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<document id="9FAE89122F8B2D3B8685339EE61041E2" ID-CLB-Dataset="6214" ID-DOI="10.11646/zootaxa.4974.3.2" ID-GBIF-Dataset="5b4eaeb3-5722-4021-84ad-125ccd80855f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4777839" ID-ZooBank="1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621588618725" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="4772526A0044FFC0FF1828DC108032FC" docLanguage="en" docName="zootaxa.4974.3.2.pdf" docOrigin="Zootaxa 4974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus brianbrowni Makhan" docType="treatment" docVersion="7" lastPageNumber="474" masterDocId="BB4B2A12004AFFCFFF8F2D18151D3030" masterDocTitle="Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="503" masterPageNumber="459" pageNumber="473" updateTime="1698940277359" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="6B88F70E2F6B69CE9951D2E8A7037116">
<mods:title id="577A41A8E6A11FB12171FB8C69155D5F">Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="831E8C5698F3B6E7E8556C1F4D130AF5">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="5B1A23053F00CC299FCBCF3ACDDACFE4">text</mods:typeOfResource>
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<mods:title id="0843DFA2DB3D3FA4795DB7E793AD057B">Zootaxa</mods:title>
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<mods:part id="2C3A3D12CE12F59EAA121AE1A77D774D">
<mods:date id="DD327672E4145DDF7BEC8C31CCBC4F96">2021</mods:date>
<mods:detail id="B2BDEBB0E0BCF21F161434864628B265" type="pubDate">
<mods:number id="1841AF5AD04F6A3B47753BCC0891B6D2">2021-05-21</mods:number>
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<mods:detail id="F1957C75944A7F8DAB0140C31FFC3171" type="volume">
<mods:number id="C2261EAC3D3850E33E1356A24B8EBD61">4974</mods:number>
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<mods:number id="88ADDD54EE50C251FA4502B7DDD26855">3</mods:number>
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<mods:start id="1174197756209BEDA98640785C6D45DF">459</mods:start>
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<mods:classification id="C1E6462E395FFE727B571C6FB624B988">journal article</mods:classification>
<mods:identifier id="B5D62E4950288D093C54B980DE989A8D" type="CLB-Dataset">6214</mods:identifier>
<mods:identifier id="B163FB6EA96E0B1E29C0EF5420C6177A" type="DOI">10.11646/zootaxa.4974.3.2</mods:identifier>
<mods:identifier id="5B900EBD39FE54A07C7638F8102A1DBC" type="GBIF-Dataset">5b4eaeb3-5722-4021-84ad-125ccd80855f</mods:identifier>
<mods:identifier id="BA9FD70FD326809DE0430D85541A57C6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8494623329DA951944AA5D67DFA0C642" type="Zenodo-Dep">4777839</mods:identifier>
<mods:identifier id="E134EFF75EFCB0BB064286E9975C6AB0" type="ZooBank">1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8</mods:identifier>
</mods:mods>
<treatment id="4772526A0044FFC0FF1828DC108032FC" ID-DOI="http://doi.org/10.5281/zenodo.4777906" ID-GBIF-Taxon="182642570" ID-Zenodo-Dep="4777906" LSID="urn:lsid:plazi:treatment:4772526A0044FFC0FF1828DC108032FC" httpUri="http://treatment.plazi.org/id/4772526A0044FFC0FF1828DC108032FC" lastPageId="15" lastPageNumber="474" pageId="14" pageNumber="473">
<subSubSection id="87C1B0F70044FFC1FF1828DC173635EF" box="[151,555,1476,1503]" pageId="14" pageNumber="473" type="nomenclature">
<paragraph id="CF64E37C0044FFC1FF1828DC173635EF" blockId="14.[151,555,1476,1539]" box="[151,555,1476,1503]" pageId="14" pageNumber="473">
<heading id="942C54100044FFC1FF1828DC173635EF" bold="true" box="[151,555,1476,1503]" fontSize="11" level="1" pageId="14" pageNumber="473" reason="1">
<taxonomicName id="08DB98FF0044FFC1FF1828DC173635EF" ID-CoL="6MJVF" authority="Makhan" authorityName="Makhan" box="[151,555,1476,1503]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="473" phylum="Arthropoda" rank="species" species="brianbrowni">
<emphasis id="FDAF3F6E0044FFC1FF1828DC173635EF" bold="true" box="[151,555,1476,1503]" pageId="14" pageNumber="473">
<emphasis id="FDAF3F6E0044FFC1FF1828DC14A735EF" bold="true" box="[151,442,1476,1503]" italics="true" pageId="14" pageNumber="473">Hydrochus brianbrowni</emphasis>
Makhan
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F70044FFC1FF1828F114343633" box="[151,297,1513,1539]" pageId="14" pageNumber="473" type="description">
<paragraph id="CF64E37C0044FFC1FF1828F114343633" blockId="14.[151,555,1476,1539]" box="[151,297,1513,1539]" pageId="14" pageNumber="473">
<figureCitation id="57E0FFF90044FFC1FF1828F115F83633" box="[151,229,1513,1539]" captionStart="FIGURE 2" captionStartId="5.[151,250,1989,2013]" captionTargetBox="[181,1393,181,1953]" captionTargetId="figure-15@5.[158,1429,181,1965]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus of Central American Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4777847" httpUri="https://zenodo.org/record/4777847/files/figure.png" pageId="14" pageNumber="473">Figs. 2</figureCitation>
,
<figureCitation id="57E0FFF90044FFC1FF7E28F1141C3633" box="[241,257,1513,1539]" captionStart="FIGURES 9–10" captionStartId="26.[151,264,1944,1968]" captionTargetBox="[163,1430,185,1901]" captionTargetId="figure-22@26.[151,1435,181,1920]" captionTargetPageId="26" captionText="FIGURES 9–10. Habitus and male genitalia. 9. Hydrochus brianbrowni Makhan. 10. Hydrochus nayaritensis n. sp. holotype." figureDoi="http://doi.org/10.5281/zenodo.4777861" httpUri="https://zenodo.org/record/4777861/files/figure.png" pageId="14" pageNumber="473">9</figureCitation>
,
<figureCitation id="57E0FFF90044FFC1FE8228F114343633" box="[269,297,1513,1539]" captionStart="FIGURES 31–33" captionStartId="36.[151,264,1943,1967]" captionTargetBox="[170,1417,181,1920]" captionTargetId="figure-22@36.[170,1417,181,1920]" captionTargetPageId="36" captionText="FIGURES 31–33. Geographical distribution maps." figureDoi="http://doi.org/10.5281/zenodo.4777908" httpUri="https://zenodo.org/record/4777908/files/figure.png" pageId="14" pageNumber="473">31</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F70044FFC1FF182B3511F136BD" pageId="14" pageNumber="473" type="reference_group">
<paragraph id="CF64E37C0044FFC1FF182B3517543675" blockId="14.[151,585,1581,1605]" box="[151,585,1581,1605]" pageId="14" pageNumber="473">
<treatmentCitationGroup id="EFCBC4520044FFC1FF182B3517543675" box="[151,585,1581,1605]" pageId="14" pageNumber="473">
<treatmentCitation id="4E7AC56D0044FFC1FF182B3517583675" author="Makhan, D." box="[151,581,1581,1605]" page="63" pageId="14" pageNumber="473" year="2005">
<taxonomicName id="08DB98FF0044FFC1FF182B3517583675" ID-CoL="6MJVF" authority="Makhan 2005: 63" authorityName="Makhan" authorityPageNumber="63" authorityYear="2005" box="[151,581,1581,1605]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="473" phylum="Arthropoda" rank="species" species="brianbrowni">
<emphasis id="FDAF3F6E0044FFC1FF182B3514973675" box="[151,394,1581,1605]" italics="true" pageId="14" pageNumber="473">Hydrochus brianbrowni</emphasis>
<bibRefCitation id="AB4A9E8D0044FFC1FE1F2B3517583675" author="Makhan, D." box="[400,581,1581,1605]" pageId="14" pageNumber="473" pagination="63 - 65" refId="ref25126" refString="Makhan, D. (2005) Hydrochus brianbrowni sp. nov. (Coleoptera: Hydrochidae), a new American species. Baltic Journal of Coleopterology, 5, 63 - 65." type="journal article" year="2005">Makhan 2005: 63</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="CF64E37C0044FFC1FF182B6B11F136BD" blockId="14.[151,1437,1651,2038]" box="[151,1260,1651,1678]" pageId="14" pageNumber="473">
Originally described from southern Arizona. New locality data records for
<collectingCountry id="B7CCA3EC0044FFC1FC522B6B112E36BD" box="[989,1075,1651,1677]" name="Mexico" pageId="14" pageNumber="473">Mexico</collectingCountry>
are given below
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F70044FFC1FF482B8F17FD37C6" pageId="14" pageNumber="473" type="materials_examined">
<paragraph id="CF64E37C0044FFC1FF482B8F17FD37C6" blockId="14.[151,1437,1651,2038]" pageId="14" pageNumber="473">
<emphasis id="FDAF3F6E0044FFC1FF482B8F14F53681" bold="true" box="[199,488,1687,1713]" pageId="14" pageNumber="473">Material examined (52):</emphasis>
<materialsCitation id="7FB3E9210044FFC1FE602B8F149F36E6" ID-GBIF-Occurrence="3128194473" collectingDate="1959-01-23" collectionCode="CAS" collectorName="Las Cruces, E. &amp; H. B. Leech" country="Mexico" location="La Paz" municipality="Arroyo Saltito" pageId="14" pageNumber="473" specimenCount="1" stateProvince="Baja California">
<emphasis id="FDAF3F6E0044FFC1FE602B8F175B3681" bold="true" box="[495,582,1687,1713]" pageId="14" pageNumber="473">
<collectingCountry id="B7CCA3EC0044FFC1FE602B8F175B3681" box="[495,582,1687,1713]" name="Mexico" pageId="14" pageNumber="473">Mexico</collectingCountry>
</emphasis>
:
<collectingRegion id="0D1F2D9E0044FFC1FDDA2B8F161E3681" box="[597,771,1687,1713]" country="Mexico" name="Baja California" pageId="14" pageNumber="473">Baja California</collectingRegion>
,
<collectingMunicipality id="2F0079060044FFC1FC822B8016B33681" box="[781,942,1687,1714]" pageId="14" pageNumber="473">Arroyo Saltito</collectingMunicipality>
nr.
<collectorName id="622E86AA0044FFC1FC572B8011613682" box="[984,1148,1688,1714]" pageId="14" pageNumber="473">Las Cruces, E.</collectorName>
of
<location id="CA04B5A70044FFC1FB2E2B8011ED3682" LSID="urn:lsid:plazi:treatment:4772526A0044FFC0FF1828DC108032FC:CA04B5A70044FFC1FB2E2B8011ED3682" box="[1185,1264,1688,1714]" country="Mexico" municipality="Arroyo Saltito" name="La Paz" pageId="14" pageNumber="473" stateProvince="Baja California">La Paz</location>
,
<date id="BB65C5BC0044FFC1FB742B8F107A3682" box="[1275,1383,1687,1714]" pageId="14" pageNumber="473" value="1959-01-23">
<collectingDate id="AB213C540044FFC1FB742B8F107A3682" box="[1275,1383,1687,1714]" pageId="14" pageNumber="473" value="1959-01-23">23.i.1959</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0044FFC1FF182BA4140136E5" box="[151,284,1723,1750]" pageId="14" pageNumber="473">H. B. Leech</collectorName>
(1
<collectionCode id="A9CA7BB90044FFC1FEB02BA4146736E6" box="[319,378,1724,1750]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="473" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E9210044FFC1FE022BA3112236CA" ID-GBIF-Occurrence="3128194546" collectingDate="1968-10-30" collectingDateMax="1971-08-02" collectingDateMin="1968-10-30" collectionCode="FSCA, CAS" collectorName="H. G. Real &amp; R. E. Main" country="Mexico" county="Canyon San Bernardo" elevation="457" latitude="23.466667" location="Sa. Victoria" longLatPrecision="1255" longitude="-109.833336" municipality="Potrero" pageId="14" pageNumber="473" specimenCount="1" stateProvince="Baja California Sur">
<collectingRegion id="0D1F2D9E0044FFC1FE022BA3177B36E6" box="[397,614,1723,1750]" country="Mexico" name="Baja California Sur" pageId="14" pageNumber="473">Baja California Sur</collectingRegion>
,
<quantity id="08234E990044FFC1FDFF2BA417B436E5" box="[624,681,1723,1750]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="14" pageNumber="473" unit="mi" value="9.0">9 mi.</quantity>
up
<collectingCounty id="26059BF00044FFC1FD5F2BA416D836E5" box="[720,965,1723,1750]" pageId="14" pageNumber="473">Canyon San Bernardo</collectingCounty>
fr.
<collectorName id="622E86AA0044FFC1FC662BA411B836E5" box="[1001,1189,1723,1750]" pageId="14" pageNumber="473">Boca de la Sierra</collectorName>
(
<collectingMunicipality id="2F0079060044FFC1FB3C2BA4101636E6" box="[1203,1291,1724,1750]" pageId="14" pageNumber="473">Potrero</collectingMunicipality>
), elev.
<quantity id="08234E990044FFC1FAD82BA3108136E6" box="[1367,1436,1723,1750]" metricMagnitude="2" metricUnit="m" metricValue="4.57" pageId="14" pageNumber="473" unit="m" value="457.0">
<elevation id="44F6044F0044FFC1FAD82BA3108136E6" box="[1367,1436,1723,1750]" metricMagnitude="2" metricUnit="m" metricValue="4.57" pageId="14" pageNumber="473" unit="m" value="457.0">457 m</elevation>
</quantity>
(
<geoCoordinate id="AAEF85BB0044FFC1FF2F2BF8141936CA" box="[160,260,1760,1786]" degrees="23" direction="north" minutes="23" orientation="latitude" pageId="14" pageNumber="473" precision="925" value="23.383333">23°23’N</geoCoordinate>
,
<geoCoordinate id="AAEF85BB0044FFC1FE812BC7149A36CA" box="[270,391,1759,1786]" degrees="109" direction="west" minutes="57" orientation="longitude" pageId="14" pageNumber="473" precision="925" value="-109.95">109°57’W</geoCoordinate>
),
<date id="BB65C5BC0044FFC1FE152BC7170836C9" box="[410,533,1759,1785]" pageId="14" pageNumber="473" value="1971-08-02">
<collectingDate id="AB213C540044FFC1FE152BC7170836C9" box="[410,533,1759,1785]" pageId="14" pageNumber="473" value="1971-08-02">2.viii.1971</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0044FFC1FDC02BF817D836C9" box="[591,709,1759,1786]" pageId="14" pageNumber="473">H. G. Real</collectorName>
&amp;
<collectorName id="622E86AA0044FFC1FD682BF8167D36C9" box="[743,864,1759,1786]" pageId="14" pageNumber="473">R. E. Main</collectorName>
(1
<collectionCode id="A9CA7BB90044FFC1FC0C2BF816D136CA" box="[899,972,1760,1786]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="14" pageNumber="473" type="Museum">FSCA</collectionCode>
, 37
<collectionCode id="A9CA7BB90044FFC1FC772BF8112E36CA" box="[1016,1075,1760,1786]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="473" type="Museum">CAS</collectionCode>
);
</materialsCitation>
<materialsCitation id="7FB3E9210044FFC1FBC92BF81112372E" ID-GBIF-Occurrence="3128194358" collectingDate="1968-10-30" collectingDateMax="1971-08-02" collectingDateMin="1968-10-30" collectionCode="FSCA, CAS" collectorName="E. L. Sleeper &amp; F. J. Moore" country="Mexico" county="Canyon San Bernardo" elevation="457" latitude="23.466667" location="Sa. Victoria" longLatPrecision="1255" longitude="-109.833336" municipality="Potrero" pageId="14" pageNumber="473" specimenCount="1" stateProvince="Baja California Sur">
<location id="CA04B5A70044FFC1FBC92BF811D636C9" LSID="urn:lsid:plazi:treatment:4772526A0044FFC0FF1828DC108032FC:CA04B5A70044FFC1FBC92BF811D636C9" box="[1094,1227,1759,1786]" country="Mexico" county="Canyon San Bernardo" latitude="23.466667" longLatPrecision="1255" longitude="-109.833336" municipality="Potrero" name="Sa. Victoria" pageId="14" pageNumber="473" stateProvince="Baja California Sur">Sa. Victoria</location>
,
<locationDeviation id="DED6FA5E0044FFC1FB5A2BF8108536C9" box="[1237,1432,1759,1786]" location="Santiago" pageId="14" pageNumber="473">
<quantity id="08234E990044FFC1FB5A2BF8101336C9" box="[1237,1294,1759,1786]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="14" pageNumber="473" unit="mi" value="7.0">7 mi.</quantity>
W Santiago
</locationDeviation>
, (
<geoCoordinate id="AAEF85BB0044FFC1FF2F2A1C1419372E" box="[160,260,1796,1822]" degrees="23" direction="north" minutes="28" orientation="latitude" pageId="14" pageNumber="473" precision="925" value="23.466667">23°28’N</geoCoordinate>
,
<geoCoordinate id="AAEF85BB0044FFC1FE802A1B1495372E" box="[271,392,1795,1822]" degrees="109" direction="west" minutes="50" orientation="longitude" pageId="14" pageNumber="473" precision="925" value="-109.833336">109°50’W</geoCoordinate>
),
<date id="BB65C5BC0044FFC1FE132A1B1729372D" box="[412,564,1795,1821]" pageId="14" pageNumber="473" value="1968-10-30" valueMax="1968-10-31" valueMin="1968-10-30">
<collectingDate id="AB213C540044FFC1FE132A1B1729372D" box="[412,564,1795,1821]" pageId="14" pageNumber="473" value="1968-10-30" valueMax="1968-10-31" valueMin="1968-10-30">30-31.x.1968</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0044FFC1FDFF2A1C161E372D" box="[624,771,1795,1822]" pageId="14" pageNumber="473">E. L. Sleeper</collectorName>
&amp;
<collectorName id="622E86AA0044FFC1FCA82A1C16BA372E" box="[807,935,1796,1822]" pageId="14" pageNumber="473">F. J. Moore</collectorName>
(1
<collectionCode id="A9CA7BB90044FFC1FC432A1C111A372E" box="[972,1031,1796,1822]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="473" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E9210044FFC1FB942A1B17E63772" ID-GBIF-Occurrence="3128194520" collectingDate="1994-06-15" collectionCode="IEXA" collectorName="G. Nogueira" country="Mexico" elevation="120" location="Cajon de Penas" municipality="Mpio. Tomatlan" pageId="14" pageNumber="473" specimenCount="1" stateProvince="Jalisco">
<collectingRegion id="0D1F2D9E0044FFC1FB942A1B1177372D" box="[1051,1130,1795,1821]" country="Mexico" name="Jalisco" pageId="14" pageNumber="473">Jalisco</collectingRegion>
,
<collectingMunicipality id="2F0079060044FFC1FBF92A1B1031372D" box="[1142,1324,1795,1821]" pageId="14" pageNumber="473">Mpio. Tomatlán</collectingMunicipality>
,
<location id="CA04B5A70044FFC1FAB72A1B15C63772" LSID="urn:lsid:plazi:treatment:4772526A0044FFC0FF1828DC108032FC:CA04B5A70044FFC1FAB72A1B15C63772" country="Mexico" municipality="Mpio. Tomatlan" name="Cajon de Penas" pageId="14" pageNumber="473" stateProvince="Jalisco">Cajón de Peñas</location>
,
<quantity id="08234E990044FFC1FF672A30142F3772" box="[232,306,1832,1858]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="473" unit="m" value="120.0">
<elevation id="44F6044F0044FFC1FF672A30142F3772" box="[232,306,1832,1858]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="473" unit="m" value="120.0">120 m</elevation>
</quantity>
,
<date id="BB65C5BC0044FFC1FEB02A3F14A73772" box="[319,442,1831,1858]" pageId="14" pageNumber="473" value="1994-06-15">
<collectingDate id="AB213C540044FFC1FEB02A3F14A73772" box="[319,442,1831,1858]" pageId="14" pageNumber="473" value="1994-06-15">15.vi.1994</collectingDate>
</date>
,
<collectorName id="622E86AA0044FFC1FE492A30174F3771" box="[454,594,1831,1858]" pageId="14" pageNumber="473">G. Nogueira</collectorName>
col. (1
<collectionCode id="A9CA7BB90044FFC1FD232A3017E93772" box="[684,756,1832,1858]" pageId="14" pageNumber="473">IEXA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E9210044FFC1FC872A3F16C33772" ID-GBIF-Occurrence="3128194385" box="[776,990,1831,1858]" collectorName="Pto" country="Mexico" location="Quintana Roo" pageId="14" pageNumber="473" specimenCount="1" stateProvince="Quintana Roo">
<collectingRegion id="0D1F2D9E0044FFC1FC872A3F16B53772" box="[776,936,1831,1858]" country="Mexico" name="Quintana Roo" pageId="14" pageNumber="473">Quintana Roo</collectingRegion>
,
<collectorName id="622E86AA0044FFC1FC3A2A3016C33772" box="[949,990,1832,1858]" pageId="14" pageNumber="473">Pto</collectorName>
</materialsCitation>
.
<materialsCitation id="7FB3E9210044FFC1FC662A3F15E73756" ID-GBIF-Occurrence="3128194591" collectingDate="1982-04-29" collectionCode="UNAM" collectorName="O. Comul" country="Mexico" location="Morelos" pageId="14" pageNumber="473" specimenCount="1" stateProvince="Morelos">
<collectingRegion id="0D1F2D9E0044FFC1FC662A3F11543771" box="[1001,1097,1831,1857]" country="Mexico" name="Morelos" pageId="14" pageNumber="473">Morelos</collectingRegion>
,
<date id="BB65C5BC0044FFC1FBD92A3F11D33771" box="[1110,1230,1831,1857]" pageId="14" pageNumber="473" value="1982-04-29">
<collectingDate id="AB213C540044FFC1FBD92A3F11D33771" box="[1110,1230,1831,1857]" pageId="14" pageNumber="473" value="1982-04-29">29.iv.1982</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0044FFC1FA822A3010613771" box="[1293,1404,1831,1858]" pageId="14" pageNumber="473">O. Comul</collectorName>
(1
<collectionCode id="A9CA7BB90044FFC1FF182A5415EF3756" box="[151,242,1868,1894]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="14" pageNumber="473">UNAM</collectionCode>
)
</materialsCitation>
;
<collectingRegion id="0D1F2D9E0044FFC1FE892A5414A13755" box="[262,444,1867,1894]" country="Mexico" name="San Luis Potosi" pageId="14" pageNumber="473">San Luis Potosi</collectingRegion>
,
<quantity id="08234E990044FFC1FE462A5417183755" box="[457,517,1867,1894]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="14" pageNumber="473" unit="mi" value="2.0">2 mi.</quantity>
S.
<materialsCitation id="7FB3E9210044FFC1FDA32A5411833756" ID-GBIF-Occurrence="3128194567" box="[556,1182,1867,1894]" collectingDate="1948-11-21" collectionCode="CAS" collectorName="H. B. Leech" country="Mexico" location="San Luis Potosi" pageId="14" pageNumber="473" specimenCount="1" stateProvince="San Luis Potosi">
<collectingRegion id="0D1F2D9E0044FFC1FDA32A5417FF3755" box="[556,738,1867,1894]" country="Mexico" name="San Luis Potosi" pageId="14" pageNumber="473">San Luis Potosi</collectingRegion>
,
<date id="BB65C5BC0044FFC1FD602A5316753755" box="[751,872,1867,1893]" pageId="14" pageNumber="473" value="1948-11-21">
<collectingDate id="AB213C540044FFC1FD602A5316753755" box="[751,872,1867,1893]" pageId="14" pageNumber="473" value="1948-11-21">21.xi.1948</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0044FFC1FC272A54112F3755" box="[936,1074,1867,1894]" pageId="14" pageNumber="473">H. B. Leech</collectorName>
(1
<collectionCode id="A9CA7BB90044FFC1FBD42A54118B3756" box="[1115,1174,1868,1894]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="473" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E9210044FFC1FB242A53173937BA" ID-GBIF-Occurrence="3128194571" collectingDate="1922-07-08" collectionCode="CAS" collectorName="C. T. Dodds" country="Mexico" location="Los Mochis" pageId="14" pageNumber="473" specimenCount="1" stateProvince="Sinaloa">
<collectingRegion id="0D1F2D9E0044FFC1FB242A53101F3755" box="[1195,1282,1867,1893]" country="Mexico" name="Sinaloa" pageId="14" pageNumber="473">Sinaloa</collectingRegion>
,
<location id="CA04B5A70044FFC1FA802A5410853755" LSID="urn:lsid:plazi:treatment:4772526A0044FFC0FF1828DC108032FC:CA04B5A70044FFC1FA802A5410853755" box="[1295,1432,1867,1894]" country="Mexico" name="Los Mochis" pageId="14" pageNumber="473" stateProvince="Sinaloa">Los Mochis</location>
,
<date id="BB65C5BC0044FFC1FF182A77141637B9" box="[151,267,1903,1929]" pageId="14" pageNumber="473" value="1922-07-08">
<collectingDate id="AB213C540044FFC1FF182A77141637B9" box="[151,267,1903,1929]" pageId="14" pageNumber="473" value="1922-07-08">8.vii.1922</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0044FFC1FECD2A6814DD37B9" box="[322,448,1903,1930]" pageId="14" pageNumber="473">C.T. Dodds</collectorName>
(1
<collectionCode id="A9CA7BB90044FFC1FE6E2A68170137BA" box="[481,540,1904,1930]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="473" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E9210044FFC1FDA22A68108537BA" ID-GBIF-Occurrence="3128194551" box="[557,1432,1903,1930]" collectingDate="1980-09-28" collectionCode="NMNH" country="Mexico" location="Moctezuma" pageId="14" pageNumber="473" specimenCount="1" stateProvince="Sonora">
<collectingRegion id="0D1F2D9E0044FFC1FDA22A68176237BA" box="[557,639,1904,1930]" country="Mexico" name="Sonora" pageId="14" pageNumber="473">Sonora</collectingRegion>
,
<quantity id="08234E990044FFC1FD072A6817D337B9" box="[648,718,1903,1930]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="14" pageNumber="473" unit="mi" value="20.0">20 mi.</quantity>
W.
<location id="CA04B5A70044FFC1FD7A2A68166137BA" LSID="urn:lsid:plazi:treatment:4772526A0044FFC0FF1828DC108032FC:CA04B5A70044FFC1FD7A2A68166137BA" box="[757,892,1904,1930]" country="Mexico" name="Moctezuma" pageId="14" pageNumber="473" stateProvince="Sonora">Moctezuma</location>
,
<date id="BB65C5BC0044FFC1FC0A2A7716E237B9" box="[901,1023,1903,1929]" pageId="14" pageNumber="473" value="1980-09-28">
<collectingDate id="AB213C540044FFC1FC0A2A7716E237B9" box="[901,1023,1903,1929]" pageId="14" pageNumber="473" value="1980-09-28">28.ix.1980</collectingDate>
</date>
, leg. no collector on label (2
<collectionCode id="A9CA7BB90044FFC1FABA2A68108D37BA" box="[1333,1424,1904,1930]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="473" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E9210044FFC1FF182A8C163C379E" ID-GBIF-Occurrence="3128194338" box="[151,801,1939,1966]" collectingDate="1971-11-27" collectionCode="NMNH" collectorName="K. Stephan" country="Mexico" location="Alamos" pageId="14" pageNumber="473" specimenCount="1">
<quantity id="08234E990044FFC1FF182A8C15CF379D" box="[151,210,1939,1966]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="14" pageNumber="473" unit="mi" value="7.0">7 mi.</quantity>
SE
<location id="CA04B5A70044FFC1FF702A8B1444379D" LSID="urn:lsid:plazi:treatment:4772526A0044FFC0FF1828DC108032FC:CA04B5A70044FFC1FF702A8B1444379D" box="[255,345,1939,1965]" country="Mexico" name="Alamos" pageId="14" pageNumber="473">Alamos</location>
,
<date id="BB65C5BC0044FFC1FEEA2A8B14C2379D" box="[357,479,1939,1965]" pageId="14" pageNumber="473" value="1971-11-27">
<collectingDate id="AB213C540044FFC1FEEA2A8B14C2379D" box="[357,479,1939,1965]" pageId="14" pageNumber="473" value="1971-11-27">27.xi.1971</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0044FFC1FD932A8C1785379D" box="[540,664,1939,1966]" pageId="14" pageNumber="473">K. Stephan</collectorName>
(2
<collectionCode id="A9CA7BB90044FFC1FD302A8C1607379E" box="[703,794,1940,1966]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="473" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E9210044FFC1FCA22A8C1020379E" ID-GBIF-Occurrence="3128194344" box="[813,1341,1939,1966]" collectingDate="1986-11-06" collectionCode="UNAM" collectorName="F. Arias" country="Mexico" location="La Pintada" pageId="14" pageNumber="473" specimenCount="1">
<location id="CA04B5A70044FFC1FCA22A8C16B4379D" LSID="urn:lsid:plazi:treatment:4772526A0044FFC0FF1828DC108032FC:CA04B5A70044FFC1FCA22A8C16B4379D" box="[813,937,1939,1966]" country="Mexico" name="La Pintada" pageId="14" pageNumber="473">La Pintada</location>
,
<date id="BB65C5BC0044FFC1FC3B2A8B113D379D" box="[948,1056,1939,1965]" pageId="14" pageNumber="473" value="1986-11-06">
<collectingDate id="AB213C540044FFC1FC3B2A8B113D379D" box="[948,1056,1939,1965]" pageId="14" pageNumber="473" value="1986-11-06">6.xi.1986</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0044FFC1FBD22A8C11A9379D" box="[1117,1204,1939,1966]" pageId="14" pageNumber="473">F. Arias</collectorName>
(3
<collectionCode id="A9CA7BB90044FFC1FB552A8C1028379E" box="[1242,1333,1940,1966]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="14" pageNumber="473">UNAM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E9210044FFC1FAC72A8C17C037C6" ID-GBIF-Occurrence="3128194321" collectingDate="2004-01-31" collectionCode="NMNH" collectorName="V. Gomez &amp; J. L. Navarrete-Heredia" country="Mexico" elevation="1718" latitude="21.612806" location="Huitzila" longLatPrecision="1" longitude="-103.60616" municipality="Teul de Gonzalez Ortega" pageId="14" pageNumber="473" specimenCount="1" stateProvince="Zacatecas">
<collectingRegion id="0D1F2D9E0044FFC1FAC72A8C15A037E2" country="Mexico" name="Zacatecas" pageId="14" pageNumber="473">Zacatecas</collectingRegion>
,
<collectingMunicipality id="2F0079060044FFC1FF452AAF14F737E2" box="[202,490,1975,2002]" pageId="14" pageNumber="473">Teul de Gonzalez Ortega</collectingMunicipality>
,
<location id="CA04B5A70044FFC1FE782AAF174837E1" LSID="urn:lsid:plazi:treatment:4772526A0044FFC0FF1828DC108032FC:CA04B5A70044FFC1FE782AAF174837E1" box="[503,597,1975,2001]" country="Mexico" latitude="21.612806" longLatPrecision="1" longitude="-103.60616" municipality="Teul de Gonzalez Ortega" name="Huitzila" pageId="14" pageNumber="473" stateProvince="Zacatecas">Huitzila</location>
; borde arroyo, elev.
<quantity id="08234E990044FFC1FCCF2AA0168A37E2" box="[832,919,1976,2002]" metricMagnitude="3" metricUnit="m" metricValue="1.718" pageId="14" pageNumber="473" unit="m" value="1718.0">
<elevation id="44F6044F0044FFC1FCCF2AA0168A37E2" box="[832,919,1976,2002]" metricMagnitude="3" metricUnit="m" metricValue="1.718" pageId="14" pageNumber="473" unit="m" value="1718.0">1718 m</elevation>
</quantity>
(
<geoCoordinate id="AAEF85BB0044FFC1FC262AAF115337E1" box="[937,1102,1975,2001]" degrees="21" direction="north" minutes="36" orientation="latitude" pageId="14" pageNumber="473" precision="1" seconds="46.1" value="21.612806">21°36’46.1’’N</geoCoordinate>
,
<geoCoordinate id="AAEF85BB0044FFC1FBD32AAF100B37E2" box="[1116,1302,1975,2002]" degrees="103" direction="west" minutes="36" orientation="longitude" pageId="14" pageNumber="473" precision="1" seconds="22.2" value="-103.60616">103°36’22.2’’W</geoCoordinate>
),
<date id="BB65C5BC0044FFC1FAA32AAF108537E1" box="[1324,1432,1975,2001]" pageId="14" pageNumber="473" value="2004-01-31">
<collectingDate id="AB213C540044FFC1FAA32AAF108537E1" box="[1324,1432,1975,2001]" pageId="14" pageNumber="473" value="2004-01-31">31.i.2004</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0044FFC1FF482AC4143237C6" box="[199,303,2012,2038]" pageId="14" pageNumber="473">V.Gomez</collectorName>
&amp;
<collectorName id="622E86AA0044FFC1FEDC2AC4174937C5" box="[339,596,2011,2038]" pageId="14" pageNumber="473">J.L. Navarrete-Heredia</collectorName>
(1
<collectionCode id="A9CA7BB90044FFC1FDF52AC417C837C6" box="[634,725,2012,2038]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="473" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F70045FFC0FF482D8E15E232B5" pageId="15" pageNumber="474" type="diagnosis">
<paragraph id="CF64E37C0045FFC0FF482D8E1657312C" blockId="15.[151,1437,150,717]" pageId="15" pageNumber="474">
<emphasis id="FDAF3F6E0045FFC0FF482D8E14D03080" bold="true" box="[199,461,150,176]" pageId="15" pageNumber="474">Differential diagnosis.</emphasis>
Among Central American species, recognized by the combination of moderate size (ca.
<quantity id="08234E990045FFC0FF182DA2141D30E5" box="[151,256,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.94" pageId="15" pageNumber="474" unit="mm" value="2.94">2.94 mm</quantity>
), the deep pronotal depressions, and the male genitalia (
<figureCitation id="57E0FFF90045FFC0FCF82DA216A130E4" box="[887,956,186,212]" captionStart="FIGURES 9–10" captionStartId="26.[151,264,1944,1968]" captionTargetBox="[163,1430,185,1901]" captionTargetId="figure-22@26.[151,1435,181,1920]" captionTargetPageId="26" captionText="FIGURES 9–10. Habitus and male genitalia. 9. Hydrochus brianbrowni Makhan. 10. Hydrochus nayaritensis n. sp. holotype." figureDoi="http://doi.org/10.5281/zenodo.4777861" httpUri="https://zenodo.org/record/4777861/files/figure.png" pageId="15" pageNumber="474">Fig. 9</figureCitation>
; described below). This species has a generalized
<taxonomicName id="08DB98FF0045FFC0FF782DC6146C30C8" authorityName="Leach" authorityYear="1817" box="[247,369,222,248]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="FDAF3F6E0045FFC0FF782DC6146C30C8" box="[247,369,222,248]" italics="true" pageId="15" pageNumber="474">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males. The aedeagus is very narrow, for comparison refer to the diagnosis of
<taxonomicName id="08DB98FF0045FFC0FDD62C1B1618312C" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[601,773,259,284]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="474" phylum="Arthropoda" rank="species" species="nayaritensis" status="sp. nov.">
<emphasis id="FDAF3F6E0045FFC0FDD62C1B1618312C" box="[601,773,259,284]" italics="true" pageId="15" pageNumber="474">H. nayaritensis</emphasis>
</taxonomicName>
<emphasis id="FDAF3F6E0045FFC0FC842C1A1657312C" bold="true" box="[779,842,258,284]" pageId="15" pageNumber="474">
<taxonomicNameLabel id="E69C82150045FFC0FC842C1A1657312C" box="[779,842,258,284]" pageId="15" pageNumber="474" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="CF64E37C0045FFC0FF482C3E11D231C4" blockId="15.[151,1437,150,717]" pageId="15" pageNumber="474">
Male genitalia (
<figureCitation id="57E0FFF90045FFC0FEF72C3E14A13170" box="[376,444,294,320]" captionStart="FIGURES 9–10" captionStartId="26.[151,264,1944,1968]" captionTargetBox="[163,1430,185,1901]" captionTargetId="figure-22@26.[151,1435,181,1920]" captionTargetPageId="26" captionText="FIGURES 9–10. Habitus and male genitalia. 9. Hydrochus brianbrowni Makhan. 10. Hydrochus nayaritensis n. sp. holotype." figureDoi="http://doi.org/10.5281/zenodo.4777861" httpUri="https://zenodo.org/record/4777861/files/figure.png" pageId="15" pageNumber="474">Fig. 9</figureCitation>
). General characters: paramere tips slender and pointed, narrower in lateral view than in dorsal/ventral views, in dorsal/ventral views with medial margins at apex parallel or nearly so, then arcuate away from midline, slightly narrowed proximal to plsa; aedeagus in dorsal/ventral views very slender, parallel-sided or nearly so for entire length, apex pointed; in lateral view wider at base and slightly wider for remainder of length; lateral margins of aedeagus sclerotized; basal piece in dorsal/ventral views tapering from distal to proximal, in lateral view slightly wider proximally than distally, shorter than parameres, orifice strongly sclerotized.
</paragraph>
<paragraph id="CF64E37C0045FFC0FF482CE6105B320C" blockId="15.[151,1437,150,717]" pageId="15" pageNumber="474">Dorsal surface: pdmm widely separated, overlapping aedeagus only very slightly at base; adbl moderately large; adtl at about distal 1/6; each paramere with prominent pwsa; paramere tips without sclerotized ridge.</paragraph>
<paragraph id="CF64E37C0045FFC0FF482F5E15E232B5" blockId="15.[151,1437,150,717]" pageId="15" pageNumber="474">Ventral surface: pvmm well separated, except narrowly separated where overlapping aedeagus basally; avtl not apparent.</paragraph>
</subSubSection>
<subSubSection id="87C1B0F70045FFC0FF482F96108032FC" pageId="15" pageNumber="474" type="discussion">
<paragraph id="CF64E37C0045FFC0FF482F96108032FC" blockId="15.[151,1437,150,717]" pageId="15" pageNumber="474">
<emphasis id="FDAF3F6E0045FFC0FF482F9614263298" bold="true" box="[199,315,654,680]" pageId="15" pageNumber="474">Remarks.</emphasis>
I have examined the
<typeStatus id="10605DDE0045FFC0FDA02F9617893298" box="[559,660,654,680]" pageId="15" pageNumber="474" type="holotype">holotype</typeStatus>
, including the male genitalia (LANHM). The specimen imaged (
<figureCitation id="57E0FFF90045FFC0FAE02F9615BB32FC" captionStart="FIGURES 9–10" captionStartId="26.[151,264,1944,1968]" captionTargetBox="[163,1430,185,1901]" captionTargetId="figure-22@26.[151,1435,181,1920]" captionTargetPageId="26" captionText="FIGURES 9–10. Habitus and male genitalia. 9. Hydrochus brianbrowni Makhan. 10. Hydrochus nayaritensis n. sp. holotype." figureDoi="http://doi.org/10.5281/zenodo.4777861" httpUri="https://zenodo.org/record/4777861/files/figure.png" pageId="15" pageNumber="474">Fig. 9</figureCitation>
) was collected at this locality:
<collectingRegion id="0D1F2D9E0045FFC0FD8D2FAA17C132FC" box="[514,732,690,716]" country="Mexico" name="Baja California Sur" pageId="15" pageNumber="474">Baja California Sur</collectingRegion>
,
<quantity id="08234E990045FFC0FD672FAA163F32FC" box="[744,802,690,716]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="15" pageNumber="474" unit="mi" value="9.0">9 mi.</quantity>
up Canyon San Bernardo fr. Boca de la Sierra (Potrero).
</paragraph>
</subSubSection>
</treatment>
</document>
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