNumber="466">Campeche</collectingRegion>
,
<quantity id="08234E99004DFFC8FBB22C1B1164312E" box="[1085,1145,259,286]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="466" unit="m" value="50.0">
<elevation id="44F6044F004DFFC8FBB22C1B1164312E" box="[1085,1145,259,286]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="466" unit="m" value="50.0">50 m</elevation>
</quantity>
,
<date id="BB65C5BC004DFFC8FB0A2C1B11E5312D" box="[1157,1272,259,285]" pageId="7" pageNumber="466" value="2008-07-13">
<collectingDate id="AB213C54004DFFC8FB0A2C1B11E5312D" box="[1157,1272,259,285]" pageId="7" pageNumber="466" value="2008-07-13">13.7.2008</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FABA2C1B1081312D" box="[1333,1436,259,285]" pageId="7" pageNumber="466">Wewalka</collectorName>
(Mex 10)” (
<collectionCode id="A9CA7BB9004DFFC8FE952C3F14753171" box="[282,360,295,321]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="466">NMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="7FB3E921004DFFC8FEF62C3F17D03156" ID-GBIF-Occurrence="3128194476" collectingDate="1986-05-23" collectionCode="NMNH" collectorName="Paul J Spangler &amp; Robiin A. Faitoute" country="Belize" location="San Igmacio" pageId="7" pageNumber="466" specimenCount="1" stateProvince="Cayo" typeStatus="paratype">
<emphasis id="FDAF3F6E004DFFC8FEF62C3F17963171" bold="true" box="[377,651,295,321]" pageId="7" pageNumber="466">
<typeStatus id="10605DDE004DFFC8FEF62C3F14EC3171" box="[377,497,295,321]" pageId="7" pageNumber="466" type="paratype">Paratypes</typeStatus>
(123):
<collectingCountry id="B7CCA3EC004DFFC8FDCC2C3F17963171" box="[579,651,295,321]" name="Belize" pageId="7" pageNumber="466">Belize</collectingCountry>
</emphasis>
:
<collectingRegion id="0D1F2D9E004DFFC8FD172C3F162D3171" box="[664,816,295,321]" country="Belize" name="Cayo" pageId="7" pageNumber="466">Cayo District</collectingRegion>
,
<location id="CA04B5A7004DFFC8FCB52C3F16DB3171" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FCB52C3F16DB3171" box="[826,966,295,321]" country="Belize" name="San Igmacio" pageId="7" pageNumber="466" stateProvince="Cayo">San Igmacio</location>
(
<quantity id="08234E99004DFFC8FC5A2C3F11103171" box="[981,1037,295,321]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="466" unit="km" value="8.0">8 km</quantity>
S); stock pond,
<date id="BB65C5BC004DFFC8FB4F2C3F102C3171" box="[1216,1329,295,321]" pageId="7" pageNumber="466" value="1986-05-23">
<collectingDate id="AB213C54004DFFC8FB4F2C3F102C3171" box="[1216,1329,295,321]" pageId="7" pageNumber="466" value="1986-05-23">23.v.1986</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FAE52C3F14103155" pageId="7" pageNumber="466">Paul J Spangler</collectorName>
&amp;
<collectorName id="622E86AA004DFFC8FEBE2C53171C3155" box="[305,513,331,357]" pageId="7" pageNumber="466">Robiin A. Faitoute</collectorName>
(16) (29
<collectionCode id="A9CA7BB9004DFFC8FDE52C5417D83156" box="[618,709,332,358]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="466" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="7FB3E921004DFFC8FD582C53145331BA" ID-GBIF-Occurrence="3128194353" collectingDate="1965-07-25" collectionCode="NMNH" collectorName="Paul J. Spangler" country="Costa Rica" location="Liberia" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<emphasis id="FDAF3F6E004DFFC8FD582C5316443155" bold="true" box="[727,857,331,357]" pageId="7" pageNumber="466">
<collectingCountry id="B7CCA3EC004DFFC8FD582C5316443155" box="[727,857,331,357]" name="Costa Rica" pageId="7" pageNumber="466">Costa Rica</collectingCountry>
</emphasis>
:
<quantity id="08234E99004DFFC8FCE72C5316AD3155" box="[872,944,331,357]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="7" pageNumber="466" unit="mi" value="10.0">10 mi.</quantity>
NW
<location id="CA04B5A7004DFFC8FC632C53115D3155" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FC632C53115D3155" box="[1004,1088,331,357]" country="Costa Rica" name="Liberia" pageId="7" pageNumber="466">Liberia</location>
,
<date id="BB65C5BC004DFFC8FBC42C5311D03156" box="[1099,1229,331,358]" pageId="7" pageNumber="466" value="1965-07-25">
<collectingDate id="AB213C54004DFFC8FBC42C5311D03156" box="[1099,1229,331,358]" pageId="7" pageNumber="466" value="1965-07-25">25.vii.1965</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FA862C5315DE31B9" pageId="7" pageNumber="466">Paul J. Spangler</collectorName>
(1
<collectionCode id="A9CA7BB9004DFFC8FF642C68145B31BA" box="[235,326,368,394]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="466" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FED52C77112E31BA" ID-GBIF-Occurrence="3128194523" box="[346,1075,367,394]" collectingDate="1965-07-13" collectionCode="NMNH" collectorName="Paul J. Spangler" country="Costa Rica" elevation="16" location="La Cruz" pageId="7" pageNumber="466" specimenCount="16" typeStatus="paratype">
<specimenCount id="D9DD28F5004DFFC8FED52C77148131BA" box="[346,412,367,394]" pageId="7" pageNumber="466" type="generic">
<quantity id="08234E99004DFFC8FED52C77148531BA" box="[346,408,367,394]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="466" unit="m" value="16.0">
<elevation id="44F6044F004DFFC8FED52C77148531BA" box="[346,408,367,394]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="466" unit="m" value="16.0">16 m</elevation>
</quantity>
.
</specimenCount>
S.
<location id="CA04B5A7004DFFC8FE4C2C68173F31B9" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FE4C2C68173F31B9" box="[451,546,367,394]" country="Costa Rica" name="La Cruz" pageId="7" pageNumber="466">La Cruz</location>
,
<date id="BB65C5BC004DFFC8FDA02C7717AC31BA" box="[559,689,367,394]" pageId="7" pageNumber="466" value="1965-07-13">
<collectingDate id="AB213C54004DFFC8FDA02C7717AC31BA" box="[559,689,367,394]" pageId="7" pageNumber="466" value="1965-07-13">13.vii.1965</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FD7F2C7716B431B9" box="[752,937,367,394]" pageId="7" pageNumber="466">Paul J. Spangler</collectorName>
(1
<collectionCode id="A9CA7BB9004DFFC8FC5E2C68113131BA" box="[977,1068,368,394]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="466" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FBCF2C77170B319E" ID-GBIF-Occurrence="3128194505" collectingDate="1965-07-24" collectionCode="NMNH" collectorName="Paul J. Spangler" country="Costa Rica" location="Liberia" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<quantity id="08234E99004DFFC8FBCF2C77116631B9" box="[1088,1147,367,394]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="7" pageNumber="466" unit="mi" value="5.0">5 mi.</quantity>
SW
<location id="CA04B5A7004DFFC8FB3A2C77101431B9" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FB3A2C77101431B9" box="[1205,1289,367,393]" country="Costa Rica" name="Liberia" pageId="7" pageNumber="466">Liberia</location>
,
<date id="BB65C5BC004DFFC8FA992C77108531BA" box="[1302,1432,367,394]" pageId="7" pageNumber="466" value="1965-07-24">
<collectingDate id="AB213C54004DFFC8FA992C77108531BA" box="[1302,1432,367,394]" pageId="7" pageNumber="466" value="1965-07-24">24.vii.1965</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FF472C8B1462319D" box="[200,383,403,430]" pageId="7" pageNumber="466">Paul J. Spangler</collectorName>
(17
<collectionCode id="A9CA7BB9004DFFC8FE3C2C8C1713319E" box="[435,526,404,430]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="466" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FDAE2C8B11F7319E" ID-GBIF-Occurrence="3128194392" box="[545,1258,403,430]" collectingDate="1962-07-21" collectionCode="NMNH" country="Costa Rica" location="Santa Cruz" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<location id="CA04B5A7004DFFC8FDAE2C8B17BD319D" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FDAE2C8B17BD319D" box="[545,672,403,429]" country="Costa Rica" name="Santa Cruz" pageId="7" pageNumber="466">Santa Cruz</location>
,
<date id="BB65C5BC004DFFC8FD232C8B1633319D" box="[684,814,403,429]" pageId="7" pageNumber="466" value="1962-07-21">
<collectingDate id="AB213C54004DFFC8FD232C8B1633319D" box="[684,814,403,429]" pageId="7" pageNumber="466" value="1962-07-21">21.vii.1962</collectingDate>
</date>
, leg. no collector on label (24
<collectionCode id="A9CA7BB9004DFFC8FB082C8C11FF319E" box="[1159,1250,404,430]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="466" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FB7B2C8B178931E2" ID-GBIF-Occurrence="3128194573" collectingDate="1998-08-24" collectionCode="NMW" collectorName="Drechsel" country="Costa Rica" location="Cano Negro" pageId="7" pageNumber="466" specimenCount="1" stateProvince="Alajuela" typeStatus="paratype">
<collectingRegion id="0D1F2D9E004DFFC8FB7B2C8B104B319D" box="[1268,1366,403,429]" country="Costa Rica" name="Alajuela" pageId="7" pageNumber="466">Alajuela</collectingRegion>
,
<location id="CA04B5A7004DFFC8FAEE2C8B15FD31E2" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FAEE2C8B15FD31E2" country="Costa Rica" name="Cano Negro" pageId="7" pageNumber="466" stateProvince="Alajuela">Cano Negro</location>
,
<date id="BB65C5BC004DFFC8FF632CAF146B31E1" box="[236,374,439,465]" pageId="7" pageNumber="466" value="1998-08-24">
<collectingDate id="AB213C54004DFFC8FF632CAF146B31E1" box="[236,374,439,465]" pageId="7" pageNumber="466" value="1998-08-24">24.viii.1998</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FE3B2CAF170531E1" box="[436,536,439,465]" pageId="7" pageNumber="466">Drechsel</collectorName>
(1
<collectionCode id="A9CA7BB9004DFFC8FDB02CA0179131E2" box="[575,652,440,466]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="466">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FD2F2CAF142131C5" ID-GBIF-Occurrence="3128194482" collectingDate="1965-03-23" collectionCode="CAS" collectorName="P. Cook" country="Costa Rica" location="Hacienda Tagoba" pageId="7" pageNumber="466" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<collectingRegion id="0D1F2D9E004DFFC8FD2F2CAF163931E1" box="[672,804,439,465]" country="Costa Rica" name="Guanacaste" pageId="7" pageNumber="466">Guanacaste</collectingRegion>
,
<locationDeviation id="DED6FA5E004DFFC8FCBF2CAF16ED31E1" box="[816,1008,439,466]" location="Canas" pageId="7" pageNumber="466">
<quantity id="08234E99004DFFC8FCBF2CAF166531E1" box="[816,888,439,466]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="466" unit="km" value="15.0">15 km</quantity>
SE Canas
</locationDeviation>
,
<location id="CA04B5A7004DFFC8FC732CAF11DE31E1" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FC732CAF11DE31E1" box="[1020,1219,439,465]" country="Costa Rica" name="Hacienda Tagoba" pageId="7" pageNumber="466" stateProvince="Guanacaste">Hacienda Tagoba</location>
,
<date id="BB65C5BC004DFFC8FB402CAF105731E2" box="[1231,1354,439,466]" pageId="7" pageNumber="466" value="1965-03-23">
<collectingDate id="AB213C54004DFFC8FB402CAF105731E2" box="[1231,1354,439,466]" pageId="7" pageNumber="466" value="1965-03-23">23.iii.1965</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FA062CA015C931C5" pageId="7" pageNumber="466">P. Cook</collectorName>
(1
<collectionCode id="A9CA7BB9004DFFC8FF752CC3142831C5" box="[250,309,475,501]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="7" pageNumber="466" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FEC72CC315E7322A" ID-GBIF-Occurrence="3128194467" collectingDate="1973-06-04" collectionCode="NMNH" collectorName="Erwin &amp; Hevel" country="Costa Rica" latitude="10.65" location="Rio Carana" longLatPrecision="1297" longitude="-85.36667" municipality="Hwy" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<locationDeviation id="DED6FA5E004DFFC8FEC72CC3175C31C6" box="[328,577,475,502]" location="Bagaces" pageId="7" pageNumber="466">
<quantity id="08234E99004DFFC8FEC72CC314B831C5" box="[328,421,475,501]" metricMagnitude="4" metricUnit="m" metricValue="1.8699999999999999" pageId="7" pageNumber="466" unit="km" value="18.7">18.7 km</quantity>
NW Bagaces
</locationDeviation>
,
<collectingMunicipality id="2F007906004DFFC8FDC32CC4179F31C6" box="[588,642,476,502]" pageId="7" pageNumber="466">Hwy</collectingMunicipality>
1,
<location id="CA04B5A7004DFFC8FD292CC3163531C5" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FD292CC3163531C5" box="[678,808,475,501]" country="Costa Rica" latitude="10.65" longLatPrecision="1297" longitude="-85.36667" municipality="Hwy" name="Rio Carana" pageId="7" pageNumber="466">Rio Carana</location>
, (
<geoCoordinate id="AAEF85BB004DFFC8FCB32CC316BD31C5" box="[828,928,475,501]" degrees="10" direction="north" minutes="39" orientation="latitude" pageId="7" pageNumber="466" precision="925" value="10.65">10°39’N</geoCoordinate>
,
<geoCoordinate id="AAEF85BB004DFFC8FC242CC3110B31C6" box="[939,1046,475,502]" degrees="85" direction="west" minutes="22" orientation="longitude" pageId="7" pageNumber="466" precision="925" value="-85.36667">85°22’W</geoCoordinate>
),
<date id="BB65C5BC004DFFC8FBA52CC3118B31C5" box="[1066,1174,475,501]" pageId="7" pageNumber="466" value="1973-06-04">
<collectingDate id="AB213C54004DFFC8FBA52CC3118B31C5" box="[1066,1174,475,501]" pageId="7" pageNumber="466" value="1973-06-04">4.vi.1973</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FB5D2CC3100B31C5" box="[1234,1302,475,501]" pageId="7" pageNumber="466">Erwin</collectorName>
&amp;
<collectorName id="622E86AA004DFFC8FAB42CC3106331C5" box="[1339,1406,475,501]" pageId="7" pageNumber="466">Hevel</collectorName>
(1
<collectionCode id="A9CA7BB9004DFFC8FF182F1815EF322A" box="[151,242,512,538]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="466" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FE8A2F18102C3229" ID-GBIF-Occurrence="3128194444" box="[261,1329,511,538]" collectingDate="2002-11-27" collectionCode="SEMC" collectorName="W. Shepard" country="Costa Rica" elevation="61" location="Barra Honda" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
F.N.
<location id="CA04B5A7004DFFC8FEB42F1814D33229" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FEB42F1814D33229" box="[315,462,511,538]" country="Costa Rica" name="Barra Honda" pageId="7" pageNumber="466">Barra Honda</location>
, pond and outflow stream, elev.
<quantity id="08234E99004DFFC8FCB82CE7166C322A" box="[823,881,511,538]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="7" pageNumber="466" unit="m" value="61.0">
<elevation id="44F6044F004DFFC8FCB82CE7166C322A" box="[823,881,511,538]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="7" pageNumber="466" unit="m" value="61.0">61 m</elevation>
</quantity>
,
<date id="BB65C5BC004DFFC8FCF22CE716EA3229" box="[893,1015,511,537]" pageId="7" pageNumber="466" value="2002-11-27">
<collectingDate id="AB213C54004DFFC8FCF22CE716EA3229" box="[893,1015,511,537]" pageId="7" pageNumber="466" value="2002-11-27">27.xi.2002</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FBBD2F1811AE3229" box="[1074,1203,511,538]" pageId="7" pageNumber="466">W. Shepard</collectorName>
(3
<collectionCode id="A9CA7BB9004DFFC8FB572CE710343229" box="[1240,1321,511,537]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="7" pageNumber="466" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FAB32CE714BA3251" ID-GBIF-Occurrence="3128194306" collectingDate="2002-06-27" collectionCode="SEMC" collectorName="W. Shepard &amp; R. Roughley" country="Costa Rica" county="Nandayure" elevation="61" latitude="10.180834" location="Lake" longLatPrecision="21" longitude="-85.32972" municipality="Barra Honda" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<collectingCounty id="26059BF0004DFFC8FAB32CE715A3320E" pageId="7" pageNumber="466">Nandayure</collectingCounty>
;
<collectingMunicipality id="2F007906004DFFC8FF442F3C1440320D" box="[203,349,547,574]" pageId="7" pageNumber="466">Barra Honda</collectingMunicipality>
N.P.,
<location id="CA04B5A7004DFFC8FE2C2F3B14C6320D" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FE2C2F3B14C6320D" box="[419,475,547,573]" country="Costa Rica" county="Nandayure" latitude="10.180834" longLatPrecision="21" longitude="-85.32972" municipality="Barra Honda" name="Lake" pageId="7" pageNumber="466">Lake</location>
at trailhead, elev.
<quantity id="08234E99004DFFC8FD212F3B17F5320E" box="[686,744,547,574]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="7" pageNumber="466" unit="m" value="61.0">
<elevation id="44F6044F004DFFC8FD212F3B17F5320E" box="[686,744,547,574]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="7" pageNumber="466" unit="m" value="61.0">61 m</elevation>
</quantity>
(
<geoCoordinate id="AAEF85BB004DFFC8FD762F3B1694320E" box="[761,905,547,574]" degrees="10" direction="north" minutes="10" orientation="latitude" pageId="7" pageNumber="466" precision="15" seconds="51" value="10.180834">10°10’51’’N</geoCoordinate>
,
<geoCoordinate id="AAEF85BB004DFFC8FC192F3B1130320E" box="[918,1069,547,574]" degrees="85" direction="west" minutes="19" orientation="longitude" pageId="7" pageNumber="466" precision="15" seconds="47" value="-85.32972">85°19’47’’W</geoCoordinate>
),
<date id="BB65C5BC004DFFC8FBCC2F3B11A0320D" box="[1091,1213,547,573]" pageId="7" pageNumber="466" value="2002-06-27">
<collectingDate id="AB213C54004DFFC8FBCC2F3B11A0320D" box="[1091,1213,547,573]" pageId="7" pageNumber="466" value="2002-06-27">27.vi.2002</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FB742F3C1063320D" box="[1275,1406,547,574]" pageId="7" pageNumber="466">W. Shepard</collectorName>
&amp;
<collectorName id="622E86AA004DFFC8FF182F50143B3251" box="[151,294,583,610]" pageId="7" pageNumber="466">R. Roughley</collectorName>
(6
<collectionCode id="A9CA7BB9004DFFC8FEC12F5F14823251" box="[334,415,583,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="7" pageNumber="466" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FE3B2F5F10433251" ID-GBIF-Occurrence="3128194320" box="[436,1374,583,610]" collectingDate="1993-06-04" collectionCode="SEMC" collectorName="Steve Lingafelter" country="Costa Rica" location="Palo Verde Biol. Station" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<location id="CA04B5A7004DFFC8FE3B2F5F17D43251" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FE3B2F5F17D43251" box="[436,713,583,609]" country="Costa Rica" name="Palo Verde Biol. Station" pageId="7" pageNumber="466">Palo Verde Biol. Station</location>
, ex. blacklite,
<date id="BB65C5BC004DFFC8FCE02F5F16C63251" box="[879,987,583,609]" pageId="7" pageNumber="466" value="1993-06-04">
<collectingDate id="AB213C54004DFFC8FCE02F5F16C63251" box="[879,987,583,609]" pageId="7" pageNumber="466" value="1993-06-04">4.vi.1993</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FB952F5F11C13251" box="[1050,1244,583,609]" pageId="7" pageNumber="466">Steve Lingafelter</collectorName>
(2
<collectionCode id="A9CA7BB9004DFFC8FA8A2F5F104B3251" box="[1285,1366,583,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="7" pageNumber="466" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FAE52F5F106D32B5" ID-GBIF-Occurrence="3128194603" collectingDate="2004-01-11" collectionCode="SEMC" collectorName="Palo Verde N. P." country="Costa Rica" location="Birding" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<collectorName id="622E86AA004DFFC8FAE52F5F140E32B6" pageId="7" pageNumber="466">Palo Verde N.P.</collectorName>
: nr.
<location id="CA04B5A7004DFFC8FECD2F73148532B5" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FECD2F73148532B5" box="[322,408,619,645]" country="Costa Rica" name="Birding" pageId="7" pageNumber="466">Birding</location>
tower; lagoon margin,
<date id="BB65C5BC004DFFC8FD2F2F73161632B5" box="[672,779,619,645]" pageId="7" pageNumber="466" value="2004-01-11">
<collectingDate id="AB213C54004DFFC8FD2F2F73161632B5" box="[672,779,619,645]" pageId="7" pageNumber="466" value="2004-01-11">11.i.2004</collectingDate>
</date>
, leg. AEZShort &amp; DJLebbin (AS-04-030a) (1
<collectionCode id="A9CA7BB9004DFFC8FA982F73107532B5" box="[1303,1384,619,645]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="7" pageNumber="466" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FAF32F7315ED32FD" ID-GBIF-Occurrence="3128194561" collectingDate="2004-01-11" collectionCode="SEMC" country="Costa Rica" county="Rd" elevation="35" location="Rt." municipality="Palo Verde" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<collectingCounty id="26059BF0004DFFC8FAF32F73108032B5" box="[1404,1437,619,645]" pageId="7" pageNumber="466">Rd</collectingCounty>
to
<collectingMunicipality id="2F007906004DFFC8FF3B2F97142D3299" box="[180,304,655,681]" pageId="7" pageNumber="466">Palo Verde</collectingMunicipality>
;
<quantity id="08234E99004DFFC8FEB42F97148B3299" box="[315,406,655,681]" metricMagnitude="4" metricUnit="m" metricValue="1.14" pageId="7" pageNumber="466" unit="km" value="11.4">11.4 km</quantity>
past
<location id="CA04B5A7004DFFC8FE5F2F8814EF329A" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FE5F2F8814EF329A" box="[464,498,656,682]" country="Costa Rica" county="Rd" municipality="Palo Verde" name="Rt." pageId="7" pageNumber="466">Rt.</location>
1; roadside ditch, elev.
<quantity id="08234E99004DFFC8FD762F971629329A" box="[761,820,655,682]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="466" unit="m" value="35.0">
<elevation id="44F6044F004DFFC8FD762F971629329A" box="[761,820,655,682]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="466" unit="m" value="35.0">35 m</elevation>
</quantity>
,
<date id="BB65C5BC004DFFC8FCB02F9716B73299" box="[831,938,655,681]" pageId="7" pageNumber="466" value="2004-01-11">
<collectingDate id="AB213C54004DFFC8FCB02F9716B73299" box="[831,938,655,681]" pageId="7" pageNumber="466" value="2004-01-11">11.i.2004</collectingDate>
</date>
, leg. AEZShort &amp; DJLebbin (AS-04-027) (2
<collectionCode id="A9CA7BB9004DFFC8FF182FAB15F532FD" box="[151,232,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="7" pageNumber="466" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FF732FAB167B32C2" ID-GBIF-Occurrence="3128194491" collectingDate="1996-06-01" collectionCode="NMW" collectorName="R. S. Hanley" country="Costa Rica" elevation="80" latitude="10.416667" location="La Selva" longLatPrecision="1298" longitude="-84.0" pageId="7" pageNumber="466" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
<collectingRegion id="0D1F2D9E004DFFC8FF732FAB144532FD" box="[252,344,691,717]" country="Costa Rica" name="Heredia" pageId="7" pageNumber="466">Heredia</collectingRegion>
,
<location id="CA04B5A7004DFFC8FEEB2FAC14DA32FD" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FEEB2FAC14DA32FD" box="[356,455,691,718]" country="Costa Rica" latitude="10.416667" longLatPrecision="1298" longitude="-84.0" name="La Selva" pageId="7" pageNumber="466" stateProvince="Heredia">La Selva</location>
(
<locationDeviation id="DED6FA5E004DFFC8FE572FAB17DF32FE" box="[472,706,691,718]" location="Puerto Viejo" pageId="7" pageNumber="466">
<quantity id="08234E99004DFFC8FE572FAB170F32FD" box="[472,530,691,717]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="466" unit="km" value="3.0">3 km</quantity>
S Puerto Viejo
</locationDeviation>
), at UV blacklight in station nr. 2nd forest, elev.
<quantity id="08234E99004DFFC8FB652FAB103932FE" box="[1258,1316,691,718]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="466" unit="m" value="80.0">
<elevation id="44F6044F004DFFC8FB652FAB103932FE" box="[1258,1316,691,718]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="466" unit="m" value="80.0">80 m</elevation>
</quantity>
(
<geoCoordinate id="AAEF85BB004DFFC8FABB2FAB108532FE" box="[1332,1432,691,718]" degrees="10" direction="north" minutes="25" orientation="latitude" pageId="7" pageNumber="466" precision="925" value="10.416667">10°25’N</geoCoordinate>
,
<geoCoordinate id="AAEF85BB004DFFC8FF182FCF15EE32C1" box="[151,243,727,753]" degrees="84" direction="west" minutes="0" orientation="longitude" pageId="7" pageNumber="466" precision="925" value="-84.0">84°0’W</geoCoordinate>
),
<date id="BB65C5BC004DFFC8FE862FCF146832C1" box="[265,373,727,753]" pageId="7" pageNumber="466" value="1996-06-01">
<collectingDate id="AB213C54004DFFC8FE862FCF146832C1" box="[265,373,727,753]" pageId="7" pageNumber="466" value="1996-06-01">1.vi.1996</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FE3A2FC0175A32C1" box="[437,583,727,754]" pageId="7" pageNumber="466">R. S. Hanley</collectorName>
(RSHCR012) (1
<collectionCode id="A9CA7BB9004DFFC8FC9E2FC0164332C2" box="[785,862,728,754]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="466">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FCFC2FC015F1330A" ID-GBIF-Occurrence="3128194537" collectingDate="2009-03-01" collectingDateMax="2009-03-03" collectingDateMin="2009-03-01" collectionCode="NMW" collectorName="Qu. Negra &amp; M. Brojer" country="Costa Rica" elevation="90" latitude="8.701667" location="Field Station La Gamba" longLatPrecision="21" longitude="-83.20194" municipality="La Gamba" pageId="7" pageNumber="466" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
<collectingRegion id="0D1F2D9E004DFFC8FCFC2FC016EC32C2" box="[883,1009,728,754]" country="Costa Rica" name="Puntarenas" pageId="7" pageNumber="466">Puntarenas</collectingRegion>
,
<collectorName id="622E86AA004DFFC8FC722FCF116B32C2" box="[1021,1142,727,754]" pageId="7" pageNumber="466">Qu. Negra</collectorName>
, ca.
<locationDeviation id="DED6FA5E004DFFC8FB232FCF106A32C1" box="[1196,1399,727,754]" location="Golfito" pageId="7" pageNumber="466">
<quantity id="08234E99004DFFC8FB232FCF11FB32C1" box="[1196,1254,727,753]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="466" unit="km" value="8.0">8 km</quantity>
NW Golfito
</locationDeviation>
nr.
<collectingMunicipality id="2F007906004DFFC8FF182FE414133325" box="[151,270,763,790]" pageId="7" pageNumber="466">La Gamba</collectingMunicipality>
(vill.), at
<location id="CA04B5A7004DFFC8FE0D2FE317883325" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FE0D2FE317883325" box="[386,661,763,790]" country="Costa Rica" latitude="8.701667" longLatPrecision="21" longitude="-83.20194" municipality="La Gamba" name="Field Station La Gamba" pageId="7" pageNumber="466" stateProvince="Puntarenas">Field Station La Gamba</location>
, elev.
<quantity id="08234E99004DFFC8FD512FE316053326" box="[734,792,763,790]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="466" unit="m" value="90.0">
<elevation id="44F6044F004DFFC8FD512FE316053326" box="[734,792,763,790]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="466" unit="m" value="90.0">90 m</elevation>
</quantity>
(
<geoCoordinate id="AAEF85BB004DFFC8FCA52FE316803325" box="[810,925,763,789]" degrees="8" direction="north" minutes="42" orientation="latitude" pageId="7" pageNumber="466" precision="15" seconds="6" value="8.701667">8°42’6’’N</geoCoordinate>
,
<geoCoordinate id="AAEF85BB004DFFC8FC242FE3112E3325" box="[939,1075,763,789]" degrees="83" direction="west" minutes="12" orientation="longitude" pageId="7" pageNumber="466" precision="15" seconds="7" value="-83.20194">83°12’7’’W</geoCoordinate>
),
<date id="BB65C5BC004DFFC8FBC62FE311D03325" box="[1097,1229,763,789]" pageId="7" pageNumber="466" value="2009-03-01" valueMax="2009-03-03" valueMin="2009-03-01">
<collectingDate id="AB213C54004DFFC8FBC62FE311D03325" box="[1097,1229,763,789]" pageId="7" pageNumber="466" value="2009-03-01" valueMax="2009-03-03" valueMin="2009-03-01">1-3.iii.2009</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FA822FE410603325" box="[1293,1405,763,790]" pageId="7" pageNumber="466">M. Brojer</collectorName>
(1
<collectionCode id="A9CA7BB9004DFFC8FF182E3815F9330A" box="[151,228,800,826]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="466">NMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="7FB3E921004DFFC8FF782E0716DE3309" ID-GBIF-Occurrence="3128194552" box="[247,963,799,826]" collectingDate="1966-05-20" collectionCode="FSCA" collectorName="J. M. Matta" country="Honduras" location="Casas Viejas" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<emphasis id="FDAF3F6E004DFFC8FF782E0714723309" bold="true" box="[247,367,799,825]" pageId="7" pageNumber="466">
<collectingCountry id="B7CCA3EC004DFFC8FF782E0714723309" box="[247,367,799,825]" name="Honduras" pageId="7" pageNumber="466">Honduras</collectingCountry>
</emphasis>
:
<location id="CA04B5A7004DFFC8FEF12E0717123309" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FEF12E0717123309" box="[382,527,799,825]" country="Honduras" name="Casas Viejas" pageId="7" pageNumber="466">Casas Viejas</location>
,
<date id="BB65C5BC004DFFC8FD952E0717963309" box="[538,651,799,825]" pageId="7" pageNumber="466" value="1966-05-20">
<collectingDate id="AB213C54004DFFC8FD952E0717963309" box="[538,651,799,825]" pageId="7" pageNumber="466" value="1966-05-20">20.v.1966</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FD472E381657330A" box="[712,842,800,826]" pageId="7" pageNumber="466">J. M. Matta</collectorName>
(1
<collectionCode id="A9CA7BB9004DFFC8FCFF2E0716A63309" box="[880,955,799,825]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="7" pageNumber="466" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="7FB3E921004DFFC8FC412E071637336E" ID-GBIF-Occurrence="3128194470" collectingDate="2008-07-13" collectionCode="NMW" collectorName="Wewalka &amp; Mex" country="Mexico" elevation="50" location="10 km S Campeche" pageId="7" pageNumber="466" specimenCount="1" stateProvince="Campeche" typeStatus="paratype">
<emphasis id="FDAF3F6E004DFFC8FC412E0711383309" bold="true" box="[974,1061,799,825]" pageId="7" pageNumber="466">
<collectingCountry id="B7CCA3EC004DFFC8FC412E0711383309" box="[974,1061,799,825]" name="Mexico" pageId="7" pageNumber="466">Mexico</collectingCountry>
</emphasis>
:
<collectingRegion id="0D1F2D9E004DFFC8FBBB2E0711B03309" box="[1076,1197,799,825]" country="Mexico" name="Campeche" pageId="7" pageNumber="466">Campeche</collectingRegion>
,
<location id="CA04B5A7004DFFC8FB362E0710853309" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FB362E0710853309" box="[1209,1432,799,825]" country="Mexico" name="10 km S Campeche" pageId="7" pageNumber="466" stateProvince="Campeche">
<quantity id="08234E99004DFFC8FB362E07101C3309" box="[1209,1281,799,825]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="466" unit="km" value="10.0">10 km</quantity>
S
<collectingRegion id="0D1F2D9E004DFFC8FA902E0710853309" box="[1311,1432,799,825]" country="Mexico" name="Campeche" pageId="7" pageNumber="466">Campeche</collectingRegion>
</location>
, elev.
<quantity id="08234E99004DFFC8FF5D2E5B1413336E" box="[210,270,835,862]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="466" unit="m" value="50.0">
<elevation id="44F6044F004DFFC8FF5D2E5B1413336E" box="[210,270,835,862]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="466" unit="m" value="50.0">50 m</elevation>
</quantity>
,
<date id="BB65C5BC004DFFC8FE952E5B1481336D" box="[282,412,835,861]" pageId="7" pageNumber="466" value="2008-07-13">
<collectingDate id="AB213C54004DFFC8FE952E5B1481336D" box="[282,412,835,861]" pageId="7" pageNumber="466" value="2008-07-13">13.vii.2008</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FE572E5B1722336D" box="[472,575,835,861]" pageId="7" pageNumber="466">Wewalka</collectorName>
(
<collectorName id="622E86AA004DFFC8FDC12E5B179E336D" box="[590,643,835,861]" pageId="7" pageNumber="466">Mex</collectorName>
10) (7
<collectionCode id="A9CA7BB9004DFFC8FD592E5C163E336E" box="[726,803,836,862]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="466">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FCB92E5B15E733B2" ID-GBIF-Occurrence="3128194489" collectingDate="1964-07-07" collectionCode="NMNH" collectorName="P. J. Spangler" country="Mexico" location="Patzcuaro" pageId="7" pageNumber="466" specimenCount="1" stateProvince="Michoacan de Ocampo" typeStatus="paratype">
<collectingRegion id="0D1F2D9E004DFFC8FCB92E5B16A8336D" box="[822,949,835,861]" country="Mexico" name="Michoacan de Ocampo" pageId="7" pageNumber="466">Michoacan</collectingRegion>
,
<location id="CA04B5A7004DFFC8FC4F2E5C112C336E" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FC4F2E5C112C336E" box="[960,1073,836,862]" country="Mexico" name="Patzcuaro" pageId="7" pageNumber="466" stateProvince="Michoacan de Ocampo">Patzcuaro</location>
,
<date id="BB65C5BC004DFFC8FBB22E5B11AC336D" box="[1085,1201,835,861]" pageId="7" pageNumber="466" value="1964-07-07">
<collectingDate id="AB213C54004DFFC8FBB22E5B11AC336D" box="[1085,1201,835,861]" pageId="7" pageNumber="466" value="1964-07-07">7.vii.1964</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FB622E5C1063336D" box="[1261,1406,835,862]" pageId="7" pageNumber="466">P.J. Spangler</collectorName>
(6
<collectionCode id="A9CA7BB9004DFFC8FF182E7015EF33B2" box="[151,242,872,898]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="466" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FE892E7F11BE33B1" ID-GBIF-Occurrence="3128194477" box="[262,1187,871,898]" collectingDate="1948-11-19" collectionCode="CAS" collectorName="H. B. Leech" country="Mexico" location="Maiz" pageId="7" pageNumber="466" specimenCount="1" stateProvince="San Luis Potosi" typeStatus="paratype">
<collectingRegion id="0D1F2D9E004DFFC8FE892E7F14A733B1" box="[262,442,871,897]" country="Mexico" name="San Luis Potosi" pageId="7" pageNumber="466">San Luis Potosi</collectingRegion>
,
<locationDeviation id="DED6FA5E004DFFC8FE492E7F174333B2" box="[454,606,871,898]" location="East" pageId="7" pageNumber="466">
<quantity id="08234E99004DFFC8FE492E7F173B33B1" box="[454,550,871,897]" metricMagnitude="4" metricUnit="m" metricValue="2.2530816" pageId="7" pageNumber="466" unit="mi" value="14.0">14 miles</quantity>
East
</locationDeviation>
C. del
<location id="CA04B5A7004DFFC8FD3E2E7F17F033B1" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FD3E2E7F17F033B1" box="[689,749,871,897]" country="Mexico" name="Maiz" pageId="7" pageNumber="466" stateProvince="San Luis Potosi">Maiz</location>
,
<date id="BB65C5BC004DFFC8FD762E7F166933B1" box="[761,884,871,897]" pageId="7" pageNumber="466" value="1948-11-19">
<collectingDate id="AB213C54004DFFC8FD762E7F166933B1" box="[761,884,871,897]" pageId="7" pageNumber="466" value="1948-11-19">19.xi.1948</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FC3E2E70112433B1" box="[945,1081,871,898]" pageId="7" pageNumber="466">H. B. Leech</collectorName>
(1
<collectionCode id="A9CA7BB9004DFFC8FBEE2E7F118133B1" box="[1121,1180,871,897]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="7" pageNumber="466" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FB3F2E7F14F83395" ID-GBIF-Occurrence="3128194375" collectingDate="1950-01-03" collectionCode="SEMC" collectorName="L. D. Beamer" country="Mexico" location="El Naranjo" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<location id="CA04B5A7004DFFC8FB3F2E7F103333B1" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FB3F2E7F103333B1" box="[1200,1326,871,897]" country="Mexico" name="El Naranjo" pageId="7" pageNumber="466">El Naranjo</location>
,
<date id="BB65C5BC004DFFC8FAB62E7F108533B2" box="[1337,1432,871,898]" pageId="7" pageNumber="466" value="1950-01-03">
<collectingDate id="AB213C54004DFFC8FAB62E7F108533B2" box="[1337,1432,871,898]" pageId="7" pageNumber="466" value="1950-01-03">3.i.1950</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FF462E9414793396" box="[201,356,908,934]" pageId="7" pageNumber="466">L. D. Beamer</collectorName>
(3
<collectionCode id="A9CA7BB9004DFFC8FE032E9314C03395" box="[396,477,907,933]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="7" pageNumber="466" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FE7E2E9310493395" ID-GBIF-Occurrence="3128194519" box="[497,1364,907,934]" collectingDate="1985-01-02" collectionCode="SEMC" collectorName="James Adams" country="Mexico" latitude="21.266666" location="Tamazunchale" longLatPrecision="1264" longitude="-98.8" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<location id="CA04B5A7004DFFC8FE7E2E9317883395" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FE7E2E9317883395" box="[497,661,907,933]" country="Mexico" latitude="21.266666" longLatPrecision="1264" longitude="-98.8" name="Tamazunchale" pageId="7" pageNumber="466">Tamazunchale</location>
, (
<geoCoordinate id="AAEF85BB004DFFC8FD242E9316133395" box="[683,782,907,933]" degrees="21" direction="north" minutes="16" orientation="latitude" pageId="7" pageNumber="466" precision="925" value="21.266666">21°16’N</geoCoordinate>
,
<geoCoordinate id="AAEF85BB004DFFC8FC942E93169B3395" box="[795,902,907,933]" degrees="98" direction="west" minutes="48" orientation="longitude" pageId="7" pageNumber="466" precision="925" value="-98.8">98°48’W</geoCoordinate>
),
<date id="BB65C5BC004DFFC8FC142E9316E43396" box="[923,1017,907,934]" pageId="7" pageNumber="466" value="1985-01-02">
<collectingDate id="AB213C54004DFFC8FC142E9316E43396" box="[923,1017,907,934]" pageId="7" pageNumber="466" value="1985-01-02">2.i.1985</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FBB72E9411CF3395" box="[1080,1234,907,934]" pageId="7" pageNumber="466">James Adams</collectorName>
(1
<collectionCode id="A9CA7BB9004DFFC8FB752E9310563395" box="[1274,1355,907,933]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="7" pageNumber="466" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FAEF2E9416DE33FA" ID-GBIF-Occurrence="3128194478" collectingDate="1965-08-27" collectionCode="NMNH" collectorName="W. Poza Rica &amp; Paul J. Spangler" country="Mexico" location="Veracruz" pageId="7" pageNumber="466" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<collectingRegion id="0D1F2D9E004DFFC8FAEF2E9415D733FA" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="7" pageNumber="466">Veracruz</collectingRegion>
,
<quantity id="08234E99004DFFC8FF592EB7140D33F9" box="[214,272,943,969]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="7" pageNumber="466" unit="mi" value="9.0">9 mi.</quantity>
<collectorName id="622E86AA004DFFC8FE982EA814AC33F9" box="[279,433,943,970]" pageId="7" pageNumber="466">W. Poza Rica</collectorName>
,
<date id="BB65C5BC004DFFC8FE322EB7175A33FA" box="[445,583,943,970]" pageId="7" pageNumber="466" value="1965-08-27">
<collectingDate id="AB213C54004DFFC8FE322EB7175A33FA" box="[445,583,943,970]" pageId="7" pageNumber="466" value="1965-08-27">27.viii.1965</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FD0B2EB7162733F9" box="[644,826,943,970]" pageId="7" pageNumber="466">Paul J. Spangler</collectorName>
(6
<collectionCode id="A9CA7BB9004DFFC8FCEF2EA816A633FA" box="[864,955,944,970]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="466" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FC412EB714A533DD" ID-GBIF-Occurrence="3128194506" collectingDate="1998-12-02" collectionCode="IEXA" collectorName="A. Gomez" country="Mexico" location="Almagres" municipality="Sayula de Aleman" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="2F007906004DFFC8FC412EB7118233F9" box="[974,1183,943,969]" pageId="7" pageNumber="466">Sayula de Alemán</collectingMunicipality>
,
<location id="CA04B5A7004DFFC8FB272EB7100533F9" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FB272EB7100533F9" box="[1192,1304,943,969]" country="Mexico" municipality="Sayula de Aleman" name="Almagres" pageId="7" pageNumber="466">Almagres</location>
,
<date id="BB65C5BC004DFFC8FAAB2EB7108533F9" box="[1316,1432,943,969]" pageId="7" pageNumber="466" value="1998-12-02">
<collectingDate id="AB213C54004DFFC8FAAB2EB7108533F9" box="[1316,1432,943,969]" pageId="7" pageNumber="466" value="1998-12-02">2.xii.1998</collectingDate>
</date>
,
<collectorName id="622E86AA004DFFC8FF182ECB141133DD" box="[151,268,979,1005]" pageId="7" pageNumber="466">A. Gómez</collectorName>
col. (1
<collectionCode id="A9CA7BB9004DFFC8FEE72ECB14AD33DD" box="[360,432,979,1005]" pageId="7" pageNumber="466">IEXA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="7FB3E921004DFFC8FE4B2ECB102933DE" ID-GBIF-Occurrence="3128194465" box="[452,1332,979,1006]" collectingDate="1971-07-01" collectingDateMax="1971-07-30" collectingDateMin="1971-07-01" collectionCode="NMNH" collectorName="J. Maldonado" country="Nicaragua" location="Puerto" pageId="7" pageNumber="466" specimenCount="1" typeStatus="paratype">
<emphasis id="FDAF3F6E004DFFC8FE4B2ECB175D33DD" bold="true" box="[452,576,979,1005]" pageId="7" pageNumber="466">
<collectingCountry id="B7CCA3EC004DFFC8FE4B2ECB175D33DD" box="[452,576,979,1005]" name="Nicaragua" pageId="7" pageNumber="466">Nicaragua</collectingCountry>
</emphasis>
:
<location id="CA04B5A7004DFFC8FDDD2ECC178633DE" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FDDD2ECC178633DE" box="[594,667,980,1006]" country="Nicaragua" name="Puerto" pageId="7" pageNumber="466">Puerto</location>
Cabezas,
<date id="BB65C5BC004DFFC8FC9E2ECB16B733DD" box="[785,938,979,1005]" pageId="7" pageNumber="466" value="1971-07-01" valueMax="1971-07-30" valueMin="1971-07-01">
<collectingDate id="AB213C54004DFFC8FC9E2ECB16B733DD" box="[785,938,979,1005]" pageId="7" pageNumber="466" value="1971-07-01" valueMax="1971-07-30" valueMin="1971-07-01">1-30.vii.1971</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FC652ECC119833DD" box="[1002,1157,979,1006]" pageId="7" pageNumber="466">J. Maldonado</collectorName>
C. (2
<collectionCode id="A9CA7BB9004DFFC8FB5E2ECC103133DE" box="[1233,1324,980,1006]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="466" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FACE2ECB10033421" ID-GBIF-Occurrence="3128194499" collectingDate="2002-08-05" collectingDateMax="2002-08-10" collectingDateMin="2002-08-05" collectionCode="SEMC" collectorName="W. D. Shepard" country="Nicaragua" latitude="10.983334" location="Refugio Bartola" longLatPrecision="1296" longitude="-84.333336" pageId="7" pageNumber="466" specimenCount="1" stateProvince="Rio San Juan" typeStatus="paratype">
<collectingRegion id="0D1F2D9E004DFFC8FACE2ECB15D03422" country="Nicaragua" name="Rio San Juan" pageId="7" pageNumber="466">Rio San Juan</collectingRegion>
,
<collectingRegion id="0D1F2D9E004DFFC8FF582EEF14763422" box="[215,363,1015,1042]" country="Nicaragua" name="Rio San Juan" pageId="7" pageNumber="466">Rio San Juan</collectingRegion>
,
<location id="CA04B5A7004DFFC8FEFA2EEF17373421" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FEFA2EEF17373421" box="[373,554,1015,1041]" country="Nicaragua" latitude="10.983334" longLatPrecision="1296" longitude="-84.333336" name="Refugio Bartola" pageId="7" pageNumber="466" stateProvince="Rio San Juan">Refugio Bartola</location>
, (
<geoCoordinate id="AAEF85BB004DFFC8FDB32EEF17BD3422" box="[572,672,1015,1042]" degrees="10" direction="north" minutes="59" orientation="latitude" pageId="7" pageNumber="466" precision="925" value="10.983334">10°59’N</geoCoordinate>
,
<geoCoordinate id="AAEF85BB004DFFC8FD252EEF16083421" box="[682,789,1015,1041]" degrees="84" direction="west" minutes="20" orientation="longitude" pageId="7" pageNumber="466" precision="925" value="-84.333336">84°20’W</geoCoordinate>
),
<date id="BB65C5BC004DFFC8FCA82EEF16D53422" box="[807,968,1015,1042]" pageId="7" pageNumber="466" value="2002-08-05" valueMax="2002-08-10" valueMin="2002-08-05">
<collectingDate id="AB213C54004DFFC8FCA82EEF16D53422" box="[807,968,1015,1042]" pageId="7" pageNumber="466" value="2002-08-05" valueMax="2002-08-10" valueMin="2002-08-05">5-10.viii.2002</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FB8E2EE011BF3421" box="[1025,1186,1015,1042]" pageId="7" pageNumber="466">W. D. Shepard</collectorName>
(2
<collectionCode id="A9CA7BB9004DFFC8FB4B2EEF10083421" box="[1220,1301,1015,1041]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="7" pageNumber="466" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921004DFFC8FAA82EEF14AD346A" ID-GBIF-Occurrence="3128194330" collectingDate="2013-02-13" collectionCode="NMW" collectorName="M. Brojer" country="Nicaragua" county="Rio Ochomogo" elevation="60" latitude="11.657136" location="Paso Real de Ochomogo" longLatPrecision="1" longitude="-85.97083" municipality="Ochomogo" pageId="7" pageNumber="466" specimenCount="1" stateProvince="Rivas" typeStatus="paratype">
<collectingRegion id="0D1F2D9E004DFFC8FAA82EEF10773421" box="[1319,1386,1015,1041]" country="Nicaragua" name="Rivas" pageId="7" pageNumber="466">Rivas</collectingRegion>
,
<collectingCounty id="26059BF0004DFFC8FAFB2EEF140B3405" pageId="7" pageNumber="466">Rio Ochomogo</collectingCounty>
, in
<collectingMunicipality id="2F007906004DFFC8FECD290314A33405" box="[322,446,1051,1077]" pageId="7" pageNumber="466">Ochomogo</collectingMunicipality>
at
<location id="CA04B5A7004DFFC8FE6B290417E23405" LSID="urn:lsid:plazi:treatment:4772526A004DFFC7FF182D8F11883229:CA04B5A7004DFFC8FE6B290417E23405" box="[484,767,1051,1078]" country="Nicaragua" county="Rio Ochomogo" latitude="11.657136" longLatPrecision="1" longitude="-85.97083" municipality="Ochomogo" name="Paso Real de Ochomogo" pageId="7" pageNumber="466" stateProvince="Rivas">Paso Real de Ochomogo</location>
, elev.
<quantity id="08234E99004DFFC8FCC62903169E3406" box="[841,899,1051,1078]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="466" unit="m" value="60.0">
<elevation id="44F6044F004DFFC8FCC62903169E3406" box="[841,899,1051,1078]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="466" unit="m" value="60.0">60 m</elevation>
</quantity>
(
<geoCoordinate id="AAEF85BB004DFFC8FC1B2903115A3406" box="[916,1095,1051,1078]" degrees="11" direction="north" minutes="39" orientation="latitude" pageId="7" pageNumber="466" precision="1" seconds="25.69" value="11.657136">11°39’25.69’’N</geoCoordinate>
,
<geoCoordinate id="AAEF85BB004DFFC8FBDB290310133406" box="[1108,1294,1051,1078]" degrees="85" direction="west" minutes="58" orientation="longitude" pageId="7" pageNumber="466" precision="1" seconds="15.01" value="-85.97083">85°58’15.01’’W</geoCoordinate>
),
<date id="BB65C5BC004DFFC8FAAB290310853405" box="[1316,1432,1051,1077]" pageId="7" pageNumber="466" value="2013-02-13">
<collectingDate id="AB213C54004DFFC8FAAB290310853405" box="[1316,1432,1051,1077]" pageId="7" pageNumber="466" value="2013-02-13">13.ii.2013</collectingDate>
</date>
, leg.
<collectorName id="622E86AA004DFFC8FF482958142B3469" box="[199,310,1087,1114]" pageId="7" pageNumber="466">M. Brojer</collectorName>
(2
<collectionCode id="A9CA7BB9004DFFC8FED4295814B5346A" box="[347,424,1088,1114]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="466">NMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7004DFFC8FF48297B16513639" pageId="7" pageNumber="466" type="diagnosis">
<paragraph id="CF64E37C004DFFC8FF48297B16513639" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="466">
<emphasis id="FDAF3F6E004DFFC8FF48297B14D0344D" bold="true" box="[199,461,1123,1149]" pageId="7" pageNumber="466">Differential diagnosis.</emphasis>
Among Central American species, recognized by the combination of moderate size (ca.
<quantity id="08234E99004DFFC8FF18299F15E23492" box="[151,255,1159,1186]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="466" unit="mm" value="3.2">3.20 mm</quantity>
), the moderately low and relatively impunctate pronotal reliefs, the dull, minutely roughened dorsum, and the male genitalia (
<figureCitation id="57E0FFF9004DFFC8FEE129B314AE34F6" box="[366,435,1195,1222]" captionStart="FIGURES 5–6" captionStartId="24.[151,264,1934,1958]" captionTargetBox="[161,1425,192,1893]" captionTargetId="figure-22@24.[151,1435,181,1910]" captionTargetPageId="24" captionText="FIGURES 5–6. Habitus and male genitalia of holotypes. 5. Hydrochus biexcavatus n. sp. 6. Hydrochus mesoamericanus n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777855" httpUri="https://zenodo.org/record/4777855/files/figure.png" pageId="7" pageNumber="466">Fig. 5</figureCitation>
; described below). This species has a generalized
<taxonomicName id="08DB98FF004DFFC8FC6829B3117C34F5" authorityName="Leach" authorityYear="1817" box="[999,1121,1195,1221]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="466" phylum="Arthropoda" rank="genus">
<emphasis id="FDAF3F6E004DFFC8FC6829B3117C34F5" box="[999,1121,1195,1221]" italics="true" pageId="7" pageNumber="466">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males. Differences of the male genitalia from that of the somewhat superficially similar
<taxonomicName id="08DB98FF004DFFC8FF6129EC144E353D" box="[238,339,1268,1293]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="466" phylum="Arthropoda" rank="species" species="vagus">
<emphasis id="FDAF3F6E004DFFC8FF6129EC144E353D" box="[238,339,1268,1293]" italics="true" pageId="7" pageNumber="466">H. vagus</emphasis>
</taxonomicName>
include, in
<emphasis id="FDAF3F6E004DFFC8FE5129EC1790353D" box="[478,653,1267,1293]" italics="true" pageId="7" pageNumber="466">
<taxonomicName id="08DB98FF004DFFC8FE5129EC1794353D" authorityName="Perkins" authorityYear="2021" box="[478,649,1267,1293]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="466" phylum="Arthropoda" rank="species" species="biexcavatus">H. biexcavatus</taxonomicName>
,
</emphasis>
the wider aedeagus with a much larger distal membranous area, the differently shaped dorsal margin of the aedeagus, the larger and differently shaped adbl, the more widely separated pdmm, and the differently shaped paramere tips, each with excavate dorsal surface (
<figureCitation id="57E0FFF9004DFFC8FBCB2823118E3566" box="[1092,1171,1339,1366]" captionStart="FIGURES 5–6" captionStartId="24.[151,264,1934,1958]" captionTargetBox="[161,1425,192,1893]" captionTargetId="figure-22@24.[151,1435,181,1910]" captionTargetPageId="24" captionText="FIGURES 5–6. Habitus and male genitalia of holotypes. 5. Hydrochus biexcavatus n. sp. 6. Hydrochus mesoamericanus n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777855" httpUri="https://zenodo.org/record/4777855/files/figure.png" pageId="7" pageNumber="466">Figs. 5</figureCitation>
,
<figureCitation id="57E0FFF9004DFFC8FB10282311A33566" box="[1183,1214,1339,1366]" captionStart="FIGURES 15–16" captionStartId="29.[151,264,1944,1968]" captionTargetBox="[181,1432,185,1916]" captionTargetId="figure-15@29.[151,1435,181,1920]" captionTargetPageId="29" captionText="FIGURES 15–16. Habitus and male genitalia. 15. Hydrochus foveatus Haldeman, non-type. 16. Hydrochus foveatus Haldeman, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4777875" httpUri="https://zenodo.org/record/4777875/files/figure.png" pageId="7" pageNumber="466">16</figureCitation>
). Differences of the male genitalia from that of the somewhat superficially similar
<taxonomicName id="08DB98FF004DFFC8FCD6287816EA3549" box="[857,1015,1375,1401]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="466" phylum="Arthropoda" rank="species" species="vanvondeli">
<emphasis id="FDAF3F6E004DFFC8FCD6287816EA3549" box="[857,1015,1375,1401]" italics="true" pageId="7" pageNumber="466">H. vanvondeli</emphasis>
</taxonomicName>
include, in
<emphasis id="FDAF3F6E004DFFC8FBF2287810363549" box="[1149,1323,1375,1401]" italics="true" pageId="7" pageNumber="466">
<taxonomicName id="08DB98FF004DFFC8FBF22878103A3549" authorityName="Perkins" authorityYear="2021" box="[1149,1319,1375,1401]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="466" phylum="Arthropoda" rank="species" species="biexcavatus">H. biexcavatus</taxonomicName>
,
</emphasis>
the wider aedeagus with a larger distal membranous area and having the agw nearer the aedeagal base, the longer aedeagus, compared to the length of the parameres, the aedeagus tip extending beyond the level of the plsa (in
<taxonomicName id="08DB98FF004DFFC8FB7028B0108135F1" box="[1279,1436,1447,1473]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="466" phylum="Arthropoda" rank="species" species="vanvondeli">
<emphasis id="FDAF3F6E004DFFC8FB7028B0108135F1" box="[1279,1436,1447,1473]" italics="true" pageId="7" pageNumber="466">H. vanvondeli</emphasis>
</taxonomicName>
the tip only attains the same level as the plsa), the larger and differently shaped adbl, and the differently shaped paramere tips, each with excavate dorsal surface (
<figureCitation id="57E0FFF9004DFFC8FD4B28F71609363A" box="[708,788,1519,1546]" captionStart="FIGURES 5–6" captionStartId="24.[151,264,1934,1958]" captionTargetBox="[161,1425,192,1893]" captionTargetId="figure-22@24.[151,1435,181,1910]" captionTargetPageId="24" captionText="FIGURES 5–6. Habitus and male genitalia of holotypes. 5. Hydrochus biexcavatus n. sp. 6. Hydrochus mesoamericanus n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777855" httpUri="https://zenodo.org/record/4777855/files/figure.png" pageId="7" pageNumber="466">Figs. 5</figureCitation>
,
<figureCitation id="57E0FFF9004DFFC8FCAF28F7165D3639" box="[800,832,1519,1545]" captionStart="FIGURES 23–24" captionStartId="33.[151,264,1898,1922]" captionTargetBox="[166,1428,185,1843]" captionTargetId="figure-15@33.[151,1435,181,1874]" captionTargetPageId="33" captionText="FIGURES 23–24. Habitus and male genitalia of holotypes. 23. Hydrochus vanvondeli Makhan. 24. Hydrochus beeneni Makhan." figureDoi="http://doi.org/10.5281/zenodo.4777897" httpUri="https://zenodo.org/record/4777897/files/figure.png" pageId="7" pageNumber="466">23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7004DFFC7FF782B0B143931C5" lastPageId="8" lastPageNumber="467" pageId="7" pageNumber="466" type="description">
<paragraph id="CF64E37C004DFFC8FF782B0B167636D1" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="466">
<emphasis id="FDAF3F6E004DFFC8FF782B0B149C361D" bold="true" box="[247,385,1555,1581]" pageId="7" pageNumber="466">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="57E0FFF9004DFFC8FD052B0B17D3361E" box="[650,718,1555,1582]" captionStart="FIGURES 5–6" captionStartId="24.[151,264,1934,1958]" captionTargetBox="[161,1425,192,1893]" captionTargetId="figure-22@24.[151,1435,181,1910]" captionTargetPageId="24" captionText="FIGURES 5–6. Habitus and male genitalia of holotypes. 5. Hydrochus biexcavatus n. sp. 6. Hydrochus mesoamericanus n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777855" httpUri="https://zenodo.org/record/4777855/files/figure.png" pageId="7" pageNumber="466">Fig. 5</figureCitation>
). Size:
<typeStatus id="10605DDE004DFFC8FCAE2B0B169E361D" box="[801,899,1555,1581]" pageId="7" pageNumber="466" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 3.17/1.10; head width 0.73; pronotum l/w 0.83/0.73; PA 0.68; PB 0.61; elytra 2.05/1.10. Body size moderately large (ca.
<quantity id="08234E99004DFFC8FE852B4314693646" box="[266,372,1627,1654]" metricMagnitude="-3" metricUnit="m" metricValue="3.17" pageId="7" pageNumber="466" unit="mm" value="3.17">3.17 mm</quantity>
). Dorsum golden brown, with very little iridescence, cuticle minutely roughened, dull, elytra without black spots. Punctation on elytra ca. 2–3x that of pronotum. Elytra interstriae ca. 0.5–1× strial puncture diameter. Interstria 9
<superScript id="38AE4E34004DFFC8FE0E2BBB14933681" attach="left" box="[385,398,1699,1713]" fontSize="6" pageId="7" pageNumber="466">th</superScript>
more convex than others, overhanging 10
<superScript id="38AE4E34004DFFC8FCE52BBB166A3681" attach="left" box="[874,887,1699,1713]" fontSize="6" pageId="7" pageNumber="466">th</superScript>
interstria. Usual area of callus on 5
<superScript id="38AE4E34004DFFC8FA852BBB100A3681" attach="left" box="[1290,1303,1699,1713]" fontSize="6" pageId="7" pageNumber="466">th</superScript>
moderately raised, moderately elongate, ca. 5 punctures in row on each side.
</paragraph>
<paragraph id="CF64E37C004DFFC8FF482BF41440377D" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="466">Pronotum slightly wider than long, widest slightly behind anterior margin, narrowed at base, sides arcuate, minutely denticulate; anterior margin bisinuate; densely moderately coarsely punctate, except less dense on reliefs; depressions deep.</paragraph>
<paragraph id="CF64E37C004DFFC8FF482A4F166637A5" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="466">Elytra with spaces between strial punctures ca. 0.5–1x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with slight angulation.</paragraph>
<paragraph id="CF64E37C004DFFC8FF482A87112C37ED" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="466">Ventral characters: Mentum coarsely densely punctate, with large median fovea. Submentum with two small foveae. Legs testaceous, femora and tibiofemoral joints slightly darker than tibiae.</paragraph>
<paragraph id="CF64E37C0042FFC7FF182D8F101F3155" blockId="8.[151,1436,150,537]" pageId="8" pageNumber="467">
Male genitalia (
<figureCitation id="57E0FFF90042FFC7FEC62D8F14913081" box="[329,396,150,177]" captionStart="FIGURES 5–6" captionStartId="24.[151,264,1934,1958]" captionTargetBox="[161,1425,192,1893]" captionTargetId="figure-22@24.[151,1435,181,1910]" captionTargetPageId="24" captionText="FIGURES 5–6. Habitus and male genitalia of holotypes. 5. Hydrochus biexcavatus n. sp. 6. Hydrochus mesoamericanus n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777855" httpUri="https://zenodo.org/record/4777855/files/figure.png" pageId="8" pageNumber="467">Fig. 5</figureCitation>
) general characters: paramere tips slender and pointed, narrower in lateral view than in dorsal/ventral views, in dorsal/ventral views with medial margins at apex parallel or nearly so, then arcuate away from midline, immediately proximal to plsa with dorsal surface excavate; aedeagus in dorsal/ventral views lanceolate, widest slightly proximal to midlength; in lateral view slightly narrower in distal ½, and wider in proximal 1/2, with dorsal margin slightly undulating; lateral margins of aedeagus sclerotized; basal piece length equal to or very slightly shorter than parameres, tapering slightly from distal to proximal, orifice strongly sclerotized.
</paragraph>
<paragraph id="CF64E37C0042FFC7FF182C7711F0319D" blockId="8.[151,1436,150,537]" pageId="8" pageNumber="467">Dorsal surface: pdmm widely separated, overlapping aedeagus only near base; adbl large; adtl at about midlength; each paramere with small excavation just proximal to psla, paramere tips without sclerotized ridge.</paragraph>
<paragraph id="CF64E37C0042FFC7FF182CAF143931C5" blockId="8.[151,1436,150,537]" pageId="8" pageNumber="467">Ventral surface: pvmm approximating lateral margin of aedeagus, except overlapping basally; avtl very near base of aedeagus.</paragraph>
</subSubSection>
<subSubSection id="87C1B0F70042FFC7FF482CE711883229" box="[199,1173,511,537]" pageId="8" pageNumber="467" type="etymology">
<paragraph id="CF64E37C0042FFC7FF482CE711883229" blockId="8.[151,1436,150,537]" box="[199,1173,511,537]" pageId="8" pageNumber="467">
<emphasis id="FDAF3F6E0042FFC7FF482CE714533229" bold="true" box="[199,334,511,537]" pageId="8" pageNumber="467">Etymology.</emphasis>
Named in reference to the excavations of the male genitalia paramere tips.
</paragraph>
</subSubSection>
</treatment>
</document>
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